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Studies on Continental Late Triassic Tetrapod Biochronology. I. the Type Locality of Saturnalia Tupiniquim and the Faunal Succession in South Brazil
Journal of South American Earth Sciences 19 (2005) 205–218 www.elsevier.com/locate/jsames Studies on continental Late Triassic tetrapod biochronology. I. The type locality of Saturnalia tupiniquim and the faunal succession in south Brazil Max Cardoso Langer* Departamento de Biologia, FFCLRP, Universidade de Sa˜o Paulo (USP), Av. Bandeirantes 3900, 14040-901 Ribeira˜o Preto, SP, Brazil Received 1 November 2003; accepted 1 January 2005 Abstract Late Triassic deposits of the Parana´ Basin, Rio Grande do Sul, Brazil, encompass a single third-order, tetrapod-bearing sedimentary sequence that includes parts of the Alemoa Member (Santa Maria Formation) and the Caturrita Formation. A rich, diverse succession of terrestrial tetrapod communities is recorded in these sediments, which can be divided into at least three faunal associations. The stem- sauropodomorph Saturnalia tupiniquim was collected in the locality known as ‘Waldsanga’ near the city of Santa Maria. In that area, the deposits of the Alemoa Member yield the ‘Alemoa local fauna,’ which typifies the first association; includes the rhynchosaur Hyperodapedon, aetosaurs, and basal dinosaurs; and is coeval with the lower fauna of the Ischigualasto Formation, Bermejo Basin, NW Argentina. The second association is recorded in deposits of both the Alemoa Member and the Caturrita Formation, characterized by the rhynchosaur ‘Scaphonyx’ sulcognathus and the cynodont Exaeretodon, and correlated with the upper fauna of the Ischigualasto Formation. Various isolated outcrops of the Caturrita Formation yield tetrapod fossils that correspond to post-Ischigualastian faunas but might not belong to a single faunal association. The record of the dicynodont Jachaleria suggests correlations with the lower part of the Los Colorados Formation, NW Argentina, whereas remains of derived tritheledontid cynodonts indicate younger ages. -
Tiago Rodrigues Simões
Diapsid Phylogeny and the Origin and Early Evolution of Squamates by Tiago Rodrigues Simões A thesis submitted in partial fulfillment of the requirements for the degree of Doctor of Philosophy in SYSTEMATICS AND EVOLUTION Department of Biological Sciences University of Alberta © Tiago Rodrigues Simões, 2018 ABSTRACT Squamate reptiles comprise over 10,000 living species and hundreds of fossil species of lizards, snakes and amphisbaenians, with their origins dating back at least as far back as the Middle Jurassic. Despite this enormous diversity and a long evolutionary history, numerous fundamental questions remain to be answered regarding the early evolution and origin of this major clade of tetrapods. Such long-standing issues include identifying the oldest fossil squamate, when exactly did squamates originate, and why morphological and molecular analyses of squamate evolution have strong disagreements on fundamental aspects of the squamate tree of life. Additionally, despite much debate, there is no existing consensus over the composition of the Lepidosauromorpha (the clade that includes squamates and their sister taxon, the Rhynchocephalia), making the squamate origin problem part of a broader and more complex reptile phylogeny issue. In this thesis, I provide a series of taxonomic, phylogenetic, biogeographic and morpho-functional contributions to shed light on these problems. I describe a new taxon that overwhelms previous hypothesis of iguanian biogeography and evolution in Gondwana (Gueragama sulamericana). I re-describe and assess the functional morphology of some of the oldest known articulated lizards in the world (Eichstaettisaurus schroederi and Ardeosaurus digitatellus), providing clues to the ancestry of geckoes, and the early evolution of their scansorial behaviour. -
THE GENERA of REPTILES. By
T he Genera of Re pt il e s. By BARON FRANCIS NOPCSA (Budapest). (Eingelangt am 11. Mai 1927.) Among all the Paleontologists living none has dealt with the recent and fossil reptiles with a wider grasp than Prof. L. D o l l o and at his anniversary it seems quite appropriate to review their whole array. The classification used in the following enumeration of all genera of reptiles is much the same as in my hook „Die Familien der Reptilien“; at this instance however an effort has been made to give a precise definition of every systematic unit. Alteration of the classification became necessary among the Dino- cephalians, the Nothosaurians, the Lacertilians (called here Sauroidea), the Coelurosauroidea and the Crocodilia. The new classification of the Sauroidea is intermediate between the classifications proposed by B o u l e n g e r and C a m p . Naturally as a rule only those fossil genera are referred to, that are more than simple catalogue numbers that facilitate the finding of the respective piece in a collection; indeterminable problematic genera of small interest have mostly been omitted. Extinct units and genera are marked in the lists with a cross ("j*). Difficulties of classification have been encountered in the Lacertilia and Ophidia, for in the new system the conventional families of recent Lizards and Serpents has been given subfamily rank and conse quently the different recent subfamilies had to be dropped. Instead of these subfamilies the minor units were separated by greek letters. — The tedious and complicated revision of the genera of recent Lizards and Snakes was done by Prof. -
A New Sphenodontian (Lepidosauria: Rhynchocephalia)
View metadata, citation and similar papersDownloaded at core.ac.uk from http://rspb.royalsocietypublishing.org/ on February 13, 2017 brought to you by CORE provided by CONICET Digital A new sphenodontian (Lepidosauria: Rhynchocephalia) from the Late Triassic of Argentina and the early origin of the herbivore opisthodontians rspb.royalsocietypublishing.org Ricardo N. Martı´nez1, Cecilia Apaldetti1,2, Carina E. Colombi1,2, Angel Praderio1, Eliana Fernandez1,2, Paula Santi Malnis1,2, 1,2 1 1 Research Gustavo A. Correa , Diego Abelin and Oscar Alcober 1Instituto y Museo de Ciencias Naturales, Universidad Nacional de San Juan, Avenida Espan˜a 400 Norte, Cite this article: Martı´nez RN, Apaldetti C, 5400 San Juan, Argentina Colombi CE, Praderio A, Fernandez E, Malnis 2Consejo Nacional de Investigaciones Cientı´ficas y Te´cnicas, CONICET, Buenos Aires, Argentina PS, Correa GA, Abelin D, Alcober O. 2013 A new sphenodontian (Lepidosauria: Rhyncho- Sphenodontians were a successful group of rhynchocephalian reptiles that dominated the fossil record of Lepidosauria during the Triassic and Jurassic. cephalia) from the Late Triassic of Argentina Although evidence of extinction is seen at the end of the Laurasian Early and the early origin of the herbivore Cretaceous, they appeared to remain numerically abundant in South America opisthodontians. Proc R Soc B 280: 20132057. until the end of the period. Most of the known Late Cretaceous record in http://dx.doi.org/10.1098/rspb.2013.2057 South America is composed of opisthodontians, the herbivorous branch of Sphenodontia, whose oldest members were until recently reported to be from the Kimmeridgian–Tithonian (Late Jurassic). Here, we report a new sphenodontian, Sphenotitan leyesi gen. -
Callibrachion and Datheosaurus, Two Historical and Previously Mistaken Basal Caseasaurian Synapsids from Europe
Callibrachion and Datheosaurus, two historical and previously mistaken basal caseasaurian synapsids from Europe FREDERIK SPINDLER, JOCELYN FALCONNET, and JÖRG FRÖBISCH Spindler, F., Falconnet, J., and Fröbisch, J. 2016. Callibrachion and Datheosaurus, two historical and previously mis- taken basal caseasaurian synapsids from Europe. Acta Palaeontologica Polonica 61 (3): 597–616. This study represents a re-investigation of two historical fossil discoveries, Callibrachion gaudryi (Artinskian of France) and Datheosaurus macrourus (Gzhelian of Poland), that were originally classified as haptodontine-grade sphenaco- dontians and have been lately treated as nomina dubia. Both taxa are here identified as basal caseasaurs based on their overall proportions as well as dental and osteological characteristics that differentiate them from any other major syn- apsid subclade. As a result of poor preservation, no distinct autapomorphies can be recognized. However, our detailed investigations of the virtually complete skeletons in the light of recent progress in basal synapsid research allow a novel interpretation of their phylogenetic positions. Datheosaurus might represent an eothyridid or basal caseid. Callibrachion shares some similarities with the more derived North American genus Casea. These new observations on Datheosaurus and Callibrachion provide new insights into the early diversification of caseasaurs, reflecting an evolutionary stage that lacks spatulate teeth and broadened phalanges that are typical for other caseid species. Along with Eocasea, the former ghost lineage to the Late Pennsylvanian origin of Caseasauria is further closed. For the first time, the presence of basal caseasaurs in Europe is documented. Key words: Synapsida, Caseasauria, Carboniferous, Permian, Autun Basin, France, Intra-Sudetic Basin, Poland. Frederik Spindler [[email protected]], Dinosaurier-Park Altmühltal, Dinopark 1, 85095 Denkendorf, Germany. -
The Palaeontology Newsletter
The Palaeontology Newsletter Contents 89 Editorial 2 Association Business 3 Association Meetings 28 Palaeontology at EGU 33 News 34 From our correspondents Legends of Rock: Murchison 43 Behind the scenes at the Museum 46 Buffon’s Gang 51 R: Regression 58 PalAss press releases 68 Blaschka glass models 69 Sourcing the Ring Master 73 Underwhelming fossil of the month 75 Future meetings of other bodies 80 Meeting Reports 87 Obituaries Percy Milton Butler 100 Lennart Jeppsson 103 Grant and Bursary reports 106 Book Reviews 116 Careering off course! 121 Palaeontology vol 58 parts 3 & 4 123–124 Papers in Palaeontology, vol 1, part 1 125 Reminder: The deadline for copy for Issue no 90 is 12th October 2015. On the Web: <http://www.palass.org/> ISSN: 0954-9900 Newsletter 89 2 Editorial Here in London summer is under way, although it is difficult to tell just by looking out of the window; once you step outside, however, the high pollen counts are the main giveaway. It is largely the sort of ‘summer’ weather that confuses foreign tourists, makes a mud bath out of music festivals and stops play at Wimbledon. The perfect sort of weather to leave Britain behind and jet off to sunnier climes for a nice long field campaign: as I wistfully gaze at the grey skies I wish I were packing my notebook and field gear ready for departure. Those of you who are setting out for fieldwork during this Northern Hemisphere summer will probably be aware that it is not a walk in the park, as my initial rose-tinted memories lead me to believe. -
Dentary Morphological Variation in Clevosaurus Brasiliensis (Rhynchocephalia, Clevosauridae) from the Upper Triassic of Rio Grande Do Sul, Brazil
RESEARCH ARTICLE Dentary Morphological Variation in Clevosaurus brasiliensis (Rhynchocephalia, Clevosauridae) from the Upper Triassic of Rio Grande do Sul, Brazil Paula Rosario Romo de Vivar Martínez*, Marina Bento Soares Laboratório de Paleontologia de Vertebrados, Departamento de Paleontologia e Estratigrafia, Instituto de Geociências, Universidade Federal do Rio Grande do Sul, Porto Alegre, Rio Grande do Sul, Brazil * [email protected] Abstract Clevosaurus was a cosmopolitan rhynchocephalian genus, known from the Late Triassic to the Early Jurassic. In South America this genus is represented by C. brasiliensis, an impor- OPEN ACCESS tant component of the Linha São Luiz taphocoenosis, on the top of the Norian Santa Maria Citation: Romo de Vivar Martínez PR, Bento Soares 2 Sequence of Southern Brazil. The best preserved and most abundant bone elements of M (2015) Dentary Morphological Variation in C. brasiliensis are dentaries, in which variations of shape and size are observed. The aim of Clevosaurus brasiliensis (Rhynchocephalia, this study is to describe and evaluate the variation, using geometric morphometrics meth- Clevosauridae) from the Upper Triassic of Rio Grande do Sul, Brazil. PLoS ONE 10(3): e0119307. ods. Geometric morphometric analysis of 10 specimens highlights variations in relative size doi:10.1371/journal.pone.0119307 of the dentary. Most of the variation observed for PC1 (83.3%) is likely related to ontogeny, Academic Editor: David Carrier, University of Utah, and PC2 (10.0%) is likely related to taphonomic signatures. The development patterns ob- UNITED STATES served, such as the growth of the dentary, consists of differential growth in length between Received: February 24, 2014 the posterior portion of the dentary, that grows at a higher rate, regarding the anterior portion of the element. -
Review of Fossil Collections in Scotland Review of Fossil Collections in Scotland
Detail of the Upper Devonian fishHoloptychius from Dura Den, Fife. © Perth Museum & Art Gallery, Perth & Kinross Council Review of Fossil Collections in Scotland Review of Fossil Collections in Scotland Contents Introduction 3 Background 3 Aims of the Collections Review 4 Methodology 4 Terminology 5 Summary of fossil material 6 Influences on collections 14 Collections by region Aberdeen and North East 17 Elgin Museum (Moray Society) 18 Falconer Museum (Moray Council) 21 Stonehaven Tolbooth Museum 23 The Discovery Centre (Live Life Aberdeenshire) 24 Arbuthnot Museum (Live Life Aberdeenshire) 27 Zoology Museum (University of Aberdeen Museums) 28 Meston Science Building (University of Aberdeen Museums) 30 Blairs Museum 37 Highlands and Islands 38 Inverness Museum and Art Gallery (High Life Highland) 39 Nairn Museum 42 West Highland Museum (West Highland Museum Trust) 44 Brora Heritage Centre (Brora Heritage Trust) 45 Dunrobin Castle Museum 46 Timespan (Timespan Heritage and Arts Society) 48 Stromness Museum (Orkney Natural History Society) 50 Orkney Fossil and Heritage Centre 53 Shetland Museum and Archives (Shetland Amenity Trust) 56 Bute Museum (Bute Museum Trust) 58 Hugh Miller’s Birthplace Cottage and Museum (National Trust for Scotland) 59 Treasures of the Earth 62 Staffin Dinosaur Museum 63 Gairloch Museum (Gairloch & District Heritage Company Ltd) 65 Tayside, Central and Fife 66 Stirling Smith Art Gallery and Museum 67 Perth Museum and Art Gallery (Culture Perth and Kinross) 69 The McManus: Dundee’s Art Gallery and Museum (Leisure -
Universidade Federal Do Rio Grande Do Sul Instituto De Geociências
UNIVERSIDADE FEDERAL DO RIO GRANDE DO SUL INSTITUTO DE GEOCIÊNCIAS Bruno de Amorim Arantes NOVOS MATERIAIS DE RHYNCHOCEPHALIA DA FORMAÇÃO CATURRITA (TRIÁSSICO SUPERIOR) DO RIO GRANDE DO SUL, BRASIL: ESTUDO ANATÔMICO E IMPLICAÇÕES FILOGENÉTICAS. Porto Alegre 2011 Arantes, Bruno de Amorim Novos materiais de Rynchocephalia da Formação Caturrita (triássico superior) do Rio Grande do Sul, Brasil: estudo anatômico e implicações filogenéticas. / Bruno de Amorim Arantes. - Porto Alegre : IGEO/UFRGS, 2011. [92 f.] il. Dissertação (Mestrado). - Universidade Federal do Rio Grande do Sul. Instituto de Geociências. Programa de Pós-Graduação em Geociências. Porto Alegre, RS - BR, 2011. Orientação: Profª. Drª. Marina Bento Soares 1. Rhynchocephalia. 2. Clevosauridae. 3. Clevosaurus. 4. Formação Caturrita. 5. Triássico Superior. Título _____________________________ Catalogação na Publicação Biblioteca Geociências - UFRGS Miriam Alves CRB 10/1947 Bruno de Amorim Arantes NOVOS MATERIAIS DE RHYNCHOCEPHALIA DA FORMAÇÃO CATURRITA (TRIÁSSICO SUPERIOR) DO RIO GRANDE DO SUL, BRASIL: ESTUDO ANATÔMICO E IMPLICAÇÕES FILOGENÉTICAS. Dissertação de Mestrado apresentada ao Programa de Pós-Graduação em Geociências do Instituto de Geociências da Universidade Federal do Rio Grande do Sul, como requisito parcial para a obtenção do Título de Mestre em Geociências. Orientadora: Profa. Dra. Marina Bento Soares Porto Alegre 2011 AGRADECIMENTOS A minha família e amigos pelo apoio durante os estudos. A professora Marina Bento Soares, por toda a orientação e confiança em meu trabalho. Ao Dr. Juan Carlos Cisneros, pela ajuda na identificação e descrição das amostras e pelos artigos emprestados. Ao Dr. Victor Hugo Reynoso, por disponibilizar seus artigos para consulta. Dra. Ana Maria Ribeiro, pelo acesso à coleção de esfenodontes do Museu de Ciências Naturais da FZBRS. -
Universidade Federal Do Rio Grande Do Sul Instituto De Geociências Programa De Pós-Graduação Em Geociências
UNIVERSIDADE FEDERAL DO RIO GRANDE DO SUL INSTITUTO DE GEOCIÊNCIAS PROGRAMA DE PÓS-GRADUAÇÃO EM GEOCIÊNCIAS UM ESTUDO SOBRE A VARIAÇÃO MORFOLÓGICA NO DENTÁRIO DE CLEVOSAURUS BRASILIENSIS BONAPARTE E SUES, 2006 (RHYNCHOCEPHALIA, CLEVOSAURIDAE), DO TRIÁSSICO SUPERIOR DO RIO GRANDE DO SUL, BRASIL, UTILIZANDO A MORFOMETRIA GEOMÉTRICA. PAULA ROSARIO ROMO DE VIVAR MARTÍNEZ ORIENTADORA – Profa. Dra. Marina Bento Soares Porto Alegre, 2014 UNIVERSIDADE FEDERAL DO RIO GRANDE DO SUL INSTITUTO DE GEOCIÊNCIAS PROGRAMA DE PÓS-GRADUAÇÃO EM GEOCIÊNCIAS UM ESTUDO SOBRE A VARIAÇÃO MORFOLÓGICA NO DENTÁRIO DE CLEVOSAURUS BRASILIENSIS BONAPARTE E SUES, 2006 (RHYNCHOCEPHALIA, CLEVOSAURIDAE), DO TRIÁSSICO SUPERIOR DO RIO GRANDE DO SUL, BRASIL, UTILIZANDO A MORFOMETRIA GEOMÉTRICA. PAULA ROSARIO ROMO DE VIVAR MARTÍNEZ ORIENTADORA – Profa. Dra. Marina Bento Soares BANCA EXAMINADORA Profa. Dra. Annie Schmaltz Hsiou- Universidade de São Paulo, Faculdade de Filosofia Ciências e Letras Prof. Dr. Nilton Cáceres – Universidade Fedaral de Santa Maria, Departamento de Biologia, CCNE Prof. Dr. Richard A. Fariña - Facultad de Ciencias, Universidad de la República Dissertação de Mestrado apresentada como requisito parcial para a obtenção do Título de Mestre em Geociências Porto Alegre, 2014 AGRADECIMENTOS Nesta parte do caminho que está quase por terminar, achei muitas pessoas que jamais imaginei conhecer. Em grande parte estou muito agradecida a estas pessoas, porque elas me ajudaram de algum jeito a concluir este processo, sem também esquecer daqueles que desde muitos anos atrás sempre fizeram parte de minha vida, e que agora simplesmente estão longe em distância, mas eu sei que sempre estão perto de mim. Eu quero agradecer à minha orientadora Dra. Marina Bento Soares, pelo apoio dado nestes quase dois anos, a confiança depositada em minhas loucuras (pensando como primeira loucura de atravessar o continente para vir estudar). -
A New Rhynchocephalian from the Late Triassic of Southern Brazil Enhances Eusphenodontian Diversity
Journal of Systematic Palaeontology ISSN: 1477-2019 (Print) 1478-0941 (Online) Journal homepage: https://www.tandfonline.com/loi/tjsp20 A New Rhynchocephalian from the Late Triassic of Southern Brazil Enhances Eusphenodontian Diversity Paulo R. Romo de Vivar, Agustín G. Martinelli, Annie Schmaltz Hsiou & Marina Bento Soares To cite this article: Paulo R. Romo de Vivar, Agustín G. Martinelli, Annie Schmaltz Hsiou & Marina Bento Soares (2020): A New Rhynchocephalian from the Late Triassic of Southern Brazil Enhances Eusphenodontian Diversity, Journal of Systematic Palaeontology, DOI: 10.1080/14772019.2020.1732488 To link to this article: https://doi.org/10.1080/14772019.2020.1732488 View supplementary material Published online: 20 Mar 2020. Submit your article to this journal View related articles View Crossmark data Full Terms & Conditions of access and use can be found at https://www.tandfonline.com/action/journalInformation?journalCode=tjsp20 Journal of Systematic Palaeontology, 2020 http://dx.doi.org/10.1080/14772019.2020.1732488 A New Rhynchocephalian from the Late Triassic of Southern Brazil Enhances Eusphenodontian Diversity aà b c a,d Paulo R. Romo de Vivar , Agustın G. Martinelli , Annie Schmaltz Hsiou and Marina Bento Soares aPrograma de Pos-Graduac ¸ao~ em Geoci^encias, Instituto de Geoci^encias, Universidade Federal Do Rio Grande Do Sul, Avenida Bento Gonc¸alves, 9500 Agronomia, Porto Alegre, RS, Cep 91501-970, Brazil; bCONICET-Seccion Paleontologıa de Vertebrados, Museo Argentino de Ciencias Naturales ‘Bernardino Rivadavia’, Av. Angel Gallardo 470, Buenos Aires, C1405DJR, Argentina; cDepartamento de Biologia, Faculdade de Filosofia, Ci^encias e Letras de Ribeirao~ Preto, Universidade de Sao~ Paulo, Ribeirao~ Preto, Sao~ Paulo, Brazil; dDepartamento de Geologia e Paleontologia, Museu Nacional, Universidade Federal Do Rio De Janeiro, Quinta da Boa Vista s/n, Sao~ Cristovao,~ Rio De Janeiro, RJ, Cep 20940-040, Brazil (Received 4 September 2018; accepted 17 February 2020) We describe a new eusphenodontian, Lanceirosphenodon ferigoloi gen. -
Classification and Phylogeny of the Diapsid Reptiles
zoological Journal Ofthe Linnean Society (1985), 84: 97-164. With 17 figures Classification and phylogeny of the diapsid reptiles MICHAEL J. BENTON Department of zoology and University Museum, Parks Road, Oxford OX1 3PW, U.El. * Received June 1983, revised and accepted for publication March 1984 Reptiles with two temporal openings in the skull are generally divided into two groups-the Lepidosauria (lizards, snakes, Sphenodon, ‘eosuchians’) and the Archosauria (crocodiles, thecodontians, dinosaurs, pterosaurs). Recent suggestions that these two are not sister-groups are shown to be unproven, whereas there is strong evidence that they form a monophyletic group, the Diapsida, on the basis of several synapomorphies of living and fossil forms. A cladistic analysis of skull and skeletal characters of all described Permo-Triassic diapsid reptiles suggests some significant rearrangements to commonly held views. The genus Petrolacosaurus is the sister-group of all later diapsids which fall into two large groups-the Archosauromorpha (Pterosauria, Rhynchosauria, Prolacertiformes, Archosauria) and the Lepidosauromorpha (Younginiformes, Sphenodontia, Squamata). The pterosaurs are not archosaurs, but they are the sister-group of all other archosauromorphs. There is no close relationship betwcen rhynchosaurs and sphenodontids, nor between Prolacerta or ‘Tanystropheus and lizards. The terms ‘Eosuchia’, ‘Rhynchocephalia’ and ‘Protorosauria’ have become too wide in application and they are not used. A cladistic classification of the Diapsida is given, as well as a phylogenetic tree which uses cladistic and stratigraphic data. KEY WORDS:-Reptilia ~ Diapsida - taxonomy ~ classification - cladistics - evolution - Permian - Triassic. CONTENTS Introduction ................... 98 Historical survey .................. 99 Monophyly of the Diapsida ............... 101 Romer (1968). ................. 102 The three-taxon statements .............. 102 Lmtrup (1977) and Gardiner (1982) ...........