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A Bollettino della Società Paleontologica Italiana, 57 (3), 2018, 217-250. Modena

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Middle Triassic ammonoid fauna from the Recoaro and Tretto areas (NE Italy) and its stratigraphic and paleobiogeographic evidence

Paolo Mietto, Stefano Manfrin & Manuel Rigo

P. Mietto, Dipartimento di Geoscienze, Via Gradenigo 6, I-35131 Padova, Italy; [email protected] S. Manfrin, Dipartimento di Geoscienze, Via Gradenigo 6, I-35131 Padova, Italy. M. Rigo, Dipartimento di Geoscienze, Via Gradenigo 6, I-35131 Padova, Italy; [email protected]

“Questo lavoro sul Triassico del Recoarese, argomento che è stato alla base di molte nostre comuni ricerche, è dedicato a Vittorio De Zanche, indimenticabile Maestro e Amico che troppo prematuramente il destino ha voluto sottrarre alla famiglia e alla comunità scientifica”

“This paper on the Triassic of the Recoaro Area, which was the starting topic of our common research, is dedicated to Vittorio De Zanche, unforgettable Maestro and Friend, who was prematurely stolen by destiny from his family and scientific community”

KEY WORDS - Anisian, Ladinian, Recoaro area, Ammonoids, Upper Muschelkalk, Paleobiogeography.

ABSTRACT - Between the mid 19th and the beginning of the 20th centuries, several Austrian and German geologists such as Beyrich, Mojsisovics and Tornquist, studied the geology of the Recoaro area (NE Italy). They described important ammonoid faunas, the majority being collected from a stratigraphic unit known as the Nodosus Formation. Among these faunas are problematic species that have been the focus of long discussion both from a biostratigraphic and taxonomic point of view. Unfortunately, most of the historical material has been either destroyed or lost, and recent studies in the field provided only a few scarce specimens thus making taxonomic revision difficult. One of the unresolved issues is related to the possible occurrence of the ceratitids peculiar to the Upper Muschelkalk of the Central Europa (Germanic) Basin and the Sephardic Triassic province among the Alpine ammonoids. An unexpected discovery of an old ammonoid collection stored at the Museo Geologico e Paleontologico of Padova University (MGP-PD) permitted confirmation of the presence of a distinctive ammonoid species Alloceratites tornquisti (Philippi) within the Triassic Recoaro domain. Two other species of Alloceratites have also been found in the Val Gola section near Trento. These occurrences are significant taxonomically, biostratigraphically and in relation to Middle Triassic sequence stratigraphy.

RIASSUNTO - [Le faune ad ammonoidi medio triassici di Recoaro e Tretto (Italia nord orientale) ed il loro significato stratigrafico e paleobiogeografico] - Fra la seconda metà del 1800 e gli inizi del secolo successivo, vari geologi di lingua tedesca (fra i quali Beyrich, Mojsisovics e Tornquist) studiarono la geologia del Recoarese, descrivendo fra l’altro importanti faune ad ammonoidi, quasi totalmente provenienti da una unità stratigrafica nota come Formazione a Nodosus. Si tratta di un insieme di specie assai problematico, costituito prevalentemente da forme di difficile collocazione tassonomica e biostratigrafica. Purtroppo il materiale storico andò quasi completamente distrutto o disperso, rendendo così impossibile una completa revisione tassonomica; le ricerche condotte in tempi recenti nel Recoarese, peraltro, hanno fornito solo scarsi elementi faunistici di confronto. Uno dei problemi aperti riguardava la possibile presenza, fra le forme alpine, di ceratitidi caratteristici del Muschelkalk superiore dei Bacini dell’Europa Centrale (bacini Germanico e Sefarditico). Tuttavia, il ritrovamento inaspettato di una piccola e antica collezione di ammonoidi presso il Museo Geologico e Paleontologico dell’Università di Padova ha permesso di confermare la presenza nel Triassico del Recoarese di una specie tipica del Muschelkalk superiore, Alloceratites tornquisti (Philippi), presenza che assume particolare significato non solo dal punto di vista tassonomico ma anche biostratigrafico oltre che in termini di stratigrafia sequenziale, anche perché confortato dal rinvenimento di altri ceratitidi riferibili al genereAlloceratites nella sezione della Val Gola presso Trento.

INTRODUCTION in particular with his monograph about the Triassic of the Recoaro and Tretto areas (Tornquist, 1901c). Successively, The Triassic of the Recoaro area (NE Italy) has been other authors, among which Fabiani (1920a, b), Rosenberg studied by many paleontologists since the 19th century (1959, 1961), De Boer (1963), Epting et al. (1976), (Arduino, 1769; Buch, 1848; Schauroth, 1855a, b, 1859; Mostler (1976), Barbieri et al. (1977), further described Benecke, 1866, 1868; Beyrich, 1875a, b; Mojsisovics, the geology of the whole Recoaro area. A deep revision 1876; Bittner, 1879a, b, 1880, 1881a, b, 1883, 1884; of the stratigraphy and geology of this area is referable to Tommasi, 1904; Trevisan, 1932; Selli, 1938), because Barbieri et al. (1980, 1982), De Zanche & Mietto (1981) of the large amount of fossils yielded by the Anisian and Gianolla et al. (1998), who also described in detail (Middle Triassic) stratigraphic units, such as the Gracilis the Triassic volcanism of the Recoaro area. Formation, the Voltzia beds, and the Recoaro Limestone. Beyrich (1875a) first described the occurrence The first relevant studies on the Triassic stratigraphy of this of ammonoids in a basinal stratigraphic unit of the area were conducted by Tornquist (1898b, 1899, 1900c), Recoaro area, originally considered equivalent to

ISSN 0375-7633 doi:10.4435/BSPI.2018.16 218 Bollettino della Società Paleontologica Italiana, 57 (3), 2018 the “Buchensteiner Schichten” of the Balaton and originally described in the Recoarese area by Tornquist Dolomites. However, Mojsisovics (1882) was the (1898a) as Ceratites sp. ind. aff. nodosus Bruguiere. For first author who described these ammonoid faunas, these two species documented both in the Recoarese area establishing new and biostratigraphically important and in the Upper Muschelkalk, Wenger (1957) established taxa such as the ammonoid “Trachyceras recubariense” the subgenus Ceratites (Austroceratites), which was and “Trachyceras margaritosum”. Later, Tornquist accepted as a real genus by Tozer (1980b). Nevertheless, (1896, 1898a) described new ammonoids from the same Urlichs (1997) considered dubious Austroceratites stratigraphic unit, apparently recognizing the ammonoid thuringiacus and also the presence of Austroceratites (now species Ceratites nodosus (Bruguiere), peculiar of the Alloceratites) in the Germanic Basin. Germanic Muschelkalk, along with a not well defined Along with these debatable taxa, Tornquist (1898b) species, probably related to the “nodose Ceratiten” of the described new problematic species, most of which came Germanic Basin. It has to be mentioned that at this time from the Tretto area. Unfortunately, the whole collection only three valid species of Ceratites were known from the by Tornquist stored at the Strasbourg Museum was Germanic Upper Muschelkalk, C. nodosus (Bruguiere, destroyed by fire on the th8 February 1967. 1792), C. semipartitus (Buch, 1849), and C. enodis Also the holotype of Trachyceras recubariense by (Quenstedt, 1845). For this reason, many authors used the Mojsisovics (1882: tav. VII, fig. 1), which was deposited name C. nodosus for almost all Ceratites species that were in the Museum of the Berlin University, has been named in subsequent years following Philippi (1901b) and lost. Fortunately, the type specimens of Trachyceras Riedel (1916). Since this finding, the basinal stratigraphic margaritosum and Arpadites trettensis, described by unit of Recoaro has been named “Nodosus Schichten” Mojsisovics (1882), and stored at the Geologische or Nodosus Formation. The Illyrian to Longobardian Bundesanstalt of Wien (GBA) are still available. age of this formation has been constrained by conodont We invested many years of work on the revision biostratigraphy (Mietto & Petroni, 1979, 1980). of the Anisian ammonoid faunas from Recoaro. We The unexpected record of Upper Muschelkalk looked for fossil localities, but we also tried to identify ammonoids in the Alps arose bewilderments among specimens from this area in the collections of several other contemporary research workers (Arthaber, 1896a, museum in Berlin, Königsberg, Jerusalem, and Strasbourg. b; Diener, 1900; Philippi, 1901a), including Tornquist Only few specimens belonging to species described by (1900a, b, 1901a, b), who referred his specimens to the Mojsisovics and Tornquist were found. Unexpectedly, ammonoid species Ammonites subnodosus Münster, 1831 a small collection of ammonoids from the Tretto area and Ammonites nodosus Bruguiere, after a comparison was recovered at the Museo Geologico e Paleontologico between his ammonoid collection from the Recoaro dell’Università di Padova (MGP-PD). The name of area and similar specimens from the Romania and the collector and the year of collection are not known, Sardinia terrains (Tornquist 1900a, b, 1901a, b). In fact, probably this collection was deposited in the Museum Tornquist (1898a, b) classified the ammonoid specimens at the beginning of the 20th century. A well-preserved he collected from the Recoaro and Germanic basins as specimen of “ammoniti nodose” from the Recoaro area Ceratites subnodosus (Mstr) Tornq. and Ceratites sp. ind. was identified among this fauna. This specimen, referred aff.nodosus Bruguiere, respectively. During his thorough to Austroceratites tornquisti (Philippi, 1901b) by Mietto reclassification work, Tornquist (1898a) realised that the & Manfrin (1995a), confirmed the occurrence of Upper emended Ceratites subnodosus was not corresponding Muschelkalk ammonoids within the Alpine basin. As to the Ceratites subnodosus described by Mojsisovics results this finding is of great significance for the Triassic (1882) from the Schreyer Alpe, establishing thus the new sequence stratigraphy (De Zanche et al., 1993; Gianolla species Ceratites mojsisovicsi Tornquist (1898a) for these et al., 1998), as well as on the still poorly constrained specimens. Diener (1900) rejected the new classification correlation between Germanic and Tethyan provinces proposed by Tornquist, by considering the Ammonites (Brack et al., 1999; Balini et al., 2010; Jenks et al., 2015). subnodosus Münster as a nomen nudum and thus no longer valid. Furthermore, Diener re-established the species described by Mojsisovics (Ceratites subnodosus), GEOLOGICAL SETTING AND STRATIGRAPHY and erected the new species Ceratites muensteri (Diener, 1900) for all the specimens described as Ceratites By a geological point of view, the Recoaro area is subnodosus (Münster) by Tornquist. Tornquist (1900b) meant as a vast region of the north-western Vicenza (Fig. renamed, without any description, “nodose Ceratiten” 1) Prealps, in which the Hercynian crystalline basement from Recoaro as Ceratites haani, but this taxon has been is covered by the Permo-Triassic stratigraphic succession. considered as nomen nudum (Urlichs, 2016). Philippi From West to East, the Recoaro region includes (Fig. 1c) (1901b) agreed with Diener on the value of Ceratites the area between the Agno Creek valley and the Recoaro subnodosus Mojsisovics and accepted the species Terme hollow; the Leogra Creek and the relative Tretto Ceratites muensteri described by Diener, but moved hollow; and the Posina Creek basin with the Laghi hollow. the specimens from the Recoaro area to the new species The Recoaro Terme village (Fig. 1) is located in the Ceratites tornquisti. high portion of the Agno creek valley, and it is easily Riedel (1916) went through the Germanic ceratitoids accessible from Vicenza. Westwards, from Schio begins taxonomy, including all the taxa from Vicentinian area the Leogra valley that ends to the Pian delle Fugazze described by Tornquist. He confirmed the speciesCeratites mountain Pass (1162 m). This mountain Pass separates tornquisti, but established the new species Ceratites the Sengio Alto Mountain (in the South) from the Pasubio thuringiacus dubitatively including also the species group (in the North). The Tretto hollow extends to the P. Mietto et alii - Middle Triassic ammonoids from the Recoaro and Tretto areas (NE Italy) 219

Fig. 1 - Location map of the Recoaro and Tretto areas indicating the fossiliferous sites. 1A: Rotolon Valley; 1B: Contrada Ronchi; 2C: Contrada Pozzani; 2D: Nogare; 3E: Recoaro 1000; 4G: Creme Valley; 4H: Molino Mazzega; 4I: Orco Valley; 4J: San Rocco; 4K: San Ulderico; 4L: Timonchio Creek. 220 Bollettino della Società Paleontologica Italiana, 57 (3), 2018

North of Schio, west of Leogra and it is delimited to the tectonic depressions on the carbonate platform of Monte East and North by the low Astico creek. Behind the Tretto Spitz Limestone with abundant volcano-sedimentary hollow, there is a road going from Arsiero to the Posina supply. Above the volcanic complex which can be creek basin and to the Laghi hollow. The Recoaro area locally eroded, Carnian siliciclastic-carbonatic sediments also includes the highest sector of the Leno di Vallarsa occurred (De Zanche & Mietto, 1977), in transitional creek valley (Trento), which flows from the Pian delle succession with the onset of the carbonate platform of Fugazze Pass westwards, ending into the Adige river close the Dolomia Principale. to the Rovereto town. The last sector of the Recoaro area consists in the left branch of the Leno di Vallarsa creek, representing by the Sinello valley (west of the Sengio Alto MATERIALS AND METHODS Mountain), which begins at Spèccheri village, and going to the Recoaro Terme hollow through the Campogrosso All the studied material belongs to a small historical pass (1457 m). collection recently discovered and stored at the MGP-PD The first description of the Recoaro Permo-Triassic and to other specimens collected during field trips (Tab. succession was given by Tornquist (1901c), and 1). The latter specimens are also stored at the MGP-PD. successively reviewed by Barbieri et al. (1980) and De These specimens have been compared with historical Zanche & Mietto (1981). Gianolla et al. (1998) identified specimens stored at the GBA - Geologische Bundesanstalt different sequence stratigraphic units in the Recoaro (Wien, Austria) and with a specimen at the MCR - Museo successions that permitted a precise correlation to the Civico di Rovereto (Trento, Italy), a cast of which is stored surrounded areas, in particular with the Trento area and the at the MGP-PD. For biostratigraphical, paleogeographical Dolomites (Fig. 2). The Triassic succession of the Recoaro and sequence stratigraphy relationships, other specimens area differs from that of the surrounding areas, being stored at the MGP-PD from the Val Gola (near Trento) and characterised by a reduced thickness, sometimes with gaps the Rite Mountain (Cadore) have been analysed (Tab. 1). and hiatuses. The succession was deposited in shallow marine enviroments affected by volcanism, Illyrian to Longobardian in age. This peculiar volcanism changed FOSSIL-BEARING LOCALITIES from acid to basic, ending with the appearance of basalts. During the Triassic, the Recoaro area was located close Considering the stratigraphic succession of the to a landmass in which the exposed basement complex, Recoaro area proposed by De Zanche & Mietto (1981), known as Southern Mobile Belt (Viel, 1981), supplied all ammonoids are from the following formations (Fig. 2): abundant metamorphic clasts into the sedimentary or volcano-sedimentary units of Recoaro, from the Spathian 1. Recoaro Limestone (Pelsonian). to Norian (De Zanche & Farabegoli, 1981; De Zanche & 2. Sturia Limestone (Illyrian). Mietto, 1984). This scenario is also supported by AGIP 3. Monte Spitz Limestone (Illyrian-Fassanian). cores drilled to the North of Vicenza (Pozzo Villaverla 4. Nodosus Formation (Illyrian-Longobardian). 1) and offshore Venice (Pozzo Assunta 1) (Gianolla et al., 1998). 1. Recoaro Limestone Above the metamorphic basement, the stratigraphic Only few and small ammonoids have been documented succession consists in continental red sandstones from the Recoaro Limestone (Meneguzzo, 1868), along belonging to the Val Gardena Sandstones, in transition the Rotolon Valley. with calcareous and dolomitic sabkha deposits of the Bellerophon Formation, Late Permian in age. A. Rotolon Valley The Lower Triassic succession is common throughout Geographical coordinates - WGS84 N 45° 43’ 09.4”; the entire Southern Alps, and is characterised by the E 11° 10’ 00.3”; altitude 1137 m. deposition of silty to dolomitic facies belonging to the Werfen Formation and Lower Serla Dolomite. The Anisian Accessibility - From the center of Recoaro Terme succession is instead peculiar of the Recoaro area and (Fig. 1), along the road passing through Merendaore to consists in stratigraphic order of the Gracilis Formation, Campogrosso, reaching the Rifugio alla Guardia (1038 Voltzia beds, Recoaro Limestone, Tretto Conglomerate m). From this mountain hut, just after the followed hairpin and Sturia Limestone (Fig. 2). The Voltzia beds and the bend, there is a mule path going to an incised valley that is Tretto Conglomerate are very important by a sequential exposed on the NW side of the Obante Mountain (named interpretation and they represent a temporary emersion Guglie del Fumante). This incised valley is characterised of the area, documented also by the presence of tetrapod by several debris flow events involving the stratigraphic footprints and plant remains in this unit (Gianolla et succession between the Gracilis Formation to the Dolomia al., 1998; Avanzini & Mietto, 2008). The Monte Spitz Principale Fm (De Zanche & Mietto, 1981). Limestone is a unique carbonate platform of the Recoaro area, Illyrian to Fassanian in age. During this time Fossils - A specimen of Proarcestes sp., specimen interval, portions of the carbonate platform of Monte Spitz R(al)1 (MGP-PD 29155), was collected from the Recoaro Limestone drowned differentially due to syndepositional Limestone debris. tectonics and completely emerged in the Late Ladinian, just before the acme of the volcanic activity. The first B. Contrada Ronchi evidences of the Middle Triassic volcanic activity are Geographical coordinates - WGS84 N 45° 41’ 30”; E documented in the Nodosus Formation, by the infilling of 11° 11’ 57.3”; altitude 773 m. P. Mietto et alii - Middle Triassic ammonoids from the Recoaro and Tretto areas (NE Italy) 221

Fig. 2 - Stratigraphic column of the Triassic of Recoaro area and correlated sections in Southern Alps (from Gianolla et al., 1998, modified).

Accessibility - From Recoaro Terme, along the road Fossils - From these outcrops, an abundant fauna reaching the Cesare Battisti hut (Agno di Lora) to the Storti composed by brachiopods, crinoids, gastropods and locality (Fig. 1). From this locality, along a broken path to bivalves is exposed by dissolution/alteration of the rock contrada Ronchi (770 m), some outcrops of gray yellow- surface (cf. De Zanche & Mietto, 1981). Among this fauna, spotted limestones belonging to the Recoaro Limestone only one specimen of ammonoid ascribed to Bulogites are easily recognizable. zoldianus (Mojsisovics, 1882) (specimen EA12, MGP- 222 Bollettino della Società Paleontologica Italiana, 57 (-), 2018

Repository Biostratigraphic units Sample Taxa Locality Stratigraphic units Age legit Previous finding MGP-PD (Mietto & Manfrin, 1995)

R(al)1 29155 Proarcestes sp. Rotolon Valley - Recoaro Terme Recoaro Limestone Pelsonian ? A. Lovato NO EA 12 31919 Bulogites zoldianus (Mojsisovics) Contrada Ronchi - Recoaro Terme Recoaro Limestone Pelsonian binodosus Subzone Gb. Cracco NO

PZ 3.1 29152 cf. Megaceratites sp. Contrada Pozzani - Tretto - Schio Sturia Limestone Illyrian trinodosus Subzone P. Mietto & S. Manfrin NO PZ 3.2 29153 Megaphyllites cf. oenipontanus Mojsisovics Contrada Pozzani - Tretto - Schio Sturia Limestone Illyrian trinodosus Subzone P. Mietto & S. Manfrin NO CLT.1 29154 Norites cf. falcatus Arthaber Nogare - Tretto - Schio Sturia Limestone Illyrian trinodosus Subzone A. Monegato NO

R(al)2 31921 Proarcestes sp. Recoaro 1000 - Recoaro Terme Monte Spitz Limestone Illyrian ? P. Mietto NO

SRT.dt.11 31967 Hungarites sp. San Rocco - Tretto - Schio Nodosus Formation Illyrian ? Nevadites Zone P. Mietto YES SRT.dt.3 29243 Lecanites misanii (Mojsisovics) San Rocco - Tretto - Schio Nodosus Formation Illyrian ? Nevadites Zone P. Mietto NO SRT.dt.4a 29244 cf. Beyrichites sp. San Rocco - Tretto - Schio Nodosus Formation Illyrian ? Nevadites Zone P. Mietto NO SRT.dt.4b-c 29245a, b cf. Beyrichites sp. San Rocco - Tretto - Schio Nodosus Formation Illyrian ? Nevadites Zone P. Mietto NO SRT.dt.5 31924 Epigymnites sp. San Rocco - Tretto - Schio Nodosus Formation Illyrian ? Nevadites Zone P. Mietto NO SRT.dt.6 31925 Epigymnites sp. San Rocco - Tretto - Schio Nodosus Formation Illyrian ? Nevadites Zone P. Mietto NO SRT.dt.7 31923 Proacestes sp. San Rocco - Tretto - Schio Nodosus Formation Illyrian ? Nevadites Zone P. Mietto YES DD(1) 29231 Proacestes sp. San Rocco - Tretto - Schio Nodosus Formation Illyrian ? Nevadites Zone ex old coll. MGP-PD YES DD(2) 29232 Proarcestes sp. San Ulderico - Tretto - Schio Nodosus Formation Illyrian ? Nevadites Zone ex old coll. MGP-PD YES AA(1) 29229 Parasturia sp. San Ulderico - Tretto - Schio Nodosus Formation Illyrian ? Nevadites Zone ex old coll. MGP-PD NO CC(1) 29230 Eoprotrachyceras curionii (Mojsisovics) San Ulderico - Tretto - Schio Nodosus Formation Fassanian curionii Subzone ex old coll. MGP-PD YES ORC.1 31922 Eoprotrachyceras cf. curionii (Mojsisovics) Orco Valley - Tretto - Schio Nodosus Formation Fassanian curionii Subzone E. Di Lallo YES CRM.1 29239 Epigymnites credneri (Mojsisovics) Creme Valley - Recoaro Terme Nodosus Formation Fassanian recubariensis Subzone Gb. Cracco & A. Lovato NO CRM.2a, b 29240a, b Falsanolcites recubariensis (Mojsisovics) Creme Valley - Recoaro Terme Nodosus Formation Fassanian recubariensis Subzone Gb. Cracco & A. Lovato YES VO.4 29236 Falsanolcites gortanii (Pisa) Molino Mazzega - Tretto - Schio Nodosus Formation Fassanian recubariensis Subzone P. Mietto NO VO.6 29237 Eoprotrachyceras gervasuttii Fantini Sestini Molino Mazzega - Tretto - Schio Nodosus Formation Fassanian recubariensis Subzone L. Giusberti NO VO.5 29238 Proacestes sp. Molino Mazzega - Tretto - Schio Nodosus Formation Fassanian recubariensis Subzone L. Giusberti & A. Toller YES HH(1) 29235 Falsanolcites recubariensis (Mojsisovics) Timonchio creek - Tretto - Schio Nodosus Formation Fassanian recubariensis Subzone ex old coll. MGP-PD YES FO47 cast 31920 Falsanolcites recubariensis (Mojsisovics) Trambileno - S. Colombano - Trento M. Spitz Limestone Fassanian recubariensis Subzone ex coll. MCR YES FF(1) 29234 Alloceratites tornquisti (Philippi) San Ulderico - Tretto - Schio Nodosus Formation Illyrian recubariensis Subzone ex old coll. MGP-PD YES SRT.dt.2 29242 Protrachyceras margaritosum (Mojsisovics) San Rocco - Tretto - Schio Nodosus Formation Longobardian margaritosum Subzone P. Mietto YES SUT.dt.1 31928 Arpadites arpadis (Mojsisovics) San Ulderico - Tretto - Schio Nodosus Formation Longobardian ? longobardicum Subzone P. Mietto YES SRT.dt.8 31927 Arpadites cf. telleri (Mojsisovics) San Rocco - Tretto - Schio Nodosus Formation Longobardian ? longobardicum Subzone P. Mietto YES SRT.dt.1 29241 Arpadites sp. San Rocco - Tretto - Schio Nodosus Formation Longobardian ? longobardicum Subzone P. Mietto YES SRT.dt.9 31926 Arpadites sp. San Rocco - Tretto - Schio Nodosus Formation Longobardian ? longobardicum Subzone P. Mietto YES SRT.dt.10 31932 Arpadites aff.bracki Fantini Sestini San Rocco - Tretto - Schio Nodosus Formation Longobardian ? longobardicum Subzone P. Mietto NO 59A(GG) 29228 Arpadites cf. ventisettembris Tornquist San Ulderico - Tretto - Schio Nodosus Formation Longobardian ? longobardicum Subzone ex old coll. MGP-PD YES

G 4.1 31961 Alloceratites toulonensis (Riedel) Val Gola section - Trento Val Gola Limestones Illyrian chiesense Subzone F. Cramerotti NO VG 155.1 31962 Alloceratites thuringiacus (Riedel) Val Gola section - Trento Val Gola Limestones Illyrian chiesense Subzone P. Mietto NO MRT 1.1 31885 Noetlingites sp. Mt. Rite section - Cibiana - Belluno Dont Formation Pelsonian balatonicus Subzone P. Mietto & P. Gianolla NO

Tab. 1 - List of the Triassic ammonoids from the Recoaro area and correlated sections housed in the Geological and Paleontological Museum of Padova University (MGP-PD). P. Mietto et alii - Middle Triassic ammonoids from the Recoaro and Tretto areas (NE Italy) 223

PD 31919) (legit G. Cracco) was found. This species D. Nogare is peculiar of the binodosus Subzone (sensu Mietto & Geographical coordinates - WGS84 N 45° 44’ 43.4”; Manfrin, 1995a; for a sightly different correlation, see E 11° 21’ 03.7”; altitude 568 m. Vörös, 2003; Monnet et al., 2008; Jenks et al., 2015), and it is known from the Pelsonian of Southern Alps to Accessibility - After the crossroad going to Pozzani, the Balaton highlands. The identification of this specimen a rock wall consisting of canalised conglomeratic banks confirms the Pelsonian age of the Recoaro Limestone, in crops out along the main road, just after a left sharp turn agreement with the brachiopod age (see Barbieri et al., in locality Nogare (after the indication to Casa Laita). 1980). Lithology and fossils - The outcrop of Nogare (Fig. Another specimen in the available collection at Museo 4) consists in the stratigraphic transitional interval Geologico “D. Dal Lago” (Valdagno, Vicenza) with the between the Tretto Conglomerate and Sturia Limestone, number (MGV 25 (ig 292953), is from the same Contrada as illustrated in De Zanche et al. (1979b). Five cm above Ronchi area. However, it was collected from the Rotzo the last conglomeratic bank, at the base of the Sturia Formation moraine debris (legit O. Gonzo) and belongs to Limestone, a specimen of Norites cf. falcatus Arthaber, Juraphyllites libertus (Gemmellaro, 1884), Early Jurassic 1896a was documented from the black marls rich in in age (Mietto, 1985). organic matter (specimen CLT.1, MGP-PD 29154), Illyrian in age (Gianolla et al. 1998). 2. Sturia Limestone Tornquist (1900c) collected Sturia sansovinii 3. Monte Spitz Limestone (Mojsisovics, 1869) along with other undetermined Even if Tornquist (1899) wrote an entire book on the ceratitoids from the “Sturia-Kalk” (i.e. Sturia Limestone), documented fauna from the Monte Spitz Limestone, no which crops out along a path close to San Ulderico locality, ammonoid was found from this stratigraphic unit. From in the Tretto area. Further research in this area permitted the sequence stratigraphic point of view (Fig. 2), this to identify the Sturia Limestone but no other ammonoid unit is equivalent to the carbonate platforms belonging specimens were found. However, ammonoids from this to the Contrin Formation, to the “Lower Edifice” (sensu stratigraphic unit were collected at two other localities in De Zanche et. al., 1995) and to the Sciliar 1 and Sciliar 2 the Tretto area (Fig. 1). cropping out in the Dolomites (Gianolla et al., 1998). This means that the Monte Spitz Limestone is late Illyrian to C. Contrada Pozzani latest Fassanian in age (Gianolla et al., 1998). Following Geographical coordinates - WGS84 N 45° 45’ 22.6”; this stratigraphic interpretation, it is possible to justify the E 11° 21’ 00.4”; altitude 537 m. huge thickness variation of the Monte Spitz Limestone on the Recoaro area, probably due to synsedimentary tectonic Accessibility - After two km from Schio, towards events and to sudden drowning or emersion phases Piovene, it is possible to reach the Tretto hollow, keeping (Barbieri et al., 1977, 1980; De Zanche & Mietto, 1982). left to Santorso. From Timonchio locality, keeping left, there are two roads going up to San Rocco di Tretto or to San Ulderico di Tretto, both joining in Bosco del Tretto locality. Driving up to San Ulderico, there is a crossroad on the right to Pozzani, going direct to a bentonite quarry, known as “Caolino del Tretto” (Mietto, 2003). From Pozzani locality, there is a path to Valle dell’Orco, at the end of which a small outcrop is present, belonging to the Monte Spitz Limestone.

Lithology and fossils - This outcrop consists of a breccia bank with sharp clasts of Monte Spitz Limestone, laying on 50 cm thick coarse calcarenites with dasycladaceae algae, which overlie 70 cm of bituminous limestones (Fig. 3). In the Recoaro area, this latter lithotype has not been yet described as member of the Monte Spitz Limestone, and it can be equivalent to the Dark Margon Limestones in Valsugana and Adige Valleys (De Zanche & Mietto, 1986, 1989), or to the Moena Formation of the western Dolomites (Masetti & Neri, 1980). At Pozzani locality, this bituminous stratigraphic unit overlain a fetid black limestone (20 cm thick) with dasycladaceae algae belonging to the Sturia Limestone. At the top of this fetid black bank, ammonoid cf. Megaceratites sp. (specimen PZ3.1, MGP-PD 29152) and Megaphyllites cf. oenipontanus Mojsisovics, 1882 (PZ 3.2, MGP- PD 29153) were collected, indicating an Illyrian age Fig. 3 - Stratigraphic log of the Contrada Pozzani section (Tretto (trinodosus Subzone sensu Mietto & Manfrin, 1995a). area). Scale column in cm. 224 Bollettino della Società Paleontologica Italiana, 57 (3), 2018

beginning of the Vallarsa Valley, from the Spèccheri Lake area to the Alpe di Campogrosso which are connected with the high Agno Valley (Barbieri et al., 1980), the Monte Spitz Limestone 2 crops out.

Fossils - An external mould of Falsanolcites recubariensis (Mojsisovics, 1882) was collected from a loose block of Monte Spitz Limestone 2 (specimen Fo 47) and is now stored in the Museo Civico di Rovereto (one cast of FO 47 is stored in the MGP-PD, n. 31920). This species is index of the recubariensis Subzone, Fassanian in age (sensu Mietto & Manfrin, 1995a).

4. Nodosus Formation This stratigraphic unit is the richest in ammonoids in the Recoaro area. This formation consists in volcanorudites/ arenites, or volcanosiltites in its upper portion, intercalated to cherty or nodular micritic limestones. It is variable both in thickness and lithofacies, depending on the locality. These variations are consisting with the interpretation of this unit, that represents the infilling of syntectonic troughs cutting the carbonate platform represented by the Monte Spitz Limestone (Fig. 2). On the western Tretto area, above the Monte Spitz Limestone, this formation is represented by thin bituminous-carbonate laminites, which are very similar to the Plattenkalk member of the Buchenstein Formation of the Dolomites. In the Nodosus Formation, the “rosso ammonitico” facies are present, documenting local drowning event of the carbonate platform, such as those recorded in the Fongara and Campogrosso area (Barbieri et al., 1980; Mietto & Petroni, 1980). Furthermore, the Nodosus Formation testifies the Fig. 4 - The outcrop of Nogare (Tretto area) with emphasised beginning of the acid volcanic activity well known in the Tretto Conglomerate and Sturia Limestone lithostratigraphic the Recoaro area, named as Southern Mobile Belt (Viel, boundary and the position (with star) of the finding of Norites cf. falcatus Arthaber. 1981). Clearly, the name of the Nodosus Formation is not correct, because Ceratites nodosus (Bruguiere) is not documented in this unit. However, the original name is E. Recoaro 1000 preserved for historical reason (cf. Barbieri et al., 1980). Geographical coordinates - WGS84 N 45° 41’ 26.8”; The age of the Nodosus Formation is comprised between E 11° 13’ 27.1”; altitude 936 m. the Illyrian to the early Longobardian, but ammonoid documentation is restricted from the Hungarites Zone to Accessibility - About three km after Valdagno towards probably the longobardicum Subzone. Urlichs (2016: p. Recoaro Terme (Fig. 1), on the right of the San Quìrico 31) noted that “The age of the “Nodosus Beds” is Early locality, it is possible to reach Fongara village driving to Late Fassanian/Early Ladinian (sic! = Longobardian) along the southern side of the Monte Spitz. This road goes in the sense of Balini et al. (2010), because the three directly to the Valdagno hutte and to the Monte Falcone subsequent index species Eoprotrachyceras curionii, ski lifts, named Recoaro 2000. Just after Fongara, in the Protrachyceras margaritosum, and Eoprotrachyceras Chempele locality, it is possible to reach the top of the gredleri occur in these beds. This age is indubitable, ski lift directly from Recoaro, which is named Recoaro because the two former species have been introduced 1000. From this point, it is easy to follow a wide path by Mojsisovics (1882) from the silicified Buchenstein along the northwestern side of the Monte Spitz, passing Formation of Schilpario/Vicenza (sic! = Lombardy), through a forest, and reaching a small rock wall of ca. 10 which were later re-named “Nodosus Beds” by Tornquist m, representing the Monte Spitz Limestone. (1898a, b)”. Apart from the mistakes the latter statement is Fossils - From this outcrop a specimen of Proarcestes inconsistent, because Tornquist (1898a, b) re-named sp. (specimen R(al)2 [MGP-PD 31921]) was collected. “Subnodosus Beds” the previous “Nodosus Beds” Unfortunately, this taxon is not biostratigraphically useful. of Recoaro and Tretto/Vicenza and not the silicified Buchenstein Formation of Schilpario. Also the occurrence F. San Colombano of the quoted three index species arises some problems. Accessibility - San Colombano locality is situated As noted by Balini (1994), Eoprotrachyceras curionii within the Leno di Vallarsa creek basin (Trentino-Alto was defined by Mojsisovics (1882) from the Buchenstein Adige region), and it is easily accessible driving ca. 1.5 Formation of Schilpario (Bergamo) and later noted by km along Vallarsa valley from Rovereto (Trento). At the Tornquist (1898b) also from the “Nodosus Beds” of P. Mietto et alii - Middle Triassic ammonoids from the Recoaro and Tretto areas (NE Italy) 225

Tretto area. Protrachyceras margaritosum was defined Lithology and fossils - gray-greenish nodular and by Mojsisovics (1882) from the Valle dell’Orco in the cherty limestones belonging to the Nodosus Formation Tretto area (Vicenza). (Eopro) Trachyceras gredleri was are exposed in the riverbed. A single specimen of defined by Mojsisovics (1882) from the Wengen Beds of Eoprotrachyceras cf. curionii (Mojsisovics, 1882) Alpe di Siusi (Bolzano) and from the condensed facies was collected (secimen ORC.1, MGP-PD 31922) from of M. Clapsavon in Carnia (Udine). Nevertheless, the calcareous beds cropping out in this locality, This choice of the specimen illustrated by Mojsisovics (1882: specimen is early Fassanian in age (curionii Subzone). pl. 17), and selected as lectotype by Krystyn et al. (2004), In addition to this specimen, we have studied a type is problematic by a taxonomic point of view and for its specimen of Trachyceras margaritosum Mojsisovics, biostratigraphic meaning, as exposed in the following 1882, illustrated on plate 82, fig. 1 of Mojsisovics (1882), Biostratigraphic Appendix. In any case, specimens now stored at the GBA (GBA 1882/003/0164). This referable to Protrachyceras gredleri were never found in specimen, together with a second one that is presently the Recoaro area, in disagreement with the statement of missing, was collected (Mojsisovics, 1882; p. 128) “... Urlichs (2016: pp. 31-32). in rothen, Tuffbeimengungen enthaltenden Buchesteiner Kalk von Val dell’Orco im Tretto bei Schio...”. G. Val di Crème Geographical coordinates - WGS84 N 45° 41’ 22.7”; J. San Rocco di Tretto E 11° 10’ 37.1”; altitude 1009 m. Geographical coordinates - WGS84 N 45° 45’ 23.2”; E 11° 21’ 27.8”; altitude 636 m. Accessibility - After Contrada Ronchi (Fig. 1) towards Asnicar locality, there is a wild path going westward to Accessibility - The locality San Rocco di Tretto was Malga Creme (Creme hutte). first described by Tornquist (1898b, 1901c) and yielded a rich and known fauna from Tretto area (Fig. 1). This Fossils - During maintenance road works, along locality is situated along a path starting from the San the main axis of the Creme valley, some nodular Rocco di Tretto village to the Orco Valley, ca. 100 m blocks belonging to the upper part of the Nodosus below the village. Fm yielded ammonoid specimens of Falsanolcites recubariensis (Mojsisovics) (specimen CRM.2, MGP-PD Lithology and fossils - Reddish micritic limestones 29240), and Epigymnites credneri (Mojsisovics, 1882) belonging to the Nodosus Formation crop out along (specimen CRM.1, MGP-PD 29239), both referable to the the path, but all the ammonoid specimens collected by recubariensis Subzone sensu Mietto & Manfrin (1995a), Tornquist (1898b) are from debris and blocks accumulated latest Fassanian in age. by farmers to clean and prepare the fields for cultivations. The San Rocco del Tretto fauna described by Tornquist H. Molino Mazzega quarry (1898b, 1901c) consisted of Ceratites subnodosus (Mstr) Geographical coordinates - WGS84 N 45° 45° 14.3”; Tornquist (1898a) vel Ceratites muensteri Diener in E 11° 21’ 35.3”; altitude 526 m Tornquist (1901) [= Alloceratites tornquisti (Philippi)], Ceratites vicentinus Tornquist, 1898b [= Paraceratites Accessibility - In the Tretto area, there is a road vicentinus (Tornquist) in Urlichs & Mundlos (1980) connecting Timonchio with San Rocco di Tretto (Fig. 1). but the suture line suggests a possible attribution to the About 500 m after Sbrogliaoro houses locality (370 m), genus Schreyerites], Ceratites beneckei (Mojsisovics, at the second U-shaped turn, a white road going up to the 1882) [= Beyrichites beneckei (Mojsisovics)], Arpadites right side of the Orco valley leads to an abandoned quarry cinensis Mojsisovics, 1882, A. telleri Mojsisovics, of shale, known as Molino Mazzega quarry. 1882, A. arpadis (Mojsisovics, 1870), A. ventisettembris Tornquist, 1898b, Hungarites sanroccensis Tornquist, Lithology and fossils - This quarry was used for the 1898b, Beyrichites reuthense Mojsisovics [= Beyrichites “kaolin” production because of the alteration of the acid reuttensis (Beyrich, 1867)], Ptychites uhligi Mojsisovics, volcanoclastic rocks of the Nodosus Formation, very 1882, Protrachyceras margaritosum (Mojsisovics, 1882), abundant in this area (cf. De Zanche et al., 1979a). Several and Proarcestes pannonicus (Mojsisovics, 1882). This calcarenitic and micritic limestones are intercalated with is a clearly heterogeneous fauna, in which some specific these volcanoclastic layers. They yielded few specimens attributions cannot be unfortunately confirmed (Bagnoli of Eoprotrachyceras gervasuttii Fantini Sestini, 1994 et al., 1997). A stock of species referable to a time interval (specimen VO.6, MGP.PD 29237), Falsanolcites gortanii between Fassanian and early Longobardian is identifiable in (Pisa, 1966) (specimen VO.4, MGP-PD 29236) and this fauna. However, two ammonoid assemblages cannot fit Proarcestes sp. (specimen VO.5, MGP-PD 29238). All these with this time interval. The first, represented bySchreyerites taxa are typical of upper Fassanian (recubariensis Subzone). vicentinus, Beyrichites beneckei, Beyrichites reuttensis and Ptychites uhligi, is early Illyrian in age. The second I. Valle dell’Orco assemblage is instead Longobardian in age, composed by Geographical coordinates - WGS84 N 45° 45’ 46.2”; Arpadites cinensis, A. telleri and A. arpadis, most probably E 11° 21’ 10.8”; altitude 504 m. longobardicum Subzone (sensu Mietto and Manfrin, 1995a). This time interval is generally not documented in field and Accessibility - After the abandoned Molino Mazzega probably partly in gap in the Recoaro area (cf. Gianolla quarry (Fig. 1), along a small path, it is possible to reach et al., 1998). Only few specimens were directly collected the Orco creek riverbed. from the San Rocco di Tretto area or studied form the 226 Bollettino della Società Paleontologica Italiana, 57 (3), 2018 collection stored at the MGP-PD. They are all latest Illyrian Longobardian (margaritosum Subzone). Notably, some to Longobardian in age. In fact, these few ammonoids are problematic species described by Tornquist (1898b) such represented by Proarcestes sp. [specimen DD(1) (MGP- as Ceratites prettoi and C. vicarius, which are probably PD 29231), SRT.dt.7 (MGP-PD 31923)], Epigymnites sp. referred to the early Illyrian, are not consistent with the (SRT.dt.5, MGP-PD 31924; SRT.dt.6, MGP-PD 31925), cf. stratigraphy. Beyrichites sp. (SRT.dt.4a, MGP-PD 29244; SRT.dt.4b-c, The ammonoid specimens from the old collection MGP-PD 29245), Lecanites misanii (Mojsisovics, 1882) stored at the MGP-PD and examined by the writers, (SRT.dt.3, MGP-PD 29243), Hungarites sp. (SRT.dt.11, were found from the old path (by labels). They are MGP-PD 31967), upper Illyrian (probably Nevadites Zone); classified by Proarcestes sp. [DD(2), MGP-PD 29232], Protrachyceras margaritosum (Mojsisovics) (SRT.dt.2, Parasturia sp. [AA(1), MGP-PD 29229], = upper Illyrian- MGP-PD 29242), which are Longobardian (margaritosum Fassanian; Eoprotrachyceras curionii [CC(1), MGP-PD Subzone); Arpadites sp. (SRT.dt.1, MGP-PD 29241; 29230] = Fassanian (curionii Subzone); Alloceratites SRT.dt.9, MGP-PD 31926), Arpadites aff. bracki Fantini tornquisti (Philippi) [FF(1), MGP-PD 29234] = Fassanian Sestini, 1994 (SRT.dt.10, MGP-PD 31932), Arpadites (recubariensis Subzone); Arpadites cf. ventisettembris cf. telleri Mojsisovics (SRT.dt.8, MGP-PD 31927) is still Tornquist [59A(GG), MGP-PD 29228] = Longobardian Longobardian but possibly belonging to the longobardicum (probable longobardicum subzones). In addition to this Subzone (sensu Mietto and Manfrin, 1995a). fauna, we studied only one specimen of Arpadites arpadis Mojsisovics (specimen SUT.dt.1, MGP-PD 31928), K. San Ulderico di Tretto collected from the field in the new path, Longobardian in Geographical coordinates - WGS84 N 45° 45° 04.8”; age (longobardicum Subzone?). E 11° 20’ 42.7”; altitude 698 m. The original specimen of Arpadites trettensis illustrated by Mojsisovics (1882: tav. LXXXI, fig. 4) and examinated Accessibility - San Ulderico di Tretto is the second by the writers is stored at the GBA (GBA1882/003/0063). fossiliferous locality (after the San Rocco one) described by Tornquist (1898b, 1901), and it is located at the L. Torrente Timonchio beginning of the old path going from San Ulderico di Timonchio creek originates by the confluence of the Tretto to Palle and Soggio localities (Fig. 1). Valle dell’Orco creek and Acquasaliente creek, gathering all the materials coming from the Orco valley and San Fossils - The new path, which corresponds to the Rocco di Tretto area (Fig. 1). San Ulderico di Tretto section, is traced in the Nodosus Formation, and it was investigated for conodont Fossils - From this locality, Mojsisovics (1882) illustrated biostratigraphy (Mietto & Petroni, 1979). At present, the some specimens of Protrachyceras recubariense Mojsisovics new path is paved and the section is not exposed anymore. (= Falsanolcites recubariensis [Mojsisovics]), and During the conodont investigation, no ammonoid Protrachyceras curionii Mojsisovics (= Eoprotrachyceras was collected in situ. Instead, only one specimen of curionii [Mojsisovics]), which were successively confirmed Arpadites arpadis Mojsisovics (specimen SUT.1, MGP- by Tornquist (1898b). Only one specimen from this area is PD 31928) was collected from a volcanoclastic block stored at the MGP-PD (Mietto, 1995), which is a not well in debris of a near field. However, a rich ammonoid preserved body chamber of Falsanolcites recubariensis fauna was described by Mojsisovics (1882) and then (Mojsisovics), (specimen HH(1), MGP-PD 29235), by Tornquist (1898b) from the San Ulderico area. Fassanian in age (recubariensis Subzone). This fauna consists of Ceratites subnodosus (Mstr) Tornquist (1898a) vel Ceratites muensteri Diener in M. Other localities Tornquist (1901c) [= Alloceratites tornquisti (Philippi)], Some ammonoids described by Mojsisovics (1882) Ceratites sp. ind. aff. nodosus Bruguiere [= ? Ceratites are from other localities of the Recoaro Terme area, in (Austroceratites) thuringiacus Riedel in Wenger, 1957: 95 particular from the southwestern side of the Mount Spitz, = Alloceratites cf. tornquisti (Philippi) in Urlichs, 2016: which however proved to be barren during recent field 30], Ceratites prettoi Tornquist,1898b, Ceratites vicarius investigations. These localities are: Tornquist, 1898b (pro Ceratites simplex Arthaber, 1896a), 1. “Zwischen Chempele und Fantoni bei Fongara”. Arpadites ventisettembris Tornquist, Arpadites trettensis From this locality, a specimen of Arpadites sp. ind. Mojsisovics, Hungarites mojsisovicsi (Böckh) Mojsisovics ex aff. A. Arpadis was described. This specimen was [= Hungarites zalaensis (Böckh, 1873)], Hungarites n. sp. subsequently ascribed to Arpadites ventisettembris by ind. ex aff.mojsisovicsi Mojsisovics (sic!) [= Hungarites Tornquist (1898b). At Chempele locality (see Recoaro zalaensis (Böckh)], Hungarites sanroccensis Tornquist 1000), in an abandoned quarry, it is possible to recognise [= “Hungarites” sanroccensis Tornquist], Protrachyceras lithofacies peculiar of the drowning of Monte Spitz recubariense Mojsisovics [= Falsanolcites recubariensis Limestone. These lithofacies, which are similar to those (Mojsisovics)], Protrachyceras margaritosum Mojsisovics cropping out at Campogrosso (cf. Mietto & Petroni, 1980), [= Protrachyceras margaritosum (Mojsisovics)], and are assigned to the Nodosus Formation. Proarcestes pannonicus Mojsisovics [= Proarcestes 2. “... am Wege von Castagna nach Caili bei Recoaro”. pannonicus (Mojsisovics)]. An adult but incomplete specimen of Trachyceras recubariense was collected here by Beyrich and later As the San Rocco fauna, these ammonoids are not illustrated by Mojsisovics (1882: Tav. VII, fig. 1). In this homogeneous and/or not identifiable (Bagnoli et al., 1997) area there are no visible outcrops but the debris blocks but all of them are upper Illyrian (Nevadites Zone) to lower document the presence of nodular limestones, sometimes P. Mietto et alii - Middle Triassic ammonoids from the Recoaro and Tretto areas (NE Italy) 227 with red to greenish chert, along with colored tuffs. No Remarks - The specimen could be referred to ammonoids have been found from these blocks. Norites falcatus Arthaber instead of Norites gondola 3. “Unweit Cainallo bei Pornaro nächst Fongara auf (Mojsisovics). The former species is characterised by dem Monte Spitz”. From this locality, only a problematic a slender whorl section, a greater involution due to the specimen of Protrachyceras mascagni (sic!) was illustrated higher increasing in height of the whorl respect to N. by Tornquist (1898b). The lithofacies cropping out here gondola. If the test is preserved, N. gondola shows, at are known as Fongara Breccia, they are similar to those of least from intermediate growth stages, a diagnostic keeled the Chempele locality. The Fongara Breccia represents the umbilical rim. Hence, we refer the quoted specimen to the erosional phase affecting the drowning facies of the Monte Arthaber’s species even if in open nomenclature. Spitz Limestone. Considering the original descriptions and illustration, Protrachyceras mascagnii Tornquist shows Occurrence and Age - The specimen has been morphological features similar to Anolcites laczkói var. found in situ in the basal part of the Sturia Limestone falcoidea Salopek (1936: pp. 216-217, fig. 9). Moreover, interfingered with the latest conglomeratic beds of the because of its ceratitic suture line, this taxon cannot be Tretto Conglomerate. At present, the genus Norites is referred to Protrachyceras laczkoi (Diener, 1901), which documented only in the Mediterranean area, e.g. in the instead shows a subammonitic suture line (cf. Vörös, Balaton area (Vörös, 2003). In the Southern Alps its age 1998). Notably, Protrachyceras mascagnii is comparable is comprised between the latest Bithynian of the Cima with Protrachyceras lermani, illustrated by Parnes (1986), Valmedan section (Mt. Cucco area, Carnian Prealps, which was collected from Har Gevanim locality (Israel). Udine) (unpublished datum) and the latest Fassanian Instead, partially in disagreement with Parnes (1986) we (Fantini Sestini, 1994). deem diagnostic the venter morphology which appears quite similar in both species. Superfamily Ceratitoidea Mojsisovics, 1879 Family Longobarditidae Spath, 1951 SYSTEMATIC DESCRIPTIONS Subfamily Noetlingitinae Parnes, 1975

Morphological parameters used in this study are Genus Noetlingites Hyatt, 1900 detailed as follows: D: diameter (mm) = H+h+U; H: Type species Ceratites strombecki Griepenkerl, 1860 max. whorl height in D (mm); h: min. whorl height in D (mm); U: umbilicus in D (mm); W: max. whorl width Noetlingites sp. in H (mm); w: min. whorl width in H (mm); SGR(%): (Figs 5, 6a) [(H-h)/h]×100, involution rate. If not otherwise specified, all measurements are referred to specimens preserved with Material - One specimen MRT 1.1 (MGP-PD 31885) test and considering their ornamentation. The external from ex Forte Monte Rite section (Valle di Cadore, Belluno). tubercles mark the boundary of the ventral area. Description - The specimen consists of half of a very involute shell, included in a dark grey marly limestone Order Ceratitida Hyatt, 1884 from the bed MRT1 of the Dont Formation. The whorl Superfamily Noritoidea Karpinsky, 1889 section, even if only part of the specimen can be examined, Family Noritidae Karpinsky, 1889 is lanceolate with maximum width in the inner third of the flank, but above the umbilical rim. The flank at the Genus Norites Mojsisovics, 1879 beginning is smooth, then barely visible sigmoid ribs Type species Ammonites gondola Mojsisovics, 1869 are present in its latter part. The external margin is well marked, smooth, without any evidence of nodes. The Norites cf. falcatus Arthaber, 1896a keeled venter is sharp. (Pl. 1, figs 1a-c) The suture (Fig. 6a) line is partially visible and shows three fairly broad entire saddles which covers ¾ of the Material - One specimen CLT 1 (MGP-PD 29154) flank. The umbilical suture is not preserved. from the Sturia beds of Nogare (San Ulderico nel Tretto, Schio, Vicenza). Dimensions (mm) -

Description - The specimen, partly included in a dark- D H h U W w H/D U/D SGR grey matrix, is an internal mould crushed and badly preserved. MRT 1.1 43 22 16.5 4.5 9* - 0.51 0.10 33.3 A little portion of ventrolateral margin, angular in shape, and * estimated the general morphological outline fit well withNorites . The shell is clearly highly involuted, as for this genus, but the Remarks - The morphological framework and the umbilical area is not preserved. The suture line is not visible. suture line perfectly fit with the genus Noetlingites. The incompleteness of the specimen prevents from the Dimensions (mm) - attribution to N. strombecki (Griepenkerl), the unique species of Noetlingites so far known. D H h U W w H/D U/D SGR * CLT 1 - - - - 9 - - - - Occurrence and Age - The quoted specimen has been * estimated found in the Dont Formation from the ex Forte Monte Rite 228 Bollettino della Società Paleontologica Italiana, 57 (3), 2018 section (Eastern Dolomites), bed MRT 1, associated with 1898b Arpadites Arpadis Mojs. - Tornquist, p. 650. Balatonites zitteli Mojsisovics, 1882, Laeviornites sp., 1903 Arpadites Arpadis Mojs. - Frech, Pl. 10, fig. 8a-b. Bulogites sp. and Acrochordiceras sp. This assemblage 1914 Arpadites Arpadis Mojs. - De Toni, p. 130-131, Pl. 10, is referred to the pelsonian balatonicus Subzone (Mietto fig. 6. & Manfrin, 1995a). 1915 A. Arpadis v. Mojsisovics - Diener, p. 54. non 1927 Arpadites Arpadis Mojsisovics - Ogilvie Gordon, p. Noetlingites is a typical Lower Muschelkalk genus but 63, Pl. 7, fig. 14. it was found also outside the German Basin: Grossreifling 1931 Arpadites arpadis Mois. - Zivkovic, p. 93-94. in Northern Calcareous Alps, Austria (Tatzreiter, 2001), 1933 Arpadites arpadis Mojs. - Kutassy, p. 427. Aszöfö II section in the Balaton Highland, Hungary 1998 Arpadites arpadis (Mojsisovics) - Vörös, Pl. 10, figs (Vörös, 2003). The Hungarites sp. illustrated by Gu et al. 5-7. (1980: plate 2, figs 6-7) from the Chaqupu Formation in ? 1998 Arpadites toldyi Mojsisovics - Vörös, Pl. 11, fig. 5 (non Dequen (Doilungdequen country, Tibet, China), because fig. 4). of the lack of ornamentation on the flank, the well marked and smooth ventrolateral margin and sharp ventral Material - One fragmentary specimen SUT.dt.1 (MGP- keel, is here assigned to Noetlingites. On the contrary, PD 31928) from San Ulderico nel Tretto (Schio, Vicenza). the occurrence of Noetlingites in the sub-metamorphic limestones cropping out at Punta Bianca in La Spezia Gulf Description - The two poorly preserved fragmnents area (Northwestern Apennines) (Venturi & Tenneroni, of the specimen (a, b) preserved as internal moulds, 1982) is not confirmed, due to the lack of exaustive are included in a reddish hybrid limestone. The shell description and illustrations. is evolute, with a very slow increasing in height. The whorl is characterised by narrow, dense ribs, slightly sigmoid in shape. The ribs are chiefly primary, only a Family Danubitidae Spath, 1951 long intercalatory rib can be observed. Some primary Subfamily Arpaditinae Hyatt, 1900 ribs show barely visible swellings in correspondence of the umbilical area. Along the flank no nodes are present. Genus Arpadites Mojsisovics, 1879 The flank shows a suture line characterised by three entire Type species Ammonites Arpadis Mojsisovics, 1870 saddles with the first lateral lobe slightly indented.

Arpadites arpadis (Mojsisovics, 1870) Dimensions (mm) - (Pl. 1, figs 2a-b) D H h U W w H/D U/D SGR * 1870 Ammonites Arpadis Mojsisovics, p.109-110, Pl. 5, fig. 6. SUT dt.1b - 8.4 ------1873 Ammonites Arpadis Mojs. - Böckh, p. 167-170, Pl. 9, figs 12-14. Remarks - Despite of the fragmentary and poor 1882 Arpadites Arpadis E. v. Mojsisovics - Mojsisovics, p. preservation, the morphological characters of the 54-55, Pl. 25, fig. 29. specimen fit well with those ofArpadites arpadis.

EXPLANATION OF PLATE 1 Middle Triassic ammonoids from the Recoaro area. Fig. 1 - Norites cf. falcatus Arthaber; specimen CLT.1 (MGP-PD 29154); Sturia Limestone, Nogare (Tretto area, Schio, Vicenza); 1a: ventral view, 1b: right lateral view; 1c: partial left lateral view. Fig. 2 - Arpadites arpadis (Mojsisovics); specimen SUT.dt.1 (MGP-PD 31928); Nodosus Formation, San Ulderico (Tretto area, Schio, Vicenza); 2a: external print with part of internal mould; 2b: fragment of internal mould, right lateral view. Fig. 3 - Arpadites aff. bracki Fantini Sestini; specimen SRT.dt.10 (MGP-PD 31932); Nodosus Formation, San Rocco (Tretto area, Schio, Vicenza); 3a: right lateral view; 3b: oral view. Fig. 4 - Arpadites cf. telleri (Mojsisovics); specimen SRT.dt.8 (MGP-PD 31927); Nodosus Formation, San Rocco; 4a: right lateral view; 4b: ventral view. Fig. 5 - Arpadites cf. ventisettembris Tornquist; specimen 59A(GG) (MGP-PD 29228); Nodosus Formation, San Ulderico (Tretto area, Schio, Vicenza); right lateral view. Fig. 6 - cf. Beyrichites sp.; specimen SRT.dt.4a (MGP 29244); Nodosus Formation, San Rocco (Tretto area, Schio, Vicenza); 6a: trasversal section of the conch; 6b: right lateral view. Fig. 7 - Bulogites zoldianus (Mojsisovics); specimen EA12 (MGP.PD 31919); Recoaro Limestones, Contrada Ronchi (Recoaro Terme, Vicenza); 7a: left lateral view, 7b: oral view, 7c: ventral view, 7d: right lateral view. Fig. 8 - Falsanolcites gortanii (Pisa); specimen VO.4 (MGP-PD 29236); Nodosus Formation, Molino Mazzega (Tretto area, Schio, Vicenza); right lateral view. Fig. 9 - cf. Megaceratites sp.; specimen PZ3.1 (MGP-PD 29152); Sturia Limestone; Contrada Pozzani (Tretto area, Schio, Vicenza); 9a: longitudinal section, 9b: internal side of the volution. Scale bar is equal to 1 cm. P. Mietto et alii - Middle Triassic ammonoids from the Recoaro and Tretto areas (NE Italy) Pl.229 1 230 Bollettino della Società Paleontologica Italiana, 57 (3), 2018

are well rounded in section and as broad as the inter-rib spaces. On the venter a median shallow furrow is bordered by tiny keels. Only at approximate diameter of 2.7 cm the ventral keel are bordered outward by faint furrows. The suture line is not visible.

Dimensions (mm) - D H h U W w H/D U/D SGR * * Fig. 5 - Noetlingites sp., specimen MRT 1.1 (MGP-PD 31885); Dont SRT.dt.10 27.5 6.9 - - 6.4 - 0.25 - - Formation, Monte Rite section (Cadore area, Belluno). a) Lateral * estimated view (scale bar is equal to 1 cm); b) magnification of suture lines (scale bar is equal to 0.5 cm). Remarks - The species which shows better morphological similarity with the specimen under study is Arpadites Occurrence and Age - The specimen has been found bracki Fantini Sestini, 1994 that exhibits the occurrence of in debris from the Nodosus Formation. Arpadites arpadis broad, rounded, primary ribs only. Nevertheless, A. bracki is surely known in the Longobardian of the Southern Alps differs by the presence of ribs that decrease in strenght and the Balaton Highland (e.g., Mojsisovics, 1882; Frech, towards the ventral area and for the subovoidal whorl 1903; De Toni, 1914; Vörös, 1998). section instead of subquadrate as in A. aff.bracki .

Occurrence and Age - A. aff. bracki has been found Arpadites aff.bracki Fantini Sestini, 1994 in debris from the Nodosus Formation, while A. bracki is (Pl. 1, fig. 3a-b) known from the margaritosum-longobardicum subzones (Longobardian) from the Esino Limestone of Val Parina, Material - One specimen SRT.dt.10 (MGP-PD 31932) Central Lombardy (cf. Fantini Sestini, 1994). from San Rocco di Tretto (Schio, Vicenza).

Description - We have examined one silicified Arpadites cf. telleri Mojsisovics, 1882 specimen mostly included in the matrix, made of reddish (Pl. 1, fig. 4a-b) hybrid limestone. The specimen shows an evolute serpenticone coiling, subquadrate in section, characterised Material - One specimen SRT.dt.8 (MCP-PD 31927) by flanks bearing primary, proverse, projected ribs. They from San Rocco di Tretto (Schio, Vicenza).

Fig. 6 - Suture lines of ammonoids emigrated from the Central European Muschelkalk basins into the Southern Alps. a) Noetlingites sp.; specimen MRT 1.1 (MGP-PD 31885); Dont Formation, Monte Rite section (Cadore area, Belluno). b) Alloceratites toulonensis (Riedel); specimen G 4.1 (MGP-PD 31961); Val Gola Limestones, Val Gola section (Trento). c) Alloceratites tornquisti (Philippi); specimen FF(1) (MGP-PD 29234); Nodosus Formation, San Ulderico (Tretto area, Schio, Vicenza). d) Alloceratites thuringiacus (Riedel); specimen VG 155.1 (MGP-PD 31962); Val Gola Limestones,Val Gola section (Trento). Scale bars are equal to 5 mm. P. Mietto et alii - Middle Triassic ammonoids from the Recoaro and Tretto areas (NE Italy) 231

Description - We have examined one silicified Occurrence and Age - At present A. ventisettembris is specimen mostly included in the matrix, made of reddish known only from the Nodosus Formation of the Tretto area. hybrid limestone. The estimated diameter of the shell, from which only a part of the whorl is visible, is about 2 cm. The specimen shows tiny but well pointed external Family Ceratitidae Mojsisovics, 1879 nodes. Faint traces of radial ribs are present below the Subfamily Beyrichitinae Spath, 1934 nodes. On the venter a median furrow bordered by well marked keels occurs. The suture line is not visible. cf. Beyrichites sp. (Pl. 1, fig. 6a-b) Dimensions (mm) - 1898b Ceratites Beneckei v. Mojsisovics - Tornquist, p. 643-645, D H h U W w H/D U/D SGR Pl. 20, fig. 2. SRT.dt.8 - - - - 3.8 - - - - Material - One specimen SRT.dt.4a, b, c (MGP-PD Remarks - The observed features are consistent with 29244, 29245a, b) from the Nodosus Formation of San Arpadites telleri, however, the incompleteness of the Rocco (Tretto, Schio, Vicenza). specimen prevents from a sure attribution to this species. Description - SRT.dt.4a is an external print while SRT. Occurrence and Age - A. cf. telleri has been found dt.4b and dt.4c are two fragments of the internal mould in debris from the Nodosus Formation, while A. telleri included in the matrix, made of reddish hybrid limestone. Mojsisovics is documented in the longobardicum Only the fragments .4b and .4c are described. Subzone (Longobardian) from the Esino Limestone of The former, about 18 mm in diameter, is characterised Val Parina, Central Lombardy (cf. Fantini Sestini, 1994). by a flank bearing numerous, thin, sometimes fasciculated The species is also present in the Longobardian of Val di ribs. The same are sigmoid in shape. The fasciculated Cino (Mojsisovics, 1882) and probably in Zlatar Mounts ribs bear, in the inner third of the flank, tiny nodes. In the in Croatia (Zivkovic, 1931). cross section of specimen SRT dt.4c the specimen shows an oval then slightly trapezoidal whorl section while the visible part portion of the body chamber appears elliptical Arpadites cf. ventisettembris Tornquist, 1898b in shape. The suture line is not visible. (Pl. 1, fig. 5) Dimensions (mm) - Material - One specimen 59A(GG) (MGP-PD 29228) from San Ulderico nel Tretto (Schio, Vicenza). D H h U W w H/D U/D SGR SRT.dt.4b - - - 3.5* 4* - - - - Description - We have examined one specimen SRT.dt.4c 23.4* 11.1* 8.8* - 7.6* - 0.47* 0.15* 26.1* included in the matrix, made of reddish biocalcarenite. * The specimen shows a complete flank, and a diameter of estimated 1.9 cm. The shell is evolute, characterised by ribs as broad as the interspaces, proverse, slightly projected near the Remarks - The quoted specimen, on the basis of external margin. Primary ribs are prevalent but frequent visible features, is consistent with “Ceratites Beneckei intercalatory ribs also occur. Only two bifurcated ribs in v. Mojsisovics” (sensu Tornquist, 1898b: pl. 20, fig. 2). correspondence of the umbilical rim are present. Venter as Nevertheless, true representatives of Beyrichites beneckei for the genus, characterised by a median furrow bordered (Mojsisovics, 1882) exhibit, on the phragmocone, a by keels. The occurrence of a narrow furrow outside the subtriangular or high ovoidal whorl section. Moreover, keels occurs. The suture line is not visible. the latter species is nearly smooth in the lower part of the flank. Otherwise, regularly developed and broad ribs Dimensions (mm) - occur in its upper part. D H h U W w H/D U/D SGR * Occurrence and Age - Beyrichites beneckei is a tipical 59A(GG) 19 5.5 4.5 9 4 - 0.29 0.47 22.2 lower Illyrian species (Mojsisovics, 1882; Mietto & * estimated Manfrin, 1995a; Monnet et al., 2008). On the contrary, the reddish hybrid limestones of the Nodosus Formation of the Remarks - Tornquist (1898b: p. 651-652, pl. 20, fig. Tretto area, from which belongs the specimen SRT dt.4, 7) depicted as new species Arpadites venti-settembris a has been checked by conodonts (Mietto & Petroni, 1980) specimen of Arpadites about 4.7 cm in diameter in which showing a time-span comprised from middle Illyrian is preserved only part of the last whorl. The specimen (avisianum Subzone) to lower Longobardian. 59A(GG) shows a pattern of ribbing fairly comparable with the type of Tornquist, despite the absence of its inner whorls. Subfamily Bulogitinae Mietto & Manfrin in Manfrin, Moreover, it must be taken into account that the section Mietto & Preto, 2005 of A. venti-settembris drowned by Tornquist (1898b) is clearly a mistake for the presence of a central keel which Genus Bulogites Arthaber, 1912 not exists in any species of Arpadites. Type species Ceratites multinodosus Hauer, 1892 232 Bollettino della Società Paleontologica Italiana, 57 (3), 2018

Bulogites zoldianus (Mojsisovics, 1882) Mojsisovics, 1882; Gaetani, 1969, Balini et al., 1993) and (Pl. 1, fig. 7a-d) in the Balaton Highland (Vörös, 1987, 2003; Tatzreiter & Vörös, 1991). Mietto & Manfrin (1995a) reported the p 1851 A. (Ceratites) binodosus Hauer, p. 114-115, Pl. 19, fig. occurrence of Bulogites zoldianus within the ammonoid 3 (non figs. 1-2, 4). assemblage of the binodosus Subzone, then in the p 1865 Ceratites binodosus Hauer - Hauer, pp. 19-23. uppermost part of the Pelsonian Substage. In the Balaton * 1882 Ceratites zoldianus Mojsisovics, p. 39, Pl. 10, fig. 6 (non Highland Bulogites zoldianus marks the homonymous fig. 5) (vidimus). Subzone of Vörös (1987) which is comprised in the 1905 Ceratites zoldianus Mojs. - Airaghi, p. 241-242. 1906 Ceratites zoldianus E. v. Mojsisovics - Mariani, p. 8. binodosus Subzone of Mietto & Manfrin (1995a), as stated 1915 Ceratites zoldianus v. Mojsisovics - Diener, p. 92. also by Vörös (2003: table S-2, p. 33). ? 1926 Ceratites zoldianus E. v. Mojsisovics - Assmann, p. 526, Pl. 9, fig. 5 (non fig. 6). Subfamily Paraceratitinae Silberling, 1962 ? 1928 C. zoldianus v. Mojsisovics - Schmidt, p. 296, fig. 812. 1931 Ceratites cf. zoldianus v. Mojs. - Voelcker, p. 457. 1933 Ceratites zoldianus Mojs. - Kutassy, p. 461. Genus Megaceratites Balini, 1993 ? 1937 Ceratites zoldianus E. v. Mojs. - Assmann, p. 104, Pl. 22, Type species Megaceratites fallax Balini, 1993 fig. 2 (non fig. 3). p 1963 Bulogites zoldianus (Mojsisovics) 1882 - Assereto, p. cf. Megaceratites sp. 61-64 (non Text-fig. 22), Pl. 6, fig. 3 (non fig. 4-6). (Pl. 1, fig. 9a-b) p 1969 Bulogites aff. zoldianus (Mojsisovics, 1882) - Gaetani, p. 519-520, Pl. 36, fig. 9 (non fig. 8), ?10. Material - One specimen PZ 3.1a,b (MGP-PD non 1986 Bulogites ex gr. zoldianus (Mojsisovics, 1882) - Rakus, 29152a,b) from the Sturia Limestone of Contrada Pozzani p. 82-83, Text-fig. 9, Pl. 1, fig. 6. (Tretto, Schio, Vicenza). 1987 Bulogites zoldianus (Mojsisovics, 1882) - Vörös, Pl. 3, fig. 1. 1991 Bulogites zoldianus (Mojsisovics, 1882) - Tatzreiter & Description - The specimen is completely included in a Vörös, Pl. 2, fig. 1. matrix made of a dark grey limestone. As a consequence, p 2003 Bulogites zoldianus (Mojsisovics, 1882) - Vörös, p. 90-91 the rock sample has been cut to see part of longitudinal (non Text-fig. A-17), Pl. A-6, fig. 1. and transversal sections, marked a and b. Half of the last whorl width suggesting a trapezoidal whorl section is Material - One specimen EA 12 (MGP-PD 31919) visible on the transversal section . The maximun width from the Recoaro Limestone of Contrada Ronchi (Recoaro is located in the inner third of the flank. The umbilical Terme, Vicenza). rim is slightly angular. With reference to the longitudinal section (PZ 3.1b) the suture line has been exposed by Description - The specimen is preserved as one third mechanical and chemical methods, and shows three broad of isolated shell, probably provided with test. The shell entire saddles alternated by three finely indented lobes. is highly trapezoidal in section, with the maximum width On the longitudinal section (PZ 3.1a) the external area is at the lateral row of nodes. The venter is low rounded, characterised by an undulated external margin obviously and bordered by a continuous row of external nodes. The due to the occurrence of nodose endings of lateral ribs. latter are of the same strength, obliquely elongated and equidistant each other. The flank bear proverse, marked, Dimensions (mm) - slightly sigmoid ribs. The ribs are wide as the intercostal spaces or slightly narrower. Primary and long intercalatory D H h U W w H/D U/D SGR ribs are regularly arranged on the flank. Only the primary PZ 3.1a 71.7 23 - - 14.8* - 0.32 - - ribs bear a row of lateral nodes which are located on the * estimated flank one third from the umbilical rim. Umbilical nodes are absent while the umbilical rim is angular. Remarks - Unfortunately, the preservation of the The suture line is not visible. specimen prevents from a sure generic attribution. Nevertheless, on the basis of the above described Dimensions (mm) - observation, we refer with open nomenclature this strongly ribbed ceratitoid to the genus Megaceratites. D H h U W w H/D U/D SGR * * * * EA 12 33 17 - 10 15 - 0.51 0.30 - Occurrence and Age - The genus Megaceratites is * estimated documented in the Southern Alps (Balini, 1993; Mietto & Manfrin, 1995a), and in Hungary (Vörös, 1998) in the Remarks - The portion of the shell at disposal, made lower part of the Illyrian. of a juvenile exemplar, exhibits the morphology of the Ceratites zoldianus of Mojsisovics (1882: pl. 10, fig. 6) and the Bulogites zoldianus of Vörös (1987: pl. 3, fig. 1), Subfamily Ceratitinae Mojsisovics, 1879 Tatzreiter & Vörös (1991: pl. 2, fig. 1) and Vörös (2003: pl. A-VI, fig. 1). Genus Alloceratites Spath, 1934 (= Austroceratites Wenger, 1957; Occurrence and Age - At present, Bulogites zoldianus = Austroceratities in Tozer, 1980b) is documented in the Southern Alps (e.g., Hauer, 1851; Type species Ceratites Schmidi Zimmermann, 1883 P. Mietto et alii - Middle Triassic ammonoids from the Recoaro and Tretto areas (NE Italy) 233

Dimensions (mm) - D H h U W w H/D U/D SGR G 4.1 - 32* - - 29* - - - - * estimated

Remarks - The above description is consistent with the diagnosis of A. toulonensis given by Riedel (1916) and Urlichs (1997), especially as regard the outline of the body chamber. The suture line (Fig. 6b), despite the bad preservation, shows also elements characteristic of some representatives of Ceratitinae.

Occurrence and Age - Alloceratites toulonensis has been found in the Toulon area (Provence, SE France) by Riedel (1916), Wenger (1957) and Urlichs (1997). Urlichs (1997, 2016) correlates the Upper Muschelkalk Ceratites-bearing beds of Provence with the enodis Zone of the German basin. It is very interesting that the occurrence of Chieseiceras cf. pemphix (Merian, 1857) (formerly Protrachyceras recubariense) is in the middle part of the spinosus Zone of SW Deutschlands (Urlichs, 1978). In the Southern Alps, the range of Ch. pemphix is comprised between the chiesense and basal part of the curionii subzones. In Val Gola both Alloceratites toulonensis and A. thuringiacus have been collected from the chiesense Fig. 7 - Schematic and partial log of the Val Gola section (Trento) Subzone. On the basis of new data from the Southern with the position of Alloceratites species (from De Zanche & Mietto, Alps, the chiesense Subzone is well correlatable with the 1986 modified). Scale column is in meters. spinosus Zone, rather than with the following enodis Zone.

Alloceratites tornquisti (Philippi, 1901b) Alloceratites toulonensis (Riedel, 1916) (Pl. 3, fig. 2a-c; Fig. 6c) (Pl. 3, fig. 1a-b; Fig. 6b) 1896 Ceratites nodosus aut. - Tornquist, p. 21. * 1916 Ceratites toulonensis Riedel, p. 68-69, Pl. 17, fig. 2. p 1898a Ceratites subnodosus (emend. Mnstr.) Tornq. (non v. 1934 Ceratites toulonensis Riedel - Spath, p. 488-489. Mojs) = nodosus aut. - Tornquist, p. 210-221, Pl. 8, fig. 1 1957 C. (Austroceratites) toulonensis Riedel - Wenger, p. 95, Pl. (non figs 2-3); Pl. 9, figs 1-2. 19, fig. 6. ? 1898a Ceratites sp. ind. aff. nodosus Brug. - Tornquist, p. 221- 1957 C. (Progonoceratites) raricostatus Riedel - Wenger, Pl. 19, 222, Pl. 10, fig. 1 (non fig. 2). fig. 5 (non p. 77-78, Pl. 9, figs 7, 9). 1900b Ceratites Haani - Tornquist, p. 94. 1997 Ceratites (Austroceratites) toulonensis Riedel - Urlichs, * 1901b Ceratites Tornquisti Philippi, p. 387-392, Pl. 39, fig. 2. pp. 4, 7, figs 2-4. 1916 Ceratites Tornquisti Phil. - Riedel, p. 65-66, Pl. 17, fig. 1. 2016 Alloceratites toulonensis (Riedel, 1916) - Urlichs, p. 31, 1934 Ceratites Tornquisti Philippi - Spath, p. 35, 488. figs 1.4-1.6 1957 C. (Austroceratites) tornquisti Philippi - Wenger, p. 95. 1995a Austroceratites tornquisti - Mietto & Manfrin, p. 556. Material - One specimen G 4.1 (MGP-PD 31961) from 2016 Alloceratites tornquisti (Philippi, 1901) - Urlichs, p. 30- the Val Gola Limestone (De Zanche & Mietto, 1986) of 31, fig. 1.2-1.3. the Val Gola section near Trento (Fig. 7). The specimen was found in debris from grey nodular lithofacies typical Material - One specimen, labelled FF(1) (MGP-PD of the “chiesense groove” (i.e. Brack & Rieber, 1986). 29234) from the Nodosus Formation of San Ulderico (Tretto area, Schio, Vicenza), more precisely from a Description - The specimen consists of the first locality at 150 m NW the church of San Ulderico (original portion of the body chamber with the last suture line label) along the old path for Palle and Soggio localities. that is somewhat visible.The body chamber shows a Urlichs (2016) indicated as holotype, by monotypy, the barely visible, slightly flattened venter but bordered specimen illustrated by Tornquist (1896) as Ceratites by a faint external margin. The preserved flank bears nodosus on which Philippi (1901b) introduced the quoted quite spaced lowered ribs that end with broad blunt species. Nevertheless, Urlichs (2016) pointed out that the nodes. The nodes are located somewhat below the specimens of Tornquist, housed at the Geological Institute external margin. Both flanks show little portions of the of the Strasbourg University, have been lost, due to the umbilical area suggesting a gently rounded outline. The destruction by fire of the Institute on 1967. At present, suture line is partly deformed but traces of broad entire the unique specimen of Alloceratites tornquisti (Philippi, saddles (3-4), followed by a denticulated umbilical 1901b) at disposal, is the specimen FF(1) (MGP-PD suture, are visible. 29234), coming from the type locality and type horizon 234 Bollettino della Società Paleontologica Italiana, 57 (3), 2018 of the original material. Consequently, this specimen is depicted by Tornquist (1896, 1898a, b) from the Nodosus here suggested as neotype. Formation of the Tretto area. The matrix of our specimen suggests a comparison with the stratigraphic interval Description - The specimen is an internal mould of the in which Falsanolcites recubariens is documented. body chamber (but with traces of the test in the umbilical Philippi (1901b: 390), in the original diagnosis of the area), about half a whorl, in which the last suture line is species, referred it as occurring also from the Germanic preserved. The shell is involute but slowly increasing Muschelkalk and, more precisely, from a bed 14 m above in height. The whorl is subexagonal in section if the the Trochitenkalke, in the locality named “Schöningen ornamentation pattern is considered. Hence, the midflank am Elm” (Lower Saxony). Later on, Riedel (1916: 66) is inflated bearing a lateral row of spines. These spines described and illustrated (pl. 17: fig. 1) a conspecific are spaced (five in half a whorl), radially elongated, and specimen, in his opinion coming from the Germanic Basin, protruded over blunt, faint, barely visible rectiradiate ribs, but without any reference of the site of finding. Wenger in one case apparently bifurcated from a lateral spine. The (1957) does not seem to confirm this assumption, so the umbilical rim is tapered, well rounded without any traces presence of A. tornquisti should not be confirmed in the of nodes. The venter is fairly narrow, low arched bordered Germanic Basin. by external row of spaced spines (eight in half a whorl), flattened in the coiling direction and in alternating position respect those of the other side. The last suture line (Fig. Alloceratites thuringiacus (Riedel, 1916) 6c) is characterised by four broad entire saddles while a (Pl. 3, fig. 3a-b; Fig. 6d) fifth very tiny saddle is inferred. 1898a Ceratites sp. ind. aff. nodosus Brug. - Tornquist, p. 221- Dimensions (mm) - 222, Pl. 10, fig. 2 (non fig. 1). * 1916 Ceratites thuringiacus Riedel, p. 66-68, Pl. 17, fig. 3. D H h U W w H/D U/D SGR 1957 Ceratites (Austroceratites) thuringiacus Riedel - Wenger, p. 95. 78.7* 32.6* 26.3 19.8 25.6* - 0.39* 0.25* 24* FF(1) 2002 Ceratites (Austroceratites) cf. toulonensis Riedel - * estimated Urlichs & Posenato, p. 199, fig. 2. 2016 Alloceratites cf. tornquisti (Philippi, 1901) - Urlichs, Figs Remarks - Alloceratites tornquisti differs from A. 1, 3. toulonensis in the lack of strong radial ribs which bear lateral spiny nodes in marginal position. Moreover, the Material - One specimen, VG 155.1 (MGP-PD 31962), former species shows a more involuted coiling, due to found in bed 155 of the Val Gola Limestone (Fig. 7) from the faster increasing in height of the whorl, quite slender the Val Gola section (Trento). in section. Description - The specimen at disposal is an internal Occurrence and Age - Besides the new specimen here mould included in a matrix made of gray limestone, described, A. tornquisti is known from the two specimens yellowish in the surface. This ammonoid is fairly

EXPLANATION OF PLATE 2 Middle Triassic ammonoids from the Recoaro area. Figs 1-3 - Falsanolcites recubariensis (Mojsisovics). 1 - specimen CRM.2a (MGP-PD 29240a), neotype; Nodosus Formation, Creme Valley (Recoaro Terme, Vicenza); 1a: right lateral view; 1b: left lateral view. 2 - specimen HH(1) (MGP-PD 29235); Nodosus Formation, Timonchio creek (Tretto area; Schio, Vicenza); 2a: left lateral view, 2b: ventral view, 2c: right lateral view. 3 - Falsanolcites recubariensis (Mojsisovics); cast of the specimen FO47 (MGP-PD 31920); M. Spitz Limestone, Trambileno (San Colombano, Rovereto, Trento); 3a: external print, 3b: internal mould. Fig. 4 - Eoprotrachyceras curionii (Mojsisovics); specimen CC(1) (MGP-PD 29230); Nodosus Formation, San Ulderico (Tretto area, Schio, Vicenza); left lateral view. Fig. 5 - Eoprotrachyceras cf. curionii (Mojsisovics); specimen ORC.1 (MGP-PD 31922); Nodosus Formation, Orco Valley (Tretto area, Schio, Vicenza); right lateral view. Fig. 6 - Eoprotrachyceras gervasuttii Fantini Sestini; specimen VO.6 (MGP-PD 29237); Nodosus Formation, Molino Mazzega (Tretto area, Schio, Vicenza); 6a: oral view; 6b: ventral view, 6c: right lateral view. Fig. 7 - Protrachyceras margaritosum (Mojsisovics); specimen SRT.dt.2 (MGP-PD 29242); Nodosus Formation, San Rocco (Tretto area, Schio, Vicenza); 7a: left lateral view, 7b: ventral view. Fig. 8 - Epigymnites credneri (Mojsisovics); specimen CRM.1 (MGP-PD 29239); Nodosus Formation, Creme Valley (Recoaro Terme, Vicenza); left lateral view. Fig. 9 - Megaphyllites cf. oenipontanus Mojsisovics; specimen PZ3.2 (MGP-PD 29153); Sturia Limestone, Contrada Pozzani (Tretto area, Schio, Vicenza); 9a: left lateral view, 9b: trasversal section. Scale bar is equal to 1 cm. P. Mietto et alii - Middle Triassic ammonoids from the Recoaro and Tretto areas (NE Italy) Pl.235 2 236 Bollettino della Società Paleontologica Italiana, 57 (3), 2018 preserved, only the last part of the phragmocone and the the Germanic Basin (Schlotheim in Thüringen) (Riedel, first part of the body chamber are checked. Despite the 1916; Wenger, 1957). incomplete preservation of the whorl, a rounded whorl section is inferred. The whorl section looks subexagonal in correspondence of the lateral spiny nodes. Umbilical Superfamily Clydonitoidea Hyatt in Meek, 1877 rim is well rounded and smooth, without nodes. On Family Trachyceratidae Hyatt in Meek, 1877 the body chamber, the midflank is inflated due to Subfamily Anolcitinae Mietto & Manfrin the occurrence of broad, quite spaced, residual part in Mietto et al., 2008 of the damaged spines. On the preserved part of the phragmocone spiny nodes occur in the midflank. Ribs Genus Falsanolcites Rieber & Brack, 2004 are not well visible or absent, but the last one, which Type species Anolcites (Trachyceras) furcosus occurs on the body chamber and fairly elevated, ends Mojsisovics, 1893 at two/third of the flank before reaching the base of the damaged spine, showing an inferred egression. An Falsanolcites gortanii (Pisa, 1966) external row of blunt spiny nodes also occur, these nodes (Pl. 1, fig. 8) are about twice with respect to those in the midflank. Both the lateral and external spiny nodes become more v* 1966 Trachyceras (Protrachyceras) gortanii Pisa, p. 633-638, spaced during ontogeny. Among the visible suture lines, text-pl. fig. 4, Pl. 81, figs 1-6. the last one is particularly well preserved (Fig. 6d). Five 1993a Protrachyceras gortanii Pisa - Brack & Rieber, p. 486, broad entire saddles, decreasing in shape towards the Pl. 13, figs 8-9. umbilicus, are present. First and second lateral lobes 1993b Protrachyceras gortanii Pisa - Brack & Rieber, Pl. 9, figs 7, 10. are finely indented. ? 2007 Falsanolcites sp. - Brack et al., Fig. 5.5.

Dimensions (mm) - Material - One specimen VO.4 (MGP-PD 29236) from D H h U W w H/D U/D SGR the Nodosus Formation of Molino Mazzega in the Orco VG 155.1 59.6 25 18.6 16 - - 0.42 0.27 34.4 Valley (Tretto, Schio, Vicenza).

Remarks - Alloceratites thuringiacus differs from Description - The specimen is juvenile and consists of A. tornquisti in having a slow increasing of the whorl only half of the flank. It is probably preserved with test, height and then a more evolute coiling. The whorl section and included in a matrix made of reddish hybrid micritic shows more spaced external spines. Moreover, the lateral limestone. The shell is involute, somewhat increasing in spines show an abrupt egression in marginal position. height. The preserved portion of the flank bears about From A. toulonensis, the quoted taxon differs by the 22 sigmoid ribs, serried and rounded in section, broader lack of strong radial ribs and of lateral spiny nodes in than the interspaces. The ribs are mainly bifurcated at the marginal position also in the phragmocone. Moreover, umbilical rim or in the inner third of the flank or just a little the ribs are quite stronger in the body chamber. For the over. Primary and intercalatory ribs are apparently absent. general features of the body chamber and, in particular Three rows of rounded nodes (umbilical, marginal and the evolute coiling and the strong egression of the row of external) occur along the flank, as for the quoted species the lateral nodes, the specimen is well comparable with (cf. Pisa, 1966). The suture line is not visible. Ceratites thuringiacus Riedel (1916: pl. 17, fig. 3) and also with the specimen from the Tretto area indicated Dimensions (mm) - by Tornquist (1898a: pl. 10, fig. 1 only) as dubitatively suggested by Wenger (1957: p. 95). Urlichs (1997: p. 7) D H h U W w H/D U/D SGR does not consider C. thuringiacus as valid species for VO.4 34* 15.2* 10.9* 7.9* - - 0.45* 0.23* 39.4* the bad and incomplete preservation of the holotype (by * estimated monotypy). Urlichs (2016) classified both the specimens of Tornquist (1898a: pl. 10, figs 1-2) asAlloceratites cf. Remarks - The specimen, for its general morphological tornquisti. As exposed above, the writers do not agree features perfectly fits to the original description of the with this interpretation. Urlichs & Posenato (2002) quoted species. documented Alloceratites cf. toulonensis in Sardinia, in the “Muschelkalk” of Punta del Lavatoio locality Occurrence and Age - At present, Falsanolcites near Alghero. Due to the quik shift of the lateral nodes gortanii is documented in the Southern Alps and towards the ventral area during growth, and the general particularly in the Red Ammonitic Limestones of Mt. morphological features, we deem reasonable to refer this Clapsavon at Fienili Chiasteons and Clap di Val in specimen to A. thuringiacus. Carnia (Pisa, 1966) and in the Buchenstein Formation at Seceda in Gardena Valley, Dolomites (Brack & Rieber, Occurrence and age - Alloceratites thuringiacus was 1993a, b). The quoted species most probably occurs in found associated with representatives of Chieseiceras the “caotico” west of Forcella Giau in Dolomites (Brack chiesense (Mojsisovics, 1882) in the homonymous et al. 2007). Unfortunately, the ventral area of this Subzone of the Val Gola section near Trento (De Zanche specimen is not illustrated. From writers’ unpublished & Mietto, 1986; Mietto & Manfrin, 1995a). This taxon is data, the species is also documented at Punta Zonia present also in the Tretto area (Tornquist, 1898a) and in section in Dolomites. The species is recorded from the P. Mietto et alii - Middle Triassic ammonoids from the Recoaro and Tretto areas (NE Italy) 237 recubariensis Subzone of the Fassanian Substage (Mietto mould (similarly to the original specimen), coming & Manfrin, 1995a). from Val di Crème (Recoaro Terme, Vicenza) from an isolated carbonate block ascribable of the Nodosus Formation, was examined (Pl. 2, fig. 1). Probably the Falsanolcites recubariensis (Mojsisovics, 1882) outer preserved whorl is the adapical portion of the body (Pl. 2; figs 1a-b, 2a-c, 3a-b) chamber. Another specimen (specimen HH1 – MGP- PD n° 29235), characterised by a large fragment of a * 1882 Trachyceras recubariense Mojsisovics, p. 114, Pl. 7, body chamber (Mietto 1995: p. 37), is also referred to fig. 1. Falsanolcites recubariensis. It was collected from the non 1882 Trachyceras recubariense E. v. Mojsisovics - type area (Timonchio Creek, Tretto area). The choice Mojsisovics, p. 114, Pl. 5, fig. 3. of a juvenile specimen as neotype is due to its better non 1886 Trachyceras cf. recubariense Mojs. - Polifka, p. 76-77, preservation, showing the morphological characters of Pl. 8, figs 1-2. 1898b Protrachyceras recubariense Mojs. - Tornquist, p. the quoted species. Even if the general preservation of the 661-662. specimen is not very good, some portions of the selected 1898 Protrachyceras recubariense E. v. M. - Geyer, p. 133, neotype (MGP-PD n° 29840a) allow a sure diagnosis. fide Pisa, 1966, Pl. 73, fig. 2. The neotype shows a fairly evolute coiling with an v 1900 Protrachyceras cfr. recubariense Mojs. - Tommasi, p. ovoidal whorl section and rounded umbilical wall. The 25, Pl. 3, fig. 4. flanks exhibit marked sigmoid ribs, rounded in section non 1901 Protrachyceras recubariense v. Mojs. - Diener, p. 24-25. when well preserved, well elevated and as wide as the non 1912 Protrachyceras ex aff. recubariense Mojs. - Salopek, inter-rib spaces. Intercalatory ribs are apparently absent. p. 17, Pl. 1, fig. 5. Bifurcated ribs are much more frequent than primary ? 1913 Anolcites furcosus Mojs. - Simionescu, pp. 299-300, pl. ribs. The branching point of ribs is at the first lateral row 2, fig. 7. 1914 Protrachyceras recubariense Mojs. - Horn, p. 20-21. of tubercles. Four rows of nodes are present: umbilical, ? 1922 Protrachyceras nov. spec. ex aff.recubariensis v. Mojs. lateral, marginal and external. The umbilical nodes appear - Welter, p. 157, Pl. 156(2), figs 17-19. as rounded swellings whereas the lateral nodes, located 1927 Protrachyceras cf. recubariense Mojsisovics - Ogilvie- near the umbilical ones, are blunt and barely visible. The Gordon, p. 60. marginal nodes, weak and rounded, are located near to the v 1966 Trachyceras (Protrachyceras) recubariense Mojs. - external ones. The latter are well rounded and elevated, Pisa, p. 646-652, Pl. 72, figs 1-2; Pl. 73, figs 1-2; Pl. respect to the marginal nodes, and in alternating position n.t., figs 3, 5. with those of the other side. vp 1966 Trachyceras (Protrachyceras) gredleri Mojs. - Pisa, p. The suture line is not visible. 639-642, Pl. 70, fig. 3 (not figs 2, 4). ? 1973 Protrachyceras sp. aff. recubariense Mojs. - reitzi Boechk - Collignon, p. 156-157, Pl. 8, fig. 1. Dimensions of the neotype (mm) - non 1978 Protrachyceras recubariense (Mojsisovics) - Urlichs, p. 6-7, Fig. 1, Pl. 1, fig. 1. D H h U W w H/D U/D SGR 1986 Eoprotrachyceras recubariense (Mojs.) - Brack & MGP-PD 29840a - 20.5 - - 17.4 - - - - Rieber, Pl. 5, fig. 5. 1993b Eoprotrachyceras recubariensis (Mojs.) - Brack & Other material - Two specimen, the first HH(1) Rieber, Pl. 8, fig. 3; Pl. 9, figs11-12. (MGP-PD 29235) from debris of the Nodosus Formation v 1995 “Anolcites” Recubariensis - Mietto, p. 37. (Timonchio creek, Tretto, Schio); a second specimen FO v 1995a “Anolcites” recubariensis (Mojsisovics, 1882) - Mietto 47(cast) (MGP-PD 31920) from debris of the Calcare di & Manfrin, Pl. 4, figs 1, 3. Monte Spitz 2 (Trambileno, San Colombano, Trento). ? 1998 Eoprotrachyceras cf. recubariense (Mojsisovics) - Vörös, Pl. 9, fig. 3. The original specimen is stored in the Museo Civico di v 2004 “Anolcites” recubariensis - Gervasi, p. 18. Rovereto (Trento). 2007 Falsanolcites recubariense (Mojs.) - Brack et al., Fig. 5.4. Description - The specimen HH(1) consists of a large badly preserved weathered portion of a body chamber Designation of the neotype - MGP-PD 29840a (Mietto, 1995). The ventral area is broad, characterised (CRM.2a). The original material (Mojsisovics, 1882: pl. by external nodes fairly flattened in the coiling direction 7, fig. 1), stored in Berlin, unfortunately has been lost. and in alternating position with those of the other side. At present, it is possible to define a neotype from the The venter exhbits a rounded furrow. The whorl section type area (Recoaro, NE Italy), justified for the critical is subquadrangular. The visible part of the flank bears biostratigraphical and taxonomical significance of this numerous strong primary proverse ribs, about fifteen for species (see also the suggestion of Pisa, 1966), according a third of the preserved whorl. to art. 75 of the ICZN. The specimen FO 47 is an external print of a little portion of a flank in which marked proverse slightly Stratum typicum and locus typicus - Nodosus projected ribs occur. These ribs bear three rows of nodes, Formation in Val di Creme locality (Recoaro area, namely the first lateral, the marginal and the external one. Southern Alps, NE Italy). The umbilical portion is not preserved.

Description of the neotype - A fragment of a juvenile Remarks - Despite the bad preservation of the quoted representative of Falsanolcites recubariensis (specimen specimens, their morphological features fit well with the CRM.2a - MGP-PD n° 29840a) preserved as internal diagnosis of Falsanolcites recubariensis. 238 Bollettino della Società Paleontologica Italiana, 57 (3), 2018

Dimensions (mm) - 1914 Protrachyceras Curionii Mojs. - Horn, p. 19-20, Pl. 1, figs 3-4. D H h U W w H/D U/D SGR 1927 Protrachyceras Curionii Mojsisovics - Ogilvie- HH(1) - 41.4* - - 27.1* - - - - Gordon, p. 51, Pl. 6, fig. 11. FO 47 ------1928 Protrachyceras Curionii, Mojsisovics - OgilvieGordon, Pl. 2, fig.10. * estimated non 1935 Protrachyceras curionii v. Mojs. - Schmidt, p. 110, Pl. 6, figs 6-7. Occurrence and Age - At present, in addiction to the non 1962 Trachyceras (Protrachyceras) curionii Mojs. var. Recoaro area (Mojsisovics, 1882; Tornquist, 1898b), ramonensis n.var. Parnes, p. 42-43, Pl. 8, figs 1-4. Falsanolcites recubariensis is surely documented in the 1964 Trachyceras curionii Mojs. mut. rubra Frech. Boda, p. 166. Southern Alps and probably in the Balaton upland (Vörös, v ?1966 Trachyceras (Protrachyceras) curionii Mojs. - Pisa, p. 1998) and in Greece (Epidaurus: fide Jacobshagen, 1967). 630-633, Pl. 70 (I), fig. 1. The species is recorded in the condensed levels of the 1986 Eoprotrachyceras curionii (Mojs.) - Brack & Rieber, Red Ammonitic Limestones of Mt. Clapsavon in Carnia Pl. 5, fig. 4. (Tommasi, 1897, 1900; Pisa, 1966; Mietto & Manfrin, p 1993 Eoprotrachyceras curionii Mojs.) - Brack & Rieber, 1995a); in the Sappada area (Geyer, 1898), and Punta Pl. 8, fig. 7 (not fig. 8). Zonia in Cadore; in the Knollenkalke of the Buchenstein non 1993 Eoprotrachyceras cf. curionii - Vörös, Pl. 6, fig. 5. Beds of Lombardy, Giudicarie and Dolomites (Horn, 1994 Trachyceras curionii Mojsisovics, 1882 - Balini, Pl. 2, 1914; Ogilvie-Gordon, 1927; Brack & Rieber, 1986, 1993; figs 1-2; Pl. 3, fig. 1. Brack et al., 2007); and in the Val Gola Limestone in the v 1995a Eoprotrachyceras curionii (Mojsisovics 1882) - Mietto & Manfrin, Pl. 3, fig. 5; Pl. 4, fig. 6. surroundings of Trento (De Zanche & Mietto, 1989). ? 1995 Eoprotrachyceras curionii (Mojsisovics, 1878) - Goy, Falsanolcites recubariensis is index of the homonymous Pl. 2, fig. 1. Subzone of the Fassanian substage (Mietto & Manfrin, p 1998 Eoprotrachyceras curionii (Mojsisovics) - Vörös, Pl. 1995a). 9, figs ?1, 2, ?4. ? 1998 Eoprotrachyceras cf. curionii (Mojsisovics) - Vörös, Pl. 9, fig. 5. Subfamily Protrachyceratinae Tozer, 1971 v 2004 Eoprotrachyceras curionii Gervasi, p. 18. non 2004 Eoprotrachyceras curionii (Mojs., 1882) - Rieber & Genus Eoprotrachyceras Tozer, 1980a Brack, p. 162, fig. 8. Type species Eoprotrachyceras matutinum Tozer, 1980a ? 2008 Eoprotrachyceras cf. curionii (Mojsisovics, 1882) - Vörös, Pl. 1, fig. 5. ? 2016 Eoprotrachyceras curionii (Mojsisovics) - Perez- Eoprotrachyceras curionii (Mojsisovics, 1882) Valera et al., Fig. 3A. (Pl. 2, fig. 4) Material - One specimen CC(1) (MGP-PD 29230) * 1882 Trachyceras Curionii Mojsisovics, p.116-117, Pl. 14, from the Nodosus Formation of San Ulderico nel Tretto fig. 4. 1887 Trachyceras Curionii Mojs. - Mojsisovics, p. 328. (Schio, Vicenza). 1898b Protrachyceras Curioni Mojs. - Tornquist, p. 662- 663. Description - The small specimen (D about 24 mm) is 1898 Protrachyceras Curioni Mojs. - Toula, Pl. 1, fig. 2. partially included in a matrix made of reddish calcarenite. 1911 Trachyceras Curionii Mojs. mut. nov. rubra It shows bifurcations of the ribs in correspondence of (Protrachyceras) - Frech, p. 22-24, Figs 7a, 9, Pl. 4, the second lateral row of nodes which is a diagnostic fig. 1 character of Eoprotrachyceras curionii. Along the flank 1911 Trachyceras Curionii Mojs. (Protrachyceras) typ. - five rows of nodes, as for the species, are visible while Frech, Pl. 4, fig. 2. those in external position, if well preserved, are flattened 1912 Protrachyceras Curionii Mojs. var. - Salopek, p. 17- in the coiling direction. The ribs are rectilinear but 18, Pl. 5, fig. 1. 1913 Protrachyceras Curionii Mojs. - Simionescu, p. 290- quickly proverse, broad as the inter-rib spaces or less. 291 and 351, fig. 16. The ventral area is very partially exposed but shows v 1914 Protrachyceras cf. Curionii Mojs. - De Toni, p. 137- external nodes in alternating position. The suture line is 138. not preserved.

EXPLANATION OF PLATE 3 Ammonoids of the genus Alloceratites, emigrated from the Central European Muschelkalk basins into the Southern Alps. Fig. 1 - Alloceratites toulonensis (Riedel); specimen G 4.1 (MGP-PD 31961); Val Gola Limestones, Val Gola section (Trento); 1a: left lateral view; 1b: magnification of the suture line. Fig. 2 - Alloceratites tornquisti (Philippi); specimen FF(1) (MGP-PD 29234); Nodosus Formation, San Ulderico (Tretto area, Schio, Vicenza). 2a: right lateral view, 2b: ventral view, 2c: magnification of the last suture line. Fig. 3 - Alloceratites thuringiacus (Riedel); specimen VG 155.1 (MGP-PD 31962); Val Gola Limestones,Val Gola section (Trento); 3a: left lateral view, 3b: magnification of the last suture lines. Scale bar is equal to 1 cm. Bar is 5 mm only for figs. 1b, 2c and 3b. P. Mietto et alii - Middle Triassic ammonoids from the Recoaro and Tretto areas (NE Italy) Pl.239 3 240 Bollettino della Società Paleontologica Italiana, 57 (3), 2018

Dimensions (mm) - Eoprotrachyceras Zone of the Fassanian of the Tretto area (Schio, Vicenza). D H h U W w H/D U/D SGR CC(1) 24 9.9 5.3* 8.8 - - 0.4 0.36 86.8* * estimated Eoprotrachyceras gervasuttii Fantini Sestini, 1994 (Pl. 2, fig. 6a-c) Remarks - The specimen shows features which 1992 Eoprotrachyceras sp.n. A - Jadoul, Gervasutti & Fantini perfectly fit with the diagnosis of the species. Sestini, Pl. 23, fig. 2. * 1994 Eoprotrachyceras gervasuttii Fantini Sestini, p. 257-258, Occurrence and Age - Eoprotrachyceras curionii is Text-fig. 4.a, Pl. 5, figs 1-2. widely documented in all the Mediterranean domain. In ? 1998 Eoprotrachyceras gervasuttii Fantini Sestini - Vörös, Pl. 8, the Southern Alps it is documented in the Esino Limestone fig. 9. (Mietto & Manfrin, 1995a) and in Buchenstein Formation of Lombardy (Mojsisovics 1882; Brack & Rieber 1986), in the Material - One specimen VO.6 (MGP-PD 29237) Zwischenbildungen (Val Gola Limestone) of Val Gola in the from the Nodosus Formation of the Molino Mazzega in surroundings of Trento (Brack & Rieber, 1986; De Zanche & the Orco Valley (Tretto area, Schio, Vicenza). Mietto, 1986, 1989), in the Nodosus Formation of Recoaro (Mojsisovics, 1882; Tornquist, 1898b), in the Knollenkalk of Description - The fragmentary specimen is included in Frommerbach and Puflatsch (Alpe di Siusi: Ogilvie-Gordon a matrix consisting of gray hybrid limestone, weathered 1927, 1928). The species was also found in Val de Pena (De in brown. It shows only part of a flank and relative ventral Toni, 1914) and in Carnia, at Clap di Val in Carnia by the area. The flank is nearly flattened and bears rectilinear, authors and, probably, at Fienili Chiasteons (Pisa, 1966). primary ribs, more or less as broad as the interspaces. Ribs The species is recorded in the Balaton region (Frech, 1911; are slightly projected, starting from the last (second) row ?Vörös, 2008), in the Dinarids (Salopek, 1912) and in in of lateral nodes. The marginal nodes occur in intermediate Romania (Simionescu, 1913). Mojsisovics’s report (1887) position between the latter row of nodes and the external for the Balearic Island cannot be confirmed. The species one. Traces of the first of lateral nodes are inferred, has a precise stratigraphic position between the underlying quite below of the second lateral. The whorl section is chiesense Subzone and the overlying recubariensis Subzone. subrectangular with maximum width in correspondence It marks the homonymous curionii Subzone/Zone at the base of the pointed second row of lateral nodes, due to the bad of the Ladinian Substage (Brack et al., 2005). preservation of the inner part of the flank. The ventrolateral area is broad with marginal and external nodes prevalent in strength. The suture line is not visible. Eoprotrachyceras cf. curionii (Mojsisovics, 1882) (Pl. 2, fig. 5) Dimensions (mm) - D H h U W w H/D U/D SGR Material - One specimen ORC.1 (MGP-PD 31922) * found in situ (bed R 327) in the Nodosus Formation of VO.6 - - - - 21.3 - - - - the Orco Valley (Tretto, Schio). * estimated

Description - The badly preserved and deformed Remarks - The specimen, despite its incompleteness, specimen, about 125 mm in diameter, is included in a shows the general morphological features well comparable matrix made of greenish hybrid limestone, yellowish with those of Eoprotrachyceras gervasuttii, as described in alteration. Only the last portion of the phragmocone by Fantini Sestini (1994). and the first part of the body chamber are inferred. Ribs are not particularly elevated, broad as the interspaces Occurrence and Age - Eoprotrachyceras gervasuttii or less, bearing true spines in umbilical position and in was found in the Esino Limestone of Val Parina in the correspondence of the first lateral row of nodes. At least Bergamasc Prealps (Jadoul et al., 1992; Fantini Sestini, other two/three rows of nodes are inferred along the 1994), and possibly in the Balaton Higland (Vörös, flank. Rib bifurcations, when present, occur only at the 1998). From writers’ unpublished data, this species also firstlateral nodes. The suture line is not visible. occurs in the Clap di Val locality of the Carnian Prealps and in the Punta Zonia section in Central Dolomites. E. Dimensions (mm) - gervasuttii is a typical species from the upper Fassanian (recubariensis Subzone). D H h U W w H/D U/D SGR ORC.1 12.5* ------* estimated Genus Protrachyceras Mojsisovics, 1893 Type species Trachyceras archelaus Laube, 1869 Remarks - The lack of rib bifurcations at the second lateral row of nodes suggests to consider this specimem Protrachyceras margaritosum (Mojsisovics, 1882) in open nomenclature. (Pl. 2, fig. 7a-b)

Occurrence and Age - Despite its finding in situ, v * 1882 Trachyceras margaritosum Mojsisovics, p. 127-128, Pl. this specimen can be generically referred to the 82, fig.1. P. Mietto et alii - Middle Triassic ammonoids from the Recoaro and Tretto areas (NE Italy) 241

1898 Trachyceras margaritosum v. Mojsisovics - Tornquist, * 1882 Gymnites Credneri Mojsisovics, p. 237, pl. 59, figs 1-3. p. 663-664. 1900 Gymnites Credneri Mojs. - Tommasi, p. 40-41. 1993a (Eo-)Protrachyceras margaritosum (Mojs., 1882) - 1903 Gymnites Credneri Mojs. - Frech, p. 35, Pl. 7, fig. 1. Brack & Rieber, p. 484, Text-fig.17h, Pl. 13, figs 6-7. non 1904 Gymnites Credneri Mojsisovics 1882 - Martelli, p. 113- 1994 Protrachyceras cf. margaritosum (Mojsisovics) - Fantini 114. Sestini, Pl. 2, fig. 3. v. 1914 Gymnites Credneri Mojs. - De Toni, p. 162-163. 1995a Protrachyceras margaritosum (Mojsisovics, 1882) - 1915 G. Credneri v. Mojsisovics - Diener, p. 138. Mietto & Manfrin, Pl. 4, figs 2, 4. Material - One specimen, CRM.1 (MGP-PD 29239), Material - One specimen, SRT.dt.2 (MGP 29242), from the Nodosus Formation of Val di Creme (Recoaro from debris of the Nodosus Formation from San Rocco Terme, Vicenza). (Tretto, Schio, Vicenza). Description - The specimen is juvenile and is nearly Description - The somewhat wheatered half specimen, complete. It is made of white calcite and included in is about 6.5 cm in diameter, and it is included in hybrid reddish micritic limestone. The shell coiling is very calcarenites, brownish in color. The shell is slightly involute, and characterised by an high elliptical whorl involuted, with an ovoidal whorl section showing the section with the maximum width in the midflank. The maximum width at the periumbilical area. Despite of the outline of the flank outline is gently rounded. The venter bad preservation of the specimen, the pattern of ribbing is narrow and well rounded. The shell surface is smooth and nodes is consistent with that of the type specimen, without any traces evidence of ornamentation. The suture that is characterised by numerous (at least six for the line is not visible. considered diameter) and equidistant rows of nodes of the same strenght. The ribs are slightly sigmoid and proverse. Dimensions (mm) - The suture line is not visible. D H h U W w H/D U/D SGR Dimensions (mm) - CRM.1 58 31.2 20.6 6.2 9.5 7.1 0.54 0.11 51.4 * estimated D H h U W w H/D U/D SGR * * * * * SRT.dt.2 65 - 19 22.4 - 11.9 - 0.34 - Remarks - It is well known that juveniles specimens * estimated of diverse species of Epigymnites are quite similar in morphology cause because of the lack of ornamentation. Remarks - The specimen at disposal has been The examined specimen under study could be referred, compared with the syntypeus stored at the GBA in particular, to the similar species Epigymnites moelleri (GBA 1882/003/0164). At the same diameter the shell (Mojsisovics, 1882). Nevertheless, the latter species morphology, as the whorl section, the involution degree exhibits flanks more flattened, maintaining the maximum and the ornamentation of the flank ornamentation, are fit width in correspondence of the midflank. well comparable. Occurrence and Age - The specimen at disposal Occurrence and Age - Up to date, P. margaritosum has been found in the same sample with Falsanolcites is documented in the Tretto area (Mojsisovics, 1882; recubariensis. From the literature this species has been Tornquist, 1898), in Lombardy (Val Parina: Fantini reported from Hungary (Frech, 1903) and Southern Alps Sestini, 1994; Mietto & Manfrin, 1995a; Bagolino: (Carnia: Mojsisovics, 1882; Tommasi, 1900; Valdepena: Brack & Rieber, 1993), in the Trento area and Dolomites De Toni, 1914). From the unpublished data this species (Brack & Rieber, 1993; Mietto & Manfrin, 1995a), in occurs also in the La Grea section (Marmolada area) Carnia (Mietto & Manfrin, 1995a). From unpublished associated with Ticinites crassus (Hauer, 1896). At data the quoted species also occurs at Valdepena locality present, Epigymnites credneri is documented from (Cadore area). For historical meaning, as the first the uppermost Anisian (crassus Subzone) to the basal representative of the genus Protrachyceras, Mietto & Longodardian (margaritosum Subzone). Manfrin (1995a) deem this species as marker of the base at the Longobardian (Upper Ladinian: Protrachyceras Zone: margaritosum Subzone). This species prevents Superfamily Megaphyllitoidea Mojsisovics, 1882 the appearance of Protrachyceras archelaus, which is Family Megaphyllitidae Mojsisovics, 1896 historically considered the marker of the Longobardian. Genus Megaphyllites Mojsisovics, 1879 Type species Ammonites jarbas Münster Superfamily Pinacoceratoidea Mojsisovics, 1879 in Wissmann & Münster, 1841 Family Gymnitidae Waagen, 1895 Subfamily Gymnitinae Waagen, 1895 Megaphyllites cf. oenipontanus Mojsisovics, 1882 Genus Epigymnites Diener, 1916 (Pl. 2, fig. 9a-b) Type species Gymnites Ecki Mojsisovics, 1882 Material - One specimen, PZ 3.2 (MGP 29153), from Epigymnites credneri (Mojsisovics, 1882) the bed PZ 3 of the Sturia Limestone from Contrada (Pl. 2, fig. 8) Pozzani (Tretto, Schio, Vicenza). 242 Bollettino della Società Paleontologica Italiana, 57 (3), 2018

Description - The specimen is A badly preserved and immigrants from the Upper Muschelkalk European basins included in gray-brownish limestone. Coiling is very through the Burgundian Gate. involute, the whorl section has been obtained by sawing The genus Alloceratites has been recorded by Xu et al. the specimen. The whorl section is elliptical with the (2003) from the Early Ladinian of the Zhuganpo Fm. in maximun width in the middle flank. The visible portion Gualing (China), but this record is far to be confirmed due of the flank is smooth. The suture line is not visible. to the very bad preservation of the illustrated specimens (pl. II, figs 12- 13). Dimensions (mm) - In the Southern Alps, A. toulonensis and A. thuringiacus have been collected from a stratigraphic section located in D H h U W w H/D U/D SGR Gola Valley (cf. De Zanche & Mietto, 1986, 1989), close PZ 3.2 16.3 10.3 - - 6.1 - 0.6 - - to Trento (layer VG 155 of the Val Gola Limestones), associated with Chieseiceras chiesense (Mojsisovics), Remarks - Pisa (1966) deems Megaphyllites Parasturia sp., and Arpadites sp., thus representing the oenipontanus as a junior synonym of M. jarbas (Münster in uppermost Illyrian (chiesense Subzone). On the other Wissmann & Münster, 1841) Despite that, we agree with the hand, A. tornquisti, on the basis of the matrix of the assumption of Fantini Sestini (1994, 1996) to maintain both unique specimen at disposal, most probably belongs to taxa as distinct species. In fact, respect the latter species, M. the lithostratigraphic interval of the Nodosus Formation of oenipontanus shows a more slender and regularly elliptical the Tretto area in which also Falsanolcites recubariensis whorl section, as observed in the specimen PZ 3.1. is documented, and related, in the writer’s opinion, with the enodis-sublaevigatus zones interval of the Upper Occurrence and Age - M. oenipontanus was found in Muschelkak. the Wettersteinkalk from the mountains north of Innsbruck Urlichs (1978) illustrated a Protrachyceras and near Idrija in Slovenia (Mojsisovics, 1882). The recubariense (Mojsisovics) from the spinosus Zone of species also occurs in Dobrogea (Romania: Simionescu, the Upper Muschelkalk (Württemberg, Germanic Basin). 1913), in the Southern Alps (Val de Pena: De Toni, 1914; The examination of the figure suggests the attribution Forni di Sotto and Clapsavon areas in Carnia (Pisa, 1966) of this ammonoid to Chieseiceras, most probably to C. in the Esino Limestone of Val Parina in Lombardy (Fantini pemphix (Merian). The occurrence of recurrent, well Sestini, 1994, 1996) and perhaps in Balaton Highlands marked radially elongated umbilical swellings support (Frech, 1903). The quoted species is documented in a this change of classification. The range of C. pemphix is large time-span interval from the late Illyrian to most of included in the interval between the chiesense Subzone the Ladinian. and the basal portion of the curionii Subzone. Afterwards, Brack & Rieber (1996), Urlichs & Kurzweil (1997) and Brack et al. (1999) referred the same ammonoid DISCUSSION AND CONCLUSIONS to the genus Nevadites. However, this specimen does not show the row of marginal tubercles which occurs in The genus Alloceratites is represented by four species: the representatives of the genus Nevadites, at the same Alloceratites tornquisti (Philippi), A. toulonensis (Riedel), diameter. Instead, the presence of only two rows of nodes A. thuringiacus (Riedel) and A. schmidi (Zimmermann). in external and periumbilical position, associated with a As expressed before, A. tornquisti is surely documented subrectangular whorl section and a tabulate venter, are in the Triassic of the Recoaro area. Philippi (1901b) and diagnostic characters of the genus Chieseiceras. Riedel (1916) consider this species occurring also in the As referred before, the co-occurrence of Alloceratites Germanic Muschelkalk but Wenger (1957) does not seems toulonensis (Riedel), A. thuringiacus (Riedel) and to confirm this occurrence.A. toulonensis is documented Chieseiceras cf. pemphix (Merian) allows to correlate the in the Upper Muschelkalk of the Provence (Riedel, 1916; Chieseiceras chiesense Subzone (and possibly the basal Urlichs, 1997, 2016) and now also in the Triassic of Eoprotrachyceras curionii Subzone) of the Tethys Realm, southern Trentino region. A. thuringiacus has been found which correspond to the Anisian-Ladinian boundary in the Upper Muschelkalk of the Thuringia (Riedel, 1916) interval, with the Acanthoceratites spinosus Zone of the and also in the “Muschelkalk” of the Sardinia; recently, in Upper Muschelkalk of the Germanic Basin (Fig. 8). This the Triassic of southern Trentino region, too. A. schmidi, correlation is in disagreement with the biostratigraphic is from the Germanic Basin, in particular from the conclusions of Kozur (1974, 1975), Urlichs & Mundlos Lettenkeuper (Erfurt Formation) of Thuringia (see Urlichs, (1980) and Urlichs (1997, 2016). Also Rein & Ockert 2016). Alloceratites shows a ceratitic suture line, which (2000) consider unrealistic the correlation between is characterised by five brachyphyllic entire saddles, well Provence and Germanic basin proposed by Urlichs comparable to that of other representatives of Ceratitinae (1997). from the Triassic of the Germanic and Sephardic basins but Many other authors (e.g., Brack & al., 1999; it differs from the suture line of the representatives of the Bachmann & Kozur, 2004; Klug et al. 2005) deem that subfamily Paraceratitinae, which instead is typical of the ammonoids living in the Muschelkalk basins of the central Tethys Realm. Hagdorn et al. (2015) and Urlichs (2016) Europe were stenohaline. Even if the connections between deem that Alloceratites migrated from the Peritethys the Peritethys basins and the Tethyan Alpine areas were (Provence area of Sephardic Province) towards both the guaranteed throught the Burgundian Gate on the west Germanic Basin and Southern Alps. A. tornquisti, A. and at least by the Silesian Gate on the east during the toulonensis and A. thuringiacus are now known in the Middle Triassic, few ecological barriers (e.g., salinity) Southern Alps and it is reasonable to consider them as might have prevented the ammonoid migrations between P. Mietto et alii - Middle Triassic ammonoids from the Recoaro and Tretto areas (NE Italy) 243

Fig. 8 - Falsanolcites gortanii (Pisa); specimen VO.4 (MGP-PD 29236); Nodosus Formation, Molino Mazzega (Tretto area, Schio, Vicenza); right lateral view. the two domains (cf. Hagdorn et al., 2015). However, A similar scenario occurred also during the two episodes of (maybe eurihaline) ammonoid migration transgressive peak of a 2nd order cycle, Pelsonian in age, have been documented during the Anisian-Ladinian which was equivalent to the “maximum flooding surface” boundary interval (chiesense-curionii subzones) and in of the An3 depositional sequence (sensu De Zanche et the Fassanian (recubariensis Subzone). It is worth noting al., 1993; Gianolla et al., 1994), corresponding to the that only the species belonging to the genus Alloceratites balatonicus Subzone (Mietto & Manfrin, 1995a). In migrated in the Alpine basins. fact, in the Southern Alps, in the old Forte of Monte Rite Besides the passive post-mortem transport, the stratigraphic section (Eastern Dolomites), a specimen migration can be related to the tectonic control on sea- of Noetlingites, which is typical of the Germanic Basin, level change in both domains (cf. Brack et al., 1999, Klug was collected from the bed MRT1 (MRT 1.1, MGP-PD et al., 2005). Nevertheless, in the writers’ opinion this 31885) (Fig. 3). Another specimen of Noetlingites was event may be explained as due to an eustatic fluctuation also found in the Balaton Highlands (Vörös, 2003). of the sea level, well recognised in the Southern Alps Moreover, Noetlingites strombecki was collected from a as the “maximum flooding surface” of the 3rd order coeval bed at Rahnbauerkogel, around Grossreifling in the depositional sequence named La1, which is equivalent Northern Calcareous Alps (Tatzreiter, 2001). Moreover, to the transgressive peak of a 2nd order cycle and of the the specimen ascribed to the genus Hungarites from 3rd order depositional sequence named La2 (De Zanche Dequen in Tibet (Gu et al., 1980: pl. 2, figs. 6, 7) can et al., 1992, 1993, 1994a, 1994b; Gianolla et al., 1994, instead be referred to the genus Noetlingites (Mietto & 1998; Mietto & Manfrin, 1995a). Manfrin, 1995a). It is however noteworthy that in this These global events, in fact, could have enlarged the time interval corresponding to the lower Muschelkalk connections between the Alpine and the central-European the paleoecological and paleoenvironmental conditions basins for short periods and at the same time significantly of Central European basins were normal. This deposition changed the salinity of the surrounding areas close to the evolved, partly at least, to normal condition in the Upper gates, such as the Burgundian Gate that was close to the Muschelkalk, after the salinity crisis of the Middle Provence and Recoaro-Trento areas (see Brack et al., Muschelkalk (Klug et al., 2005). Furthermore, during the 1999: fig. 8). deposition of the Lower Muschelkalk the communications 244 Bollettino della Società Paleontologica Italiana, 57 (3), 2018 between the Germanic Basins and the Tethys were Balini M. (1993). New Genera of Anisian Ammonoids from the more active and wide through the East Carpathian and Prezzo Limestone (Southern Alps). Atti Ticinensi di Scienze Silesian-Moravian gates. In fact, a huge migration of della Terra, 35: 179-193. Alpine-Mediterranean ammonoids such as Balatonites, Balini M. (1994). The Triassic cephalopods of the Curioni collection housed in the “Servizio Geologico Nazionale”: historical and Bulogites, and Acrochordiceras has been documented taxonomical reappraisal. Bollettino del Servizio Geologico in the south-eastern portion of the Germanic basin (cf. d’Italia, 111: 55-76. Urlichs & Mundlos, 1985, Brack et al., 1999; Klug et al. Balini M., Gaetani M. & Nicora A. (1993). Day 2 Tuesday 29 June. 2005), confirming that the migration of the species was In Gaetani M. (ed.), Anisian/Ladinian boundary field workshop. possible in both directions. Southern Alps - Balaton Highlands. 27 June - 4 July 1993. I.U.G.S. Subcomm. Trias. Strat., 43-54. Balini M., Lucas S.G., Jenks J.F. & Spielmann J.A. (2010). Triassic ACKNOWLEDGEMENTS ammonoid biostratigraphy: an overwiew. In Lucas S.G. (ed.), The Triassic Timescale. Geological Society of London, Special The authors are especially indebted with Mariagabriella Pubblication, 334: 221-262. Fornasiero and Letizia Del Favero of the Geological and Barbieri G., De Vecchi Gp., De Zanche V., Di Lallo E., Frizzo Paleontological Museum of the Padova University for their usual P., Mietto P. & Sedea R. (1980). Note illustrative della Carta and appreciable kind assistance. May thanks also to Irene Zorn Geologica dell’area di Recoaro alla scala 1:20.000. Memorie of the Geologische Bundesanstalt of Wien for her usual kind di Scienze Geologiche, 34: 23-52. availability. Barbieri G., De Vecchi Gp., De Zanche V., Mietto P. & Sedea R. Marco Balini, Roberto Gatto, Hans Hagdorn, Claude Monnet, (1982). Stratigrafia e petrologia del magmatismo triassico Nereo Preto and Attila Vörös are thanked for their suggestions, that nell’area di Recoaro. In Castellarin A. & Vai G.B. (eds), improved the manuscript significantly. Guida alla Geologia del Sudalpino centro-occidentale. 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BIOSTRATIGRAPHIC APPENDIX The biostratigraphic meaning of the gredleri Zone

When Krystyn (1983) described a succession with Later on, despite this inadequate definition, thegredleri conodonts from the Epidaurus condensed section, he ammonoid biozone was generally accepted (Brack & suggested a new ammonoid biostratigraphic scale for the Rieber, 1993a, 1993b, 1994; Fantini Sestini, 1994, 1996; Tethys. One of the new contributions of this scale was the Nicora & Brack, 1995; Brack et al., 1995; Vörös, 1998; introduction of the gredleri Zone in the lower portion of Brack et al., 1999; Bachmann & Kozur, 2004; Budai & the Longobardian, that was previously comprised in the Vörös, 2006; Hounslow et al., 2007; Lucas, 2010; Balini archelaus Zone. The gredleri Zone was evidently defined et al., 2010). Mietto & Manfrin (1995a) split the gredleri by the occurrence of Protrachyceras cf. gredleri which, Zone into a margaritosum Subzone (lower part) and a however, was not illustrated. gredleri Subzone (upper part). Moreover, they replaced

Fig. 9 - About the stratigraphic position of Protrachyceras gredleri (Mojsisovics) and the gredleri Zone sensu Krystyn (1983). a) The stratigraphic arrangement of the Alpe di Siusi/Seiser Alm area is exemplified by the Frombach section (Alpe di Siusi/Seiser Alm area); with star the realistic position of the lectotype of P. gredleri (from Mietto et al., 2012 modified). Scale column is in meters. b) Relative diverse position of the gredleri Zone/Subzone and the lectotype of P. gredleri in the biostratigraphic scheme of Mietto & Manfrin (1995a). 250 Bollettino della Società Paleontologica Italiana, 57 (3), 2018 the archelaus Zone of Krystyn (1983) with the new Examples of Protrachyceras of similar size and preservation longobardicum and neumayri subzones. have been found in the same area at the base of the Rio The original designation of Trachyceras gredleri by Frommer/Frommer Bach section 1 (Fig. 9a) at the same Mojsisovics (1882) comprises one gerontic fragmentary locality (Mietto & Manfrin 1995b, Mietto et al. 2008). specimen from the Alpe di Siusi/Seiser Alm in Dolomites The fauna occurring in these volcanoarenites belongs to and some specimens from the red ammonitic limestones the neumayri Subzone, i.e., the upper archelaus Zone of cropping out from the M. Clapsavon in Carnia. For its Krystyn (1983). In the Alpe di Siusi/Seiser Alm area these morphology, the former specimen is clearly identifiable volcanoarenites cropping out above several hundred meters as Protrachyceras, but the specimen illustrated in of Upper Ladinian volcanites (see Mietto et al. 2012). Mojsisovics (1882) at pl. 34, fig. 7 from M. Clapsavon It must be noted that the gredleri Zone of Krystyn has shows a morphological framework that perfectly fits always been placed in the lower part of the archelaus Zone with a specimen from Clap di Val illustrated by Mietto & of the previous authors, rather than in its upper part to Manfrin, 1995a (pl 5, fig. 8), which also shows a ceratitic which the specimen from Alpe di Siusi/Seiser Alm belongs. suture line, and assigned the species to Eoprotrachyceras. Hence, the designation of the lectotype of upper archelaus The designation of the specimen illustrated by age (Krystyn et al., 2004) as marker of a pre-archelaus Mojsisovics (1882: pl. 17) as lectotype of Trachyceras biozone becomes paradoxical and makes the gredleri gredleri by Krystyn et al. (2004: p. 48) introduces a Zone inconsistent and useless (Fig. 9b). Consequently, also substantial problem. This specimen, stored in the GBA the gredleri Subzone as defined by Mietto and Manfrin (GBA1882/003/0151/02) and examined by the writers, (1995a) loses its significance, so that the corresponding was found at Alpe di Siusi/Seiser Alm in Longobardian stratigraphic interval is now considered as equivalent to volcanoarenites of the Wengen Group (Viel, 1979). the upper part of the margaritosum Subzone.