Dinosaur Origins
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Ischigualasto Formation. the Second Is a Sile- Diversity Or Abundance, but This Result Was Based on Only 19 of Saurid, Ignotosaurus Fragilis (Fig
This article was downloaded by: [University of Chicago Library] On: 10 October 2013, At: 10:52 Publisher: Taylor & Francis Informa Ltd Registered in England and Wales Registered Number: 1072954 Registered office: Mortimer House, 37-41 Mortimer Street, London W1T 3JH, UK Journal of Vertebrate Paleontology Publication details, including instructions for authors and subscription information: http://www.tandfonline.com/loi/ujvp20 Vertebrate succession in the Ischigualasto Formation Ricardo N. Martínez a , Cecilia Apaldetti a b , Oscar A. Alcober a , Carina E. Colombi a b , Paul C. Sereno c , Eliana Fernandez a b , Paula Santi Malnis a b , Gustavo A. Correa a b & Diego Abelin a a Instituto y Museo de Ciencias Naturales, Universidad Nacional de San Juan , España 400 (norte), San Juan , Argentina , CP5400 b Consejo Nacional de Investigaciones Científicas y Técnicas , Buenos Aires , Argentina c Department of Organismal Biology and Anatomy, and Committee on Evolutionary Biology , University of Chicago , 1027 East 57th Street, Chicago , Illinois , 60637 , U.S.A. Published online: 08 Oct 2013. To cite this article: Ricardo N. Martínez , Cecilia Apaldetti , Oscar A. Alcober , Carina E. Colombi , Paul C. Sereno , Eliana Fernandez , Paula Santi Malnis , Gustavo A. Correa & Diego Abelin (2012) Vertebrate succession in the Ischigualasto Formation, Journal of Vertebrate Paleontology, 32:sup1, 10-30, DOI: 10.1080/02724634.2013.818546 To link to this article: http://dx.doi.org/10.1080/02724634.2013.818546 PLEASE SCROLL DOWN FOR ARTICLE Taylor & Francis makes every effort to ensure the accuracy of all the information (the “Content”) contained in the publications on our platform. However, Taylor & Francis, our agents, and our licensors make no representations or warranties whatsoever as to the accuracy, completeness, or suitability for any purpose of the Content. -
The Origins of Dinosauria: Much Ado About Nothing
View metadata, citation and similar papers at core.ac.uk brought to you by CORE provided by RERO DOC Digital Library [Palaeontology, Vol. 57, Part 3, 2014, pp. 469–478] FRONTIERS IN PALAEONTOLOGY THE ORIGINS OF DINOSAURIA: MUCH ADO ABOUT NOTHING by MAX C. LANGER Departamento de Biologia, FFCLRP, Universidade de S~ao Paulo, Av. Bandeirantes 3900,14040-901, Ribeir~ao Preto, SP Brazil; e-mail: [email protected] Typescript received 19 February 2014; accepted in revised form 7 March 2014 Abstract: Research this century has greatly improved our early members of the main dinosauromorph lineages and knowledge of the origin and early radiation of dinosaurs. requires a more meticulous assessment of characters and The unearthing of several new dinosaurs and close out- homologies than those recently conducted. Presently, the groups from Triassic rocks from various parts of the oldest uncontroversial dinosaur records come from Late world, coupled with improved phylogenetic analyses, has Triassic (Carnian) rocks of South America, southern Africa set a basic framework in terms of timing of events and and India, hinting at a south-western Pangaea origin of the macroevolutionary patterns. However, important parts of group. Besides, macroevolutionary approaches suggest that the early dinosauromorph evolutionary history are still the rise of dinosaurs was a more gradual process than pre- poorly understood, rendering uncertain the phylogenetic viously understood. Obviously, these tentative scenarios position of silesaurids as either non-dinosaur Dinosaurifor- need to be tested by new fossil finds, which should also mes or ornithischians, as well as that of various early help close the major gaps recognized in the fossil record of saurischians, such as Eoraptor lunensis and herrerasaurs, as Triassic dinosauromorphs. -
The Anatomy of Asilisaurus Kongwe, a Dinosauriform from the Lifua
THE ANATOMICAL RECORD (2019) The Anatomy of Asilisaurus kongwe,a Dinosauriform from the Lifua Member of the Manda Beds (~Middle Triassic) of Africa 1 2 3 STERLING J. NESBITT , * MAX C. LANGER, AND MARTIN D. EZCURRA 1Department of Geosciences, Virginia Tech, Blacksburg, Virginia 2Departamento de Biologia, Universidade de Sao~ Paulo, Ribeirao~ Preto, Brazil 3Sección Paleontología de Vertebrados CONICET—Museo Argentino de Ciencias Naturales “Bernardino Rivadavia”, Buenos Aires, Argentina ABSTRACT The diagnosis of Dinosauria and interrelationships of the earliest dino- saurs relies on careful documentation of the anatomy of their closest rela- tives. These close relatives, or dinosaur “precursors,” are typically only documented by a handful of fossils from across Pangea and nearly all speci- mens are typically missing important regions (e.g., forelimbs, pelves, skulls) that appear to be important to help resolving the relationships of dinosaurs. Here, we fully describe the known skeletal elements of Asilisaurus kongwe, a dinosauriform from the Middle Triassic Manda Beds of the Ruhuhu Basin of Tanzania. The taxon is known from many disarticulated and partially articulated remains and, most importantly, from a spectacularly preserved associated skeleton of an individual containing much of the skull, pectoral and pelvic girdles, forelimb and hindlimb, and parts of the vertebral column including much of the tail. The unprecedented detail of the anatomy indi- cates that Asilisaurus kongwe had a unique skull that was short and had both a premaxillary and dentary edentulous margin, but retained a number of character states plesiomorphic for Archosauria, including a crocodylian- like ankle configuration and a rather short foot with well-developed meta- tarsals I and V. -
The Origin and Early Evolution of Dinosaurs
Biol. Rev. (2010), 85, pp. 55–110. 55 doi:10.1111/j.1469-185X.2009.00094.x The origin and early evolution of dinosaurs Max C. Langer1∗,MartinD.Ezcurra2, Jonathas S. Bittencourt1 and Fernando E. Novas2,3 1Departamento de Biologia, FFCLRP, Universidade de S˜ao Paulo; Av. Bandeirantes 3900, Ribeir˜ao Preto-SP, Brazil 2Laboratorio de Anatomia Comparada y Evoluci´on de los Vertebrados, Museo Argentino de Ciencias Naturales ‘‘Bernardino Rivadavia’’, Avda. Angel Gallardo 470, Cdad. de Buenos Aires, Argentina 3CONICET (Consejo Nacional de Investigaciones Cient´ıficas y T´ecnicas); Avda. Rivadavia 1917 - Cdad. de Buenos Aires, Argentina (Received 28 November 2008; revised 09 July 2009; accepted 14 July 2009) ABSTRACT The oldest unequivocal records of Dinosauria were unearthed from Late Triassic rocks (approximately 230 Ma) accumulated over extensional rift basins in southwestern Pangea. The better known of these are Herrerasaurus ischigualastensis, Pisanosaurus mertii, Eoraptor lunensis,andPanphagia protos from the Ischigualasto Formation, Argentina, and Staurikosaurus pricei and Saturnalia tupiniquim from the Santa Maria Formation, Brazil. No uncontroversial dinosaur body fossils are known from older strata, but the Middle Triassic origin of the lineage may be inferred from both the footprint record and its sister-group relation to Ladinian basal dinosauromorphs. These include the typical Marasuchus lilloensis, more basal forms such as Lagerpeton and Dromomeron, as well as silesaurids: a possibly monophyletic group composed of Mid-Late Triassic forms that may represent immediate sister taxa to dinosaurs. The first phylogenetic definition to fit the current understanding of Dinosauria as a node-based taxon solely composed of mutually exclusive Saurischia and Ornithischia was given as ‘‘all descendants of the most recent common ancestor of birds and Triceratops’’. -
A Phylogenetic Analysis of the Basal Ornithischia (Reptilia, Dinosauria)
A PHYLOGENETIC ANALYSIS OF THE BASAL ORNITHISCHIA (REPTILIA, DINOSAURIA) Marc Richard Spencer A Thesis Submitted to the Graduate College of Bowling Green State University in partial fulfillment of the requirements of the degree of MASTER OF SCIENCE December 2007 Committee: Margaret M. Yacobucci, Advisor Don C. Steinker Daniel M. Pavuk © 2007 Marc Richard Spencer All Rights Reserved iii ABSTRACT Margaret M. Yacobucci, Advisor The placement of Lesothosaurus diagnosticus and the Heterodontosauridae within the Ornithischia has been problematic. Historically, Lesothosaurus has been regarded as a basal ornithischian dinosaur, the sister taxon to the Genasauria. Recent phylogenetic analyses, however, have placed Lesothosaurus as a more derived ornithischian within the Genasauria. The Fabrosauridae, of which Lesothosaurus was considered a member, has never been phylogenetically corroborated and has been considered a paraphyletic assemblage. Prior to recent phylogenetic analyses, the problematic Heterodontosauridae was placed within the Ornithopoda as the sister taxon to the Euornithopoda. The heterodontosaurids have also been considered as the basal member of the Cerapoda (Ornithopoda + Marginocephalia), the sister taxon to the Marginocephalia, and as the sister taxon to the Genasauria. To reevaluate the placement of these taxa, along with other basal ornithischians and more derived subclades, a phylogenetic analysis of 19 taxonomic units, including two outgroup taxa, was performed. Analysis of 97 characters and their associated character states culled, modified, and/or rescored from published literature based on published descriptions, produced four most parsimonious trees. Consistency and retention indices were calculated and a bootstrap analysis was performed to determine the relative support for the resultant phylogeny. The Ornithischia was recovered with Pisanosaurus as its basalmost member. -
Chapter 10, the Mistaken Extinction, by Lowell Dingus and Timothy Rowe, New York, W
Chapter 10, The Mistaken Extinction, by Lowell Dingus and Timothy Rowe, New York, W. H. Freeman, 1998. CHAPTER 10 Dinosaurs Challenge Evolution Enter Sir Richard Owen More than 150 years ago, the great British naturalist Richard Owen (fig. 10.01) ignited the controversy that Deinonychus would eventually inflame. The word "dinosaur" was first uttered by Owen in a lecture delivered at Plymouth, England in July of 1841. He had coined the name in a report on giant fossil reptiles that were discovered in England earlier in the century. The root, Deinos, is usually translated as "terrible" but in his report, published in 1842, Owen chose the words "fearfully great"1. To Owen, dinosaurs were the fearfully great saurian reptiles, known only from fossil skeletons of huge extinct animals, unlike anything alive today. Fig. 10.01 Richard Owen as, A) a young man at about the time he named Dinosauria, B) in middle age, near the time he described Archaeopteryx, and C) in old age. Dinosaur bones were discovered long before Owen first spoke their name, but no one understood what they represented. The first scientific report on a dinosaur bone belonging was printed in 1677 by Rev. Robert Plot in his work, The Natural History of Oxfordshire. This broken end of a thigh bone, came to Plot's attention during his research. It was nearly 60 cm in circumference--greater than the same bone in an elephant (fig.10.02). We now suspect that it belonged to Megalosaurus bucklandii, a carnivorous dinosaur now known from Oxfordshire. But Plot concluded that it "must have been a real Bone, now petrified" and that it resembled "exactly the figure of the 1 Chapter 10, The Mistaken Extinction, by Lowell Dingus and Timothy Rowe, New York, W. -
University of Birmingham a New Dinosaur With
University of Birmingham A new dinosaur with theropod affinities from the Late Triassic Santa Maria Formation, South Brazil Marsola, Julio; Bittencourt, Jonathas; Butler, Richard; Da Rosa, Atila; Sayao, Juliana; Langer, Max DOI: 10.1080/02724634.2018.1531878 License: None: All rights reserved Document Version Peer reviewed version Citation for published version (Harvard): Marsola, J, Bittencourt, J, Butler, R, Da Rosa, A, Sayao, J & Langer, M 2019, 'A new dinosaur with theropod affinities from the Late Triassic Santa Maria Formation, South Brazil', Journal of Vertebrate Paleontology, vol. 38, no. 5, e1531878. https://doi.org/10.1080/02724634.2018.1531878 Link to publication on Research at Birmingham portal Publisher Rights Statement: Checked for eligibility: 25/07/2018 This is the accepted manuscript for a forthcoming publication in Journal of Vertebrate Paleontology. General rights Unless a licence is specified above, all rights (including copyright and moral rights) in this document are retained by the authors and/or the copyright holders. The express permission of the copyright holder must be obtained for any use of this material other than for purposes permitted by law. •Users may freely distribute the URL that is used to identify this publication. •Users may download and/or print one copy of the publication from the University of Birmingham research portal for the purpose of private study or non-commercial research. •User may use extracts from the document in line with the concept of ‘fair dealing’ under the Copyright, Designs and Patents Act 1988 (?) •Users may not further distribute the material nor use it for the purposes of commercial gain. -
Integrating Gross Morphology and Bone Histology to Assess Skeletal Maturity in Early Dinosauromorphs: New Insights from Dromomeron (Archosauria: Dinosauromorpha)
Integrating gross morphology and bone histology to assess skeletal maturity in early dinosauromorphs: new insights from Dromomeron (Archosauria: Dinosauromorpha) Christopher T. Griffin1, Lauren S. Bano2, Alan H. Turner3, Nathan D. Smith4, Randall B. Irmis5,6 and Sterling J. Nesbitt1 1 Department of Geosciences, Virginia Tech, Blacksburg, VA, USA 2 Department of Biology, Virginia Tech, Blacksburg, VA, USA 3 Department of Anatomical Sciences, Stony Brook University, Stony Brook, NY, USA 4 The Dinosaur Institute, Natural History Museum of Los Angeles County, Los Angeles, CA, USA 5 Natural History Museum of Utah, University of Utah, Salt Lake City, UT, USA 6 Department of Geology and Geophysics, University of Utah, Salt Lake City, UT, USA ABSTRACT Understanding growth patterns is central to properly interpreting paleobiological signals in tetrapods, but assessing skeletal maturity in some extinct clades may be difficult when growth patterns are poorly constrained by a lack of ontogenetic series. To overcome this difficulty in assessing the maturity of extinct archosaurian reptiles—crocodylians, birds and their extinct relatives—many studies employ bone histology to observe indicators of the developmental stage reached by a given individual. However, the relationship between gross morphological and histological indicators of maturity has not been examined in most archosaurian groups. In this study, we examined the gross morphology of a hypothesized growth series of Dromomeron romeri femora (96.6–144.4 mm long), the first series of a non- dinosauriform dinosauromorph available for such a study. We also histologically sampled several individuals in this growth series. Previous studies reported that Submitted 7 August 2018 D. romeri lacks well-developed rugose muscle scars that appear during ontogeny in Accepted 20 December 2018 closely related dinosauromorph taxa, so integrating gross morphology and Published 11 February 2019 histological signal is needed to determine reliable maturity indicators for early Corresponding author Christopher T. -
Comparative Bone Microstructure of Three Archosauromorphs from the Carnian, Late Triassic Chañares Formation of Argentina
Comparative bone microstructure of three archosauromorphs from the Carnian, Late Triassic Chañares Formation of Argentina JORDI ALEXIS GARCIA MARSÀ, FEDERICO L. AGNOLÍN, and FERNANDO E. NOVAS Marsà, J.A.G., Agnolín, F.L., and Novas, F.E. 2020. Comparative bone microstructure of three archosauromorphs from the Carnian, Late Triassic Chañares Formation of Argentina. Acta Palaeontologica Polonica 65 (2): 387–398. The Chañares Formation exhibits one of the most important archosauriform records of early Carnian ecosystems. Here we present new data on the palaeohistology of Chañares archosauriforms and provide new insights into their paleobiology, as well as possible phylogenetically informative traits. Bone microstructure of Lagerpeton chanarensis and Tropidosuchus romeri is dominated by fibro-lamellar tissue and dense vascularization. On the other hand, Chanaresuchus bonapartei is more densely vascularized, but with cyclical growth characterized by alternate fibro-lamellar, parallel-fibered and lamellar-zonal tissues. Dense vascularization and fibro-lamellar tissue imply fast growth and high metabolic rates for all these taxa. These histological traits may be tentatively interpreted as a possible adaptative advantage in front of Chañares Formation environmental conditions. Key words: Archosauromorpha, Lagerpeton, Tropidosuchus, paleobiology, paleohistology, Mesozoic, South America. Jordi Alexis Garcia Marsà [[email protected]] and Fernando E. Novas [[email protected]], Labora- torio de Anatomía Comparada y Evolución de los Vertebrados, -
New Perspectives on Pterosaur Palaeobiology
Downloaded from http://sp.lyellcollection.org/ by guest on September 24, 2021 New perspectives on pterosaur palaeobiology DAVID W. E. HONE1*, MARK P. WITTON2 & DAVID M. MARTILL2 1Queen Mary University of London, Mile End Road, London E14NS, UK 2School of Earth and Environmental Sciences, University of Portsmouth, Burnaby Road, Portsmouth PO1 3QL, UK *Correspondence: [email protected] Abstract: Pterosaurs were the first vertebrates to evolve powered flight and occupied the skies of the Mesozoic for 160 million years. They occurred on every continent, evolved their incredible pro- portions and anatomy into well over 100 species, and included the largest flying animals of all time among their ranks. Pterosaurs are undergoing a long-running scientific renaissance that has seen elevated interest from a new generation of palaeontologists, contributions from scientists working all over the world and major advances in our understanding of their palaeobiology. They have espe- cially benefited from the application of new investigative techniques applied to historical speci- mens and the discovery of new material, including detailed insights into their fragile skeletons and their soft tissue anatomy. Many aspects of pterosaur science remain controversial, mainly due to the investigative challenges presented by their fragmentary, fragile fossils and notoriously patchy fossil record. With perseverance, these controversies are being resolved and our understand- ing of flying reptiles is increasing. This volume brings together a diverse set of papers on numerous aspects of the biology of these fascinating reptiles, including discussions of pterosaur ecology, flight, ontogeny, bony and soft tissue anatomy, distribution and evolution, as well as revisions of their taxonomy and relationships. -
University of Birmingham the Earliest Bird-Line Archosaurs and The
University of Birmingham The earliest bird-line archosaurs and the assembly of the dinosaur body plan Nesbitt, Sterling; Butler, Richard; Ezcurra, Martin; Barrett, Paul; Stocker, Michelle; Angielczyk, Kenneth; Smith, Roger; Sidor, Christian; Niedzwiedzki, Grzegorz; Sennikov, Andrey; Charig, Alan DOI: 10.1038/nature22037 License: None: All rights reserved Document Version Peer reviewed version Citation for published version (Harvard): Nesbitt, S, Butler, R, Ezcurra, M, Barrett, P, Stocker, M, Angielczyk, K, Smith, R, Sidor, C, Niedzwiedzki, G, Sennikov, A & Charig, A 2017, 'The earliest bird-line archosaurs and the assembly of the dinosaur body plan', Nature, vol. 544, no. 7651, pp. 484-487. https://doi.org/10.1038/nature22037 Link to publication on Research at Birmingham portal Publisher Rights Statement: Checked for eligibility: 03/03/2017. General rights Unless a licence is specified above, all rights (including copyright and moral rights) in this document are retained by the authors and/or the copyright holders. The express permission of the copyright holder must be obtained for any use of this material other than for purposes permitted by law. •Users may freely distribute the URL that is used to identify this publication. •Users may download and/or print one copy of the publication from the University of Birmingham research portal for the purpose of private study or non-commercial research. •User may use extracts from the document in line with the concept of ‘fair dealing’ under the Copyright, Designs and Patents Act 1988 (?) •Users may not further distribute the material nor use it for the purposes of commercial gain. Where a licence is displayed above, please note the terms and conditions of the licence govern your use of this document. -
A Re-Evaluation of the Enigmatic Dinosauriform Caseosaurus Crosbyensis from the Late Triassic of Texas, USA and Its Implications for Early Dinosaur Evolution
A re-evaluation of the enigmatic dinosauriform Caseosaurus crosbyensis from the Late Triassic of Texas, USA and its implications for early dinosaur evolution MATTHEW G. BARON and MEGAN E. WILLIAMS Baron, M.G. and Williams, M.E. 2018. A re-evaluation of the enigmatic dinosauriform Caseosaurus crosbyensis from the Late Triassic of Texas, USA and its implications for early dinosaur evolution. Acta Palaeontologica Polonica 63 (1): 129–145. The holotype specimen of the Late Triassic dinosauriform Caseosaurus crosbyensis is redescribed and evaluated phylogenetically for the first time, providing new anatomical information and data on the earliest dinosaurs and their evolution within the dinosauromorph lineage. Historically, Caseosaurus crosbyensis has been considered to represent an early saurischian dinosaur, and often a herrerasaur. More recent work on Triassic dinosaurs has cast doubt over its supposed dinosaurian affinities and uncertainty about particular features in the holotype and only known specimen has led to the species being regarded as a dinosauriform of indeterminate position. Here, we present a new diagnosis for Caseosaurus crosbyensis and refer additional material to the taxon—a partial right ilium from Snyder Quarry. Our com- parisons and phylogenetic analyses suggest that Caseosaurus crosbyensis belongs in a clade with herrerasaurs and that this clade is the sister taxon of Dinosauria, rather than positioned within it. This result, along with other recent analyses of early dinosaurs, pulls apart what remains of the “traditional” group of dinosaurs collectively termed saurischians into a polyphyletic assemblage and implies that Dinosauria should be regarded as composed exclusively of Ornithoscelida (Ornithischia + Theropoda) and Sauropodomorpha. In addition, our analysis recovers the enigmatic European taxon Saltopus elginensis among herrerasaurs for the first time.