The Turonian - Coniacian Boundary in the United States Western Interior
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Stratigraphy and Paleontology of Mid-Cretaceous Rocks in Minnesota and Contiguous Areas
Stratigraphy and Paleontology of Mid-Cretaceous Rocks in Minnesota and Contiguous Areas GEOLOGICAL SURVEY PROFESSIONAL PAPER 1253 Stratigraphy and Paleontology of Mid-Cretaceous Rocks in Minnesota and Contiguous Areas By WILLIAM A. COBBAN and E. A. MEREWETHER Molluscan Fossil Record from the Northeastern Part of the Upper Cretaceous Seaway, Western Interior By WILLIAM A. COBBAN Lower Upper Cretaceous Strata in Minnesota and Adjacent Areas-Time-Stratigraphic Correlations. and Structural Attitudes By E. A. M EREWETHER GEOLOGICAL SURVEY PROFESSIONAL PAPER 1 2 53 UNITED STATES GOVERNMENT PRINTING OFFICE, WASHINGTON 1983 UNITED STATES DEPARTMENT OF THE INTERIOR JAMES G. WATT, Secretary GEOLOGICAL SURVEY Dallas L. Peck, Director Library of Congress Cataloging in Publication Data Cobban, William Aubrey, 1916 Stratigraphy and paleontology of mid-Cretaceous rocks in Minnesota and contiguous areas. (Geological Survey Professional Paper 1253) Bibliography: 52 p. Supt. of Docs. no.: I 19.16 A. Molluscan fossil record from the northeastern part of the Upper Cretaceous seaway, Western Interior by William A. Cobban. B. Lower Upper Cretaceous strata in Minnesota and adjacent areas-time-stratigraphic correlations and structural attitudes by E. A. Merewether. I. Mollusks, Fossil-Middle West. 2. Geology, Stratigraphic-Cretaceous. 3. Geology-Middle West. 4. Paleontology-Cretaceous. 5. Paleontology-Middle West. I. Merewether, E. A. (Edward Allen), 1930. II. Title. III. Series. QE687.C6 551.7'7'09776 81--607803 AACR2 For sale by the Distribution Branch, U.S. -
30. Turonian–Santonian Benthic
Mascle, J., Lohmann, G.P., and Moullade, M. (Eds.), 1998 Proceedings of the Ocean Drilling Program, Scientific Results, Vol. 159 30. TURONIAN–SANTONIAN BENTHIC FORAMINIFER ASSEMBLAGES FROM SITE 959D (CÔTE D’IVOIRE-GHANA TRANSFORM MARGIN, EQUATORIAL ATLANTIC): INDICATION OF A LATE CRETACEOUS OXYGEN MINIMUM ZONE1 Ann E.L. Holbourn2,3 and Wolfgang Kuhnt2 ABSTRACT Turonian–Santonian organic-rich fissile black claystones with laminated intervals from Hole 959D on the Côte d’Ivoire- Ghana Transform Margin, drilled during Ocean Drilling Program Leg 159, contain benthic foraminifer assemblages dominated by buliminid associations. The lower Turonian assemblage from Core 159-959D-68R is strongly dominated by Bolivina anam- bra, Praebulimina sp. 1, Praebulimina sp. 2, Praebulimina sp. 3, and Gavelinella spp. The upper Turonian to lower Coniacian assemblage from Core 159-959D-67R displays low abundance and low diversity and consists mostly of organically cemented agglutinated taxa and/or some corroded tests of Lenticulina, Bolivina, Gyroidinoides ex gr. nitidus. The middle Coniacian to lower Santonian assemblage from Core 159-959D-66R and from the base of Core 159-959D-65R contains high numbers of Praebulimina robusta, Praebulimina fang, Neobulimina subregularis, and Buliminella cf. gabonica, but shows marked fluctua- tions in abundance and diversity, which appear to be related to changes in total organic carbon. The distinct composition of the two buliminid associations in Core 159-959D-68R and Core 159-959D-66R suggests that endemism was stronger during the early Turonian, when circulation was probably more restricted and connections between equatorial Atlantic basins were lim- ited. We interpret the late Coniacian–early Santonian depositional environment to be an oxygen minimum zone in a more open marine outer shelf or upper slope setting. -
Cenomanian Turonian Coniacian Santonian Campanian
walteri aff. aff. spp. spp. imperfectus spp. (prisms) Chronostratigraphy Offshore Norway sp. 1 Geologic Time Scale 2012 Zonation (Gradstein et al., 1999, and this study) Allomorphina halli / pyriformis Sigmoilina antiqua Textularia Gavelinella intermeda gracillima Valvulineria Bulbobaculites problematicus Caudammina ovuloides Nuttallinella florealis Stensioeina granulata polonica Inoceramus Rzehakina minima Rzehakina epigona Fenestrella bellii Gaudryina filiformis Trochamminoides Haplophragmoides Gavelinella usakensis Caudammina ovula Coarse agglutinated spp. LCO dubia Tritaxia Plectorecurvoides alternans Reussella szajnochae Recurvoides Hippocrepina depressa Psammosphaera sphaerical radiolarians Ma Age/Stage Lingulogavelinella jarzevae elegans Lt NCF19 Maastrichtian volutus LCO 70 NCF18 E szajnochae dubia Lt 75 LCO of NCF17 Campanian Deep Water M Agglutinated 80 Foraminifera E bellii NCF16 Lt Inoceramus LCO NCF15 85 Santonian M E polonica NCF14 Lt Coniacian M E Marginotruncana NCF13 90 Lt Turonian M E Dicarinella NCF12 95 Lt brittonensis M NCF11 Cenomanian delrioensis LCO NCF10 E antiqua NCF9 100 Figure 2.8c. Stratigraphic ranges of Upper Cretaceous benthic foraminifera, and miscellaneous index taxa, oshore mid-Norway, with the foraminiferal zonation established in this study. s.l. Chronostratigraphy Offshore Norway Geologic Time Scale 2012 Zonation (Gradstein et al., 1999, and this study) Abathomphalus mayaroensis Pseudotextularia elegans Hedbergella planispira Hedbergella hoelzi Praeglobotruncana delrioensis Praeglobotruncana stephani -
Upper Cenomanian •fi Lower Turonian (Cretaceous) Calcareous
Studia Universitatis Babeş-Bolyai, Geologia, 2010, 55 (1), 29 – 36 Upper Cenomanian – Lower Turonian (Cretaceous) calcareous algae from the Eastern Desert of Egypt: taxonomy and significance Ioan I. BUCUR1, Emad NAGM2 & Markus WILMSEN3 1Department of Geology, “Babeş-Bolyai” University, Kogălniceanu 1, 400084 Cluj Napoca, Romania 2Geology Department, Faculty of Science, Al-Azhar University, Egypt 3Senckenberg Naturhistorische Sammlungen Dresden, Museum für Mineralogie und Geologie, Sektion Paläozoologie, Königsbrücker Landstr. 159, D-01109 Dresden, Germany Received March 2010; accepted April 2010 Available online 27 April 2010 DOI: 10.5038/1937-8602.55.1.4 Abstract. An assemblage of calcareous algae (dasycladaleans and halimedaceans) is described from the Upper Cenomanian to Lower Turonian of the Galala and Maghra el Hadida formations (Wadi Araba, northern Eastern Desert, Egypt). The following taxa have been identified: Dissocladella sp., Neomeris mokragorensis RADOIČIĆ & SCHLAGINTWEIT, 2007, Salpingoporella milovanovici RADOIČIĆ, 1978, Trinocladus divnae RADOIČIĆ, 2006, Trinocladus cf. radoicicae ELLIOTT, 1968, and Halimeda cf. elliotti CONARD & RIOULT, 1977. Most of the species are recorded for the first time from Egypt. Three of the identified algae (T. divnae, S. milovanovici and H. elliotti) also occur in Cenomanian limestones of the Mirdita zone, Serbia, suggesting a trans-Tethyan distribution of these taxa during the early Late Cretaceous. The abundance and preservation of the algae suggest an autochthonous occurrence which can be used to characterize the depositional environment. The recorded calcareous algae as well as the sedimentologic and palaeontologic context of the Galala Formation support an open-lagoonal (non-restricted), warm-water setting. The Maghra el Hadida Formation was mainly deposited in a somewhat deeper, open shelf setting. -
40Ar/39Ar Dating of the Late Cretaceous Jonathan Gaylor
40Ar/39Ar Dating of the Late Cretaceous Jonathan Gaylor To cite this version: Jonathan Gaylor. 40Ar/39Ar Dating of the Late Cretaceous. Earth Sciences. Université Paris Sud - Paris XI, 2013. English. NNT : 2013PA112124. tel-01017165 HAL Id: tel-01017165 https://tel.archives-ouvertes.fr/tel-01017165 Submitted on 2 Jul 2014 HAL is a multi-disciplinary open access L’archive ouverte pluridisciplinaire HAL, est archive for the deposit and dissemination of sci- destinée au dépôt et à la diffusion de documents entific research documents, whether they are pub- scientifiques de niveau recherche, publiés ou non, lished or not. The documents may come from émanant des établissements d’enseignement et de teaching and research institutions in France or recherche français ou étrangers, des laboratoires abroad, or from public or private research centers. publics ou privés. Université Paris Sud 11 UFR des Sciences d’Orsay École Doctorale 534 MIPEGE, Laboratoire IDES Sciences de la Terre 40Ar/39Ar Dating of the Late Cretaceous Thèse de Doctorat Présentée et soutenue publiquement par Jonathan GAYLOR Le 11 juillet 2013 devant le jury compose de: Directeur de thèse: Xavier Quidelleur, Professeur, Université Paris Sud (France) Rapporteurs: Sarah Sherlock, Senoir Researcher, Open University (Grande-Bretagne) Bruno Galbrun, DR CNRS, Université Pierre et Marie Curie (France) Examinateurs: Klaudia Kuiper, Researcher, Vrije Universiteit Amsterdam (Pays-Bas) Maurice Pagel, Professeur, Université Paris Sud (France) - 2 - - 3 - Acknowledgements I would like to begin by thanking my supervisor Xavier Quidelleur without whom I would not have finished, with special thanks on the endless encouragement and patience, all the way through my PhD! Thank you all at GTSnext, especially to the directors Klaudia Kuiper, Jan Wijbrans and Frits Hilgen for creating such a great project. -
Sucesión De Amonitas Del Cretácico Superior (Cenomaniano – Coniaciano) De La Parte Más Alta De La Formación Hondita Y De L
Boletín de Geología Vol. 33, N° 1, enero-junio de 2011 SUCESIÓN DE AMONITAS DEL CRETÁCICO SUPERIOR (CENOMANIANO – CONIACIANO) DE LA PARTE MÁS ALTA DE LA FORMACIÓN HONDITA Y DE LA FORMACIÓN LOMA GORDA EN LA QUEBRADA BAMBUCÁ, AIPE - HUILA (COLOMBIA, S. A.) Pedro Patarroyo1 RESUMEN La sección de la quebrada Bambucá (Aipe - Huila) posee una buena exposición de los depósitos del Cretácico del Valle Superior del Magdalena. De la parte alta de la Formación Hondita se recolectaron Acanthoceras sp. y Rhynchostreon sp. del Cenomaniano superior. Dentro del segmento inferior de la Formación Loma Gorda se hallaron Choffaticeras (C.) cf. segne, Fagesia cf. catinus, Neoptychites cf. andinus, Mitonia gracilis, Morrowites sp., Nannovascoceras ? sp., Quitmaniceras ? sp., Benueites ? sp. junto con Mytiloides kossmati, M. goppelnensis y Anomia sp. del Turoniano inferior. Estratigráficamente arriba aparecen Paramammites ? sp., Hoplitoides sp. H. ingens, H. cf. lagiraldae, Codazziceras ospinae, Allocrioceras sp., que pueden estar representando entre el Turoniano inferior y medio. Para la parte alta de este segmento se encontraron Prionocycloceras sp. P. guayabanum, Reesidites subtuberculatum, Subprionotropis colombianus, Mytiloides scupini, Dydimotis sp., Gauthiericeras sp., Anagaudryceras ? sp., Eulophoceras jacobi, Paralenticeras sieversi, Hauericeras cf. madagascarensis, Peroniceras (P.) subtricarinatum, Forresteria (F.) sp., Barroisiceras cf. onilahyense, Ankinatsytes venezolanus que abarcan entre el Turoniano superior y el Coniaciano. Con base en la fauna colectada no es posible establecer los límites Cenomaniano/Turoniano y Turoniano/Coniaciano. Palabras clave: Amonitas, Cretácico superior, Valle Superior del Magdalena, Aipe-Huila-Colombia. UPPER CRETACEOUS AMMONITE SUCCESSION (CENOMANIAN – CONIACIAN) RELATED TO THE UPPER HONDITA AND LOMA GORDA FORMATIONS ALONG THE BAMBUCÁ CREEK, AIPE - HUILA (COLOMBIA, S.A.) ABSTRACT The Bambucá creek section (Aipe - Huila) shows a very good exposition of the Upper Magdalena Valley Cretaceous deposits. -
Upper Cretaceous Stratigraphy and Biostratigraphy of South-Central New Mexico Stephen C
New Mexico Geological Society Downloaded from: http://nmgs.nmt.edu/publications/guidebooks/63 Upper Cretaceous stratigraphy and biostratigraphy of south-central New Mexico Stephen C. Hook, Greg H. Mack, and William A. Cobban, 2012, pp. 413-430 in: Geology of the Warm Springs Region, Lucas, Spencer G.; McLemore, Virginia T.; Lueth, Virgil W.; Spielmann, Justin A.; Krainer, Karl, New Mexico Geological Society 63rd Annual Fall Field Conference Guidebook, 580 p. This is one of many related papers that were included in the 2012 NMGS Fall Field Conference Guidebook. Annual NMGS Fall Field Conference Guidebooks Every fall since 1950, the New Mexico Geological Society (NMGS) has held an annual Fall Field Conference that explores some region of New Mexico (or surrounding states). Always well attended, these conferences provide a guidebook to participants. Besides detailed road logs, the guidebooks contain many well written, edited, and peer-reviewed geoscience papers. These books have set the national standard for geologic guidebooks and are an essential geologic reference for anyone working in or around New Mexico. Free Downloads NMGS has decided to make peer-reviewed papers from our Fall Field Conference guidebooks available for free download. Non-members will have access to guidebook papers two years after publication. Members have access to all papers. This is in keeping with our mission of promoting interest, research, and cooperation regarding geology in New Mexico. However, guidebook sales represent a significant proportion of our operating budget. Therefore, only research papers are available for download. Road logs, mini-papers, maps, stratigraphic charts, and other selected content are available only in the printed guidebooks. -
Abhandlungen Der Geologischen Bundesanstalt in Wien
ZOBODAT - www.zobodat.at Zoologisch-Botanische Datenbank/Zoological-Botanical Database Digitale Literatur/Digital Literature Zeitschrift/Journal: Abhandlungen der Geologischen Bundesanstalt in Wien Jahr/Year: 2017 Band/Volume: 71 Autor(en)/Author(s): Summesberger Herbert, Kennedy William James, Skoumal Peter Artikel/Article: Early and middle Santonian Cephalopods from the Gosau Group (Upper Cretaceous, Austria) 1. Nautiloidea and non-heteromorph Ammonoidea 5-99 ABHANDLUNGEN DER GEOLOGISCHEN BUNDESANSTALT Abh. Geol. B.-A. ISSN 0378-0864 ISBN 978-3-85316-093-0 Band 71 S. 5–99 Wien, Oktober 2017 Early and middle Santonian Cephalopods from the Gosau Group (Upper Cretaceous, Austria) 1. Nautiloidea and non-heteromorph Ammonoidea HERBERT SUMMESBERGER1, WILLIAM J. KENNEDY2 & PETER SKOUMAL3 13 Text-Figures, 16 Tables, 23 Plates Österreichische Karte 1:50.000 BMN / UTM 76 Wiener Neustadt / NL 33-03-01 Wiener Neustadt revision 89 Angath / NL 32-03-18 Kundl description 90 Kufstein / NL 33-01-13 Kufstein Cephalopoda 95 Sankt Wolfgang im Salzkammergut / NL 33-01-17 Hallstatt Santonian 96 Bad Ischl / NL 33-01-11 Bad Ischl Gosau Group 98 Liezen / NL 33-02-07 Windischgarsten 99 Rottenmann / NL 33-02-08 Spital am Pyhrn 120 Wörgl Contents Abstract ................................................................................................. 5 Zusammenfassung ......................................................................................... 6 Introduction.............................................................................................. -
Redalyc.SUCESIÓN DE AMONITAS DEL CRETÁCICO SUPERIOR (CENOMANIANO – CONIACIANO) DE LA PARTE MÁS ALTA DE LA FORMACIÓN HONDIT
Boletín de Geología ISSN: 0120-0283 [email protected] Universidad Industrial de Santander Colombia Patarroyo, Pedro SUCESIÓN DE AMONITAS DEL CRETÁCICO SUPERIOR (CENOMANIANO – CONIACIANO) DE LA PARTE MÁS ALTA DE LA FORMACIÓN HONDITA Y DE LA FORMACIÓN LOMA GORDA EN LA QUEBRADA BAMBUCÁ, AIPE - HUILA (COLOMBIA, S. A.) Boletín de Geología, vol. 33, núm. 1, enero-junio, 2011, pp. 69-92 Universidad Industrial de Santander Bucaramanga, Colombia Disponible en: http://www.redalyc.org/articulo.oa?id=349632022005 Cómo citar el artículo Número completo Sistema de Información Científica Más información del artículo Red de Revistas Científicas de América Latina, el Caribe, España y Portugal Página de la revista en redalyc.org Proyecto académico sin fines de lucro, desarrollado bajo la iniciativa de acceso abierto Boletín de Geología Vol. 33, N° 1, enero-junio de 2011 SUCESIÓN DE AMONITAS DEL CRETÁCICO SUPERIOR (CENOMANIANO – CONIACIANO) DE LA PARTE MÁS ALTA DE LA FORMACIÓN HONDITA Y DE LA FORMACIÓN LOMA GORDA EN LA QUEBRADA BAMBUCÁ, AIPE - HUILA (COLOMBIA, S. A.) Pedro Patarroyo1 RESUMEN La sección de la quebrada Bambucá (Aipe - Huila) posee una buena exposición de los depósitos del Cretácico del Valle Superior del Magdalena. De la parte alta de la Formación Hondita se recolectaron Acanthoceras sp. y Rhynchostreon sp. del Cenomaniano superior. Dentro del segmento inferior de la Formación Loma Gorda se hallaron Choffaticeras (C.) cf. segne, Fagesia cf. catinus, Neoptychites cf. andinus, Mitonia gracilis, Morrowites sp., Nannovascoceras ? sp., Quitmaniceras ? sp., Benueites ? sp. junto con Mytiloides kossmati, M. goppelnensis y Anomia sp. del Turoniano inferior. Estratigráficamente arriba aparecen Paramammites ? sp., Hoplitoides sp. -
GEOLOGIC TIME SCALE V
GSA GEOLOGIC TIME SCALE v. 4.0 CENOZOIC MESOZOIC PALEOZOIC PRECAMBRIAN MAGNETIC MAGNETIC BDY. AGE POLARITY PICKS AGE POLARITY PICKS AGE PICKS AGE . N PERIOD EPOCH AGE PERIOD EPOCH AGE PERIOD EPOCH AGE EON ERA PERIOD AGES (Ma) (Ma) (Ma) (Ma) (Ma) (Ma) (Ma) HIST HIST. ANOM. (Ma) ANOM. CHRON. CHRO HOLOCENE 1 C1 QUATER- 0.01 30 C30 66.0 541 CALABRIAN NARY PLEISTOCENE* 1.8 31 C31 MAASTRICHTIAN 252 2 C2 GELASIAN 70 CHANGHSINGIAN EDIACARAN 2.6 Lopin- 254 32 C32 72.1 635 2A C2A PIACENZIAN WUCHIAPINGIAN PLIOCENE 3.6 gian 33 260 260 3 ZANCLEAN CAPITANIAN NEOPRO- 5 C3 CAMPANIAN Guada- 265 750 CRYOGENIAN 5.3 80 C33 WORDIAN TEROZOIC 3A MESSINIAN LATE lupian 269 C3A 83.6 ROADIAN 272 850 7.2 SANTONIAN 4 KUNGURIAN C4 86.3 279 TONIAN CONIACIAN 280 4A Cisura- C4A TORTONIAN 90 89.8 1000 1000 PERMIAN ARTINSKIAN 10 5 TURONIAN lian C5 93.9 290 SAKMARIAN STENIAN 11.6 CENOMANIAN 296 SERRAVALLIAN 34 C34 ASSELIAN 299 5A 100 100 300 GZHELIAN 1200 C5A 13.8 LATE 304 KASIMOVIAN 307 1250 MESOPRO- 15 LANGHIAN ECTASIAN 5B C5B ALBIAN MIDDLE MOSCOVIAN 16.0 TEROZOIC 5C C5C 110 VANIAN 315 PENNSYL- 1400 EARLY 5D C5D MIOCENE 113 320 BASHKIRIAN 323 5E C5E NEOGENE BURDIGALIAN SERPUKHOVIAN 1500 CALYMMIAN 6 C6 APTIAN LATE 20 120 331 6A C6A 20.4 EARLY 1600 M0r 126 6B C6B AQUITANIAN M1 340 MIDDLE VISEAN MISSIS- M3 BARREMIAN SIPPIAN STATHERIAN C6C 23.0 6C 130 M5 CRETACEOUS 131 347 1750 HAUTERIVIAN 7 C7 CARBONIFEROUS EARLY TOURNAISIAN 1800 M10 134 25 7A C7A 359 8 C8 CHATTIAN VALANGINIAN M12 360 140 M14 139 FAMENNIAN OROSIRIAN 9 C9 M16 28.1 M18 BERRIASIAN 2000 PROTEROZOIC 10 C10 LATE -
Report 2015–1087
Chronostratigraphic Cross Section of Cretaceous Formations in Western Montana, Western Wyoming, Eastern Utah, Northeastern Arizona, and Northwestern New Mexico, U.S.A. Pamphlet to accompany Open-File Report 2015–1087 U.S. Department of the Interior U.S. Geological Survey Chronostratigraphic Cross Section of Cretaceous Formations in Western Montana, Western Wyoming, Eastern Utah, Northeastern Arizona, and Northwestern New Mexico, U.S.A. By E.A. Merewether and K.C. McKinney Pamphlet to accompany Open-File Report 2015–1087 U.S. Department of the Interior U.S. Geological Survey U.S. Department of the Interior SALLY JEWELL, Secretary U.S. Geological Survey Suzette M. Kimball, Acting Director U.S. Geological Survey, Reston, Virginia: 2015 For more information on the USGS—the Federal source for science about the Earth, its natural and living resources, natural hazards, and the environment—visit http://www.usgs.gov or call 1–888–ASK–USGS. For an overview of USGS information products, including maps, imagery, and publications, visit http://www.usgs.gov/pubprod/. Any use of trade, firm, or product names is for descriptive purposes only and does not imply endorsement by the U.S. Government. Although this information product, for the most part, is in the public domain, it also may contain copyrighted materials as noted in the text. Permission to reproduce copyrighted items must be secured from the copyright owner. Suggested citation: Merewether, E.A., and McKinney, K.C., 2015, Chronostratigraphic cross section of Cretaceous formations in western Montana, western Wyoming, eastern Utah, northeastern Arizona, and northwestern New Mexico, U.S.A.: U.S. Geological Survey Open-File Report 2015–1087, 10 p. -
A New Middle Jurassic Diplodocoid Suggests an Earlier Dispersal and Diversification of Sauropod Dinosaurs
ARTICLE DOI: 10.1038/s41467-018-05128-1 OPEN A new Middle Jurassic diplodocoid suggests an earlier dispersal and diversification of sauropod dinosaurs Xing Xu1, Paul Upchurch2, Philip D. Mannion 3, Paul M. Barrett 4, Omar R. Regalado-Fernandez 2, Jinyou Mo5, Jinfu Ma6 & Hongan Liu7 1234567890():,; The fragmentation of the supercontinent Pangaea has been suggested to have had a profound impact on Mesozoic terrestrial vertebrate distributions. One current paradigm is that geo- graphic isolation produced an endemic biota in East Asia during the Jurassic, while simul- taneously preventing diplodocoid sauropod dinosaurs and several other tetrapod groups from reaching this region. Here we report the discovery of the earliest diplodocoid, and the first from East Asia, to our knowledge, based on fossil material comprising multiple individuals and most parts of the skeleton of an early Middle Jurassic dicraeosaurid. The new discovery challenges conventional biogeographical ideas, and suggests that dispersal into East Asia occurred much earlier than expected. Moreover, the age of this new taxon indicates that many advanced sauropod lineages originated at least 15 million years earlier than previously realised, achieving a global distribution while Pangaea was still a coherent landmass. 1 Key Laboratory of Evolutionary Systematics of Vertebrates, Institute of Vertebrate Paleontology & Paleoanthropology, Chinese Academy of Sciences, 100044 Beijing, China. 2 Department of Earth Sciences, University College London, Gower Street, London WC1E 6BT, UK. 3 Department of Earth Science and Engineering, Imperial College London, South Kensington Campus, London SW7 2AZ, UK. 4 Department of Earth Sciences, Natural History Museum, Cromwell Road, London SW7 5BD, UK. 5 Natural History Museum of Guangxi, 530012 Nanning, Guangxi, China.