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• JOINT Cent re for NAT UR E Ecology & Hydrology CONSERVATION COMMITTEE N AT U RA L EN V IR O N M EN T R ES EA R C H C O U N C IL • • • • - • • • • 4 0 9 * ••• 0 0 0 • • Pr o visio n al atlas o f t h e • acu le ate Hy m e n o p te r a • enr,::. • fl e a 84 40041. 00 • a • • • • o f Br ita in an d Ir elan d • IP 0 * : 1480 444: Par t 6 • • • • e-• : • • 50 on •• • 0 0 0 UV. 0 0 0 0 11 e a 0 Ro b in E !w a r d s (Ed ito r ) • ,.,• I • • B e e N , W a S p S a n d An ts Re co rd in g Sc %I. 4. f a • • • DO* _ a 9 6 le • • • • • • es G i l V i A/ Br o ad (Ed it o r ) • it • • • • • 0 Bio lo g ic a l, Re co r d s ( ;en tr e • • • • • se n a• o 9- 11 on t eget ili el e t ' • • • tan e s e . : In seel tt l eso, eo • • • mimes • • • • 444•• * * •• of - • • • • • • •• • •" • . * • • • • • -• • • : 50 ::.::r 4::.814.1- - . .. -1 p r. 3._.r. •.• :..• • ::. •• • . • 41.4- • • Cent re fo r Ecology and Hydrology N at ural Environrnenc Research Council Ci tp ‘ itig hi I tIit h in in d fl 21- mg) K lm.k St lit e [HEI N f -ifi dfR U I l i me Ce nnic S ir l r nli imv mud lv d n ilo m 15 1 111 ic El m L k s U mm re e l ,.., ull un c i t.ThicArch in d ie li mel Ired w, ater t_in v iro m neidi d 'Je n ( l I En Te mui 1 Nock di ul t in i in t i l s ld ci tl i tiLli thst sk il ls in :1 \. \ it11 lJ !i El u. i i! env irt r um :y i n g th e <ysitritni: ( )Hi Ctidi filt Hui .1, :it finci ri.tdini li •it:, i d hi di/ die _Ind ON: u nt itid it i a :d i, :Hien id tti l in d i( iat of Im mitn ic mi l uc mm lidtm .11 J I M I N) I l e N.Il e \A t 0 /N ; H S_ So N I I H I N, I i I O LI : 11 I r i 11 1111 :2TI L I I itIt o a kfidlik n ei dd i‘ mind t r i ( a i l e k e mi c e mi N TNN I n h u m[ d i n e d i n I i i . N ,ffi j Ve / d i k e : d e I lie ottst il l. Cid Ii rcncmnei i zEn aez ikil ule 1c m milk h e i l l p. :1 vid e ritni_N: gc rle rid in ic uesl puh li cdi to Jilt ! f I "tire. Ntiii k in der in k iiim au riii i Jil L1 it C. 1 . 11 DLitt:tic ti cc h .dt ir k li to L ga .11 Re< idt cl i, Ctintry t M i tt t turdit i Nil di / \\I (Hid \Ad , c m:N y.1 1yd in p h, 4 ii rl d li mm Li. in Hi, N an d 11,--di ,e,dit O L I N Hird to ‘ vi off-. ci miii d in • ry titti t li ng co m il nin ir t thirit iughn ut Lind tadim ck cr unc Li-141.MT, l e I 1111( I V I SL I FIN l a w L' I l e l r i i m n e n r .i l in hmmmnne - mci it ytiit , d itdr iNi ii it in PiN t it tutiiiit - mitt irt.i Wi n i ii . mo l e tl ittu J4J.irmtm, , I K 'C l e S . P k I e, ‘ W W W R 1 .1m milu li d HO' !SRC. I, it tindii. Fund ed th e I Ntuti fy Ci..insc n ttnt in f li dn illifit mi n d n key ct im po ne ni i ii the Nato mi l No vi t ick FIN- Pat \\ id spiiii. :In d Suciety nt ie me i. ttni cid ioimicmll v ilid thit> s P U I 11/^Th e id C c I r e (t it. I e t k \\,-(),x 1 [fil m in I I w il in gt Te l i d tic-I— 7 - 2 O ) : %. A li r LIP. Provisional atlas of the A culea te Hy men op tera of B rita in a nd Irela nd Pa rt 6 Ro b in Edw ar ds (Ed ito r ) Bees, Wasps a nd A n ts Record ing Soc iety Gav in Br o ad (Ed ito r) Biolog ica l Reco rds Centre Centre for Ecology and Hydrology Natural Environment Research Council Citation information: Edwar ds, R. and Br oad, G., eds. 2006. Provisional atlas of the aculeate Hymenoptera of Britain and Ireland. Part 6. Huntingdon: Biological Records Centre. CON TEN TS Page Acknowledgements Introduction Systematic list of species mapped k Distribution maps and species profiles Chrysididae 10 Formicidae 24 Pompilidae 34 Crabronidae 40 Apidae 60 Bibl iography 129 List of plant names 135 Cumulative index to species in Provisional atlas parts to I 6 139 I L. A CK N O WLE D GE M E N TS Again, this sixth part of the Bees,Wasps and Ants Provisional Atlas is the result of much hard wor k by many people.The species for this Atlas were carefully selected by Target Species Coordinator, Stuart Roberts, together with the following specialist group compilers: Geoff Allen (Sphecidae and Crabronidae); Michael Archer (Chrysididae); Graham Collins (Pompilidae); Mike Edwards (Apidae); George Else (Apidae);Adrian Knowles (Sphecidae and Crabronidae); Glenda Orledge (Formicidae) and Matthew Smith (Sphecidae and Crabronidae). The species profiles written by the compilers have been edited by Robin Edwards (BWARS) and Gavin Broad (BRC). Next must be mentioned the large number of entomologists and naturalists who submitted records for the target species, and last, but by no means least, Mike Edwards and Stuart Roberts have also undertaken the huge task of checking all the data and producing the maps.These were drawn on computer using the DMAP mapping package developed by Alan J Morton. O ur thanks go to all these contributors. We are specially indebted to The Trustees of the Natural Histor y Museum, London, who have made their collections of aculeates available for study by our recorders. The completed draft was read in its entirety by Mike Edwards, George Else and Stuart Roberts (BWARS).The Editors are grateful for their help in reducing errors to a minimum. Finally, we are indebted to Henry Arnold at the Biological Records Centre, CEH Monks Wood and Julie Gaunt in CEH Knowledge Management for organising, computer sett ing, printing and distributing the Atlas. 3 l i IN TR OD UCTI ON Five parts of the Provisional Atlas have been published so far: Part 1 with 55 species (Edwards 1997); Part 2 with 55 species (Edwards 1998); Part 3 with 60 species in 59 maps (Edwards & Telfer 200 1); Part 4 with 55 species (Edwards &Telfer 2002); Part 5 with 60 species (Edwards & Broad 2005) W ith this sixth part , another 59 species are added, taking the total number mapped to 344.The atlas coverage by family is shown in Table I . Table I. Breakdown of the British and Irish aculeate fauna by family, showing the coverage of the first six part s of the atlas. Figures based on t he BWARS checklist published in the Members' Handbook (BWARS 2005). Total no. No. spp No. left Family of spp mapped to map Dryinidae 34 0 34 Embolemidae 1 1 0 Bethylidae 22 2 20 Chrysididae 33 24 9 Tiphiidae 3 3 0 Mutillidae 3 3 0 Sapygidae 2 2 0 Scoliidae I 0 I Formicidae 64 27 37 Pompilidae 46 30 16 Vespidae 34 31 3 Sphecidae 7 4 3 Crabronidae 123 85 38 Apidae 268 132 136 Total 64 1 344 297 6 S YS TE M A TI C LI S T OF SP E CI E S MA P P E D T he classification here is as published in the BW ARS Mem bers' Handbook. (BW A RS 2005). Map num bers are given for each species. HYMEN O PT ERA ACU LEATA CH RYSIDO IDEA C H RYSIDIDA E Cleptinae 286 Cleptes nitidulus 287 Cleptes semiauratus Elampinae 288 Holopyga ovata Chrysidinae 289 Chrysis bicolor 290 Chrysis gracillima 291 Chrysis illigeri 292 Trichrysis cyanea VESPO ID EA FO RMIC IDA E Formicinae 293 Plagiolepis taurica Myrmicinae 294 Temnothorax interrupcus 295 Temnothorax nylanderi 296 Temnothorax unifasciatus 297 Myrmecina graminicola PO MPILI DA E Pompilinae 298 Arachnospila anceps 299 Arachnospila consobrina 300 Arachnospila trivialis A PO IDEA CRA BRON IDA E Crabroninae 30 1 Crossocerus annulipes 302 Crossocerus megacephalus 303 Crossocerus wesmaeli 304 Lindenius albilabris 305 Lindenius panzeri 306 Rhopalum clavipes 307 Rhopalum coarctatum 308 Rhopalurn gracile Pemphredoninae 309 Stigmus pendulus 310 Stigmus solskyi A PIDA E Andreninae 311 Andrena bicolor 312 And rena denticulata 313 And rena fuscipes 314 And rena labial is 315 Andrena nitida 316 Andrena simillima 317 Andrena thoracica 318 Andrena tridentata Halictinae 319 Lasioglossum fratellum 320 Lasioglossum fulvicor ne 321 Lasioglossum sexnotatum 322 Lasioglossum villosulum 323 Lasioglossum zonulus 324 Sphecodes reticulatus 325 Sphecodes rubicundus 326 Sphecodes scabricollis Ant hophor inae 327 Anthophora bimaculata 328 Ant hophora furcata 329 Ant hophora plumipes 330 Ant hophora quadrimaculata 331 Ant hophora retusa 332 Eucera longicornis 333 Eucera nigrescens 334 Melecta albifrons 335 Melecta luctuosa 336 Nomada fabriciana 337 Nomada ra pes 338 Nomada sexfasciata Apinae 339 Bombus jonellus 340 Bombus prator um 34 1 Bombus soroeensis 342 Bombus barbutellus 343 Bombus campestris 344 Bombus sylvestris 8 • D IS TR IB UTI ON M AP S A N D SP E CIES P R OFILES Maps 286 to 344 show the recorded distribution of the individual species.
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  • Bees and Wasps of the East Sussex South Downs

    Bees and Wasps of the East Sussex South Downs

    A SURVEY OF THE BEES AND WASPS OF FIFTEEN CHALK GRASSLAND AND CHALK HEATH SITES WITHIN THE EAST SUSSEX SOUTH DOWNS Steven Falk, 2011 A SURVEY OF THE BEES AND WASPS OF FIFTEEN CHALK GRASSLAND AND CHALK HEATH SITES WITHIN THE EAST SUSSEX SOUTH DOWNS Steven Falk, 2011 Abstract For six years between 2003 and 2008, over 100 site visits were made to fifteen chalk grassland and chalk heath sites within the South Downs of Vice-county 14 (East Sussex). This produced a list of 227 bee and wasp species and revealed the comparative frequency of different species, the comparative richness of different sites and provided a basic insight into how many of the species interact with the South Downs at a site and landscape level. The study revealed that, in addition to the character of the semi-natural grasslands present, the bee and wasp fauna is also influenced by the more intensively-managed agricultural landscapes of the Downs, with many species taking advantage of blossoming hedge shrubs, flowery fallow fields, flowery arable field margins, flowering crops such as Rape, plus plants such as buttercups, thistles and dandelions within relatively improved pasture. Some very rare species were encountered, notably the bee Halictus eurygnathus Blüthgen which had not been seen in Britain since 1946. This was eventually recorded at seven sites and was associated with an abundance of Greater Knapweed. The very rare bees Anthophora retusa (Linnaeus) and Andrena niveata Friese were also observed foraging on several dates during their flight periods, providing a better insight into their ecology and conservation requirements.
  • Bee-Plant Networks: Structure, Dynamics and the Metacommunity Concept

    Bee-Plant Networks: Structure, Dynamics and the Metacommunity Concept

    Ber. d. Reinh.-Tüxen-Ges. 28, 23-40. Hannover 2016 Bee-plant networks: structure, dynamics and the metacommunity concept – Anselm Kratochwil und Sabrina Krausch, Osnabrück – Abstract Wild bees play an important role within pollinator-plant webs. The structure of such net- works is influenced by the regional species pool. After special filtering processes an actual pool will be established. According to the results of model studies these processes can be elu- cidated, especially for dry sandy grassland habitats. After restoration of specific plant com- munities (which had been developed mainly by inoculation of plant material) in a sandy area which was not or hardly populated by bees before the colonization process of bees proceeded very quickly. Foraging and nesting resources are triggering the bee species composition. Dis- persal and genetic bottlenecks seem to play a minor role. Functional aspects (e.g. number of generalists, specialists and cleptoparasites; body-size distributions) of the bee communities show that ecosystem stabilizing factors may be restored rapidly. Higher wild-bee diversity and higher numbers of specialized species were found at drier plots, e.g. communities of Koelerio-Corynephoretea and Festuco-Brometea. Bee-plant webs are highly complex systems and combine elements of nestedness, modularization and gradients. Beside structural com- plexity bee-plant networks can be characterized as dynamic systems. This is shown by using the metacommunity concept. Zusammenfassung: Wildbienen-Pflanzenarten-Netzwerke: Struktur, Dynamik und das Metacommunity-Konzept. Wildbienen spielen eine wichtige Rolle innerhalb von Bestäuber-Pflanzen-Netzwerken. Ihre Struktur wird vom jeweiligen regionalen Artenpool bestimmt. Nach spezifischen Filter- prozessen bildet sich ein aktueller Artenpool.