THE JEPSON GLOBE A Newsletter from the Friends of The Jepson Herbarium

VOLUME 18 NUMBER 3 January 2008

Director’s Column - A Jepson by Brent D. Mishler Herbarium Workshop Recent Herbaria Retiree’s Text and Photos by Joan Eiger Continue Research Efforts Gottlieb You know you’re in Dr. Alan R. Smith and Dr. heaven when you land in New Zealand Barbara Ertter, two of the herbaria’s and see silver fern logos on everything dedicated curators, have recently re- from sport shirts to the sides of inter- tired. Fortunately, each will remain ac- island ferries. This global hot spot for tive in their research endeavors. Dr. ferns is home to about 200 species scat- Smith came to Berkeley in 1969 as a Mehrabadi, Share Doust, Doran, Mishler, Rahmani, and Tahbaz tered through coastal, forest, low mon- research botanist specializing in Pteri- tane, and even alpine habitats. About dophytes. He was curator of this major Visiting Iranian officials seek to 40% are endemic – a result of the long group and also worked closely with geographic isolation of this 3-island graduate students, leading monthly enhance scientific exchange with UC botanists nation (the size of California) follow- “forays” to explore the collection. Bar- ing its separation from the Gondwana by Staci Markos bara Ertter came to Berkeley in 1985, supercontinent about 80 million years was Curator of Western North Ameri- ago. Since then, its flora and fauna can Flora, and served as Administrative High-level Iranian officials have been shaped by tectonic plate col- Curator until 2006. She also participated made a rare visit to the University and lisions, mountain uplifts, volcanic ac- in the American-Iranian Botanical pro- Jepson Herbaria in late December, tivity, sea level fluctuations, a series of gram. Below, each describes a handful 2007. Three distinguished visitors, Dr. Pleistocene ice ages, and biotic ex- of their ongoing projects. Mostafa Rahmani, (Director of Inter- changes across the Tasman and South Dr. Smith: In January 2007, est Section of the Islamic Republic of Pacific waters. Spores disperse espe- soon after teaching a 10-day fern work- Iran’s office at the Pakistan Embassy cially easily; about 13% of New shop on the of New in Washington D.C.), Dr. Abolfazl Zealand’s fern species are shared with Zealand, I retired. It was a great field Mehrabadi, (Deputy Director of Inter- Tasmania and Australia, ~1,500 km trip and wonderful experience with kin- est Section of the Islamic Republic of (930 miles) to the west. There have dred spirits (See New Zealand Ferns Iran at D.C.), and Dr. Ali Asghar Share been extinctions and adaptations as article, this issue), but it was time, I Doust, (Professor & General Secretary both original and migrant species wanted to pursue other activities in of the Iranian Counsel Abroad, Continued on page 5. addition to continuing research in fern President’s Office), came to discuss the systematics. I continue working in the American-Iranian Botanical program ALSO IN THIS ISSUE herbarium three or four days a week, that was initiated in 1999 at the Uni- to work on ongoing and long-deferred versity Herbarium by Dr. Fosiee Univ. Herbarium Research projects. Often, the research is collabo- Tahbaz. The ongoing scholarly collabo- rative with individuals at other institu- ration between scientists and students C. Perrine moves to SLO tions, for example, the ferns of Bolivia from Iran, UC Berkeley, and other U.S. (with a German colleague), or those of institutions has resulted in three expe- Donald Kaplan, In Memoriam Continued on page 2. Continued on page 4. Director’s Column, continued from page 1. Venezuela, each of which contains covery prompts new questions, sug- the herbaria, I will be rehired on 20% more than a thousand fern species. gests a new avenue of research, or in- time to continue as Curator of Western Undescribed taxa and taxonomic prob- volves me with colleagues. North American Flora. This will allow lems abound in the wet tropics, and me to prepare updated versions of my there are no modern fern treatments for Annotated Checklist of the East Bay most neotropical countries. I continue Flora and The Flowering and to collaborate on phylogenetic work Ferns of Mount Diablo, and to work (evolutionary relationships), especially with UC entomologists Gordon Frankie on the family Polypodiaceae, with and Robbin Thorp on a UC Press field colleagues in Europe and South guide to urban bees and their preferred America. Occasionally, I look at plants. I will also continue my work Californian and western North American on the history of California botany, my ferns, for instance as a contributor to interest in Juncus, and my involvement the second edition of The Jepson in the American-Iranian Botanical Pro- Manual. gram coordinated by Dr. Fosiee Tahbaz. Last March, a new opportunity My immediate focus is to com- arose – a McBryde Fellowship at the plete treatments needed for volume 9 National Tropical Botanical Garden in of Flora of North America North of Kauai, Hawai'i. I spent four weeks an- Drs. Smith and Ertter, January 2008. Mexico, as well as the second edition notating specimens in the herbarium, Photos by Bridget Wessa of The Jepson Manual. In collabora- mainly from Pacific islands, going in tion with Reidar Elven, Jim Reveal, and the field with colleagues, and involv- David Murray, I am responsible for gen- ing myself in projects already under- era in tribe Potentilleae, notably Poten- way there: ferns of the Marquesas Is- tilla s.l., Ivesia, and Horkelia. I am also lands, and from Rapa in the Austral Is- assisting Walter Lewis in the treatment lands, Polynesia. New species, prob- of Rosa. The last continent-wide treat- lematic genera, and phytogeographic ments of Potentilla and Rosa were in relationships are the central focus of my the early 1900’s, and numerous changes interest in ferns of these areas. have accumulated since then with no One of my passions over the consensus existing across regional flo- years has been fern identification, es- ras. As an essential preliminary to the pecially in the Neotropics; this is a treatment itself, I have produced ten niche served by fewer than a handful publications during the last year con- of individuals around the world. Lately, taining nearly a dozen new species and I have expanded my interests in identi- varieties, numerous new combinations, fication to such exotic localities as the Dr. Ertter: After 22 years at several lectotypifications, and other islands of Sulawesi (Indonesia), UC Berkeley, I opted for early retire- important clarifications of the species Pohnpei, the Society Islands (Moorea), ment in September 2007, after passing and genera to be treated. Far from New Caledonia, and Madagascar, and administrative responsibilities into the wrapping up my research on to fern-rich areas of Vietnam. In any capable hands of Andrew Doran. This Potentilleae and Rosa once volume 9 is given year, the number of specimens I was coupled with moving my primary completed, I am already making plans identify exceeds 5,000, and this also residence back to my roots in Boise, to address multiple questions that re- involves the curation of these speci- Idaho, where most of my relatives still main unanswered, which will not only mens into the collections at Berkeley. live and where I could have easy ac- involve field work throughout western Most recently, a shipment of nearly 500 cess to vast mountains, remote deserts, North America but also collaborators in specimens arrived for identification and the amenities of a vibrant small Alaska, the Czech Republic, Sweden, from the richest areas of southern city. and Germany. China, in Yunnan Province. By no means am I retiring While I look forward to in- Diversity in life is a good thing, from botanical research, which remains creased opportunity for foreign collabo- both in helping maintain my interest in my lifelong passion, nor am I abandon- ration and travel, my primary focus re- ferns, and as I reach those "Golden ing my ties to the University and mains western North America, with its Years.” I plan to stay active as long as Jepson Herbaria. Indeed, thanks to wealth of uncatalogued biodiversity, the spark is there, and each new dis- Mary L. Bowerman’s generous gift to complex biogeography, and awesome Ertter, continued on page 7. 2 New Research at the University Herbarium

Bryophyte collecting in Costa Rica Professor Paul Wilson of Cal State Northridge is visit- by Dan Norris, Research Botanist ing the herbarium on sabbatical. He has done research on pollination in Penstemon, Calochortus, and several other From 3-17 July, 2007, I was a guest at Nectandra wildflower groups, but this year he is studying mosses. Al- Gardens, about 80 km northwest of San Jose, Costa Rica. I ready he has brought in was sponsored by the Nectandra Institute (a non-profit Oligotrichum parallelum and organization based in Alameda, California). Nectandra Warnstorfia pseudostra- Gardens is open to the public and is part of the Nectandra minea from the Sierra Ne- Cloud Forest Preserve, a parcel of about 130 hectares of vada, which are range exten- cloud forest, owned by David and Evelyne Lennette since sions with the nearest previ- 1999. They are developing an ecological education and ous records from Washing- conference center there, and they co-founded the Nectandra ton. He is collaborating with Institute. UC botanists on a compari- My role in the project was to develop a list of species son of the moss florulas of 14 of bryophytes and lichens found in the reserve. About 648 natural areas around Califor- separate collections were made during my two weeks of nia. field studies. The collections were received at the herbarium in Berkeley in November and are now being analyzed. Some of the species found were epiphylls (lichens, liverworts, and Don R. Reynolds, Research Botanist a few mosses growing on the of various vascular In November 2007, I made a trip to the Singapore taxa). A typical may have 10-15 different epiphytic Botanical Garden (SBG), with the sponsorship of Dr. Benito species growing on it! The fraction of leafy liverworts is Tan. My interest is twofold: epifoliar fungi and high, amounting to about 85% of all bryophytes collected. Mycomycetes. I have been collecting these groups since 1963 Nectandra Gardens and the Nectandra cloud forest in the neo- and paleotropics and particularly in the SE Asian preserve are located only 15 km from the town of San area, and undertaking taxonomic studies on ascomycetes that Ramon. Nectandra Gardens is an ideal destination for bio- occur in the tropical forest canopy as well as Myxomycetes logically interested individuals. Five miles of trails circle in general. On this recent trip, I focused on the collections through the housed at the SBG and made collections from there as well pristine cloud as adjacent forests in the Singapore vicinity. Newly collected forest with specimens will be incorporated in the UC Herbarium and placards ident- utilized in ongoing monographic and biodiversity studies. ifying a large number of the Mo-Mei Chen, Research Associate woody species In a project supported by Save-the-Redwoods on the prop- League, I am continuing a project to produce a unique erty. The trails collected work that brings together seven previously are in superb published works on the Redwoods of the world. These works condition and will be translated from English to Chinese for the first time can be used and therefore, the new publication will be of worldwide scope even by phys- for an English/Chinese readership and will serve to educate ically limited more than a billion people all over the globe about the individuals. importance of preserving redwoods. This project has been developed as an international collaboration between the University and Jepson Herbaria, the Beijing Botanical Institute, and the Chinese Academy of Sciences. Joint expeditions to the area in China where Dawn Redwoods occur have taken place over many years, with more then 100 botanists participating from the US and China. www.nectandra.org

3 From Berkeley to San Luis Obispo new community and are becoming by Cynthia Perrine more familiar with the central coast’s abundant outdoor opportunities. I’m As announced in the last issue, thankful that we had Dr. Keil teach the Anna Larsen has taken the helm of the Flora of San Luis Obispo County last weekend workshops. The 2008 season year; come spring I might be able to is full of intriguing topics and new re- name the flowers that bloom on those gions to explore (see back cover for a hillsides! list of workshops). New opportunities I want to take this opportunity have led my family further down to thank each and every workshop in- California’s coast, to San Luis Obispo. structor, participant, and volunteer who We knew that what brought helped make the past four years of our family to Berkeley 4 years ago weekend workshops so amazing. I’m would eventually lead us away again. especially grateful for interim coordi- While I coordinated the workshop pro- nators Anna Larsen and Heather gram, my husband, John, completed Charlotte, Cynthia and Shannon Perrine, Driscoll who filled in for me during my his Ph.D. and then a postdoctoral fel- near Bishop’s Peak maternity leaves. I will hold close all lowship with the Museum of Verte- It was an exciting summer for of the great memories I have of “20- brate Zoology’s Grinnell Resurvey us! We welcomed our second daugh- minute” hikes, taking 12-passenger Project. He has accepted a Conserva- ter, Charlotte Elizabeth, on July 15th vans 4-wheeling, rare species sightings, tion Biology professorship at Califor- (while participants enjoyed the Flora of keying by lamplight, flashlight botaniz- nia Polytechnic University. He started the Great Basin workshop in Nevada). ing, cooking enchiladas, and so much teaching in January and we look for- A month later, we relocated from Ber- more. It’s true that we do it for the love ward to continuing his mammal re- keley to San Luis Obispo. The girls are of plants but it’s really the people who search throughout California. growing fast, and we are enjoying our make our efforts so rewarding.

Iran, continued from page 1. ditions to Iran and multiple visits of can-Iranian Botanical program seeks to versity. The lecture will be part of our Iranian scholars to the U.S. Recently, do. Brent Mishler, Director of the Her- “Botany Lunch” series which offers Berkeley undergraduate students were baria, and Dr. Tahbaz have proposed a informal lectures every Friday at noon hired to mount and file all of the speci- collaborative project that will investi- in the herbaria’s meeting room. mens collected from Iran and then find gate threats to biodiversity all existing Iranian material that is al- from development and glo- ready part of the University bal warming. During the Herbarium’s collection. In total, over meeting, Mishler said 3,000 specimens will soon be ready for “Biodiversity transcends use by researchers around the world. politics” and he received nods of agreement from ev- Dr. Tahbaz has continued to eryone. seek funding and support for this To learn more project and hence the reason for the about the American-Iranian visit in late December. During a for- Botanical program, please mal reception that included dialogue in visit http://ucjeps.berkeley. both English and Farsi, Dr. Rahmani edu/main/research/iran/. said “Politics is going to fluctuate, but Readers are also invited to science is something that is not going attend a lecture on March 7, to change with the president.” He then 2008, “A comparison of added, “Animosity is not going to go range management in Iran away, unless these two countries en- and the U.S.” presented by hance their interaction in peaceful are- Eahsan Shahriary, Research Scholar Mishler, Rahmani, Share Doust, Tahbaz, nas.” from University of Tehran and Depart- and Mehrabadi. Photo by Cathy Cockrell, That is exactly what the Ameri- ment of Botany, Oklahoma State Uni- UC Berkeley Public Affairs.

4 Ferns of New Zealand, continued from page 1.

Waipoua Kauri Forest, then east to a whole new level. Psilotum nudum to Kerikeri on the Pacific’s Bay grows out of sheltered rock crevices in of Islands, where we were joined North Island thermal areas. Four spe- by Dr. Barbara Parris, a local cies of Tmesipteris are in NZ, one of fern researcher. From there, we which – T. tannensis – is endemic. All headed southeast to Te Aroha at are epiphytic, dangling from old leaf the base of the Coromandel Pen- bases of tree ferns, especially insula, south again and a bit in- dicksonias. We saw three of the four land to Rotorua in the Taupo species and there was no forest that was Volcanic Zone, and then west to lacking a fork fern or two. We even the Pureora Forest en route to found excellent specimens in New Plymouth – gateway city Whakarewarewa Forest in Rotorua, a to Mt. (Mt. Egmont) strange place planted almost exclusively Fern enthusiasts in front of Tane Mahuta Kauri tree, l to r: National Park. Finally, we were with North American conifers like Cali- (back) Paul Silva, Joan Smith, Janice Forbis, Richard southbound to the hilly capital fornia REDWOOD! Despite the floris- Beidleman, Dr. Alan R. Smith (instructor), Cynthia Perrine (coordinator), Linda Beidleman, Tara Forbis, Jan of Wellington, with its Karori tic disconnect, lots of native NZ ferns, Nachlinger, (front) Milton Gottleib, Joan Gottleib (author), Wildlife Sanctuary, the Otari- including tree ferns and epiphytes, Wendy Born, Liz Parsons, Melanie Denninger Wilton Bush, and a superb, behind-the- thrive under the alien canopy. struggled to keep pace with this dy- scenes program at Te Papa Museum NZ is home to other ancient namic land. and Herbarium. pteridophyte families. The Ophio- The greatest challenge for the During the Wellington seg- glossaceae (recently assigned to the 1 biota began about 1,200 years ago with ment, we met Patrick Brownsey , and same order and class as the Psilotaceae) Maori (Polynesian) settlers. They were John Dawson2, authors of the reference is represented by three species of followed by Europeans about 200 years books that were our trip “bibles.” Botrychium and two of Ophioglossum, ago, after the historic explorations of For this report there is only including the tiny, easily overlooked O. Abel Tasman, James Cook, and others. enough space to touch on a few of New coriaceum that was seen on a New Burning and clearing, urban develop- Zealand’s fascinating ferns and lyco- Zealand foray to the a de- ment, and alien species introductions pods. They are grouped here into pat- cade ago. have restricted the original flora and terns of scientific and/or aesthetic in- Marattia salicina is the stun- fauna to 10% of New Zealand’s land terest. The first such grouping would ning, sole representative of the mass. Fully half of the indigenous birds be the “oldies,” those from families at Marattiaceae in NZ. Its huge, twice- are extinct, the most dramatic being the the bottom of phylogenetic clades3, pinnate fronds bear synangia that edge large, flightless moas. Even the dating back 300 million years or more. its pinnules like a perfectly stitched country’s signature kiwis are now rare That would put Psilotum (whisk fern) hem. and local. Except for a couple of bat and Tmesipteris (fork fern) in first species there were no extant mammals The endemic “mangemange,” place. They are the sole remaining articulatum (Lygodiaceae), in New Zealand until humans brought genera of what was recently thought everything from rats and cats to the is a common climbing fern in lowland to be the earth’s oldest forests. Along open areas on many omnipresent sheep. Aside from humans family – the Psilotaceae. themselves (now numbering 4+ mil- “bush” trails one of the tangle or um- lion), the Australian possum, imported The entire sporophyte body of brella ferns (Gleicheniaceae) is sure to for the fur trade, has probably been the these amazing plants consists of un- show. Gleichenia microphylla, most destructive, having both the abil- branched or dichotomously branched Sticherus flabellatus, and the endemic ity to climb and a voracious appetite for stems (both subterranean and aerial) S. cunninghamii spread their forking nearly everything green. and veinless or single-veined, vestigial rachises and stunning symmetries in “leaves” with synangia (fused sporan- eye-catching arrays, despite their At the beginning of December, gia) in their axils. There are no roots, thicket-like growth habit. The 2006, fourteen of us met in Auckland and brown, underground are is represented by 3 gen- for a Jepson Herbarium workshop virtually indistinguishable from the era. Todea barbara is a large, impres- “Ferns of New Zealand,” expertly led mycorrhizal gametophytes (except for sive, clump-former that generates tall, by Dr. Alan R. Smith. Our route fol- gametangia on the latter). They bring leathery, bi-pinnate fronds from mas- lowed the Tasman Coast north to the “alternation of generations” debate sive, fibrous bases. Naked sporangia

Continued next page. 5 Ferns of New Zealand, continued from page 5. cover the rachis ends of lower pinnules The NZ flora is notable for and NZ species listed under this name on the basal pinnae. Osmunda regalis has several true climbing ferns. These are more accurately placed in the been introduced to NZ and is naturalizing are unrelated genera that begin life pantropical genus Alsophila (includ- in some areas. has two en- on the ground, often in juvenile ing the famed silver fern A. dealbata) demic species in NZ, one of which gets form with small, simple, strap- or in the Old World genus the prize as “most beautiful” fern of the shaped leaves. They reach matu- Sphaeropteris (e.g., black ponga S. trip. That would be Leptopteris superba, rity as their long-creeping rhizomes medullaris with sleek, dark stipes)3. aptly named for the furry feel of its el- make their way up a support. In- Not all “tree” ferns are especially tall. egant, tapered fronds, segments of which cluded in this group are A. colensoi is accurately dubbed the project upward at a 90o angle from the tenella (), “creeping” tree fern. It seldom gets plane of the leaf – a 3-D sensation. Blechnum filiforme, Lygodium more than 1 m (3.3’) tall, normally Epiphytic and climbing ferns con- articulatum (endemic), Micro- growing somewhat prostrate. stitute another interesting and significant sorum scandens, and Pyrrosia There was a lot of interest in group in NZ. The filmy ferns, eleagnifolia (endemic). the composition of tree fern trunks Hymenophyllaceae, dominate the epi- Other marvels of NZ pteri- because they are commonly lashed to- phytic niche, with 27 species in NZ, and dology are the majestic tree ferns gether in NZ to make decorative also qualify as another of New Zealand’s that dot the landscape and are such fences, new plants often sprouting ancient fern lineages. Cardiomanes significant parts of the forest under- from axillary buds. Cross-sections reniforme (endemic) is well-named for its story. DNA technology is finally confirm that tree ferns are not “woody” large, kidney-shaped fronds. Far-creep- clarifying the relationships within in the sense of cambium-generated sec- ing rhizomes can form spreading mats on this group of eight related families, ondary xylem tissue. Rather, the bulk the ground or up on branches and stumps. only some of which have species of their girth is composed of fibrous Its delicate, translucent look belies its re- that could be described as “trees”. frond bases and adventitious roots. markable resurrection capability. Drought All NZ tree ferns are in the fami- Primary vascular bundles and pith oc- resistance is clearly important to epiphytes lies Cyatheaceae and Dickson- cupy a small, central area. It is a WOW since they cannot grow down to deeper iaceae. Most of the NZ species are moment when one realizes that ar- water sources. Thus, Hymenophyllum endemic and can reach impressive borescent ferns arise from fertilization ferrugineum is covered densely with stel- heights of 20 m (65’). Their colos- events on the same types of tiny, late hairs, the endemic H. flexuosum has sal fronds can be 5 m (16’) long and ephemeral gametophytes that generate a seersucker surface all the way to the base form dense skirts that characterize grammitids! of each frond, and H. pulcherrimum, also the dicksonias. The genus Cyathea The three endemic fern endemic, is crinkly-leafed. Trichomanes is found in the New World tropics genera in NZ are worthy of special venosum, with the classic, tubular indu- note. They are Anarthropteris sia that mark the genus, looks like strips (Polypodiaceae), Leptolepia of pale-green, tissue paper hugging moist (), and Loxoma tree trunks. Other epiphytic/epipetric NZ (Loxomataceae). Anarthropteris ferns include cunninghamii lanceolata is a grammitid fern and a (endemic), Asplenium flaccidum, A. relatively common forest epiphyte, polyodon, and the grammitids growing at or below eye level on trunks (Polypodiaceae) Anarthropteris or rocks. It looks like short straps lanceolata (an endemic genus), hanging limply from their supports. Ctenopteris heterophylla, Grammitis Bulging, round sori seem oversized on billardierii, G. ciliata (endemic), and G. the small fronds. As with most NZ pseudociliata. The dryopterid Rumohra ferns, the sori have no protective in- adiantiformis (cultivated for long-lasting dusia. Leptolepia novae-zelandiae, fronds used in flower arrangements) and which Dr. Smith saw often on the the lycopod Huperzia varia also thrive on South Island, may also have been seen arboreal perches in NZ. And, of course, during our Te Aroha trek. It favors all species of Tmesipteris are epiphytic. stable, dark forest (unusual for a Dr. Smith noted that conifers are gener- dennstaedtioid) and has jagged-edged, ally free of epiphytes (something to do triangular indusia that open at the pinna with their bark or resins?). Article author Joan Gottlieb poses next to New Zealand’s national symbol, the silver tree fern. margins. Broad, deltate fronds are very

6 Loxoma pinna detail. Ertter, continued from page 2. primeval” species. Rem- beauty. In addition to significant field nants of these remarkable work, my goals include fostering inter- “gumland” forests are actions among regional herbaria, land now fully protected. management agencies, botanical gar- In the “goblin” dens, conservation organizations, and cloud forests on Mt. other institutions and individuals with Taranaki – as close to botanical interests. Towards this goal I heaven as a fern lover gets have become a research associate at the – an unusual angiosperm, Snake River Plains herbarium (SRP) of with a strong fern connec- Boise State University and the Harold tion, grows. The smooth- M. Tucker herbarium (CIC) of the Col- barked kamahi lege of Idaho (my alma mater), as well (Weinmannia racemosa – as the Missouri Botanical Garden her- finely dissected and have conspicuously Cunoniaceae) starts life as an epi- barium (MO). reduced basal, basiscopic pinnules. This phyte on the fibrous base of a tree- I am also very much enjoying field image is important, because fern trunk. Roots soon grow down getting a garden started at my new Leptolepia resembles the dryopterid ge- into soil and multiple trunks grow up, abode, successfully incorporating some nus Lastreopsis, whose three NZ species eventually reaching 25 m (80’). This favorites from Berkeley (including have reniform indusia and greatly exag- often obliterates the original fern Shasta snow-wreath), while experi- gerated basal and basiscopic pinnules. host, but sometimes merely sur- menting with a new palette in an area Last, but hardly least, is Loxoma rounds it – an interesting association, where competitive gardening has re- cunninghamii, a magnificent, sizable fern but generally not to the benefit of the cently taken hold. I have already had of clearings, open forests and banks, thriv- fern. the pleasure of hosting friends from ing in poor, heavy soils. Distinctive, tu- California, and continue to treasure We meditated on the impor- bular indusia fully cup their contained both long-established and newly tance of all these native, often unique sporangia and project outward, like nar- formed friendships in a life-style deci- plants in NZ while enjoying lunch row thimbles, from notches at pinnule sion that I am enjoying immensely. in Wellington under a huge, metallic margins. Loxoma is a Northland fern, still sphere with a cutout silver fern mo- abundant on the Coromandel Peninsula, As you gear up for spring tif. It was suspended between the but its wider range is giving way to de- botanizing, don’t forget art museum and shops across Gal- velopment. We saw it at The Fernery, a to take the lery Square. This is a country that glass-roofed greenhouse complex set into Second Edition truly appreciates its fabulous ferns, an excavated hillside in Pukekura Park, treatments for a as did we, with Dr. Alan Smith’s gen- New Plymouth. test run ! erous encouragement and expertise. One cannot leave NZ without REFERENCES Second Edition of marveling at some of its non-pterido- The Jepson Manual phyte flora – especially its antipodean, Text reprinted with permission from broadleaf conifers. On the North Island Fiddlehead Forum, Volume 34, num- Treatments for public viewing massive kauri trees ( australis – ber 1 available at: Araucariaceae), with their metallic-grey, 1 Brownsey, Patrick J. and John C. resinous bark and parallel-sided trunks Smith-Dodsworth, 2000, New http:// – 6 m (20’) in diameter – can rise un- Zealand Ferns and Allied Plants, ucjeps.berkeley.edu/ branched for 20 m (65’) before spread- Auckland NZ, David Bateman Ltd jepsonmanual/review/ ing their crowns (actually the lower 2 Dawson, John and Rob Lucas, branches are shed early and the scars are 2000, Nature Guide to the New Please forward any comments sealed over). These giants can live 2000+ Zealand Forest, Auckland NZ, you may have about these years and have a direct lineage going Random House New Zealand. treatments to the Scientific back 100 million years. Until recently, 3 Smith, Alan R. et al., 2006, “A clas- Editor, they dominated extensive forests in sification for extant ferns.” Taxon Dr. Thomas Rosatti Northland, along with podocarps like 55:705-731. 705-731. ([email protected]). rimu (Dacrydium cupressinum), totara . (Podocarpus totara), and other “forest

7 In Memoriam

Donald Kaplan (1938-2007). Faculty, staff, and students of the Herbaria will remember Dr. Kaplan as a de- voted teacher, colleague, world-renowned plant morphologist, author, and railroad en- thusiast. Dr. Kaplan earned many honors and awards and much has been written about him including a portrait in The Jepson Globe (2002, V13#2). He described himself as a “communicator for the plants” and we truly valued his ideas, research, and fellowship. We will miss him. Dr. Kaplan is survived by his wife, two sons, and sister. A campus memorial is being planned for early 2008. To be included on the mailing list for this event, please contact Staci Markos ([email protected]).

LLLawrence R. Heckard Fund Awards

The purpose of the Lawrence R. Heckard Fund of The Jepson Herbarium is to encourage research in the systematics of the vascular plants of California and their close relatives in North America. The Heckard Fund Committee received 10 proposals for 2007 (not including mini-grant proposals received off-cycle during the year for The Jepson Manual revi- sion). Below is the list of awards made in response to our regular call for submissions this year:

Investigator Subject of proposal Ackerly, David Digitization of Baker seed data Baldwin, Bruce Tarweed systematics Bolmgren, Kjell Rhamnus/Frangula comparative ecology Driscoll, Heather Californian Allium systematics & Chelsea Specht Ertter, Barbara Potentilleae floristic treatments Moore, Abigail Grindelia systematics Park, Michael Collinsia systematics Shuldman, Michael Comparative ecology of woody California endemics Simms, Ellen Phylogenetic and population studies in Lupinus Stuart, Stephanie Comparative studies of cold adaptation in Sierra

The Herbarium and all of the grant recipients are very grateful to the late Larry Heckard for his generous bequest, it will continue to fund research projects such as those listed above in perpetuity. Mark Your Calendar!

East Bay Science Cafe Don’t live in the meets monthly in the evening. For details, see: Bay Area? Visit http://bnhm.berkeley.edu/ Saturday, Sciencecafes.org about/sciencecafe.php April 12th to find a cafe near you!

8 THE JEPSON HERBARIUM PROJECTS & RESOURCES

Support the Second Edition! The Jepson Flora Project Second Edition of The Jepson Manual Have you been thinking about making a gift to Online Interchange for California Floristics Jepson Desert Manual support the second edition of The Jepson Manual? Online Horticultural Database Electronic Publication of Jepson’s A Flora of California Would you like to be acknowledged or honor a Ecological Flora of California loved one or friend in the front pages of the Publications & Research Projects Constancea: University of California new Manual? electronic publications in botany DeCew's Guide to the Seaweeds Flora of Mount Diablo If so, please consider joining our sponsorship Unravelling the dynamics of mating-system program. All gifts of $1,000 or more will be evolution in tribe Collinsieae Building the Tree of Life ⎯ A National Re- acknowledged in the new Manual (must be source for Phyloinformatics and Computa- received in full by December 31, 2008). tional Phylogenetics Deep Green Plant Phylogenetics: Novel Analytical Methods for Scaling Data from With acknowledgment in the new Manual, Genomics to Morphology Beyond "Deep Green": Towards an Integra- your gift will serve as a lasting tribute to your tion of Plant Phylogenetics and Plant dedication to the conservation and Genomics Demography and Germination Ecology of understanding of the flora. the Endangered Santa Cruz Tarplant Phylogeny and evolution of the true thistles, genus Cirsium (Compositae) Sponsorship opportunities are exclusive and are Niche conservatism, functional trait evolution, available on a first-come, first-served basis. and the diversification of the California vernal pool flora Educational Services & Resources Please see our web site: Botanical Workshops & Courses http://ucjeps.berkeley.edu/jeps/friends/sponsored.html Plant Identification 2,200,000+ Worldwide Plant Specimens for a complete list of genera, families, and subregions Botanical Library and Slide Collection that have already been sponsored. Administration Trustees: Vice Chancellor Emeritus Roderic Park, Chairman; Vice Chancellor Beth Burnside (on leave); UC Botanical Garden California Native Plant Society Director, Paul Licht; Professors John Taylor and Brent Mishler (ex officio) Director: Professor Brent Mishler CNPS 2009 Conservation Conference Curator: Professor Bruce Baldwin Research Associate: Bridget Wessa January 17 - 20, 2009 Jepson Flora Project Staff: Sacramento Convention Center Project Research Specialists: Jeff Greenhouse, Scott Simono & Sheraton Hotel Project Manager: Staci Markos Database Admin.: Richard Moe This conference provides a unique forum Scientific Editor: Tom Rosatti for California’s diverse audience to learn Managing Editor: Margriet Wetherwax Administrative Curator: Andrew Doran about science and policy-based solutions Senior Museum Preparator: Ana Penny for improving native plant conservation. Assistant Museum Scientist: Kim Kersh Development Coordinator: Staci Markos See www.cnps.org for details! Administrative Assistant: Edith Summers Public Programs: Anna Larsen Globe Editors: Perrine & Markos

9 Jepson ManualVolunteers Fundraising Needed! / Consulting Firms

VOLUNTEER OPPORTUNITIES IN THE UNIVERSITY AND JEPSON HERBARIA Curatorial Volunteers Needed!

Are you interested in learning more about the California flora, gaining firsthand experience with herbarium techniques, and socializing with fellow native plant enthusiasts? Then have we got a deal for you! Selected Saturdays of each month are Group Volunteer Days in the Herbaria. What better way to spend those rainy winter weekends! Group Volunteer Saturdays begin at 10 am and finish up by 5 PM (participants need not stay the full time). We also welcome individual volunteers who can come in during our regular hours (M-F 8-5). We will try to match your unique interests and abilities.

2008 Volunteer Saturdays

February 9

March 8 April 5 May 10

For more information, please call or write to Ana Penny (510) 642-2465, [email protected]

California Botanical Society

Annual Banquet February 23 - UC Santa Cruz Arboretum, Santa Cruz, CA - Speaker: John N. Thompson, Department of Ecology and Evolutionary Biology, UC Santa Cruz “An Ecologist’s View of How to Build a Plant.” *Annual Banquet Requires Purchase of Advance Registration, see web site for details*

Free Lectures - 2063 Valley Life Sciences Building, UC Berkeley - 7:00 PM. March 20 - Science-Based Policy of Genetically Engineered Organisms at the Environmental Protection Agency - Richard Whitkus, Department of Biology, Sonoma State University, www.sonoma.edu/users/w/ whitkus/

April 17 - California Alliums: Insights into a Biogeographic Radiation and Edaphic Evolution, Chelsea Specht, Plant and Microbial Biology, University of California, Berkeley, pmb.berkeley.edu/~specht

Refreshments will be served before (6:45 PM) and after the seminars. For additional information please call (510) 643-7008 or visit www.calbotsoc.org

10 FRIENDS OF THE JEPSON HERBARIUM CATEGORIES OF GIVING

Name(s) ______GIFTS TO SUPPORT THE SECOND EDITION $25,000 Honor the contributions and Address ______founding principles of W. L. City, State Zip ______Jepson, former Jepson Trustees Lincoln Constance and Robert Telephone / email ______Ornduff, and former Jepson Curators Rimo Bacigalupi and I would like to join the Friends / renew my membership Lawrence R. Heckard (contribution to the annual fund). $10,000 Support taxonomic efforts in an organizing unit of the Manual I would like to support the Second Edition of The Jepson Manual with my Ferns, Gymnosperms, Dicots, gift of ______. or Monocots $5,000 Support floristic effort for a Enclosed is _____ of a total pledge of _____ to be paid over ____ years. particular bioregion Please acknowledge me as a sponsor of ______(Twenty-four listed in (indicate genus name, e.g., Lilium, family name, e.g., Poaceae, or other the Manual) category) by printing my name in The Jepson Manual $2,500 Support taxonomic work in a (for gifts of $1,000 or more, see side bar). particular family See the Herbarium web site for Please acknowledge my gift as anonymous. an up-to-date, complete list My or my spouse’s employer will match this gift. $1,000 Show enthusiasm for your (Please enclose company form) favorite genus (pledge $200 / 5 years) This gift is ___ in honor of ___ in memory of ______

ANNUAL SUPPORT Please make your check payable to UC Regents $500 Contribute to the illustration of or charge your gift. a new species ___ Visa ___ Mastercard $250 Help accession specimens from Account # ______the backlog Exp. Date ______$100 Support taxonomic research at Signature ______the species level $35/$50 Basic membership in Friends MAIL TO: of the Jepson Herbarium The Jepson Herbarium 1001 VLSB #2465 SPONSORSHIP OPPORTUNITIES are University of California exclusive and will be available on a Berkeley, CA 94720-2465 first-come, first-served basis. With ap-

The information you provide will be used for University business and will not be released unless required proval from the donor, gifts at the by law. A portion of all gifts is used to defray the costs of administering the funds. $1,000 level and above will be ac- All gifts are tax deductible as prescribed by law. knowledged in the front pages of The Jepson Manual (gifts must be received by December 31, 2008).

11 FRIENDS OF THE JEPSON HERBARIUM Nonprofit Organization The Jepson Globe, Vol. 18 No. 3 1001 Valley Life Sciences Building #2465 U.S. Postage PAID University of California, Berkeley University of California Berkeley, CA 94720-2465

ADDRESS SERVICE REQUESTED

Workshops remaining in the 2008 series March April, continued July Intro to Bryophytes Compositae Juncaceae March 1-2 April 26-27 July 18-20 Morphology and ID March 8-9 May September Bryo Inventory Shasta-Trinity Forest Climate Change March 15-16 May 1-4 September 10-14 South Diablo Range Poaceae *wait list only March 27-30 May 10-11 October Fifty Families *wait list only Medicinal Plants Insects & Coevolution March 29-30 & April 5-6 May 17-18 October 18 Sierra Valley April May 29-June 1 November Fifty-one Families *wait list only Lithocarpus densiflorus April 5-6 & 12-13 June November 23 Plant ID ~ San Diego Co. Bee Pollination April 10 -13 June 6-8 Santa Cruz Island *wait list only Paleobotany April 17-20 June 14 Desert Mountains April 24-27

Please visit our web site at: http://ucjeps.berkeley.edu/workshops for full descriptions and registration information, or contact Anna Larsen, (email preferred) [email protected] or (510) 643-7008