Bollettino della Società Paleontologica Italiana, 47 (1), 2008, 51-69. Modena, 30 aprile 200851

Parkinsonia parkinsoni Zone (upper Bajocian, Middle Jurassic) ammonites from Monte Kumeta (Western Sicily)

András GALÁCZ

A. Galácz, Department of Paleontology, Eötvös L. University, H-1117 Budapest, Pázmány Péter sétány 1/c, Hungary; [email protected]

KEY WORDS - Western Sicily, Upper Bajocian, Ammonites, Stratigraphy, Systematic descriptions.

ABSTRACT - Middle Jurassic ammonites are yielded richly by the lower Rosso Ammonitico above the massive carbonates and below the upper Rosso Ammonitico in several places in Western Sicily. One of the best localities is near the top of Monte Kumeta, near Piana degli Albanesi. Here several levels with distinct fossil assemblages have been investigated and collected in the past years. A prominent level is with upper Bajocian, Parkinsonia parkinsoni Zone ammonites, which was separated from another level with Garantiana garantiana Zone ammonites below, and Bathonian ones above. The level gave a comparatively rich assemblage with seemingly reworked specimens. The ammonites, of which several species are documented from Sicily first time now, are described systematically. Their stratigraphic evaluation suggested a narrow, probably uppermost Parkinsoni Zone, i.e. uppermost Bajocian age, and their paleogeographic affinity pointed to connections with the Southern Alps and Bakony Mts., Hungary, i.e. areas representing the true pelagic regions of the Jurassic Mediterranean Tethys.

RIASSUNTO - [Ammoniti della Zona a Parkinsonia parkinsoni (Bajociano superiore, Giurassico Medio) nel Rosso Ammonitico di Monte Kumeta, Sicilia Occidentale] - In diverse località della Sicilia occidentale i terreni mediogiurassici del Rosso Ammonitico inferiore, interposti fra gli spessi carbonati di piattaforma e i depositi del Rosso Ammonitico superiore, contengono ricche associazioni ad ammoniti. Una delle località dove questi depositi sono ben esposti è Monte Kumeta, vicino a Piana degli Albanesi. In questa località sono stati studiati alcuni livelli, ciascuno caratterizzato da una diversa associazione ad ammoniti. Un livello significativo contiene ammoniti della Zona a Parkinsonia parkinsoni, riferibile al Bajociano superiore. Esso sormonta un livello con ammoniti della Zona a Garantiana garantiana, mentre al di sopra è presente un livello con ammoniti batoniane. Il livello riferibile alla zona a Parkinsonia parkinsoni ha fornito una ricca associazione contenente anche esemplari probabilmente rimaneggiati. Il rimaneggiamento ha prodotto l’accumulo di clasti di calcari fossiliferi e ammoniti riesumate e con incrostazioni ferromanganesifere, in una tasca prodottasi nei sottostanti calcari a crinoidi, foderata da uno spesso hardground. L’associazione ad ammoniti è dominata da Phylloceratidi. Le Ammonitinae presenti sono rappresentate da Parkinsonidi, Lobosphinctes macro e microconchi, Strigoceras, Phlycticeras, Dimorphinites che sono qui documentate e dettagliatamente descritte per la prima volta in Sicilia. L’intervallo stratigrafico indicato dall’associazione ad ammoniti è ristretto alla parte superiore della Zona a Parkinsonia parkinsoni corrispondente alla parte sommitale del Bajociano. Dal punto di vista paleogeografico l’associazione rinvenuta sembra differire dalle coeve associazioni della vicina Africa settentrionale, mentre presenta affinità con quelle delle Alpi Meridionali e dei Monti Bakony (Ungheria) che rappresentano le vere regioni pelagiche della Tetide Giurassica.

INTRODUCTION western Sicily, thus a new record deserves a detailed description. The present account is one of a planned Western Sicily, as a classic area of Mediterranean series of papers describing further Middle Jurassic Jurassic fossils, is best known for Aalenian, Bathonian- ammonite assemblages from this same locality. Callovian and Upper Jurassic ammonites. Localities The here described specimens were collected in the yielding Bajocian ammonites are rare, and even some of Cerniglia Quarry and in the so-called Upper Quarry in those recorded earlier (e.g. Favara, near Agrigento; the period 1983-2002. The specimens are kept in the Gemmellaro, 1877) are now inaccessible (see Wendt, collection of the Natural History Museum (TTKM) of 1964; Pavia & Cresta, 2002). Wendt (1971) described the Eötvös L. University, Budapest. two upper Bajocian assemblages from the fissure fillings of Rocca Busambra, one (Fauna 9) from the Niortense and one (Fauna 10) from the Garantiana Zone. Other UPPER BAJOCIAN AMMONITES AT MONTE localities of upper Bajocian ammonites are Cozzo di KUMETA Cugno and Bellolampo, both near Palermo in the Panormide Domain (Vörös et al., 1987). Recently a The first to mention Bajocian ammonites from Monte limestone lens yielding upper Bajocian (and lower Kumeta (Fig. 1) was Wendt (1964, pp. 84-85), who listed Bathonian) ammonites within the main hardground in the fossils from the Concavum to Humphriesianum Zones Rocca chi Parra succession was studied (Pavia et al., (i.e. upper Aalenian to lower Bajocian) interval. In his 2002; Martire & Pavia, 2004). subsequent work (Wendt, 1965, p. 290) he gave a more The here described occurrence from Monte Kumeta extended ammonite list, in which indications of further (Fig. 1) represents a narrower interval, because only upper levels: the Bajocian Sauzei Zone, middle Bathonian and Parkinsoni Zone ammonites appear. Nevertheless, the lower Callovian also appear. Later (Wendt, 1969, fig. 3) Bajocian is an interval of which palaeontological he indicated “cephalopod limestone (massive, bedded, documentation, as compared to that of the higher Middle nodular limestone)” also from the whole upper Bajocian. Jurassic or Upper Jurassic, is rather incomplete in Jenkyns & Torrens (1971) dated the first level in the

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also to study the source fossiliferous level in situ in the quarries. Putting the history of collecting ammonites on Monte Kumeta into a perspective, a pattern is shown where older collections indicated more and older dates, including ones never found again, while recent studies gave fewer and stratigraphically younger horizons. This suggests that, along with the northward thinning of the Rosso Ammonitico limestone, the older fossiliferous levels disappear as the condensation horizons wedge out in this direction, and the quarrying reveal nowadays the uppermost, prevailing levels only (Fig. 3).

Fig. 1 - The locality Monte Kumeta in Western Sicily STRATIGRAPHY

The general stratigraphy of the Monte Kumeta Jurassic was recently treated in detail (Di Stefano et al., 2002a, b; Chiari et al., 2004). The here described ammonites came Rosso Ammonitico limestone above the Toarcian from the lowermost prevailing fossiliferous level of the hardground as Humphriesianum Zone on Monte Kumeta. Rosso Ammonitico limestone covering the main Our collecting of Jurassic, mainly Toarcian and hardground. The ammonites were collected in two Middle Jurassic, ammonites on Monte Kumeta has been localities: in “Cava Cerniglia” - the main working quarry started in the early 1980’s and went on more or less on the hill - and in the “Upper Quarry”, an old, abandoned regularly since then. Firstly a short note on the lower quarry about 150 m above, on the faulted southern slope Bathonian was published (Galácz, 1985), then data on the of the Kumeta top. general ammonite stratigraphy were included in The position of the uppermost Bajocian fossil beds is publications about the locality (Di Stefano et al., 2001, slightly different in the two places. While in “Cava 2002a, b). These data indicated that the quarries near the Cerniglia” there is a fossil horizon today with at least top of the hill expose roughly the same sequence: massive one (Garantiana Zone) assemblage between the Toarcian shallow-water carbonates at the base, a main main hardground and the uppermost Bajocian level, in the ferromanganese hardground with mainly Toarcian “Upper Quarry” the Parkinsoni Zone ammonites, which ammonites in the thick, black ferromanganese crust, and are embedded in a ferro-manganiferous crust, follow red, nodular limestone with several fossiliferous levels, immediately, almost inseparably the ferromanganese then radiolarite, which is followed by Upper Jurassic blanket with Toarcian ammonites. In both places the limestones in the higher cover. The ammonite-rich levels Bajocian fossils came well below levels with Bathonian, of the red nodular limestone (Rosso Ammonitico then Callovian ammonites within the Rosso Ammonitico inferiore) are well visible in the walls exposed by the followed by the radiolarite with a basal belemnite bed wire-cut technology used in the quarries, and could be dated as late Callovian-middle Oxfordian (Mariotti, 2002; studied on the detachment surfaces of the sawn blocks Chiari et al., 2004). lying in and around the quarries. Specimens clearly show reworking. Many are In the main quarries, at least four Middle Jurassic embedded in the reddish limestone or in the fossiliferous levels were identified, and there are ferromanganese crust as fragments, and the infilling of indications for some more. Immediately above the thick the shells is similar to the calcareous matrix. Actually, manganese crust covering the jagged, irregular surface the reworking was seemingly limited to the very early of the Carixian crinoidal limestone, the first level with burial phase, without “taphonomic” condensation. The ammonites in the Rosso Ammonitico gave different fossil specimens show single, one-phase breakage. This latter assemblages (Fig. 2). Most common is the upper means that seemingly only one part of the previously Bajocian, but remnants of older fossiliferous levels were infilled and lithified specimens is usually missing. There also identified. In 1983 we found a single, big specimen is neither sign of breaking off the body-chambers, nor of Skirroceras sp., which suggests the Sauzei or tendency of loss or worn of prominent sculptural Humphriesianum Zone, and is in accordance with earlier elements as sharp ribs or pointed tubercles. records (see above) of these levels from Monte Kumeta. As reconstructing the process immediately before the Similarly, a single specimen of Strenoceras sp. found final burial, it is suggested an elevated submarine place in 1990 in the Cerniglia Quarry suggests the presence of nearby, where the just lithified calcareous sediments, with the Niortense Zone. The Garantiana Zone was indicated the embedded biogenic elements, were removed from, by a smaller ammonite assemblage with Ermoceras and redeposited in the sediment-poor ferromanganese- (Galácz, 1999a). precipitation site. The history of the ammonites is most In recent years one of the levels most commonly similar to that of the broken-off and redeposited pillars occurring on the surfaces of the cut blocks is that with from the jagged Liassic surface which are visible in the Parkinsoni Zone ammonites. The here described so-called “Cimitero dei pinnacoli” in another, nearby ammonites came mostly from these blocks, but temporal exposure (Di Stefano et al., 2002a, b). The time of the expositions during the last decades gave opportunities redeposition was probably very close to the age of the

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assemblage, i.e. possibly latest Bajocian, because there Holcophylloceras zignodianum (d’Orbigny, 1848) are no Bathonian elements in the collected material. The Holcophylloceras privasense Joly, 2000 very similar process of resedimentation of Middle Lytoceras eudesianum adeloides (Kudernatsch, 1852) Jurassic ammonites just above the Toarcian pinnacled Nannolytoceras tripartitiforme (Gemmellaro, 1877) surface in the Rocca chi Parra quarry (Martire & Pavia, Strigoceras truellei (d’Orbigny, 1845) 2004) has been taken place a little later, because there Phlycticeras buckmani (Brasil, 1894) lower Bathonian ammonites are mixed together with Oppelia plicatella Gemmellaro, 1877 upper Bajocian ones. Cadomites (Cadomites) daubenyi (Gemmellaro, 1877) A significant part of the collected ammonites is made Cadomites (Polyplectites) sp. indet. up by fragmented, indeterminable specimens - mostly Parkinsonia (Durotrigensia) ferruginea Nicolesco, Phylloceratids. The determined ammonite assemblage, 1928 of which important taxa are described below, is as Dimorphinites dimorphus (d’Orbigny, 1846) follows: Vermisphinctes (Vermisphinctes) vermiformis Buckman, 1920 Phylloceras kudernatschi (Hauer, 1854) Vermisphinctes (Vermisphinctes) martiusi (d’Orbigny, Calliphylloceras disputabile (Zittel, 1868) 1845)

Fig. 2 - Middle Jurassic fossil assemblages in the quarries near the top of Monte Kumeta. Enlarged is the upper Bajocian fossiliferous level above the ferromanganese main hardground. The layers with the Parkinsoni Zone assemblage is shaded grey.

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Fig. 3 - Reconstruction of development of quarrying and exposing the Middle Jurassic rocks with wedging out of fossiliferous horizons in the Monte Kumeta quarries. 1 = Pliensbachian crinoidal limestone; 2 = the main hardground with mostly Toarcian ammonites; 3 = lower Rosso Ammonitico limestone with fossiliferous horizons; 4 = Belemnite marl; 5 = radiolarite.

Planisphinctes (Lobosphinctes) intersertus Buckman, western Mediterranean region of the Tethys. Within this 1923 region, wherever Parkinsoni Zone assemblages appear, Planisphinctes (Planisphinctes) tenuissimus the taxonomic composition is similar: Nannolytoceras, (Siemiradzki, 1898) Oppelia, Cadomites, and Dimorphinites are the common associates to the Phylloceratids, and they occur together This is a Parkinsoni Zone assemblage, indicating most with Strigoceras and early Perisphinctinae (Vermi- probably the upper, Parkinsonia bomfordi Subzone. This sphinctes, Lobosphinctes), while Parkinsoniids appear age assignment is suggested by the presence of usually very rare. However, a more detailed analysis may Planisphinctes (M+m) species, with Dimorphinites indicate closer palaeogeographic affinities. dimorphus, which latter seems to be a good indication The Kumeta upper Bajocian assemblage is most for the uppermost Bajocian in the Mediterranean region similar to those described from the Venetian Prealps by (see Sturani, 1964; Galácz 1980; Sandoval, 1983, 1990). Sturani (1964), from the Lessini Veronesi by Clari et al. Other fossils are rather rare. As additional elements, (1984), and from the Bakony Mountains, Hungary a single nautiloid (Paracenoceras sp.) and some (Galácz, 1980). In these places the represented genera brachiopods were collected. The brachiopods, which were are those mentioned above, with Parkinsoniids as very determined by A. Vörös, are as follows: Apringia cf. atla rare elements. (Oppel), A. cf. alontina (Di Stefano), and Karadagithyris On the other hand, in the geographically nearer Atlas gerda (Oppel), indicating also the upper Bajocian. range (in the Folded Middle Atlas and in the High Atlas) This fossil assemblage is very similar to the Bajocian Parkinsoni Zone assemblages are dominated by components of the mixed upper Bajocian-lower Thambites and Oppelia (du Dresnay, 1967). Later studies Bathonian assemblage discovered in the nearby “Rocca (Enay et al., 1987) recorded the presence of Dimor- chi Parra” quarry (Martire et al., 2000; Martire & Pavia, phinites dimorphus and associated Parkinsoniids below 2002, 2004). That ammonite assemblage includes a very rich Zigzag Zone assemblage in the same area. A Lobosphinctes, Planisphinctes, Dimorphinites, Cadomites species, close to C. daubenyi (Gemmellaro) Parkinsonia spp., similarly indicating the top of the is also present here, just as in Algeria (loc. cit.). Parkinsoni Zone. Dimorphinites dimorphus was recorded also from Tunisia (Arkell, 1956, p. 278). However, Cadomites and Dimorphinites seem to be rare elements in the PALEOBIOGEOGRAPHY Maghrebian Bajocian. From northern Tunisia a similar Parkinsoni Zone assemblage was recorded (Soussi et al., Middle Jurassic ammonite assemblages with high 2000) with Cadomites, Prorsisphinctes and representation of Phylloceratids are characteristic in the Dimorphinites.

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The benthic elements, i.e. brachiopods show Distribution - P. kudernatschi is a characteristic, considerable differences between northwest Africa and common ammonite in the Mediterranean upper Bajocian Sicily. Vörös (1984) demonstrated that the Bajocian to Callovian. The species was recorded from several brachiopods of Morocco and the Callovian brachiopods localities and levels from Western Sicily. of Tunisia and Libya have clear NW European affinities in contrast to the strictly Mediterranean brachiopods of western Sicily. The same pattern is shown by the Middle Calliphylloceras disputabile (Zittel, 1868) Jurassic gastropod distribution in the western Tethys. As Pl.1, figs. 1, 3 detailed studies indicated (Conti & Monari, 2001), the gastropod assemblages from the Atlas range are close to 1868 Phylloceras disputabile Zitt. - ZITTEL, p. 606. the assemblages in northwest European epicontinental 1980 Calliphylloceras disputabile (Zittel, 1868) - GALÁCZ, p. 37, areas, while significantly different from those in the Pl. 5, figs. 3, 6, text-figs. 25-29 (cum syn.) 1999 Calliphylloceras disputabile (Zittel, 1868) - GALÁCZ, 1999b, “Intratethyan Region”, including western Sicily. These p. 152, Pl. 1, fig. 1. suggest a wide pelagic (oceanic?) water mass as barrier 2002 Calliphylloceras disputabile (Zittel, 1868) - D’ARPA in PAVIA between Middle Jurassic Sicily and the North African & CRESTA, p. 48. Gondwanian shelf. However, the nektonic ammonites were probably less limited in their distribution by this Material - Several, mainly fragmentary specimens. pelagic barrier. This is how elements (Parkinsoniids, Most are small or medium-size fragments, the biggest Morphoceratids), having been migrated from NW Europe one is a partially preserved body-chamber of a ca. 180 along the NW African shelf could arrive to western Sicily, mm diameter specimen. while sporadically some Arabian forms moving in opposite direction on the same shelf migration route Description - On the middle and outer whorls it is a could reach this region (Galácz, 1999a). narrowly-umbilicated species with well-rounded umbilical side and shoulder, convex flank and highly- arched venter, with a general ‘fat’ aspect in cross-section. SYSTEMATIC DESCRIPTIONS While the internal mould is smooth, the shell bears fine, forwardly curved striation. The whorls exhibit 6 to 8 In the synonymies references are made to the first constrictions on the internal mould and corresponding descriptions and figures of the species, to relevant recent rounded, radial swellings on the shell. The constrictions descriptions in monographs and previous mentions of the start prorsiradially at the umbilical margin, then curve species from Sicily. Detailed synonymies are presented forward ca. at the middle of the flank. only in cases of species without recent detailed discussions. Remarks - Joly (2000, p. 79) included Ammonites heterophylloides Oppel, 1856 into C. disputabile on Phylloceras kudernatschi (Hauer, 1854) subspecific level. However, the two forms, as it was Pl. 1, fig. 2 clearly indicated by Neumayr (1871, p. 33l) differ in features enough to distinguish them on species level 1852 Ammonites heterophyllus Sow. var. - KUDERNATSCH, p. 6, Pl. within the generally character-poor group of 1, figs. 5-9. Phylloceratids. 1854 Ammonites Kudernatschi v. Hauer - HAUER, p. 902. 1980 Phylloceras kudernatschi (Hauer, 1854) - GALÁCZ, p. 31, Pl. 3, figs. 1-2, text-figs. 17-20 (cum syn.). Distribution - C. disputabile, this common Middle 2000 Phylloceras kudernatschi (Hauer, 1852) - JOLY, p. 45, Pl. 7, Jurassic Tethyan ammonite was repeatedly recorded from figs. 2-4, text-figs. 82-85. Sicily. The earlier Sicilian records were recently revised 2002 Phylloceras (Phylloceras) kudernatschi (Hauer, 1852) - by D’Arpa in Pavia & Cresta (2002, p. 48) and new ones D’ARPA in PAVIA & CRESTA, p. 33. are those by Galácz, 1999b, and Martire et al., 2000. Material - Three fragmentary specimens, internal and middle whorls, all are phragmocones, the biggest Holcophylloceras zignodianum (d’Orbigny, 1848) one is of 65 mm maximum preserved diameter. Pl. 1, figs. 4, 5; Pl. 2, fig. 1

Description - This is a narrowly-umbilicated 1848 Ammonites Zignodianus d’Orb. - D’ORBIGNY 1842-49, p. 493, Phylloceratid with high-oval whorl section. The funnel- Pl. 182, figs. 1-3. shaped umbilical area is followed by convex sides and 1871 Phylloceras mediterraneum sp. nov. - NEUMAYR, p. 340, Pl. highly arched venter. The sculpture is a dense ribbing that 17, figs. 2-5. appears only on the shell, the internal mould being almost 1980 Holcophylloceras zignodianum (d’Orbigny, 1848) - GALÁCZ, p. 41, Pl. 5, figs. 4-5, Pl. 6, fig. 1, Pl. 7, fig. 1, text-figs. 30- smooth. The narrow ribs run radially on the inner part of 32 (cum syn.). the flank, then curve forward slightly. While nearly equal 2002 Holcophylloceras zignodianum (d’Orbigny, 1848) - PAVIA in in strength, the ribs form folds on the outer third of the PAVIA & CRESTA, p. 61, fig. 240-b. flanks, which appear as radial undulations at every third or fourth rib. Material - Four fragmentary specimens, all are The visible parts of the suture-line show the deep, incomplete phragmocones; the biggest one is of ca. 70 tapering first lateral lobe and the high, diphyllic lobes. mm diameter.

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Description - A relatively narrow-umbilicated the hitherto known H. privasense specimens do not Phylloceratid with compressed, high-oval whorl-section. show the short ventral ribs between the constrictions The low, vertical umbilical wall and the gently convex so common in H. zignodianum. flank form a rounded, but distinct margin, while the lateral side rounds gradually into the highly-arched venter. The Distribution - Joly (2000, p. 103) selected the type distinguishing feature is the presence of the characteristic form the lower Bathonian material of Privas (Ardèche), constrictions (6-7 per whorls): arising at the umbilical and he mentioned Bajocian (S. humphriesianum Zone) margin, they curve forward, then form a sharp break just representatives also. above the middle of the flank, and run out to the venter with another falcoid curve. At the sharp break there is a lip-like forward extension. The constrictions are Lytoceras eudesianum adeloides (Kudernatsch, 1852) especially strong and deep on the internal mould. The Pl. 2, figs. 13-14 bigger specimens bear short simple outer ribs arising at the ventrolateral shoulder and continuing out onto the 1852 Ammonites Adeloides Kd. - KUDERNATSCH, p. 9, Pl. 2, figs. venter. These ribs run parallel to the outer portion of the 14-16. curved constriction. 1964 Lytoceras adeloides (Kudernatsch, 1852) - PUGIN, p. 20, text- fig. 4 (cum syn.). The suture-line shows the characteristic diphyllic and triphyllic saddles. Material - A single, partially preserved specimen.

Remarks - Phylloceras mediterraneum of Neumayr Description - A loosely coiled ammonite with (1871) is a junior synonym, but some authors regard it circular cross-section. The sculpture is formed by fine, as a separate species (see e.g. Joly, 2000). Even this radial, rounded riblets with periodic sharp collars bearing limited material shows well that the most variable crincles on the upper part of the flanks and on the venter. character of the species is the incision of the ventral ribs: The specimen is wholly septate showing some portions there are specimens (e.g. on Pl. 1, figs. 4, 5) showing of the characteristic Lytoceratid suture-line. smooth outer parts at 45 mm diameter, while others (e.g. on Pl. 2, fig. 1) have well-developed short ribs on the Remarks - Though not very well preserved, the ventrolateral shoulder and on the venter at approximately species shows the minute sculptural elements on the the same size. recrystallized shell, and can be identified at subspecific level. Distribution - As usual in Mediterranean Middle H. zignodianum Jurassic assemblages, is one of the most Distribution - This form, being a common element common elements in the Kumeta assemblage. Other in Mediterranean Bajocian and Bathonian assemblages, Sicilian occurrences were recorded earlier by several is commonly recorded from wide areas. However, those authors (e.g. Floridia, 1931; Wendt, 1964). Gemmellaro’s previously figured forms from Sicilian localities (see Sicilian records were recently revised by Pavia (in Pavia Gemmellaro, 1873, Pl. 5, figs. 4-5; Floridia, 1931, Pl. 3, & Cresta, 2002). fig. 4) could be better ranged into the subspecies eudesianum. Holcophylloceras privasense Joly, 2000 Pl. 2, fig. 2 Nannolytoceras tripartitiforme (Gemmellaro, 1877) Pl. 2, figs. 3-4 1935 Holcophylloceras mediterraneum Neumayr - ROMAN, p. 29, Pl. 2, fig. 4, Pl. 4, fig. 6. 2000 Holcophylloceras privasense nov. sp. - JOLY, p. 103, Pl. 26, 1877 Lytoceras tripartitiforme, Gemm. - GEMMELLARO, p. 61, Pl. figs. 1-3, text-figs. 209-211. 4, fig. 9. 2002 Nannolytoceras tripartitiforme (Gemmellaro, 1877) - PAV I A Material - Two incomplete, fragmentary specimens. in PAVIA & CRESTA, p. 103, figs. 57-58. Material - Numerous fragmentary specimens, mainly Description - The phragmocone fragments show partially preserved portions of whorls. slightly convex flanks and highly arched venter, thus the cross-section is oval. The single sculptural element is Description - The fragments indicate evolute coiling the periodically appearing constrictions, showing a of whorls with rounded, higher than wide, rectangular prorsiradiate inner and a gently forwardly-curved outer cross-section. The whorls are smooth, only constrictions portion, with a knee-like break above the middle of the appear. The forwardly arched constrictions are slightly flank. deeper on the umbilical corner, but remain distinct on Suture-line cannot be seen. the flanks and also on the venter. The suture-line shows the characteristic Remarks - This species differs from H. zignoidanum Nannolytoceras-features, just as those figured by Pavia in its constrictions (see Joly, 2000, p. 103), and in its (2002, fig. 58). whorl-section. This latter is oval-lenticular in H. privasense , while rather rectangular with well-defined Remarks - The here described incomplete material, ventrolateral shoulders in H. zignoidanum. Furthermore, while it points with all visible characters to Gemmellaro’s

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form, has made possible only to identify the species, in 1909-30, Pl. 585). The main difference between the further remarks cannot be made. typical and the latter forms is the narrower whorl-section, but this is a strongly variable character, not enough to Distribution - N. tripartitiforme has been recorded regard it as of specific value (see Sturani, 1964, p. 14). so far only from Sicily. The type material came from Favara, of which Middle Jurassic assemblage was Distribution - Strigoceras truellei is most common probably of late Bajocian age. in the upper part of the Parkinsoni Zone, but earlier occurrences (Niortense and Garantiana Zones) were also recorded (see Rioult et al. in Fischer, 1994, p. 103). The Strigoceras truellei (d’Orbigny, 1845) geographic distribution of the species is wide: there are Pl. 1, figs. 7-8 good records from the Caucasus (Rostovtsev, 1985) and Central Asia (Turkmenistan; Besnosov & Mitta, 1993, 1845 Ammonites Truellei d’Orbigny - D’ORBIGNY 1842-49, p. 2000) to all Western-Mediterranean and NW European 361, Pl. 117, figs. 1-3, Pl. 129, figs. 1-2. regions. 1980 Strigoceras truellei (d’Orbigny, 1845) - GALÁCZ, p. 54, Pl. 9, fig. 4 (cum syn.). 1983 Strigoceras (Strigoceras) truellei (d’Orbigny, 1845) - SANDOVAL, p. 98, Pl. 2, figs. 3-4, text-figs. 2-3. Phlycticeras buckmani (Brasil, 1894) pars 1985 Strigoceras truellei (Orbigny) - ROSTOVTSEV, p. 129, Pl. Pl. 1, fig. 6 34, fig. 9 (only). 1985 Strigoceras truellei (d’Orbigny) 1845 - FERNÁNDEZ LÓPEZ, 1894 Strigoceras Buckmani nov. sp. - BRASIL, p. 42, Pl. 4, figs. 4- p. 127, Pl. 12, fig. 6, text-figs. 11-12. 5. 1990 Strigoceras truellei (d’Orbigny) - SANDOVAL, p. 160, Pl. 1998 Phlycticeras buckmani (Brasil) [M] - SCHWEIGERT & DIETZE, 4, fig. 7. p. 7, Pl. 1, fig. 2. pars 1993 Strigoceras truellei (d’Orbigny), 1845 - BEZNOSOV & 2003 Phlycticeras buckmani (Brasil) - SCHWEIGERT & DIETZE, p. 5, MITTA, p. 23, Pl. 1, figs. 4-5 (only). Pl. 2, fig. 1. 1994 Strigoceras truellei (d’Orbigny, 1845) - RIOULT ET AL. in FISCHER, p .102, Pl. 50, figs. 1-2. Material - A single, partial specimen of which 1997 Strigoceras truellei (d’Orb.) - RIOULT ET AL., Pl. 15, fig. 7. preserved part indicates ca. 105 mm diameter, but the pars 2000 Strigoceras truellei (d’Orbigny, 1845) - BEZNOSOV & actual size of the specimen must have been far bigger, MITTA, p. 51, Pl. 1, figs. 1- 2 (only). because it is wholly septate. 2001 Strigoceras truellei (d’Orb. 1845) - ARP, p. 214, Pl. 6, fig. 4. Description - A relatively big form with narrow whorl- 2005 Strigoceras truellei (d’Orbigny, 1845) - SCHLÖGL ET AL., section formed by flattened, convergent flanks and lowly- p. 343, Pl. 3, fig. 4. arched venter at the given size of the phragmocone. The 2006 Strigoceras truellei (d’Orbigny) [M] - DIETZE & DIETL, p. 10, fig. 6. preservation cannot makes possible to discern the width and the features of the umbilicus, and also the sculpture Material - Two incomplete, fragmentary specimens. can be seen incompletely. There are low, rounded, prorsiradiate inner ribs and long, slightly forward curved Dimensions - The bigger specimen is of ca. 140 mm outer ribs, which latter terminate abruptly on the maximum preserved diameter and wholly septate, while ventrolateral margin. The ventral keel has been broken the smaller one, showing the middle whorls, is also septate off when the specimen was freed from the rock. up to its ca. 90 mm diameter. Entire suture-line cannot be seen, but the portions visible on different parts of the specimen indicate very Description - This is a big ammonite, with high, complicated sutures. lanceolate whorl-section, and relatively narrow umbilicus. The carinate venter is narrow, flattened on the Remarks - Schweigert & Dietze (1998, 2003) inner and becomes acute on the outer whorls. The middle recently suggested that the previously known, but never whorls (Pl. 1, fig. 8) bear short, regular outer ribs arising re-recorded species of Brasil should be ranged into at about the upper third of the flanks, then the outer ribs Phlycticeras. In their 2003 work they figured (Pl. 2, fig. stretch on the flanks down to ca. middle-height, and 1) a body-chamber fragment from the Inferior Oolite of become obsolete toward the end of the phragmocone. Dorset, which matches closely the here described Sicilian There are three wide, low, but well-distinguished spiral form. This latter, being a poorly-preserved internal mould, ridges on the lateral side: one on the umbilical margin, does not show the strigation, however, the other features one at the middle of the flank and one in between. At bigger (coiling, ribbing, etc.) are quite close to those of the size the spiral ridges smooth out, and the ribs as well, specimen from England. only faint, wedge-shaped outer ribs remain on the outer quarter of the whorl. The strigation is well visible on the Distribution - Brasil (1894, p. 43) indicated the upper specimens. Bajocian Oolithe ferrugineuse of Sully as the source of his specimen, and Schweigert & Dietze (1998, p. 7) gave Remarks - The morphological variability of S. truellei the closer Parkinsoni Zone as the type level. Their Dorset is wide. The here described specimens are close to the specimen (Schweigert & Dietze, 2003, p. 5) came from form what Quenstedt (1886-87) distinguished as var. the Truellei Bed (Parkinsoni Zone, Truellei horizon) of gracilis and elevated to species level by Buckman (1925 Burton Bradstock. The most probable level of the Kumeta

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upper Bajocian assemblage within the upper Bajocian Distribution - The type material is most probably is also topmost Parkinsoni Zone. late Bajocian in age, but indications of younger and older ages are also cited (see Pavia in Pavia & Cresta, 2002, p. 206). Oppelia plicatella Gemmellaro, 1877 Pl. 2, fig. 5; Pl. 3, fig. 3 Cadomites (Cadomites) daubenyi (Gemmellaro, 1877 Oppelia plicatella, Gemm. - GEMMELLARO, p. 62, Pl. 3, figs. 1877) 6-7 (only). Pl. 2, fig. 8; Pl. 4, fig. 3. 1980 Oxycerites plicatella (Gemmellaro) - GALÁCZ 1980, p. 62, Pl. 13, fig. 4, Pl. 14, figs. 1-2, text-figs. 50-51. pars1877 Stephanoceras Daubenyi Gemm. - GEMMELLARO, p. 67, 2002 Oppelia plicatella Gemmellaro - PAVIA in PAVIA & CRESTA, p. Pl. 4, figs. 4-5 (only). 203, figs. 134-137a, b. 2002 Cadomites daubenyi (Gemmellaro, 1877) - PAVIA in PAVIA & CRESTA, p. 236, figs. 161, 162, 163a, b (cum syn.). Material - Several fragmentary specimens, phragmocone and body-chamber portions alike. The Material - Several fragments, mainly inner and middle phragmocone fragments are of 40 to 70 mm diameter, phragmocone whorls, the biggest specimen is being 55 the biggest fragment is of a specimen of ca. 120 mm mm and still septate. diameter, with septation up to about 90 mm. Description - This is a medium-size Cadomites, with Description - Medium-size Oppeliid with narrow narrow umbilicus on the inner and middle whorls, slightly umbilicus, vertical but low umbilical wall, tightly rounded rounded umbilical wall and high, weakly angular venter. umbilical margin, slightly convex flanks and narrow, keeled The sculpture consists of dense, sharp ribbing with venter. The venter becomes rounded and unkeeled on the forwardly arched inner and radial outer ribs. At the body-chamber. The ribbing consists of short lunulate outer maximum width of the whorl small, pointed tubercles ribs on the outer third of the middle whorls. Between the form a row. lunulate ribs short, very distinct, regular, dense tertiary Suture-lines cannot be seen. riblets appear on the ventrolateral shoulder and a little below. On the last whorl these tertiary ribs disappear, only Remarks - This is a well-known species, of which type the outer lunulae remain, but they reach down almost to was refigured and synonymy discussed in detail recently the middle of the flank. On the middle whorls a shallow (Pavia in Pavia & Cresta 2002, pp. 236-239). The spiral groove runs on the flank at ca. 1/3 height, which is distinguishing specific characters listed by Pavia can be resulted by the elevated umbilical margin and the feeble all recognised in the Kumeta specimens: the deep, crater- spiral ridge at the middle of the flank. like umbilicus, the high, subangular venter, and the dense, Suture-lines cannot be seen. sharp ribs which are significantly weaker on the internal mould. Remarks - The species was just recently revised by Pavia (in Pavia & Cresta, 2002) on the basis of Distribution - Despite the several misinterpretations, Gemmellaro’s original material. The revision gave a good genuine C. daubenyi is a widespread species in the aspect of the species, which seems to be a common form Mediterranean region and beyond. All records, including in upper Bajocian and basal Bathonian assemblages. that of the type material, indicate late Bajocian, Parkinsoni Zone age.

EXPLANATION OF PLATE 1

figs. 1, 3 - Calliphylloceras disputabile (Zittel, 1868). 1 - Wholly septate specimen, TTKM A0750, lateral view. 3 - Wholly septate specimen, TTKM A0749, lateral view.

fig. 2 - Phylloceras kudernatschi (Hauer, 1854), fragmentary incomplete specimen, TTKM A0751, lateral view.

figs. 4-5 - Holcophylloceras zignodianum (d’Orbigny, 1848), wholly septate specimen, TTKM A0743. 4 - Lateral view. 5 - Ventral view.

fig. 6 - Phlycticeras buckmani (Brasil, 1894), wholly septate, fragmentary, incomplete specimen, TTKM A0769, lateral view.

figs. 7-8 - Strigoceras truellei (d’Orbigny, 1845). 7 - Wholly septate specimen, TTKM A0761, lateral view. 8 - Wholly septate, fragmentary specimen, TTKM A0762, lateral view.

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Cadomites (Polyplectites) sp. indet. 1931 Parkinsonia parkinsoni var. pseudoferruginea Nic. - SCHMIDTILL & KRUMBECK, p. 869, Pl. 83, fig. 7, Pl. 84, Material - A single, incomplete specimen. figs. 4, 6, Pl. 85, fig. 7. 1951 Parkinsonia pseudoferruginea Nicolesco - MAUBEUGE, p. 91, Pl. 8, fig. 4, Pl. 12, fig. 6. Description - The specimen is only a broken body- non 1972 Parkinsonia pseudoferruginea Nicolesco - AMANNIAZOV, chamber suggesting a microconch of 32-35 mm diameter p. 25, Pl. 5, fig. 1, Pl. 9, fig. 6, Pl. 10, fig. 6. adult size. The width/breadth ratio of the whorl near the 1973 Parkinsonia (Durotrigensia?) pseudoferruginea Nicolesco aperture is 16 mm/10 mm (=1.6) on a portion with highly - PAVIA, p. 123, Pl. 23, fig. 2, Pl. 24, fig. 1. arched venter, high, pointed tubercles and dense outer ribs. ? 1993 Gonolkites pseudoferrugineus (Nicolesco) - BESNOSOV & The poorly visible aperture has a narrow, smooth ventral MITTA, p. 206, Pl. 45, fig. 4. 1994 Parkinsonia pseudoferruginea Nicolesco - PAVIA, Pl. 2, collar which rounds into the elongated lateral lappets. fig. 2. ? 1998 Gonolkites pseudoferrugineus (Nicolesco, 1927) - Remarks - The specimen is very close to the BESNOSOV & MITTA, p. 29, Pl. 25, figs. 2-3. “Polyplectites? venetus” specimen described and figured by Wendt (1964, p. 132, Pl. 21, fig. 2) from the Bathonian Material - A single, incomplete fragment of a of Monte Inici. Sturani (1971, p. 136) excluded this form medium-size specimen. from the species designated as Stephanoceras venetum by Parona (1896), and the ammonite described by Description - The preserved parts represent a portion Gemmellaro as Stephanoceras? nov. sp. (1877, p. 147, of the phragmocone with about 1/4 part of the body- Pl. 19, fig. 2) is also different. A very similar specimen chamber. The umbilicus is moderately narrow, bordered is figured from the upper Bajocian (?Niortense Zone) of by widely-rounded umbilical wall, which arches into the Digne (Cadomites (Polyplectites) sp. ind. in Pavia 1973, convex flank. The venter is narrow and rounded. The Pl. 15, fig. 3). Without additional, better-preserved ribbing consists of low, slightly prorsiradiate inner ribs, specimens one cannot state anything on closer species which fade out just above the middle of the flanks, then affinity. bifurcate. There are additional intercalatory ribs also, so the ratio of the primary and secondary ribs is close to 1:3. The secondaries do not cross the venter: they end Parkinsonia (Durotrigensia) pseudoferruginea alternately at the middle. Nicolesco, 1928 Suture-lines cannot be seen. Pl. 2, figs. 6-7 Remarks - This is a commonly-recorded species, and 1886-87 Ammonites Parkinsoni - QUENSTEDT, Pl. 71, fig. 22 probably one of the few that would pass the test of a long (only). overdue revision of the upper Bajocian Parkinsoniids. 1927 Parkinsonia Parkinsoni Sow. var. pseudoferruginea n. var. - NICOLESCO 1927-28, p. 37, Pl. 9, Figs. 2-3, text-figs. 15-16. Distribution - All previous records refer to upper Bajocian, Parkinsoni Zone occurrences. These are mainly

EXPLANATION OF PLATE 2

fig. 1 - Holcophylloceras zignodianum (d’Orbigny, 1848), wholly septate, incomplete, fragmentary specimen, TTKM A0744, lateral view.

fig. 2 - Holcophylloceras privasense Joly, 2000, wholly septate, incomplete, fragmentary specimen, TTKM A0756, lateral view.

figs. 3-4 - Nannolytoceras tripartitiforme (Gemmellaro, 1877). 3 - Wholly septate middle whorls, TTKM A0764, lateral view. 4 - Fragment of a septate whorl, TTKM A0765, lateral view.

fig. 5 - Oppelia plicatella Gemmellaro, 1877, wholly septate, incomplete, fragmentary specimen, TTKM A0766, lateral view.

figs. 6-7 - Parkinsonia (Durotrigensia) pseudoferruginea Nicolesco, 1928, body-chamber fragment, TTKM A0772. 6 - Ventral view. 7 - Lateral view.

fig. 8 - Cadomites (Cadomites) daubenyi (Gemmellaro, 1877), wholly septate middle whorls fragment, TTKM A0757, lateral view.

figs. 9-12 - Dimorphinites (Dimorphinites) dimorphus (d’Orbigny, 1846). 9-10 - Wholly septate inner whorls, TTKM A0771. Ventral (9) and lateral view (10). 11-12 - Nearly complete specimen (asterisk indicates the end of phragmocone), TTKM A0770. Ventral (11) and lateral view (12).

figs. 13-14 - Lytoceras eudesianum adeloides (Kudernatsch, 1852), wholly septate specimen, TTKM A0768. 13 - Lateral view. 14 - Ventral view.

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from NW European localities, because Parkinsoniids are most common species of the group, in which recently a rare or rather subordinate in Mediterranean assemblages. small, new form was ranged as Dimorphinites centriglobus n. sp. by Rioult (in Fischer, 1994, p. 131, Pl. 50, fig. 8). This remarkable ammonite shows Dimorphinites (Dimorphinites) dimorphus similarities to ‘Morphoceras’ dimorphoides of Parona (d’Orbigny, 1846) (1896, Pl. 2, fig. 8) which was refigured later by Sturani Pl. 2, figs. 9-12 (1971, Pl. 14, fig. 19) as Dimorphinites? dimorphoides. While the form described by Rioult came from the pars1846 Ammonites dimorphus - D’ORBIGNY 1842-49, p. 410, Pl. Parkinsoni Zone, and Parona’s specimen is from a 141, figs. 1-2, 5-8 (only). material from the Niortense Zone, Sturani mentioned that 1980 Dimorphinites (Dimorphinites) dimorphus (d’Orbigny, this single specimen differs from the other Niortense 1846) - GALÁCZ, p. 100, Pl. 23, figs. 1-3, 5, text-figs. 81-83 (cum syn.). Zone ammonites from Monte Meletta by its infilling 1983 Dimorphinites (Dimorphinites) dimorphus (d’Orbigny, limestone. Thus it is not impossible that the two small 1845) - SANDOVAL, p. 330, Pl. 27, figs. 1-2, text-figs. examples represent the same tiny Dimorphinites species. 112A-C, 113A, 114. 1985 Dimorphinites dimorphus (Orbigny) - ROSTOVTSEV, p. 162, Distribution - Dimorphinites dimorphus is widely Pl. 44, figs. 7-10. distributed in Parkinsoni Zone assemblages from the 1990 Dimorphinites dimorphus (d’Orbigny) - SANDOVAL, p. 160, Pl. 4, fig. 5. Caucasus (Rostovtsev, 1985) to England (Buckman, 1923 1994 Dimorphinites dimorphus (d’Orbigny, 1846) - RIOULT in in 1909-30). The genus itself seems to be restricted to FISCHER, p.130, Pl. 50, fig. 7. the topmost Bajocian, however interesting ammonites, 1999 Dimorphinites (Dimorphinites) dimorphus (d’Orbigny, under the new generic name Pseudodimorphinites were 1846) - WIERZBOWSKI ET AL., p. 48, figs. 17/4-5, 18/1. documented from the Zigzag Zone (basal Bathonian) of 2005 Dimorphinites (Dimorphinites) dimorphus (d’Orbigny, Iran by Seyed-Emami et al. (1989). Despite the apparent 1846) - SCHLÖGL ET AL., p. 344, Pl. 13, fig. 2. similarities to Dimorphinites (mainly the lack of the ventral groove), they are closer to the group of Material - Two incomplete, fragmentary specimens. Asphinctites pinguis, where ribs without ventral The bigger one is of 53 mm diameter and incomplete, interruption also occur (see Galácz, 1999, p. 159). because the last portion of the body-chamber is missing. Recently in the quarry of the nearby “Rocca chi Parra” It is septate up to 40 mm diameter. The smaller specimen a rich Dimorphinites-assemblage, with D. dimorphus is a wholly-septate inner whorls. and other congeneric species (including microconchs) was discovered in a condensed topmost Bajocian-basal Description - This early Morphoceratid has Bathonian fossiliferous level (Martire et al., 2000; Pavia sphaerocone inner whorls with a nearly closed umbilicus, et al., 2002). rounded flanks and highly-arched venter. The coiling changes near the end of the phragmocone, with sudden widening of the umbilicus and contraction of the body- Vermisphinctes vermiformis Buckman, 1920 chamber, which results in an evolute coiling on the last Pl. 3, fig. 2 part of the conch. The whorls bear dense ribbing, with low, rounded primary and similar secondary ribs which 1920 Vermisphinctes vermiformis, nov. - BUCKMAN in 1909- arise as branching off or intercalating between the 30, Pl. 162. primaries. Costal interruption occurs on the venter, 1920 Vermisphinctes subdivisus, nov. - BUCKMAN in 1909- especially on the inner whorls. The most distinct feature 30, Pl. 190. is the appearance of constrictions from the innermost ? 1923 Perisphinctes Martiusi d’Orb. sp. - FALLOT & BLACHET, whorls. The constrictions are narrow and deep, and cross p.105, Pl. 1, fig. 10. the ribbing in an angle. ? 1973 Leptosphinctes (Vermisphinctes) cfr. vermiformis (S. Buckman) - PAVIA, p. 132, Pl. 27, fig. 2. Only details of the suture-line are visible. 1983 Vermisphinctes (Vermisphinctes) vermiformis Buckman, 1920 - SANDOVAL, p. 387, Pl. 33, figs. 2-4, Remarks - D. dimorphus is an easily-distinguishable text-figs. 117M, 118I. ammonite, of which microconch is also known (D. non1985 Leptosphinctes vermiformis (Buckman) - ROSTOVTSEV, (Vigoriceras) defrancii (d’Orbigny), see Galácz, 1980, p. 166, Pl. 45, figs. 5-6, Pl. 46, figs. 3-4, text-fig. 24. pp. 100-103; Rioult in Fischer, 1994, p. 115). This is the ? 2006 Vermisphinctes ex gr. vermiformis Buckman [?M] - DIETZE & DIETL, p. 8, Pl. 2, fig. 4.

EXPLANATION OF PLATE 3

fig. 1 - Planisphinctes (Lobosphinctes) intersertus Buckman, 1923, fragmentary specimen with preserved aperture, TTKM A0775, lateral view. fig. 2 - Vermisphinctes vermiformis Buckman, 1920, wholly septate middle whorls, TTKM A0773, lateral view. fig. 3 - Oppelia plicatella Gemmellaro, 1877, fragmentary, incomplete specimen (asterisk indicates the end of phragmocone), TTKM A0767, lateral view.

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Material - A single specimen with a well-preserved Vermisphinctes martiusi (d’Orbigny, 1845) side. This is a specimen of 93 mm preserved diameter, Pl. 4, fig. 4 but the traces of the next whorl’s umbilical seam indicates one more whorl, which was broken off. 1845 Ammonites Martiusi d’Orbigny - D’ORBIGNY 1842-1849, p. 381, Pl. 125, figs. 1-4. Description - A medium-size ammonite with dense, 1980 Vermisphinctes (Vermisphinctes) martinsi (d’Orbigny, 1845) - GALÁCZ, p. 107, Pl. 25, fig. 1 (cum syn.). regular ribbing. It is septate up to 60 mm diameter, there 1990 Vermisphinctes martiusi (d’Orbigny) - SANDOVAL, p. 160, Pl. is 3/4 whorl of preserved body-chamber, but the trace of 4, fig. 3. the umbilical seam of the next whorl indicates a whole 1994 Vermisphinctes martiusi (d’Orbigny, 1845) - RIOULT ET AL. in whorl of broken-off body-chamber parts. The coiling is FISCHER, p. 111, Pl. 49, figs. 1a-c. serpenticone, with wide, shallow umbilicus. The umbilical wall is low, the umbilical corner is rounded, the flanks Material - A single, incomplete specimen as part of a are flattened on the inner, and slightly convex on the outer big ammonite of ca. 115 mm preserved diameter, which whorls, the ventral side is highly arched. The ribbing was still incomplete, because it is septate nearly to the consists of long, slightly prorsiradiate, narrow ribs which end (up to ca. 110 mm). have a slight curvature on the umbilical margin. There are 57 primary ribs on the last, and 50 on the penultimate Description - A widely-umbilicated form with whorl- whorl. The primaries branch into 2 or 3 secondaries very section with low, rounded umbilical side, which becomes high on the flanks, thus they are not visible in the oblique on the last preserved whorl. The flanks are umbilicus. There are 3 deep, strongly prorsiradiate flattened on the inner, and become slightly convex on the constrictions on each whorl. They cut the ribbing in a middle and outer whorls. The venter is narrow and highly well-distinguished angle. rounded. The sculpture is formed by ribbing with sharp Suture-line cannot be seen. primary ribs on the inner and somewhat rounded inner ribs on the outer whorls. The secondaries branch high on Remarks - V. vermiformis and the slightly later the flanks. There are 2 to 3 secondary ribs by one primary. designated V. subdivisus of Buckman seem to be The ribs are straight radial on the inner whorls but become synonymous, because no significant differences are slightly forwardly-arched on the penultimate and last shown in the morphology. Buckman (1920, Pl. 190) was whorl. The constrictions (3 per whorls) appear regularly apparently uncertain about the level of the type of V. and follow the run of the primary ribs. subdivisus, indicating the Garantiana hemera, but he Suture-lines cannot be seen. later (1927, p. 33) listed it in the next (Vermisphinctes) hemera above, which belongs to the Parkinsoni Zone. Remarks - The lectotype (Rioult et al. in Fischer, 1994, Pl. 49, fig. 1), which was designated by Buckman Distribution - Genus Vermisphinctes is a common (1920, p. 26), is a macroconchiate specimen, septate up element in upper Bajocian assemblages in NW Europe to ca. 112 mm, and with stronger sculpture on the shell and the Mediterranean realm (see Galácz 1980, p. 105), than on the internal mould. The Kumeta specimen and the few well-localised occurrences of V. shows very good agreement, the only difference is the vermiformis indicate the Parkinsoni Zone. Sandoval slightly fewer ribs. (1983, p. 388) pointed out that the species occurs The difference between V. martiusi and the other together with Dimorphinites in the uppermost Bajocian congeneric forms (incl. V. vermiformis) is the bigger size of the Betic Cordilleras. and the body-chamber, which becomes a little larger,

EXPLANATION OF PLATE 4

figs. 1-2 - Planisphinctes (Lobosphinctes) intersertus Buckman, 1923, nearly complete specimen, TTKM A0774. 1 - Ventral view. 2 - Lateral view.

fig. 3 - Cadomites (Cadomites) daubenyi (Gemmellaro, 1877), wholly septate inner whorls, TTKM A0758, lateral view.

fig. 4 - Vermisphinctes martiusi (d’Orbigny, 1845), incomplete, fragmentary specimen, (asterisk indicates end of phragmocone), TTKM A0776, lateral view.

figs. 5-6 - Planisphinctes (Planisphinctes) tenuissimus (Siemiradzki, 1898), wholly septate, incomplete, fragmentary specimen. TTKM A 0777. 5 - Lateral view. 6 - Ventral view.

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resulting in a slight narrowing of the umbilicus on the Remarks - Arkell (1951-59, p. 173) was undecided last part of the conch. Further difference is the change about the generic status of Lobosphinctes, but later of ribbing from straight radial to slight prorsiradiate on authors (e.g. Sturani, 1967) kept its independence, at least the outer whorls. as a subgenus, on the basis of the evolute coiling, the suture-line with very tight lobes, and the strong Distribution - The lectotype came from around the constrictions. According to Torrens (1987, p. 104) these boundary of the Garantiana and Parkinsoni Zones (Rioult features and the lack of parabolic nodes on the inner et al. in Fischer, 1994), but most of the well-localized whorls suggest that Lobosphinctes is a Leptosphinctid specimens in the literature (e.g. Galácz, 1980; Sandoval, with homoeomorphic similarities to Procerites. 1990) are recorded from the Parkinsoni Zone. Additional distinct character is the ribbing, with the equally strong primaries and secondaries, which do not fade on the body chamber. The Kumeta specimens differ Planisphinctes (Lobosphinctes) intersertus Buckman, from the type in having lower, narrower last whorl. 1923 Pl. 3, fig. 1; Pl. 4, figs. 1-2 Distribution - The holotype of L. intersertus came from the Parkinsoni Zone, Truellei Subzone (see Torrens, 1923 Lobosphinctes intersertus, nov. - BUCKMAN 1909-30, Pl. 1987, p. 104), but the species ranges up into the basal 447, figs. 1-2. Bathonian Zigzag Zone. It is a widely-distributed form, 1964 Procerites (Lobosphinctes) intersertus S. Buckman - having been recorded from NW European and STURANI, p. 34, Pl. 4, fig. 5, text-fig. 30. 1967 Procerites (Lobosphinctes) intersertus (S. Buckman) - Mediterranean localities, and from Central Asia as well STURANI, p. 44, Pl. 16, figs. 1-3, 6, Pl. 17, fig. 1. (see the synonymy above). The exotic occurrence of a 1973 Procerites (Lobosphinctes) cfr. intersertus (S. Buckman) very similar Perisphinctid in the Argentine Rotundum - PAVIA, p. 139, Pl. 29, figs. 1-2, 4. Zone (deeper upper Bajocian) indicated the probable 1983 Planisphinctes (Lobosphinctes) intersertus (Buckman, presence of uppermost Bajocian ammonites in the 1923) - SANDOVAL, p. 413, Pl. 39, fig. 7, text-fig. 121G. Southern Andes (see Riccardi et al., 1989, p. 559, 1984 Lobosphinctes intersertus S. Buckman - NASTASEANU & Westermann in Hillebrandt et al., 1992, p. 258). BADALUTA, p. 86, Pl. 5, fig. 1. 1987 Lobosphinctes gr. intersertus S.S. Buckman - TORRENS, p. 104, Pl. 4, figs. 3-4. ? 1989 Lobosphinctes intersertus Buckman- RICCARDI ET AL., p. Planisphinctes (Planisphinctes) tenuissimus 567, Pl. 1, fig. 6. (Siemiradzki, 1898) ? 1992 Lobosphinctes intersertus Buck. - WESTERMANN in Pl. 4, figs. 5-6 HILLEBRANDT ET AL., p. 258, Pl. 61, fig. 1. 1993 Prorsisphinctes intersertus (Buckman), 1923 - BESNOSOV 1898 Perisphinctes tenuissimus n.sp. - SIEMIRADZKI, p. 85, Pl. & MITTA, p. 127, Pl. 16, fig. 1. 21, fig. 19, text-fig. 3. 1998 Prorsisphinctes (Prorsisphinctes) intersertus (Buckman, 1927 Perisphinctes perspicuus Parona - DORN, p. 244, Pl. 7, fig. 1923) - BESNOSOV & MITTA, p. 27, Pl. 12, fig. 6, Pl. 13, 2. figs. 1-5. 1966 Planisphinctes tenuissimus (Siemiradzki) - STURANI, p. 40, 2000 Prorsisphinctes (Prorsisphinctes) intersertus (Buckman) Pl. 12, figs. 1,4. - BESNOSOV & MITTA, Pl. 7, fig. 1. 1972 Siemiradzkia (Planisphinctes) incognita sp.n. - STEPHANOV, 2000 Lobosphinctes intersertus Buckman - FERNÁNDEZ-LÓPEZ, p. 35. p. 48, Pl. 1, fig. 2. non 1982 Planisphinctes tenuissimus (Siemiradzki) - AZARIAN, p. 157, ? 2005 Planisphinctes (Lobosphinctes) ex gr. intersertus Pl. 33, figs. 2-3. Buckman, 1923 - SCHLÖGL ET AL., p. 344, Pl. 10, figs. 1- 1983 Planisphinctes (Planisphinctes) tenuissimus (Siemiradzki, 2. 1898) - SANDOVAL, p. 408, Pl. 38, fig. 2, Pl. 39, fig. 5, text-figs. 121F, 123B. Material - Two well-preserved, but incomplete big 1985 Planisphinctes tenuissimus (Siemiradzki) - ROSTOVTSEV, specimens. The smaller one is of 150 mm diameter, the p. 168, Pl. 45, figs. 5-7. other, somewhat bigger specimen is about 165 mm and non 1993 Planisphinctes tenuissimus (Siemiradzki), 1898 - BESNOSOV shows the aperture. & MITTA, p. 131, Pl. 19, fig. 2. 1997 Planisphinctes tenuissimus (Siemiradzki) - RIOULT ET AL., p. 52, Pl. 15, fig. 11. Description - Comparatively big, serpenticone non 1998 Prorsisphinctes (Planisphinctes) tenuissimus (Siemiradzki, ammonites with wide, shallow umbilicus. The umbilical 1898) - BESNOSOV & MITTA, p. 27, Pl. 15, figs. 1, 2. sides are low and obliquely rounded, the flanks are ? 2005 Planisphinctes (Planisphinctes) ex gr. tenuissimus slightly convex, and the venter is rounded and half- (Siemiradzki, 1898) - SCHLÖGL ET AL., p. 344, Pl. 9, figs. circular. The whorls show regular ribbing both on the 3-4. phragmocone and on the body-chamber. There are 53 rounded, straight, slightly prorsiradiate inner ribs on the Material - A single, incomplete, wholly septate whorl penultimate whorl, and 38 primaries on the last half-whorl fragment. on the smaller specimen. It means that the number of ribs increases significantly on the body-chamber. Each Description - A widely-umbilicated septate whorl with primaries branch into two or three secondaries, and rounded, higher than wide cross-section. The umbilical intercalatories also appear. Constrictions occur mainly wall is low, the umbilical margin is blunt, the flanks are in the inner and middle whorls. convex and round into the wide, arched venter. The Suture-lines cannot be visible. sculpture is dense ribbing with slightly curved, radial inner ribs and prorsiradiate, slightly forwardly curved secondary

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ribs. The primary/secondary rib ratio is 1:2, with some Oxfordium, Süddeutschland). Berliner geowissenschaftliche trifurcate and few single primaries. Abhandlungen, E 36: 189-241. Azarian N.R. (1982). [Jurassic Ammonites of the Armenian Soviet Republic]. 191 pp. Academy of Science of Armenia, Jerevan [in Remarks - It is a general opinion to regard Russian]. Planisphinctes as the microconch of Lobosphinctes (see Besnosov N.V. & Mitta V.V. (1993). [Upper Bajocian and Bathonian e.g. Torrens, 1987, p. 104). Beznosov & Mitta (1993, ammonites of the Northern Caucasus and Central Asia]. Bulletin 1998) figured fine Siemiradzkia specimens under this of VNIGNI, Nedra: 347 pp. Moscow [in Russian]. name. Besnosov N.V. & Mitta V.V. (1998). Catalogue of Ammonitida and key sections of the Upper Bajocian-Lower Bathonian of North Caucasus. Bulletin of K.F. VNIGNI, 1: 1-70. Distribution - Planisphinctes tenuissimus and its Besnosov N.V. & Mitta V.V. (2000). Jurassic geology and ammonites most probable macroconch pair Lobosphinctes of Great Balkhan (Western Turkmenistan). Bulletin of K.F. intersertus indicate the upper, Parkinsonia bomfordi VNIGNI, 5: 1-115. Subzone of the Parkinsoni Zone (see Pavia, 2000). As Brasil L. (1894). Céphalopodes nouveaux ou peu connus des étages shown by the synonymy, the geographic distribution of Jurassiques de Normandie. Bulletin de la Société géologique de P. tenuissimus is wide, just as that of its probable Normandie, 16 [1892-93]: 27-46. macroconch. Martire & Pavia (2004, p. 29) recorded the Buckman S. S. (1909-30). (Yorkshire) Type Ammonites. vols. I- VII, pls. 1-790, Wheldon & Wesley, London. species from the “Rocca chi Parra” quarry nearby. Chiari M., Baldanza A. & Parisi G. (2004). Integrated stratigraphy (radiolarians and calcareous nannofossils) of the Jurassic siliceous sediments from Monte Kumeta (western Sicily). Rivista CONCLUSIONS Italiana di Paleontologia e Stratigrafia, 110 (1): 129-140. Clari P.A., Marini P., Pastorini M. & Pavia G. (1984). Il Rosso The studied Parkinsoni Zone ammonites from Monte Ammonitico Inferiore (Baiociano-Calloviano) nei Monti Lessini settentrionali (Verona). Rivista Italiana di Paleontologia e Kumeta represent a typical Mediterranean assemblage, Stratigrafia, 90 (1): 15-86. with Phylloceratids as the most common, and Conti M.A. & Monari S. (2001). Middle Jurassic gastropods from Perisphinctids as the most characteristic elements. The the Central High Atlas, Morocco. Geobios, 34 (2): 183-214. here described fossil assemblage is very similar to those Di Stefano P., Galácz A., Mallarino G., Mindszenty A., Vörös A., mentioned from other Mediterranean areas: northern Italy Baudo F., Lantos Z. & Görög Á. (2001). Facies, geometrie (Sturani, 1964, Clari et al., 1984), Bakony Mts., Hungary deposizionali e superfici di discontinuitá associate alla genesi di una paleoscarpata giurassica. Il caso di Monte Kumeta (Sicilia): (Galácz, 1980), thus palaeogeographically probably GEOITALIA 3o Forum FIST, Sessione 8: 214-215. belongs to the pelagic realm of the Western Di Stefano P., Galácz A., Mallarino G., Mindszenty A. & Vörös A. (Mediterranean) Tethys. (2002a). Birth and evolution of a Jurassic escarpment: Monte The indications from the nearby Rocca chi Parra Kumeta, Western Sicily. Facies, 46: 273-298. locality (Pavia et al., 2002) suggest a very similar Di Stefano P., Galácz A., Mallarino G., Mindszenty A. & Vörös A. taxonomic composition; however the depositional (2002b). Stop 14 - Birth and dynamics of a Jurassic submarine history there is different, with upper Bajocian and lower escarpment at Monte Kumeta. In Santantonio M. (ed.), 6th International Symposium on the Jurassic System, Palermo, Italy, Bathonian forms occurring in the same condensed bed. 12-22 September 2002, General Field Trip Guidebook: 102-110. In the Monte Kumeta quarries the represented Middle Dietze V. & Dietl G. (2006). Feinstratigraphie und Ammoniten- Jurassic fossiliferous levels all seem to yield Faunen-Horizonte im Ober-Bajocium und Bathonium des Ipf- uncondensed ammonites, in spite of the fact that many Gebietes (Schwäbische Alb, Südwestdeutschland). Stuttgarter show traces of reworking before burial. Beiträge des Naturkunde, Serie B (Geologie und Paläontologie), 360: 1-51. Dorn P. (1927). Die Ammonitenfauna der Parkinsonienschichten bei Thalmässig (Frankenalb). Jahrbuch der Preussischen ACKNOWLEDGEMENTS Geologischen Landesanstalt, 48: 225-251. Dresnay R. du. (1967). Relations paleogéographiques entre sillons The collecting of the above-described ammonites, and the atlasiques (Haut et Moyen-Atlas) et plate-forme des Hautes- studying of the general and detailed stratigraphy of the Monte Kumeta Plateaux (Maroc oriental) pendant l’Aalenien et le Bajocien. Jurassic were realised through cooperation with colleagues and Mémoire de B.R.G.M, Fr., 75, [1971]: 147-161. friends from the Dipartimento di Geologia e Geodesia, Università di Enay R., Mangold C., du Dresnay R. & Rakus M. (1987). Arrivals Palermo. I am particularly indebted to Prof. Piero di Stefano, who of Arabian origin among the ammonite faunas of Morocco during helped the studies in all possible ways. Prof. Andrea Mindszenty the Bajocian-Bathonian. Palaeogeography, Palaeoclimatology, and Dr. Attila Vörös, my friends from Budapest gave support on the Palaeoecology, 61: 107-120. field and in the time of drawing conclusions on stratigraphy. The Fallot P. & Blachet F. (1923). Observations sur la faune des Terrains beautiful times we shared in western Sicily remain unforgettable. I Jurassiques de la region de Cardó et de Tortosa (Province de am grateful to László Makádi for help in preparing the text-figures. Tarragone). Treballs de l’Institució Catalana d’História Natural, 1921-1922 (11): 73-261. Fernández-López S. (1985). El Bajociense en la Cordillera Iberica. REFERENCES Depart. Paleont., Univ. Compl. Madrid: 1-848. Fernández-López S. (2000). Lower Bathonian ammonites of Serra Amanniazov K. (1972). [North Asian Parkinsoniids]: 34 pp. de la Creu (Tivissa, Catalan Basin, Spain). Revue de Turkmenian State Univ. Press, Ashabad [in Russian]. Paléobiologie, Vol. Spéc. 8: 45-52. Arkell W.J. (1951-59). Monograph of the English Bathonian Fischer J.-C. (1994). Révision critique de la Paléontologie française Ammonoidea: Palaeontographical Society, 104-112: 1-264. d’Alcide d’Orbigny. Vol. I, Céphalopodes jurassiques: 344 pp. Arkell W.J. (1956). Jurassic Geology of the World. xiv + 806 pp, Masson, Paris. Oliver & Boyd, Edinburgh and London. Floridia G. (1931). Osservazioni geologiche sul M. Inici (Trapani). Arp G. (2001). 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