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September 2003

Pentatomidae () of : a checklist with description of a new ochlerine

Nolberto Arismendi Escuela Agricola Panamericana, Zamorano, Apartado Postal 93, , Honduras, C.A.

Donald B. Thomas U.S. Department of Agriculture, Agricultural Research Service, Kika de la Garza Subtropical Agricultural Research Center

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Arismendi, Nolberto and Thomas, Donald B., " (Heteroptera) of Honduras: a checklist with description of a new ochlerine genus" (2003). Insecta Mundi. 31. https://digitalcommons.unl.edu/insectamundi/31

This Article is brought to you for free and open access by the Center for Systematic Entomology, Gainesville, Florida at DigitalCommons@University of Nebraska - Lincoln. It has been accepted for inclusion in Insecta Mundi by an authorized administrator of DigitalCommons@University of Nebraska - Lincoln. INSECTA MUNDI, Vol. 17, No. 3-4, September-December, 2003 219

Pentatomidae (Heteroptera) of Honduras: a checklist with description of a new ochlerine genus

Nolberto Arismendi Escuela Agricola Panamericana, Zamorano Apartado Postal 93, Tegucigalpa, Honduras, C.A.

Donald B. Thomas U.S. Department of Agriculture, Agricultural Research Service, Kika de la Garza Subtropical Agricultural Research Center, 2413 E. Hwy 83, Weslaco, TX 78596

Abstract. Through collecting, surveys of museum collections, and search of the literature, we are able to list 181 species of Pentatomidae as occurring within the boundaries of the Republic of Honduras. Most of these, 129, around 70%, are widespread in the American tropics. Twenty-nine species are new country records, reported for Honduras for the first time. Four species of pentatomids are endemic to Honduras including a new genus and species of ochlerine (Discocephalinae) herein described. Although a few species extend from South America into Honduras (the Gondwanan element), and a few from North America extend into Honduras (the Nearctic element), the most important faunal element is one which is native to nuclear Central America.

Resumen. Con colectas, revisiones de las colecciones de museos, y revisiones de literaturas, se puede enumerar 181 especies de Pentatomidae distribuidas dentro de los límites de la república de Honduras. La mayoría de éstos, 129, alrededor del 70%, se extienden en las zonas tropicales americanas. Veinte y nueve especies son nuevos registros en el país, reportados en Honduras por primera vez. Cuatro especies de pentatómidos son endémicas de Honduras incluyendo un nuevo género y especie de Ochlerini (Discocephalinae) adjunto descrito. Aunque algunas especies se extienden de Sudamérica a Honduras (el elemento de Gondwana), y algunas de Norteamérica se extienden dentro Honduras (el elemento neárctico), el elemento más importante de la fauna es aquel que es nativo del núcleo de la región centroamericana.

Introduction contribution to the knowledge of Honduran Het- eroptera, and to an eventual national biotic inventory. Faunal surveys of Honduras date from the Brit- The Republic of Honduras is the largest country ish explorations that culminated in the Biologia in Central America with an area of 112,090 square Centrali-Americana, which included two volumes on km; about the size of Virginia. Geographically, it is the Heteroptera (Distant 1880-1893). bordered by the countries of , , Although the family Pentatomidae is one of the and , and bounded by ocean on two sides. largest in the order, it was underrepresented in that Although located in the tropics, the climate is strong- survey with only five species listed for Honduras. ly modified by elevation. On the constricted Pacific According to Selander and Vaurie (1962), Honduras side, a flat, swampy strip of lowland bounds the Gulf was the least collected country of those represented in of Fonseca. The Caribbean lowlands are much more the “Biologia.” Through a literature search, based extensive and consist of two distinctive areas. The primarily on genus level revisionary studies, we found northwestern coast has the most intensive agricul- published Honduran Records for a total of 63 species tural land use in Honduras. The original banana of pentatomids, while another 85 species were found to republic, the area has diversified into pineapple, palm have distributional ranges that included countries oil, citrus and mango production. The northeastern both to the north and south, without specific Hondu- coast, known as Mosquitia, is the least inhabited and ran localities being cited. For example, in the revision least developed region, consisting of rain forest, trop- of Alitocoris, Sailer (1950) cited collection records for ical savannah and mangrove swamp. During heavy A. schraderi Sailer for Guatemala and . rainfall much of the area is covered by shallow water, Consequently, we believe that the Honduran insect and rain falls year round. Because of the limited fauna may be undercollected and its diversity largely accessibility, we were unable to collect in Mosquitia. undocumented. The present work is offered as a 220 Volume 17, No. 3-4, September-December, 2003, INSECTA MUNDI

Figure 1. Map with the Departments of Honduras.

The rugged, mountainous interior comprises about Materials and Methods 80% of the national terrain and is climatically, and therefore, ecologically diverse. The easterly mountain To extend our knowledge of the diversity and ranges have dense, broad-leaf tropical forest. Those in distribution of pentatomids from this region we sur- central and western Honduras have open woodland, veyed the holdings of two insect collections in Hondu- mainly pine forest interspersed with oak and scrub. ras. The small collection at the Universidad Nacional Cloud forest or remnants of cloud forest can be found Autonoma de Honduras in Tegucigalpa and the larger on the highest ridges and peaks. Although situated at collection at the Escuela Agricola Panamericana (EAP), tropical latitudes, there is a pronounced dry season also called Zamorano. Both were studied and their from November to April, with freezing temperatures specimens determined to species. Additionally, the and frosts occurring on the high peaks, the highest junior author surveyed the entomological collections being Cerro de Las Minas at 2,850 m. For the most at Texas A&M University, Colorado State University, part the valleys of central Honduras have been cleared Brigham Young University, the University of Ne- for pasture and subsistence agriculture or have re- braska, the University of California Riverside, and turned to secondary growth. A strong natural conser- the Carnegie Museum of Natural History, for Hondu- vation effort is underway in Honduras to preserve the ran specimens. Efforts were then directed at acquir- larger plots of native forest and their indigenous ing material by active collecting in Honduras with the faunas. We were able to obtain permission to collect in assistance and collaboration of Dr. Ronald D. Cave, these protected areas, many of which are now national Professor of Entomology at Zamorano. The two prima- parks. ry methods of collecting were by hand netting the seen on the vegetation by day, and mercury vapor light collecting by night. These collections were carried out mainly in the vicinity of Zamorano (De- INSECTA MUNDI, Vol. 17, No. 3-4, September-December, 2003 221 partment of Francisco Morazán), the Centro Univer- sitario de la región Litoral Atlántico University Re- search station near (Atlántida), at Parque Nacional Pico Bonito (Atlántida), at Parque Nacional Cusuco (Cortés), Parque Nacional Pico Pijol (Yoro), and Parque Nacional La Muralla (Olancho), under permit from the Honduran government obtained through Zamorano. The Departments of Honduras are depicted in Fig. 1. The species of Pentatomidae are enumerated in a checklist with departmental records for each, and specific localities within departments for new country records. For the biogeographic analysis, the recorded distribution is cited and scored as follows: Endemic (E) for those species known only from Honduras, Central American (C) for those species known only from nuclear Central America (tropical to the uplands of Nicaragua), northern (N) for those species occurring north of and into nuclear Central America, southern (S) for those species occurring from Hondu- ras southward into isthmian Central America (Pan- ama and Costa Rica), and widespread (W) for those species occurring in the regions both north and south of Honduras. In preparing the checklist a traditional was followed. In recent years there have been propos- als to split larger genera, such as Fieber and Amyot and Serville, into smaller genera (Ahmad 1996 and Lariviere 1994, respective- Figure 2. Hondocoris cavei, new genus, new species; dorsal ly). But these proposals have not been generally habitus. accepted (e.g. Frey-da-Silva and Grazia 2001), or are undergoing further review (e.g. Ahmad and McPher- Description: Brachypterous; membranes of wings son 1998). absent, hind wings absent; ocelli absent. Eyes contig- uous with pronotum (Fig. 2). Humeral angles weakly Results and Discussion and anterior pronotal angles not produced, pronotal margins not explanate. Scutellum short and broad, As a result of our collecting and work in collec- lateral margins straight, without frenal constriction tions, we found 29 species of pentatomids not previ- or discontinuity; apex of scutellum attaining mid- ously recorded in Honduras. Added to the literature abdomen (5th tergite). Meso- and metasternum thinly records, this results in a total list of 181 species of the carinate on midline. Scent gland orifice round, with- family Pentatomidae occurring within the geographic out spout, but located on caudal side of short raised boundaries of the country. For comparison, there are auricle which extends laterally about one-tenth dis- 85 species listed for Nicaragua (Maes 1994), 194 tance to metasternal margin. Paired trichobothria species for (Froeschner 1999), 317 species in located on each abdominal sternite behind and about Mexico (Thomas 2000) and 224 species in the United half-way between each spiracle and lateral margin. States (Froeschner 1988). During this survey we Antennae five-segmented. Labium arising slightly discovered a genus and species new to science in the before anterior limit of eyes; intercalary segment tribe Ochlerini collected by Professor Cave, which is absent; segment projecting caudad of bucculae described herein, and named in his honor. and attaining middle of prosternum. Pro- and me- sotibiae prismatic in cross-section; metatibiae sulcate Hondocoris Thomas, new genus on planar surface. Superior surface of third metatar- (Figs. 2, 3) 222 Volume 17, No. 3-4, September-December, 2003, INSECTA MUNDI

Thorax. Anterolateral margins of pronotum car- inate, irregular, corrugate; anterior angle not project- ing into a tooth. Pronotal length at midline, 2.3 mm; width across humeri 6.6 mm. Apex of scutellum subsinuately notched. Length of scutellum 4.7, width, 4.3 mm. Evaporatorium not extensive, covering only about one quarter of metapleural surface. Abdomen. Apices of connexiva not produced. First gonocoxites strongly convex, declivitous posteri- Figure 3. Female terminalia of H. cavei: Gx1 = first gonocoxite,

Pt8 = eighth paratergite, Pt9 = ninth paratergite. orly; mesial margins thickened, labiate, lateral angle lobate. Posterior margin of eighth paratergite sinu- sal segment with shallow, ovoid groove. Eighth ately projected at middle, spiracles present. Ninth paratergites of female with spiracle present. paratergites foliate, contiguous posteriorly, surface rugose, apices not attaining posterior margin of eighth Diagnosis: In Rolston’s (1992a) revision of Ochleri- paratergite. Fused second gonocoxites strongly con- ni, Hondocoris will key to Miopygium Breddin, but vex, barely exposed (Fig. 3). differs from that genus in having 5-segmented anten- nae (4 in Miopygium), ocelli absent (present in Miopy- Holotype: female; labeled: Honduras: Yoro. Parque gium), the labium arising before the anterior limit of Nac. Pico Pijol, 14 June 1998, rcol. R. Cave. (Depos- the eyes (behind in Miopygium), by the longer ros- ited with U.S. National Museum of Natural History, trum and the broader scutellum. Washington D.C.)

Type-species: Hondocoris cavei, Thomas, new spe- Etymology: the generic name is a combination of the cies first syllable of Honduras and the greek word for bug. The specific epithet honors its discoverer, Professor Hondocoris cavei, Thomas, new species Ronald D. Cave, formerly of Zamorano.

Description: Body roundly-oval in outline, widest Remarks: The type locality is a remnant of cloud across abdomen; dorso-ventrally compressed. Length: forest at 1,450 m elevation. The specimen was collect- 13.8, width 9.2 mm. Color of body black, except for ed by Professor Cave under a fallen tree trunk. The thick, , carina adorning middle of each corium, time of collection was about one year after the trees just ental and parallel to embolar suture. Dorsal had been cut down. A deliberate search of the same surface of head, pronotum, scutellum and corium site in August 2002 failed to recover further speci- strongly rugose, the rugae oriented transversely. mens, although aradids were common. Notably, the Head. Juga occluding at terminus of tylus, apices tree trunks at this date were in advanced decomposi- divergent, forming v-shaped notch at apex of head. tion. Perhaps this bug species only colonizes newly Length: 2.2 mm, width across eyes 2.9 mm. Anteoc- fallen tree trunks, or persists in the trees for such ular processes robust, obtuse, not projecting but time after they have fallen as to reach an advanced producing strong excavation in margin of head just state of decay. anterior to eye. Anteocular width 1.9 mm. Each antennifer with short angular tooth at apex. Anten- New Distributional Records for Honduras nae five segmented (fifth segment missing in holo- type, but apex of 4th segment is socketed). First For published distributional records we relied antennal segment long, exceeding apex of head; sec- primarily on genus level revisionary studies. Such ond segment short, about half length of first; third studies are available for the majority of genera in the segment longest, about 3x the fourth; fourth segment New World. The notable exception is the genus Edes- about equal in length to first. Antennal segments sa, which happens to be the largest genus in the black except distal two-thirds of fourth segment pale. family. Fernandes et al (2001) estimate that there are Bucculae elevated, evanescent posteriorly, projecting 260 nominal species in this genus. The only available into strong tooth anteriorly. Rostrum long, apex key is that by Stål (1872) which predates the Biologia attaining middle of 6th abdominal sternite in repose; Centrali-Americana wherein the largest number are segments II, III and IV subequal in length, I shortest, described. The 21 species of this genus appearing in about two-thirds length of II. our checklist are derived from the ranges given in INSECTA MUNDI, Vol. 17, No. 3-4, September-December, 2003 223

Kirkaldy’s (1909) catalogue, which in turn are derived assemblage that dominates the highlands of western mainly from the older records of Distant and Stål. We North America (Graham 1973). It also forms the mid- feel this literature survey probably underestimates section of the dispersal corridor for lowland tropical the number of species that actually occur in elements extending northward from the rain forests of Honduras. But in the absence of a revision we are South America (Rich and Rich 1983, Simpson and unable to associate the majority of specimens with Neff 1985). specific names. Of the 181 species known or likely to occur in Of the 29 new distributional records, the majori- Honduras, the bulk of the species, 129, about 71%, are ty, 20 species, are southeastward range extensions of widespread in distribution in the sense that they species previously known to occur in Mexico or Gua- range over an area well to the north and to the south temala. These include the following species: Adevopli- of Honduras. According to the generally accepted tus caseae (Thomas), Arocera melanopyga (Stål), theory, the Neotropical fauna is primarily of Gond- totonaca Thomas, Banasa calva Say, Disde- wanan origin (Savage 1982, Halffter 1987), that is, ria decorata Distant, chiapus Rolston, genera that evolved in South America and dispersed Euschistus comptus Walker, into Central America. With regard to pentatomids, Rolston, Euschistus strenuus Stål, guate- the Nearctic influence is clearly minimal, arguably malensis Rider, Mormidea laevigata Distant, Neotibi- represented only by the genera Brochymena, Cos- lis chiapensis (Thomas and Brailovsky), Rhyssoceph- mopepla, and perhaps . Presumably, the ala rufonotata Stål, Rio obscuratus Ruckes, Rio large Neotropical genera well represented in South testaceus Ruckes, Odmalea schaefferi (Barber), America, such as Edessa, Euschistus, , and Ochlerus handlirschi Breddin, dichrous Banasa, evolved in South America and invaded Cen- (Herrich-Schaeffer), Oplomus mutabilis (Stål) and tral America. It has never been clear to us why the Pantochlora vivida Stål. Five new records for Hondu- Central American fauna has to have evolved else- ras are species previously reported in the Central where. It is established from both tectonic and fossil American countries south of Honduras: Euschistus evidence that nuclear Central America has existed as achatadus Rolston, Banasa dolobrata (Thomas), a land mass since Paleozoic times, long before the biconicus (Thomas), Glaucioides engel- modern flora and fauna evolved. Moreover, during mani Thomas and Antiteuchus panamensis (Ruck- most of that history, nuclear Central America was not es). Four new records are range extensions of South connected to South America. It is true that only four American species not previously known to occur in species on our checklist are endemic to Honduras, or Central America. These include the following species: at least, known only to occur in Honduras. These are Chloropepla lenti Grazia, Rhyncholepta grandical- Priapismis pini Rolston, Alitocoris brunneus Sailer, losa Bergroth, Odmalea concolor (Walker), and Di- Edessa phoenicopus Dallas and Hondocoris cavei nocoris lineatus (Dallas). Thomas. But if one excludes the faunal components which are either endemic or widespread, and there- Biogeographic Analysis of the Honduran fore of low information content, the largest component Pentatomidae of the fauna is the nuclear Central American element. That is, 26 species have a distribution from southern Tectonically, Honduras rests on the Chortis block Mexico to Nicaragua, as opposed to only seven species (Malfait and Dinkelman 1972, Donnelly et al. 1990), with an isthmian origin, that is, species indigenous to a geologically stable area that has been land positive Panama and Costa Rica and extending into Hondu- since at least the end of the Paleozoic (Delevoryas and ras. Only eight species on the list occur from South Srivastava 1981). The connection to North America America up to but stopping in Honduras. It seems to dates from around the end of the Mesozoic, a time us that if the Central American fauna, which is before modern faunas evolved. Conversely, the isth- clearly Neotropical, originated by invasion from South mian connection to South America is geologically America, then a greater proportion of the fauna recent, established not until the Pliocene about 3.5 should be shared with the areas to the south, whereas mya. Biogeographically, Honduras is an integral part the distributional evidence indicates the opposite. An of nuclear Central America, roughly equivalent to the instructive example is the distribution of the edessine Chortis block, an area which extends from northern genus Brachystethus Laporte. The ten known species Nicaragua to the state of Chiapas, Mexico (Ryan 1963, are dispersed over all of the Neotropical region from Raven and Axlerod 1975). Floristically, Honduras lies Mexico to Argentina. But, a cladistic analysis by at the southernmost extent of the pine-oak forest Barcellos and Grazia (2003) indicates that the Central 224 Volume 17, No. 3-4, September-December, 2003, INSECTA MUNDI

American group of species is basal. The distributional Oplomus mundus Stål pattern at the genus level from our Honduran study Honduran Records: Atlántida, El Paraíso, also suggests that nuclear Central America was an Yoro. April to November. important center in the evolution of the Neotropical Distribution: (W) USA to Panamá (Thomas pentatomid fauna. 1992). Oplomus mutabilis (Stål) Honduran Records: El Paraíso: Danlí, 9 Checklist of Pentatomidae of Honduras October 1977, J. Mankins coll. Distribution: (C) México and Guatemala Subfamily (Thomas 1992). Oplomus pulcher Dallas (Dallas) Honduran Records: Santa Bárbara: Lago Honduran Records: Atlántida, , Yojoa, 2 August 1977, J. Mankins coll.; Siquat. Cortés. March and May. 12 September 1977, J. Mankins coll.; Taulabe, Distribution: (W) USA to Argentina (Thomas 7 October 1977, J. Mankins coll. 1992). Distribution: (W) México to Panamá (Thomas spinidens (Fabricius) 1992). Honduran Records: Comayagua, El Paraíso, confluens (Herrich-Schaeffer) Francisco Morazán. May to November. Honduran Records: Francisco Morazán: Valle Distribution: (W) Circumtropical, USA to de Angeles, August 1993, S. Fernandez coll.; Costa Rica, (Thomas 1992). also Comayagua. July and August. lineolatus (Herrich-Shaeffer). Distribution: (W) USA to Costa Rica (Thomas Honduran Records: Ortega (1997). El Paraíso, 1992). Olancho, Francisco Morazán, Lempira, Yoro. aenescens (Stål) January to August. Honduran Records: none. Distribution: (W) USA to Venezuela (Thomas Distribution: (W) México to Argentina 1992). (Thomas 1992). Coryzorhaphis cruciata Stål Podisus affinis Distant 1880 Honduran Records: Thomas (1992). Atlántida, Honduran Records: Olancho: Montaña El Francisco Morazán. June. Boquerón, 12 June 2003, J. Torres coll.; also, Distribution: (W) México to Venezuela Copán, El Paraíso and Yoro. May to November. (Thomas 1992). Distribution: (C) México, Guatemala (Thomas Coryzorhaphis egeri Thomas 1992), Nicaragua (Maes 1994). Honduran Records: none. Podisus congrex (Stål) Distribution: (W) México to Costa Rica Honduran Records: Olancho: Parque La (Thomas 1992). Muralla, 14 km N. La Unión, 7 May 2003, J. (Linnaeus) Torres and N. Arismendi colls. Also, Yoro: Pico Honduran Records: Thomas (1992); Francisco Pijól, 17 June 2004, D. Thomas and D. Morazán, El Paraíso, Lempira, Yoro. March to Robacker colls. November. Distribution: (W) México to (Thomas Distribution: (W) USA to Brazil (Thomas 1992). 1992). Podisus falcatus (Distant) Heteroscelis lepida (Stål) Honduran Records: El Paraíso, Olancho. June Honduran Records: Francisco Morazán: La and November. Lima, 18 June 1978, J. Mankins coll. Distribution: (W) Guatemala, Costa Rica Distribution: (W) USA to Ecuador (Thomas (Thomas 1992). 1992). Podisus nigriventris Distant Oplomus dichrous (Herrich-Schaeffer) Honduran Records: Olancho, May. Honduran Records: Comayagua, 21 July 1978, Distribution: (W) México to (Thomas J. Mankins coll. 1992). Distribution: (N) USA (Arizona) and México (Thomas 1992). INSECTA MUNDI, Vol. 17, No. 3-4, September-December, 2003 225

Podisus sagitta (Fabricius) Antiteuchus tripterus (Fabricius) Honduran Records: Thomas (1992), Francisco Honduran Records: Atlántida, Comayagua, El Morazán, Atlántida, Comayagua, El Paraíso Paraíso, Francisco Morazán, Yoro. February to and Yoro. January to November. October. Distribution: (W) USA to Venezuela, West Distribution: (W) México to (Ruckes Indies (Thomas 1992). 1964). Podisus trucidatus Thomas Dinocoris lineatus Dallas Honduran Records: Francisco Morazán: Honduran Records: Comayagua: La Soledad, Parque Nacional La Tigra. January. May. Distribution: (S) Costa Rica and Panama Distribution: (S) Brazil (Becker and Grazia (Thomas 1992). 1985). anchorago (Fabricius) Dinocoris rufitarsus Ruckes Honduran Records: Thomas (1992), Distant Honduran Records: Ruckes (1958) Paratypes (1880); also, Atlántida, Francisco Morazán. from Honduras: Brus Lagoon 25 April 1947, January to December. C.W. Cook coll. Also, El Paraíso and Francisco Distribution: (W) USA to Panamá (Thomas Morazán. January to July. 1992). Distribution: (S) Honduras to Brazil (Becker Supputius typicus (Distant). and Grazia 1985). Honduran Records: Atlántida, March. Discocephalessa humilis (Herrich-Schaeffer) Distribution: (W) México to Ecuador (Thomas Honduran Records: Copán, Atlántida, 1992). Comayagua, , Santa Bárbara. Tylospilus cloelia (Stål) January to December. Honduran Records: none. Distribution: (W) Guatemala to Colombia Distribution: (W) México to Argentina (Ruckes 1966a). (Thomas 1992). Discocephalessa notulata (Stål) Tylospilus acutissimus (Stål) Honduran Records: none. Honduran Records: Francisco Morazán: Distribution: (W) México to Costa Rica (Ruckes Tegucigalpa, Nov. 1981, M.A. Valle coll. 1966a). Distribution: (W) USA to Colombia (Thomas Lineostethus clypeatus (Stål) 1992). Honduran Records: El Paraíso, Atlántida. Tyrannocoris jole (Stål) May. Honduran Records: Thomas (1992); also, Distribution: (W) México, , Panamá Francisco Morazán. June. (Ruckes 1966a). Distribution: (N) México to Honduras, Cuba, Pelidnocoris stalii Haglund Hispaniola (Thomas 1992). Honduran Records: none. Distribution: (W) México to Panamá (Ruckes 1966b, Froeschner 1999). Subfamily Discocephalinae Phoeacia erubescens (Distant). Honduran Records: Cortés, Olancho, Yoro. Tribe Discocephalini April to September. Distribution: (W) Guatemala to Panamá Antiteuchus innocens Engleman (Distant 1880). Honduran Records: Paratypes from Priapismus pini Rolston Tegucigalpa; also, Atlántida, May to July. Honduran Records: Cortés: Cerro Cusuco Distribution: (W) México to Costa Rica (Type locality), 26 January 1991, R. Cameron. (Engleman and Rolston 1983). Distribution: (E) Honduras (Rolston 1992b). Antiteuchus panamensis (Ruckes) Honduran Records: El Paraíso, April. Tribe Ochlerini Distribution: (S) Panamá (Ruckes 1964). Antiteuchus piceus (Palisot de Beauvois) Alitocoris brunneus Sailer Honduran Records: None Honduran Records: Holotype specimen Distribution: (W) México to Argentina, West intercepted on orchids from Honduras (Sailer Indies (Ruckes 1964). 226 Volume 17, No. 3-4, September-December, 2003, INSECTA MUNDI

1950); also, Atlántida, Francisco Morazán. Distribution: (W) México to Panamá, West May. Indies (Rolston 1983). Distribution: (E) Honduras (Sailer 1950). Acrosternum scutellata (Distant) Alitocoris maculosus Sailer Honduran Records: Atlántida, Lempira, Honduran Records: one paratype intercepted Olancho. March to August. on Bananas from Honduras (Sailer 1950). Distribution: (W) México to Panamá (Rolston Distribution: (C) Guatemala to Honduras 1983). (Sailer 1950). Adevoplitus caseae (Thomas) Alitocoris schraderi Sailer Honduran Records: Francisco Morazán, Honduran Records: none. a Dios. April to July. Distribution: (W) Guatemala and Costa Rica Distribution: (C) México and Guatemala (Sailer 1950). (Brailovsky and Barrera 1982). Hondocoris cavei Thomas Arocera aequinoxialis (Westwood). Honduran Records: Yoro: Pico Pijól, 14 June Honduran Records: None. 1998, R. Cave coll. Distribution: (W) Guatemala to (Rider Distribution: (E) Honduras. 1992). Macropygium reticulare (Fabricius) Arocera melanopyga (Stål) Honduran Records: Francisco Morazán, Honduran Records: Atlántida: Rio Zecate, 18 Gracias a Dios, Lempira, Olancho, El Paraíso May 2001, D. Thomas, D. Robacker, and R. Yoro, Santa Bárbara. April to July. Cave, colls. also, Yoro. March to November. Distribution: (W) México to Colombia (Distant Distribution: (C) México, Belize, Guatemala 1880). (Rider 1992). Ochlerus handlirschi Breddin Arocera placens (Walker) Honduran Records: Atlántida, Comayagua, Honduran Records: Santa Bárbara (Rider March, May and November. 1992); also, Colón, Cortés, Comayagua and Distribution: (C) México (Breddin 1909). Yoro. April to June. Ochlerus cinctus (Spinola) Distribution: (W) México to Argentina, West Honduran Records: El Paraíso, August. Indies (Rider 1992). Distribution: (W) México to Brazil (Distant albopunctatus (De Geer) 1880). Honduran Records: Copán, Francisco Morazán, Stalius tartareus (Stål) El Paraíso. March to October. Honduran Records: Comayagua, 2.8 km N. Distribution: (W) USA to Argentina, West Los Planes, 25 May 2002, R. Cave. (on Indies (Brailovsky 1981). Rhamnus sphaerosperma); also, Olancho and Arvelius porrectispinus Breddin Santa Barbara. May to June. Honduran Records: San Antonio (Brailovsky Distribution: (W) México to Colombia (Rolston 1981); also, El Paraíso, Olancho, Yoro. June to 1992a). September. Distribution: (W) México to Brazil, West Indies (Brailovsky 1981). Subfamily Banasa calva (Say) Honduran Records: Francisco Morazán: Cerro Tribe Uyuca, 5 May 2002, D.Thomas and J. Burne colls.; Cortés, Parq. Nac. Cusuco, 21 August Acrosternum marginata (Palisot de Beavois) 1998, R. Cave Coll. Also, La Paz. April to Honduran Records: Yoro, Atlántida, Choluteca, August. Comayagua, Cortés, El Paraíso, Francisco Distribution: (N) USA to Guatemala (Thomas Morazán, Lempira, Olancho. January to and Yonke 1988). December. Banasa centralis Sailer Distribution: (W) USA to Ecuador, West Indies Honduran Records: Atlántida, Cortés, Olancho. (Rolston 1983). May and June. Acrosternum montivaga (Distant) Distribution: (W) México to Colombia (Thomas Honduran Records: El Paraíso, Francisco and Yonke 1988). Morazán. October. INSECTA MUNDI, Vol. 17, No. 3-4, September-December, 2003 227

Banasa dolobrata Thomas Distribution: (W) Guatemala to Panamá Honduran Records: Cortés. May. (Thomas and Yonke 1988). Distribution: (S) Costa Rica to Ecuador Banasa totonaca Thomas (Thomas and Yonke 1988). Honduran Records: Atlántida, Cortés, El Banasa excavata Thomas Paraíso, Francisco Morazán, La Paz, Olancho, Honduran Records: Francisco Morazán, Santa Bárbara and Yoro. February to Olancho, Santa Bárbara, Yoro. April to September. September. Distribution: (C) Mexico (Thomas and Yonke Distribution: (W) Guatemala to Panamá 1988). (Thomas and Yonke 1988). Banasa zeteki Sailer Banasa lacertosa Thomas Honduran Records: Thomas and Yonke (1988); Honduran Records: Olancho: Montaña also, Atlántida, El Paraíso, Cortés, Olancho, Botaderos, NW San Esteban, 7 August 2002, Yoro, Santa Bárbara and Francisco Morazán. Wells et al. coll. April to September. Distribution: (S) Panamá (Thomas and Yonke Distribution: (W) México to Colombia, West 1988). Indies (Thomas and Yonke 1988). Banasa lenticularis Uhler Berecynthus hastator (Fabricius) Honduran Records: Thomas and Yonke (1988); Honduran Records: Olancho, Catacamas; also, Atlántida, El Paraíso and Francisco Valle, Comayagua, Francisco Morazán. April to Morazán. February to August. Nobember. Distribution: (W) USA to Colombia, West Distribution: (W) México to Brazil (Grazia Indies (Thomas and Yonke 1988). 1982). Banasa minor Sailer Capivaccius bambusii Brailovsky and Rolston. Honduran Records: none. Honduran Records: none. Distribution: (W) Guatemala to Panamá Distribution: (W) México to Panamá (Thomas and Yonke 1988). (Froeschner 1999). Banasa panamensis Sailer Chlorocoris biconicus Thomas Honduran Records: Olancho: Moñtana del Honduran Records: Cortés, Olancho, Yoro. Malacate, 18 June 2001, B.Ratcliffe, M.Jameson May to August. and R.Cave colls. Distribution: (S) Nicaragua to Panamá Distribution: (W) Belize to Colombia (Thomas (Thomas 1985). and Yonke 1988). Chlorocoris distinctus Signoret Banasa salvini Distant Honduran Records: Thomas (1985); also, Honduran Records: Thomas and Yonke (1988). Atlántida, Comayagua, El Paraíso, Francisco Also, El Paraíso, Cortés, Olancho, Santa Morazán, Olancho, Yoro. March to December. Bárbara and Yoro. May to July. Distribution: (W) USA to Ecuador (Thomas Distribution: (W) México to Venezuela 1985). (Thomas and Yonke 1988). Chlorocoris rufispinus Dallas Banasa santarosana Thomas Honduran Records: Thomas (1985). Cortés, El Honduran Records: none. Paraíso, Francisco Morazán, Yoro and Distribution: (W) México to Costa Rica Olancho. March to September. (Thomas 2000). Distribution: (W) México to Panamá (Thomas Banasa sleeperi Thomas 1985). Honduran Records: none. Chloropepla lenti Grazia Distribution: (W) Guatemala, Costa Rica Honduran Records: Atlántida. August. (Thomas and Yonke 1988). Distribution: (S) Venezuela (Grazia 1968). Banasa stalii Distant decorata (Hahn) Honduran Records: Cortés. May. Honduran Records: Comayagua: Taulabe, 2 Distribution: (W) México to Panamá (Thomas April 1980, J. Mankins coll. and Yonke 1988). Distribution: (W) USA to Venezuela (Bonatto Banasa stigmosa Distant and Grazia 1985). Honduran Records: none. Cosmopepla coeruleata Montandon Honduran Records: none. 228 Volume 17, No. 3-4, September-December, 2003, INSECTA MUNDI

Distribution: (W) USA to Colombia (McDonald Euschistus emoorei Rolston 1986). Honduran Records: El Paraíso, Francisco Cyptocephala antiguensis (Westwood) Morazán, Olancho and Valle. January to Honduran Records: Comayagua, Francisco December. Morazán, Olancho. April to September. Distribution: (W) México to Colombia (Rolston Distribution: (W) USA to Peru (Rolston 1986). 1974). Disderia decorata Distant Euschistus leonensis Rolston Honduran Records: El Paraíso, Francisco Honduran Records: Rolston (1974); also, Valle, Morazán. May to September. Atlántida. January to July. Distribution: (C) México, Belize, Guatemala Distribution: (S) Honduras to Costa Rica (Brailovsky 1986). (Rolston 1974). Euschistus achatadus Rolston Euschistus nicaraguensis Rolston Honduran Records: Francisco Morazán: Cerro Honduran Records: Rolston (1974); Valle, Uyuca, 17 May 2000, A.B. Smith and F. Choluteca, Francisco Morazán. May to Ocampo colls. September. Distribution: (C) Nicaragua (Rolston 1974). Distribution: (S) Nicaragua to Panamá Euschistus bifibulus (Palisot de Beauvois) (Rolston 1974). Honduran Records: Valle, Cortes, Atlantida (Palisot de Beauvois) and Francisco Morazan. February to November. Honduran Records: none. Distribution: (W) México to Colombia, West Distribution: (N) USA to Nicaragua (Maes Indies (Rolston 1974). 1994). Euschistus biformis Stål Euschistus quadrator Rolston Honduran Records: Choluteca, El Paraíso, Honduran Records: Atántida, Comayagua, Francisco Morazán, Lempira, Yoro and Francisco Morazán. January to December. Olancho. January to December. Distribution: (N) USA and México (Rolston Distribution: (W) USA to Panamá (Rolston 1974). 1974). Euschistus strenuus Stål. Euschistus chiapus Rolston Honduran Records: Choluteca, Comayagua. Honduran Records: El Paraíso. April and April and June. October. Distribution: (C) México (Rolston 1974). Distribution: (C) México and Guatemala Euschistus sulcacitus Rolston (Rolston 1974). Honduran Records: none Euschistus comptus Walker Distribution: (C) México to Nicaragua (Maes Honduran Records: Francisco Morazán. 1994). February. Euschistus zafadus Rolston Distribution: (C) México and Guatemala Honduran Records: El Paraíso, October. (Rolston 1974). Distribution: (W) Belize to Costa Rica (Rolston Euschistus corcovacitus Rolston 1974). Honduran Records: Rolston (1971) paratype Glaucioides engelmani Thomas 1980 from Puerto Castillo; also, Atlántida, Cortés, El Honduran Records: Atlántida: Parque Nacional Paraíso, Francisco Morazán, Santa Bárbara. Pico Bonito, March. January to December. Distribution: (S) Panamá to Brazil (Rolston et Distribution: (C) México to Honduras (Rolston al. 1980). 1971). Hypatropis rolstoni Fernandes and Grazia Euschistus crenator (Fabricius) Honduran Records: one paratype intercepted Honduran Records: Isla Ruatan (Distant in bananas from Honduras (Fernandes and 1880); Punta Castilla (Rolston 1974); also, Grazia 1996). Atlántida, Cortés, El Paraíso, Francisco Distribution: (S) Honduras to Venezuela Morazán, Olancho, Santa Bárbara, Comayagua (Fernandes and Grazia 1996). and Choluteca. January to December. imbuta (Walker) Distribution: (W) México to Peru, West Indies Honduran Records: Atlántida, Francisco (Rolston 1974). Morazán, Olancho, Yoro. May to October. INSECTA MUNDI, Vol. 17, No. 3-4, September-December, 2003 229

Distribution: (W) México to Panamá (Distant Mormidea lunara Rolston 1880). Honduran Records: Comayagua (Rolston flavicollis (Drury) 1978a); also, Choluteca, El Paraíso, Francisco Honduran Records: none. Morazán, Santa Bárbara. January to November. Distribution: (W) USA, México, Costa Rica, Distribution: (W) México to Panama (Rolston West Indies (Arnold 1995). 1978a). Loxa virescens Amyot and Serville Mormidea notulata (Herrich-Schaeffer) Honduran Records: Atlántida, El Paraíso, Honduran Records: Cortés, El Paraíso, Francisco Morazán, Olancho, Yoro and Santa Francisco Morazán, Olancho, Santa Bárbara Bárbara. January to September. and Yoro. June to October. Distribution: (W) México to Uruguay (Eger Distribution: (W) México to Peru (Rolston 1978). 1978a). Loxa viridis (Palisot de Beauvois) Mormidea pama Rolston Honduran Records: Atlántida, Choluteca, Honduran Records, Atlántida: P.N. Pico Comayagua, Colón, Francisco Morazán, Bonito, Rio Zacate, 5 May 2000, B. Ratcliffe, A. Olancho, Santa Bárbara and Yoro. January to Smith, F.Ocampo and R.Cave colls.; also, September. Santa Bárbara, Francisco Morazán and Yoro. Distribution: (W) USA to Argentina (Eger March to November. 1978). Distribution: (W) USA to Colombia, West Mayrinia variegata (Distant) Indies (Rolston 1978a). Honduran Records: El Paraíso: 8.3 Km SE Mormidea pictiventris Stål Capire, 24 May 2002, R. Cave coll.; also, Honduran Records: Atlántida, Comayagua, El Olancho and Francisco Morazán. May to July. Paraíso, Francisco Morazán, Olancho, Yoro. Distribution: (W) México to Brazil (Grazia January to October. 1972, Thomas 2000). Distribution: (W) México to Venezuela (Rolston Mormidea angustata Stål 1978a). Honduran Records: Olancho. May. Mormidea ypsilon (Linnaeus) Distribution: (W) México to Brazil, West Indies Honduran Records: Distant (1880); Cortés, (Rolston 1978a). Santa Bárbara. Atlántida, Yoro. February to Mormidea collaris Dallas December. Honduran Records: Olancho: Montaña del Distribution: (W) México to Uruguay (Rolston Malacate, 18 June 2001, B.Ratcliffe, M.Jameson 1978a). and R. Cave colls.; also, Atlantida, El Paraiso, histrionica Hahn 1834 Francisco Morazán, Yoro. February to Honduran Records: Atlántida, El Paraíso, November. Francisco Morazán, Ocotepeque, Yoro. January Distribution: (W) México to Colombia (Rolston to December. 1978a). Distribution: (W) México to Costa Rica (Distant Mormidea cubrosa (Dallas) 1880). Honduran Records: DEPT El Macuelizo, 13 Murgantia varicolor (Westwood) Jan 95 (no collr); also, Francisco Morazán. Honduran Records: none. January. Distribution: (W) México to Brazil, West Indies Distribution: (W) USA to Colombia, West (Distant 1880). Indies (Rolston 1978a). Neotibilis chiapensis (Thomas and Brailovsky) Mormidea guatemalensis Rider Honduran Records: Cortés: Cerro Cusuco, 20 Honduran Records: Yoro: Parque Pico Pijol, 3 May 2001, D.Robacker and D.Thomas colls. June 2003, N. Arismendi coll. Distribution: (C) México (Thomas and Distribution: (C) México and Guatemala (Rider Brailovsky 1993). and Rolston 1989). viridula (Linnaeus) Mormidea laevigata Distant Honduran Records: El Paraíso, Francisco Honduran Records: Atlantida. July. Morazán. February to December. Distribution: (C) Mexico (Rolston 1978a). Distribution: (W) Cosmopolitan (Kirkaldy 1909). 230 Volume 17, No. 3-4, September-December, 2003, INSECTA MUNDI

Oebalus insularis Stål Distribution: (W) México to Guyana (Rolston Honduran Records: Sailer (1944); also, 1984). Atlántida, Choluteca, Comayagua, El Paraíso, Pharypia nitidiventris (Stål). Francisco Morazán, Olancho. May to October. Honduran Records: none. Distribution: (W) USA to Panamá, West Indies Distribution: (W) México to Panamá (Sailer 1944). (Froeschner 1999). pugnax (Fabricius) Pharypia pulchella (Drury) Honduran Records: Distant (1880); also, Honduran Records: Distant 1880 (type locality Comayagua, El Paraíso, Francisco Morazán. is Bay of Honduras); also, La Paz. July. January to October. Distribution: (W) México to Brazil (Distant Distribution: (W) USA to Brazil (Sailer 1944). 1880). Padaeus teapensis (Distant) guildinii Westwood Honduran Records: El Paraíso, Francisco Honduran Records: Francisco Morazán, El Morazán. April to November. Paraíso. May to July. Distribution: (W) México to Costa Rica Distribution: (W) USA, to Paraguay, West (Kirkaldy 1909). Indies (Kirkaldy 1909). Padaeus trivittatus Stål albopunctulatus (Palisot de Beauvois) Honduran Records: none. Honduran Records: None. Distribution: (W) México to Costa Rica (Maes Distribution: (W) USA? to Brazil, West Indies 1994). (Froeschner 1988, Arnold 1995). Padaeus verrucifer Stål (Palisot de Beauvois) Honduran Records: Santa Bárbara, Montaña Honduran Records: Atlántida, Choluteca, El La Cumbre, 8 June 2003, N. Arismendi and J. Paraíso, Francisco Morazán, Lempira, Torres colls. Santa Bárbara and Olancho. January to Distribution: (C) México and Guatemala November. (Kirkaldy 1909). Distribution: (W) USA to Venezuela (Distant Padaeus viduus (Vollenhoven) 1880, Grazia 1984). Honduran Records: El Paraíso, Francisco (Fabricius) Morazán, Ocotepeque. January to November. Honduran Records: Atlántida, Yoro. February Distribution: (W) México to Venezuela (Grazia to November. 1984, Distant 1880). Distribution: (W) México to Panamá (Distant Pallantia diffusa Walker 1880). Honduran Records: Comayagua: 5 August Ramosiana insignis (Blanchard) 1979, J. Mankins coll. Honduran Records: El Paraíso, Francisco Distribution: (W) México to Brazil (Grazia Morazán. January to September. 1989). Distribution: (W) México to Panamá (Kormilev Pallantia maculosa Grazia 1951). Honduran Records: none. Rhyncholepta grandicallosa Bergroth Distribution: (W) México to Costa Rica (Grazia Honduran Records: Olancho: Montaña del 1989). Malacate, 18 June 2001, B.Ratcliffe, M.Jameson Paratibilis confusa Ruckes and R.Cave colls.; also, Atlántida, Yoro. June to Honduran Records: none. September. Distribution: (W) México to Peru (Grazia and Distribution: (S) Panamá to Brazil and Barcellos 1991). (Becker and Grazia 1971). santarosensis Rolston Rhyssocephala infuscata Rider. Honduran Records: Olancho: Montaña El Honduran Records: Atlantida (Rider 1991); Bosqueron, 12 June 2003, J. Torres coll.; also, also, Olancho, Yoro and El Paraíso. January to El Paraiso. June. September. Distribution: (W) México to Costa Rica Distribution: (W) México to Costa Rica (Rider (Thomas 2000). 1991). Pellaea stictica (Dallas) Rhyssocephala macdonaldi Rider Honduran Records: Atlántida, Francisco Honduran Records: none. Morazán. May to July. INSECTA MUNDI, Vol. 17, No. 3-4, September-December, 2003 231

Distribution: (W) Guatemala to Ecuador (Fabricius) (Froeschner 1999). Honduran Records: (Rider and Chapin 1992), Rhyssocephala rufonotata (Stål) also Comayagua, El Paraíso, Francisco Honduran Records: Atlántida, Francisco Morazán, Yoro. January to October. Morazán, Olancho, Santa Bárbara, Yoro. May Distribution: (W) USA to Panamá, West Indies to September. (Rider and Chapin 1992). Distribution: (C) México and Guatemala (Rider Vulsirea violacea (Fabricius) 1991). Honduran Records: Francisco Morazán. May Rhyssocephala splendens (Blanchard) to July. Honduran Records: El Paraíso, Francisco Distribution: (W) México to Venezuela (Distant Morazán. May to July. 1880, Grazia 1984). Distribution: (W) México to Panamá, Jamaica (Rider 1991). Tribe Procleticini Rio obscuratus Ruckes Honduran Records: Atlántida, P.N. Pico Odmalea concolor (Walker) Bonito, Est. Curla, 2 June 2002, R. Cave.; also, Honduran Records: Atlántida. May and June. Francisco Morazán. May to July. Distribution: (S) Panamá to Brazil (Rolston Distribution: (C) México (Grazia and Fortes 1978b). 1995). Odmalea schaefferi (Barber) Rio testaceus Ruckes Honduran Records: Atlántida. May to August. Honduran Records: Francisco Morazán: Distribution: (N) USA and México (Rolston Zamorano, 2 October 1993, R. Turnbow Coll. 1978b). Distribution: (C) Guatemala and El Salvador (Grazia and Fortes 1995). Tribe Rio variegatus Ruckes Honduran Records: none Brochymena haedula Stal Distribution: (W) Guatemala to Costa Rica Honduran Records: Francisco Morazán: Cerro (Grazia and Fortes 1995). Uyuca, Siguat. 13-10-77, J. Mankins coll.; also, Roferta marginalis (Herrich-Schaefer) Comayagua. April to October. Honduran Records: None. Distribution: (W) México to Costa Rica Distribution: (W) Belize and Panamá; Cuba, (Lariviere 1994). Trinidad and Brazil (Barber and Bruner 1946). Brochymena inbio Lariviere Sibaria englemani Rolston Honduran Records: Cedros, Tegucigalpa. Honduran Records: El Paraíso: Yuscarán, 26 July. October 1977, J.V. Mankins coll.; also, Distribution: (S) Honduras and Costa Rica Atlántida. February to September. (Lariviere 1994). Distribution: (W) México to Colombia (Rolston 1975). Tribe Mecidiini Taurocerus edessoides Spinola Honduran Records: Yoro, Atlántida. January Mecidea major Sailer to September. Honduran Records: none. Distribution: (W) México to Brazil (Distant Distribution: (N) USA to Nicaragua (Maes 1880). 1994). Thyanta maculata (Fabricius) Mecidea minor Ruckes Honduran Records: Choluteca, Cortés, Honduran Records: none. Comayagua (Rider and Chapin (1992); also, Distribution: (W) USA to Costa Rica (Arnold Francisco Morazán and Santa Bárbara. March 1995). to May. Distribution: (W) USA to Panama (Rider and Subfamily Edessinae Chapin 1992). Edessa abdominalis Erichson Honduran Records: none. 232 Volume 17, No. 3-4, September-December, 2003, INSECTA MUNDI

Distribution: (W) México to Guyana (Kirkaldy Edessa olivacea Stål 1909). Honduran Records: none. Edessa affinis Dallas Distribution: (W) México to Panama Honduran Records: Kirkaldy (1909). (Froeschner 1999). Distribution: (W) Belize and Guatemala to Edessa patricia Stål Brazil (Kirkaldy 1909). Honduran Records: none. Say Distribution: (W) México to Panamá Honduran Records: Santa Bárbara: Lago (Froeschner 1999). Yojoa, 5 July 1977, J. Mankins coll.; also, El Edessa phoenicopus Dallas Paraíso, Atlántida, Francisco Morazán and Honduran Records: type locality in Honduras Olancho. January to October. (Dallas 1851). Distribution: (W) USA to Panamá (Distant Distribution: (E) Honduras (Distant 1880). 1880). Edessa privata (Walker) Edessa cornuta Burmeister 1835 Honduran Records: none. Honduran Records: Kirkaldy (1909). Distribution: (C) México to Nicaragua (Maes Distribution: (W) México to Brazil (Distant 1994). 1880). Edessa pudibunda Stål Edessa gentilitia Distant 1890 Honduran Records: none. Honduran Records: none. Distribution: (W) México, Guatemala, Costa Distribution: (W) México to Costa Rica Rica (Distant 1880). (Kirkaldy 1909). Edessa reticulata Dallas Edessa helix Erichson Honduran Records: Francisco Morazán. Honduran Records: none. December. Distribution: (W) México to Guyana (Kirkaldy Distribution: (C) México (Kirkaldy 1909). 1909, Froeschner 1999). Edessa rufomarginata De Geer Edessa jugata Westwood Honduras: (Maes 1994); also, Francisco Morazán, Honduran Records: none. Atlántida, Olancho, El Paraíso and Yoro. Distribution: (W) México to Bolivia (Kirkaldy January to December. 1909). Distribution: (W) México to Argentina (Distant Edessa junix Stål 1880). Honduran Records: none. Edessa trifurca Dallas Distribution: (W) México to Panamá Honduran Records: type locality, Isla Ruatan (Froeschner 1999). (Dallas 1851). Edessa jurgiosa Stål Distribution: (C) México to Honduras (Kirkaldy Honduran Records: none. 1909). Distribution: (W) México to Panamá Edessa vinula Stål (Froeschner 1999). Honduran Records: none. Edessa lineigera Stål Distribution: (W) México to Panama (Kirkaldy Honduran Records: Atlántida: P.N. Pico 1909). Bonito, Rio Zecate. 5 May 2000, B. Ratcliffe, A. Olbia caprina Stål Smith, F.Ocampo and R.Cave colls.; also, Honduran Records: none. Cortés. May to September. Distribution: (W) México to Brazil (Distant Distribution: (W) México to Panamá (Fernandes 1880). et al. 2001). Pantochlora vivida Stål Edessa nigricornis Stål Honduran Records: Cortés, Olancho, Yoro, Honduran Records: none. May to August. Distribution: (W) México to Panamá Distribution: (C) México and Guatemala (Froeschner 1999). (Kirkaldy 1909). Edessa nigrispina Dallas Peromatus notatus Burmeister Honduran Records: type locality in Honduras Honduran Records: Atlántida: Parque Nacional (Dallas 1851). Pico Bonito, Estación Curla, 17 February 2002. Distribution: (W) México to Panama (Kirkaldy Distribution: (C) México (Kirkaldy 1909). 1909). INSECTA MUNDI, Vol. 17, No. 3-4, September-December, 2003 233

Peromatus robustus Haglund manuscript. The habitus illustration of Hondocoris Honduran Records: Olancho: El Boqueron, 8 cavei was prepared by Ms. Angela Fox of the Nebraska June 2002. State Museum. Distribution: (W) México, Guatemala, Costa Rica, Panamá (Distant 1880). Literature Cited Peromatus truncatus Distant Honduran Records: none Ahmad, I. 1996. A revision of the Distribution: (W) México to Panama (Kirkaldy tribe Pentatomini Leach (: Pentatomi- 1909). dae: Pentatominae) from Indo-Pakistan subcon- Brachystethus improvisus Breddin tintent with special reference to their . Honduran Records: Cortés: Santa Cruz de Proceedings of the Pakistani Congress of Zoology Yojoa, 4 Nov 1994, R. Cordero; Francisco 16: 41-86. Morazán: Zamorano, 8 June 1984, R. Parada; Ahmad, I. and J.E. McPherson. 1998. Additional Cerro Uyuca, 24 April 1984, R. Caballero. information on male and female genitalia of Distribution: (W) Guatemala to Brazil Parabrochymena Lariviere and Brochymena (Barcellos and Grazia 2003). Amyot and Serville (Hemiptera: Pentatomidae). Brachystethus rubromaculatus Dallas Annals of the Entomological Society of America Honduran Records: Barcellos and Grazia 91: 800-807. (2003); also, Atlántida and Yoro. January to Arnold, K.A. 1995. Fragmenta Heteroptera Neotro- November. pica III, fauna costaricana I (Insecta: Heteroptera). Distribution: (W) México to Panamá (Distant Faunistische Abhandlungen, Staatliche Muse- 1880). um fur Tierkunde, Dresden 20: 9-13. Barber, H.G. and S.C. Bruner. 1946. Records and descriptions of miscellaneous Cuban Hemiptera. Subfamily Tessaratominae Bulletin of the Brooklyn Entomological Society, 41: 52-62. Piezosternum subulatum (Thunberg) Barcellos, A. and J. Grazia. 2003. Cladistic anal- Honduran Records: none ysis and biogeography of Brachystethus Laporte Distribution: (W) México to Brazil (Distant (Heteroptera, Pentatomidae, Edessinae). Zootaxa 1880). 256: 1-14. Becker, M. and J. Grazia-Vieira. 1971. Sobre o genero Rhyncholepta Bergroth, 1911, com a de- Acknowledgments scricâo de uma nova epecie. (Hemiptera, Pentato- midae, Pentatominae). Revista Brasileira de Bio- The authors are indebted to Joel Meras of Parque logia 31: 389-399. Nacional La Muralla, Celio Gutierrez of Parque Na- Becker, M. and J. Grazia. 1985. Revisâo do genero cional Pico Pijól, Fernando Martinez of Parque Nacio- Dinocoris Burmeister, 1835 (Heteroptera, Pen- nal Pico Bonito and Alberto Ponce of Parque Nacional tatomidae, Discocephalinae). Revista Brasileira Cusuco, for permission to collect insects. The authors de Zoologia 3: 65-158. also thank the following individuals and their institu- Bonatto, S.L. and J. Grazia, 1985. Redescricâo de tions for access to Honduran material for study: Brett Cosmopepla decorata (Hahn, 1834) com o estudo C. Ratcliffe, University of Nebraska; Shawn Clark, da genitalia de ambos sexos (Heteroptera, Pen- Brigham Young University; Edward C. Riley, Texas tatomidae). Anais da Sociedad Entomologica do AandM University; John E. Rawlins, Carnegie Muse- Brasil 14: 289-301. um of Natural History; Boris Kondratieff, Colorado Brailovsky, H. 1981. Revisión del genero Arvelius State University; and Douglas Yanega, University of Spinola (Hemiptera - Heteroptera -Pentatomidae California, Riverside. But we are especially grateful to - Pentatomini). Anales del Instituto de Biologia. those generous individuals that donated material for Universidad Nacional Autónoma de México 51: study: Robert Turnbow, Henry Howden, Bruce Gill, 239-298. Jim Wappes and Jeff Huether. Julio Torres, Roberto Brailovsky, H. 1986. Hemiptera-Heteroptera de Cordero, Jeff Burne, Dave Robacker and Ron Cave Mexico XXXVII: tres nuevas especies y nuevos provided assistance and companionship on collecting registros de la familia Pentatomidae. Anales del trips. Ron Cave and Joseph E. Eger reviewed the 234 Volume 17, No. 3-4, September-December, 2003, INSECTA MUNDI

Instituto de Biologia. Universidad Nacional Au- eroptera, or True Bugs, of Canada and the conti- tónoma de México. 57(2): 281-298. nental United States. E.J. Brill, Leiden. 958 pp. Brailovsky, H. and E. Barrera. 1982. Hemiptera- Froeschner, R.C. 1999. True Bugs (Heteroptera) of Heteroptera de Mexico XXII. Nuevos registros de Panama: a synoptic catalog as a contribution to la tribu Pentatomini y descripción de una nueva the study of Panamanian biodiversity. American especie del genero Pseudevoplitus Ruckes (Pen- Entomological Institute, Gainesville FL. 393 pp. tatomidae). Anales del Instituto de Biologia. Uni- Graham, A. 1973. History of the arborescent tem- versidad Nacional Autónoma de México 52: 231- perate element in the northern Latin American 246. biota. Pp. 301-314, in, A. Graham (ed.). Vegeta- Breddin, G. 1909. Beitrage zur systematik der pen- tion and Vegetational History of Northern Latin tatomiden sudamerikas. Sitzungs Berichten Ge- America. Elsevier, Amsterdam. sellschaft Naturforshender Freunde zu Berlin. Grazia, J. 1968. Sobre o género Chloropepla Stål, 1909: 615-631. 1867, com descricâo de uma nova especie (Hemi- Dallas, W.S. 1851. List of Specimens of Hemipterous ptera, Pentatomidae, Pentatominae). Revista Insects in the Collection of the British Museum, Brasileira de Biologia 28: 193-206. Pt. 1, Taylor and Francis, London. 368 pp. Grazia, J. 1972. Sobre o género Mayrinia, Horvath Delevoryas, T. and S.C. Srivastava. 1981. Juras- 1925 (Heteroptera, Pentatomidae, Pentatomini). sic plants from the Department of Francisco Revista Peruana de Entomologia 15: 117-124. Morazan, Central Honduras. Review of. Palaeo- Grazia, J. 1982. Revisâo do género Berecynthus Stål, botany and Palynology. 34: 345-357. 1862 (Heteroptera, Pentatomidae, Pentatomini). Distant, W.L. 1880-1893. Insecta. Rhynchota, Hemi- Revista Brasileira de Entomologia 26: 173-182. ptera - Heteroptera. In, F.D Godman, and O. Grazia, J. 1984. Pentatominae da Venezuela (Het- Salvin, (eds). Biologia Centrali-Americana. Lon- eroptera, Pentatomidae). Anais da Sociedad Ento- don. Pt. 1: 462 pp. mologica do Brasil 13: 71-81. Donnelly, T.W., G.S. Horne, R.C. Finch and E. Grazia, J. 1989. Revisâo do género Pallantia Stål, Lopez-Ramos. 1990. Northern Central-Ameri- 1862 (Heteroptera: Pentatomidae). Revista ca: the Maya and Chortis blocks. Chapt. 3, In: The Brasileira de Entomologia 24: 15-27. Geology of North America, vol. H, the Caribbean Grazia, J. and A. Barcellos. 1991. Sobre o género region. G. Dengo and J.E.Case [eds.] Geological Paratibilis Ruckes (Heteroptera: Pentatomini). Society of America, Boulder CO. Anais da Sociedad Entomologica do Brasil 20: 209- Eger, J.E. 1978. Revision of the genus Loxa (Hemi- 216. ptera: Pentatomidae). Journal of the New York Grazia, J. and N. Fortes, 1995. Revisâo do género Entomological Society 86: 224-259. Rio Kirkaldy, 1909 (Heteroptera, Pentatomidae). Engleman, H.D. and L.H. Rolston. 1983. Eight Revista Brasileira de Entomologia 39: 409-430. new species of Antiteuchus Dallas (Hemiptera: Halffter, G. 1987. Biogeography of the montane Pentatomidae). Journal of the Kansas Entomo- entomofauna of Mexico and Central America. logical Society 56: 175-189. Annual Review of Entomology 32: 95-114. Fernandes, J.A. and J. Grazia, 1996. Revisâo do Kirkaldy, G.W. 1909. Catalogue of the Hemiptera genero Hypatropis Bergroth, 1891 (Heteroptera, (Heteroptera) with biological and anatomical ref- Pentatomidae). Revista Brasileira de Entomolo- erences, lists of food plants and parasites, etc. Vol. gia 40: 341-352. 1. . Berlin. 392 pp. Fernandes, J.A., P.H. van Doesburg and C. Kormilev, N.A. 1951. Sobre los generos Vulsirea Greve. 2001. The Edessa collaris - group of Spinola (1837), Ramosiana Kormilev (1950) y Edessa Fabricius, 1803 (Heteroptera: Pentatomi- Adoxoplatys Breddin (1903), con la descripcion de dae: Edessinae). Zoologische Mededelingen, Leiden tres especies nuevas (Hemipt. Pentat.). Revista 75: 239-250. de la Sociedad Entomologica de Argentina 15: 83- Frey-da-Silva, A. and J. Grazia. 2001. Novas 95. espécies de Acrosternum subgênero Lariviere, M.C. 1994. Parabrochymena Lariviere (Heteroptera, Pentatomidae, Pentatomini). Iher- (Hemiptera: Pentatomidae): systematics, natu- ingia (Séries Zoologia), Porto Alegre 90: 107-126. ral history, chorological affinities, and evolution- Froeschner, R.C. 1988. Family Pentatomidae Leach, ary relationships, with a biogeographic analysis 1815: the stink bugs. Pp. 544-597, in, T. Henry of Parabrochymena and Brochymena Amyot and and R. Froeschner (eds.). Catalog of the Het- INSECTA MUNDI, Vol. 17, No. 3-4, September-December, 2003 235

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