Mosses of the Mediterranean, an Annotated Checklist
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Cryptogamie, Bryologie, 2013, 34 (2): 99-283 © 2013 Adac. Tous droits réservés Mosses of the Mediterranean, an annotated checklist Rosa M. ROS a*, Vicente MAZIMPAKA b Usama ABOU-SALAMA c, Michele ALEFFI d, Thomas L. BLOCKEEL e, Montserrat BRUGUÉS f, Rosa Maria CROS f, Maria Giovanna DIA g, Gerard M. DIRKSE h, Isabel DRAPER b, Wagieh EL-SAADAWI c, Adnan ERDA∆ i, Anna GANEVA j, Rosalina GABRIEL k, Juan M. GONZÁLEZ-MANCEBO l, Camille GRANGER m, Ilana HERRNSTADT n, Vincent HUGONNOT o, Kamel KHALIL p, Harald KÜRSCHNER q, Anna LOSADA-LIMA l, Leena LUÍS r, Stefan MIFSUD s, Maria PRIVITERA t, Marta PUGLISI t, Marko SABOVLJEVI± u, Cecilia SÉRGIO r, Haanaa M. SHABBARA c, Manuela SIM-SIM r, André SOTIAUX v, Roberta TACCHI w, André VANDERPOORTEN v, Olaf WERNER a In Memoriam to A.J.E. SMITH aUniversidad de Murcia, Facultad de Biología, Departamento de Biología Vegetal, Campus de Espinardo, 30100-Murcia, Spain. E-mail: [email protected], [email protected] bUniversidad Autónoma de Madrid, Facultad de Ciencias, Departamento de Biología, Ctra. de Colmenar Viejo Km 15, 28049 Madrid, Spain. E-mail: [email protected], [email protected] cBotany Department, Faculty of Science, Ain Shams University, Abbasia, Cairo, Egypt. E-mail: [email protected], [email protected], [email protected] dScuola di Scienze Ambientali, Laboratorio di Briologia, Università degli Studi di Camerino, Via Pontoni, 5 - 62032 Camerino (MC), Italy. E-mail: [email protected] eAshfurlong Close, Dore, Sheffield S17 3NN, United Kingdom. E-mail: [email protected] fDepartament de Biologia Animal, Vegetal i Ecologia, Unitat de Botànica, Universitat Autònoma de Barcelona, 08193 Bellaterra, Barcelona, Spain. E-mail: [email protected], [email protected] gDipartimento di Biologia Ambientale e Biodiversità, Università degli Studi di Palermo, Via Archirafi, 38,90133 Palermo, Italy. E-mail: [email protected] hNatuurmuseum Nijmegen, Gerard Noodtstraat 121, NL-6155 ST Nijmegen, The Netherlands. E-mail: [email protected] iAdnan Menderes University, Faculty of Arts & Sciences, Biology Department, 09010, Aydin,Turkey. E-mail: [email protected] jInstitute of Biodiversity and Ecosystem Research, 23 Acad. G. Bonchev Str., 1113 Sofia, Bulgaria. E-mail: [email protected] * Corresponding author: [email protected] doi/ 10.782/cryb.v34.iss2.2013.99 100 R.M. Ros, V. Mazimpaka et al. kDepartamento de Ciências Agrárias, Azorean Biodiversity Group- CITAA, Universidade dos Açores, P-9702 Angra do Heroísmo Codex, Açores, Portugal. E-mail: [email protected] lUniversidad de La Laguna, Departamento de Biología Vegetal (Botánica), Calle Astrofísico Francisco Sánchez s/n, 38071-La Laguna, Tenerife, Canary Islands, Spain. E-mail: [email protected], [email protected] m24, rue Camille Roy, 69007 Lyon, France. E-mail: [email protected] nThe Hebrew University of Jerusalem, Israel. E-mail: [email protected] oConservatoire botanique national du Massif central, le bourg, 43270 Chavaniac Lafayette, France. E-mail: [email protected] pEcology and Forestry Department, Faculty of Agriculture, University of Aleppo, Aleppo, Syria. E-mail: [email protected] qInstitut für Biologie der Freien Universität Berlin, Systematische Botanik und Pflanzengeographie, Altensteinstraße 6, D-14195 Berlin, Germany. E-mail: [email protected] rUniversidade de Lisboa, Faculdade de Ciências de Lisboa, DBV/Centro de Biologia Ambiental, Campo Grande, 1749-016 Lisboa, Portugal; Museu Nacional de História Natural e da Ciência, Jardim Botânico, Rua da Escola Politécnica, 58, 1250-102 Lisboa, Portugal. E-mail: [email protected], [email protected], [email protected] s123 Lampuka Str, Paola, Malta. E-mail: [email protected] tDipartimento di Scienze Biologiche, Geologiche e Ambientali, Università degli Studi di Catania, via A. Longo, 19, 95125 Catania, Italy. E-mail: [email protected] uInstitute of Botany and Botanical Garden, Faculty of Biology, University of Belgrade, Takovska 43, 11000 Belgrade, Serbia. E-mail: [email protected] vUniversité de Liège, Institut de Botanique, B22 Sart Tilman, B-4000 Liège1, Belgium. E-mail: [email protected], [email protected] wUniversità di Camerino, Orto botanico “Carmela Cortini”, Viale Oberdan, 62032 Camerino (MC), Italy. E-mail: [email protected] Abstract – The names of all mosses published up to the end of August 2011 in the countries of the Mediterranean basin, the Macaronesian Islands and Bulgaria are compiled in an annotated checklist. The list comprises accepted names and synonyms, and provides explanatory annotations for ambiguous and disputed names. Literature references supporting the reports in each individual area are given only for taxa reported once or in a single locality. A total of 1168 accepted species and 81 infraspecific taxa are reported from the whole area. Mosses / distribution / nomenclature / checklist / Mediterranean Mosses of the Mediterranean, an annotated checklist 101 INTRODUCTION In the course of the meeting held in Pisa in autumn 2007, the year which saw the publication of the checklist of Mediterranean liverworts and hornworts (Ros et al., 2007a), the OPTIMA Bryophyte Commission decided to continue with the second step of the project approved in Palermo six years previously, i.e. the preparation of a checklist of Mediterranean mosses. The reasons for this were exactly the same as those concerning the liverworts and hornworts: the lack of a critical list of taxa published in the Mediterranean region to date, and the need to fill this gap. There had been some important publications at national and insular level, including both floras (Cortini Pedrotti, 2001a, 2006; Casas et al., 2003, 2006; Heyn & Herrnstadt, 2004; Guerra et al., 2006, 2010c; Brugués et al., 2007c; Kürschner & Frey, 2011) and lists (Dia et al.,1987; Frey & Kürschner, 1991; Düll et al., 1999; El-Saadawi et al., 1999; Nizam & Al-Araj, 1999; Ros et al., 1999; Sabovljevi† & Stevanovi†, 1999; Martinªiª, 2003; Sérgio & Carvalho, 2003; Losada-Lima et al., 2004, 2010; Casas, 2005; Cekova, 2005; Gabriel et al., 2005; Kürschner & Erda∑, 2005; Natcheva & Ganeva, 2005; Colacino & Sabovljevi†, 2006; Dragi†evi† & Velji†, 2006; Sabovljevi†, 2006; Sérgio et al., 2006c, Sotiaux et al., 2007; Aleffi et al., 2008, Cros et al., 2008; Sabovljevi† et al., 2008a; Sérgio et al., 2008; González-Mancebo et al., 2008a; Mifsud, 2012), and at continental level a European bryophyte flora (Frey et al., 2006) and a checklist of the European mosses had appeared (Hill et al., 2006). However, none of these publications covered the whole Mediterranean basin, and it seemed expedient to produce a complete floristic and taxonomic update of the mosses of the entire Mediterranean region in the form of a critical checklist, incorporating currently accepted names and the synonyms used in this region in the past. The methods, criteria and processes were the same as those used for the checklist of liverworts and hornworts (Ros et al., 2007a). The authors responsible for individual countries and islands would prepare a list of accepted names and synonyms, and subsequently Rosa María Ros (Secretary of the OPTIMA Bryophyte Commission) and Vicente Mazimpaka (member of the Commission) would integrate all the checklists into a master list including accepted names and synonyms. For this new checklist it was decided that literature references would be included for each country and island, but only for the taxa reported once or in a single locality. After an interval of five years, we now present the resulting checklist, incorporating the names compiled at the beginning of the project, together with those published during the intervening years. The numerous scattered bibliographic sources, the large amount of data accumulated during three centuries of taxonomic and floristic work on Mediterranean mosses, and the difficulties derived from working in remote locations have sometimes been an obstacle, but the conviction of providing a useful tool to colleagues and to all those interested in the bryophytes of the Mediterranean region has been a great stimulus. Although a great effort has been made to trace and check all the names included in the list, it is unrealistic to expect a perfect result considering the scope and complexity of the work involved. Therefore the authors would appreciate any comments on errors or omissions observed in the lists in order to improve future revisions. 102 R.M. Ros, V. Mazimpaka et al. METHODOLOGY Criteria for the list of accepted names The list includes published names and records up to the end of August 2011. The checklist by Hill et al. (2006) has been the basic reference work for accepted names. As this is limited to Europe and Macaronesia, additional reference works for the territories outside this area have been followed, namely Ros et al. (1999) and El-Saadawi et al. (1999) for North Africa, Heyn & Herrnstadt (2004), Frey & Kürschner (1991) and Kürschner & Frey (2011) for the Near East and Kürschner & Erda∑ (2005) for Turkey. We have made some exceptions to these lists in the light of more recently published taxonomic papers or other criteria, and these are documented in the annotations. In cases where infraspecific taxa have not been distinguished in the literature for a given territory, then the relevant occurrences have been indicated in the checklist at species level. For the most part, however, we have been restrictive in the inclusion of infraspecific taxa. Thus, if the majority of recent authors have not recognised a taxon, then we have also omitted it here. We have not recognised the category of “form”. Taxa whose presence in a given area is doubtful have been excluded at the discretion of the author responsible for that area. We also have omitted names previously rejected in published papers, but explanatory annotations have been included for these rejections. Doubtful taxa have not been included in the general list but are mentioned in annotations. The names of the accepted taxa and their occurrences in individual territories are presented in tabular format. Taxa are arranged in alphabetical order of genera and species mainly following the classification of Bryophyta proposed by Goffinet et al.