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- ISPH 2015 logo (front cover) designed by Jasmina Wiemann. The logo highlights several aspects well suited for the Bonn, 2015 meeting. The dwarf sauropod dinosaur, Europasaurus, stands in front of a histology-filled silhouette of the main dome of Poppelsdorf Palace, the main venue for ISPH 2015. This Late Jurassic sauropod is a fitting representative, as it was discovered in Lower Saxony, Germany, and its dwarf status was verified with histological investigations. - Cover, program and abstract book designed by Aurore Canoville and Jessica Mitchell. - Program and abstract book editors: Aurore Canoville, Jessica Mitchell, Koen Stein, Dorota Konietzko-Meier, Elzbieta Teschner, Anneke van Heteren, and P. Martin Sander. ISPH 2015 – Bonn, Germany ISPH 2015 – Bonn, Germany Table of Contents TABLE OF CONTENTS Symposium Organizers and Acknowledgements ...................................................... 4 Welcome Address ........................................................................................................ 5 Program ........................................................................................................................ 6 Main Events ........................................................................................................ 6 Venues ................................................................................................................ 8 Scientific Sessions / Oral Presentations ........................................................... 11 Scientific Sessions / List of Posters .................................................................. 18 Abstracts ..................................................................................................................... 21 Oral Presentations ............................................................................................. 21 Posters ............................................................................................................... 80 Workshops and Lab Tour ...................................................................................... 108 Dinner Cruise ........................................................................................................... 110 Fieldtrip Guide ........................................................................................................ 111 List of Participants .................................................................................................. 116 ISPH 2015 – Bonn, Germany 3 Symposium Organizers and Acknowledgements 3rd International Symposium on Paleohistology July 2-5, 2015 – Bonn, Germany Steinmann Institute, University of Bonn Nussallee 8, 53115 Bonn, Germany Symposium Convenor P. Martin Sander Host Committee Jessica Mitchell & Aurore Canoville Program Committee (Co-Chairs) Aurore Canoville Filippo Bertozzo Dorota Konietzko-Meier Anneke van Heteren Jessica Mitchell Dorota Konietzko-Meier Koen Stein P. Martin Sander Christen Shelton Field Trip Committee Koen Stein Anneke van Heteren Ella Teschner Dorota Konietzko-Meier Katja Waskow Koen Stein Jasmina Wiemann Elzbieta Teschner Tzu-Ruei Yang Volunteers Benjamin Gutwasser, Jelle Heijne, Philipp Knaus, Marie-Claire Koschowitz, Jens Lallensack, Sebastian Pommerening, Adun Samathi, Michael Scheil, Kayleigh Wiersma, Tanja Wintrich, Timo Yigit, Ivette Weitz Acknowledgements Special thanks go to Beate Mühlens-Scaramuzza, Steinmann Institute, for providing administrative assistance. The Rheinische Friedrich-Wilhelms-Universität Bonn is thanked for providing the conference venues. We are grateful to the Deutsche Forschungsgemeinschaft (DFG) for providing funding. 4 ISPH 2015 – Bonn, Germany Welcome Address WELCOME ADDRESS It is our great pleasure to welcome you to the 3rd International Symposium on Paleohistology at the University of Bonn in Bonn, Germany, from July 2nd to July 5th, 2015. The University of Bonn has a rich history in paleontological and evolutionary research, making it a befitting venue. In fact, Bonn is arguably the birthplace of paleobiology, manifested in the first-ever published life reconstruction of an extinct animal in its environment by Georg August Goldfuß in 1831. In this seminal study on the pterosaur Scaphognathus crassirostris, Goldfuß recognized the presence of hair and a vascularized wing membrane, arguing that the animal was an active flyer and had an endothermic metabolism. Paleohistology has much to contribute to such research, as we are all aware of, and it is in this spirit that we hope to hold the meeting. We are looking forward to 60 talks and 26 posters, an ever- growing number of presentations from ISPH meeting to ISPH meeting! In addition, there will be plenty of time for informal discussion, and we are also very excited about the thin sectioning workshops offered by Ellen-Thérèse Lamm and the trouble shooting session and lab tour at our department. The venue will be the very building that Goldfuß reached his paleobiological conclusions in, the Poppelsdorf Palace. This converted Rococo summer palace of Clemens August, Elector of Cologne, dating from 1740, was given to the University of Bonn in 1818. We have a lecture hall for 200 people and will take our breaks and exhibit posters in two historic halls overlooking the botanical garden of the university. Bonn is a medium-sized city of 300 000 inhabitants and the former capital of Germany. It is also the birthplace of Ludwig van Beethoven. His presence is felt everywhere, including in the motto of the city “Freude. Joy. Joie. Bonn”. Our region, the Rhineland, is very open to the world, and once an event has taken place twice, it is already considered a “tradition”. It is in this spirit that we invite you to participate in another successful International Symposium on Paleohistology, ISPH 3, after the meetings in Sabadell and Bozeman. This symposium would not have been possible without the energetic and creative members of the host committee, in particular Jessica Mitchell and Aurore Canoville, and student helpers to whom I extend my heartfelt thanks for their hard work with never wavering cheerfulness. We also acknowledge the generous funding by the German Research Foundation (DFG) that made the participation of many students possible. P. Martin Sander, Symposium Convenor ISPH 2015 – Bonn, Germany 5 Program – Main Events PROGRAM - MAIN EVENTS THURSDAY, JULY 2, 2015 Time Event Location 8:00 AM – 12:00 PM Beginner's Histology Übungsraum & Labor, Steinmann- Workshop Institut (Nussallee 8, rear building) 1:00 PM – 5:00 PM Advanced Histology Workshop Übungsraum & Labor, Steinmann- Institut (Nussallee 8, rear building) 5:00 PM – 6:30 PM Lab Tour with preparator Olaf Labor, Steinmann-Institut (Nussallee 8, Dülfer rear building) 6:00 PM – 7:30 PM Registration Entrance to Goldfuß Museum, (Nussallee 8, front building) 7:00 PM – 10:00 PM Ice Breaker Goldfuß Museum (Nussallee 8, front building) FRIDAY, JULY 3, 2015 Time Event Location 8:00 AM – 9:00 AM Registration Gartensaal, Poppelsdorf Palace 8:00 AM – 9:00 AM Upload Talks Hörsaal Zoologie, Poppelsdorf Palace 8:00 AM – 9:00 AM Poster Setup Stucksaal, Poppelsdorf Palace 9:00 AM – 9:15 AM Welcome Address Hörsaal Zoologie, Poppelsdorf Palace 9:15 AM - 9:45 AM Keynote Session I Hörsaal Zoologie, Poppelsdorf Palace (H. Woodward) 9:45 AM – 10:30 AM Session I Hörsaal Zoologie, Poppelsdorf Palace 10:30 AM – 11:00 AM Coffee Break Gartensaal and Stucksaal, Poppelsdorf Palace 11:00 AM – 12:00 PM Session I (continued) Hörsaal Zoologie, Poppelsdorf Palace 12:00 PM - 1:30 PM Lunch Break (Upload Talks) 1:30 PM – 2:00 PM Keynote Session II Hörsaal Zoologie, Poppelsdorf Palace (K. Mikhailov) 2:00 PM – 3:15 PM Session II Hörsaal Zoologie, Poppelsdorf Palace 3:15 PM – 3:45 PM Coffee Break Gartensaal and Stucksaal, Poppelsdorf Palace 3:45 PM – 4:15 PM Keynote Session III Hörsaal Zoologie, Poppelsdorf Palace (W. Dirks) 4:15 PM – 5:30 PM Session III Hörsaal Zoologie, Poppelsdorf Palace 6:00 PM – 8:00 PM Troubleshooting Workshop and Labor and Übungsraum, Steinmann- Core Drilling Demonstration Institut (Nussallee 8, rear building) 6 ISPH 2015 – Bonn, Germany Program – Main Events SATURDAY, JULY 4, 2015 Time Event Location 8:00 AM – 9:00 AM Upload Talks Hörsaal Zoologie, Poppelsdorf Palace 8:00 AM – 9:00 AM Poster Setup Stucksaal, Poppelsdorf Palace 9:00 AM – 9:30 AM Keynote Session IV Hörsaal Zoologie, Poppelsdorf Palace (E.M. Griebeler) 9:30 AM - 10:45 AM Session IV Hörsaal Zoologie, Poppelsdorf Palace 10:45 AM – 11:15 AM Coffee Break Gartensaal and Stucksaal, Poppelsdorf Palace 11:15 AM – 12:45 PM Session IV (continued) Hörsaal Zoologie, Poppelsdorf Palace 12:45 PM – 2:15 PM Lunch Break (Upload Talks) 2:15 PM – 3:30 PM Session V Hörsaal Zoologie, Poppelsdorf Palace 3:30 PM – 5:30 PM Poster Session Stucksaal, Poppelsdorf Palace SUNDAY, JULY 5, 2015 Time Event Location 8:00 AM – 9:00 AM Upload Talks Hörsaal Zoologie, Poppelsdorf Palace 9:00 AM – 9:30 AM Keynote Session VI Hörsaal Zoologie, Poppelsdorf Palace (S. Sanchez) 9:30 AM - 10:15 AM Session VI Hörsaal Zoologie, Poppelsdorf Palace 10:15 AM – 10:45 AM Coffee Break Gartensaal and Stucksaal, Poppelsdorf Palace 10:45 AM – 11:15 AM Keynote Session VII Hörsaal Zoologie, Poppelsdorf Palace (J. Vinther) 11:15 AM – 12:15 AM Session VII Hörsaal Zoologie, Poppelsdorf Palace 12:15 PM – 1:45 PM Lunch Break (Upload Talks) 1:45 PM – 3:30 PM Session VIII Hörsaal Zoologie, Poppelsdorf Palace 3:30 PM – 4:00 PM Coffee Break Gartensaal and Stucksaal, Poppelsdorf Palace 4:00 PM – 5:30 PM Session IX Hörsaal Zoologie, Poppelsdorf Palace 6:15 PM Meet
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