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Tribul Cochylini (Lepidoptera, Tortricidae) În România. Partea III

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Tribul Cochylini (Lepidoptera, Tortricidae) În România. Partea III Bul.inf. Entomol., 17: 5-56, 2006 ISSN 1221 - 5244 Tribul Cochylini (Lepidoptera, Tortricidae) în România. Partea III. (Agapeta, Ceratoxanthis, Fulvoclysia, Eugnosta, Commophila, Prochlidonia, Eupoecilia) Zoltán KOVÁCS & Sándor KOVÁCS Abstract The Tribus Cochylini (Lepidoptera, Tortricidae) in Romania Results of the revision of the tribe Cochylini of Romania are presented. The presences of 89 species belonging to 16 genera are confirmed in the Romanian fauna. The first part of the series dis- cussed the general description of the tribe and the first two genera: Phtheochroa and Hysterophora (Bul. inf. Soc. lepid. rom. 12(1-4): 5-45, 2001), the second part discussed the genera Cochylimorpha, Phalonidia and Gynnidomorpha (Bul. inf. Entomol. 14-15: 57-145, 2003-2004). This third part of the series discusses the genera Agapeta, Ceratoxanthis, Fulvoclysia, Eugnosta, Commophila, Prochlidonia and Eupoecilia. The presences of 12 species belonging to 6 genera are confirmed in the Romanian fauna. The presence of Eugnosta hydrargyrana hydrargyrana (EV E RSMANN , 1842), Commophila ae- neana (HÜBN E R , [1800]) and Eupoecilia sanguisorbana (HE RRIC H - SC H ÄFF E R , 1856) are considered doubtful in the Romanian fauna. Keys to all treated and further one expected species, based on exter- nal characters, male and female genitalia are provided. Fore wings of adults, male and female genitalia are figured. Short description, biological and distributional data are given. The examined material is presented. Key words: Cochylini, Romania, revision, Agapeta, Ceratoxanthis, Fulvocly- sia, Eugnosta, Commophila, Prochlidonia, Eupoecilia. 5 Genul Agapeta HÜBN E R , [1825] Diagnostic: ADULT. Palpii labiali sunt relativ lungi, segmentul al doi- lea este mult lăţit în partea sa subterminală, segmentul al treilea este scurt, dar proeminent. Antenele ajung până la jumătatea aripilor anterioare. Aripa anterioară (Fig. 143-145) este lată, marginea costală la baza aripii este îndoită, în continuare este uşor convexă, plica costală lipseşte. Vârful aripii este scurt. Marginea externă este concavă. Toate nervurile sunt inde- pendente. Chorda este subţire, porneşte la mijlocul distanţei dintre emergenţa nervurilor R1-R2. Originea nervurii R1 este la mijlocul celulei, nervura R5 se termină pe marginea externă sub vârful aripii. Nervura CuP este rudimentară. Media este absentă. Aripa posterioară este lată, apexul este proeminent şi rotunjit. Nervura Sc+R1 este lungă, ajunge la marginea costală aproape de vârful aripii. Ner- vurile Rs şi M1 sunt lung pedunculate, M3-CuA1 scurt pedunculate. Dimorfismul sexual puţin evident, femelele sunt mai mari decât mas- culii. Armătura genitală masculă (Fig. 146-148): Tegumen este lat, slab scle- rotizat, cu pedunculi scurţi. Uncus lipseşte. Tuba analis are un sclerit ven- tral mare. Socii sunt mici şi triunghiulari sau lungi şi înguşti. Vinculum este puternic, ramurile sale ventral sunt sudate numai la A. zoegana L., la celelalte specii nu sunt sudate, capetele lor sunt unite de o membrană fină. Valva este scurtă, dar lată, la baza marginii costale este o apofiză lată spinoasă, pulvinus, care lipseşte şi ea la A. zoegana L. Costa este îngustă, sacculus de formă şi lăţime variabilă. La baza valvei este o apofiză voluminoasă, care lipseşte la A. zoegana L. Transtilla are părţile laterale doar uşor lăţite, partea mediană este mică. Juxta are o apofiză mediană îngustă şi lungă, care la A. zoegana L. este scurtă. Aedeagus este lung şi îngust, apofiza terminală poate să fie foarte lungă. Cornutus lipseşte. Caulis este scurt. Armătura genitală femelă (Fig. 149-151): Ovipozitorul este normal conformat şi scurt. Apofizele posterioare şi anterioare sunt în general scurte. Tergitul VIII este scurt. Sterigma este de formă variabilă, sudată cu peretele posterior al antrumului. Antrum este larg şi complex, bine sclerotizat. Bursa copulatrix are ductus bursae şi corpus bursae bine diferenţiaţi. Ductus bursae este de obicei îngust şi membranos, dar la A. largana RBL . este larg şi bine sclerotizat. Corpus bursae este membranos, are signa formate din sclerite den- tiforme. Observaţii: Desenul şi talia exemplarelor poate fi foarte variabilă, în special la A. hamana L. Răspândire: Genul este răspândit în vestul palearcticului, sunt cuno- scute 4 specii, toate prezente şi în Europa, dintre care 3 au fost semnalate şi 6 din România. Lista speciilor genului Agapeta din România Agapeta HÜBN E R , [1825] Agapete HÜBN E R , [1825] Euxanthis HÜBN E R , [1825] Xanthosetia ST E P he NS , 1829 hamana (LINNA E US , 1758) diversana HÜBN E R , 1793 periphragmella SC H AW E RDA , 1911 albana SC H ILL E , 1917 depuncta OBRAZTSOV , 1943 albana DUFRAN E , 1960 deficiens DUFRAN E , 1960 largana (RE B E L , 1906) vicolana că p u ş e , 1964 zoegana (LINNA E US , 1767) ferrugana HAWORT H , [1811] brunneocycla RAZOWS K I , 1961 Cheia de determinare a speciilor genului Agapeta din România bazată pe morfologia externă a adultului 1 (2) - desenul aripii anterioare este extins asupra apexului şi a marginii ex- terne, care împreună cu banda transversală subapicală delimitează complet zona apicală a aripii ............................................. zoegana L. 2 (1) - desenul aripii anterioare se extinde numai parţial asupra apexului şi a marginii externe 3 (4) - desenul este format din două benzi transversale complete ......................................................................................... largana RBL . 4 (3) - benzile transversale nu sunt complete, desenul este foarte variabil, poate fi extins şi complex sau redus ................................... hamana L. Cheia de determinare a speciilor genului Agapeta din România bazată pe structura armăturii genitale mascule 1 (4) - la baza marginii costale a valvei este o apofiză spinoasă 2 (3) - apofiza este lungă, îndoită, cu vârful ascuţit ...................... hamana L. 3 (2) - apofiza este lată, dreaptă, cu vârful rotunjit .................... largana RBL . 4 (1) - la baza marginii costale a valvei nu este apofiză ............... zoegana L. 7 Cheia de determinare a speciilor genului Agapeta din România bazată pe structura armăturii genitale femele 1 (4) - ductus bursae este membranos 2 (3) - antrum şi sterigma sunt foarte complexe ........................... hamana L. 3 (2) - antrum este complex, sterigma simplă ............................... zoegana L. 4 (1) - ductus bursae este bine sclerotizat .................................. largana RBL . Descrierea speciilor Agapeta hamana (LINNA E US , 1758), (Fig. 143, 146, 149) ADULT: Anvergura aripilor anterioare: 19-23 mm. Ceafa, fruntea, palpii la- biali, tegulele şi toracele sunt de culoare galbenă, partea laterală şi inferioară al palpilor labiali este maro. Aripa anterioară (Fig. 143) lungă şi lată are cu- loarea de fond galbenă. Desenul maro-ruginiu deschis este foarte variabil, format: dintr-o linie îngustă de-a lungul jumătăţii bazale a marginii costale; o bandă transversală mediană îngustă, care frecvent este fragmentată în mai multe pete sau redusă la o pată la mijlocul aripii deasupra marginii posterio- are; o bandă transversală subapicală îngustă, care este şi ea foarte variabilă, porneşte de la marginea costală, în zic-zac ajunge până la celulă, dar poate să fie întreruptă sub costa, iar de la celulă se întinde oblic până la tornus; la apex şi de-a lungul marginii externe a aripii este o linie îngustă, care poate să fie confluentă cu banda transversală subapicală. Pe desenul din jurul ce- lulei şi pe pata marginii posterioare sunt suprapuşi numeroşi solzi de culoare plumburie strălucitoare. Franjurile sunt de culoare galbenă. Aripa posterioară este cenuşie-maronie închisă, franjurile sunt de culoare albă-gălbuie cu o linie bazală maro închisă. Armătura genitală masculă (Fig. 146): Tegumen este lat şi scurt, rotunjit. Socii sunt mici, triunghiulari. Tuba analis are un sclerit ventral mare, bine sclerotizat. Vinculum este îngust, ventral nu este sudat. Transtilla are părţile laterale late şi partea mediană numai uşor lăţită. Valva este lată cu marginea costală mai lungă decât sacculus, pulvinus este puternic sclerotizat, de forma unei gheare. Sacculus este lat la bază, se îngustează treptat şi se termină cu o apofiză terminală ascuţită. La baza valvei este o apofiză voluminoasă, bine sclerotizată, partea lui costală este scurtă şi triunghiulară, partea lui de pe sac- culus este lungă, arcuită, cu dinţi mici, cu vârful ascuţit. Juxta are o apofiză mediană mare, triunghiulară. Aedeagus este lung şi lat, are o apofiză terminală relativ groasă cu vârful recurbat, nu are cornutus. Armătura genitală femelă (Fig. 149): Papillae anales sunt scurte şi în- guste. Tergitul VIII este scurt. Apofizele posterioare şi anterioare sunt scurte 8 Fig. 143–145: Aripile anterioare ale speciilor genului Agapeta: 143 – A. hamana (LINNA E US , 1758) ♀; 144 – A. largana (RE B E L , 1906) ♀; 145 – A. zoegana (LINN - A E US , 1767) ♂ (original). 9 Fig. 146–148: Armăturile genitale ♂♂ ale speciilor genului Agapeta: 146 – A. hamana (LINNA E US , 1758); 147 – A. largana (RE B E L , 1906); 148 – A. zoegana (LINNA E US , 1767) (original). 10 şi subţiri. Sterigma şi antrum sunt foarte complexe, sudate posterior. Ductus bursae este scurt şi îngust, membranos. Corpus bursae este mare, ovoid-alun- git. Signa sunt formate de numeroase sclerite dentiforme mari grupate lângă marginea corpului bursae, pătrund şi în ductus bursae. OBSERVAŢII: Specia este asemănătoare cu A. angelana (KE NN E L , 1919), care este cunoscută numai din Spania şi cu A. largana RBL ., la care benzile transversale
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