Transactions and Proceedings

of the Palaeontological Society of Japan

New Series No. 89

Palaeontological Society of Japan April 20, 1973 Editor: Hiroshi UJIIE Associate editors: Ikuo OB ATA, Yasuji SAITO, Kazuhiko UEMURA

Officers for 1973 - 1974

Honorary President: Teiichi KOB AYAS HI Presiden t : Tatsuro MATSUMOTO Councillors (*Executive): Kazuo ASAMA,* Kiyoshi ASANO, Kiyotaka CHI NZE I,* Taka­ shi HAMADA,* Tetsuro HA NA I,* Kotora HATA I, Itaru HAY AM I,* Kametoshi KANMERA,* Tamio KOTAK A, Tatsuro MATSUMOTO,* Tokio SHIKAM A, Tsugio SHUTO, Yokichi TAKA YANAGI,* Toshimasa TANAI, Hiroshi UJIIE* Secretaries: Wataru HASHIMOTO, Saburo KANNO Executive Committee : General Affairs: Kiyotaka CHINZEI, Yasuhide IWASAKI Membership: Kazuo ASAMA, Ienori FUJIY AMA Finance: Takashi HAMADA, Naoaki AOKI Planning: Tetsuro HAN AI, Saburo KANNO Publications Transactions: Hiroshi, UJIIE, Ikuo OBATA, Yasuji SAITO, Kazuhiko UEMURA Special Papers: Kametoshi KANMERA, Itaru HAY AMI, Tomowo OZAWA "Fossils": Yokichi TAKAYANAGI, Toshiaki TAKAYAMA

Fossil on the cover is Bellerophon jonesianus DE KONINCK from the Late Permian Akasaka Series in Gifu Prefecture.

All communications relating to this journal should be addressed to the PALAEONTOLOGICAL SOCIETY OF JAPAN c/ o Business Center for Academic Societies, japan. Yayoi 2-4-16, Bunkyo-ku, Tokyo 113, japan Sole agent: University of Tokyo Press, Hongo, Tokyo Trans. Proc. Palaeont. Soc. japan, N.S., No. 89, pp. 1-14, pis. 1-4, April 20, 1973

61l. BIOSTRATIGRAPHIC STUDY OF THE JURASSIC TOYORA G ROUP* PART II

HIROMICHI HIRANO

Department of Geology, Faculty of Science, Kyushu University

.:;; ='- ;7 *cl!\WI"ii'Jil\fO)itT:1I\"ijf;~h'I"J1iJf)t - k"0) 2: j!g:imNillfJ:: 'Jiff-r.Q 7 /-t T -1 r 0) oj 'G, 3::c. L-"Cgg~I"II.iI\1t;j:"J:V':lfX!IlTffi1r}1I\O) Whitbian .toJ:V' Yeovilian ~lf,T Hildocerat­ idae 0) 6 ~ 11 Hne -? ~ ijr.W& L- t-:o YOKOYAMA (1904) fe J:.Q Hildoceras chrysanthemum c. H. densicostatum vi, k"O)5J~fiO)HHjI!c. ~h"Cv'.QJI1JWXfe-?~tfriML-t-:f.if*, [iij-l'illc. 'l'IJJE L- jJlf:ffO)15f!Ji~ c. -? t-:o '* t-:k"O) ~ i'iff: G < nl)iill"(:' ji, -? t-:;Q;, IllHHI.to J: V'/J{rJJm O)~;l'i~;Q' G Harpoceras (s. str.) fe1'iJ}~h.Q"'~tO)C.!FUWiL-t-:o U1..c.viJjljfC, jfrt-:fe Hildoceras felf1ti!IHl\IjJm~ftflj~~Tto)~ Hildoceras sp. c. L-"Ci-IHIj-L-t-: o fIt*' Haugia aff.japon­ ica c.~h"Cv't-:to)fi, Haugia C:fit.J:< Phymatoceras ;Q;*~"(:·ji,.Qo Lillia toyorana c. ~ h"C v' t-: fill fe -? v'"C t ~~iO)fi'i*, Phymatocerasfefl1,T.Q to) c. 'l'UIf)i L- t-:o 3::~/!.- - r

"(:'f~ Ght-:Jl:lff:!Jl-} I\'J, ggg:qLJlm, llfX!IlT I\"i 0)1'±1K!Zl C1tTiJ21:li\ml:~ {, RHm felf, L- "C ji,.Q 0 Jf Y!f 1Ji,. ill

Introduction Subgenus Harpoceras W AAGEN, 1869

This article is Part II of the biostrati­ Harpoceras (Harpoceras) chrysanthemum graphic study of the Jurassic Toyora (YOKOYAMA) Group. In Part I lithostratigraphy was explained and Domerian ammonites were PI. 1, Figs. 1-3; PI. 2, Figs. 1-3; described (HIRANO, 1971). In this part 6 PI. 3, Figs. 6-9. genera including 11 species of Toarcian 1904. Hildoceras chrysanthemum YOKOYAMA; Hildoceratidae and Hammatoceratidae YOKOYAMA, Jour. Coll. Sci., Imp. Univ. are described. Tokyo, vol. 19, art. 20, pp. 11-12, pI. 2, figs. 1-4. 1904. Hildoceras densicostatum YOKOYAMA; Palaeontology (continued from Part I) YOKOYAMA, ibid., pp. 12-13, pI. 3, figs. 1-2. Subfamily Harpoceratinae 1947. Hildoceras (s.l.) chrysanthemum YOKO­ NEUMAYR, 1875 YAMA; MATSUMOTO in MATSUMOTO & ONO, Sci. Rep. Fac. Sci., Kyushu Univ., Genus Harpoceras WAAGEN, 1869 Ceol., vol. 2, no. 1, pp. 27-28. 1947. Hildoceras (s.l.) densicostatum YOKO­ Type-species :-Ammonites falcifer Sow­ YAMA; MATSUMOTO in MATSUMOTO & ERBY, 1820 (designated by ARKELL, 1951, ONO, ibid., vol. 2, no. 1, p. 28. l. C. Z. N. Opinion 363, 1954). 1956. Hildoceras chrysanthemum YOKOYAMA; • Received Feburuary 12, 1973; read jan. ARKELL, jurassic Geology of the World, 23, 1971, at Tokyo. Oliver & Boyd Ltd., p. 421.

1 2 Hiromichi HI RAND

1956. Hildoceras densicostatum YOKOYAMA; long as inner one-third of the flank. ARKELL, op. cit., p. 421. Description:-The whorls enlarge mod­ 1962. Hildoceras chrysanthemum YOKOYAMA; eratelyand are somewhat evolute. The SATO, Mem. Soc. Ceol. France, no. 94, umbilical periphery is more or less sharp p. 59. and the wall is steep but not so high. 1962. Hildoceras densicostatum YOKOYAMA; SATO, ibid., no. 94, p. 59. The venter is unicarinate-bisulcate. The 1963. Hildoceras chrysanthemum YOKOYAMA; ribs spring on the umbilical seam and go TAKAI, MATSUMOTO and TORIYAMA on forward. At about one-third or two­ ed., Geology of Japan, Univ. Tokyo fifths of the flank the ribs sharply curve Press, p. 83. backward. Near the ventral shoulder the 1963. Hildoceras densicostatum YOKOYAMA; ribs recurve forward and more or less TAKAI, MATSUMOTO and TORIYAMA projected on the ventral shoulder but ed., op. cit., p. 83. soon fade away. The detailed curvature of the ribs varies one by one, as illus­ Types :-Here I designate one of the trated by many figures in the plates. illustrated four specimens by YOKOYA­ The ribs become coarser toward outer MA (1904, pI. 2, fig. 1) as the lectotype one-third of the flank, but they are not of H. chrysanthemum, MM. 7058a, and the very weak on the inner part of the flank. three other specimens as the paralecto­ The number of ribs per whorl varies to types (YOKOYAMA, 1904, pI. 2, figs. 2-4), some extent. Sometimes there are striae, MM. 7059-7061. They were obtained at lirae, bifurcation and insertion. Bifur­ loco 47. cation arises at one-third of the flank At the same time I designate one of where the ribs curve backward. In some the illustrated two specimens by YOKO­ specimens ribs become weaker toward Y AMA (1904, pI. 3, fig. 1) as the lectotype the aperture and many fine riblets are of H. densicostatum, MM. 7064a, and the there instead of the major ribs. In one other (pI. 3, fig. 2) as the paralectotype, specimen such a change occur at 90 mm. MM. 7065. The former was obtained at diameter but in another specimen already Nishinagano and the latter at loco 47. at 34 mm. diameter. In a small stage Material.·-Two hundreds and thirty­ (e. g., diameter 10 mm.) the curve of ribs eight specimens are examined, of which is generally weak and more or less simi­ seventy-two specimens, MM. 7058-7061, lar to that of Fuciniceras nakayamenses. GK. G. 2431-2441, 2443-2455, 2457-2492, In a few specimens the sutures are in­ 2501-2508, were previously regarded as completely preserved, showing a narrow Hildoceras chrysanthemum, eighty-eight and deep lateral lobe. specimens, MM. 7064-7065, GK. G. 2551- Measurements :- 2565A, B, 2566-2577, 2579-2584A, B, 2585- 2610A, B-2633, as Hildoceras densicosta­ No. D. U. H. U/D HID Rn~ tum, thirty-nine specimens, GK. G. 2509- MM. 7058a 87.0 38.9 26.2 0.45 0.31 25 2519, 2521-2548, as an intermediate form MM. 7059 42.8 15.1 16.7 0.35 0.39 22 and thirty-nine specimens, GK. G. 11318- MM. 7060 40. 1 14.8 15. 1 0.37 0.38 19 MM. 7065 66.5 27.0 22. 4 0.41 O. 34 33 11322, 11356-11379, GYU. M. 1058-1067, GK. G. 2552 46. 2 20.0 22.0 0.43 O. 48 39 are newly examined. GK. G. 2557 39.8 13. 8 16. 4 0.35 0.41 32 Diagnosis ,'-'-A species of Harpoceras (s. str.) whose ribs have somewhat clear Statistics :-Abou t seventy specimens relief, moderate curve and the haft as are available for counting the ribs per 611. Jurassic Toyora Group-II 3 whorl at a given diameter. Here the By the test as mentioned above no assumption is made that the sample is significant difference is found from the collected at random from one fossil pop­ normal distribution. The frequency dis­ ulation with the normal distribution of tribution of the number of ribs at each the characters. diameter seems to be unimodal. The results of the chi-square test are Remarks :-Hitherto several palaeont­ as follows. ologists and biostratigraphers have had a question concerning the taxonomic po­ sition of the present species in the gen­ Table 1. The number of ribs per half a whorl at 20 mm diameter. Data classified eric rank. For example MATSUMOTO (in for the chi-square test. The classes be­ MATSUMOTO & ONO, 1947) dealt it as tween 27.5 and 31.5 are evaluated as one Hildoceras (s. 1.) and SATO (1967, p. 391) class. suggested that it might belong to Harpo­ Class Oi Ei ceras. In this paper the present species 13.5-15.5 2 2.44 is referred to the genus Harpoceras. 15.5-17.5 6 6.06 Hildoceras HYATT, 1867, was establish­ 17.5-19.5 16 10. 74 ed on the basis of the two species, H. 19.5-21. 5 13 14.45 21. 5-23. 5 12 13.98 bifrons and H. walcotti. Later ZITTEL 23.5-25.5 7 9. 72 (1885, pp. 458-463) dealt Hildoceras as a 25.5-27.5 8 5. 13 subgenus of Harpoceras W AAGEN, 1869 27.5-29.5 1 1. 93 under his Arieticeras-series and from that 29.5-31. 5 1 0.53 time the diagnoses of Hildoceras and Total 66 64. 98 Harpoceras become wider. HAUG (1885, N=66, x=21.29, s=3.53, V=16.58, X2=5.53, pp. 632-646) spread the scope of Hildo­ ;.:20•05 ("=5)=11.07, P»0.05, Not significant. ceras without clear definition and his genus Hildoceras included Lillia bayle, Harpoceras levisoni, Hildaites borealis, Table 2. The number of ribs per half a Harpoceras serpentinum, etc. GEMMEL­ whorl at 30 mm diameter. Data classified LARO (1885) and FUCINI (1900, 1904, 1905, for the chi-square test. The two classes 1923-28) used Hildoceras without clear between 17.5 and 21.5 are evaluated as one class and likely two classes between 29.5 definition in much wider meaning than and 33.5 are done. that of HYATT. FUCINI's diagnoses of Class Oi Ei Harpoceras, Grammoceras and Hildoceras were confusing in his articles. BETTONI 17.5-19.5 2 1. 76 (1900, pp. 53-66) also used genus Hildo­ 19.5-21. 5 6 4.06 21. 5-23. 5 6 6.87 ceras in a wide sense. The diagnosis 23.5-25.5 7 8.39 given by him was different from that of 25.5-27.5 8 7. 86 HYATT. Hildoceras chrysanthemum and 27.5-29.5 6 5.34 were establish­ 29.5-31. 5 2 2.64 Hildoceras densicostatum 31. 5-33. 5 2 1. 00 ed in such an age. In the present paper I employ the diagnosiS given by ARK ELL Total 39 37.92 (1957). N=39, x=24.95, s=3.59, V=14.39, X2 =1.28, The present species has a unicarinate­ X20.05("=3) = 7.81, P »0.05, Not significant. bisulcate venter and no longitudinal The number of ribs per half a whorl at 50 mm. diameter. groove on the flank, and therefore it is N=ll, x=30.45, s=3.45, V=11.31. clearly excluded from Hildoceras. 4 Hiromichi HIRANO

The present species is closely allied shoulders. On the other hand the venter to Hildaites borealis (BUCKMAN) as Yo­ of the present species is unicarinate­ KOY AMA (1904) mentioned. Hildaites bisulcate and not acute but has gentle belongs to the subfamily Hildoceratinae ventrolateral shoulders. as was treated in the "Treatise" by Harpoceras mulgravium (YOUNG & ARKELL (1957). The present species, BIRD) is somewhat similar to the pre­ however, does not belong to Hildoceras sent species, but the former has longer as mentioned above. On the other hand, hafts of the ribs and more acute curve the characters of the present species of ribs on the middle of the flank. are not inconsistent with the diagnosis Occurrence :-Loc. 3: 113, loco 30: 29, of the Harpoceratinae, and Harpoceras loco 26: 12, loco 47: 10, loco 5: 8, loco 28' : is suitable for its generic position. 8, loco 9; 5, loco 2: 4, loco 4: 3, loco 6: The present species has been hitherto 3, loco 22: 3, loco 40: 3, loco 1: 2, loco 35 : divided into Hildoceras (s. 1.) chrysanthe­ 2, loco 47B: 2, loco 18E: 1, loco 41: 1, un­ mum and H (s. 1.) densicostatum. YOKO­ known loco of the Sakuraguchi-dani: 6, YAMA (1904) described that the latter was Tabe district: 1, unknown loc.: 22. distinguishable from the former by the greater number of ribs. MATSUMOTO (in Harpoceras (Harpoceras) inouyei MATSUMOTO & ONO, 1947) revised partly the diagnoses of the two species and (YOKOYAMA) pointed out that there are some speci­ PI. 1, Figs. 4-5; PI. 3, Fig. 5. mens which show intermediate charac­ 1904. Hildoceras inouyei YOKOYAMA; YOKO· ters. YAMA, Jour. Coli. Sci., Imp. Univ. To­ I have examined more than 200 spe­ kyo, vol. 19, art. 20, pp. 13-14, pI. 2, cimens including type specimens of the figs. 5-6. two nominal species and biometrically 1947. Hildoceras (s.l.) inouyei YOKOYAMA; treated about 70 specimens. On the basis MATSUMOTO in MATSUMOTO & 01'\0, of the results of statistics as well as Sci. Rep. Fac. Sci., Kyushu Univ., vol. other observations, all the specimens of 2, no. 1, p. 28. the sample are regarded as belonging to 1956. Hildoceras inouyei YOKOYAMA; ARKELL, one species. Considering the page pre­ Jurassic Geology of the World, Oliver cedancy in YOKOYAMA'S paper, I choose & Boyd Ltd., p. 421. the former as the specific name. 1962. Hildoceras inouyei YOKOYAMA; SATO, Mem. Soc. Ceol. France, no. 94, p. 59. Comparison:-The present species oc­ 1962. Hildoceras inouyei YOKOYAMA; T AKAI, curs together with Harpoceras (s. str.) MATSUMOTO and TORIYAMA ed., Geolo­ sp. cf. H. exaratum, but is distinguish­ gy of Japan, Univ. Tokyo Press, p. 83. able by its less strongly curved ribs. YOKOYAMA (1904) said that the present Types:-As YOKOYAMA (1904) estab­ species was closely allied to Hildaites lished this species on several syntypes, borealis (BUCKMAN). Indeed in the mode one of his illustrated specimens (1904, of the ribs Hildaites borealis is quite pI. 2, fig. 6) is here designated as the similar to the present species, but the lectotype, MM. 7063, and another illus­ hafts of the ribs are shorter in the trated specimen (1904, pI. 2, fig. 5) as the former than in the latter. The venter paralectotype, MM. 7062. They were of H. borealis is unicarinate-tabulate and obtained at loco 47. has somewhat subangular ventro-lateral Material :-Seventy-two specimens, 611. Jurassic Toyora Group-II 5

Types, GK. G. 2641-2652, 2654-2666, 2668- Comparison:-The present species is 2676, 2679-2681, 2683-2685,11299,11316-7, closely allied to Harpoceras chrysanthe­ 11342-11349, GYU. M. 1028-1047. mum (YOKOYAMA) in general characters, Diagnosis :-The diagnosis of the pre­ especially in the quite similar curvature sent species, as noted by YOKOYAMA of ribs. The former has, however, dis­ (1904) and supplemented by MATSUMOTO tinctly involute shell and sometimes show (in MATSUMOTO & ONO, 1947), is: A spe­ fasciculation and inserted characters of cies of Harpoceras (s. str.) which has a the ribs. fairly narrow umbilicus (usually less The present species is also very similar than 1/3 of diameter in every size) and to Hildaites borealis (BUCKMAN) (WRIGHT, bifurcated, indistinctly fasciculated andl 1883, p. 438) in the mode of ribs. The or occasionally inserted ribs. present species, however, is characteri­ Description:-The whorls enlarge rap­ zed by the involute shell and the uni­ idly, with a fairly narrow umbilicus. The carinate-bisulcate venter. venter is probably unicarinate-bisulcate, The mode of insertion and bifurcation although the specimens with a complete­ of the ribs is somewhat similar to that ly preserved venter are not available. of Lioceratoides yokoyamai (MATSUMO­ The umbilical periphery is rounded, but TO), but the latter shows more eminent the wall is steep and low. The ribs spring and regular bifurcation and insertion. on the umbilical wall and go on for­ Occurrence :-Loc. 3: 25, loco 5: 12, loco ward. At a point one-third or two-fifths 30: 11, loco 1 : 3, loco 26: 3, loco 47: 5, loco of the flank the ribs curve backward 2: 1, loco 4: 2, loco 18D: 1, loco 18E: 1, with increasing coarseness, drawing arcs, loco 27: 1, loco 28: 1, loco 35: 1, loco 42: and then recurve forward near the ven­ 1, unknown: 4. tral shoulder. They usually are very weak and often inperceptible on the Harpoceras (Harpoceras) sp. d. H. (H.) umbilical periphery. The bifurcation occurs at one"third or two-fifths of the mulgravium (YOUNG & BIIm) flank where the ribs curve backward. PI. 1, Fig. 6. The fasciculation is indistinctly visible on the innermost part of the flank. In­ Material.·-A fragmentary specimen, serted ribs occasionally occur on the GK. G. 11355. outer two-thirds of the flank. The lirae Description:-The whorls enlarge rath­ and striae are rare, but, if present, they er rapidly, and the umbilicus seems to are visible along the ribs. In some spe­ be somewhat narrow. The venter seems cimens the sutures are partly preserved, to be unicarinate-bisulcate, though it is but the preservation is too poor for poorly preserved. Ribs are strongly fal­ tracing details. An example is shown coid and somewhat prorsiradiate. They in Pl. 3, Fig. 5. are straight and inclined forward on the Measurements :- inner half of the flank. At the middle of the flank they abruptly curve back­ No. D. U. H. UjD HID Rn~ MM. 7062 26.1 8. 6 10.4 0.33 0.40 28 ward and near the ventral shoulder they MM. 7063 39.1 11. 0 16.8 0.28 0.43 29 recurve forward, moderately projecting GK. G. 2642 41. 0 14.3 16. 2 0.35 0.40 37 on the venter. They are accompanied GK. G. 2668 42. 9 15. 4 19.2 0.36 0.45 36 by lirae on the preserved last stage. The GK. G. 2672 52.3 12.2 25. 7 0.23 0.49 28 ribs are numerous and separated by in- Hiromichi HIRANO

Fig. 1. Map showing the studied area and the fossil localities. terspaces narrower than the ribs. They characters of the young stage is not are strong on the outer half of the flank observable. Therefore I tentatively de­ and somewhat weak on the inner half. scribe it under Harpoceras cf. mulgravium Measurements :.-H: 46.1 (YOUNG & BIRD). Comparison:-The present specimen H. mulgravium (YOUNG & BIRD) is al­ is similar to the British specimens of lied to H. falcifer (SOWERBY) (PINNA, Harpoceras mulgravium (YOUNG & BIRD) 1968, pp. 37-39), but is distinguished in (WRIGHT, 1882, p. 433, BUCKMAN, 1909, its wider umbilicus, less involute shell p. 4b) in the mode of volution, venter and straight hafts of the ribs_ and ribs, but it is fragmentary and its Occurrence :-Loc. 28: 1. 611. Jurassic Toyora Group-II 7

Harpoceras (Harpoceras) sp. cf. H. (H.) falciform ribs and the involute shell. According to BUCKMAN (1909, p. 5b) three exaratum (YOUNG & BIRD) ontogenetic stages are discerned in H. PI. 2, Fig. 4; PI. 3, Fig. 1. (H.) exaratum, of which later two stages, "(2) the single ribs, falciform stage; (3) 1904. Harpoceras sp. ; YOKOYAMA, Jour. ColI. the finer rib stage, less falciform" are Sci., Imp. Univ. Tokyo, vol. 19, art. 20, confirmed in the Toyora specimens. As p. 15, pI. 4, fig. 2. the detailed nature of venter is not 1947. Harpoceras (s. str.) sp. aff. H. exaratum visible, I treat the present specimens (YOUNG & BIRD); MATSUMOTO in MA· as Harpoceras (H.) sp. ct. H. exaratum TSUMOTO & ONO, Sci. Rep. Fac. Sci., Kyushu Univ., Ceol., vol. 2, no. 1, p. 28. (YOUNG & BIRD). 1962. Harpoceras sp. aff. H. exaratum (YOUNG The present species is similar to Har­ & BIRD) ; SATO, Mem. Soc. Ceol. France, poceras (H.) okadai (YOKOYAMA) but is no. 94, p. 59. distinguished by its wider umbilicus and stronger curve and longer hafts of the Material:-Twelve specimens, GK. G. ribs. 2711, 2716, 11336-11341, GYU. M. 1025- Occurrence:-Loc. 28: 3, loco 2: 2, loco 1027, MM. 7071. 47: 3, loco 1: 1, loco 22: 1, loco 36: 1, loco Description :-The whorls enlarge rap­ 36': 1. idly and the umbilicus is narrow. The venter bears a less stout keel. Whether the venter has sulci or not is not con­ Subgenus Harpoceratoides firmable. The mode of ribs is falciform BUCKMAN, 1909 and is characteristic of Harpoceras (s. Type-species :-Ammonites alternatus str.). The ribs are usually simple. The SIMPSON in BUCKMAN, 1909. angle of the curved ribs near the middle of the flank is about 120· to 150·. The space of the ribs varies to some extent, Harpoceras (Harpoceratoides) nagatoensis being wider than the interspace in most sp. nov. [MATSUMOTO, MS.] specimens. Much finer ribs are occasion­ ally visible along the major ribs, and in PI. 3, Figs. 2-4. a large fragmentary specimen numerous 1947. Crammoceras (PleydeUia) sp.; MATSU­ fine ribs are characteristic. One speci­ MOTO in MATSUMOTO & ONO, Sci. Rep. men has its aptychus which may be re­ Fac. Sci., Kyushu Univ., Ceol., vol. 2, ferred to Cornaptychus. no. 1, p. 27, pI. 1, fig. 7. Measurements :- 1953. Harpoceratoides nagatoensis MATSUMO­ TO (nom. nud.); MATSUMOTO in MA­ No. D. U. H. UjD H/D Rn! TSUMOTO et al. ed., "Historical Geolo­ GK. G. 2711 ca.45 10.9 21. 1 0.24 0.47 40 gy" (in Japanese), Asakura Book Ltd., p. 359. Comparison: -The present specimens 1962. Harpoceratoides nagatoensis MATSUMO­ have been regarded as Harpoceras (s. TO; SATO (nom. nud.) Mem. Soc. Ceol. str.) sp. aft. H. exaratum (YOUNG & BIRD). France, no. 94, p. 59. Indeed the present specimens are very similar to the European specimens of Types:-The specimen illustrated· by Harpoceras (s. str.) exaratum (YOUNG & MATSUMOTO (in MATSUMOTO & ONO, BIRD) in the dense ribbing, the mode of 1947, pI. 1, fig. 7) is designated here ~s the 8 HirO?nichi HIRANO

Sandy shale, thick ly bedded, nol plaly, not so liss i!£', somewhat compact.

Silt-silty shale, thickly bedded, not platy, calcareous nodule bearing.

Sandy shal~. th ic kly bedded, sometimes massive.

Sandy shale. bedding plane IIat. thickly platy, sometimes massive. U' Silt-silty shale, !issile, generally platy.

Clay-silty clay shale, thinly bedded, lis sile like paper.

Alternation of shale and sand ston e.

Sa.ndy shale-shal)' sandsione.

Fine sandstone. Uh

Meium sandstone.

Coarse sandstone.

Ub ogogogo Conglomerate. 11-'"'' 0000

loc. 2& ir~:: Na - - loe. 27

loc.26 _ loc, f.!. - Nss - IDe. 25 Nm - loc. 21, _',,· 30 - lot. 23 _ "4 1 - loc.22 _-" 40 - - 7- - loc. 21 _ 10c: . 3.L?_ Nm loc. 20 - Nm _loc.~?_ - :g~ .' It- - loc.17 _ loc.32 Ncs __ tOC . 16 _loc. 3L­ ~-loc: . 29 Nsh -_._ .- 1 - 11-·-·.-· ll ~m. Na a e Fig. 2. Columnar sections showing localities. The horizontal bars are the boundaries of the formations (left side of the column) and the boundaries of the zones (right side). The locali ty numbers correspond to the numbers in the systematic description. a: Column along the sequence in the Sakuraguchi-dani; b: Column along the Anda-d ani; c: Column along the Kogo-dani; d : Column along the Chuzankei Ravine, Higashinakayama; e: Column along the River Era.

Explanation of Plate 1 (Natural size)

Figs. 1- 3. Harpoceras (s. str.) chrysanthemum (YOKOYAMA) 1. MM. 7058a, Lectotype, loco 47; 2. MM. 7059, Para lectotype, loco 47; 3. MM. 7060, Para lectotype, loco 47. Figs. 4-5. Harpoceras (s. str.) inouyei (YOKOYAMA) 4. MM. 7062, Lectotype, loco 47; 5. MM. 7063, Para lectotype, loco 47. Fig. 6. Harpoceras (s. str.) sp. d. H. muigravium (YOUN G & BIRD) GK. G. 11355, loco 28. Fig. 7. Dumortieria ? sp. Rubber cast of GK. G. 2797, loco 58. Fig_ 8. Cram11!oceras (s. str.) sp. aff. C. obesum (BUCKM AN) GK_ G. 2781, loco 55 . HIRANO: Jurassic Toyora Group-II Plate 1 611. Jurassic Toyora Group-II 9

holotype, GK. G. 2007, which was obtain­ weaker curve of ribs. ed at loco 9. Five specimens, GK. G. Occurrence :-Loc. 9: 4, loco 22: 1, loco 2381-2384, 2391, are designated as para­ 30: 1. types. Material:-The above mentioned six Genus Pseudolioceras BUCKMAN, 1889 specimens. Etymology:-"Nagato" is the old name Type-species:- Ammonites compactilis of the southwestern part of Yamaguchi SIMPSON in BUCKMAN, 1889. Prefecture. Diagnosis :-A species of Harpocera­ Pseudolioceras sp. to ides which has fine, numerous and PI. 3, Figs. 10-11. moderately curved ribs and proportion­ ally wide umbilicus. 1947. Hyperlioceras sp. ; MATSUMOTO in MA­ Description:-The whorls enlarge mod­ TSUMOTO & ONO, Sci. Rep. Fac. Sci., erately or somewhat rapidly. The volu­ Kyushu Univ., vol. 2, no. 1, p. 29, pI. tions are moderately tight in most of the 2, fig. 2. specimens and in others somewhat in­ 1956. Pseudolioceras? sp.; ARKELL, Jurassic volute. A keel is visible on the venter Geology of the World, Oliver & Boyd Ltd., p. 421. and may be unicarinate. The ribs are 1962. Pseudolioceras sp.; SATO, Mem. Soc. so fine that they look like lirae. Their Geol. France, no. 94, p. 59. curve is falciform and is characteristic 1963. Pseudolioceras sp. ; T AKAI, MATSUMOTO of Harpoceras. At inner one-third of the and TORIYAMA ed., Geology of Japan, flank, they curve backward moderately. Univ. Tokyo Press, p. 83. They are numerous, crowded and fasci­ culated on the inner lateral and the Material :-Ten unfavourably pre­ umbilical wall. They are wider than served specimens, GK. G. 2012, 2772- their interspace. The apertural part of 2776, 2782, 2792, 2793, 2798. a large specimen (PI. 3, Fig. 3) is cov­ Description:-The whorls enlarge very ered by numerous lirae instead of the rapidly and the umbilicus is very nar­ ribs. row. The venter is unicarinate-tabulate. Measurements :- The umbilical wall is low and more or less steep. The ribs are moderately \"0. D. U. H. U/D falciform. They spring on the umbilical GK. G. 2007 26. 0 8. 9 10.3 0.34 wall and go on somewhat forward. At GK. G. 2383 32. 4 14. 2 10.4 0.44 the middle of the flank they curve mod­ GK. G. 2382 58.6 22.9 21. 5 0.39 erately backward and then recurve for­ Comparison :-The present species is ward near the ventral shoulder. The similar to Protograml1loceras nipponicum ribs are wide and rather low. The in­ (MATSUMOTO) in its curvature of ribs terspaces of the ribs are also wide. In but the latter does not show the fasci­ one specimen, GK. G. 2773, lirae are pre­ culation of the ribs. sent on the ribs of the last whorl. The present species is similar to Har­ Aleasurements :- poceratoides serotinum BETTONI (1900, pp. No. D. U. U/D H. 65-66; HAAS, 1913, pp. 106-109; MONES­ GK. G. 2010 64. 1 1l.5 O. 18 33.9 TIER, 1931, pp. 31-32) but is distinguished GK. G. 2772 44.7 ca.9 0.20 24. 4 by its more evolute shell and rather GK. G. 2773 ca. 34. 7 ca. 10. 7 0.31 ca. 13.5 10 Hiromichi HIRANO Comparison:-The present species is ceras. The ribs curve backward at the similar to Pseudolioceras andromacoi middle of the flank. Usually they are FUCINI (1923, pp. 61-62) in the tight in­ very weak or sometimes imperceptible volution and falciform ribs, but is dis­ on the inner half of the flank but stout tinguished by its less dense ribs. It is on the outer half. They are widely in­ somewhat similar to Pseudolioceras sub­ terspaced. Lirae are present on the lythense (HAAS) (1913, pp. 113-114), but inner half of the flank. is distinguished by its wider ribs. Measurements :-Height of the whorl As the available specimens are un­ of the last stage: GK. G. 11351=28.1, favourably preserved, precise specific 11352=32.1,11353=24.8,11354=33.2, GYU. identification is hardly done. M. 1068=51.7. Occurrence :-Loc. 55: 8, loco 53: 1, loco Comparison:-The present specimens 54: 1. are similar to the British specimens of Hildoceras bifrons (BRUGUIERE) (WRIGHT, Subfamily Hildoceratinae HYATT, 1867 1882, p.436, pi. 59, figs. 1-2) in the mode of volution, venter and stout ribs, but Genus Hildoceras HYATT, 1867 the curvature of ribs in the former is Type-species:-Ammonites bifrons BRu­ not so abrupt as in the latter. In the GUIERE, 1789 (subsequently designated by present specimens the inner part of the BUCKMAN, 1889). flank is not grooved but flattened and this may be a diagnostic character of Hildoceras sp. aff. H. bifrons (BRUGUIERE) the present specimens. Because the character of the small stage is unknown PI. 4, Figs. 4-5; PI. 2, Fig. 10 and only the fragmentary specimens are in Part III. available, I tentatively treat the present Material.·-Five specimens, GYU. M. specimens under Hildoceras sp. aff. H. 1068, GK. G. 11351-11345, of incomplete bifrons, although they could possibly preservation. represent a new species. Description:-The whorls enlarge Occurrence :-Loc. 7: 1, loco 25: 1. loco slowly and the umbilicus is proportion­ 27: 1, loco 40: 1, loco 44: 1. ally wide. The venter is tricarinate­ bisulcate. The umbilical periphery is Subfamily Grammoceratinae rounded but the wall is nearly perpen­ BUCKMAN, 1904 dicular and somewhat high. Therefore the whorl-section is roughly rectangular. Genus Grammoceras HYATT, 1867 The flank is flat. The curvature of a Type-species :-Ammonites striatulus rib is falcoid and characteristic of Hildo- SOWERBY, 1823 (subsequently designated

Explanation of Plate 2 (Natural size)

Figs. 1-3. Harpoceras (s. str.) chrysanthemum (YOKOYAMA) 1. MM. 7065, loco 47; 2. MM. 7061, Para lectotype, loco 47; 3. MM. 7064, Rubber cast, Nishinagano Fig. 4. Harpoceras (s. str.) sp. d. H. exaratum (YOUNG & BIRD) MM. 7071, loco 47. HIRANO: J~~TaissiG ToyoTa Group-II Plate 2

1 611. Jurassic Toyora Group-II 11 by BUCKMAN, 1890). Genus Dumortieria HAUG, 1885

Subgenus Grammoceras HYATT, 1867 Type-species :-Ammonites levesquei O'ORBIGNY, 1884 (subsequently designat­ Grammoceras (Grammoceras) sp. aff. ed by BUCKMAN, 1890). G. (G.) obesum (BUCKMAN)

PI. 1, Fig. 8. Dumortieria? sp.

1947. Cf. Crammoceras sp.; MATSUMOTO in PI. 1, Fig. 7. MATSUMOTO & ONO, Sci. Rep. Fac. Sci., (?) sp.; MATSUMOTO Kyushu Univ., Ceol., vol. 2, no. 1, p. 27. 1947. Dumortieria in MATSUMOTO & ONO, Sci. Rep. Fac. Sci., Material :-Five specimens, GK. G. 2781, Kyushu Univ., Ceol., vol. 2, no. 1, p. 29. 2783, 2787, 2784, 2795. 1956. ? Dumortieria sp.; ARKELL, Jurassic Description:-The whorls enlarge mod­ Geology of the World, Oliver & Boyd Ltd., p. 421. erately and the venter has a small keel. 1962. Dumortieria? sp. ; SATO, Mem. Soc. Ceol. The umbilical periphery is rounded and France, no. 94, p. 59. the wall is low. The ribs are moderately falcoid. They spring on the umbilical Material:-A single fragmentary spe­ seam and go on forward with flexuosity. cimen, GK. G. 2797. After they proceed one-fourth or one­ Description :-The whorls enlarge third of the flank, they curve backward somewhat rapidly and consequently the and recurve forward near the ventral umbilicus is rather narrow. The venter shoulder. They are simple, somewhat is invisible. The ribs, which spring at elevated and separated by wider inter­ the umbilical seam, are somewhat flexu­ spaces. ous, slightly falcoidal, fine, simple and Measurements :- widely interspaced. Measurements :- No. D. U. U/D H. Rn! GK. G. 2781 59. 4 20. 4 O. 34 22. 7 ca. 21 No. D. U. U/D Rn! GK. G. 2797 ca. 18 ca. 7 ca. 0.39 17 Comparison:-The present species re­ sembles Grammoceras obesum (BUCKMAN) Comparison:-The present species (1904, pp. 150-151) in the mode of ribs, somewhat resembles Dumortieria tabu­ but has a tighter volution and somewhat lata BUCKMAN (1905, p. 185) in the mode stronger curvature of ribs. of ribs and volution, but it has finer The present species is somewhat simi­ ribs. It is similar to Dumortieria leves­ lar to Grammoceras thouarsense (O'ORBIG­ quei (O'ORBIGNY) (HAUG, 1885, pp. 662- NY) in the curvature of ribs, but is dis­ 663) in the somewhat fine and widely tinguished by its tighter volution and interspaced ribs, but has more flexuous sparser ribbing. ribs. On the other hand it is apparently As the available specimens are not similar to Tmetoceras in its simple and numerous and unfavourably preserved, fine ribs. Because only a single speci­ I provisionally treat the present species men with poorly preserved venter is as Grammoceras sp. aff. G. obesum (BUCK­ available, a precise identification cannot MAN). be done. Occurrence :-Loc. 55: 5. Occurrence :-Loc. 58: 1. 12 Hiromichi HIRANO

Family Hammatoceratidae 2764, 2765, 2766, 2768, 2768, 2770, 2771, BUCKMAN, 1887 2777, as paralectotypes, of which one specimen, GK. G. 20n was previously Subfamily Phymatoceratinae illustrated by MATSUMOTO (1947, pI. 2, HYATT, 1867 fig. 1). Material :-The above mentioned Genus Phymatoceras HYATT, 1867 twelve specimens. Diagnosis :-A species of Phymatoceras Type-species:-Phymatoceras robustum which has slightly to moderately sig­ Hy A TT, 1867. moidal, widely interspaced ribs and no tubercles. Phymatoceras toyoranum (MATSUMOTO) Description:-The whorls enlarge somewhat rapidly and consequently the PI. 4, Fig. 3. umbilicus is narrow in the young stage of about 20 mm. in diameter. In the 1947. Lillia toyorana MATSUMOTO; MATSU­ MOTO in MATSUMOTO & ONO, Sci. Rep. later stages the whorls enlarge some­ Fac. Sci., Kyushu Univ., Ceoi., vol. 2, what slowly and the umbilicus is pro­ no. 1, p. 29, pI. 2, fig. 1. portionally moderate. The venter is 1956. Phymatoceras toyoranum (MATSUMOTO); unicarinate-bisulcate. The ribs spring ARKELL, Jurassic Geology of the World, at the umbilical seam and go on forward. Oliver & Boyd Ltd., p. 421. At a point inner one-third of the flank 1962. Phymatoceras toyoranum (MATSUMOTO); they curve backward and near the ven­ SATO, Mem. Soc. Ceoi. France, no. 94, tral shoulder they recurve forward. p. 59. They are, thus, slightly sigmoidal and 1962. Phymatoceras toyoranum (MATSUMOTO); they become stouter toward the ventral TAKAI, MATSUMOTO and TORIYAMA ed., shoulder. They are interspaced widely Geology of Japan, Univ. Tokyo Press, p. 83. and often bifurcate at the umbilical periphery. In some specimens the ribs Types :-Here I designate one of the are slightly elevated at the umbilical twelve syntypes, GK. G. 2763, as the periphery where the bifurcation arises, lectotype and others, GK. G. 2011, 2761, but the elevation is not so strong as to

Explanation of Plate 3 (Natural size) Fig. 1. Harpoceras (s. str.) sp. d. H. exaratum (YOUNG & BIRD) Rubber cast of GK. G. 2711, loco 2. Figs. 2-4. Harpoceras (Harpoceratoides) nagatoensis sp. nov. 2. GK. G. 2383, Para type, loco 9; 3. GK. G. 2381, paratype, loco 30; 4. GK. G. 2007, Holotype, loc. 9. Fig. 5. Harpoceras (s. str.) inouyei (YOKOYAMA) GK. G. 2642, loc. 3. Figs. 6-9. Harpoceras (s. str.) chrysanthemum (YOKOYAMA) 6. GK. G. 2572, loc. 3; 7. GK. G. 2552, loc. 3; 8. GK. G. 2557, lac. 3; 9. GK. G. 250-1, loco 3. Figs. 10-11. Pseudoiioceras sp. 10. GK. G. 2773, loc. 55; 11. GK. G. 2012, loc. 55. HIRANO: Jurassic Toyora Group-II Plate 3 611. Jurassic Toyora Group-II 13

be called tubercles. One specimen, GK. Oliver & Boyd Ltd., p. 421. G. 2763, with a diameter of 64 mm., has 1962. Haugia aff. japonica (NEUMAYR) ; SATO, some lirae on the apertural part. Mem. Soc. Ceol. France, no. 94, p. 59. Measurements :- Material:-Two specimens, GK. G. 2801, No. D. U. UjD Rn! 11350. GK. G. 2763 ca. 61 21.6 ca. 0.35 20 Description:-The whorls enlarge !'Ilowly and consequently the umbilicus Comparison:-The present species is IS proportionally wide. The venter has similar to Phymatoceras pulcher MERLA a low keel. The presence of sulci is not (1933, pp. 30-31) in the mode of volution confirmed. The umbilical periphery is and the curvature of ribs, but is distin­ rounded and the wall is steep. Some­ guished by its stouter ribs and indis­ what prominent tubercles are present tinctness of tubercles at the umbilical between the umbilical wall and the um­ periphery. bilical shoulder, and two ribs spring The present species is similar to Phy­ from every tubercle in the growth stages matoceras malagma (DUMORTIER) (BUCK­ less than 50 mm. in diameter. The ribs MAN, 1899, p. 21) in the curvature of ribs, go on backward on the flank and near the bifurcation and the volution. The the ventral shoulder they curve forward. unmistakable distinction between the In the late growth stage with diame­ two species is the considerably dominant ters between ca. 50 mm. and ca. 100 mm., tubercles in the latter. the tubercles are weak or very weak, The present species resembles Phy­ and the twin becomes obscure. The ribs matoceras obtecta (BUCKMAN) (1899, p. 21) are sigmoid, very weak on the inner in the mode of ribs. The latter is dis­ one-third of the flank and somewhat tinguished in its more dominant and strong on the ventral shoulder. The numerous tubercles in younger stages curvature of the rib in this stage is the and more rapidly enlarging whorls. same as that in the younger stages. In The present species is also similar to one specimen, GK. G. 2801, lirae occur Phymatoceras narbonensis (BUCKMAN) in the part with diameters between ca. (MONESTIER, 1931, p. 21) in the mode of 70 mm. and ca. 85 mm. and they are bifurcation. They are distinguished in prominent on the umbilical shoulder. that the ribs are more flexuous and con­ The tubercles and ribs become very siderably elevated at the umbilical pe­ weak and finally fade away in the last riphery and the whorls enlarge more stage over 100 mm. in diameter. rapidly in the latter than in the former. Measurements :- Occurrence:-Loc. 55: 13. No. D. D. H. DID Rn! GK. G. 2801 ca.83 38. 9 ca. 23. 7 ca. O. 47 ca. 27 Phymatoceras sp. GK. G. 11350 - 54. 8 30. 9 25 PI. 4, Figs. 1-2 Remarks:-The present species has been considered as being allied to 1947. Haugia aff. japonica (NEUMAYR); MA­ Haugia japonica (NEUMAYR) from Shi­ TSUMOTO in MATSUMOTO & ONO, Sci. Rep. Fac. Sci., Kyushu Univ., Ceol., vol. koku for a long time. The taxonomic 2, no. 1, p. 29, pI. 2, fig. 5. position of Haugia japonica has so far 1956. Haugia aff. japonica (NEUMAYR); AR­ been discussed by many students (NEU­ KELL, Jurassic Geology of the World, MAYR in NAUMANN & NEUMAYR, 1890, 14 Hiromichi HIRANO p. 35; YEHARA, 1927, pp. 37-39; SHIMIZU, The present species is not referred to 1928, pp. 219-222; KOBA YASH!, 1935, pp. the genus Haugia, because it has a low 79-83) and recently ZEISS (1956, p. 76) keel and sigmoid ribs. pointed out that it is closely allied to Considering all the observable charac­ Hecticoceras (Putealiceras) mathayense ters it is better to ascribe the present (KILIAN). HAY AMI & MATSUMOTO (MA­ species to Phymatoceras. TSUMOTO et al., 1963, p. 35) regarded it Comparison:-The present species is as Hecticoceras? (Putealiceras) japonicum. similar to Phymatoceras pulcher MERLA H.? (P.?) japonicum has less sigmoidal (1933, pp. 30-31) in the mode of volution ribs and tubercles at a point about one­ and ribs, but is distinguished by its less third from the umbilical margin to the persistent tubercles. venter, but the present species has more The present species is similar to Phy­ sigmoid ribs and tubercles on the um­ matoceras robustum HYATT (ARKELL et bilical shoulder. There are unmistak­ al., 1956, p. L265) in the mode of tuber­ able differences between them. cles and volution. In the former, twin­ Although the present species is repre­ ned ribs arises from every tubercle in sented only by two specimens, it is not the younger stages. On the other hand, referred to the genus Hecticoceras by in the latter the ribs are simple, twin­ the following ground. Ribs are strong ned or triploid from tubercles and the in Hecticoceras but moderate Or weak in mode is irregular. the present species. In Hecticoceras the The present species is similar to Phy­ ribs end at the ventrolateral tubercles, matoceras toyoranum in the mode of vo­ but such a character is not shown by lution and ribbing, but is distinguished the present species. Hecticoceras has by its dominant tubercles at the umbili­ tubercles through the growth stage, but cal shoulder in the younger stages and in the present species the tubercles are somewhat weaker ribs. weakened in a late stage with diameters Occurrence :-Loc. 56: 2. between 50 to 100 mm. and finally fade away. (to be continued)

Explanation of Plate 4 (Natural size unless otherwise indicated) Figs. 1-2. Phymatoceras sp. 1. Gypsum cast of GK. G. 11350, lac. 56, x 0.8; 2. Gypsum cast of GK. G. 2801, lac. 56, xO.8. Fig. 3. Phymatoceras toyoranum (MATSUMOTO) Rubber cast of GK. G. 2763, Lectotype, lac. 55. Figs. 4-5. Hildoceras (s. str.) sp. aff. H. bi/rons (BRUGUIERE) 4. GK. G. 11351, lac. 40; 5. GK. G. 11354, lac. 27. HIRANO.' J~wassic ToyoTa G'l'"oup-III Plate 4-

5 Trans. Proc. Palaeont. Soc. Japan, N.S., No. 89, pp. 15-23, pIs. 5, 6, April 20, 1973

612. CORAL FOSSILS FROM THE KOSODE FORMATION, YAMAN ASHI PREFECTURE*

KATSUHIKO NISHIMIY A

Institute of Geosciences, Yamanashi University

and

NOBUO Y AMAGIW A

Institute of Geosciences, Osaka Kyoiku University

IlI~WL IHm~I\'lO)-if' /' ::I'1I:E: llI~g\'l;O)~t*i1ffiH;::n-1nT Q/Hdl~~ip!?:>ilIl±', L.t.: -if' /' ::I'1I:Em~tUjL., 10fm~~)jIJL.1lc.iI!1(L.t.:o .:(-0)$ 2{11HHfrflli'"C-if'/'::I·1I:Ef,'F O)t1lf1lX:~*0)~ "'Lff'1~L.t.:o i1§1.n\i:~· llIr.r,ul!;*:

The coral fauna may be regarded as Introduction and acknowledgements very valuable in order to discuss the geological age, correlation and palaeo­ The Kosode Formation is a character­ environment of the Kosode Formation, istic member composing the zonal struc­ so far as other leading fossils are indeed ture in the Kanto-Mountainland, and is lacking. Ten coral species are dis­ extending from Kosode toward Aoiwa­ criminated from the formation, and de­ dani in the surveyed area, the north­ scribed in the present paper. In addi­ eastern part of Yamanashi Prefecture tion, the Jurassic Torinosu Series is (Text-fig. 1). The stratigraphical and briefly mentioned, making special ref­ structural studies of the formation have rence to the Kosode Formation. been carried out by FUJIMOTO (1939), On publishing the present paper, the FUJIMOTO and SUZUKI (1969), Yamanashi writers wish to express their sincere Prefecture Government (1970), and NISHI­ thanks to Professor Keiji NAKAZAWA MIY A and Y AMAGIW A (1971). There has of the Kyoto University for his contin­ been, however, published no palaeonto­ uous suggestions. Thanks are due to logical report, although corals are rather Mr. Toyohiko Iw AHASHI of the Osaka dominant in limestone lenses at Kosode Kyoiku University for his photographic and Aoiwadani, and occur sometimes in work, and also due to Mr. and Mrs. association with stromatoporoids. Yasuo NOGAMI of the Kyoto University * Received Oct. 20, 1972; read June 27, for their helps in preparation for the 1971, at Nara. manuscript. 15 16 Katsuhiko NISHIMIYA and Nobuo Y AMAGIWA

N IIiiI ItXfU)\1 II ~III ~IV \ ~v 1._1 VI

_-- VII

VIII

o 1000 2000m L'__ --'-, __ ---',

Kobotoke Group

Fig. 1. Geological map of the northeastern part of Yamanashi Prefecture. A: Kumotoriyama Formation, B: Kosode Formation, C: Aoiwadani Formation, D: Kamo­ sawa Formation, E: Oonari Formation, I: shale or slate, II: sandstone, III: conglomerate, IV: schalstein, V: chert, VI: limestone, VII: fault, VIII: strike and dip, 1-21; Sample loco

and by FUJIMOTO and SUZUKI (1969) in Stratigraphy the Oborogawa District. They are sub­ The Jurassic strata were studied by divided as follows: FUJIMOTO (1939) in the Kogochi District

Kogochi District Oborogawa District Kogochi Group (Jurassic) Otaki Group (Jurassic) Takanosuyama Formation Tochimoto Formation Kawamata Formation Kosode Formation Sogoyadani Formation Kosode Formation Oonari Formation Koreisan Formation

The stratigraphy of the surveyed Torinosu Series is separately distrib­ Aoiwadani-Kosode area is summarized, uted in three belts, namely, the Oonari as shown in Table 1. The Jurassic Formation in the southern belt, the 612. Corals from Kosode Formation 17

Table 1. Stratigraphical subdivision in the surveyed area.

Aoiwadani-Kosode Area

-~------~,------I Cretaceous Shimanto Series Kobotoke Group ------1------(fault) ----,------1 Jurassic Torinosu Series Kosode F. I Kamosawa F. Oonari F. 1- (fault) _-'______~ Triassic? Aoiwadani Formation

(fault) --~------I Permian Chichibu Series Kumotoriyama Formation

Kamosawa Formation in the middle mation and the presumably Triassic belt, and the Kosode Formation in the Aoiwadani Formation is marked by a northern belt. The boundaries of these fault, which is cut off by NE-SW faults. formations are bounded by fault. The Kosode Formation: Typically The Oonari Formation: Typically exposed along the upper stream of the developed at Oonari, well cropping out Kosodegawa River and at Aoiwadani; along the middle stream of the Taba­ composed of slate (often changed into gawa River; mainly composed of sand­ phyllite) and sandstone, occasionally stone, shale and slate, subordinately intercalating blackish grey limestone made up of schalstein and chert, inter­ lenses, which yield hexacorals and calating blackish grey limestone lenses; stromatoporoids; about 500 m thick. about 2000 m thick. The formation showing a general The formation showing a general trend of NW -SE, dipping to northeast trend of NW -SE, dipping to northeast at angles 40-70". A considerable num­ and southwest at angles 50-80°, general­ ber of faults trending to NW-SE and ly characterized by homoclinal struc­ NE-SW. The formation is bounded on ture. Several faults trending to NW-SE the north by the Permian Kumotori­ and NE-SW. The boundary between yama Formation and on the south by the present formation and the Kamo­ the Aoiwadani Formation. These for­ saw a Formation in the middle belt is mations are in contact with each other marked by fault, which is cut off by by faults, which are cut off by NE-SW NE-SW faults. faults. The Kamosawa Formation: Typically The limestone lenses intercalated in cropping at Kamosawa; mainly com­ the Kosode Formation yield corals, as posed of shale and slate, intercalating shown in Table 2. sandstone, thin limestone and chert; The fauna of corals listed in Table 2 about 1200 m thick. are equivalent or similar to those from The formation showing a strike of the Upper Jurassic in Japan, and the N45-800W, dipping to northeast or south­ Kosode Formation can be safely cor­ west at angles 40-80°. Several faults related with the Torinosu Series in the trending to NW-SE and NE-SW. The Sagawa basin, Southwest Japan. The boundary between the present for- Kamosawa Formation and the Oonari 18 Katsuhiko NISHIMIY A and Nobuo Y AMAGIWA

Table 2. List of the Jurassic corals from the Kosode Fcrrmation.

Aoiwadani Kosodegawa ~ "',..... ::=lii ~ 0> 0 ...... N M ~ In ...... N N (fl-"'''0 Species o::l U U U u .- 0 u u u u 0 0 0 0 '<::.0 0 0 0 0 ..4 ..4 ..4 ..4 [/) '-J ..4 ..4 ..4 ..4 ~ ------Actinastrea? sp. indet. x ------Caiamophyllia sp. indet. x x x ------Dimorphoseries tabayamensis x ------Enallhelia sp. indet. x x ------I sastrea? sp. indet. x ------Stylina kantoensis x ------Stylina? sp. indet. x ------Thamnasteria huzimotoi x x ------T. aff. huzimotoi x ------T. sp. A x ------T. sp. B x ------Trocharea sp. indet. x ------Hexacoral gen. et sp. indet. x x

Formation, from which any valuable Genus Thamnasteria LESAUVAGE, 1823 fossil has never been obtained, are tentatively regarded as the Jurassic, Thamnasteria huzimotoi EGUCHI judging from their lithic resemblance Plate 5, figs. 2a-c to the well ascertained Torinosu Series in the surveyed area. The details of 1951. Thamnasteria huzimotoi EGUCHI, Sci. the correlation will be discussed by the Rep. Tohoku Univ., 2nd ser. (Geol.), writers in the near future. vol. 24, p. 79, pI. 26, fig. 10. Corallum massive and thamnasterioid. Corallites small and confluent, without Systematic description distinct wall; central distance 2.0-4.0 Order Scleractinia BOURNE, 1900 mm. Septa slightly sinuous, confluent with those of the adjacent corallites, Suborder Astrocoeniina VAUGHAN 30-45 in number, 10-13 of them extend­ & WELLS, 1943 ing to center; laterally by synapticulae. Perforations generally scarce. Col­ Family Thamnasteriidae VAUGHAN umella styliform, but in some corallites & WELLS, 1943 rudimentary. 612. Corals from Kosode Formation

Comparison: The present form is Compariso'n: The present fonn is closely allied to the type specimen of very similar to Thamnaster.ia huzimotoi Thamnasteria huzimotoi EGUCHI (1951, p. EGUCI-II (1951, p.79, pI. 26, fig. 10; NISHI­ 79, pI. 26, fig. 10) from the Torinosu MIYA & Y AMAGIWA, the present paper) group in Itsukaichi, Kanto-Mountainland from the Upper Jurassic in the Kanto­ in many common characters. However*, Mountainland, but differs from the the former's perforations are more latter in having less numerous syn­ scarce than the latter's. It is similar apticulae and longer central distance. to T. torinosuensis EGUCHI (1951, p. 67, It resembles T. torinosuensis EGUCHI pI. 24, fig. 9; MORI, 1963, p. 58, pI. 22, (1951, p. 67, pI. 24, fig. 9; MORI, 1963, p. fig. 3) from the Upper Jurassic in the 58, pI. 22, fig. 3). However, the former Sagawa basin and the Soma area, but has more numerous septa. the former differs from the latter in Occurrence: Loc. 2, Aoiwadani, Taba­ having more numerous septa. It re­ yama-mura, Kitatsuru-gun, Yamanashi sembles T. crateriformis GREGORY, (1900, Prefecture. pp. 135-136, pI. 17, figs. 4, 5, 7) from the Geological horizon: Upper Jurassic. Upper Cutchum formation in Cutch in many points. However, the former can be distinguished from the latter by its Thamnasteria sp. B indet. smaller coralli tes and less numerous Plate 5, figs. la, b septa. Occurrence: Loc. 2 at Aoiwadani and Corallum thamnasterioid. Corallites Loc. 20 at Kosodegawa, Tabayama­ relatively large and united by septa, mura, Kitatsuru-gun, Yamanashi Prefec­ about 8.0 mm between calicural centers. ture. Septa slightly sinuous and less nu­ Geological horizon: Upper Jurassic. merous (20+?); perforate, about 12 of them extending to center. Columella distinct and styliform. Synapticulae Thamnasteria sp. A indet. present. No longitudinal section. Plate 5, figs. 3a, b Comparison: The present species somewhat resembles Thamnasteria abu­ Corallum massive and thamnasterioid. kumaensis MORI (1963, p. 58, pI. 21, figs. Corallites somewhat irregular in size, 6, 7) from the Upper Jurassic in the withou t distinct wall. Corallite centers Soma area. However, the former is 3.5-6.0 mm. Septa gently flexous, some­ separable from the latter in having what perforate, confluent between coral­ perforate septa and a styliform col­ lites. Corallites 30-45 septa; 11-15 of umella. them (first and second cycles) extending Occurrence: Loc. 5, Aoiwadani, Taba­ to center and some of them joint with a yama-mura, Kitatsuru-gun, Yamanashi columella. Synapticulae present. Col­ Prefecture. umella styliform, but in some corallites Geological horizon: Upper Jurassic. rudimentary. No longitudinal section.

* According to the writers' observations, Family Astrocoeniidae KOBY, 1890 the septa of the type specimen are about 40 in number. Subfamily Astrocoeniinae KOBY, 11890 20 Katsuhiko NISHIMIYA and Nobuo YAMAGIWA

Genus Actinastrea D'ORBIGNY 1849 Genus Stylina LAMARCK, 1816

Actinastrea ? sp. indet. Stylina kantoensis, sp. nov.

Plate 6, fig. 6 Plate 6, fig. 1

Corallum massive. Corallites sub- Corallum massive. Corallites circular polygonal in transverse section, united in transverse section, subequal in size; by wall; generally 1.2-1.5 mm in dia­ generally 1.3-1.5 mm in diameter. Coral­ meter. Septa about 24, straight; 6 of lites united by coastae. Corallite cen­ first cycle long and reach to columella, teres 2.2-3.0 mm. Septa thick, 12 in 6 of second cycle slightly shorter than number, arranged in two hexameral cy­ the first and reach to near the columella. cles; 6 of first cycles extending to near Septa of third cycle slightly developed. columella. Columella styliform, circular No longitudinal section. in transverse section; 0.2-0.3 mm in dia­ Comparison: The present form is meter. Wall thick. Intercorallite area represented by a single transverse sec­ well developed, formed of costae. Cos­ tion. It is similar to the species belong· tae sub equal, 24 in number. No longi­ ing to the genus Stylina, but differs tudinal section. from the latter in having subpolygonal Comparison: The present form re­ corallites. It resembles Astrocoenia sp. sembles Stylina sugiyamai EGUCHI (1951, (MORI, 1963, p. 56, pI. 21, fig. 8) from p. 60, pI. 22, fig. 7) from the Upper the Upper Jurassic in the Soma area, Jurassic Torinosu group in the Sagawa Northeast Japan. However, the calicural basin, Southwest Japan, but differs from wall of the former is subpolygonal in the latter, having longer central dis­ transverse section and the corallites of tance. It can be distinguished from S. the former are not well preserved. higoensis EGUCHI (1951, p.74, pI. 12, figs. Occurrence: Loc. 3, Aoiwadani, Taba· 6-9) from the Sakamoto formation in yama-mura, Kitatsuru-gun, Yamanashi Kuma Mountainland, southwest Japan in Prefecture. having broader corallites and longer Geological horizon: Upper Jurassic. central distance. Occurrence: Loc. 20, Kosodegawa, Family Stylinidae D'ORBIGNY, 1851 Tabayama-mura, Kitatsuru-gun, Yama­ nashi Prefecture. Subfamily Stylininae D'ORBIGNY, 1851 Geological horizon: Upper Jurassic.

Explanation of Plate 5

All figs. x3

Fig. 1. Thamnasteria sp. B indet. a) Transverse section, reg. no. 72092401a; b) Longitudinal section, reg. no. n092401b. Fig. 2. Thamnasteria huzimotoi EGUCHI a) Transverse sectioh, reg. no. 72092402a; b) Longitudinal section, reg. no. 72092402b; c) Transverse (somewhat oblique) section, reg. no. 72092403. Fig. 3. Thamnasteria sp. A indet. a) Transverse section, reg. no. 72092404; b) Transverse section, reg. no. 72092404. NISHI MIY A and YAMAGI WA: Corals !1"Om Kosode Fo'rmation Plate 5

lb 612. Corals from Kosode Formation 21

Subfamily Euheliinae numerous, 19 of them extending to center. Secondary (marginal) calice only DE FROMENTEL, 1861 one. In this calice, about 12 of septa Genus Enallhelia EDWARDS extending to center. Septa slightly sinuous and somewhat perforate. Dis­ & HAl ME, 1850 tance of primary and secondary centers Enallhelia sp. indet. about 6.5 mm. Wall relatively thin. Col­ umella trabecular in central calice, but Plate 6, figs. 7a, b rudimentary in marginal one. Syn­ apticulae numerous and distinct. The present species almost agrees Comparison: The present species re­ with the genus Enallhelia in many sembles Dimorphoseris oolitic a DUNCAN important characters. It is closely (1872, pt. 3, pp. 22-23, pi. 4, figs. 1-4) from similar to E. nipponica somaensis EGU­ the Oolites series in Europe in many CHI (1942, pp. 143-144, pi. 6, figs. 8, 9; common characters. However, it differs MORI, 1963, pp. 56-57, pi. 23, fig. 3) from from the latter in having smaller coral­ the Upper Jurassic in the Soma area in lum, trabecular columella and less nu­ many common characters. However, it merous septa extending to columella. is separable from the latter in having Occurrence: Loc. 5, Aoiwadani, Taba­ a compressed columella. It is related to yama-mura, Kitatsuru-gun, Yamanashi E. nipponica EGUCHI (1942, pp. 142-143, Prefecture. pi. 6, figs. 4-7) from the Lower Creta­ Geological horizon: Upper Jurassic. ceous in Miyagi Prefecture, but the former's calices are smaller than the latter's. Occurrence: Loc. 20 at Kosodegawa Superfamily Agariciicae GRAY, 1847 and a boulder at Shiosawa, Tabayama­ Family Calamophyllidae VAUGHAN mura, Kitatsuru-gun, Yamanashi Prefec­ ture. & WELLS, 1943 Geological horizon: Upper Jurassic. Genus Calamophyllia BLAINVILLE, 1830

Calamorphyllia sp. indet.

Suborder Fungiina VERRILL, 1865 Plate 6, fig. 3

Superfamily Fungiicae DANA, 1846 The present form is not well pre­ Family Synastreidae ALLOITEAU, 1952 served. It may belong to the genus Calamophyllia" than the genus Stylosmilia Genus Dimorphoseris DUNCAN, 1872 in having no styliform columella. It resembles Rhabdophyllia Qr Stylosmilia Dimorphoseris tabayamensis, sp. nov. (EGUCHI, 1951, pi. 23, fig. 5) from the Plate 6, figs. 2a, b Upper Jurassic Torinosu group at Oga­ wa, Ogawa-mura, Takaoka-gun, Kochi Coralum thamnasterioid and turbi­ Prefecture in many respects. However, nate; about 18 mm in diameter. Colony the former has somewhat larger coral­ formation by circumoral budding. There lites than the latter. is a large central primary calice; septa Katsuhiko NISHIMIYA and"Nobuo YAMAGIWA

Occurrence: ~oc. 2 and 4 at Aoiwa­ Superfamily Poriticae GRAY, 1842 dani ?-nd Loc. 20" ~t Kosodegawa, Taba­ yama-mura, Kitatsuru-gun, Yamanashi Family Microsolenidae KOBY, 1890 Prefecture. Geological horizon: Upper Jurassic. Genus Trocharea ETALLON, 1864 Trocharea sp. indet.

Genus Isastrea EDWARDS Plate 6, fig. 5 & HArME, 1851 Corallum simple. Calicural diameter about 15mm in transverse section. Thin I sastrea ? sp. indet. wall developed, although it is not rec­ Plate 6,fig. 4 ognizable in part. Septa numerous, 70- 80 in number. Septa slightly sinuous The present specimens may belong to in part and geniculate in part, with per­ the genus Isastrea in having the follow­ forations. Columella trabecular. Syn­ ing characters. 1) cerioid colonies, 2) apticulae present but a few. No longi­ corallite wall present, but with septa tudinal section. confluent partly absent, 3) monocentric Comparison: The present form is corallites, but rarely dicentric, 4) tra­ closely related to Trocharea sp. (EGUCHI, becular columella, but feeble. It is simi­ 1951, pI. 25, fig. 1) from the Torinosu lar to some species of the genus Lato­ group at Ogawa, Ogawa-mura, Sagawa mtwndra. "The corallites of the latter, basin, Southwest Japan in having however, are mostly di- or tricentric. many common" characters. However, Occurrence: Loc. 2, Aoiwadani, Taba­ the former one has less numerous septa yama-muni, Kitatsuru-gun, Yamanashi than the .latter. Prefecture. Occurrence: Loc. 2, Aoiwadani, Taba­ Geological horizon: Upper Jurassic. yama -mura, Kitatsuru -gun, Yamanashi

Explanation of Plate 6

Fig. 1. Stylina kantoensis, sp. nov. Transverse section of the holotype, x 3, reg. no. 72092405. Fig. 2. Dimorphoseris tabayamensis, sp. nov. a) Transverse section of the holotype, x 3, reg. no. 72092406a; b) Longitudinal section of the holotype, x 3, reg. no. 72092406b. Fig. 3~ Calamophyllia sp. indet. Transverse section, x 3, reg. no. 72092407. Fig". 4. Isastriw? sp. indet. Transverse section, x 3, reg. no. 72092408. Fig~ 5. Trocharea sp. indet. Transverse section, x 3, reg. no. 72092409. Fig. 6. Actinastrea? sp. indet." Transverse section, x 5, reg. no. 72092410. Fig. 7. Enallhelia sp. indet. a) Transverse secfion, x 5, reg. no. 72092411; b) Transverse section, x 5, reg. no. 72092412. NISHIMIY A and YAMAGIWA: Corals from Kosode Formation Plate 6 612. Corals from Kosode Formation 23

Prefecture. ries of the Kwanto Mountain. jub. Publ. Geological horizon: Upper Jurassic. Comm. Prof. Yabe's 60th Birthday, vol. 1, pp. 457-479. All the figured specimens are collected -- & SUZUKI, M. (1969): Geology of the by Katsuhiko NISHIMIY A, and are de­ basin of the River Oboro-gawa, a tribu­ posited at the Institute of Geoscience, tary of the River Arakawa. Bull. Chi­ Osaka Kyoiku University; reg. nos. chibu Mus. Nat. Hist., no. 15, pp. 1-18. GREGORY, J. W. (1900): Jurassic fauna of 72092401 to 72092412. Cutch. Ceol. Surv. India, ser. 9, vol. 2, pt. 2, pp. 1-195, pIs. 2a-27. References MORI, K. (1963): Geology and Palaeontology of the Jurassic Somanakamura Group, DUNCAN, P. M. (1872): A monograph of Fukushima Prefecture, Japan. Sci. Rep. British fossil corals. Palaeontogr. Soc., Tohoku Univ., 2nd ser. (Ceol.), vol. 35, London, Monogr., 2nd ser., pt. 3, pp. 1-24, no. 1, pp. 33-65, pIs. 21-23. pIs. 1-7. NISHlMIY A, K. & YAMAGIW A, N. (1971): EGUCHI, M. (1924): Recent and fossil corals Discovery of coral fossils from the Ao­ of the family Oculinidae from Japan. yuwadani, Tabayama-mura, Yamanashi Ceol. Soc. japan, jour., vol. 49, no. 583, Prefecture. Ceol. Soc. japan, Jour., vol. pp. 135-l42, pI. 6 (5). 77, no. 12, pp. 791-792. (in Japanese) - (1951): Mesozoic Hexacorals from Ja­ Yamanashi Prefecture Government (1970): pan. Sci. Rep. Tohoku Univ., 2nd ser. Geologic map of Yamanashi Prefecture (Ceol.), vol. 24, pp. 1-96, pIs. 1-28. on the scale of 1: 100,000 and its ex­ FUJIMOTO, H. (1939): On the Torinosu Se- planatory text. pp. 1-240. (in Japanese)

'ill. Aoiwadani 1"1 ;5- :tl- Kumotoriyama = ~ LlJ Kitatsuru-gun -~I:; 'li[1 f1l Shiosawa tj 'll1I iR Kosode IJ' MI Tabayama-mura ft tit LlJ *1 Ushiroyamagawa 1:& LlJ }I[ Kamosawa ~ iR Oonari *- fit Tabagawa ft lit. }II Trans. Proc. Palaeont. Soc. Japan, N.S., No. 89, pp. 24-26, pI. 7, April 20, 1973

613. A TRIASSIC HEPATICA FROM THE OMINE COAL-FIELD, SOUTHWEST HONSHU, JAPAN*

KAZUO HUZIOKA

Institute of .Mining Geology, Mining College, Akita University

and

EITARO T AKAHASI

Department of Mineralogical Sciences and Geology, Faculty of Literature and Science, Yamaguchi University

*MtmIII,It::::.:llHc:aXIHr,1S: ill I::JYr'!;**J!mrl:l*Hl'fm~0)~:(,iJ\M19E J::tJID (~~nt);b -.:=. "' !l) J: fJ ffIi: t..- t-::aH!1r,:P Hepaticites oishli UfrTffi) ~~[,J~£ t..- t-:, W!lliO) Onychiopsis Series (l:.ff~:/;L :7 ~c - Tff~B!'!!!*c) 0) Marchantites yabei KRYSHTOFOVICH (Thallites yabei (KRYSHT.) HARRIS le~I6G;fc-Cv'Q) feltt..-, NliJd*O)~i\liJ:*m<, 7J~tffl1l\f,~MTIv't..-, ± IlJiHi~fJIijiJ' Gj&.fRiJ;*~ t..- -C v' Q, EJ:*J!i"I5O)~Hr,:p""(" Ji, Q, jt{~fiiJ]-93 . ;@jt{.f,~*Jl!I\

Hepaticites oishii dealt with in this KON'NO, Equisetostachys (Neocalamites?) paper occurred from the Upper Triassic pedunculatus KON'NO takahashii KON'NO, Momonoki Formation of the Mine Group Neocalamites carrerei (ZEILLER) HALLE, at the Hagimine colliery in the Omine Todites goeppertianus (MUNSTER), T. coal-field, Yamaguchi Prefecture, south­ recurvatus HARRIS, Clathropteris menis­ west Honshu. The Mine Group, a coal­ co ides BRONGNIART, C. obovata OISHI, bearing deposit, was subdivided by KA­ Dictyophyllum sp., Cladophlebis haiburn­ T AY AMA (1939) into three formations, the ensis (LINDLEY et HUTTON), C. nebbensis lower or the Hirabara Formation, the (BRONGNIART), c. sp., Taeniopteris min­ middle or the Momonoki Formation and ensis OISHI, Podozamites concinnus OISHI the upper or the Aso FOrmation. et HUZIOKA, and P. atsuensis T AKAHASI. The undermentioned fossil plants, to­ The Mesozoic hepatics are rather rare gether with Hepaticites oishii, have been in the world. Thallites yabei (KRYSHTO­ known from the Middle Carnian Momo­ FOVICH) HARRIS (syn. Marchantites yabei noki Formation at the Hagimine colliery KRYSHTOFOVICH, 1930), an only known (TAKAHASI, 1959; KON'NO, 1962): Equi­ species of the Mesozoic Hepaticae in setites minensis KON'NO, E. takahashii Japan, is one of the most characteristic elements in the Onychiopsis Series (Up­ per Jurassic to Lower Cretaceous) of * Received Nov. 30, 1972; read June 26, Japan (OISHI, 1940), and also in the Nak­ 1954. at Sendai. tong flora (Y ABE, 1905; KRYSHTOFOVICH, 24 613. Hepaticites 25

1930) of southern Korea and the Nikan broad, composed of midrib and lamina, flora (KRYSHTOFOVICH, 1930) of the Far repeatedly forked at angles of 30-80 de­ East. grees and intervals of 1.0-1.5 cm. Midrib Several form-genera of the fossil Hepa­ thick, distinct, ridged at under surface, ticae, such as Thallites, Hepaticites and with numerous rhizoids. Lamina trans­ Marchantites, have been established by versally bullate, often spotted, entire to the previous authors. According to W AL­ wavy at margin. Rhizoids borne on TON (1925) and HARRIS (1942), Marchan­ midrib, rigid, regularly and densely ar­ tites is restricted to the hepatics agree­ ranged, frequently forked. ing with the Oligocene Marchantites se­ The midrib and rhizoids are shown zannensis SAPORT A, the type species, in in fig. 1a. The spotted surface (air cham­ which marchantiaceous air chambers, ber?) and the wrinkled surface are shown ventral scales and reproductive organs in fig. lb. The thallus of this fossil have been demonstrated. The form­ hepatica is restored as shown in fig. 3. genus Thallites W ALTON (1925) is to be Comparison and remarks :-Compared used for thalloid fossils showing no char­ with Thallites yabei (KRYSHTOFOVICH) acter proving that they belong to the HARRIS (1942) (syn. Marchantites yabei Hepaticae rather than to the Algae. The KRYSHTOFOVICH, 1930) which is shown form-genus Hepaticites WALTON is used in fig. 4, a well-known Upper jurassic­ for thalloid fossils showing some definite Lower Cretaceous Hepatica in eastern character of the Hepaticae. In practice Asia, this Triassic thallus are much nar­ this is usually the rhizoids. rower, forking at shorter intervals. HAR­ The thallus of Hepaticites oishii are RIS (1931) reported two species of Hepa­ well-impressed on the dark-coloured ticites, H. globosus HARRIS and H. laevis shale, but are not sufficiently preserved HARRIS, from the Thaumatopteris Zone to permit their anatomical studies. This of Scoresby Sound, East Greenland. Of species is the oldest representative of them, H. oishii closely resembles H. laevis, the Hepaticae in japan, being of the but in the former species the midrib is Upper Triassic (Middle Carnian) age. strongly ridged at the under-surface and At this place the writers express their bears numerous rigid rhizoids which cordial thanks to Mr. N. NAMBA who are often furcated. collected the materials from the Hagi­ H. oishii is somewhat similar to but mine colliery of the amine coal-field. distinguishable from the jurassic H. arcuatus (LINDLEY et HUTTON) HARRIS (1942) (syn. Marchantites erectus (LECK.) Description of species SEWARD) from England, and the Upper Class Hepaticae Triassic H. solenotus HARRIS (1938) of England, and also Thallites uralensis Form-genus Hepaticites W ALTON, 1925 KRYSHTOFOVICH et PRINADA (1933) from eastern Urals. Hepaticites oishii HUZIOKA et T AKAHASI Occurrence and collection :-Hagimine new species colliery, amine coal-field, Yamaguchi Pre­ fecture; Momonoki Formation (Middle PI. 7, figs. 1, la, lb, 2, 3 Carnian) of the Mine Group. Holotype, Description:-Thallus regularly and di­ AKMG-7019; paratype, AKMG-7020.Coll. chotomously branched, narrow, 0.3-0.4cm N. NAMBA. 26 Kazuo HUZIOKA and Eitaro T AKAHASI

References rea and the Nikan Series of the Manchu· rian Border. Ann. Soc. Pal. Rus., Tome HARRIS, T.M. (1931): The fossil flora of 8, pp. 144-147, pI. 15. Scoresby Sound, East Greenland. Part 1. -- and V.D. PRINADA (1933): Contribution Medd. om Cn/mland, Bd. 85, no. 2, pp. 1- to the Rhaeto-Liassic flora of the Chelia­ 102, pis. 1-18. binsk Brown coal-basin, eastern Urals. -- (1938): The British Rhaetic flora. Brit. Trans. United Ceol. Prosp. Servo USSR, Mus. (Nat. Hist.) , London, pp. 1-84, pis. Fasc. 346, pp. 1-34, pis. 1-5. 1-5. OISHI, S. (1940): Mesozoic floras of Japan. -- (1942): On two species of Hepatics of jour. Fac. Sci., Hokkaido Imp. Univ., ser. the Yorkshire Jurassic flora. Ann. Mag. 4, vol. 5, nos. 2-4, pp. 123-480, pis. 1-48. Nat. Hist., ser. 11, vol. 9, pp. 393-401. T AKAHASI, E. (1959): Floral changes since KATAYAMA, M. (1939): Stratigraphical stu­ the Mesozoic Age of western Honshu, dy on the Mine Series. jour. Ceol. Soc. Japan. Sci. Rept. Yamaguchi Univ., vol. japan, vol. 46, no. 546, pp. 127-141. (in 10, pp. 181-237. (in Japanese) Japanese) WALTON, J. (1925) : Carboniferous Bryophyta. KON'NO, E. (1962): Some species of Neocala­ 1. Hepaticae. Ann. Bot., vol. 39, pp. 563- mites and Equisetites in Japan and Korea. 572. pI. 18. Sci. Rept. Tohoku Univ., 2nd ser., Special Y ABE, H. (1905): Mesozoic plants from Ko­ Volume, no. 5, pp. 21-47, pis. 9-18. rea. jour. Coli. Sci., Imp. Univ. Tokyo, KRYSHTOFOVICH, A.N. (1930): A Liverwort vol. 20, art. 8, pp. 1-59, pis. 1-4. from the Middle Daido Formation of Ko-

Aso Mine Hagimine Momonoki Hirabara Omine

Explanation of Plate 7

Figs. 1, la, 1b, 2, 3. Hepaticites oishii new species, from the Hagimine colliery. 1. Holotype, AKMG-7019. 1a and lb. Enlarged to show the detailed characters of thallus, x 3 and x 2, respectively. 2. Para type, AKMG-7020. 3. A restoration of Hepaticites oishii. Fig. 4. Thallites yabei (KRYSHTOFOVICH) HARRIS (syn. Marchantites yabei KRYSHTOFOVICH), from the Naktong Series of southern Korea. Reproduced for comparison; YABE, 1905, p. 41, pI. 3, fig. 16a; originally, Sagenopteris bilobata YABE var. major YABE. HUZIOKA and T AKAHASI: Hepaticites Plate 7

·/~'.\(-f " ! ( \ ·U J i ·. . ·1 '1 ! \ ' .. "...... '\ .f ! l' t f ,'"1 4 \ \ I I \ \ /,J t / / 1\ i , .--.V?i (-'1~ L,./ i/o _c

SHIBATA photo. Trans. Proc. Palaeont. Soc. Japan, N.S., No. 89, pp. 27-41, pI. 8, April 20, 1973

614. V ASCOCERATID AMMONITES FROM THE TURONIAN OF HOKKAIDO

(STUDIES OF THE CRETACEOUS AMMONITES FROM HOKKAIDO AND SAG HALlEN-XXVI)

T ATSURO MATSUMOTO

Department of Geology, Kyushu University, Fukuoka 812

~tffljJ[ 7-;L - j:l .::. 7 :/il!;/ A::! t: '7 Af4~::fi: Vascoceratidae i'i'f 7- AfflO)&i1iDl~lc

~,!1i:-t ~d)~, *ifi$:ti!!.~"t'·i'i ~ :h6b-C ~ nL';!i,.Q 0 i:. I:.lc~tffljJ[O)-fl}11 7-;L - j:l .::. 7 :/iJ' f? ~nlcf.!.l±\~nt.: 4 tf,*a-, Vascoceras sp. aff. V. durandi (THOMAS and PERON), Fagesia thevestensis (PERON), Fagesia sp. d. rudra (STOLICZKA) 0) 3 fllHcnrJ5E L.., ~~L..-C~ti-t.Qo ~f?~7-;L-j:l.::.7:/M~~~.QnrJM.::fi0)~~~~m~~~a-A~, ilifIiillia-ilit.7;..Qo Vascoceratids O)!j;f1ftJ:~:(!1vi, .lj3.IC~!t!mIl~"JtJ:~IJ;fd~I:rL'tJ: <, .;c-O)lg~t: '7!11 r iM*~.g.*Jl!, ~.Q i:.1t'IDl~, !tHfljO)~5jfr.;j: cO)~fr;'ICMrc.: L.. t.: ~filU:O)!j;f~M:a- 'h~ 1.\\!:-t ~~ L', :1:.,iffiJ~rIf~~ilSfiRJ:\fct1f~l1:iJ;;!i, -;,f.:o)L'vi;!i,.Q ~ It'iJ'o 13 *IC 'hJ}l";: l..t.: 3 f Lt. oj IOJ!,:hn.Qo ~iiIE O)liffi~M: 'h ~;t oj .Q iJ;, ~:(!1a-~~rr.Q Ie ~ t.: fJ, 7 :/7T~jl'tII4l-!t!HC -:J1t, -C ~QN~iJ;::fjJ/lltJ: i:. C iJ;~MiJ,n.Qo ~ * i~ ~I;

Introduction had to omit for brevity in my last paper. Before entering in the description I The Vascoceratidae are typically Te­ thank Dr. Hakuyu OKADA and Mr. Ka­ thyan in occurrence. This has been zunari TANABE who found an interest­ evidenced by numerous works of pre­ ing specimen in a field work with me decessors. In my concise note on the and provided it for this study, Mr. Ha­ palaeobiogeography of the late Cretace­ jime KOYAMA who showed me another ous Ammonoidea (MATSUMOTO, 1973), I interesting specimen, and Professor Te­ have mentioned that some immigrants tsuro HANAI who gave me facility to of the vascoceratids are found outside study some specimens preserved in the the Tethys region. University of Tokyo. Dr. Itaru HAY AMI In this paper I describe three species and Miss Mutsuko HAYASHIDA kindly of the vascoceratid ammonites which assisted me in preparing the plates and occurred rarely in the Turonian of Hok­ typescript. kaido. After the description supplemen­ tary remarks are given on the biogeo­ Description of Species graphical distribution of the vascoceratid ammonites, presenting a map which I Family Vascoceratidae SPATH, 1925

* Received Dec. 31, 1972; read Jan. 16, 1973 The classification of the Vascocerati· at Sendai. dae is more or less different between 27 2R Tatsuro MATSUMOTO authors (e. g. FUR ON, 1935, SCHNEEGANS, allied to one another and have more or 1943; REYMENT, 1954, 55; BARBER, 1957 ; less similar sutures. Typically the su­ WRIGHT, 1957; COLLIGNON, 1958; WIED­ ture of Fagesia is the most complicated MANN, 1964; FREUND and RAAB, 1969). of the family. It is more deeply incised The material from Japan is too poor to and consists of higher saddles and nar­ give comments on the problem, but I rower or deeper lobes than that of Vas­ had fortunately an experience to look coceras. through a number of specimens from In Paravascoceras FURON, 1935, there Nigeria and talked with REYMENT and are external ribs but no umbilical tuber­ BARBER, though separately, when we cles. Pachyvascoceras FURON, 1935, is were studying Cretaceous ammonites at regarded by SCHNEEGANS (1943), BARBER the British Museum (Natural History), (1957), and FREUND and RAAB (1969) as 1953-54. From my general impression, a synonym of Paravascoceras, because I was rather inclined to agree with there is a continuous series between SCHNEEGANS' attitude in treating of the their type-species, V. crassum FUR ON, material. Anyhow, it is worthwhile to 1935, and V. cauvini CHUDEAU, 1909. I note that great many varieties of vas­ would agree with FREUND and RAAB coceratid ammonites were differentiated (1969, p. 19) in regarding Broggiliceras within a short time interval in the early BENAVIDES-CAcERES, 1956, as another Turonian of the Tethys region. It would synonym of Paravascoceras. Some spe­ be interesting to study the mode of cies which were referred to Pachyvas­ evolution through examining ontogeny coceras may not automatically transfer­ and variation from population to popu­ red to Paravascoceras. V. durandi (THO­ lation. MAS and PERON) can be reckoned as To describe a small number of incom­ such an example. pletely preserved specimens from Hok­ Discovascoceras COLLIGNON, 1958 was kaido, I have to depend, for convenience commented by WIEDMANN (1959, p. 757) sake, on some previously proposed as indistinct. Its type-species, D. tes­ framework of classification. selitense COLLIGNON 1958, was regarded by WIEDMANN as a synonym of Vasco­ Genus Vascoceras CHOFF AT, 1898 ceras (Pachyvascoceras) triangulare FA­ RAUD, 1940 and then by FREUND and Type-species :-Vascoceras gamai CHOF­ RAAB (1969, p. 23) as that of Paravas­ FAT, 1898. coceras obessum (TAUBENHAUS, 1920). Remarks:-This genus has a globose COLLIGNON (1965, p. 179 [17J), however, to sub globose, involute shell, in which redefined Discovascoceras, describing the umbilical tubercles and fold like costae change of characters with growth and are distinct only in early stages and the presence of three keels. On the lost with growth. In Fagesia PERVIN­ grounds of its stratigraphic occurrence QUIERE, 1907, the shell is inflated cadi­ at the very base of the Turonian and cone with ornament similar to but more also of its morphologic characters, COL­ persistent than that of Vascoceras. In LIGNON considers to allocate Discovas­ Plesiovascoceras SPATH, 1926, the whorl coceras at the initial part where the is evolute, coronate and ornamented with Vascoceratidae and the Tissotiidae are strong obtuse tubercles at the umbilical going to branch apart (see REYMENT, margin. The three genera are closely 1954, text-fig. 8). 614. Vascoceratid ammonites 29

Vascoceras sp. aff. V. durandi 1965. Vascoceras (Pachyvascoceras) aff. dou­ (THOMAS and PERON) villei·durandi, COLLIGNON, Atlas Fossiles Caracteristiques de Madagascar (Ammo. PI. 8, Fig. 1, Text.fig. 1 nites), fasc. 12, p. 43, pI. 394, fig. 1676. 1969. Vascoceras durandi, FREu'>D and R,AAB, Compare.- Special Papers in Palaeont., no. 4, p. 29, 1889. Pachydiscus durandi TlIm,'IAs et PlmOi':, text·fig. 6h-i. Moll. Foss. Tunisie, p. 27, pI. 18, figs. 5-8. Material :-GK. H5674, from loc. R2513a, 1896. Pachydiscus durandi, PERO,\, Minn. Soc. the Obira, collected and supplied by H. Geo!. France, Pa!eont., Mem. No. 17, p. C)KADA and K. TANABE, 1971. 44, pI. 4, fig. 1 ; pI. 5, fig. 1; pI. 17, fig. 5. Description:-The specimen is fairly 1898. Vascoceras douvillei CIIOFF AT, Faune Cre!. Portugal, p. 59, pI. 10, figs. 3, 6; well preserved on the right side but pI. 11, figs. 2-5; pI. 21, figs. 13-16. badly crushed on the left. It is wholly 1907. Vascoceras durandi, PERVli'\QUIERE, septate and the body-whorl has mostly Etudes Pa!eont. Tunisienne, vol. 1, p. gone. The measurements are as fol­ 332, pI. 21, fig. 1a-b; text-fig. 125. lows:

Diameter Umbilicus I-Ieight Breadth B.;H. 115mm.(1) 26(.23) 54(.47) 33x2(.57) 1. 22

The coiling is considerably involute The mode of ribbing, if any, on the and seems to be somewhat eccentric. external part of the inner whorl is not The whorl is inflated on the flanks, known, as it is difficult to take out the broader than high and broadest near deeply overlapping outer whorl. Only the umbilical shoulder in the lower part. a faint umbilical buldge is perceptible. The flanks are convergent passing into Sutures are pseudoceratitic, with point­ a rather flattened venter which occupies ed lobules and phylloid folioles. The the breadth of the external lobe. The external lobe (E) is subrectangular in shell is as a whole subglobose in general rough outline, although its details are aspect. obscured for unfavourable preservation. The umbilicus is deep and surrounded The first lateral saddle between E and by a high and vertical wall which forms L is asymmetric, steep on the side of an angular edge with the flank. E and gently sloping towards L. The So far as the observable part is con­ first lateral lobe (L) is opened V -shaped cerned, the shell is free from distinct in rough outline, moderately deeply in­ ornament. But a very faint buldge is cised by several lobules and bipartite discernible outside the umbilical edge at the bottom. The second lateral sad­ of the outer whorl. This is probably a dle is roughly dome-shaped and shallow­ reduced umbilical tubercle, although no ly incised on top, leaving phylloid foli­ distinct umbilical nodes are observed on oles. The second lateral lobe (U2) is the visible part of the inner whorl. much narrower than L and moderately From the low buldge mentioned above incised, having a narrow and deeply runs a very indistinct, low rib, on which pointed lobule at its bottom. The sad­ several fine rib lets and lirae are super­ dle inside U2 is nearly as high as the imposed. They are somewhat prorsira­ second lateral saddle, sub quadrate in diate and gently concave on the flank. rough outl ine anr. apparently tripartite 30 Tats~i?"O MATSUMOTO

/ ,/ / I / / I { I \ // \\ ( \ \ '-_ _ _ _ I \ / I \ I \ { \ { \ \ \ \ \ \

E

u.s . f < \" c 10 mm,

Fig. 1. Vascoceras Sp. aff. ]I. durandi ( THOM AS and P ERO N ) . GK. H5674, frorn loco R2513a . Photographic latera l vie w (a), with whitening ; diagrammatic frontal view (b); external suture (c) at s .

into folioles of unequal size, with the ComjJarison :- Although the specimen narrowest one on the side of U2, the from l-lokkaido is incompletely preserved, moderate one in the middle and the the olBerved characters suggest that it broadest one on the umbilical shoulder. is very close to the extreme end of the Narrow U3 and a part of U5 are visible variation of Vascoceras durandi (THOMAS on the umbilical wall. and P ERON) (see P ERV1NQU1ERE, 1907), 614. Vascoceratid ammonites 31 which has been known from the Lower To sum up, it is difficult to identify Turonian of Tunisia, Algeria, Portugal precisely the present form, unless the and Israel. Its umbilicus is considerably characters of the early shell is more narrower than that of the measured spe· distinctly known. It is, however, highly cimens from these areas. In the dimen· possible that the described specimen is sions of the umbilicus, whorl·height and close to the extreme end of the varia­ breadth in proportion to the diameter, tion of Vascoceras durandi (THOMAS and the present specimen is closest to the PERON), although it could represent a specimen from the Turonian zone of geographically separated subspecies or Mammites conciliatum of Madagascar de· another closely allied species. scribed by COLLIGNON (1965b) under the Occurrence :-The described specimen name of Vascoceras (Pachyvascoceras) aff. was obtained at loco R2513a, from a cal­ douvillei·durandi. It differs from that careous nodule in the mudstone of the Madagascar specimen in that it has less Middle Yezo Group exposed along the distinct ornament and not so sub quad· Okufutamata, a tributary of the Obira rate whorl·section as the Madagascar [Opirashibets], by H. OKADA and K. one. It has rather a rounded subtrape· TANABE, during the field work conduct­ zoidal whorl-section. In this respect and ed by me in 1971. The mudstone of loco in the general aspects in lateral view, R2513a is allocated at the middle of unit the present specimen is apparently simi­ Mj of TANAKA (1963). No mega-fossils lar to Choffaticeras meslei (PERON, 1897), were associated with this ammonite at from the Turonian of Algeria and Israel. this locality. In the Nakakinebetsu val­ It is distinguished from that species by ley, another tributary of the Obira, about the absence of keels, more inflated and 13 km southwest from loco R2513a, a broader whorls and more complex su­ probable extension of unit Mj and sub­ tures which show dissimilar pattern jacent part down to the middle of unit (compare Text-fig. 1 with PERON'S pI. Mh are more prolific, where I obtained, 17, fig. 1). The suture of the Hokkaido among others, Inoceramus (Mytiloides) specimen is close to, if not exactly iden­ labiatus, Mammites sp., and Pseudaspido­ tical with, that of P. durandi as illus­ ceras sp. On the grounds of available trated by PERVINQUIERE (1907, text-fig. evidence I refer the sequence from the 125). upper part of Mh to the top of Mj to On account of the characteristic shell­ the Lower Turonian. form and the indistinct, almost obsolute Location of R2513a is about 850 m east ornament, I thought that the present and 3900 m south from the point, N 44 ° form could be referred to Discovasco­ 10'-E 142°0', northwest corner of the ceras COLLIGNON, 1958, which should be Horokanai sheet, geological map of provided with keels as redefined by COL­ 1: 50,000, published by the Geological LIGNON (1965b). Our species has no keel Survey of Japan (1958). It is in the and the lobes of its suture are more Rumoi district, Teshio province, north­ deeply incised and less numerous on the west Hokkaido. flank than those of D. tesselitense COL­ LIGNON and other related species. Thus Genus Fagesia PERVINQUIERE, 1907 it is not referable to Discovascoceras, if Discovascoceras is excluded from Para­ Type-species :-Olcostephan us superstes vascoceras as independent genus. KOSSMAT, 1897. 32 Tatsuro MATSUMOTO

Remarks:-This genus is well defined, Fagesia thevestensis (PERON) as mentioned by FREUND and RABB (1969, PI. 8, Fig. 2, Text-fig. 2 p. 33). They commented, however, that numerous species of very wide distri­ 1896. Mammites (?) thevestensis PERON, Mem. bution included later in this genus dif­ Soc. Ceol. France, Pateont., Mem. No. 17, fer considerably from a well-defined p. 23, pI. 7, figs. 2, 3. group formed by F. superstes, F. the­ 1904. Ammonites kotoi Y ABE, Jour. Coll. Sci., vestensis, F. rudra, F. peroni and F. Imp. Univ. Tokyo, vol. 20, art. 2, p. 26, bomba, from India, Middle East, Spain pI. 6, figs. 3 and 4. and North Africa, so that in several 1907. Fagesia thevestensis, PERVINQUIERE, Etudes Paieont. Tunisienne, vol. 1, p. cases the generic position is uncertain. 325, pI. 20, figs. 5, 6; text-figs. 123, 124. In fact, Fagesia cali/ornica ANDERSON, 1969. Fagesia thevestensis, FREUND and RAAB, 1935 and F. shastensis ANDERSON, 1935, SPecial Papers in Paiaeont., no. 4, p.35, from California, should be better trans­ text· fig. 7g. ferred to Plesiovascoceras, as I have already pointed out (MATSUMOTO, 1959, Material :-GT. 1-3249, from loco T610b­ p. 102). c, Saku area (colI. T. MATSUMOTO). GT. The species from Japan described MM 7561, from a rolled nodule of the below undoubtedly belong to the typi­ Yubari, the holotype of Ammonites kotoi, cal group of Fagesia. is also examined.

Measurements :- Specimen Diameter Umbilicus Height Breadth B./H. GT. I-3249 80(1) 20(.25) 36(.45) 45(.56) 1. 25 GT. MM7561 100(1) 27(.27) 44(.44) 53(.53) 1.2 PERON'S type 80(1) 25(.31) 34(.42) 45(.56) 1.3

Descriptive remarks :-GT. 1-3249 re­ Some difference in the size of umbilicus presents a fairly large individual, but is probably within the variation of a its outer whorl, is incomplete and shows species, as FREUND and RAAB (1969, p. eroded sutures. Therefore its inner 35) generally admitted. whorl alone is measured and illustrated. The whorl is inflated and lunate in GT. 1-3249 and GT. MM7561 are com­ cross-section. It is broader than high, plementary with each other, showing but is not so broad as in F. superstes altogether the following characteristic (KOSSMA T, 1897). The measured pro­ features. portion between height and breadth is The outer whorl, of GT. 1-3249, though of the order of that measured on the incomplete, is as large as or somewhat PERON'S type, PERVINQUIERE's specimens larger than the large specimen of PER­ I and III. The large outer whorl of GT. VINQUIERE (1907, pI. 20, fig. 6). They are 1-3249 seems to have broader whorl but both still septate and the shell must cannot be accurately measured. The have reached about 200 mm., if the adult venter of the inner whorl is moderate­ body-whorls were preserved. PERON'S ly rounded as in the holotype of PERON holotype must have been an immature and the smaller specimen of PERVIN­ shell, with which the inner whorl of GT. QUIERE (1907, pI. 20, fig. 5). The umbili· 1-3249 is well comparable in dimensions. cal wall is very steep or nearly vertical. 614. Vascoceratid ammonites 33

c

F ig. 2. Fagesia thevestensis (P ERON) . GT. MM7561 , holotype of Ammonites kotoi YABE, ,1904, from Yubari. Kyushu Univ. [1. H.] photos, with whitening.

The periumbilical tubercles are some­ To sum up the two specimens from what bullate or subrounded. Although Hokkaido are identified with Fagesia they are not completely preserved in th evestensis ( P ERON), from the Lower the Hokkaido specimens, they seem to Turonian of Algeria, Tunisia and Mid­ be similar in intensity and density to dle East. It is still highly possible that those in the specimens described by Fagesia thevestensis (P ERO N, 1896) and PERVINQUIERE. The ribs are also very Fagesia superstes (KOSSMAT, 1897) could similar in intensity, density, forward be within the variation of one and the curvature and mode of bifurcation to same species, as PERON (1897, p. 84) at those of the holotype and the inner first pointed out. Without treating a whorl of P ERVINQUIERE. In the two spe­ sufficient number of specimens from cimens from Hokkaido some of the ribs place to place (Algeria, Tunisia, Israel are slightly more elevated than others and India), I cannot decide the matter. and some of the interspaces are some­ For the time being the two species are what deeper than others, if not forming distinguished as PERVI 'QUIERE did. As distinct constrictions. On the outer there is no reason to regard the Japa­ whorl the ornament is much weakened. nese form as an independent species, The suture is of typical Fagesia pat­ Ammonites Iw to i Y ABE, 1904, falls in the tern, showing three, high, deeply incised synonym of Fagesia th evestensis ( PERON, saddles on the side and deeply incised 1896). lobes. The first lateral saddle is tri­ Occurrence :-GT. 1-3249 was obtained partite. L is nearly as deep as E and by myself from lac. T610- c, Onodera­ bifid at the bottom. The terminals of zawa, a small branch of the Sakugawa the subdivided folioles are somewhat [Sakotan-gawa], unit IIc of the Middle phylloid. Yezo Group in the Saku area, Teshio 34 TatS'lLrO MATSUMOTO province, northwest Hokkaido (for the Text-fig. 3 location see a map of pI. 12 in MATSU­ Compare.- MOTO, 1942). From another locality of 1965. Ammonites rudra STOLICZKA, Palaeon!. unit IIc about 3 km west of Saku, I have Indica, ser. 1, vol. 1, p. 122, pI. 60. recently obtained Inoceramus (Mytiloides) 1897. Olcostephanus rudra, KOSSMAT, Beitr. labiatus, which indicates Lower Turo­ Paliiont. Oesterr. Ungarns, vol. 11. p. 29 [136J. nian. 1907. Fagesia rudra, PERVINQUIERE, Etudes GT. MM7561 was obtained by Y ABE in Pateont Tunisienne, vol. 1, p. 322. a rolled nodule of the Yubari, Ishikari 1965. Fagesia rudra, COLLIGNON, Atlas Fos­ province, central Hokkaido. Although I siles Caracteristiques de Madagascar did a field work in the same area, I was (Ammonites), fasc. 12, p. 48, pI. 369, not successful to get the same species fig. 1678-A; pI. 397, fig. 1678-B. from this area. Material :-A specimen in the private collection of Hajime KOYAMA, Saku, Na­ Fagesia sp. cf. F. rudra (STOLICZKA) kagawa-machi, Hokkaido.

Measurements :- Specimen Diameter Umbilicus Height Breadth Bj.H. KOYAMA'S 120.0(1) 29.5(.24) 50.5(.42) 100(.83) 2 STOLICZKA'S 170 (1) - (.23) - (.39) 2. 12 COLLIGNON'S 180 (1) 36 (.20) 95 (.53) 136(.76) 1. 43

Descriptive remarks :-The last suture fine ribs are discernible, showing a is seen at 120 mm. in diameter and the gently forward curve on the venter. main part of the body-whorl is badly Fine lirae may be seen on the inter­ crushed. Even the septate whorl is spaces of the ribs. somewhat deformed on the left side. The ornament of the inner whorl is The above measurement of the whorl­ not well seen. breadth is approximately estimated from The suture is of typical Fagesia pat­ the little deformed right half. (see Fig. tern, showing asymmetrically tripartite, 3b). high, lateral saddles and deeply incised The proportion of the dimensions of lobes. Koy AMA's specimen is closer to STOLIC­ The observed characters indicate that ZKA'S type rather than to COLLIGNON's the described specimen is probably re­ specimen. The shell is considerably in­ ferable to Fagesia rudra (STOLICZKA), volute and subglobose in aspect. The which has been reported from the Lower whorl is about twice as broad as high, Turonian of southern India (STOLICZKA, broadest between the umbilical shoul­ 1865; KOSSMAT, 1897), Madagascar (COL­ ders, and semi-lunate in section. The LIGNON, 1965) and Spain (WIEDMANN, umbilicus is narrow and deep, with a 1964, listed only). steep and high wall and subangular Occurrence:-H. KOYAMA obtained this shoulder. The venter is broadly convex. specimen at a locality close to my loco The last septate whorl is nearly T1158 or T1159, in the mudstone of unit smoothish, but very faint, blunt umbili­ IIc, Saku-gakko-no-sawa, Abeshinai-Saku cal bulges are perceptible and also weak area, Teshio province, northwest Hok- 614. Vascoceratid ammonites 35

E

" .. ! / ,/,/:/,/,' " I' ," I •

: ':' : : ,f .' " I . ,, ,' : ~: ;' " , "

c

Q

b 20mm

Fig. 3. Fagesia sp. d. F. rudra (STOLlCZKA). Diagrammatic sketch of a specimen of H. KOYA~IA'S collection from the Saku area. Lateral view (a), whorl-section (b) and ex­ ternal suture (c) at S. (T.M. delin.) kaido. Calcareous nodules in unit IIc cated. A comprehensive discussion on often shows a particular, light greenish the palaeobiogeography has already been tint. The rock matrix of this fossil given in my last paper (MATSUMOTO, shows the same colour. Unit lIe in this 1973). Here is given a supplementary area contains Inoceramus (Mytiloides) note. labiatus and is assigned to Lower Turo­ While several Turonian genera of 'the nian. Acanthoceratidae and the Collignon i­ ceratidae, such as J(anabiceras, Watino­ ceras, Mammites, Pseudaspidoceras, Ro­ Note on the Biogeographical Distribution maniceras, Collignoniceras and Subprio­ of the Vascoceratid Ammonites nocyclus show world-wide distribution, ammonites belonging to the Vasco<::era­ Fig. 4 is a Turonian palaeogeographic tidae and also the Tissotiidae charac­ map (without restoring the relative po­ teristically occur in the Tethys Sea re­ sition of continents) in which the occur­ gion in a broad sense. rence of selected ammonites are indi- Although the generic assignment of 36 Tatsuro MATSUMOTO

• PuzQsia, Mesopuzosio • Collignoniceros. Selwynoceras .. Vascoceras, Plesiova5Coceros o Tetragoniles o Mammites, Pseudaspidocera~ '" Paravascoceras x Hyphontoceras • Watinoceros. Sumitomoceras H Nigericeras, GomheocefOS , Scophites, Otoscaphites t:. Fcgesia o Metoicaceros v Neoptychites ? Data wanted

Fig. 4. Turonian palaeobiogeographic map of selected ammonoids. the higherto described species has been southern France, Mexico and Texas. revised in the light of up-to-date classi­ The two authors state, furthermore, fication, some personal difference in that Nigericeras and Gombeoceras are identification may still remain. I am often associated with the Paravaseoeeras afraid that this may affect the conclu­ fauna and that Metoicoceras is so with sion about the distribution of the vas­ the Vascoceras fauna. It is noted, how­ co cera tid genera. ever, that Gombeoceras occurs in Tur­ According to FREUND and RAAB (1969), kestan which is certainly situated on Paravaseoeeras (in their sense) and Para­ the north side of the Tethys, that Vas­ mammites prevail in the southern shelf coceras is found in the Middle East and seas of the Tethys, such as those in the Nigeria where Paravaseoeeras prevails Middle East, Lybia, Tunisia, Algeria, and that Paravascoceras is found in Niger, Nigeria, eastern Brazil and Peru, Mexico and adjacent parts where Vas­ while Vascoeeras and Plesiovaseoeeras coeeras and Plesiovascoeeras abound. A are common in the northern shallow species of Nigericeras represented by a seas, such as those in Portugal, Spain, considerable number of specimens has 614. Vascoceratid ammonites 37 , ...... deep troughs and also rIsmg islands. Currents may have passed across some part of this incomplete barrier. I do not think that Metoicoceras of the Acanthoceratidae (Metoicoceratinae) is so intimately associated with the Vas­ coceras fauna. It is most commonly distributed in the Gulf Coast and the Interior provinces of North America, ranging from the upper Upper Ceno­ manian to the lower Lower Turonian. It is distributed fairly commonly in western Europe and Morocco, suggest­ ing an intimate faunal relation on both sides of the north Atlantic ocean, which may have been narrow in the late Cre­ taceous times. The genus is rarely found in the Middle East, southern India, Nigeria and Madagascar (?) (the latter two areas in Upper Cenomanian only), but no immigrant nor floated remnant of Metoicoceras has been found f in the Pacific region (see SASTRY and MA TSUMOTO, 1967). o c;I'?;,' It would be desirable to examine the faunal comparison by means of a quan­ titative measure, as has been demon­ strated by REYMENT and T AIT (1972, p. 322). It would be likewise necessary to observe carefully the mode of occur­ recently been reported from southeast­ rence of the fossils. For instance, it ern Colorado (COBBAN, 1971) and another has not yet been settled whether the species closely allied to N. glabrum small assemblage of the vascoceratids BARBER from somewhere (locality un­ found at a locality in southern Montana certain) in North America (MATSUMOTO (reported by REESIDE, 1923) represents and OTSUKA, 1969). Furthermore, . there an ammonite faunule migrated from the are several genera of wider distribution, south or merely implies an accumulation such as Fagesia, Neoptychites and Tho­ of dead cephalopod shells floated from masites, which are not restricted to the main distribution area of the vas­ either of the northern and southern coceratids in the south. A map of sides of the Tethys. The barrier be­ palaeocurrent in the early Turonian tween the northern and the southetn seaway in North America has already provinces, if ever existent, must not been inferred by CLOUD (1961, fig. 10). have been very rigid. It was probably This was hinted by the known distri­ represented by a geosynclinal part of bution of the straight shelled ammonoid the Tethys, which may have comprised Sciponoceras gracile (SHUMARD), but can TatsuroMATSUMOTO also serve for explaining the dispersal Turonian shelf seas of the Tethys. or migration of Plesiovascoceras up to As the genus Fagesia is generally Montana and also to California. widespread, it would not be surprising Vascoceratid ammonites indeed seem to find several fossils of this genus from to l;>e generally prolific in the shelf seas Japan. Their occurrence can be inter­ of the Tethys, but they actually occur preted as either a rare immigrant or a in particular beds which occupy a re­ float brought by current from the south. stricted part of Lower Turonian se­ The known distribution of Fagesia. the­ quence, as recorded precisely by FREUND vestensis is, however, much apart from and RAAB (1969). This is also true for Japan, unless F. superstes is proved t9 the mode of occurrence of the vasco­ be conspecific with F. thevestensis. A ceratids in southern Montana. The fact find of Vascoceras sp. aff. V. duran# is. suggests that the ammonites belonging rather puzzling, since V. durandi occurs to the Vascoceratidae which are char­ in Algeria, Tunisia, Portugal and Israel, act~rized by more or less pseudocerati­ which are situated around the pr:esent tic sutures and reducing ornament may Mediterranean Sea. The Japanese form have had a particular habitat and mod~ is, however, allied to a form from Mada­ of life. In this connexion it should be gascar called V. aff. choffati-durandi. noted that Neolobites of the Engono­ There are several species of other fami­ ceratidae, which are phylogenetically lies which are common or closely allied unrelated to the Acanthocerataceae but 'between Japan and Madagascar but not have likewise pseudoceratitic sutures, yet found elsewhere. This is probably occurs characteristically in the late caused by too poorly explored situation Cenomanian shelf seas of the Tethys, of the Cretaceous areas in southern i. e. generally the same region as the Asia. main distributional area of the Vasco­ I have reported recently (MATSUMOTO, ceratidae and the Tissotiidae. It should 1971) a find of Hourcquia from south be furthermore noted that Fagesia which Saghalien, a specialized member of the has more deeply incised sutures than Vascoceratidae in late Turonian-Conia­ other genera of the Vascoceratidae cian times. Professor W. HASHIMOTO shows wider distribution. has shown me another specimen of Now in Japan the ammonites belong­ Hourcquia obtained in the Abeshinai­ ing to the Vascoceratidae are very few Saku area, Teshio province, Hokkaido. and no examples of the Tissotiidae and According to HASHIMOTO (1973), it prob­ the Coilopoceratidae have been found. abl)' came from the Coniacian. Although Only one specimen of Vascoceras sp. aff. the age is younger than Lower Turonian, V" durandi, tw:o of Fagesia thevestensis this is another example of the genus and another of Fagesia sp. cf. F. rudra known only from Madagascar and the are known. from the Lower Turonian of Japanese biogeographic province. Hokkaido. This number of specimens To sum up, it is worthwhile to report is incomparably smaller than that of that several vascoceratid ammonites do hundreds of vascoceratid ammonites occur, though rarely, in the Japanese from Nigeria. On the other h~md des­ province. To explain this fact we need moceratids and tetragonitids, which oc­ more information about the Cretaceous wr abundantly in Japan, are distributed ammonites in southeast Asia and adja­ very sparsely or almost absent in the cent areas which are expected to have 614. Vascoceratid ammonites 39 been the nearest source of the north­ Sci. Nat. Vacucluse, vol. 3, pp. 3-24, 11 ward dispersal or migration of the vas­ pis. coceratid ammonites up to Hokkaido FREUND, Raphael and RAAB, Menahem (1969) : and south Saghalien. Lower Turonian ammonites from Israel. Special Papers in Palaeont., no. 4, pp. 1- 83, pis. 1-10. References Cited FURON, R. (1935): Le Cretace et Tertiaire du Sahara Soudanais. Arch. Mus. Hist. ANDERSON, F.M. (1935): The genus Fagesia Nat., Paris, ser. 6, vol. 13, pp. 1-97, pis. in the Upper Cretaceous of the Pacific 1-7. coast. Jour. Paleont-., vol. 5, no. 2, pp. HASHIMOTO, Wataru (1973): Hourcquia ha­ 121-126, pis. 15-17. taii HASHIMOTO, a new species of ammo­ BARBER, W. (1957) :', Lower Turonian ammo­ nite from the Upper Cretaceous System nites from north-eastern Nigeria. Ceol. of the Abeshinai region, Teshio province, Surv. Nigeria, Bull. no. 26, pp. 1-86, pis. Hokkaido. Professor Kotora Hatai Memo­ 1-34. rial Volume, pp. 315-318, pI. 35, Sendai. BENAVIDES-CAcERES, V.E. (1956): Cretace­ KOSSMAT, Franz (1897): Untersuchungen ous system in northern Peru. Bull. Amer. tiber die stidindische Kreideforma tion Museum Nat. Hist., vol. 108, no. 4, p. 353- (Zweiter Theil). Beitr. Paliiont. Oesterr.­ 494, pis. 31-66. Ungarns u. d. Orients, vol. 11, pp. 1-46 CHOFF AT, Paul (1898): Recueil d'etudes pa­ [108-153J, pis. 1-8 [12-19J. leontologiques sur la faune cretacique du MATSUMOTO, Tatsuro (1942): Fundamentals Portugal. Vol. 1, Especes nouvelle ou peu in the Cretaceous stratigraphy of Japan. connues. Commisso Trav. Ceol. Portugal, Part. I. Mem. Fac. Sci., Kyushu Univ., ser. 2, pp. 41-86, pis. 3-32. ser. D, vol. 1, no. 3, pp. 129-280, pis, 5-20. CHUDEAU, R. (1909): Ammonites du Damer­ -- (1959): Upper Cretaceous ammonites of gou (Sahara meridional). Bull. Soc. Ceol. California. Part II. Ibid., Special vol. 1, France" ser. 4, vol. 9, pp. 67-71, 5 pis. pp. 1-172, pis. 1-41. CLOUD, P.E. Jr. (1961): Paleobiogeography -- (1971): Uncommon keeled ammonites of the marine realm. In Oceanography, from the Upper Cretaceous of Hokkaido Amer. Ass. Adv. Sci., pp. 151-200. and Saghalien. Ibid., vol. 20, no. 2, pp. COBBAN, W.A. (1971): New and little-known 305-317, pis. 48-49. ammonites from the Upper Cretaceous -- (1973): Palaeobiogeography of Late (Cenomanian and Turonian) of the west­ Cretaceous Ammonoidea., In HALLAM, ern interior of the United States. U.S. A. [EditorJ : Atlas of Palaeobiogeography, Ceol. Surv. Prof. Paper 699, pp. 1-24, pis. Chapt. 38, Elsevier, Amsterdam. 1-18. MATSUMOTO, Tatsuro and OTSUKA, Hiruyuki COLLIGNON, Maurice (1958): Cephalopodes (1969): Ammonite collection donated by neocretaces du Tinrhert (Fezzan). Ann. Yukichi OGAWA to the Museum, Bureau Pateont., vol. 43 (1957), pp. 115-136 [3-24J, of Cultural Center, Kagoshima. Sci. Repts. pis. 16-18 [1-3]. Dept. Ceol., Kyushu Univ., vol. 10, no. 2, -- (1965a): Nouvelle ammonites neocreta­ pp. 51-65, pis. 7-15 [in Japanese with cees Saharienne. Ibid., vol. 51 (1965), English abstract]. pp. 165-202 [3-40J, pis. A-H. PERON, Alphonse (1889-90): Description des - (1965b): Atlas des Fossiles Caracteristi­ mollusques fossiles des terrains cretace ques de Madagascar (Ammonites), fasc. de la region des Hauts Plateaux de la 12, pp. 1-82, pIs. 376-413; Servo Geol., Tunisie. Recueillisen 1885 et 1886 par M. Tananarive. Phillipe THOMAS. Exploration scientifique F ARAUD, M. (1940): Le genre Vascoceras de la Tunisie, 405 pp., 35 pis., Paris. dans Ie Turonien du Gard. Bull. Soc. -- (1896): Les ammonites du Cretace su- 40 Tatsuro MATSUMOTO

perieur de l'Algerie. Mem. Soc. Ceol. SPATH, L.F. (1926): On new ammonites from France, Paleont., Mem. no. 17, vol. 6, fasc. the English Chalk. Ceol. Mag., vol. 63, 4, pp. 1-24, pis. 14-19 [1-6J (1896); vol. pp. 77-83. 7, fasc. 1-2, pp. 25-88, pis. 1-12 [7-18J STOLICZKA, Ferdinand (1863-66): Ammoniti­ (1897) . dae, with revision of the Nautilidae. etc. PERVINQUIERE, L. (1907) : Etudes de pale­ In BLANFORD, M.F., and STOLICZKA, F., ontologie tunisienne. I. Cephalopodes des 1861-66. The fossil Cephalopoda of the Terrains secondaires. Carte geol. Tunisie, Cretaceous rocks of southern India. Mem. 483 pp., 27 pis. Ceol. Surv. India, Palaeont. Indica, vol. REESIDE, J.B., Jr. (1923): A new fauna from 1, pp. 41-216,pls. 26-94. the Colorado Group of southern Montana. TANAKA, Keisaku (1963): A study on the U.S. Ceol. Surv. Prof. Paper 132-B, pp. Cretaceous sedimentation in Hokkaido, 25-33, pis. 11-21. Japan. Ceol. Surv. Japan, Rept. no. 197, REYMENT, R.A. (1954) : Some new Upper Cre­ 119 pp. taceous ammonites from Nigeria. Colo­ T AUBENHAUS, H. (1920): Die Ammoneen der nial Ceol. Min. Resources, vol. 4, no. 3, Kreideforma tion Palastinae und Syrien. pp. 248-270, pis. 1-5. Zeit. deutsch. Paliistina- Vereins, vol. 43, -- (1955): The Cretaceous Ammonoidea of 58 pp., 9 pis. southern Nigeria and the southern Came­ WIEDMANN, Jost (1960) : Le cretace superieur roons. Ceol. Surv. Nigeria, Bull. no. 25, de l'Espagne et du Portugal et ses cepha­ 112 pp., 24 pis. lopodes. 848 Congres Soc. savantes, 1959, REYMENT, R.A. and TAIT, E.A. (1972): Fau­ pp. 706-764. nal evidence for the orgin of the South -- (1964): Le cretace superieur de l'Es­ Atlantic. 24th Intern. Ceol. Congr., sect. pagne et du Portugal et ses cephalopodes. 7, pp. 316-323, Montreal. Estudios Ceologicos, Inst. Lucas Mallada, SASTRY, M.V.A. and MATSUMOTO, Tatsuro C.S.I.C. (Espana). vol. 20, pp. 107-148. (1967): Notes on some Cretaceous am­ WRIGHT, C.W. (1957): In MOORE, R.C. [Ed­ monites from southern India. Part II. itorJ : Treatise on Invertebrate Paleontol­ Mem. Fac. Sci., Kyushu Univ., ser. D, ogy, Part L, Mollusca, Cephalopoda, Am­ vol. 18, no. 1, pp. 1-5, pis. 1. monoidea, p. Ll-L490, Geol. Soc. Amer., SCHNEEGANS, D. (1943): Invertebres du Cre­ & Univ. Kansas Press. tace superieur du Damergou (Territoire YABE, Hisakatsu (1904): Cretaceous Cepha­ du Niger). Bull. Direc. Min. Couvern. lopoda from the Hokkaido. Part II. Jour. Cener. Afri. Ocrid. Franc., no. 7, pp. 87- Coll. Sci., Imp. Univ. Tokyo, vol. 20, art. 166, 8 pis. 2, pp. 1-45, pis. 1-6.

Explanation of Plate 8

Fig. 1. Vascoceras sp. aff. V. durandi (THOMAS and PERON) ...... Page 29 GK. H5674, from loco R2513a, Member Mh of the Obira area. Lateral (a) and rear (b) views, x 0.9. Fig. 2. Fagesia thevestensis (PERON) ...... Page 32 Inner whorl of GT. 1-3249, from loco T610 b-c, unit II b of the Saku area. Lateral (a), rear (b) and frontal (c) views, xO.9. Kyushu University [I. HAYAMIJ photos, without whitening for Fig. 1 and with whitening for Fig. 2. MATSUMOTO: Vascoceratid ammonites Plate 8 614. Vascoceratid ammonites 41

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PROCEEDINGS OF THE PALAEONTOLOGICAL SOCIETY OF JAPAN

13 *~~~~:~ 19731f. • 1f.~ • ~~v;l:, 19731f. Upper Permian fusulinaceans from Kuzuu, 1 fj1613 CY<), 17 El (7./C) I<::'JR~I:*~!!Il"jl:ftl;llk~~': Tochigi Prefecture, Japan .... H. & H. IGO Planktonic foraminiferal sequence in a ~~~*;t!<~ 1<::..t5'-' , L r#JIlli~ nt-= (~1Jn~ 681'5) 0 piston core from off Sanin district, Japan Sea ...... UJIIE, H. & M. ICHIKURA i4Ji?'~~~tHlmf,ll'ff Distribution of the Japanese Miogypsina, SCOR (Scientific Committee on Ocean Re­ with a description of a new species search) (J) Working Group 37 (J)iJ1i1liJJI<::.-? ...... UJIIE, H. '-"L ...... ·frijfJJPi$a ~2~~R~t1:l~G~km.~.~~~U(J) Committee on Pacific Neogene Stratigraphy mt'LI~"?'-"L ...... Jjt,m::tfi (J)~.lL C. ~~i1tiiID I~ -? '-', L ...... ~ IJIPi$i5 !r.l1ll!:=r:~i!f.1:lEk T'/ J 7"7 '/!1 r '/1t1:lI<::'''?'-'' ffill:J~rll~!ffIJ,*=Jl!!.g-~ 9 @]r,f,~ •...... •. iBi:J:I:4= L (Tf,~) ...... ~l.lJI~Hg·~I:ID. i$ Vascoceratid ammonites from the Turonian ~ ~IJ ~(/f i1il of Hokkaido ...... MATSUMOTO, T. Evolution of Lepidolina multiseptata (Per- ~1:::t-'F'i$!Ik!:9JG(J);tfr~ft ...... Wlr±l~-n~ mian foraminifer) in East Asia ...... OZAWA, T. Genetic polymorphism and geographic vari­ ation in Umbonium (Preliminary obser- 11M A IIIYt illl vation) ...... OZAWA, T. 1t1:l1l\:(J)i1§1t~:I'II~~ 7 II.- •••••••••••••• iiliJ]( ttl­ f:kFl3W,='/;j:H1;l1:(J)*~· hii'i=,!i'lFjJ!ffIJf.r ...... Some Lower Cretaceous plants newly found ...... ~j'W§ififrij from the Ryoseki Group, Kochi Prefec­ Growth of massive corals ...... HAMADA, T. ture, Japan ... KIMURA, T. & M. HIRATA "Cladochonus" from the red bed of Malaya Older Mesozoic plants from the Tae-Dong (Malaysia) ...... HAMADA, T. Series, Korea. Pt. 1 ...... On some fossil Scleractinia from Japan ...... KIMURA, T. & B.-K. KIM ...... EGUCHI, M . Phylogenetic system of vascular plants .. Japanese Tertiary cassidids and their re­ ...... ASAMA, K. lated species of the west coast of North i'PIiUIfI< IIII~I:!Ik1.i(J) i'i J I=l}\lfl~ M:J-:" fE~f.r~l<::' America ...... KANNO, S. "?'-"L ...... I],JR ttl- "Pecten (Amusium) ocala nus Dall" and some M**~.~mMiliKft£~~~~*(J)~._ pectinid with internal radial ribs ...... f\::1<::'-?1r'L ...... +JR *tI- ...... KANNO, S. Lower Carboniferous conodonts from Mitsu- IJ'~J.ij( • flJ:,liMilfJfMf.r?r~(J) ";I.;;t r ~ J 71'1" 1<::. sawa, north of Itsukaichi, Tokyo ...... "?'-"L ...... ;'5jl't*~ ...... IGo, H. & F. KOBAYASHI 7'/~T1 H1f~i'i:(J)~~-Gaudryceras Ii:'* Geologic age of the Mamba Group. Kanto ;f:;t c. L- L ...... " ., ...... -'jZ!I1t~l,@ Massif. Pt. 1. Corals and fusulinaceans 1=~i/]i~l'iiU1&rl~(J) r ~ t7AFoln[I*I<::'''?'-''L ...... IGo, H. & S. TAKIZAWA ··································*ttfIjUffi Ditto. Pt. 2. Conodonts ...... Faunal analysis of the Upper Miaoli (Byo- ...... TAKIZAWA, S. & H. IGO ritsu) Group in the Tunghsiao area, 43

Miaoli Province, Formosa, China ...... Late Cenozoic planktonic Foraminifera ...... KOTAKA, T. from the oil-bearing formations of Japan Proposed correlation of Neogene sediments (t-t~il) .. MAIYA, S., T. SAITO, & T. SAT a in Noto and Oga Peninsulas, Japan, Planktonic foraminifers of the Nishikuro­ using planktonic diatoms (t-t1m) ...... sawa Formation, northwestern Honshu, ...... BURCKLE, L. H. Japan (f-tRjll) ...... SAITO, T. & S. MAIYA

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New Series No. 89 April 20, 1973

CONTENTS

TRANSACTIONS

611. HIRANO, Hiromichi: Biostratigraphic study of the Jurassic Toyora Group. Part II ...... 1

612. NISHIMIYA, Katsuhiko and Y AMAGIWA, Nobuo: Coral fossils from the Kosode Formation, Yamanashi Prefecture...... 15

613. HUZIOKA, Kazuo and T AKAHASI, Eitaro: A Triassic Hepatica from the Omine coal-field, Southwest Honshu, Japan ...... 24

614. MATSUMOTO, Tatsuro: Vascoceratid ammonites from the Turonian of Hokkaido (Studies for the Cretaceous ammonites from Hokkaido and Saghalien-XXVI) ...... 27

PRODCEEDINGS ...... 42