Olericulture – Hort 320 Lesson 4, Classif, Growth & Development
Jeremy S. Cowan WSU Spokane County Extension 222 N. Havana St. Spokane, WA 99202 Phone: 509-477-2145 Fax: 509-477-2087 Email: [email protected]
Classification
Cavalier- Linnaeus Haeckel Chatton Copeland Whittaker Woese Smith 1735 1866 1925 1938 1969 1990 1998 3 2 4 5 3 6 2 kingdoms kingdoms empires kingdoms kingdoms domains kingdoms Bacteria Prokaryota Monera Monera Bacteria (not Archaea Protista treated) Protozoa Protoctista Protista Chromista Eukaryota Plantae Eucarya Plantae Vegetabilia Plantae Plantae Fungi Fungi Animalia Animalia Animalia Animalia Animalia Botanical Classification
Kingdom: Plantae
Sub-Kingdom: Land Plants (Embryophyta)
Division: Seed Plants (Magnoliophyta)
Class: Cone Bearing (Gymnospermae) Flowering (Angiospermae)
Subclass: Monocotyledon (Liliopsida) Dicotyledon (Magnoliopsida) Plant Taxonomy Monocots
Corn - 1 seed leaf
Monocot – a plant whose embryo has one cotyledon Dicots
Castor Bean 2 seed leaves Dicots Monocots Dicots Monocots
Flower parts Flower parts In 4’s or 5’s In 3’s Dicots Monocots
Tap root Fibrous root Dicots Monocots Vegetable Classification
Monocot vegetables: Araceae – arum family vegetables: taro, dasheen related: calamus, jack-in-the-pulpit
Vegetable Classification
Monocot vegetables: Dioscoreaceae – yam family vegetables: yam related: wild yams Vegetable Classification
Monocot vegetables: Gramineae – grass family vegetable: sweet corn related: grasses, sedges
Vegetable Classification
Monocot vegetables: Lilieaceae – lily family vegetables: onion, leek, garlic, shallot, chive, asparagus related: lily, camas, solomon’s seal Vegetable Classification
Dicot vegetables: Polygonaceae – buckwheat family vegetables: rhubarb, sorrel related: knotweed, smart weed, dock
Vegetable Classification
Dicot vegetables: Amaranthaceae (Chenopodiaceae) – goosefoot family vegetables: beet, swiss chard, spinach related: lambsquarter, pigweed Vegetable Classification
Dicot vegetables: Brassicaceae (Cruciferae) – mustard family vegetables: cabbage, rutabaga, turnip, radish related: wild mustards
Vegetable Classification
Dicot vegetables: Euphorbiaceae – spurge family vegetables: cassava related: poinsettia, castor bean, spurge Vegetable Classification
Dicot vegetables: Fabaceae (Leguminosae) – pea family vegetables: pea, bean, cowpea, soybean, peanut, others related: alfalfa, acacia, lupine, clover
Vegetable Classification
Dicot vegetables: Malvaceae – mallow family vegetables: okra related: hibiscus, hollyhock Vegetable Classification
Dicot vegetables: Cucurbitaceae – gourd family vegetables: watermelon, cantaloupe, squash, pumpkin, cucumber, others related: luffa, wild cucumber
Vegetable Classification
Dicot vegetables: Apiaceae (Umbelliferae) – parsley family vegetables: carrot, parsnip, parsley, celery, others related: hemlock, cow parsnip Vegetable Classification
Dicot vegetables: Convolvulaceae – morning glory family vegetables: sweet potato related: flowering morning glory, bindweed, dodder
Vegetable Classification
Dicot vegetables: Solanaceae – nightshade family vegetables: potato, tomato, pepper, eggplant, others related: nightshade, jimson- weed, physalis Vegetable Classification
Dicot vegetables: Asteraceae (Compositae) – composite (aster) family vegetables: lettuce, chicory, endive, salsify, artichoke, others related: dandelion, thistle, daisy, ragweed, sunflower Botanical Classification
Additional Taxonomy (most useful) - Family Genus Species Cultivar
Botanical Classification
Family: An assemblage of genera that closely or uniformly resemble one another in general appearance and technical characters Botanical Classification
Genus: Identifies a more or less closely related and definable group of plants that may include one or more species.
The species within a genus are usually structurally or phylogenetically related. Botanical Classification
Species: A group of similar organisms capable of interbreeding and are distinctly different in morphological or other characteristics from other species in the same genus. Botanical Classification
Variety: A subdivision of a species consisting of a population with morphological characteristics distinct from other species forms.
(considered a naturally occurring taxonomic division) Botanical Classification
Cultivar (cultivated variety): Denotes certain cultivated plants that are alike in most important aspects of growth but are clearly distinguishable from others by one or more definite characteristics. Botanical Classification
Clone: Identifies material derived from a single individual and maintained by vegetative propagation. (genetically identical)
Line: A uniform sexually reproduced population, usually self- pollinated, that is seed propagated and maintained to the desired standard of uniformity by selection. (genetically similar) Botanical Classification
Strain: A term used to identify plants of a given cultivar that possess similar characteristics but differ in some minor feature or quality Botanical Classification
Additional Taxonomy - Family Genus Species Cultivar
Complete Latin Binomial includes the name of the individual who first described the species.
Botanical Classification
Family: Brassicaceae (Cruciferae) Genus: Brassica Species: oleracea Group/Variety: capitata Cultivar: ‘Golden Acre’ Strain: ‘Golden Acre YR’
Complete Latin Binomial - cabbage Brassica oleracea L. var. capitata L. cv. Golden Acre YR
Commercially: Brassica oleracea cv. Golden Acre YR
Botanical Classification
Growth and Development
Growth stages:
Germination Establishment Vegetative growth Reproductive growth Ripening and senescence Growth and Development
Growth stages:
Germination - the beginning or resumption of growth by a spore, seed, bud, or other structure
Growth Stages
Germination
Process: Water inbibition Seed coat softening Expansion of hypocotyl and root Emergence
Growth Stages
Germination
Characteristics: Critical for plant productivity Disease susceptibility Requires near-ideal conditions Growth Stages
Establishment
Process: Early shoot growth Root elongation Growth Stages
Establishment
Characteristics: Environmental sensitivity Determines future growth rate and potential Growth Stages
Vegetative Growth
Process: Increase root mass and rooting depth Rapid leaf area increase Increase in stem and leaf mass Large leaves and succulent growth produced Factors Affecting Growth
Energy Production and Storage
Photosynthesis – CO2 + H2O light sugars (C6H12O6) ↓ Transport ↓ Respiration (energy use), structural and chemical synthesis
Net photosynthesis = photosynthesis - respiration xx x
x Growth Stages
Vegetative Growth
Characteristics: Period of rapid cell growth and expansion High rate of photosynthesis Determines reproductive potential Heavy use of water and nutrients Growth Stages
Reproductive Growth
The transition from vegetative to reproductive growth is usually marked by flowering. Leaves receive environmental stimulus for the proper timing of flowering. Growth Stages
Reproductive Growth
Process: Organ differentiation Slowing or cessation of leaf expansion Flower production Fertilization and embryo growth Fruit growth Parallel storage organ growth in some species Growth Stages
Reproductive Growth
Characteristics: Period of heavy fiber production Sensitivity for yield and quality Can be influenced by many factors including temperature, daylength, etc Growth Stages
Senescence
The latter part of plant development which leads from maturity to the ultimate complete loss of organization and function. Growth Stages
Senescence
Process: Cessation of new growth Loss of leaf area Increased susceptibility to opportunists Ripening of fruit and seed Plant death Growth Stages
Senescence
Characteristics: Yield not influenced by external factors Usually not reversible Management inputs have little impact Critical period for fruit and seed quality Factors Affecting Growth
Light Factors
Intensity – most vegetable require full sun equivalent
Duration – requirements vary by species, fruiting vegetables need more
Wavelength – wavelengths 400-450, 650-700 best for photosynthesis Factors Affecting Growth
Temperature
Optimum is the range for a crop that allows for maximum photosynthesis and normal respiration
Optimum differs by species
Optimum may change during growth period
Diurnal fluctuations as important as average Factors Affecting Growth
Temperature
Temperatures above optimum slow growth and reduce quality (pungency, fibrousness) rapid respiration, stomate closure, reduced photosynthesis
Temperatures below optimum slow growth and affect quality by reducing sugar production and storage reduce photosynthesis, transport, and respiration Factors Affecting Growth
Heat Units (Growing Degree Days, GDD)
Degree days above a crop baseline
Average of daily high and low - baseline (onions 35, tomatoes 50, eggplants 60)
Used for determining suitable environments and predicting harvest dates
Doesn’t account for early soil temps, based on daily average not actual temps, doesn’t account for higher than optimum max temps Factors Affecting Growth
Water availability
Need constant water supply periodic stress reduces growth and quality stomate closure, reduction in cell division
Soil moisture principles Water holding capacity Field capacity Optimum minimal soil moisture Permanent wilting point
Factors Affecting Growth
Fertility
Require nutrients at or near optimum sub or supra-optimum impacts yield and may severely impact quality
Seedlings need high levels of fertilizers
Seasonal applications beneficial to long-season crops
Can impact life expectancy, disease resistance, etc Factors Affecting Growth
Genetics and Physiology
Growth habit (determinate vs indeterminate)
Vernalization requirement (or problem)
Photoperiodism most crops are day neutral
Tolerance to environmental stresses
Factors Affecting Growth
Crop Management
Many growth factors can be managed Many stress and disease related problems can be ameliorated with proper management