Microfossil Paleontology and Biostratigraphy of The
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MICROFOSSIL PALEONTOLOGY AND BIOSTRATIGRAPHY OF THE EARLY MESOPROTEROZOIC BELT SUPERGROUP, MONTANA by Zachary Robert Adam A dissertation submitted in partial fulfillment of the requirements for the degree of Doctor of Philosophy in Earth Sciences MONTANA STATE UNIVERSITY Bozeman, Montana November 2014 ©COPYRIGHT by Zachary Robert Adam 2014 All Rights Reserved ii DEDICATION I would like to dedicate this work to my wife, our family in the US and Turkey, and my friends for all that they have given over the years so that I may have the comfort, care and peace of mind to focus so intently on so many tiny pieces of formerly animate matter. I also wish to dedicate this work to all eukaryotes (and complexity-equivalent organisms elsewhere in the universe) that have ever risen above the selective forces that would otherwise constrain us to an existence lacking an appreciation for the Infinite. iii ACKNOWLEDGMENTS This dissertation was made possible by the kindness and generosity of many individuals and organizations. David Mogk and Mark Skidmore guided the genesis and conclusion of the entire project with aplomb. James Schmitt and Nicholas Atwood provided insightful field interpretations of sedimentology. David Lageson illuminated many nuances of the Belt Supergroup structural environment. Nicholas Butterfield of the University of Cambridge provided invaluable micropaleontological guidance in the field and laboratory, and helpful feedback regarding the content and organization of this dissertation. Don Winston of the University of Montana introduced me to the sedimentary cycles and mysteries of the Belt Supergroup. Gerald Zieg of Tintina Resources oriented me within the Helena Embayment and provided helpful feedback regarding the history of study of the Lower Belt units. This material is based upon work supported by the National Science Foundation Graduate Research Fellowship under Grant No. 2011120064. Tuition support was provided by the NSF Integrative Graduate Education and Research Trainee (IGERT) Program. Field activities were supported by awards from the American Philosophical Society and NASA Astrobiology Institute’s Lewis and Clark Fund for Exploration Research, the Tobacco Root Geological Society, the Belt Association, and the Donald L. Smith Memorial Fund. Recep Avci, Muhammedin Deliorman and Laura Kellerman of the Imaging and Chemical Analysis Laboratory at Montana State University assisted with XRD, electron microscope and atomic force microscope data collection. iv VITA Zachary Robert Adam was born May 30th, 1981 in Wenatchee, Washington to Robert and Linda Adam. After graduating from Cashmere High School in 1999, he attended the University of Washington, earning degrees in Earth and Space Sciences and Aeronautics and Astronautics Engineering in 2004. It was during this period that Zach picked up his now-lifelong fascination with Astrobiology, and shortly thereafter he joined the University of Washington’s Astrobiology Program, earning a Master’s degree in Aeronautics and Astronautics Engineering. After volunteering for Americorps at Lewis and Clark Elementary School in Missoula, Montana, Zach worked as a life support systems engineer and launch vehicle inspector for the Office of Commercial Space Transportation at the Federal Aviation Administration, focusing on the SeaLaunch and SpaceX vehicle families. Zach then returned to academia through Montana State University, where he was presented with the opportunity to design and conduct the research project described in this dissertation. v TABLE OF CONTENTS 1. INTRODUCTION ......................................................................................................... 1 2. MICROFOSSILS FROM THE GREYSON FORMATION ....................................... 12 Introduction ................................................................................................................... 12 Geological Setting ......................................................................................................... 19 Sampling and Methods ................................................................................................. 23 Assemblage Summary .................................................................................................. 25 Systematic Paleontology ............................................................................................... 39 Biochronologic Implications ......................................................................................... 47 Conclusions ................................................................................................................... 57 3. MICROFOSSILS FROM THE CHAMBERLAIN FORMATION .............................. 59 Introduction ................................................................................................................... 59 Geological Setting ......................................................................................................... 63 Sampling and Methods ................................................................................................. 68 Assemblage Summary .................................................................................................. 70 Systematic Paleontology ............................................................................................... 79 Discussion ..................................................................................................................... 85 Conclusions ................................................................................................................... 97 4. ONTOGENETIC RECONSTRUCTION OF MICROFOSSILS BASED ON VISIBLE MORPHOLOGICAL CHARACTERS ................................... 99 Introduction ................................................................................................................... 99 Geological Setting ....................................................................................................... 101 Methods ...................................................................................................................... 103 Results ......................................................................................................................... 104 Discussion ................................................................................................................... 114 Conclusions ................................................................................................................. 124 5. CONCLUSIONS AND FUTURE WORK ................................................................. 126 APPENDICES ................................................................................................................ 131 APPENDIX A: Image and Actual Size Measurement Calibrations ........................... 132 APPENDIX B: Rock Pyrolysis Data .......................................................................... 140 APPENDIX C: Clay Mineralogy XRD Data .............................................................. 142 vi TABLE OF CONTENTS - CONTINUED APPENDIX D: Individual Microfossil Location Notes ............................................. 149 APPENDIX E: Major Element Analyses ................................................................... 161 REFERENCES CITED…………………………………………………………………163 vii LIST OF TABLES Table Page 1. Comparison of fossil taxa occurring in the SCC-34 (Chamberlain Formation) and Newlan Creek (Greyson Formation) assemblages .................................................................... 86 2. Cross-comparison of interpreted host rock paleoenvironments (top row) and fossil assemblage composition (left column) from units of the Bylot Supergroup, Belt Supergroup and Roper Group. .............................................................................................. 90 viii LIST OF FIGURES Figure Page 1. Locations of outcrops sampled for microfossil content from throughout the Belt Supergroup .......................................................................... 6 2. Map of northern margins of the Helena Embayment (inset) and fossiliferous rocks ..................................................................................... 15 3. Unit stratigraphy and geochronological constraints of the lower Belt Supergroup ..................................................................................... 16 4. WP-133 fossiliferous outcrop (Greyson Formation) unoriented and oriented clay sample spectra ...................................................................... 24 5. Summary and occurrence of microfossil taxa identified from the Newlan Creek locality, Greyson Formation, Belt Supergroup, Montana ....................................................................................... 26 6. Leiosphaeridia specimens from the Newlan Creek locality, Greyson Formation ........................................................................................... 27 7. Caudosphaera winstonii specimens from the Newlan Creek locality, Greyson Formation ............................................................................. 28 8. Pluricellular aggregations from the Newlan Creek locality, Greyson Formation ........................................................................................... 29 9. Valeria lophostriata from the Newlan Creek locality, Greyson Formation ........................................................................................... 31 10. Dictyosphaera delicata from the Newlan Creek locality, Greyson Formation ........................................................................................