Chromosome differentiation and the radiation of the subgenus Agrodiaetus (: : ) – a molecular phylogenetic approach

Dissertation

zur Erlangung des Doktorgrades (Dr. rer. nat.)

der

Mathematisch-Naturwissenschaftlichen Fakultät

der

Rheinischen Friedrich-Wilhelms-Universität Bonn

vorgelegt von

Martin Wiemers

aus

Münster (Westf.)

Bonn – September 2003

Angefertigt mit Genehmigung der Mathematisch-Naturwissenschaftlichen Fakultät der Rheinischen Friedrich-Wilhelms-Universität Bonn

1. Referent: Prof. Dr. Clas M. Naumann 2. Referent: Priv.-Doz. Dr. Bernhard Misof

Tag der Promotion: ______

2 Contents

CONTENTS

CHAPTER 1: INTRODUCTION...... 5 SCOPE AND AIMS...... 7 CHAPTER 2: NEW KARYOLOGICAL RESULTS IN AGRODIAETUS...... 9 INTRODUCTION ...... 9 MATERIAL AND METHODS...... 9 RESULTS ...... 10 Karyotype photographs of Agrodiaetus...... 11 Overview of karyotypes in the subgenus Agrodiaetus Hübner, 1822 ...... 15 DISCUSSION ...... 29 CHAPTER 3: A MOLECULAR PHYLOGENY OF AGRODIAETUS ...... 33 MATERIAL AND METHODS...... 33 Material...... 33 Selection of taxa ...... 36 DNA extraction...... 37 Selection of genes ...... 37 PCR and sequencing...... 38 Sequence alignment...... 39 Choice of methods for phylogenetic inference...... 40 Sequence comparisons and phylogenetic inference...... 41 RESULTS ...... 43 Sequences and alignment...... 43 Base composition and sequence divergence...... 43 Bayesian inference of phylogeny ...... 48 Maximum parsimony analysis ...... 60 Statistical parsimony networks...... 74 DISCUSSION ...... 87 Comparisons with traditional systematics...... 87 Congruence between gene trees and species trees in outgroups...... 90 Comparison with allozyme results...... 94 Hybridization in Agrodiaetus ...... 94 Radiation of Agrodiaetus and the colonization of Europe...... 96 CHAPTER 4: SYSTEMATICS OF AGRODIAETUS BASED ON MOLECULAR EVIDENCE ...... 97 CHAPTER 5: EVOLUTION OF MORPHOLOGICAL TRAITS IN AGRODIAETUS...... 107 INTRODUCTION ...... 107 MATERIAL AND METHODS...... 107 RESULTS ...... 108 DISCUSSION ...... 110 SUMMARY...... 112 CHAPTER 6: THE ROLE OF CHROMOSOME EVOLUTION IN THE RADIATION OF AGRODIAETUS ...... 113 INTRODUCTION ...... 113 MATERIAL AND METHODS...... 113 RESULTS ...... 114 DISCUSSION ...... 116 SUMMARY...... 119 SUMMARY ...... 120 ZUSAMMENFASSUNG ...... 120 ACKNOWLEDGEMENTS...... 121 REFERENCES...... 123 HILFSMITTEL...... 143 3 Contents

APPENDIX 1. KARYOLOGICAL RESULTS IN AGRODIAETUS...... 145 APPENDIX 2. LIST OF MATERIAL...... 149 APPENDIX 3. SEQUENCE DATA ...... 165 PLATES. AGRODIAETUS WINGS ...... 201

4 Introduction

Chapter 1: Introduction

The Holarctic butterfly genus Polyommatus Latreille, 1804 consists of approximately 120-180 species (HÄUSER & ECKWEILER 1997), many of which have been placed into subgroups which are considered separate genera according to numerous authors (e.g. the subgenera Meleageria, Cyaniris, Plebicula, Sublysandra, Neolysandra). The subgenus Agrodiaetus Hübner, 1822, which is distributed in the southern Palaearctic, is the most rich in species. ECKWEILER & HÄUSER (1997) recognize between 56 and 84 species whereas BÁLINT & JOHNSON (1997) list 102 species (including 8 species placed under Paragrodiaetus Rose & Schurian, 1977). Since 1997 alone, 23 further species rank taxa have been described from Anatolia and . The distribution of this subgenus ranges from the Iberian Peninsula to northern Pakistan and Central Mongolia, but the majority of species is found in Anatolia, Transcaucasia and Iran. The characteristic morphological feature of this subgenus is the almost complete reduction of the submarginal elements of the complex wing pattern on the underside of both pairs of wings, and a prominent white streak on the underside of the hind wings (although this can be missing in some cases). Secondly the larval foodplants of all Agrodiaetus species as far as they are known belong to only two closely related genera of Fabaceae, Onobrychis and Hedysarum which are only used by few other Polyommatus species. However, the relevance of these two features for a presumed monophylum Agrodiaetus has never been checked systematically, and therefore the delimitation of this taxon in literature remains contentious. HIGGINS (1975) and TOLMAN & LEWINGTON (1997) also include e.g. the taxa thersites Cantener, 1834 (larval foodplant Onobrychis), escheri Hübner, 1823 (larval foodplant Astragalus) and amanda Schneider, 1792 (larval foodplants and Lathyrus) within Agrodiaetus. BÁLINT & JOHNSON (1997) also place thersites with Agrodiaetus, but split off eight taxa (incl. glaucias (Lederer, 1871), erschoffii (Lederer, 1869) and dagmara (Grum-Grshimailo, 1888)) from Agrodiaetus and place them into Paragrodiaetus Rose & Schurian, 1977. This genus was synonymized with Agrodiaetus by HÄUSER & ECKWEILER (1997). Despite a rather up-to-date species inventory (ECKWEILER & HÄUSER 1997) the exact number of species is uncertain. The reasons are threefold. First, this is the only Palaearctic group of in which a high number of new species is still being discovered, especially in remote parts of Iran. 15% of the almost 250 nominal taxa described in Agrodiaetus (Fig. 1) have been named the last five years, 2/3 of them of species rank. Taxa descriptions in Agrodiaetus 250 200 150

100 Taxa descriptions 50 Total 0 1763 1803 1843 1883 1923 1963 2003 Period

Fig. 1. Taxa descriptions in Agrodiaetus within periods of 20 years

5 Chapter 1

Secondly, many taxa represent allopatric populations which differ only slightly in morphology or karyology (morphospecies and karyospecies) and a decision on their status as distinct species or subspecies is controversial and subjective. Thirdly and most importantly many Agrodiaetus taxa are extremely similar in phenotype (HESSELBARTH et al. 1995) and in contrast to other Lepidoptera taxa genitalia offer only few distinctive features. These slight differences can only be discovered by careful and time consuming preparation techniques, the results of which have been published only for some Greek taxa (COUTSIS 1986; WAKEHAM- DAWSON & SPURDENS 1994).

The discovery of different chromosome numbers in allopatric populations of various Agrodiaetus taxa has lead to the description of numerous “karyospecies” which hardly differ in phenotype. About half of all Agrodiaetus species were described within the last 40 years and differences in chromosome numbers have often been the main or even the only argument for the specific separation of taxa. Currently most authors assume that major differences in chromosome numbers usually lead to reproductive isolation of taxa although experiments in Saturniidae have shown that interspecific crossings between different karyospecies can lead to fertile offspring (NAGARAJU & JOLLY 1986). Variation in chromosome numbers between individuals or populations are also known in many other groups of , e.g. in the Polyommatus subgenus (DE LESSE 1960a & 1969), in some ancestral genera of neotropical Nymphalidae (BROWN et al. 1992), in the ant genus Leptothorax (LOISELLE et al.1990), in the Australian grasshopper Caledia (GROETERS & SHAW, 1992 & 1996; MORAN & SHAW 1977; SHAW 1976), in the New Zealand stick Hemideina (MORGAN-RICHARDS 1997), in the Japanese daddy longleg Gagrellopsis nodulifera (TSURUSAKI et al. 1991), in Australian scorpions of the genus Urodacus (SHANAHAN 1989) or the African rodent Otomys irroratus (CONTRAFATTO et al. 1992). Although karyological studies in the subgenus Agrodiaetus have been important to discover species which are not or only slightly differentiated morphologically, these studies have also lead to a profusion of names and increased the chaos in the systematics of this group. The difficulties in uniting Agrodiaetus species into monophyletic units at or above the species category stem from the small number of usable characters and the high level of homoplasy in most of them. Not surprisingly, all systematic treatments of this group (FORSTER 1956-1961; HESSELBARTH et al. 1995; ECKWEILER & HÄUSER 1997; BÁLINT & JOHNSON 1997) have remained highly unsatisfactory.

Molecular methods appear very promising to unravel the relationships within Agrodiaetus in order to understand more about its origin and radiation. Such knowledge can also help to elucidate the karyotype evolution and biogeography of Agrodiaetus. Until now, only one such study on allozyme variation in 11 taxa, most of them from the Mediterranean region, has been published (MENSI et al. 1994). In Lycaenidae, only a few molecular studies have been conducted, which is apparent from the small number of sequences in GenBank (6 on 22/09/98 when this project started; 176 on 9/9/2003, compared to 1857 in Nymphalidae, a family with approximately the same number of species worldwide (ca. 5000) as in Lycaenidae). One of the few major ones is the thesis of MEGENS (2002) who used molecular markers (COI, COII, wingless, 12s & 16s) to infer the phylogeny of the Southeast Asian butterfly genus Arhopala. This case parallels Agrodiaetus with its high number of morphologically very similar species. Molecular data suggested a rapid radiation which started already in the Miocene (7-11 Million years ago) and was completed before the onset of the Pleistocene period. Thus glaciation events (e.g. the periodical flooding of the Sunda plateau) did not appear to be the key factors for this radiation as previously suggested. Instead, life-history parameters, i.e. shifts in larval foodplants and symbiosis with ants were assumed to have played a major role in creating the diversity of Arhopala species.

6 Introduction

Life history parameters might also have been important in the radiation of Agrodiaetus. Although it is thought that the larvae only feed on two genera of Fabaceae (Onobrychis and Hedysarum), the food plant of most species is unknown. Even the few existing foodplant records can only be evaluated with great caution for various reasons: 1. Females of Agrodiaetus are often impossible to identify in the field. Therefore egg-laying records are prone to misidentification of the Agrodiaetus species. 2. Some Agrodiaetus species do not lay their eggs directly onto the foodplant but on dry plant material next to the foodplant (SCHURIAN 1976; 1999) which can easily result in erroneous records. 3. The larvae of most Agrodiaetus species are unknown and those which are known are very similar morphologically. If larvae are collected in the field they need to be reared until the eclosure of the adult butterfly for identification purposes. Apart from some European species (BOLOGNESI 2000), Agrodiaetus larvae have almost never been reared successfully. The main reason is that almost all Agrodiaetus are monovoltine, overwinter as small larvae and occur in areas with a continental climate much different from the Central European climate where most lepidopterists reside. The foodplants are also difficult to cultivate. 4. The plant genera Onobrychis and Hedysarum are very rich in species and taxonomically they pose the same puzzle to the botanist as Agrodiaetus to the zoologist. Without the help of specialist botanists, most identifications of larval foodplants will be erroneous. In contrast to many other genera of Lycaenidae which are associated with ants in a sometimes very specific way, Agrodiaetus caterpillars seem to be only tended unspecifically by ants (facultative myrmecophily, v. FIEDLER 1995c). However, hardly anything is known about ant associations in this subgenus. The only way to increase the poor knowledge on its life-history would be intensive field studies such as those carried out by DANTCHENKO (1997) on two Russian taxa. Unfortunatly there are hardly any lepidopterists in the centre of Agrodiaetus distribution who could carry out such studies.

Scope and aims

In the course of this thesis the following issues and questions will be addressed:

Chapter 2 presents the results of the karyological studies and provides data for many taxa whose karyotype was unknown previously. It includes a synopsis of all available literature data, much of it published in many different small entomological journals which are difficult to access.

The molecular genetic approach in Chapter 3 is the core of the thesis. Mitochondrial and nuclear DNA of the vast majority of Agrodiaetus species is analyzed by different methods (Maximum Parsimony, Bayesian, network) to infer the origins of and the phylogenetic relationships within Agrodiaetus. Results will be compared to current systematics which is mainly based on features of the adult phenotype. Here I will also address the question of hybridization in Agrodiaetus: Does it occur frequently causing extensive gene flow between different Agrodiaetus taxa? Another point of discussion are biogeographical issues: Where are the origins of Agrodiaetus and how old is its radiation? When was Europe colonized and how many times? Did Agrodiaetus recolonize Europe after the last glaciation or did they survive in South European refugia?

Chapter 4 aims to revise the systematics of Agrodiaetus on the basis of the molecular results, taking into account karyological data and morphology.

7 Chapter 1

The relevance of morphological features for the phylogeny of Agrodiaetus will be re- evaluated on the basis of the molecular phylogeny in Chapter 5: Is the white streak an autapomorphy of Agrodiaetus? Do the monomorphic brown Agrodiaetus form a monophyletic unit? Is the development of androconial patches correlated with the loss of iridescent coloration does it thus replace a visual sexual communication system with an olfactorial one or is it independent of wing coloration?

Chapter 6 discusses the karyological results in the light of the new phylogenetic framework: Do we find phylogenetical or biogeographical patterns in chromosome number variation? Is chromosomal evolution a directional process towards increasing numbers caused by fission or decreasing numbers by repeated fusion events? Are such events confined to certain clades? Can low and/or high chromosome numbers even have an adaptive value? Do chromosomal changes occur as byproducts of speciation or are they the cause of it? Do we even find indications that speciation occurred without a biogeographical barrier, and that chromosomal mutations provide sufficient reproductive isolation for new species to evolve in sympatry?

8 New karyological results in Agrodiaetus

Chapter 2: New karyological results in Agrodiaetus

Introduction A peculiarity of Agrodiaetus is the incredible variation in chromosome numbers ranging from 10 to 130 for the haploid set. Currently the chromosome numbers of about 65% of all Agrodiaetus species are known. The fundamental works were done in the 1950s and 60s by DE LESSE (1952, 1957, 1959a-f, 1960a-c, 1961a-c, 1962a-b, 1963a-c, 1964, 1966) and in recent years by LUKHTANOV (LUKHTANOV 1989, HESSELBARTH et al. 1995, KANDUL & LUKHTANOV 1997, LUKHTANOV et al. 1997, 1998, LUKHTANOV & DANTCHENKO 2002a-b), besides BROWN (1976b, 1977), BROWN & COUTSIS (1978), COUTSIS et al. (1999), LARSEN (1974, 1975), MUNGUIRA et al. (1995), OLIVIER et al. (1999a-b, 2000), TROIANO et al. (1979) and TROIANO & GIRIBALDI (1979). The current knowledge is best of species from Europe and Southwest Asia whereas the chromosome number of most Central Asian species (which represent less than 30% of the total number of species) is still unknown. The haploid chromosome numbers range from n=10-11 in A. birunii Eckweiler & ten Hagen, 19981 to n=128-131 in A. shahrami Skala, 2001. Compared to the modal value of n=24 in Lycaenidae (LORKOVIĆ 1990), which is only found in very few Agrodiaetus species (like A. poseidonides (Staudinger, 1886) and A. turcicus (Koçak, 1977)) these numbers represent either a reduction (less than half) or a multiplication (up to a quintuple) of the probably plesiomorph modal chromosome number of Lycaenidae. The only Lepidoptera species where even higher numbers are known are the closely related Polyommatus dorylas (n=149-151), P. nivescens (n=190-191) from Spain and finally P. atlantica (n=221-223) from Morocco which has the highest chromosome number known in Metazoa. The mechanism which leads to the change in chromosome numbers is unknown, but it seems most probable that low chromosome numbers are caused by fusion and high numbers by fission of chromosomes (LORKOVIĆ, 1990). This is also indicated by the striking reciprocal correlation between the number of chromosomes and their size. Polyploidy, which was recently suggested as a possible mechanism in some Coleophora species (LUKHTANOV & PUPLESIENE 1999), is extremely unlikely in Agrodiaetus.

Material and methods Material available for karyological studies is listed in Appendix 2. There was a total of 1155 chromosome fixations available for study (976 Agrodiaetus and 179 other specimens of Lycaenidae). These were fixed in the field by placing the testes or the posterior two thirds of the abdomens of freshly killed butterflies into Eppendorf vials (0.5ml) with a freshly prepared mixture of 100% Ethanol and 100% Acetic Acid (3:1). The vials were kept in ice water during travel and later on stored in the lab at +7°C. Karyological preparations were made with squash techniques according to the protocols in OLIVIER et al. (2000) or LUKHTANOV & DANTCHENKO (2002a) and carried out in cooperation with Dr. Jurate De Prins (Antwerpen), Karen Meusemann (Bonn) and Dr. Vladimir Lukhtanov (St. Petersburg). Digital images of preparations were obtained with an Olympus DP-50 digital camera attached to a Leica Aristoplan microscope using the software package “Analysis”. The number of chromosomes was counted and peculiarities of the karyotype were recorded. A total of 624 preparations were made (586 Agrodiaetus and 38 other species of Lycaenidae).

1 The even lower numbers of n=8-11 recorded for A. nephohiptamenos (Brown & Coutsis, 1978) have turned out to be erroneous (J. DE PRINS, pers. comm.) like those for A. aroaniensis (Brown, 1976), v. COUTSIS et al. (1999). 9 Chapter 2

Results Meiotic cells with countable chromosome numbers were found in 282 preparations (45%) and chromosome counts are given in Appendix 1. The karyotype of many species was previously unknown (see discussion). A selection of karyotype photos of such species and of those specimens which are of special scientific interest are presented here (Tab. 1; Figs. 2-27).

Tab. 1. List of figured karyotypes Species Chromosome number Figure Specimen codes* Agrodiaetus actis n=17 2 MW98166 Agrodiaetus ainsae n=ca 108 3 MW01004 Agrodiaetus birunii n=10-11 4 MW00267 Agrodiaetus caeruleus n=20 5 MW00335 Agrodiaetus dizinensis n=17 6 & 7 MW00539 Agrodiaetus ernesti n=18 8 MW98097 Agrodiaetus erschoffii n=13-14 9 MW00393 Agrodiaetus femininoides n=27 11 WE02671 Agrodiaetus gorbunovi n=20 10 MW00129 Agrodiaetus iphicarmon n=29 12 MW98104 Agrodiaetus iphigenia n=15 13 MW98106 Agrodiaetus (damocles) kanduli n=25 14 MW99465 Agrodiaetus karindus n=68 15 WE02611 Agrodiaetus klausschuriani n=56 16 MW00262 Agrodiaetus lycius n=22 17 MW98069 Agrodiaetus mofidii sorkhensis n=ca 45 27 WE02453 Agrodiaetus peilei n=39 18 WE02593 Agrodiaetus pierceae n=22 19 MW99416 Agrodiaetus phyllis n=82-86 20 MW00348 Agrodiaetus posthumus n=ca 85 21 MW00347 Agrodiaetus pseudoxerxes n=15 22 MW00330 Agrodiaetus sertavulensis n=20 25 MW98305 Agrodiaetus valiabadi n=23 (2n=46) 23 & 24 MW00498 Agrodiaetus zarathustra n=ca.22 26 WE02531 * Specimen codes refer to those used in Appendix 2.

10 New karyological results in Agrodiaetus

Karyotype photographs of Agrodiaetus

Fig. 2. A. actis, n=17 (MW98166) Fig. 3 Agrodiaetus ainsae, n=ca 108 (MW01004)

Fig. 4. Agrodiaetus birunii, n=10 Fig. 5. Agrodiaetus caeruleus (MW00267) n=20 (MW00335)

Fig. 6. Agrodiaetus dizinensis, n=17 Fig. 7. Agrodiaetus dizinensis, 2n=34 (MW00539) (MW00539)

11 Chapter 2

Fig. 8. Agrodiaetus ernesti, n=18 (MW98097) Fig. 9 Agrodiaetus erschoffii n=13-14 (MW00393)

Fig. 10. Agrodiaetus gorbunovi, n=20 (MW00129) Fig. 11 Agrodiaetus femininoides, n=27 (WE02671)

Fig. 12. Agrodiaetus iphicarmon, n=29 Fig. 13. Agrodiaetus iphigenia, n=15 (MW98104) (MW98106)

12 New karyological results in Agrodiaetus

Fig. 14. Agrodiaetus (damocles) kanduli  Fig. 15. Agrodiaetus karindus, n=68 n=25 (MW99465) (WE02611)

Fig. 16. Agrodiaetus klausschuriani Fig. 17. Agrodiaetus lycius n=56 (MW00262) n=22 (MW98069)

Fig. 18. A. peilei, n=39 (WE02593) Fig. 19. A. pierceae, n=22 (MW99416)

13 Chapter 2

Fig. 20. A. posthumus, n= ca. 85 (MW00347) Fig. 21. A. phyllis, n=82-86 (MW00348)

Fig. 22. A. pseudoxerxes Fig. 23. A. valiabadi n=15 (MW00330) n=23 (MW00498)

Fig. 24 Agrodiaetus valiabadi 2n=46 (MW00498)

Fig. 25. Agrodiaetus sertavulensis Fig. 26. Agrodiaetus zarathustra n=20 (MW98305) n=ca 22 (WE02531)

14 New karyological results in Agrodiaetus

Fig. 27. Agrodiaetus mofidii sorkhensis, n=ca 45 (WE02453)

The following list includes all species level taxa of Agrodiaetus which are included in the molecular study, even if no own karyological data are available. This list summarizes the current knowledge on their karyotypes. At this stage I adopt a splitter’s attitude and consider any taxon which might represent a separate species, whether based on different morphology or karyology and including allopatric taxa which might only represent subspecies of another taxon. An evaluation of their status will be done later in the course of this thesis, taking into account the results of the molecular genetic studies.

Overview of karyotypes in the subgenus Agrodiaetus Hübner, 1822 admetus-group (grouping according to ECKWEILER & HÄUSER, 1997) This is a group of sexually monomorph brown Agrodiaetus taxa which are often difficult or even impossible to identify without the knowledge of their karyotypes. admetus (Esper, [1783]) This species which is distributed from Hungary (type locality) to Siberia is one of the few exceptions and is easy to identify. A haploid chromosome number of n=80 is known from populations in , Western Anatolia and Armenia, and slightly lower numbers (n=77- 79) from Eastern Anatolia (DE LESSE 1960b; LUKHTANOV & DANTCHENKO 2002a). One chromosome is larger and the others are gradually decreasing in size. We can confirm this karyotype with a chromosome number of ca. n=78-80 from three preparations from Antalya and Adana Province (), the two smallest chromosomes being minute in size.

15 Chapter 2 ripartii (Freyer, 1830) This species is known to have a constant chromosome number of n=90 throughout its vast range. Chromosome counts are known from Northern Spain, France (close to the type locality Digne), , Turkey and Kazakhstan (DE LESSE 1960a, 1960b, 1961b, 1961c; LUKHTANOV & DANTCHENKO 2002a, 2002b; COUTSIS et al. 1999). There is one large and one medium sized chromosome while the others are all of similar small size. We were not able to determine the exact chromosome number in any specimen but counts around n=90 were obtained from populations in Spain (Burgos) and Turkey (Isparta, Adana, Artvin, Erzincan & Van). DE LESSE (1960b) remarks that most specimens from Turkey (although fresh) only had atypical divisions. We had the same problem and found typical divisions only in one fifth of the 17 preparations from Turkey. fabressei (Oberthür, 1910) According to DE LESSE (1960a, 1960b, 1961b) and MUNGUIRA et al. (1995) this Spanish taxon which replaces ripartii in Castile has the same chromosome number as ripartii (n=90) but a different number of large chromosomes, two large and two medium ones according to the first author, but three “macrochromosomes” according to the second author team. The metaphases in our preparations were not clear enough to establish the exact number of chromosomes or “macrochromosomes” but in cases of species with chromosomes gradually decreasing in size it is hardly possible to state a certain number of macrochromosomes. This is especially true for squash preparations (as done by MUNGUIRA (l.c.) and us), because the size depends also on how chromosomes were spread when pressing the cover slide. DE LESSE (1961d) found a contact zone of fabressei and ripartii in Teruel but both taxa have not been found sympatrically (DE LESSE 1968; MUNGUIRA et al. 1995). humedasae (Toso & Baletto, 1976) This endemic of Aosta valley (Italy) has a chromosome number of n=38 (TROIANO et al. 1979). We counted approximately the same number of chromosomes. aroaniensis (Brown, 1976) The chromosome number of this Greek taxon is n=48 (COUTSIS et al. 1999), not 15-16 as recorded by BROWN (1976b). No fixations were available to us. nephohiptamenos (Brown & Coutsis, 1978) The chromosome number of this Greek taxon was recorded as n=8-11 (BROWN & COUTSIS 1978), which would constitute the lowest number in Agrodiaetus, but according to DE PRINS (pers. comm.), the species has a very high chromosome number, possibly n=90 as in ripartii. No fixations were available to us. alcestis (Zerny, 1932) With n=19-21, this taxon has the lowest number of chromosomes of all brown Agrodiaetus (DE LESSE 1960A, 1960B; LARSEN 1975; LUKHTANOV et al. 1998; LUKHTANOV & DANTCHENKO 2002B). A haploid number of n=19 was found in Iran and Southeast Turkey, n=20(-21) in other Turkish provinces and Lebanon. Our results from Turkey and Iran confirm these results. The ssp. karacetinae Lukhtanov & Dantchenko, 2002 has been described recently from Dez valley (Hakkari) and Iran for the populations with a karyotype of n=19 chromosomes. demavendi (Pfeiffer, 1938) Variable chromosome counts of n=66-76 are known for this monomorphic Agrodiaetus species from various provinces in Iran, Turkey and Armenia (DE LESSE 1960A, 1960B;

16 New karyological results in Agrodiaetus

LUKHTANOV et al. 1998; LUKHTANOV & DANTCHENKO 2002A) and our results from East Turkey and Northwest Iran fall within this range. DE LESSE (1960b) also notes the higher number of larger chromosomes (two large and several medium ones) in comparison with ripartii with only one large and one medium-sized chromosome. From the excellent photographs of LUKHTANOV & DANTCHENKO (2002) it is apparent that the chromosomes of demavendi gradually decrease in size. Populations from Kopet Dagh (Iran) with an even higher chromosome number of n=84 (DE LESSE 1963a) have been separated as khorasanensis (Carbonell, 2001). Our material from Tehran (Samqabad) is referable by external features to the newly described taxon ahmadi (Carbonell, 2001) from Zanjan and some specimens from Azarbayjan-e Sharqi are similar to the newly described taxon urmiaensis (Schurian & ten Hagen, 2003) from Urmia Lake (Northwest Iran). Although the ranges of demavendi and ripartii overlap in Eastern Anatolia a sympatric occurrence of these two taxa, verified karyologically, is not known. interjectus (de Lesse, 1960) This taxon from Eastern Turkey has been described only on the basis of its karyotype (n=29- 32) and subsequent authors have doubted its status as a distinct species (HESSELBARTH et al. 1995; ECKWEILER & HÄUSER 1997). According to DE LESSE (1960b) it occurs sympatrically with alcestis and demavendi near Pertek. We were able to find a specimen close to the type locality near Erzincan which turned out to have n=31, confirming the results of DE LESSE (1960b). dantchenkoi Lukhtanov & Wiemers, 2003 This is a newly discovered brown Agrodiaetus species discovered in Van (Southeast Turkey) with a chromosome number of n=40-42 (LUKHTANOV, WIEMERS & MEUSEMANN, in print). Specimens MW99274, MW99319 and MW99320 constitute paratypes. Externally and genetically this taxon is similar to eriwanensis (Forster, 1960) from Armenia, which has a chromosome number of n=28-35 (LUKHTANOV & DANTCHENKO, 2002a). valiabadi (Rose & Schurian, 1977) The karyotype of this local taxon from valleys of Northern Elburs (Iran) was previously unknown. We were able to establish the chromosome number as n=23.

dolus-group mithridates (Staudinger, 1878) Variable chromosome numbers of n=21-27 have been recorded from different parts of its range in Turkey (DE LESSE 1960a, 1960b). Our result of n=23 from Malatya corresponds exactly with the result of DE LESSE (1960b) from the same place. peilei Bethune-Baker, 1921 The chromosome number of this peculiar golden brown Agrodiaetus from Lorestan (Iran) was unknown previously. We were able to determine it from three specimens as n=39. antidolus (Rebel, 1901) This is the first of a group of four apparently closely related allopatric species which is known from Eastern Turkey and has a chromosome number of n=40-41 with chromosomes gradually decreasing in size (DE LESSE 1961b; HESSELBARTH et al., 1995; LUKHTANOV et al. 1998). The taxon has been described from Kazikoparan (Iğdır Prov.) and chromosome counts are known from Pertek (Tunceli), Çatak (Van) and Bagishli (Hakkari). We tried to find the species at the

17 Chapter 2 type locality without success but we can confirm the chromosome results for four populations from the province of Hakkari. kurdistanicus (Forster, 1961) Apparently this species is indistinguishable from the former taxon by external features, but its chromosome number is higher, n=61 was recorded from Görentach Köyu N of Van (DE LESSE (1960a) and n=ca. 57-62 from Çatak (Van Province) by DE LESSE (1960a) & LUKHTANOV et al. (1998). Interestingly the latter authors also found one specimen with antidolus karyotype at Çatak. We obtained similar chromosome counts from Çatak and Erek Dağı (Van Province), with one precise result from Çatak (n=62). Erek Dağı is the type locality of kurdistanicus. Although we did not obtain precise counts from this locality we can confirm that the karyotype with high chromosome number occurs at the type locality. This is important for the stability of the name kurdistanicus because antidolus populations with low chromosome number karyotype occur in close proximity. morgani (Le Cerf, 1909) The chromosome number of this taxon from Iranian Kordestan was determined as n=25-26 by DE LESSE (1961b) from a population near Sanandaj. Our result (n=27 from Saqqez) is very close to this figure. femininoides (Eckweiler, 1987) The chromosome number of this darkest member of the antidolus species group from Northwest Iran was unknown until now. We were able to determine its chromosome number from two populations in Zanjan and Ardabil with n=27 to be the same as in morgani. sennanensis (De Lesse, 1959) The chromosome number of this taxon from Iranian Kordestan which was described as a subspecies of hopfferi was established from the type series (including the holotype from Hamakasi, Sanandaj) as n=28-30 (DE LESSE 1959c). No fixations were available to us. menalcas (Freyer, [1837]) The chromosome number of this Turkish taxon was recorded to be stable with n=85 in several populations throughout Turkey (DE LESSE 1960a, 1961a). Our results (the first ones from Fethiye and Van Province) confirm these results and we can also confirm that two of these chromosomes are much larger than the other ones (“macrochromosomes”). dolus (Hübner, [1823]) We did not have material from Southern France where DE LESSE (1960a, 1961a, 1962b) recorded a chromosome number of n=123-125, the second highest number known in Agrodiaetus, but the following two Spanish taxa are very closely related. It should be noted that the type locality of dolus is unknown and could be Southern France (as often suggested), Italy or Spain. ainsae (Forster, 1961) This taxon represents dolus in Northern Spain and was separated from it because of its lower chromosome number of n=108-110 (DE LESSE 1962b; MUNGUIRA et al. 1995). We can confirm this chromosome number. According to MUNGUIRA et al. (1995) ainsae has two “macrochromosomes” compared to six in fulgens which is the reason to consider these allopatric taxa distinct species. This number is based on a single specimen from Burgos of which the number of chromosomes could not be established due to the poor quality of the preparation and probably for this reason it is also not figured. We obtained good metaphase

18 New karyological results in Agrodiaetus plates of two specimens from Illarduya (Alava) which display 108 chromosomes gradually decreasing in size, about six of them could be called “macrochromosomes” according to MUNGUIRA et al. (1995). DE LESSE (1962b) described a new subspecies (A. dolus pseudovirgilia) from Burgos, only based on the small wing size and slight differences in coloration. fulgens (de Sagarra, 1925) This taxon from Catalonia was separated by MUNGUIRA et al. (1995) from ainsae due to the higher number of “macrochromosomes” (six compared to two in ainsae) while the total number of chromosomes was found to be very similar (n=103, judged from three specimens). As is shown in the previous paragraph (under ainsae) this difference in “macrochromosomes” does not really exist. It should also be noted that a precise count of such high chromosome numbers and the identification of an exact number of macrochromosomes which gradually decrease in size is hardly possible when using the squash technique. DE LESSE (1966) already mentioned that (also with his technique) higher numbers of good metaphase plates are necessary to establish the exact chromosome number in species with such a high number of chromosomes. This taxon cannot be separated from ainsae by external characters either. We did not get precise counts from our metaphase plates because some chromosomes seem to overlap but the chromosome number of fulgens must be between n=100 and n=110. dama-group dama (Staudinger, 1892) This South Anatolian species has only been found in a few localities in Malatya, Maraş and Mardin Province (Turkey). DE LESSE (1959f) described its karyotype from Kahramanmaraş and OLIVIER et al. (1999b) confirmed his results from the type locality Malatya. It has an asymmetric karyotype with n=41 chromosomes, about eleven of them are large, gradually decreasing in size, the others medium-sized. hamadanensis (de Lesse, 1959) The chromosome number of this Iranian taxon was found by DE LESSE (1959a) to be n=21-22 in two populations from the type locality (Araj Pass) and from Sanandaj (Kordestan). We can confirm these results for two populations from Tehran province and a population from Ardabil (Azarbayjan-e Sharqi Province). karindus (Riley, 1921) The karyotype of this Iranian taxon was previously unknown and we were able to determine the chromosome number as n=66-68 for two specimens from Saqqez (Kordestan). This taxon has been treated as a subspecies of dama (Staudinger, 1892) which is only known from Malatya (Turkey), but the latter has a chromosome number of n=41-42 (DE LESSE 1959f; OLIVIER et al. 1999). theresiae Schurian, van Oorschot & van den Brink, 1992 The karyotype of this taxon which was described from Saimbeyli in Adana Province (Turkey) was recorded as n=41-42 (HESSELBARTH et al. 1995; KANDUL & LUKHTANOV 1997), but the material came from another population near Taşkent (Konya Province). OLIVIER et al. (1999) proved with our material (specimens MW98240, MW98241, MW98243) that the chromosome number of true theresiae is n=63. The population from Taşkent was described accordingly as the following new species.

19 Chapter 2 guezelmavi Olivier, Puplesiene, van der Poorten, De Prins & Wiemers, 1999 Separated from theresiae on the basis of its karyotype which is n=41-42 (KANDUL & LUKHTANOV 1997; OLIVIER et al. 1999) and only known from its type locality (Taşkent, Konya Prov., Turkey). Our material confirms these results. Externally this taxon cannot be separated from the allopatric theresiae.

damon-group hopfferi (Herrich-Schäffer, [1851]) For this Anatolian species a stable chromosome number of n=15 was recorded by DE LESSE (1959c, 1959f, 1960a) and LUKHTANOV et al. (1998) from different Turkish Provinces. We can confirm these results for Malatya, Van and Hakkari Province. lycius (Carbonell, 1996) This recently discovered species was only known from the type locality (Elmali in Antalya Province, Turkey) until a further population was discovered on the Turkey expedition 1998 near Güneykent (Isparta Province). Its haploid chromosome number was only approximately determined by CARBONELL (1996) as ca. 20-25. We can specify this number more precisely for three specimens from the type locality with n=21-22. poseidon (Herrich-Schäffer, [1851]) DE LESSE (1963c) recorded different chromosome numbers from different localities: n=19 (Pozantı, Adana Prov.), n=20 (Kahramanmaraş) and n=21-22 (Amasya). KANDUL & LUKHTANOV (1997) recorded n=19 from Yusufeli (Artvin) and OLIVIER et al. (1999) n=20 from Malatya. We also obtained different results for populations from Zelve (Nevşehir) with n=20-21 and from Gökpinar (Sivas) with n=19. A clear geographical pattern does not emerge from these results. In one population from Ağrı DE LESSE (1963c) found a much higher chromosome number (n=24-26) but did not describe it as a new taxon. Only recently it was named as the following new karyospecies: putnami (Dantchenko & Lukhtanov, 2002) This taxon was separated from poseidon and described as a new species because of its different karyotype. The holotype from Kayabaşı (Prov. Erzurum) had a haploid chromosome number of n=26 and 4 males from Ağrı (2.VIII.1958, De Lesse leg., n=24-27) were included in the type series. We checked material from Ağrı and found n=25 in one specimen, thus specifying the results of DE LESSE (1963c). In another specimen from Yusufeli (Artvin) the chromosome number could be only approximately established as n=23-25. This specimen was attributed to putnami by LUKHTANOV (pers. comm.). If this is correct, both the karyospecies putnami and poseidon would occur in sympatry at Yusufeli, but in the author’s opinion the plates are not clear enough to rule out a lower number of only n=21 chromosomes as found in poseidon. damocles (Herrich-Schäffer, [1844]) Until recently, this species was only known from the South Ural Mts. (ssp. damocles (Herrich-Schäffer, [1844]), the Crimean Peninsula (ssp. krymaeus (Sheljuzhko, 1928) with n=25-27 according to KANDUL & LUKHTANOV (1997)) and the Volga river (ssp. rossicus Dantchenko & Lukhtanov, 1993) with n=24-26 according to LUKHTANOV et al. 1997). LUKHTANOV & DANTCHENKO (2002b) recorded this species for the first time from Turkey and described it as a new subspecies of damocles:

20 New karyological results in Agrodiaetus kanduli (Dantchenko & Lukhtanov, 2002) This taxon was described as a subspecies of damocles (Herrich-Schäffer, [1844]). The chromosome number of the holotype from Yildiz (Erzincan, Turkey) was determined as n=25. Three other males from the same locality were included in the type series but without karyological data. Specimens with similar phenotype have been found before in Eastern Turkey but were assigned to wagneri (HESSELBARTH et al., 1995). We found one specimen (MW99465) near Çatak (Van Province) which resembles this taxon closely in its phenotype and which turned out to have the same karyotype (n=25). It was assigned to damocles by LUKHTANOV (pers. comm.). caeruleus (Staudinger, 1871) The karyotype of this species was unknown until now. We were able to determine a haploid chromosome number of n=20 from two specimens of both type localities, Schakuh and Hajiabad (Prov. Golestan, Iran). transcaspicus (Heyne, [1895]) A haploid chromosome number of n=52-53 has been recorded by DE LESSE (1963a, 1963c) for this species from Kopet Dagh. The following six taxa are very similar to it in phenotype but have very different chromosome numbers: elbursicus (Forster, 1956) DE LESSE (1963c) checked material from the type locality of this taxon at Kendevan Pass (Elburs, Iran) and recorded a haploid chromosome number of n=16-17. Our results from four populations around Kendevan Pass (n=16-18) confirm these results. We also obtained a result of one specimen similar to elbursicus from Zanjan with n=18. A slightly higher chromosome number of n=19-20 was found by DE LESSE (1963c) in populations from the Eastern Elburs Mts. from Āb Ali (Demavend) and Firuzkuh. zapvadi (Carbonell, 1993) This taxon was described as a subspecies of elbursicus from Güzelsu (Van Province, Turkey) with a haploid chromosome number of n=18-19 (DE LESSE 1963c). LUKHTANOV et al. (1998) recorded a chromosome number of n=17-18 from the type locality and we can confirm a number of n=18-19 for the same locality. HESSELBARTH et al. (1995) synonymized this taxon with elbursicus. ninae (Forster, 1956) The karyotype of this taxon was recorded to be n=34 in Armenia near the type locality (Daralagez Mts.) by LUKHTANOV (1989) and LUKHTANOV & DANTCHENKO (2002a) and n=33-37 in Northeast Turkey (Doğubayazıt, Ararat, Ağrı) by DE LESSE (1963c). Chromosomes are gradually decreasing in size. Our result of a specimen from Ağrı (n=34) confirms these figures. turcicola (Koçak, 1977) This taxon was described as a subspecies of transcaspicus from Van (Turkey). Its phenotype seems to be indistinguishable from the allopatric ninae but its chromosome number is n=(19-)20 (DE LESSE 1960a, 1963c; LUKHTANOV et al. 1998). Our results confirm a chromosome number of n=20 for three populations in Van Province but one specimen from Çatak had a higher number (n=22) which is the same as in the closely related aserbaidschanus (Forster, 1956) from Azarbaijan. Sympatric occurrence with zapvadi has been reported from many locations in Van Province, and at one of them (Güzelsu)

21 Chapter 2

LUKHTANOV et al. (1998) reported a chromosome number of n=19-20 compared to n=17-18 for zapvadi at the same location. huberti (Carbonell, 1993) This species was separated from ninae based on slight differences in phenotype while the karyotype is hardly different. A chromosome number of n=33-34 was recorded by DE LESSE (1963c) from the type locality at Ağrı and LUKHTANOV & DANTCHENKO (2002a) recorded n=35-37 for Armenia. We obtained chromosome counts of n=33-36 from populations in Northeast Turkey (Artvin, Kars & Bayburt Provinces). pierceae (Lukhtanov & Dantchenko, 2002) LUKHTANOV et al. (1998) discovered that huberti from Güzeldere Geçidi (Van Prov.) has a divergent karyotype with n=22 and therefore it was described as a distinct species four years later. We obtained chromosome counts of n=ca.22 also from Çatak (Van Prov.) and Dez valley (Hakkari). carmon (Herrich-Schäffer, [1851]) DE LESSE (1960a, 1963b) investigated material of this Anatolian species from the type locality (Amasya) and from Gümüşhane and Kayseri and found a haploid chromosome number of n=81-82. One chromosome is much larger than all the other ones. We obtained only an approximate result from Artvin Province which is similar (n>=79) while two specimens from Antalya Province gave no results at all. schuriani (Rose, 1978) This taxon was described as a subspecies of carmon from Nevşehir but sunk into synonymy by HESSELBARTH et al. (1995). We found a specimen at Gezbeli Geçidi (Kayseri) which might represent this taxon and the chromosome number was only approximately identified as n=75-80 which does not exclude the possibility that the true number is n=81-82 as established by DE LESSE (1963b) from Kayseri. surakovi Dantchenko & Lukhtanov, 1994 This taxon was described as a subspecies of carmon from Azarbaijan but later raised to species status because of its different karyotype with n=50 recorded from three Armenian populations. The chromosomes gradually decrease in size (LUKHTANOV & DANTCHENKO 2002a). LUKHTANOV & DANTCHENKO (2002b) recorded this species also from Turkey and named it as a different subspecies: sekercioglui (Dantchenko & Lukhtanov, 2002) Described from Çatak (Van Province, Turkey) as a subspecies of surakovi with the same chromosome number of n=50 (LUKHTANOV & DANTCHENKO 2002b). We obtained only an approximate chromosome count from one specimen collected near the type locality which is close to that figure. pseudoxerxes (Forster, 1956) This species was described as a subspecies of carmon from Schakuh in Eastern Elburs Mts. (Iran) and at the same time kendevani (Forster, 1956) was described as another subspecies of carmon from Kendevan Pass in Central Elburs Mts. (Iran). DE LESSE (1962a) ruled out conspecifity of both taxa (as recently suggested again by ECKWEILER & HÄUSER 1997) because he found both taxa sympatric at Kendevan pass and treats pseudoxerxes as a subspecies of altivagans. He established a chromosome number of n=16 for pseudoxerxes from Kendevan Pass. The chromosome number of populations from East Elburs was

22 New karyological results in Agrodiaetus unknown until now. We were able to determine a haploid chromosome number of n=15 in a specimen from the type locality Schakuh. dizinensis (Schurian, 1982) The karyotype of this species was unknown until now. We found a haploid chromosome number of n=17 in a specimen from the type locality Dizin (Tehran, Iran), the only known locality of this species. cyaneus (Staudinger, 1899) Variable chromosome numbers of n=16-20 were found in different populations from Iran, Azarbaijan, Armenia and Turkey (Van Province) (DE LESSE 1960a, 1963b; LUKHTANOV et al. 1998; LUKHTANOV 1989; LUKHTANOV & DANTCHENKO (2002a)). Our results from Van Province (n=18) and Kars Province (n=17) fall within this range. Several subspecies of cyaneus have been described from Iran which are sometimes treated as distinct species (ssp. xerxes (Staudinger, 1899) from Elburs Mts., ssp. paracyaneus (De Lesse, 1963) from Hamadan, ssp. damalis (Riley, 1921) from Karind Gorge (Zagros Mts.), ssp. fredi Eckweiler, 1997 from Fars and ssp. kermansis (De Lesse, 1962) from Kerman). merhaba De Prins, van der Poorten, Borie, Oorschot, Riemis & Coenen, 1991 This species is only known from its type locality at Yusufeli (Artvin Province). Its chromosome number is n=16-17 (LUKHTANOV et al. 1998) and this figure can be confirmed by our data. zarathustra Eckweiler, 1997 The karyotype of this species from Dorud (Lorestan, Iran) was unknown until now. According to our results, it has a haploid chromosome number of n=ca 22. actis (Herrich-Schäffer, [1851]) OLIVIER (2000a) discovered the syntypes of this species and designated a lectotype from Tokat. This was important because the type locality of this taxon was very imprecisely stated as “Kleinasien” (= Asia minor) in its original description. Several recent authors including HESSELBARTH et al. (1995) and ECKWEILER & HÄUSER (1997) treated a different taxon under the name actis (see under sigberti). The chromosome number of actis from the type locality or surroundings is unknown and the expedition to Central Turkey in 1998 did not succeed in finding this species near Tokat or Amasya. The closest population we found was near Gürün (Sivas Province) of which we established a haploid chromosome number of n=17 from two specimens. This came as a surprise because only much higher chromosome numbers (n=24- 34) are known from phenotypically similar populations in other parts of Turkey, all of them more distant from Tokat. These populations are provisionally treated under the following name: firdussii (Forster, 1956) An extraordinary variation of chromosome numbers (n=21-34) has been recorded for this taxon and several authors suggested that it consists of different species, but there is no clear geographical pattern and populations with different karyotypes have no distinctive phenotypic features. The chromosome number of this species from the type locality Schakuh (East Elburs Mts., Iran) is unknown, but DE LESSE (1960a, 1962a) obtained counts of relatively high numbers (n=31-34) from Demavend (East Elburs). He recorded similar chromosome numbers also from Kop Dağı in Turkey (n=30-32), but in another location just 80km to the west (25 km N Erzincan) he obtained chromosome numbers of only n=24-25. This material from Erzincan was included in the type series of Agrodiaetus bilgini Dantchenko & Lukhtanov,

23 Chapter 2

2002. The description of this taxon, with the holotype from Torul (Gümüşhane Prov., Turkey) was based only on the difference in chromosome numbers (n=25). We checked a population 25 km SE Erzincan (Çağlayan) and obtained chromosome counts of n=30-32 in three specimens. LUKHTANOV et al. (1998) found low chromosome numbers also in Van Province (n=21-23 at Güzeldere Geçidi and n=25 at Çatak) and Bitlis Province (n=25 at Kuzgunkiran Geçidi). The material of the latter two populations was included in the type series of Agrodiaetus haigi Dantchenko & Lukhtanov, 2000, together with the holotype from Çatak (n=25) and further paratypes from Çatak (n=25-26), Kurubas Geçidi (n=25) and Güzeldere Geçidi (n=25). Our chromosome counts from Van Province (n=25 from Çatak and n=26 from Güzeldere Geçidi) confirm these results. The picture is complicated by the fact that intermediate chromosome numbers of n=27-29 are also known from populations in northwest Iran and eastern Turkey. DE LESSE (1960a, 1962a) found a haploid chromosome number of n=28 near (Azarbayjan-e Sharqi, Iran) and n=27-29 at Mirgemir Dağı (Ağrı, Turkey). We also found a population with n=28-29 near Zanjan (Iran) and with n=25-27 in Dez valley (Hakkari Prov., Turkey). pseudactis (Forster, 1960) This taxon which was described from Daralagez Mts. in Nachichevan (Azerbaydzhan) is very similar to firdussii and its status (species, subspecies or synonym) is in dispute. LUKHTANOV & DANTCHENKO (2002a) obtained a chromosome number of n=29 from Armenian populations. artvinensis (Carbonell, 1997) Another species which is closely related to firdussii and only known from Erzurum Prov. (type locality: Tortum) and Artvin Prov. in Turkey. Its chromosome number was found to be n=21-22 at Kilizkaya in Artvin Prov. (OLIVIER et al. 2000) and we can confirm these results from the same locality. sigberti Olivier, van der Poorten, Puplesiene & De Prins, 2000 This name was introduced to denote a taxon previously referred to under the name actis. It is mainly distributed in the Taurus Mts. and a few locations in the Pontic chain. The karyotype was described by LUKHTANOV & DANTCHENKO (2002b) who counted n=29 chromosomes in two paratypes from the type locality (Aladaglari, Niğde Prov., Turkey). We checked males from the same locality and obtained n=25 and n=28 in two different specimens. DE LESSE (1962a) found n=27 in a population from Bürücek (İçel Prov.) which probably belongs to this taxon. ernesti Eckweiler, 1989 This taxon which was described as a subspecies of firdussii was only known from the type locality (Kohu Dağı in Antalya Prov., Turkey) and its karyotype was unknown until now. We found two specimens at a new locality (Dedegöl Geçidi, Isparta Prov.) of which one revealed its karyotype which is n=18. sertavulensis (Koçak, 1979) The karyotype of this taxon described as a subspecies of pseudactis and only known from the Taurus mountains was unknown until now. We were able to establish its karyotype from two specimens from Yellibelli Geçidi (Karaman Prov.) which is 33 km West of the type locality (Sertavul Pass) as n=20.

24 New karyological results in Agrodiaetus maraschi (Forster, 1956) This taxon has been described from Kahramanmaraş as a subspecies of damone and treated as a distinct species (KOÇAK 1979), as a synonym of wagneri (s. HESSELBARTH et al. 1995) or as a subspecies of firdussii (s. ECKWEILER & HÄUSER 1997). DE LESSE (1962a) found a specimen of this taxon at Erciyes Dağı (Kayseri) and established a haploid chromosome number of n=16. We can confirm this result from a specimen found near Gürün which is 40 km closer to the type locality. altivagans (Forster, 1956) This taxon was described from the high altitudes (2800m) of Daralagez Mts. (Nachichevan, Azerbaydzhan). DE LESSE (1962a) records a variable chromosome number from Turkish populations: n=21-23 in Erzurum and Ağrı Prov., but a lower number of n=18 in two populations near Erzincan. LUKHTANOV & DANTCHENKO (2002a) confirmed the first figure for two Armenian populations (n=20-23) and we can confirm it for Güzeldere Geçidi in Van Province of Turkey (n=21-23). DE LESSE (1962a) also checked Iranian populations which he referred to as “A. altivagans ssp.” from Marand (n=19-22), Tabriz (n=18), Hamadan (n=18) and Sanandadj (n=17-18). The latter two were described as ssp. ectabanensis (de Lesse, 1963) while the first two were left undescribed due to lack of sufficient material. The one from Marand probably belongs to the following taxon: gorbunovi Dantchenko & Lukhtanov, 1994 This taxon was described as a new species which is closely related to ectabanensis from Talysh Mts. in Azerbaydzhan. Its chromosome number has not been reported, but the specimens of “altivagans ssp.” which DE LESSE (1962a) caught east of Marand (n=19-22) might represent this taxon. We collected two specimens at Dugijan (30 km East of Marand) and obtained a chromosome number of n=ca. 20. Another specimen from Pass (Azarbayjan-e Sharqi) gave the same result and a specimen from Khalkhal, Gollijeh (Ardabil) had n=19 chromosomes. wagneri (Forster, 1956) This taxon was described as a subspecies of damone and elevated to species rank by HESSELBARTH et al. (1995). DE LESSE (1960a, 1962a) karyotyped a specimen from the type locality Aksehir (Konya Prov., Turkey) and found n=16 chromosomes. Other results have not been published, yet. We found a specimen at Zelve (Nevşehir) which resembles the holotype but has n=18 chromosomes. turcicus (Koçak, 1977) The karyotype of this Eastern Anatolian species has been described by LUKHTANOV et al. (1998) of populations from Erzurum and Çatak (Van Prov.). It is one of only two species of Agrodiaetus known to have a stable haploid chromosome number of n=24 which is the modal chromosome number in Lycaenidae. (The other species is poseidonides (Staudinger, 1886)). We can confirm this result from Çağlayan (Erzincan Prov.). baytopi (de Lesse, 1959) DE LESSE (1959d) established the karyotype of this Eastern Anatolian species in his original description as n=27 from the type locality (Doğubayazıt, Ağrı Prov.). We can confirm this result from Güzeldere Geçidi (Van Prov.). DE LESSE (1959d) had problems to find eupyrene spermatogonia in this species and we also managed to find a good metaphase in only one out of 13 investigated specimens.

25 Chapter 2 rovshani Dantchenko & Lukhtanov, 1994 The karyotype of this species which has been described from Talysh mts. in Azerbaidzhan and which seems to be closely related to baytopi remains unknown. We did not obtain a result from a specimen collected in Dugijan (Azarbayjan-e Sharqi, Iran). tankeri (de Lesse, 1960) DE LESSE (1960c) described this species after he discovered (DE LESSE 1957, 1959e) that populations externally similar to iphigenia from Kop Dağı Geçidi and Mirgemir Dağı have a different karyotype with n=20-21 chromosomes and both taxa occur sympatrically on Mirgemir Dağı. The karyotype is asymmetric with nine chromosomes much larger than the other ones. We checked nine specimens from the type locality (Kopdağı Geçidi) and one from Gölyurt Geçidi (Erzurum) but were unable to find any eupyrene metaphases. iphigenia (Herrich-Schäffer, [1847]) A variable chromosome number of n=12-16 has been found in Anatolian populations. The lowest numbers (n=12) were found in NE Turkey, n=13 mainly in SE Turkey and n=15-16 in Eastern Taurus (DE LESSE 1959e; LUKHTANOV et al. 1998). We can confirm n=12 for Erzincan and found n=14-15 in Western Taurus (Fethiye and Isparta Prov.). The ssp. nonacriensis (Brown, 1977) has been described from Peleponnesos, Greece, at the Western distributional limit of this species. The separation was made partly due to its lower chromosome numbers of n=10-13 (BROWN 1977), but most other chromosome counts of this author have turned out to be erroneous, thus it should be verified again. iphicarmon Eckweiler & Rose, 1993 This taxon was described as a subspecies of iphigenia although it was found sympatrically with the latter taxon at the type locality Dedegöl Geçidi (Isparta Prov., Turkey), its only known locality (ECKWEILER & ROSE 1993). We revealed its karyotype which has a chromosome number of n=29. We were also able to check a specimen of iphigenia from the same locality which turned out to have n=15. This proves that iphicarmon cannot be conspecific with iphigenia. iphidamon (Staudinger, 1899) This North Iranian taxon is often treated as a subspecies of iphigenia (e.g. HESSELBARTH et al. 1995) because of its similar chromosome number (n=14 at Āb Ali and Firuzkuh in Eastern Elburs according to DE LESSE 1959e). We can confirm this number now for the type locality Schakuh and for Hajiabad (Golestan, Iran). damon ([Denis & Schiffermüller], 1775) The most widespread Agrodiaetus species which is found from Northern Spain to Mongolia seems to have a stable chromosome number of n=45 (with one large chromosome) throughout its range. Counts have been made in populations from the French Pyrenees, French Alps, Mt. Ararat (Turkey) and Altai Mts. (DE LESSE 1960a; LUKHTANOV 1989). We did not obtain cells with typical divisions from our four preparations from Eastern Turkey. mofidii (de Lesse, 1963) DE LESSE (1963a) described this species from Kopet Dağı, Iran with a chromosome number of n=34-35 (n=34 in the holotype). We checked two specimens from Kuh-e-Sorkh (Khorasan) of the newly described ssp. sorkhensis Eckweiler, 2003. These turned out to have a higher chromosome number of n=ca.42-45 and might therefore represent a different species.

26 New karyological results in Agrodiaetus phyllis (Christoph, 1877) DE LESSE (1959b) studied material from Kendevan Pass and Demavend (Elburs Mts., Iran) and found only a slight variation in chromosome numbers (n=79-82) and a very peculiar karyotype with one extremely large and three other large chromosomes while the rest was very small. A very similar karyotype with n=78 was found by DE LESSE (1957, 1959b) in several Eastern Anatolian populations (ssp. vanensis (de Lesse, 1957)) and by LUKHTANOV & DANTCHENKO (2002a) with n=79-80 in Armenia (ssp. sheljuzhkoi (Forster, 1960)) while the karyotype of dagestanicus (Forster, 1960) from Dagestan (Russia) turned out to be very different (n=40) and represents another species (LUKHTANOV & DANTCHENKO 2002a). We are able to confirm this karyotype not only from Kendevan Pass and Demavend but also from Schakuh, the type locality of phyllis. We found n>=82 chromosomes in specimen MW00348 and can confirm the existence of one huge outstanding bivalent, which is 10 times larger than the mid-size bivalents as well as three large bivalents which are 3-4 times larger than the mid- size bivalents. posthumus (Christoph, 1877) DE LESSE (1957, 1959b) checked the karyotype of specimens which he thought to represent this taxon and found their karyotype to be very different from phyllis with n=10-11 chromosomes. However he did not check topotypical material from Schakuh, but from other localities in Elburs Mts. (Kendevan Pass, Demavend, Firuzkuh) and these populations belong to a different taxon later named birunii (see the next paragraph). We were first to examine material from the type locality of posthumus and found a peculiar karyotype totally different from birunii with a high number of n=ca 85 chromosomes, two of them about 15 and a third about 10 times bigger than the remaining chromosomes. The karyotype of phyllis which occurs sympatrically at Schakuh has a very similar number of chromosomes but a very different karyotype (see previous paragraph). Both species are easy to identify at Schakuh whereas birunii and phyllis can easily be confused with each other (DE LESSE 1959b: 9). birunii Eckweiler & ten Hagen, 1998 This taxon was described as a subspecies of posthumus from Firuzkuh, East Elburs, but due to the totally different karyotype it must be regarded as specifically distinct from posthumus as well as from phyllis. A specimen from the type locality was karyotyped by DE LESSE (1959b) who found a haploid chromosome number of n=10 which is the lowest chromosome number known in Agrodiaetus. Further material of this taxon was karyotyped by DE LESSE (1959b) from Demavend and Kendevan Pass. We are able to confirm a chromosome number of n=10- 11 (or 2n=20-22) from Veresk (near the type locality), from Polur (Demavend), from three populations near Kendevan Pass (Pul-e Zanguleh, Gardaneh-ye Lavashm, Golestanak Nature Reserve) as well as from two new localities: Dizin Pass and Takht-e Suleyman. darius Eckweiler & ten Hagen, 1998 This taxon from Elburs Mts. (Iran) was described as a new species because it was found sympatrically with birunii which was regarded as a subspecies of posthumus by ECKWEILER & TEN HAGEN (1998). However, birunii is specifically distinct from posthumus (see previous paragraph) and thus it cannot be ruled out that darius is a subspecies of posthumus. Both taxa are allopatric and have a similar phenotype. Unfortunately the karyotype of darius remains unknown. We checked a specimen from the type locality (Polur) and from Dizin, but were unable to find typical meiotic divisions. Recently a population attributed to darius was discovered in the Iranian Talysh Mts. (-e Sharqi) near Masuleh and described as a new subspecies (masulensis ten Hagen & Schurian, 2000).

27 Chapter 2 iphigenides-group iphigenides (Staudinger, 1886) The chromosome number of this Central Asian species was found to be n=65-67 in different populations from Kirgizia and Uzbekistan (LUKHTANOV & DANTCHENKO 2002a). No chromosome fixations were available to us. poseidonides (Staudinger, 1886) The chromosome number of this Central Asian species was found to be n=24 in two populations from Kirgizia (LUKHTANOV & DANTCHENKO 2002a). It is one of only two species of Agrodiaetus known to have a stable haploid chromosome number of n=24 which is the modal chromosome number in Lycaenidae. (The other species is turcicus (Koçak, 1977). No chromosome fixations were available to us. dagmara-group dagmara (Grum-Grshimailo, 1888) The karyotype of this Central Asian species is unknown. No chromosome fixations were available to us. erschoffii-group glaucias (Lederer, 1871) The karyotype of this species which has been described from Hajiabad and Schakuh in Elburs Mts. of Iran remains unknown. We did not manage to find this species on our visit at the type localities. erschoffii (Lederer, 1869) DE LESSE (1963a) and LUKHTANOV & DANTCHENKO (2002a) recorded a chromosome number of n=13-15 from Kopetdag (Turkmenia) from where ssp. tekkeanus (Christoph, 1887) was described. We can confirm this chromosome number for nominotypical erschoffii from the type locality Hajiabad (East Elburs Mts., Iran). unknown group The following taxa have been described after 1997 and are not apparently closely related to any of the taxa listed by ECKWEILER & HÄUSER (1997): klausschuriani ten Hagen, 1999 The chromosome number of this newly discovered species was not known until now. We checked six specimens from the type locality Veresk in eastern Elburs Mts. and found a chromosome number of n=56 in three of them. tenhageni Schurian & Eckweiler, 1999 Two chromosome preparations of this newly discovered taxon from Khorasan (Iran) did not reveal typical divisions and therefore the karyotype remains unknown. SCHURIAN & ECKWEILER (1999) could not place it into any group. arasbarani (Carbonell & Naderi, 2000) The chromosome number of this new taxon from (Azarbaijan-e-Sharqi, Iran) remains unknown.

28 New karyological results in Agrodiaetus shahrami Skala, 2001 The chromosome number of this endemic from high altitudes of Zagros Mts. (Zarde-Kuh, 3000-3300m, Bakhtiari, Iran) was determined by V. Lukhtanov (SKALA 2002b) from one preparation with meiotic divisions as n=128-131, which constitutes the highest chromosome number so far known in Agrodiaetus. achaemenes Skala, 2002 Only atypical divisions with a high number of chromosomes were found in two specimens of this taxon which was recently discovered by SKALA (2002b) at very high altitudes (3800- 4000m) in the Iranian Zagros range (Kuh-e-Haft, Bakhtiari). paulae Wiemers & De Prins, 2003 At Ahar Pass (Azarbayjan-e Sharqi, Iran) a new taxon of Agrodiaetus was found (WIEMERS & DE PRINS, in print) whose phenotype has similarities to elbursicus (wing colour of the upperside) and altivagans (reduced wing markings on the underside). Its chromosome number was determined as n=17 from six specimens. At Ahar Pass this taxon was flying together with gorbunovi, which is thought to be closely related to altivagans and has n=20 chromosomes.

Tab. 2 presents a synoptic alphabetical list of the above mentionned taxa with chromosome counts. It also includes further Agrodiaetus species of which chromosome numbers are known, with corresponding references.

Discussion The correct establishment of the karyotype is a very difficult task, especially in the many Agrodiaetus species with high chromosome numbers and very small chromosomes. The only possibility to count them is in the Metaphase I of Meiosis when the bivalents are in a condensed form and well separated from each other. Even then, their size can be below 1µm which is at the absolut limit of resolution for light microscopy.Irregular (apyrene) spermatogenesis (FRIEDLÄNDER 1997) is the dominant form in adult Agrodiaetus, but correct chromosome counts can usually only be made from eupyrene spermatogonia which are much less common. Difficulties in finding “good” metaphase cells with non-overlapping bivalents can lead to erroneous counts. The presence of multivalents and macrochromosomes which can be confused with each other or possibly supernumerary chromosomes further complicates the task (LORKOVIĆ 1990). Not surprisingly, erroneous results have also been published, one unfortunate example being BROWN (1976b) and BROWN & COUTSIS (1978) with totally wrong chromosome counts (COUTSIS et al. (1999); DE PRINS, pers. comm.). Therefore caution should be applied if chromosome counts are ambiguous or cells are of poor quality. Nevertheless, a large part of the variation found in Agrodiaetus species is due to methodical difficulties, especially in the case of the squash techniques which often result in overlapping bivalents which can lead to an underestimate of chromosome numbers. Only recently, a new two-phase method of chromosome preparation was developed (LUKHTANOV & DANTCHENKO 2002a) which overcomes some of these problems, because the original position of chromosomes is less distorted. The disadvantage of this method is that no durable preparations can be made and photographs remain the only evidence. This method was applied in a part of the preparations (since 2002) and often yielded more precise chromosome counts. Despite the methodological difficulties there is some evidence that slight variation of bivalent chromosome numbers (not involving supernumerary chromosomes) within a population and even within individuals may exist. We observed such variation e.g. in A. birunii (n=10-11), A. iphigenia (n=12-14) and A. poseidon (n=19-21). More studies are needed to understand this phenomenon. More common is slight variation of chromosome numbers between populations and it is not known whether these differences are effective as genetic barriers to gene flow.

29 Chapter 2

Tab. 2. Chromosome numbers in Agrodiaetus Agrodiaetus-Taxa n Own results (haploid chromosome numbers) (new= according to Provinces (E=Spain, I=Italy, bold) TR=Turkey, IR=Iran) or references achaemenes Skala, 2002 ? see text actis (Herrich-Schäffer, [1851]) 17 TR: Sivas (17) admetus (Esper, [1783]) 78-80 TR: Antalya (80), Adana (ca78-80) ainsae (Forster, 1961) 108-110 E: Alava (ca108), Huesca (ca108) alcestis (Zerny, 1932) 19-21 IR: Zanjan (19), TR: Hakkari (19), Van (19), Karaman (20), Adana (21) altivagans (Forster, 1956) 18-23 TR: Van (ca21-23), Erzincan (ca21) antidolus (Rebel, 1901) 40-42 TR: Hakkari (42) arasbarani (Carbonell & Naderi, 2000) ? see text ardschira (Brandt, 1938) 113-115 DE LESSE (1961b) aroaniensis (Brown, 1976) 48 see text artvinensis (Carbonell, 1997) 21-22 TR: Artvin (ca21-22) aserbeidschanus (Forster, 1956) 22-23 LUKHTANOV & DANTCHENKO (2002a) baytopi (de Lesse, 1959) 27 TR: Van (ca27-28) bilgini (Dantchenko & Lukhtanov, 2002) 25 LUKHTANOV & DANTCHENKO (2002b) birunii Eckweiler & ten Hagen, 1998 10-11 IR: Tehran (10-11), Mazandaran (10-11) caeruleus (Staudinger, 1871) 20 IR: Golestan (20) carmon (Herrich-Schäffer, [1851]) 81-82 TR: Artvin (ca80) carmonides Eckweiler, 1997 15-16 LUKHTANOV (1989) ciscaucasicus Forster, 1956 16 LUKHTANOV & DANTCHENKO (2002a) cyaneus (Staudinger, 1899) 16-20 TR: Kars (17), Van (18-19) dagestanicus Forster, 1960 40 LUKHTANOV & DANTCHENKO (2002a) dagmara (Grum-Grshimailo, 1888) ? see text dama (Staudinger, 1892) 41 see text damocles (Herrich-Schäffer, [1844]) 23-26 see text damon ([Denis & Schiffermüller], 1775) 45 see text damone (Eversmann, 1841) 66-68 LUKHTANOV et al. (1997) dantchenkoi (Lukhtanov & Wiemers, 2003) 40-42 TR: Van (ca40-42) darius Eckweiler & ten Hagen, 1998 ? see text deebi (Larsen, 1974) 17 LARSEN (1974b) demavendi (Pfeiffer, 1938) 66-76 IR: Tehran (ca70), Azarbaijan (ca70-80); TR: Kars (ca66), Hakkari (ca60-70) dizinensis (Schurian, 1982) 17 IR: Tehran (17) dolus (Hübner, [1823]) 123-125 DE LESSE (1962b, 1966) ectabanensis (De Lesse, 1963) 18 DE LESSE (1964) elbursicus (Forster, 1956) 16-18 IR: Tehran (17-18), Mazandaran (16-18), Zanjan (18) eriwanensis (Forster, 1960) 32-35 LUKHTANOV & DANTCHENKO (2002a) ernesti Eckweiler, 1989 18 TR: Isparta (18) erschoffii (Lederer, 1869) 13-15 IR: Golestan (ca13-14) fabressei (Oberthür, 1910) 90 E: Soria (>75) femininoides (Eckweiler, 1987) 27 IR: Zanjan (27), Ardabil (27) firdussii (Forster, 1956) 24-32 IR: Zanjan (28-29); TR: Erzincan (ca30-32) fulgens (de Sagarra, 1925) 103 E: Tarragona (ca98-103) galloi (Baletto & Toso, 1979) 66 TROIANO & GIRIBALDI (1979) glaucias (Lederer, 1871) ? see text gorbunovi Dantchenko & Lukhtanov, 1994 19-20 IR: Azarbaijan (20), Ardabil (19) guezelmavi Olivier, Puplesiene, van der Poorten, 42 see text De Prins & Wiemers, 1999 haigi (Lukhtanov & Dantchenko, 2002) 21-25 TR: Van (24-26), Hakkari (25-27) hamadanensis (de Lesse, 1959) 21-22 IR: Tehran (21-22) hopfferi (Herrich-Schäffer, [1851]) 15 TR: Malatya (ca15-16), Van (15), Hakkari (15) huberti (Carbonell, 1993) 33-37 TR: Artvin (36), Kars (ca33-34), Bayburt (ca33) humedasae (Toso & Baletto, 1976) 38 I: Aosta (ca38) interjectus (de Lesse, 1960) 29-32 TR: Erzincan (31) iphicarmon Eckweiler & Rose, 1993 29 TR: Isparta (29)

30 New karyological results in Agrodiaetus

Agrodiaetus-Taxa n Own results (haploid chromosome numbers) (new= according to Provinces (E=Spain, I=Italy, bold) TR=Turkey, IR=Iran) or references iphidamon (Staudinger, 1899) 14 IR: Mazandaran (14), Golestan (14) iphigenia (Herrich-Schäffer, [1847]) 12-16 TR: Kars (ca11-14), Erzincan (12), Fethiye (14), Isparta (15) iphigenides (Staudinger, 1886) 65-67 LUKHTANOV & DANTCHENKO (2002a) juldusus (Staudinger, 1886) 67 LUKHTANOV & DANTCHENKO (2002a) kanduli (Lukhtanov & Dantchenko, 2002) 25 TR: Van (25) karindus (Riley, 1921) 68 IR: Kordestan (68) khorasanensis (Carbonell, 2001) 84 see text klausschuriani ten Hagen, 1999 56 IR: Mazandaran (56) kurdistanicus (Forster, 1961) 57-62 TR: Van (56-62) lorestanus Eckweiler, 1997 69-74 see text lycius (Carbonell, 1996) 21-22 TR: Antalya (21-22) maraschi (Forster, 1956) 16 TR: Sivas (16) menalcas (Freyer, [1837]) 85 TR: Fethiye (85), Antalya (>75), Van (85) merhaba De Prins, van der Poorten, Borie, 16-17 TR: Artvin (ca17) Oorschot, Riemis & Coenen, 1991 mithridates (Staudinger, 1878) 21-27 TR: Malatya (23) mofidii (de Lesse, 1963) 34-35 see text morgani (Le Cerf, 1909) 25-27 IR: Kordestan (27) nephohiptamenos (Brown & Coutsis, 1978) ca.90 see text ninae (Forster, 1956) 33-37 TR: Ağrı (34) paulae Wiemers & De Prins, 2003 17 IR: Azarbaijan (17) peilei Bethune-Baker, 1921 39 IR: Kordestan (39) pfeifferi (Brandt, 1938) 106-108 DE LESSE (1961b) phyllides (Staudinger, 1886) ca. 66 LUKHTANOV & DANTCHENKO (2002a) phyllis (Christoph, 1877) 78-82 IR: Tehran (ca76-78), Golestan (ca82-86) pierceae (Lukhtanov & Dantchenko, 2002) 21-22 TR: Van (21) poseidon (Herrich-Schäffer, [1851]) 19-22 TR: Nevşehir (21-22), Sivas (19) poseidonides (Staudinger, 1886) 24 see text posthumus (Christoph, 1877) ca. 85 IR: Golestan (ca85) pseudactis (Forster, 1960) 29 see text pseudoxerxes (Forster, 1956) 15-16 IR: Golestan (15) putnami (Lukhtanov & Dantchenko, 2002) 25-27 TR: Ağrı (25) ripartii (Freyer, 1830) 90 E: Burgos (ca90); TR: Isparta (>80), Adana (ca90), Artvin (ca90), Erzincan (>85), Van (>71) rovshani Dantchenko & Lukhtanov, 1994 ? see text schamil (Dantchenko, 2000) 17 LUKHTANOV & DANTCHENKO (2002a) schuriani (Rose, 1978) 81-82 TR: Kayseri (ca75-80) sekercioglui (Lukhtanov & Dantchenko, 2002) 50 TR: Van (>45) sennanensis (de Lesse, 1959) 28-30 see text sertavulensis (Koçak, 1979) 20 TR: Karaman (20) shahrami Skala, 2001 128-131 see text sigberti Olivier, van der Poorten, Puplesiene & 25-29 TR: Kayseri (25,28) De Prins, 2000 sorkhensis Eckweiler, 2003 ca.45 IR: Khorasan (ca45) surakovi Dantchenko & Lukhtanov, 1994 50 see text tankeri (de Lesse, 1960) 20-21 see text tenhageni Schurian & Eckweiler, 1999 ? see text theresiae Schurian, van Oorschot & van den 63 TR: Adana (59-63) Brink, 1992 transcaspicus (Heyne, [1895]) 52-53 see text turcicola (Koçak, 1977) 19-22 TR: Van (19-22) turcicus (Koçak, 1977) 24 TR: Iğdır (ca25), Erzincan (ca24) valiabadi (Rose & Schurian, 1977) 23 IR: Mazandaran (23) wagneri (Forster, 1956) 16-18 TR: Nevşehir (18, 19-21?) zapvadi (Carbonell, 1993) 18-19 TR: Van (18-19) zarathustra Eckweiler, 1997 ca. 22 IR: Lorestan (ca22)

31 Chapter 3

32 A molecular phylogeny of Agrodiaetus

Chapter 3: A molecular phylogeny of Agrodiaetus inferred from mitochondrial and nuclear DNA sequences

Material and methods

Material Material for this study was collected on trips to Turkey, Iran, Italy, France, Spain and Morocco during the years 1998-2002. While testes or abdomens were removed for karyological analysis and wings cut off for wing pattern analysis, the remaining body was placed in vials with 100% ethanol and later stored at -20°C. The parts of each specimen were coded “MW” and numbered individually. Further material was received from the following collegues:

Code Name Countries Storage of material AD Alexandre Dantchenko (Moscow) Armenia Ethanol CN Clas Naumann (Bonn) Turkey, Iran Ethanol DS Dmitriy Sobanin (Voronezh) Central Asia dried JC John Coutsis (Athens) Greece Ethanol; wings separated JM José Munguira (Madrid) Spain Ethanol; wings separated OK Otakar Kudrna (Schweinfurt) Italy, Spain frozen WE Wolfgang Eckweiler (Frankfurt) mainly Iran dried; abdomens & wings separated

A complete list of available material is found in Appendix 2.

Tab. 3. Material used for DNA extraction

Specimens Populations (COI) Specimens

Genus Spezies DNA COI MO E F I GR TR AR IR UZ TA KI ∑ ND1 Cytb ITS2 Agriades pyrenaicus 1 1 1 1 0 0 1 Agrodiaetus achaemenes 1 1 1 1 0 0 1 Agrodiaetus actinides 2 0 0 0 0 1 Agrodiaetus actis 3 1 1 1 0 0 1 Agrodiaetus admetus 3 2 1 1 2 0 1 1 Agrodiaetus ainsae 3 3 3 3 0 0 2 Agrodiaetus alcestis 7 5 4 1 5 1 0 3 Agrodiaetus altivagans 5 5 2 1 3 0 0 3 Agrodiaetus antidolus 3 3 3 3 1 0 2 Agrodiaetus arasbarani 1 1 1 1 0 0 1 Agrodiaetus aroaniensis 1 1 1 1 0 0 1 Agrodiaetus artvinensis 1 1 1 1 0 0 1 Agrodiaetus barmifiruze 1 - 0 1 0 0 0 Agrodiaetus baytopi 3 3 2 2 0 0 2 Agrodiaetus birunii 7 7 6 6 1 0 5 Agrodiaetus caeruleus 2 2 2 2 1 0 1 Agrodiaetus carmon 3 2 2 2 0 0 1 Agrodiaetus cyaneus 4 4 2 1 3 0 0 3 Agrodiaetus dagmara 1 1 1 1 0 0 0 Agrodiaetus dama 1 1 1 1 0 0 1

33 Chapter 3

Specimens Populations (COI) Specimens

Genus Spezies DNA COI MO E F I GR TR AR IR UZ TA KI ∑ ND1 Cytb ITS2 Agrodiaetus damalis 1 1 1 1 0 0 0 Agrodiaetus damocles 1 - 0 0 0 0 0 Agrodiaetus damon 2 2 1 1 2 0 0 2 Agrodiaetus damone 2 - 0 0 0 0 0 Agrodiaetus dantchenkoi 4 4 2 2 0 0 3 Agrodiaetus darius 1 1 1 1 0 0 1 Agrodiaetus demavendi 12 12 2 3 5 2 0 6 Agrodiaetus dizinensis 1 1 1 1 0 0 1 Agrodiaetus elbursicus 7 7 6 6 1 0 5 Agrodiaetus ernesti 1 1 1 1 0 0 1 Agrodiaetus erschoffii 2 2 2 2 0 0 1 Agrodiaetus fabressei 2 2 2 2 0 0 2 Agrodiaetus femininoides 2 2 2 2 0 0 2 Agrodiaetus firdussii 7 7 3 2 5 0 0 5 Agrodiaetus fulgens 1 1 1 1 0 0 1 Agrodiaetus glaucias 1 1 1 1 0 0 1 Agrodiaetus gorbunovi 4 4 3 3 0 0 4 Agrodiaetus guezelmavi 2 1 1 1 0 0 1 Agrodiaetus hamadanensis 3 3 3 3 1 0 1 Agrodiaetus hopfferi 5 2 2 2 0 0 2 Agrodiaetus huberti 4 4 3 1 4 0 0 3 Agrodiaetus humedasae 3 2 1 1 0 0 2 Agrodiaetus interjectus 1 1 1 1 1 0 1 Agrodiaetus iphicarmon 2 1 1 1 0 1 1 Agrodiaetus iphidamon 3 3 3 3 1 0 2 Agrodiaetus iphigenia 6 5 3 1 4 0 0 2 Agrodiaetus iphigenides 2 2 1 1 2 0 0 1 Agrodiaetus kanduli 1 1 1 1 0 0 1 Agrodiaetus karindus 1 1 1 1 0 0 0 Agrodiaetus khorasanensis 1 1 1 1 0 0 1 Agrodiaetus klausschuriani 2 2 1 1 0 0 1 Agrodiaetus kurdistanicus 2 2 2 2 0 0 1 Agrodiaetus lorestanus 2 1 1 1 0 0 2 Agrodiaetus lycius 3 2 2 2 0 0 1 Agrodiaetus menalcas 5 3 3 3 1 0 1 Agrodiaetus merhaba 2 2 1 1 0 0 1 Agrodiaetus mithridates 2 1 1 1 0 0 1 Agrodiaetus mofidii 2 1 1 1 0 0 1 Agrodiaetus morgani 1 1 1 1 0 0 1 Agrodiaetus nephohiptamenos 2 2 1 1 0 0 1 Agrodiaetus ninae 2 2 1 1 2 0 0 2 Agrodiaetus paulae 2 2 1 1 0 0 1 Agrodiaetus peilei 1 1 1 1 0 0 1 Agrodiaetus phyllides 3 - 0 0 0 0 0 Agrodiaetus phyllis 7 6 2 1 3 6 0 0 4 Agrodiaetus pierceae 3 2 2 2 0 0 2 Agrodiaetus poseidon 6 4 2 2 1 0 3 Agrodiaetus poseidonides 1 1 1 1 0 0 1 Agrodiaetus posthumus 2 2 1 1 0 0 1 Agrodiaetus pseudactis 1 1 1 1 0 0 1 Agrodiaetus pseudoxerxes 2 2 2 2 0 0 1 Agrodiaetus putnami 2 2 2 2 0 0 1

34 A molecular phylogeny of Agrodiaetus

Specimens Populations (COI) Specimens

Genus Spezies DNA COI MO E F I GR TR AR IR UZ TA KI ∑ ND1 Cytb ITS2 Agrodiaetus ripartii 10 9 3 1 4 8 1 0 4 Agrodiaetus rovshani 2 2 2 2 0 0 2 Agrodiaetus schuriani 1 1 1 1 0 0 1 Agrodiaetus sekercioglui 1 1 1 1 0 0 0 Agrodiaetus sennanensis 1 1 1 1 0 0 1 Agrodiaetus sertavulensis 2 1 1 1 0 0 1 Agrodiaetus shahrami 1 1 1 1 0 0 1 Agrodiaetus sigberti 3 2 1 1 0 0 1 Agrodiaetus surakovi 1 1 1 1 0 0 1 Agrodiaetus tankeri 2 2 1 1 0 0 2 Agrodiaetus tenhageni 2 1 1 1 0 0 1 Agrodiaetus theresiae 1 1 1 1 0 0 1 Agrodiaetus transcaspicus 1 - 0 0 0 0 0 Agrodiaetus turcicola 3 2 2 2 0 1 1 Agrodiaetus turcicus 3 3 3 3 1 0 2 Agrodiaetus valiabadi 2 2 2 2 0 0 2 Agrodiaetus wagneri 4 3 2 2 0 0 3 Agrodiaetus zapvadi 3 3 2 2 0 0 2 Agrodiaetus zarathustra 1 1 1 1 0 0 1 agestis 4 4 1 1 1 1 4 0 0 1 Aricia anteros 1 1 1 1 0 0 1 Aricia artaxerxes 3 3 1 1 2 1 0 2 Aricia cramera 1 1 1 1 1 0 0 1 Aricia eumedon 1 1 1 1 0 0 1 Aricia isauricus 1 1 1 1 0 0 1 Aricia torulensis 1 1 1 1 0 0 1 Cacyreus marshalli 1 1 1 1 0 0 1 Celastrina argiolus 2 2 1 1 2 0 0 1 Chilades trochylus 1 1 1 1 0 0 0 Cupido osiris 1 1 1 1 0 0 0 Cyaniris semiargus 2 2 1 1 2 0 0 2 Favonius quercus 1 1 1 1 1 0 1 Iphiclides feisthamelii 2 2 1 1 2 0 0 0 Iphiclides podalirius 1 1 1 1 0 0 1 Kretania eurypilus 1 1 1 1 0 0 1 Lampides boeticus 3 3 1 1 1 3 0 0 3 Leptotes pirithous 1 1 1 1 0 0 0 Lycaena alciphron 2 2 1 1 2 0 0 0 Lycaena asabinus 1 1 1 1 0 0 0 Lycaena candens 1 1 1 1 0 0 0 Lycaena phlaeas 1 1 1 1 0 0 0 Lycaena thersamon 3 1 1 1 0 1 0 Lycaena thetis 1 1 1 1 0 0 0 Lycaena tityrus 1 1 1 1 0 0 0 Lycaena virgaureae 1 1 1 1 0 0 0 Lysandra albicans 2 2 2 2 0 0 2 Lysandra bellargus 6 5 1 1 2 1 5 0 0 4 Lysandra caelestissimus 1 1 1 1 0 0 1 Lysandra coridon 5 5 4 1 5 0 0 4 Lysandra corydonius 5 4 4 4 0 0 2 Lysandra gennargenti 1 1 1 1 0 0 0 Lysandra ossmar 2 2 2 2 0 0 2

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Specimens Populations (COI) Specimens

Genus Spezies DNA COI MO E F I GR TR AR IR UZ TA KI ∑ ND1 Cytb ITS2 Lysandra syriaca 1 1 1 1 1 0 1 Maculinea arion 1 1 1 1 1 0 0 Meleageria daphnis 6 3 2 1 3 1 0 2 Meleageria marcida 2 2 2 2 0 0 1 Neolysandra coelestina 1 1 1 1 0 0 1 Neolysandra corona 1 1 1 1 0 0 1 Neolysandra diana 1 1 1 1 0 0 0 Neolysandra fatima 2 2 1 1 0 0 1 Plebeius argus 1 1 1 1 0 0 1 Plebeius christophi 1 1 1 1 0 0 0 Plebeius cyane 1 - 0 0 0 0 0 Plebeius loewii 3 1 1 1 0 0 1 Plebeius pylaon 1 1 1 1 0 0 1 Polyommatus aedon 1 1 1 1 0 0 1 Polyommatus amandus 2 2 1 1 2 0 0 2 Polyommatus andronicus 1 1 1 1 0 0 1 Polyommatus cornelia 5 3 3 3 0 1 3 Polyommatus dorylas 3 3 1 2 3 1 0 2 Polyommatus eroides 2 2 1 1 2 1 0 2 Polyommatus eros 2 1 1 1 0 0 0 Polyommatus escheri 2 2 1 1 2 0 0 1 Polyommatus icarus 7 5 1 1 1 1 1 5 1 0 4 Polyommatus menelaos 2 2 2 2 0 0 2 Polyommatus myrrhinus 3 3 2 2 1 0 2 Polyommatus thersites 6 4 1 2 1 4 1 0 2 Pseudophilotes abencerragus 1 1 1 1 0 0 0 Pseudophilotes vicrama 1 1 1 1 0 0 0 Satyrium esculi 1 1 1 1 0 0 0 Satyrium hyrcanicum 1 1 1 1 0 0 1 Satyrium myrtale 2 1 1 1 0 0 0 Tarucus theophrastus 1 1 1 1 0 0 1 endymion 2 1 1 1 0 0 0 Vacciniina alcedo 2 2 1 1 2 0 0 1 Vacciniina morgianus 1 1 1 1 0 0 1 Zizeeria knysna 1 1 1 1 0 0 0 Sum ∑ 370 312 12 27 2 4 13 126 8 81 1 2 1 277 24 5 209 • DNA = Number of specimens of which DNA was extracted (- = without result). • COI, ND1, Cytb, ITS2 = Number of specimens sequenced for these genes. • Populations (COI) = Number of populations sequenced for COI from Morocco (MO), Spain (E), France (F), Italy (I), Greece (GR), Turkey (TR), Armenia (AR), Iran (IR), Uzbekistan (UZ), Tajikistan (TA) and Kirgizia (KI) Nomenclature according to HESSELBARTH et al. (1995), TOLMAN & LEWINGTON (1997) & KUDRNA (2002)

Selection of taxa Material used for DNA extraction is listed in Tab. 3, sorted alphabetically according to genus and species names. Altogether 370 specimens from 158 butterfly taxa were used for DNA extraction. These include 90 ingroup taxa (subgenus Agrodiaetus of genus Polyommatus) covering 63 of the 83 species rank taxa listed by ECKWEILER & HÄUSER (1997) from all species groups, and an additional 17 species which have been described since. Of the outgroup taxa 27 are closely related taxa of the genus Polyommatus (subgenera Cyaniris, Lysandra, Meleageria, Neolysandra & Polyommatus), 16 are from the same subtribe Polyommatiti (e.g. the genera/subgenera Agriades, Aricia, Chilades, Kretania, Plebeius,

36 A molecular phylogeny of Agrodiaetus

Vacciniina), 11 from other subtribes within the tribe (Glaucopsychiti: Maculinea, Turanana; Scolitantiditi: Pseudophilotes; Celastriniti: Celastrina; Everiti: Cupido; Zizeeriti: Zizeeria; Taruciti: Tarucus; Leptotiti: Leptotes; Lampiditi: Lampides; Cacyreus), 12 from other tribes (Eumaeini: Satyrium; Theclini: Favonius; Lycaenini: Lycaena) of Lycaenidae and two taxa of another family of Papilionoidea (Papilionidae: Iphiclides). This selection covers all Polyommatus subgenera, all Polyommatini subtribes and all but one Lycaenidae tribus known to occur in Anatolia, the centre of Agrodiaetus radiation (nomenclature according to HESSELBARTH et al., 1995).

DNA extraction In most cases parts of the thorax were cut into slices and subsequently used for DNA extraction, but in a few cases only dried legs were available and thus used. DNA was extracted with QIAGEN® DNeasy Tissue Kit according to the manufacturer’s protocol for mouse tail tissue. This method turned out to give the most consistent results and was the most economical compared to other methods tried out in the beginning: • phenol-chloroform extraction (time-consuming method using hazardous chemicals) • CTAB buffer based chloroform-isoamylalcohol extraction (time-consuming method yielding a high proportion of degraded DNA) TM • BIORAD® Quantum Prep® Aqua Pure Genomic DNA Tissue Kit (fast salt- precipitation method prone to contamination with butterfly scales) Results of DNA extraction were checked with Agarose gel electrophoresis.

Selection of genes Mitochondrial genes are especially well suited for phylogenetic studies of closely related species, because of the faster rate of evolution compared to nuclear genes (1-2 times faster in ; HOY, 1994) and the shorter coalescence time due to their smaller effective population size (a quarter of an autosomal locus).

Cytochrome Oxidase I was chosen because this gene has been used successfully in phylogenetic studies of closely related Lepidoptera species, including butterflies (BELTRAN et al. (2002); BROWER (1994); CATERINO & SPERLING (1999); MONTEIRO & PIERCE (2001); RAND et al. (2000); WAHLBERG & ZIMMERMANN (2000)). Initially experiments were also carried out with cytochrome b which has only rarely been used in Lepidoptera (but recent papers on butterflies were published by AAGARD et al. (2002) and TORRES et al. (2001)). Cytochrome Oxidase I is known to evolve relatively slowly compared to other mitochondrial genes, e.g. of the ND family. Therefore part of the mitochondrial ND1 gene was also sequenced for a selection of taxa. In Lepidoptera this gene has been used in phylogenetic studies of Nymphalidae (AUBERT et al., 1999; MARTIN et al., 2000) and Geometridae (ABRAHAM et al., 2001).

Because “gene trees and species trees are not the same” (NICHOLS, 2001), conclusions drawn from a single locus can be misleading. This is especially true for mitochondrial DNA which is maternally inherited. Agrodiaetus species are thought to have evolved only recently and gene flow might still exist due to incomplete speciation and hybridization events. Hybridization can be detected if gene trees based on mtDNA and nuclear DNA are compared to each other. Therefore the internal transcribed spacer (ITS-2), a non-coding region between the nuclear ribosomal genes 5.8s and 28s was chosen for a comparative study. ITS-2 is a very variable region, while the flanking genes are very conservative, so that universal primers can be employed. ITS-2 has not been used in published phylogenetic studies of Lepidoptera yet, but was used successfully for studying the phylogeny of closely related species in other insect orders like Diptera, Hymenoptera, Coleoptera, Odonata (WEEKERS et al. 2001) and Blattodea.

37 Chapter 3

PCR and sequencing Amplification of DNA was conducted using the polymerase chain reaction (PCR). The reaction mixture (for a total reaction volume of 25µl) included: ddH20 16,80 µl 10xPCR II buffer (without MgCl2) 2,50 µl MgCl2 25mM 3,20 µl dNTP-Mix 2mM 0,50 µl Primer 1 20pm 0,375 µl Primer 2 20pm 0,375 µl Taq Polymerase 0,25 µl DNA 1,00 µl

The primers used are shown in Tab. 4.

Tab. 4. Primers Alias Designation* Sequence (5’ to 3’) References k698 TY-J-1460 TAC AAT TTA TCG CCT AAA CTT CAG CC CATERINO & SPERLING (1999) Nancy C1-N-2192 (CCC) GGT AAA ATT AAA ATA TAA ACT TC CATERINO & SPERLING (1999) Faw ND1 N1-J-12314 TAG AAT TAG AAG ATC AAC CAG C PASHLEY & KE (1992) 16sb LR-N-12866 ACA TGA TCT GAG TTC AAA CCG G VOGLER & DESALLE (1993) cb1l CB-J-10612 CCA TCC AAC ATC TCA GCA TGA TGA AA PALUMBI (1996) cb2-h CB-N-10920 CCC TCA GAA TGA TAT TTG TCC TCA PALUMBI (1996) ITS-3 GCA TCG ATG AAG AAC GCA GC WHITE et al. (1990) ITS-4 TCC TCC GCT TAT TGA TAT GC WHITE et al. (1990) * according to the positions in Drosophila yakuba (see CLARY & WOLSTENHOLME, 1985)

Fig. 28. The mitochondrial genome of Drosophila yakuba (copied from GenBank). Hatched sections of the genes Cytochrome Oxidase I (COX1), Cytochrome b (CYTB) and NADH dehydrogenase subunit 1 (ND1) were sequenced.

38 A molecular phylogeny of Agrodiaetus

For the mitochondrial genome, primers k698 and Nancy were used to amplify a 700bp fragment of cytochrome oxidase subunit 1 (COI), primers Faw ND1 and 16sb were used to amplify a 530bp fragment of NADH dehydrogenase subunit 1 (ND1) and 16s rRNA and primers cb1l and cb2-h amplified a 284bp fragment of cytochrome b gene (Cyt b).

Primers ITS-3 and ITS-4 were used to amplify the complete internal transcribed spacer 2 (ITS-2) in the nuclear genome together with fractions of the flanking regions of the genes coding for the ribosomal subunits 5.8s and 28s.

COI was sequenced for 150 taxa and the number of specimens and populations for each country are shown in Tab. 3. ITS-2 was sequenced for 125 taxa only because distantly related outgroups turned out to be non-alignable. ND1 was only sequenced for 23 representative taxa and Cytb for a mere 5 taxa.

PCR was conducted on thermal cyclers from BIOMETRA® (models UNO II or T-GRADIENT) or ABI BIOSYSTEMS® (model GENEAMP® PCR-System 2700) using the following profiles: COI & Cyt b: initial 4 minutes denaturation at 94°C and 35 cycles of 30 seconds denaturation at 94°C, 30 seconds annealing at 55°C and 1 minute extension at 72°C. ND1: Touchdown PCR with initial 3 minutes denaturation at 94°C, 15 cycles of 30 seconds denaturation at 92°C, 45 seconds annealing at 60°C with a 1°C decrease each cycle down to 45°C and 45 seconds extension at 72°C, followed by 20 cycles of 30 seconds denaturation at 92°C, 30 seconds annealing at 50°C and 45 seconds extension at 72°C. ITS-2: initial 3 minutes denaturation at 94°C and 36 cycles of 35 seconds denaturation at 94°C, 30 seconds annealing at 48°C and 1 minute extension at 72°C.

PCR products were purified using purification kits from PROMEGA® or SIGMA® and checked with agarose gel electrophoresis before and after purification.

Cycle sequencing was carried out on BIOMETRA® T-GRADIENT or ABI BIOSYSTEMS® GENEAMP® PCR-System 2700 thermal cyclers using sequencing kits of MWG BIOTECH® (for LI-COR® automated sequencer) or ABI BIOSYSTEMS® (for ABI® 377 automated sequencer) according to the manufacturers’ protocols and with the following cycling times: initial 2 minutes denaturation at 95°C and 35 cycles of 15 seconds denaturation at 95°C, 15 seconds annealing at 49°C and 15 seconds extension at 70°C. Primers used were the same as for the PCR reactions for the ABI (primer 1 was used for forward and primer 2 for independent reverse sequencing), but for LI-COR truncated and labelled primers were used with 3 bases cut off at the 5’ end and labelled with IRD-800. For ABI sequencing the products were cleaned using an ethanol precipitation protocol.

Electrophoresis of sequencing reaction products was carried out on LI-COR® or ABI® 377 automated sequencers using the manufacturer’s protocols.

Sequence alignment Alignment of the mitochondrial gene sequences COI, ND1 and Cyt b was done by hand in comparison with sequences obtained from GenBank. Some of these were also included in the analysis as outgroups (Tab. 5):

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Tab. 5. GeneBank sequences used as outgroups Gene Family Species GenBank accession number COI Lycaenidae Euphilotes bernardino AF170864 COI Nymphalidae Coenonympha tullia AF170860 COI Papilionidae Papilio machaon AF044006 COI Papilionidae Iphiclides podalirius AF170873 COI Pieridae Colias eurytheme AF044007 COI Riodinidae Apodemia mormo AF170863 COI,ND1,Cytb Drosophilidae Drosophila yakuba NC001322 Cytb Pyralidae Ostrinia nubilalis AF442957 ND1 Libytheidae Libytheana bachmanii U32463 ND1 Lycaenidae Celastrina ladon U32455 ND1 Lycaenidae Incisalia spec. U32461 ND1 Nymphalidae Danaus plexippus U32457 ND1 Papilionidae Papilio machaon AJ224107 ND1 Pieridae Phoebis sennae U25875 ND1 Riodinidae Apodemia mormo U32452

Alignment was unambiguous and straightforward, due to the conserved amino acid coding triplet pattern, and apart from one insertion event (triplet) in one Cyt b sequence, no indels were found. The 16s rRNA sections of the ND1-16s sequences were discarded from the analysis due to poor quality and alignment ambiguities.

Alignment of ITS-2 sequences was done with the help of ClustalX (THOMPSON et al., 1997), but manual re-editing was required to improve the alignment. Indel events are very common in ITS-2 regions which makes alignment difficult in distantly related taxa. Therefore ITS-2 has only been used in closely related species and genera, although the discovery of secondary structures for its transcripts might make it more versatile for phylogenic studies at deeper taxonomic levels in future (COLEMAN, 2003).

Alignment of ingroup sequences was mostly straightforward due to their similarity, but distant outgroups had so many indels that they had to be excluded from the analysis and (with one exception) only sequences of the subtribus Polyommatiti and one outgroup taxon (Tarucus theophrastus) were retained to avoid the exclusion of variable portions deemed to be important for phylogenetic reconstruction of Polyommatus and closely related genera. Only one section of ITS-2 had to be exluded from the analysis due to alignment problems (characters 566-588). Because of alignment problems no ITS2-sequences from GenBank were included in the analysis. (The only Lepidopteran ITS2-sequences in GenBank are from the butterfly family Papilionidae).

Choice of methods for phylogenetic inference

Phylogenies are usually presented in a tree and the choice of tree-building methods is a subject of heated debate between proponents of these methods. Even though different methods differ in their efficiency to reconstruct the true tree, their success depends on so many conditions that it is hardly possible to tell which method is appropriate in real data sets. Most often, the decision will have to be pragmatic and a comparison of different methods will

40 A molecular phylogeny of Agrodiaetus show that differences are confined to branches which have weak statistical support (PAGE & HOLMES 1998; NEI & KUMAR 2000). Distance methods such as UPGMA or Neighbour Joining are very fast, but less robust than other methods if underlying assumptions are violated. They also discard a lot of information because they calculate trees from distances matrices. Therefore they are not shown here.

Maximum parsimony (MP) is one of the most widely used tree-building methods in phylogenetic systematics. It uses all available information from nucleotide data to infer the shortest tree. However, it is difficult to test the robustness of the tree. The most commonly used method is the bootstrap, which generates pseudoreplicates from sequence data, but bootstrap values are difficult to interprete and values are often unreasonably low in data sets of closely related species with low divergence. Despite of these drawbacks, MP-trees are presented here.

Maximum likelihood (ML) is a more general approach than MP and requires an explicit model of evolution. It is computationally very expensive, especially if bootstrap values need to be computed, and therefore currently unsuitable for large data sets.

The Bayesian approach has only recently been applied for phylogenetic reconstruction but already proved to be very efficient. It is related to ML but much faster to compute. Its beauty lies in the statistical properties which allow the calculation of the robustness of clades very well. Therefore this method was chosen for most discussions of phylogenetic relationships.

Tree-building methods are only effective in inferring hierarchical relationships but they produce unresolved trees if recombination occurs between taxa, e.g. due to hybridization events, or if ancestral haplotypes survive in extant taxa. Such events have to be expected in radiations of closely related species such as Agrodiaetus. Network approaches are more appropriate in these cases although they have only rarely been used so far in phylogenetic systematics (POSADA & CRANDALL 2001). Different methods exist to infer networks. The Statistical parsimony approach, which is chosen here, connects haplotypes in a stepwise fashion emphasizing the similarities between them.

Sequence comparisons and phylogenetic inference

Sequence statistics (base composition, number of variable and parsimony-informative sites) were calculated with MEGA Version 2.1 (KUMAR et al., 2001).

A saturation analysis for the different positions of COI was conducted with the help of MEGA (KUMAR et al., 1995) plotting transitions against transversions (with and without AT- transversions) and AT-transversions against CG-transversions.

COI and ITS-2 datasets were tested for incongruence with the incongruence length difference test (FARRIS et al., 1995) implemented in PAUP 4.0 beta10 (SWOFFORD, 1998) as the “partition homogeneity test”.

A Bayesian phylogeny reconstruction was done with the computer program MrBayes (HUELSENBECK & RONQUIST 2001). Codon positions of the COI and ND1 datasets were partitioned to allow them to have different rates of evolution and in the combined analyses genes were also partitioned to allow for different rates. The standard 4by4 nucleotide substitution model was applied, all substitution types were allowed to be different (General

41 Chapter 3

Time Reversible model) and the rates of variable sites were drawn from a gamma distribution (INVGAMMA option) with the continuous gamma distribution broken into four categories of equal weight. The Metropolis-coupled Markov chain Monte Carlo simulation (MCMCMC or (MC)3) was run under standard settings with 1,000,000 generations and every 100th tree was saved. Trees obtained before stationarity of the chain was achieved were discarded (burn in). The programme calculates branch lenghths which were multiplied with the factor 1,000,000 to enable input into the programme TreeExplorer (written by Koichiro Tamura, Tokyo). The posterior clade probabilities can be used to infer the confidence of each clade, and these values are placed at the nodes. The tree obtained from the COI data set was rooted with Apodemia mormo (C. & R. Felder, 1859) as outgroup. This species is a representative of the family Riodinidae which is appears to be the sister family of Lycaenidae according to molecular results (CAMPBELL et al. 2000) and traditional classification (ELIOT 1973). The trees from the ITS2- and the combined data sets were rooted with Tarucus theophrastus as outgroup. This species belongs to the subtribus Taruciti which turned out to be the subtribus most closely related to Polyommatiti in the COI analysis.

For a phylogenetic reconstruction under a maximum parsimony approach a heuristic search for the most parsimonious tree (HSEARCH) was carried out with PAUP after the exclusion of uninformative characters (PAUP command: EXCLUDE UNINF) under the following settings: • Maximum number of trees per replicate restricted to 100 (SET MAXTREES=100) • Random addition of sequences (ADDSEQ=RANDOM) • Branch swapping method “Tree Bisection Reconnection” (SWAP=TBR) Due to a bug in the current PAUP version, only one replicate is calculated if the number of trees found is higher than “MAXTREES”, even if the number of replicates is set to a higher value. Therefore the PAUP analysis was repeated 1000 times to receive 1000 replicates. The restriction on the maximum number of trees was necessary to prevent the calculation of millions of rearrangements in the tips of the tree. Experiments with higher limits than 100 were conducted but did not improve the results.

This analysis was repeated for each gene separately and with the different combinations of ITS-2, COI and ND1.

Consensus trees (strict, semistrict, majority rule & Adams consensus) were calculated from the shortest trees of each replicate and from the shortest trees of all replicates together.

Bootstrap analyses were done with PAUP with the heuristic search option employed, random addition of sequences, TBR branch swapping and 1000 replicates.

Gene genealogies of closely related species groups were estimated with statistical parsimony as described by TEMPLETON et al. (1992) and implemented in the computer program TCS (CLEMENT et al., 2000). Because of the sensitivity of this approach to missing character information the analysis of the ITS-2 data set was repeated excluding 8 sequences with more than 10% missing characters (AD98001, AD98018, MW00110, MW00269, MW00316, MW01001, MW01025, MW99292). This data set was analyzed in two ways: counting gaps as missing character information and for the ingroup Agrodiaetus also counting gaps as 5th character.

42 A molecular phylogeny of Agrodiaetus

Results

Sequences and alignment The following mitochondrial gene sequences were obtained and aligned successfully: 690bp of Cytochrome Oxidase I (COI), 275bp of Cytochrome b (Cytb) and 230bp of NADH dehydrogenase subunit 1 (ND1), homologous to the following sites of Drosophila yakuba mitochondrial genome (CLARY & WOLSTENHOLME 1985): 1474-2163, 10650-10924 and 12391-12620, respectively.

Of the nuclear genome, a 120bp fraction of the 5.8s ribosomal subunit and the complete internal transcribed spacer 2 (ITS-2), an aligned 672bp, were aligned successfully. ITS-2 consists of several conservative sections with highly variable sections in between, including some sections with repetitive motifs similar to microsatellites. Indels are common and length variation in ITS-2 turned out to be considerable within Lycaenidae. The shortest sequence was found in Tarucus theophrastus Fabricius, 1793 (MW02025) with 369bp while Lysandra corydonius Herrich-Schäffer, [1852] (MW99514) had the longest with 521bp. The long sequences of many species of the subgenus Lysandra were mainly due to repetitive insertions at position 19 in the aligned sequences made up of the two motifs “GTC” (repeated up to 9 times) and “CACGGCG” (repeated up to 3 times). Within the other Polyommatus subgenera, length variation was much less pronounced, with sequences varying between 439bp in Polyommatus cornelia Freyer, [1850] and 484bp in Polyommatus thersites Cantener, [1835]. Variation within Agrodiaetus was similar. Most sequences were between 443bp (Agrodiaetus maraschi Forster, 1956) and 471bp (Agrodiaetus gorbunovi Dantchenko & Lukhtanov, 1994 & A. wagneri Forster, 1956). Only Agrodiaetus dama Staudinger, 1892 had a longer sequence of 482bp. In the distantly related outgroups (which were not included in the further analysis due to alignment problems) the longest ITS-2 sequence was found in Iphiclides podalirius L., 1758 (JC02001) from the family Papilionidae with 536bp. In this family an even more extreme length variation has been found in the genus , (sequences taken from GenBank): 687bp in Luehdorfia chinensis Leech, 1893 (AB071925) compared to only 362bp in Luehdorfia japonica Leech, 1889 (AB071910).

Base composition and sequence divergence Base composition in the insect mitochondrial genome is known to be heavily AT-biased, especially in 3rd codon positions, while the nuclear genome has a more homogenous base ratio. This also holds for the investigated gene sections in Lycaenidae (Tab. 6). In ITS-2 no base heterogeneity is found in Iphiclides podalirius L. (Papilionidae), but in Lycaenidae a slight deficiency in Adenin (A) is covered by a surplus in Guanin (G). This base composition is especially pronounced in the whole tribus Polyommatini and is remarkably constant in all taxa investigated. In the mitochondrial genes, the AT bias ranges from between 59% and 73% in 1st and 2nd codon positions (lowest in COI and highest in ND1) to 90% in 3rd codon positions of all three genes. Guanin (G) is especially rare in 3rd codon positions of genes on the (+)strand (COI, Cytb) while Cytosin (C) is rarest in ND1 which is on the (–)strand. The AT bias at 3rd codon positions of COI is less pronounced in the genus Polyommatus (90%) compared to other genera of the family Lycaenidae (94%). Thus the AT bias in Polyommatus is similar to other insect groups, but not as extreme as in the genus Arhopala (Lycaenidae) with average values over 95% (MEGENS 2002).

43 Chapter 3

Tab. 6. Base composition (Nucleotide frequencies with standard errors) Gene T C A G COI (Polyommatus) 37.22 0.65 15.48 0.60 32.39 0.52 14.90 0.44 1st Position 28.70 0.87 13.91 0.83 32.27 0.59 25.13 0.66 2nd Position 42.50 0.40 25.10 0.37 15.92 0.45 16.49 0.33 3rd Position 40.49 1.58 7.42 1.25 49.03 1.34 3.07 1.06 COI (other Lycaenidae) 39.39 1.16 13.84 0.67 35.52 0.98 14.22 0.43 1st Position 29.78 1.09 12.72 0.92 32.88 0.84 24.62 0.77 2nd Position 42.65 0.56 24.67 0.44 16.20 0.51 16.48 0.39 3rd Position 45.78 2.83 4.11 1.52 48.54 2.77 1.56 0.90 ND-1 (Polyommatus) 45.31 0.53 9.40 0.51 32.98 0.79 12.30 0.70 1st Position 41.20 0.76 11.06 0.78 31.49 1.22 16.34 1.10 2nd Position 46.76 0.61 15.57 0.50 25.99 0.33 11.78 0.28 3rd Position 48.10 1.18 1.48 1.22 41.67 2.70 8.75 2.77 Cytb (Lycaenidae) 43.18 0.67 10.88 0.51 32.20 0.57 13.83 0.68 1st Position 37.55 0.19 7.68 0.78 32.90 0.42 21.93 0.30 2nd Position 45.33 0.62 16.88 0.64 21.00 0.20 16.88 0.35 3rd Position 46.68 2.32 8.05 1.59 42.78 2.01 2.50 2.13 ITS-2 (Polyommatini) 23.15 0.58 25.95 0.63 19.67 0.86 31.23 0.69 (Polyommatus) 23.19 0.58 25.95 0.63 19.64 0.82 31.22 0.69 (other Polyommatini) 22.60 0.37 26.02 0.64 20.10 1.19 31.27 0.70 (other Lycaenidae) 23.91 1.86 25.09 1.62 22.83 1.62 28.17 1.34 (Iphiclides podalirius) 26.8 24.7 22.2 26.3

Tab. 7. Variable and parsimony-informative sites in single gene studies Gene Taxa Sequences nucleotides amino acids variable parsimony- variable parsimony- sites informative sites informative COI Papilionoidea 318 326 47.2% 274 39.7% 74 32.2% 38 16.5% Lycaenidae 310 311 45.1% 260 37.7% 68 29.6% 34 14.8% Polyommatini 297 300 43.5% 249 36.1% 63 27.4% 32 13.9% Polyommatiti 282 284 41.2% 237 34.3% 56 24.3% 32 13.9% Polyommatus 261 267 38.7% 219 31.7% 49 21.3% 29 12.6% Agrodiaetus 200 223 32.3% 172 24.9% 37 16.0% 17 7.4% ND1 Lycaenidae 25 76 33.0% 45 19.6% 18 23.7% 8 10.5% Polyommatus 22 58 25.2% 33 14.3% 10 13.2% 6 7.9% Agrodiaetus 15 38 16.5% 23 10.0% 8 10.5% 2 2.6% Cytb Lycaenidae 6 53 19.2% 15 5.4% 8 8.7% 3 3.3% ITS2 Polyommatiti 199 257 39.6% 154 23.7% (gaps= Polyommatus 185 238 36.7% 132 20.3% missing) Agrodiaetus 138 159 24.5% 76 11.7% 5.8s Polyommatini 200 9 7.5% 4 3.3%

The number and percentage of variable and parsimony-informative sites for each gene is given in Tab. 7. Figures are for different taxonomic levels, from the superfamily (Papilionoidea) down to the ingroup (subgenus Agrodiaetus). The variation in amino acids of mitochondrial genes is much lower than the variation in nucleotide sites because most variation is found in synonymous sites (especially 3rd codon positions). The section of 5.8s rRNA turned out to be almost invariable in Lycaenidae and was therefore lumped with ITS-2 44 A molecular phylogeny of Agrodiaetus in the phylogenetic analysis. The comparison of sequence variation in the combined gene analyses (Tab. 8) reveals that Cytb and ND1 evolve faster than COI. This difference is almost entirely due to the higher number of nonsynonymous substitutions.

Tab. 8. Variable and parsimony-informative sites in combined gene studies Gene Taxa Sequences nucleotides amino acids variable parsimony- variable parsimony- sites informative sites informative ND1 Lycaenidae 25 76 33.0% 45 19.6% 18 23.7% 8 10.5% COI Lycaenidae 25 200 29.0% 116 16.8% 30 13.0% 9 3.9% Cytb Lycaenidae 6 53 19.2% 15 5.4% 8 8.7% 3 3.3% COI Lycaenidae 6 109 15.8% 24 3.5% 12 5.2% 1 0.4% COI Polyommatini 200 276 40.0% 218 31.6% 51 22.2% 27 11.7% ITS-2 Polyommatini 200 277 42.7% 156 24.0% 5.8s Polyommatini 200 9 7.5% 4 3.3%

The saturation analysis showed no saturation in 1st and 2nd codon positions of COI within Lycaenidae, because the number of transitions is linearly increasing with the number of transversions (Fig. 29). In the 3rd codon positions transitions were saturated in ingroup- outgroup comparisons (Fig. 30) but there is still a high number of transversions. Due to the AT bias most of these are AT-transversions as can be seen from Fig. 31 in which AT- transversions were excluded. Fig. 32 displays the ratio of AT-transversions against CG- transversions. The steep linear increase indicates that AT-transversions are not saturated in ingroup-outgroup comparisons.

Fig. 29. Ratio of transitions (ti) and transversions (tv) in 1st and 2nd codon positions of COI in all investigated taxa

45 Chapter 3

Fig. 30. Ratio of transitions (ti) and transversions (tv) in 3rd codon positions of COI within Agrodiaetus (ingroup) and compared to the outgroups (other taxa).

Fig. 31. Ratio of transitions and transversions excluding AT-transversions in 3rd codon positions of COI within Agrodiaetus (ingroup) and compared to the outgroups (other taxa).

46 A molecular phylogeny of Agrodiaetus

Fig. 32. Ratio of AT-transversions and other transversions in 3rd codon positions of COI within Agrodiaetus (ingroup) and compared to the outgroups (other taxa).

The partition homogeneity test revealed a significant incongruence between the COI and ITS-2 data sets (p<0.001). Despite this result, both data sets were combined for a total evidence approach, but they were also analyzed separately to figure out the incongruence in an effort to elucidate deviations of the gene trees from the true species tree.

47 Chapter 3

Bayesian inference of phylogeny The phylogenetic trees inferred from the Bayesian approach are represented in Fig. 33 - Fig. 46 for the COI data set, Fig. 47 & Fig. 48 for ITS-2 and Fig. 49 - Fig. 57 for the combined data set. Only the COI data set can be used to infer the phylogeny above the level of the subtribus (Polyommatiti).

98 100 Poly ommatini Theclini 61 Eumaeini 100 Lycaenini 100 Riodinidae

500000 Fig. 33. MrBayes, COI, Lycaenidae (relationships of tribes)

The basal splits in the COI tree are on the tribus level (Fig. 33), separating the monophyletic Lycaena (Lycaenini), Satyrium (Eumaeini) and Favonius (Theclini) from the other taxa of tribus Polyommatini). Relationships within the genus Lycaena are highly resolved and shown in Fig. 34. The most basal taxa (thetis, asabinus, thersamon) belong to the thersamon-group (HESSELBARTH et al. 1995).

98 Poly ommatini 100 Favonius quercus Satyrium hyrcanicum 61 100 Satyrium esculi 84 Satyrium myrtale Lycaena thetis Lycaena asabinus 100 Lycaena thersamon 98 93 Lycaena candens 99 Lycaena virgaureae 57 Lycaena tityrus 100 100 Lycaena alciphron heracleana Lycaena alciphron melibaeus 98 Lycaena phlaeas Apodemia mormo

500000 Fig. 34. MrBayes, COI, Lycaenidae (excl. Polyommatini)

Within Polyommatini the splits follow the level of morphologically defined subtribes but their relationships are not well resolved (Fig. 35). 100 91 Poly ommatiti Taruciti 88 Celastriniti 98100 95 Scolit100antiditi 92 Glaucopsychiti 32 98 51 Lampiditi 100 Cacyreus 98 Ev er iti 100 Leptotiti 83 Zizeeriti Theclini 61 Eumaeini 100 Lycaenini 100 Riodinidae

500000 Fig. 35. MrBayes, COI, Lycaenidae (subtribes)

48 A molecular phylogeny of Agrodiaetus

The subtribus most closely related to Polyommatiti are Taruciti, Celastriniti, Scolitantiditi, Glaucopsychiti and Lampiditi (Fig. 36, Fig. 37). 100 Poly ommatiti 100 Celastriniti 95 Taruciti 98 100 Scolitantiditi Glaucopsychiti 98 98 Lampiditi 100 Cacyreus Ev er iti 100 Leptotiti Zizeeriti Theclini Eumaeini 100 Lycaenini 100 Riodinidae

Fig. 36. MrBayes, COI, Lycaenidae (subtribes), condensed tree at 95% confidence limit

100 Poly ommatiti 91 100 Celastrina argiolus mauretanica Celastrina argiolus argiolus 88 98 Tarucus theophrastus 100 Pseudophilotes abencerragus Pseudophilotes vicrama 95 92 Maculinea arion 98 Turanana endymion 32 Lampides boeticus (Spain) 100 Lampides boeticus (Morocco) 51 83 Lampides boeticus (Turkey) 98 Cacyreus marshalli Cupido osiris 100 Leptotes pirithous 83 Zizeeria knysna Theclini 61 Eumaeini 100 Lycaenini 100 Riodinidae

500000 Fig. 37. MrBayes, COI, Polyommatini excl. Polyommatiti

Within Polyommatiti, the genus Chilades and the North American Euphilotes appear as the most basal taxa in the tree and the other taxa of the two genera Polyommatus and Plebeius together form a monophyletic unit (Fig. 38, Fig. 39). 100 Polyommatus 40 Plebeius (Agriades) pyrenaicus 88 Plebeius (Aricia) anteros Plebeius (Aricia) cramera 72 28 Plebeius (Aricia) agestis complex 99 Plebeius (Aricia) torulensis 100 100 Plebeius (Aricia) isauricus 42 Plebeius (Aricia) eumedon 100 Plebeius (Plebejides) pylaon 94 96 Plebeius (Kretania) eurypilus 37 24 Plebeius (Vacciniina) alcedo (Iran) 100 Plebeius (Vacciniina) alcedo (Turkey) Plebeius (Vacciniina) morgianus 100 65 Plebeius (Plebejidea) loewii Cyaniris semiargus persica 32 100 Cyaniris semiargus maroccana Plebeius (Plebeius) argus 100 Plebeius (Plebeius) christophi Euphilotes bernardino Chilades trochylus

200000 Fig. 38. MrBayes, COI, Polyommatiti

49 Chapter 3

100 Polyommatus Plebeius (Agriades) pyrenaicus 88 Plebeius (Aricia) anteros 72 Plebeius (Aricia) cramera Plebeius (Aricia) agestis complex 99 Plebeius (Aricia) torulensis 100 100 Plebeius (Aricia) isauricus Plebeius (Aricia) eumedon 94 100 Plebeius (Plebejides) pylaon 96 Plebeius (Kretania) eurypilus Plebeius (Vacciniina) alcedo (Iran) 100 Plebeius (Vacciniina) alcedo (Turkey) Plebeius (Vacciniina) morgianus 100 65 Plebeius (Plebejidea) loewii Cyaniris semiargus persica 100 Cyaniris semiargus maroccana Plebeius (Plebeius) argus 100 Plebeius (Plebeius) christophi Euphilotes bernardino Chilades trochylus

Fig. 39. MrBayes, COI, Polyommatiti, condensed (50% level)

While the genus Polyommatus (excluding the subgenus Cyaniris) also forms a monophyletic unit, the monophyly of the genus Plebeius is not supported and the relationships of most of its taxa are not well resolved. One exception is the subgenus Aricia (excluding A. eumedon) whose monophyly and taxa relationships appear quite well resolved. This genus includes the -complex, a group of closely related taxa (agestis, artaxerxes, montensis, Fig. 40), but Aricia cramera (Eschscholtz, 1821) which is also thought to belong to this complex and phenotypically very similar, appears very distant genetically. 100 Aricia montensis (Spain) 83 Aricia montensis (Morocco) Aricia artaxerxes (Greece) Aricia agestis (Turkey) Aricia artaxerxes (Greece) 100 Aricia agestis (Greece) Aricia agestis (Iran)

Fig. 40. MrBayes, COI, Aricia agestis complex, condensed (50% level)

99 100 Agrodiaetus Neolysandra corona 69 99 Neolysandra fatima 10Po0lyommatus eroides forsteri 53 Polyommatus menelaos 40 31 100 Polyommatus eroides eroides Polyommatus eros yildizae 100 85 Polyommatus icarus Poly10om0 matus dorylas 100 Polyommatus icarus (Morocco) 36 100 Meleageria daphnis 27 100 Polyommatus escheri 41 21 45 Polyommatus thersites Polyom10m0atus amandus amandus Polyommatus amandus abdelaziz 10083 Polyommatus myrrhinus 100 Polyommatus cornelia 99 P85olyommatus aedon Neolysandra coelestina 39 93 Neolysandra diana Lysandra 100

200000 Fig. 41. MrBayes, COI, Polyommatus

50 A molecular phylogeny of Agrodiaetus

99 100 Agrodiaetus Neolysandra corona 69 99 Neolysandra fatima 100 Polyommatus eroides forsteri 53 Polyommatus menelaos 100 Polyommatus eroides eroides 85 Polyommatus eros yildizae 100 Polyommatus icarus 100 Polyommatus dorylas 100 100 Polyommatus icarus (Morocco) 100 Meleageria daphnis Polyommatus escheri Polyommatus thersites 100 Polyommatus amandus amandus Polyommatus amandus abdelaziz 100 83 Polyommatus myrrhinus 100 Polyommatus cornelia 99 85 Polyommatus aedon Neolysandra coelestina 93 Neolysandra diana Lysandra 100

Fig. 42. MrBayes, COI, Polyommatus, condensed (50% level)

99 100 Agrodiaetus Neolysandra corona 69 99 Neolysandra fatima 100Polyommatus eroides forsteri 53 Polyommatus menelaos 40 Polyommatus eroides eroides 100 Polyommatus eros yildizae 31 85 Polyommatus icarus (Spain) Polyommatus andronicus 100 Polyommatus icarus (Turkey) 69 67 Polyommatus icarus (Greece) 29 Polyommatus icarus (Iran) 100 Polyommatus dorylas (Spain) 100 Polyommatus dorylas (SW Turkey) 100 Polyommatus dorylas (NE Turkey) Polyommatus icarus (Morocco) 73 Meleageria daphnis marcida 62 Meleageria daphnis versicolor 100 27 Meleageria daphnis versicolor 36 Meleageria daphnis brandti 87 Meleageria daphnis marcida 21 100 Polyommatus escheri (Greece) Polyommatus escheri (Spain) 41 45 Polyommatus thersites (NE Turkey) Polyommatus thersites (Spain) 100 71 Polyommatus thersites (Iran) 35 Polyommatus thersites (SW Turkey) Polyommatus amandus amandus Polyommatus amandus abdelaziz 97 Polyommatus myrrhinus (Erzurum Prov.) 83 Polyommatus myrrhinus (Erzurum Prov.) 100 Polyommatus myrrhinus (Erzurum Prov.) 37 Polyommatus cornelia (Kayseri Prov.) 100 Polyommatus cornelia (Adana Prov.) 85 Polyommatus cornelia (Erzincan Prov.) 99 Polyommatus aedon aedon Neolysandra coelestina 39 93 Neolysandra diana Lysandra 100

200000 Fig. 43. MrBayes, COI, Polyommatus

Within the genus Polyommatus (Fig. 41, Fig. 42) the subgenera Agrodiaetus (sensu ECKWEILER & HÄUSER 1997) and Lysandra appear clearly as monophyletic units. The

51 Chapter 3 unification of Meleageria with Lysandra is not supported. The subgenus Neolysandra splits into two separate clusters. One of them, which consists of the species N. corona (Verity, 1936) and N. fatima (Eckweiler & Schurian, 1980), forms the sister clade to Agrodiaetus, the other (with N. coelestina (Eversmann, 1843) and N. diana Miller, 1913) clusters with the P.aedon/myrrhinus/cornelia-group. This split is also supported by genital morphology (COUTSIS 2001) which also suggests a closer relationship of N. corona and N. fatima to P. dorylas ([Denis & Schiffermüller], 1775) than to N. coelestina and N. diana. Some species (P. icarus, P. eroides, P. cornelia) do not appear as monophyletic units in the COI gene tree (Fig. 43) and P. myrrhinus, which is considered a subspecies of P. aedon by HESSELBARTH et al. (1995), clusters with P. cornelia. The genus Lysandra constitutes a parallel case compared to Agrodiaetus with closely related taxa differing in (mostly high) chromosome numbers. Lysandra syriaca appears at the basis but the relationships of the other taxa of the coridon group remains unresolved or obscure and only the populations of Lysandra bellargus form a well supported monophyletic clade (Fig. 44, Fig. 45).

Lysandra syriaca 100 Lysandra ossmar (Turkey: Isparta) Lysandra corydonius (Turkey: Erzurum) 100 Lysandra corydonius (Turkey: Igdir) 99 Lysandra corydonius (Turkey: Igdir) 46 Lysandra ossmar (Turkey: Sivas) Lysandra albicans (Spain: Huesca) Lysandra albicans (Spain: Huesca) 86 96 Lysandra albicans (Spain: Soria) 60 Lysandra coridon (Spain: Burgos) 39 Lysandra gennargenti (Italy: Sardinia) 61 100 Lysandra coridon (Spain: Barcelona) 70 Lysandra caelestissimus (Spain: Teruel) 98 53 Lysandra coridon (Spain: Tarragona) 98 Lysandra corydonius (Turkey: Erzurum) 100 Lysandra coridon (Italy: Aosta) 23 Lysandra bellargus (Iran) 21 Lysandra bellargus (Italy) 51 Lysandra bellargus (SE Turkey) Lysandra bellargus (SW Turkey) 100 Lysandra bellargus (Spain) 100000 Fig. 44. MrBayes, COI, Lysandra

Lysandra syriaca 100 Lysandra ossmar (Turkey: Isparta) Lysandra corydonius (Turkey: Erzurum) 100 Lysandra corydonius (Turkey: Igdir) 99 Lysandra corydonius (Turkey: Igdir) Lysandra ossmar (Turkey: Sivas) Lysandra albicans (Spain: Huesca) 86 Lysandra albicans (Spain: Huesca) 96 Lysandra albicans (Spain: Soria) 60 Lysandra coridon (Spain: Burgos) Lysandra gennargenti (Italy: Sardinia) 61 100 Lysandra coridon (Spain: Barcelona) 98 70 Lysandra caelestissimus (Spain: Teruel) 53 Lysandra coridon (Spain: Tarragona) Lysandra corydonius (Turkey: Erzurum) 98 100 Lysandra coridon (Italy: Aosta) Lysandra bellargus (Iran) 51 Lysandra bellargus (Italy) Lysandra bellargus (SE Turkey) Lysandra bellargus (SW Turkey) 100 Lysandra bellargus (Spain) Fig. 45. MrBayes, COI, Lysandra, condensed (50% level)

The condensed tree for the Agrodiaetus clade (with 95% confidence limit) is presented in Fig. 46. The following clades are well supported: • antidolus-group (antidolus, kurdistanicus, femininoides, morgani, peilei, karindus) • elbursicus-group (elbursicus, zarathustra, arasbarani, paulae) 52 A molecular phylogeny of Agrodiaetus

• carmon-group (carmon, surakovi, sekercioglui) • admetus-group (admetus, ripartii, nephohiptamenos, demavendi, lorestanus, khorasanensis) • iphidamon-group (iphidamon, dizinensis) • menalcas-group (menalcas, alcestis, interjectus, dantchenkoi, aroaniensis, humedasae, valiabadi, fabressei, ainsae, fulgens) • poseidonides-group (poseidonides, dagmara) • baytopi-group (baytopi, rovshani, tankeri, iphicarmon) • erschoffii-group (erschoffii, tenhageni, achaemenes, shahrami, glaucias, phyllis, klausschuriani, posthumus, darius, caeruleus, birunii) • poseidon-group (poseidon, putnami, hopfferi, lycius, actis, firdussii, sigberti, pseudactis, artvinensis, ernesti, damalis, merhaba, mithridates, wagneri, altivagans, sertavulensis, merhaba, mofidii, pseudoxerxes, gorbunovi, kanduli, sennanensis, cyaneus)

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53 Chapter 3

On a lower level of confidence (83%), the first three clades form a joint clade together with the taxa dama, schuriani and pierceae and, with only 75% confidence, the taxa ninae, turcicola, zapvadi, huberti and damon cluster together. The relationships between the different clades remain largely unresolved on a low level of confidence.

92 Agrodiaetus JC00039 P.escheri MW98235 P.cornelia MW99038 P.cornelia MW98264 P.cornelia 100 MW00326 P.aedon MW99537 P.myrrhinus MW99550 P.myrrhinus 100 100 MW02001 P.amandus 99 MW99047 P.amandus MW00504 N.corona 93 MW99301 N.fatima 100 MW00076 M.daphnis 100 MW98029 M.daphnis MW00290 M.marcida MW99013 N.coelestina MW98220 P.loewii 94 MW99018 A.pyrenaicus 99 MW99134 A.eumedon MW00024 V.alcedo MW99163 P.pylaon 100 MW99303 K.eurypilus 99 100 MW00525 C.semiargus MW02034 C.semiargus MW00517 V.morgianus 100 MW01061 A.montensis 99 MW02033 A.montensis JC00055 A.artaxerxes 100 100 MW01048 A.cramera 99 MW99097 A.isauricus 100 MW99001 A.torulensis MW98270 A.anteros MW00116 P.argus MW00469 L.bellargus 100 MW99608 L.bellargus MW01011 L.bellargus MW98278 L.bellargus 99 99 MW99514 L.corydonius MW99536 L.corydonius MW01059 L.albicans 96 MW98129 L.ossmar MW98185 L.ossmar 100 MW98228 L.syriaca MW01116 L.coridon MW01018 L.coridon 0 MW01092 L.albicans MW01034 L.albicans MW99612 L.coridon 100 OK96022 L.caelestissimus 100 MW01019 P.dorylas MW99014 P.dorylas MW00302 P.thersites 100 MW01083 P.thersites MW02006 P.icarus MW00412 P.icarus MW00530 P.eroides JC00029 P.menelaos JC00042 P.eroides MW01025 P.icarus JC00061 P.andronicus JC00063 P.icarus MW02025 T.theophrastus

Fig. 47. MrBayes, ITS2, Polyommatiti, condensed (90% confidence level)

54 A molecular phylogeny of Agrodiaetus

The Bayesian analysis of the ITS-2 dataset does not support the monophyly of Polyommatus (Fig. 47). Relationships of Polyommatus sensu strictu appear unresolved and the Plebeius subcluster which is found within Polyommatus includes not only Cyaniris but also Lysandra. The monophyly of Aricia (excl. A. eumedon) and Lysandra are very well supported and within the latter group the taxa bellargus, corydonius and ossmar appear as monophyletic units. The monophyly of Agrodiaetus is supported with a posterior probability of 92%, but the sister group is unresolved. Within Agrodiaetus (Fig. 48), Agrodiaetus damon appears as the sister taxon to all other Agrodiaetus, but the phylogeny of the remaining taxa is mainly unresolved.

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Fig. 48. MrBayes, ITS2, Agrodiaetus, 80%

Apart from several small clusters of closely related species only few larger clusters appear with a high level (>90%) of confidence: • menalcas-group (menalcas, alcestis, interjectus, dantchenkoi, aroaniensis, humedasae, fabressei, ainsae, fulgens) • iphigenia-group (iphigenia, baytopi, rovshani, tankeri, iphicarmon, turcicus) • carmon-group (carmon, surakovi, sekercioglui) • antidolus-group (antidolus, kurdistanicus, femininoides, morgani)

55 Chapter 3

The last two groups form a cluster together with the taxa dama, peilei, huberti, theresiae, guezelmavi, arasbarani, elbursicus, ninae, turcicola, pierceae, zarathustra, zapvadi and the undescribed taxon from Ahar with a posterior probability of 81% and the admetus-group (admetus, ripartii, nephohiptamenos, demavendi, lorestanus, khorasanensis) reappears with a confidence level of 84%. Relationships between these groups are unresolved.

100 Polyommatus 100 Plebeius (Plebejides) pylaon 76 Plebeius (Kretania) eurypilus 100 Plebeius (Vacciniina) alcedo Plebeius (Vacciniina) morgianus Polyommatus (Cyaniris) semiargus 59 100 Plebeius (Plebeius) argus Plebeius (Aricia) eumedon 60 Plebeius (Plebejidea) loewii 52 Plebeius (Agriades) pyrenaicus Plebeius (Aricia) anteros 100 Plebeius (Aricia) torulensis 100 Plebeius (Aricia) isauricus 100 Plebeius (Aricia) cramera Plebeius (Aricia) artaxerxes 67 97 Plebeius (Aricia) montensis (Spain) 99 Plebeius (Aricia) montensis (Morocco) Lysandra 100 Tarucus theophrastus

Fig. 49. MrBayes, ITS2&COI, Polyommatiti, condensed tree (50% confidence level)

Lysandra syriaca 100 Lysandra ossmar (Turkey: Isparta) Lysandra corydonius (Turkey: Erzurum) 97 Lysandra ossmar (Turkey: Sivas) Lysandra corydonius (Turkey: Igdir) 69 30 Lysandra bellargus (Turkey) 76 Lysandra bellargus (Italy) 76 100 Lysandra bellargus (Iran) Lysandra bellargus (Spain) 66 Lysandra albicans (Spain: Huesca) 31 Lysandra albicans (Spain: Soria) 100 Lysandra albicans (Spain: Huesca) 51 Lysandra coridon (Spain: Burgos) Lysandra coridon (Spain: Barcelona) 84 Lysandra coridon (Italy: Aosta) 98 94 Lysandra caelestissimus (Spain: Teruel)

50000 Fig. 50. MrBayes, ITS2&COI, Lysandra

Lysandra syriaca 100 Lysandra ossmar (Turkey: Isparta) Lysandra corydonius (Turkey: Erzurum) 97 Lysandra ossmar (Turkey: Sivas) Lysandra corydonius (Turkey: Igdir) Lysandra bellargus (Turkey) 100 Lysandra bellargus (Italy) 69 Lysandra bellargus (Iran) Lysandra bellargus (Spain) Lysandra albicans (Spain: Huesca) Lysandra albicans (Spain: Soria) Lysandra albicans (Spain: Huesca) 100 Lysandra coridon (Spain: Burgos) Lysandra coridon (Spain: Barcelona) Lysandra coridon (Italy: Aosta) 98 Lysandra caelestissimus (Spain: Teruel)

Fig. 51. MrBayes, ITS2&COI, Lysandra, condensed tree (95% confidence level)

In the combined data set of COI and ITS-2, the subgenus Lysandra forms a monophyletic clade at the base of the phylogram while the other Polyommatus (excluding Cyaniris) are found in a highly supported monophyletic clade at the tip of the tree (Fig. 49, Fig. 50, Fig.

56 A molecular phylogeny of Agrodiaetus

51). The sister group to this clade is a weakly supported Plebeius monophylum which includes Cyaniris. Apart from the well supported and resolved Aricia-monophylum (excluding A. eumedon) the relationships within the genus Plebeius remain poorly resolved. Lysandra syriaca appears to be the sister species of all other Lysandra species which are divided into four well-supported clades, but it is conspicuous that corydonius and ossmar do not appear as monophyletic units. The other Polyommatus species split into seven groups (Fig. 52, Fig. 53) and Neolysandra coelestina (which is separated from the other two Neolysandra species) appears as sister taxon to the remaining Polyommatus. The relationships between the other main clades, which appear with a posterior probability of 100%, remain unresolved. 100 Agrodiaetus 100 Polyommatus dorylas (Spain) 28 90 Polyommatus dorylas (NE Turkey) Polyommatus thersites (Iran) 100 Polyommatus thersites (Spain) 100 Polyommatus icarus (Morocco) 85 Polyommatus menelaos 44 100 Polyommatus eroides eroides 100 Polyommatus eroides forsteri Polyommatus icarus (Iran) 62 Polyommatus icarus (Greece) 100 Polyommatus andronicus 73 100 73 Polyommatus icarus (Spain) Polyommatus escheri 100 Meleageria daphnis brandti 100 Meleageria daphnis versicolor Meleageria daphnis marcida 94 100 Polyommatus amandus abdelaziz 31 Polyommatus amandus amandus 100 Neolysandra corona 100 Neolysandra fatima Polyommatus aedon aedon 99 Polyommatus cornelia (Adana Prov.) 100 Polyommatus cornelia (Erzincan Prov.) 100 Polyommatus cornelia (Kayseri Prov.) Polyommatus myrrhinus (Erzurum Prov.) 66 98 Polyommatus myrrhinus (Erzurum Prov.) Neolysandra coelestina

200000 Fig. 52. MrBayes, ITS2&COI, Polyommatus (excl. Lysandra)

100 Agrodiaetus 100 Polyommatus dorylas (Spain) Polyommatus dorylas (NE Turkey) Polyommatus thersites (Iran) 100 Polyommatus thersites (Spain) 100 Polyommatus icarus (Morocco) Polyommatus menelaos 100 Polyommatus eroides eroides Polyommatus eroides forsteri 100 Polyommatus icarus (Iran) Polyommatus icarus (Greece) 94 100 Polyommatus andronicus 100 Polyommatus icarus (Spain) Polyommatus escheri 100 Meleageria daphnis brandti 100 Meleageria daphnis versicolor Meleageria daphnis marcida 100 Polyommatus amandus abdelaziz Polyommatus amandus amandus 100 Neolysandra corona 100 Neolysandra fatima Polyommatus aedon aedon 99 Polyommatus cornelia (Adana Prov.) 100 Polyommatus cornelia (Erzincan Prov.) Polyommatus cornelia (Kayseri Prov.) 100 Polyommatus myrrhinus (Erzurum Prov.) 98 Polyommatus myrrhinus (Erzurum Prov.) Neolysandra coelestina

Fig. 53. MrBayes, ITS2&COI, Polyommatus (excl. Lysandra), condensed tree (95%)

57 Chapter 3

MW00177 A.gorbunovi C20 100 MW00178 A.gorbunovi 36 MW00177 A.gorbunovi C20 100 MW00129 A.gorbunovi C20 40 MW00178 A.gorbunovi WE02674 A.gorbunovi 100 MW00129 A.gorbunovi C20 100 100 MW00330 A.pseudoxerxes C15 WE02674 A.gorbunovi MW00179 A.cyaneus 100 MW00330 A.pseudoxerxes C15 MW99094 A.cyaneus C17 100 MW00179 A.cyaneus 100 MW99448 A.cyaneus C18 MW99094 A.cyaneus C17 100 40 WE02621 A.sennanensis 100 MW99448 A.cyaneus C18 99 MW98203 A.mithridates C23 90 WE02621 A.sennanensis 100 WE02454 421 A.mofidii MW98203 A.mithridates C23 94 98 MW99465 A.kanduli C25 99 100 WE02454 421 A.mofidii MW99353 A.altivagans C22 MW99465 A.kanduli C25 98 MW98313 A.sertavulensis C20 MW99353 A.altivagans C22 100 MW98136 A.wagneri C18 MW98313 A.sertavulensis C20 96 MW98170 A.maraschi C16 100 100 MW98136 A.wagneri C18 98 MW99240 A.altivagans C21 96 MW98170 A.maraschi C16 55 MW99057 A.merhaba C17 100 98 MW99240 A.altivagans C21 AD98012 A.altivagans MW99057 A.merhaba C17 100 MW98138 A.poseidon C20 AD98012 A.altivagans 100 MW98154 A.poseidon C21 100 MW98138 A.poseidon C20 MW98180 A.poseidon C19 71 100 MW98154 A.poseidon C21 100 MW99501 A.putnami C25 100 MW98180 A.poseidon C19 83 MW99408 A.hopfferi C15 MW99501 A.putnami C25 MW98189 A.hopfferi C16 72 MW99408 A.hopfferi C15 49 MW98079 A.lycius MW98189 A.hopfferi C16 MW98097 A.ernesti C18 MW98079 A.lycius MW98285 A.sigberti C25 66 MW98097 A.ernesti C18 MW99006 A.firdussii C30 37 MW98285 A.sigberti C25 100 MW00234 A.firdussii C25 67 MW99006 A.firdussii C30 AD98009 A.pseudactis 60 100 MW00234 A.firdussii C25 100 MW99058 A.artvinensis C25 AD98009 A.pseudactis 95 MW98162 A.actis C17 100 MW99058 A.artvinensis C25 MW00151 A.firdussii 95 MW98162 A.actis C17 100 MW99247 A.firdussii C24 42 MW00151 A.firdussii 95 MW99413 A.firdussii C25 100 MW99247 A.firdussii C24 95 JC00045 A.nephohiptamenos 37 MW99413 A.firdussii C25 MW00015 A.demavendi C70 JC00045 A.nephohiptamenos 100 MW00185 A.demavendi C70 80 MW00015 A.demavendi C70 MW99382 A.demavendi C70 100 MW00185 A.demavendi C70 WE02677 A.demavendi 100 MW99382 A.demavendi C70 JC01014 A.admetus 37 WE02677 A.demavendi 100 MW01014 A.ripartii C90 28 60 JC01014 A.admetus JC00043 A.ripartii MW01014 A.ripartii C90 WE02431 A.khorasanensis 52 JC00043 A.ripartii MW99196 A.ripartii C90 49 WE02431 A.khorasanensis MW99407 A.demavendi C60 49 88 MW99196 A.ripartii C90 100 WE02535 A.lorestanus 42 MW99407 A.demavendi C60 MW00189 A.demavendi 100 WE02535 A.lorestanus 100 MW99105 A.demavendi C66 72 MW00189 A.demavendi 100 WE02491 A.achaemenes 100 MW99105 A.demavendi C66 WE85001 A.shahrami 100 WE02491 A.achaemenes 100 MW00348 A.phyllis C85 WE85001 A.shahrami 100 MW00452 A.phyllis C75 100 MW00348 A.phyllis C85 AD98036 A.phyllis 100 MW00452 A.phyllis C75 99 MW99174 A.phyllis 100 AD98036 A.phyllis MW00262 A.klausschuriani C56 99 MW99174 A.phyllis 100 WE02451 A.tenhageni 52 65 MW00262 A.klausschuriani C56 WE00002 A.glaucias 35 39 WE02451 A.tenhageni MW00393 A.erschoffii C14 WE00002 A.glaucias 97 MW00101 A.darius MW00393 A.erschoffii C14 MW00347 A.posthumus C85 11 97 MW00101 A.darius MW00444 A.birunii C11 39 MW00347 A.posthumus C85 95 MW00267 A.birunii C10 95 MW00444 A.birunii C11 MW00102 A.birunii C11 MW00267 A.birunii C10 100 MW00072 A.birunii C11 100 MW00102 A.birunii C11 MW00409 A.caeruleus C20 57 MW00072 A.birunii C11 MW00547 A.birunii C10 43 MW00409 A.caeruleus C20 DS00001 A.poseidonides 24 28 MW00547 A.birunii C10 95 DS01001 A.iphigenides 56 DS00001 A.poseidonides 100 MW98049 A.iphigenia C14 DS01001 A.iphigenides MW99009 A.iphigenia C12 55 100 MW98049 A.iphigenia C14 MW99135 A.turcicus C25 78 MW99009 A.iphigenia C12 100 MW99203 A.turcicus C24 43 MW99135 A.turcicus C25 97 96 MW98103 A.iphicarmon 100 MW99203 A.turcicus C24 MW99372 A.baytopi C27 86 MW98103 A.iphicarmon MW99565 A.tankeri 97 96 MW99372 A.baytopi C27 100 MW00176 A.rovshani MW99565 A.tankeri 100 WE02662 A.rovshani MW99309 A.baytopi 100 MW00176 A.rovshani 100 100 WE02662 A.rovshani MW99559 A.tankeri 100 MW99309 A.baytopi 100 MW00064 A.valiabadi 39 MW99559 A.tankeri MW00498 A.valiabadi C23 100 MW00064 A.valiabadi JM00001 A.fabressei MW01039 A.fabressei MW00498 A.valiabadi C23 100 25 JM00001 A.fabressei MW01001 A.ainsae 41 MW01039 A.fabressei MW01053 A.ainsae 100 MW01001 A.ainsae MW01107 A.fulgens 48 100 MW01053 A.ainsae 100 100 MW99591 A.humedasae C38 100 94 MW01107 A.fulgens MW99605 A.humedasae 100 MW99591 A.humedasae C38 JC00040 A.aroaniensis 93 100 MW99605 A.humedasae MW98172 A.menalcas JC00040 A.aroaniensis MW98315 A.alcestis C20 93 MW98172 A.menalcas MW98212 A.alcestis C21 MW98315 A.alcestis C20 100 MW99164 A.interjectus C31 100 MW98212 A.alcestis C21 MW00231 A.alcestis C19 100 MW99164 A.interjectus C31 100 MW99274 A.dantchenkoi C42 75 MW00231 A.alcestis C19 MW99319 A.dantchenkoi C42 93 18 MW99274 A.dantchenkoi C42 MW99276 A.dantchenkoi C42 95 53 MW99319 A.dantchenkoi C42 MW99471 A.dantch.Xmenalcas C50 22 MW99276 A.dantchenkoi C42 MW00539 A.dizinensis C17 15 MW99471 A.dantch.Xmenalcas C50 100 MW00269 A.iphidamon C14 MW00539 A.dizinensis C17 100 MW00328 A.iphidamon MW00269 A.iphidamon C14 97 MW99292 A.pierceae C21 100 100 MW00328 A.iphidamon MW99341 A.pierceae C23 97 MW99292 A.pierceae C21 96 MW00226 A.femininoides C27 MW99341 A.pierceae C23 96 WE02671 A.femininoides C27 96 MW00226 A.femininoides C27 100 MW99393 A.antidolus C42 96 WE02671 A.femininoides C27 MW99406 A.antidolus C44 100 100 MW99393 A.antidolus C42 MW99286 A.kurdistanicus C55 MW99406 A.antidolus C44 WE02591 A.peilei C39 100 93 MW99286 A.kurdistanicus C55 100 WE02614 A.morgani C27 54 WE02591 A.peilei C39 MW98205 A.dama 100 100 WE02614 A.morgani C27 MW98261 A.schuriani C80 47 AD98001 A.surakovi MW98205 A.dama 100 MW98261 A.schuriani C80 100 MW98009 A.carmon AD98001 A.s urakovi MW00032 A.hamadanensis C22 100 MW00051 A.elbursicus C17 100 MW98009 A.carmon 100 100 MW00032 A.hamadanensis C22 MW00110 A.elbursicus C18 90 52 MW00051 A.elbursicus C17 MW00316 A.elbursicus C17 100 MW00110 A.elbursicus C18 100 MW00058 A.elbursicus C18 84 MW00316 A.elbursicus C17 MW00232 A.elbursicus C18 38 MW00058 A.elbursicus C18 WE02661 A.arasbarani 61 53 MW00232 A.elbursicus C18 MW00127 A.paulae C17 100 WE02661 A.arasbarani 100 WE02531 A.zarathustra C22 MW00127 A.paulae C17 100 MW98240 A.theresiae C59 100 WE02531 A.zarathustra C22 MW98294 A.guezelmavi 100 MW98240 A.theresiae C59 MW99479 A.turcicola C20 39 MW98294 A.guezelmavi MW99508 A.ninae C34 44 MW99479 A.turcicola C20 AD98018 A.ninae 33 100 MW99226 A.zapvadi 57 MW99508 A.ninae C34 AD98018 A.ninae 100 MW99374 A.zapvadi C19 100 MW99226 A.zapvadi AD98024 A.huberti 100 MW99374 A.zapvadi C19 MW99095 A.huberti C34 100 MW99552 A.huberti C33 78 AD98024 A.huberti MW99095 A.huberti C34 MW99546 A.damon 100 46 MW99552 A.huberti C33 100 MW99613 A.damon MW99546 A.damon 100 MW99613 A.damon 100000 Fig. 54. MrBayes. ITS2&COI. Agrodiaetus Fig. 55. MrBayes. ITS2&COI. Agrodiaetus, 95%

58 A molecular phylogeny of Agrodiaetus

The subgenus Agrodiaetus which represents one of these clades splits into 10 groups which can be named according to the taxonomically oldest species (Fig. 56, Fig. 57). One of them which includes only one species, Agrodiaetus damon, appears to be the sister group to all the others. The complete tree is presented in fig. Fig. 54 and the condensed topology (confidence limit of 95%) in Fig. 544.

60 100 poseidon-group 28 admetus-group 52 100 erschoffii-group 56100 poseidonides-group 55 iphigenides-group 48 43 iphigenia-group 97 95 dolus-group 100iphidamon-group 100 carmon-group 100 damon-group 100

100000 Fig. 56. MrBayes, ITS2&COI, Agrodiaetus

100 poseidon-group admetus-group 100 erschoffii-group 100 95 poseidonides-group iphigenides-group iphigenia-group 97 dolus-group 100 iphidamon-group 100 carmon-group 100 damon-group 100

Fig. 57. MrBayes, ITS2&COI, Agrodiaetus, condensed tree (95% confidence limit)

Apodemia mormo (Riodinidae) MW00497 Favonius quercus (Theclini) MW99045 Maculinea arion (Glaucopsychiti) JC00057 Aricia artaxerxes 0 MW00302 Polyommatus thersites MW00032 Agrodiaetus hamadanensis 100 MW00056 Agrodiaetus elbursicus MW99406 Agrodiaetus antidolus MW00072 Agrodiaetus birunii MW98154 Agrodiaetus poseidon 98 MW00076 Meleageria daphnis MW99014 Polyommatus dorylas 100 MW00412 Polyommatus icarus MW00530 Polyommatus eroides MW00409 Agrodiaetus caeruleus MW99550 Polyommatus myrrhinus 96 MW98228 Lysandra syriaca 100 MW99068 Agrodiaetus ripartii MW00189 Agrodiaetus demavendi 88 MW99105 Agrodiaetus demavendi MW98172 Agrodiaetus menalcas MW99471 A.dantchenkoiXmenalcas MW00231 Agrodiaetus alcestis 73 MW99164 Agrodiaetus interjectus MW00328 Agrodiaetus iphidamon MW99135 Agrodiaetus turcicus

Fig. 58. MrBayes, ND1, Lycaenidae, condensed tree (70% confidence limit)

For the ND1 analysis most taxa were chosen to represent the main Agrodiaetus clades which were obtained from the combined COI- and ITS2-analyses in order to check if this gene gives a better resolution of clade relationships. The tree obtained from the ND1 analysis (Fig. 58) has little resolution but is compatible with those of the COI- and ITS2-analysis. A

59 Chapter 3 combination of the ND1 dataset with the COI- and ITS2-datasets also does not improve the resolution regarding the relationships of the different Agrodiaetus clades with each other. The tree obtained from the Cytb analysis (Fig. 59) is also compatible with the results from the other genes but due to the low number of taxa sequenced does not provide additional information.

Ostrinia nubilalis AF442957 Lycaena thersamon MW98006 Polyommatus cornelia MW98225 Agrodiaetus iphicarmon MW98102 100 100 Agrodiaetus admetus MW98055 51 Agrodiaetus turcicola MW99258

0.05 Fig. 59. MrBayes, Cytb

Maximum parsimony analysis The Maximum parsimony analysis yielded 100000 trees in each of the COI-, ITS2- and the combined analysis because the maximum limit of 100 trees was reached in each replicate. The trees obtained from the COI analysis had scores between 2414 and 2433. The consensus tree calculated from the 300 shortest trees (score=2414) is shown in Fig. 60 for the outgroup & Fig. 61 for Agrodiaetus and that calculated from all trees in Fig. 62 (outgroup) & Fig. 63 (Agrodiaetus).

Scores of the ITS-2 analysis were between 1105 and 1107 (39800 of which had the lowest score of 1105). The consensus tree of all trees is shown in Fig. 64.

Scores of the combined analysis were between 2599 and 2606. A consensus tree of the 2700 shortest trees is presented in Fig. 65 and the consensus tree of all trees is shown in Fig. 66 & Fig. 67.

Bootstrap trees for the three analyses are shown in Fig. 68 (COI outgroups), Fig. 69 (COI Agrodiaetus), Fig. 70 (ITS-2), Fig. 71 (ITS-2 & COI combined, outgroups) and Fig. 72 (ITS-2 & COI combined, Agrodiaetus).

MP-trees calculated from the ND1-gene were poorly resolved and different combinations with the COI- and/or ITS-2 gene also did not provide new results and are therefore not presented here.

60 A molecular phylogeny of Agrodiaetus

A F A 1 F A 7 0 F 0 4 s A s 0 8 4 s a a i F l 44 6 a i s 0 l r l r 1 0 y y e s 0 e i r y t 70 r e r 0 r i e M C 6 h s o t 0 o h s e o e c r s t 873 W o P d c t 7 d si . d s i s . e r e e a . s s t r 0 e C P h e i e n e . o p P t f A 1 P . s a . h er I o o i r 1 9 P t z F 8 ph l P . s i M l n i 4 h e us i P 1 J 1 o 1 t e a 2 . r d C y 1 9 P h l 7 W 4 9 i i 0 t a d c s m 1 3 i 0 m 0 0 P . y 0 3 c 0 l 1 8 8 8 i I e 8 i o 2 . p 9 0 P . 2 de a 0 r s 6 f 0 0 2 0 ur 9 1 s 0 e h c 9 P e o s 3 0 1 1 02 8 . r e 2 0 s i a 0 o h W s y 0 9 0 A 1 W 0 6 P e d a M 4 t . i l s po t t a 9 03 0 p h h C W o I. M 0 e o M W I u o M P r o . a e MW W 0 8 n f p l J 3 e la W d dal l n M W 9 9 0 e o m m i 5 . e e 2 e i a M W 7 0 s d M 0200 0 P m n s M 1 0 m i e e W 5 . e 2 th a r l M 0 2 ru W 0 i l i i M W 9 P M 2 1 a a i u i 9 4 .m a s 0 r s M W 9 c s W 0 3 Z m m iu 0 P .i u u 4 M W 0 2 r c M 0 9 S .kn o e s 0 0 1 P ca ni 2 7 .p r li M 0 5 5 .i o W 0 i ysn m i C 0 0 2 r 0 0 F ri o J 0 0 P d 0 7 .q th C 0 3 n s 0 L u o a J 1 3 .a u M 1 .a e u C 0 1 P r M W 7 l r s J W 8 ca s W 9 L ci c 9 61 .i ru 9 8 .a p u M W 0 P ca 9 2 lc hr s 0 3 .i a M 51 30 ip o M 0 6 P tim W 5 L h n C 00 2 fa 0 L. .t ro J 0 41 N. 01 vi ity n C 0 1 tima 19 rg ru J 0 0 .fa MW L au s W 93 N na M 9 .c re M 9 9 oro W98094 95 an ae W 46 .c us 20 de M 99 4 N nd L. ns W 50 ma MW L.t the M 00 P.a a 00 her tis W 01 rcid 044 sam M 20 .ma M L.a on W0 0 M is W02 sab M 029 phn 009 inu W0 M.da M L.ph s M 74 is W993 lae 982 aphn 98 S as MW 9 M.d M .myrt 9802 nis W010 ale MW .daph 27 S.e 0076 M MW991 sculi MW0 cida 58 S.hyr 7 M.mar canicum MW0008 MW amandus 98002 C.osiris MW99047 P. MW01120 C.marshalli MW98228 L.syriaca MW99084 C.argiolus MW99042 L.corydonius lus MW995 MW02008 C.argio 36 L.corydonius MW phrastus 98129 L. 025 T.theo ossmar MW02 ion MW004 T.endym 69 L.b 99221 n MW ellargus MW M.ario 98278 9045 MW L.be MW9 rama 994 llargu P.vic M 35 L s 080 gus W99 .bell W99 erra M 608 argu M enc us W0 L.b s P.ab etic M 101 ella 31 .bo us W 1 L rgus 020 2 L tic M 991 .be MW 102 boe s W 40 llar W0 L. ticu M 99 L. gus M 65 oe s W 51 cor 82 .b ylu M 98 4 L ydo W9 8 L ch o W 1 .c ni M 2 ro in M 0 85 or us 20 .t rd s W 1092 L yd 0 C na ti M 0 .o on MW 25 er es s W 1 ss ius 94 b g e M 05 L. ma 9 es .a rx W 01 9 al r W ot A e tis O 0 L bic M il 2 ax s K 0 3 .a a ph 06 rt e tis M 9 10 4 lb ns u 0 g s s W 9 1 L i E 0 .a .a e e M 0 8 .a ca 4 C A A g x W 0 0 lb n 86 J 7 a r is M 1 1 L. ic s 0 5 0 e s O W 9 1 c a 7 0 2 A. x n is 9 0 L o n 1 0 0 a s M K .g r s F 0 0 8 rt te s 0 6 3 e id A C 0 2 n n u M W 9 1 1 L o J 0 .a te c is 6 1 2 . n n W 9 o i s a M W 1 c n M 9 A n r r M 9 0 L o a m u n s W 2 6 r 5 . o e M 9 9 .c r g W a e o a M W 0 2 id 5 A m l r 00116 9 L o e M 0 . s m n a M W o i u M 1 . r n 3 . e W 1 L c 0 r n M 9 i n t A a t a n MW 8 d M W . i 3 i M 3 o 0 A o r i o a 9 9 c n i a W o 0 8 t l W 4 r C . c d i a 0 8 A a d s 1 i n . . st l i W s W 2 4 7 a s 0 d J e l 0 e . 0 2 2 . 0 9 A A e e e u A p o 0 u u 0 2 N l n e l 9 5 e A 9 0 2 4 n 1 0 a r n n 9 P . y 1 n 0 0 W n 1 e . n i e i 9 2 s 9 3 r i 0 r 0 9 5 0 o r 9 0 0 3 P . e t M 6 o h 5 u 9 0 P o h 4 ar h i 1 1 c r 3 W 0 7 o P n s c r 4 m 0 . r 2 . 0 c r . 3 7 c s 9 r 6 r W 2 6 . c 0 C g a M 1 4 . 0 . y e i 9 P 0 C h y l i 8 N 3 m 2 3 us 0 P . o c M 0 V r d 9 P V s . u W 3 4 m e i u V m o 3 my . P s s W E K . e 8 . s 0 6 . . m w s 1 5 e t n M W . a m P . 3 . o M 0 a P 2 P W 3 p 0 e m i l o M 0 i p 8 l c i 9 2 y 8 u c a W 0 r 7 h 9 e i 9 l g 9 8 e a 3 r r 5 a i 9 3 d M y ia g 9 5 d r 5 o W W 5 g p o u W 9 o n 9 n 9 u i s M M W 9 l u 9 u M 9 s M s

s W

W

M

M MW Fig. 60. MP Majority Rule Consensus Tree of the shortest trees (COI) excluding Agrodiaetus

61 Chapter 3

5

5 2

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d e 7 8 17 n x 1 1 r s C a i 9

C e C 0 s k MW 1 0 x 2 0 s n 2 s s s ba s i C 2 o 2 us u e C 2 u e n v i a u d l i C C e n 99 e C e a o v h c i u v 9 n a r s M n g n o MW n o o v s 1 i e ane a n 1 3 M e a a u n o s M a n W s n 8

W y C M n m 8 y v u n 5 MW y b u n a 1 W p i M m i W c cy r C2 2 2 c e a t . e 0 . c b r . u E 9 W . l g . b l C . o r 6 3 W 0 9 s d r b e s C 0 A 0 9 A A . a a u i J W A . 2 A o r 1 9 g 1 A A . r 0 9 o n n 0 2 3 v v s C 0 . . A A g o i C 1 8 d 9 0 g t g e a C J E 0 6 8 6 4 A . a n 0 8 l s C 0 5 9 A . g t n i 5 3 8 em 59 a g a s 0 0 1 7 1 1 9 1 5 . r 0 4 A a e 4 h 0 3 4 7 A . g a e 2 3 1 A 8 7 3 0 8 e g x 0 0 2 6 w c t 4 A A 4 r 0 4 5 6 5 5 . a v a 6 0 s 4 A . 9 7 A i s a 9 6 4 9 7 0 A a 9 . t e 3 d A 9 v 6 . 3 7 7 A w l a 0 i 00179 d A . 9 1 2 1 4 A d v A 9 2 9 . i 1 e . 9 3 A t r A d . 9 9 7 a l r 5 d e e 0 6 1 ox C . m . . d 0 9 5 9 A . a d e 9 a W W h A A d d W 2 d . e W 3 m n a W 0 0 3 A . t n e m W E 3 i i a . a e ma a 0 01 1 6 m u i k d m W 0 1 A . b M . e M 9 m a M W d n a v M 0 e m m M W 0 3 3 m i M p h i M E 8 a e W 9 A . f e a v v o M 8 1 4 5 h s 6 h o a C M W 9 o i s W p e n v e o M W 2 6 A ps r l 1 M o r v n W 9 8 0 . n J h a d v e n 7 W e a 5 0 h e n n M W 9 9 1 7 3 A .m a C W M C o s d i e n 0 M m g i 1 0 1 i n d M W 9 9 1 0 A .m r 2 0 W 0 1 h p a d i C n d i M W 9 8 a a e C M 0 i t n d 2 21 A v f i C 1 1 0 0 p a i C 7 i M W 9 8 1 .d ti f r 2 W 0 0 5 t e C i M W 3 2 7 l p e n 1 1 4 a m n 7 0 M 9 2 A a o ff o C J 9 4 07 3 A m 7 0 W 4 5 . d 5 C 8 e s M W 0 2 0 6 A .h p i n 2 0 0 A 2 A . e n i 0 M 0 9 3 o e o 1 8 ri o s M E 5 2 A .h s id C 5 M 0 4 .r A .ri p n E 9 1 9 o e i 2 M W 1 ip . p a o s W 2 0 8 A .p s m C 9 W M 9 4 A a ri a r s W W 0 8 1 8 A o a i 1 M W 9 . r p ti M E 8 0 .p tn m C 9 W M 9 2 A ad ti a rt i 9 4 8 u a n 1 W 91 9 6 .a m i r ii W D 9 9 3 A .p tn o C 99 9 9 92 4 d C ti A W 9 1 4 A u id n 4 9 6 6 A m e 9 i 8 5 .p se o M 0 06 A 3 . tu 0 M W 9 1 1 A o id W 7 8 . A ri et s M W 98 06 .p e 9 A. A rip . p u 9 01 A os s W 9 de .r a rip ar s M W 9 5 3 .p iu E 10 m ip rti a tii M W 99 8 A yc s 18 M 02 2 a ar i C rt M 81 0 .l iu C W 53 A. ve tii 9 ii W 9 18 A yc sti M 00 5 de nd C 0 M W 8 79 .l e i MW W 18 A m i 90 M 9 0 A ern si 0 9 99 9 .lo a C W 98 9 A. us C3 91 14 A. re ve 60 M W 808 7 fird sii 5 05 1 A de sta nd M 9 09 A. us C2 MW A. .d ma nu i W 98 5 fird sii M 9 de em v s M W 12 A. us 25 M W9 910 ma a en M 00 06 fird ti C W9 93 4 ve ve di W 90 A. ber 92 76 A.d nd nd M 9 34 sig ctis 5 86 A A.a em i C i MW 02 A. uda C2 MW .ku ntid av 66 W0 285 se nsis 993 rdis olu end M 98 A.p vine M 93 tan s C i W 009 .art 7 W99 A.a icus 30 M 98 8 A is C1 MW9 406 ntido C5 AD 905 .act 9473 A.an lus 5 W9 62 A berti A.ku tidol C42 M 981 A.sig C25 WE0 rdista us C MW 284 ussii 2612 nicus 44 W98 A.fird C25 WE A.karin C65 M 9422 dussii 02614 dus C MW9 3 A.fir A.morg 66 W9941 rdussii WE0259 ani C2 M 151 A.fi 24 MW0 1 A.peil 7 MW00 firdussii C 0226 A.fem ei C39 99247 A. ininoides C MW chaemenes WE02671 A.fem 27 E02491 A.a ininoides C27 W .shahrami MW WE85001 A 98205 A.dama .erschoffii WE0267 MW00327 A 6 A.hamadanensis C21 MW00393 A.erschoffii C14 hamadanensis C22 MW00032 A. WE02451 A.tenhageni 00001 A.hamadanensis MW9 MW C80 9174 A.phyllis 61 A.schuriani MW9918 MW982 ae C23 7 A.phyllis 1 A.pierce AD98036 MW9934 eae C21 MW A.phyllis 2 A.pierc 18 00140 A W9929 sicus C MW00 .phyllis M A.elbur C18 M 348 A 00110 rsicus i W004 .phyllis MW A.elbu aran MW 52 A. C85 0232 .arasb lae M 0025 phyllis MW0 661 A .pau W0 9 A.k C75 E02 57 A C22 MW 0262 lauss W 001 stra 17 M 003 A.kl chur MW athu e C W0 35 auss iani C .zar aula 17 W 04 A.ca chu 57 1 A A.p us C s E00 09 eru rian 0253 27 sic icu MW 00 A.ca leus i C56 WE 001 lbur urs 18 M 00 2 A eru C2 W .e elb C W0 34 .gla leu 0 M 1 A A. cus 17 MW 03 7 A uc s C 005 56 rsi s C 6 M 0 58 .po ias 20 W0 000 lbu cu C1 W 01 A st M W .e rsi s on M 0 01 .po hum M 8 A lbu icu m 6 M W 04 A st u 05 .e rs ar 4 W 00 44 .da hu s C 00 A lbu .c i C 79 M 0 26 A ri mu 85 16 .e 9 A lu C i M W 0 7 .b us s MW 03 A 0 iog n v i W 0 54 A iru 0 59 80 rc o ko v M 04 7 .bi nii MW 00 9 e rm ra a 9 M W 00 7 A ru C 0 W ek a u lm 5 W0 0 1 6 .b ni 11 W M .s .c .s e C 4 D 0 02 A. iru i C M A A A ez e 3 D S 06 b n 1 9 60 1 u ia C 20 S 0 00 0 A.b iru ii C 0 28 0 00 .g s e 4 M 0 0 7 A n 1 9 9 8 A e a C 3 i M W 0 00 2 . iru ii 0 9 9 9 4 er in la C rt 0 1 A b n C W W D 9 h .n o ti e e M W 9 0 . iru ii 11 M M A 2 .t A ic r b a 3 M W 9 9 5 A b n C 8 A c e u n 3 W 9 1 .p ir ii 1 9 0 8 r b h i 6 W 9 5 3 A o un C 1 W 4 0 tu u . n C 3 M E 9 5 .d s 1 2 5 . h A . i 9 M 0 2 2 A a e ii 1 M 8 9 A . A rt C 1 i W 0 0 0 6 g id C 9 9 A 4 e ti C d 9 M W 2 1 A .t 1 9 2 8 r a M 9 6 7 3 u m o 1 W W 7 5 0 1 b i v 1 0 M W 9 . r a n M 4 9 0 u e n W 9 6 6 A tu c id M 9 0 8 h b p C 2 4 M 9 9 3 2 . r ic ra e 9 9 9 8 . u ad a i o A W 0 A t c 9 v C 1 4 W 9 9 5 u u s 9 D A h z d m 1 M D 5 9 A .r r ic s

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62 A molecular phylogeny of Agrodiaetus

A i r A F e 1 F t A s e 7 0 r A F a 0 4 o

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63 Chapter 3

6

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k e m a o d m e e 1 e C 5 r k d a C r v c C s b a 2 a i 4 i r b C a a e i i i M a p v a s s u t C n c t l 3 i u v c u M M M s v c i r r a r p e n . . h o s W . s h p C a s n . u e e a e i W W M W . . z a c n 7 . i A A . t a i A b b n s 2 z M 9 A . e m e s 5 9 W A . z c l n 0 A 9 A u u a a n 5 0 C 9 9 . r e 9 1 W 9 A e e 9 3 A h r d M 6 2 h u n a 6 C 0 8 8 A z s M 4 0 0 6 . . t i n 5 5 4 e a d M 0 1 5 . e e 1 0 0 2 1 6 n a s M W W 14 0 2 A A h 4 0 1 4 . a d 7 1 1 3 O 5 t u m u 9 d 9 7 A i 0 0 W 8 2 . M W 9 0 6 9 3 7 A c 9 A 0 9 4 5 g a m a o i 3 9 9 8 4 A u 9 9 . 9 A 7 9 3 9 A h a 2 9 9 5 98001 A 9 a n n W 9 A 9 9 2 . i C 5 M W . . t 9 9 7 8 A m 4 1 d 9 2 . 2 9 9 h a g W 0 6 9 d W 0 0 n 4 W a m t s 9 r A . D 9 0 A . i W M 9 6 W9 4 C e 2 D E 0 i d a r 9 8 9 4 4 h a s u 4 C 9 p A W 5 A i l 7 6 4 . o A 9 M W . m 6 M m s 9 W 0 6 a M 9 1 d C 1 r m A W 9 9 2 s a . M 9 . d o 2 9 2 8 3 i u MW W 9 0 A e r u 0 A a A r r p m M 8 2 2 f d l s u 1 9 i M D 9 A . u i c C M 5 2 A . ve A ti p o p M W 8 0 3 t o i 0 9 3 r . a o M W 9 6 k lu s 1 i r i a n A 9 0 0 5 A . n id n W W 5 5 A . p . i C r M W 7 t o e M 4 r n r p t n W 0 0 0 a a 9 A A . i a i r i M 6 6 A . n id t id E W 1 d p d p a 9 t i M W 0 2 2 t s 0 9 . a r ii M W 2 6 A .a i o 0 1 d A . e t i C a r 0 0 8 2 n d n 9 7 M 2 e lo m r ii r t M W 8 6 A a r i 3 2 4 0 0 .d r ti t ii M E 9 0 2 7 . u in 6 W W 3 1 4 m e a i C 6 i M 0 9 3 A k C C E 1 8 a e C 9 i W 3 9 . m i i 6 9 9 A m s v 0 W W 9 9 6 A e le n C 0 9 0 A .d v ta e 9 0 M 9 3 0 .f i a s 8 3 2 .k A e a n n 0 M W 9 4 3 rg u C M 8 6 h . e n v d W 9 9 7 A .pe o d i 1 W 2 7 o d m d e u i M 4 1 n n 2 M M 9 A 7 r e a i n s M W 9 9 7 A .m ri ia C 3 W 9 . A. a m v C d 9 6 1 a ur e 2 W 3 de s a en 6 i M W 2 9 A k h a C 7 M 00 0 8 m d a v M W 0 5 4 . c e 1 W 1 0 1 em n e d 6 2 1 A .s rc ae C M 0 8 18 A av e n i M E 0 6 2 A ie e s W 01 5 6 .d e a n d W E 2 1 .p rc cu 0 8 A A e n ve si i 0 26 61 A ie si s 8 0 3 .d .d m d n s W E 0 2 2 .p ur cu 1 01 A. em e a i C d W E 8 9 A lb si s C 5 de a ma ve 7 i W 9 92 1 .e ur cu 16 M A. m ve v n 0 W 9 34 A lb si C J W de a n e d M W 9 51 .e ur us C0 98 m ve di nd i M 9 0 A elb sic 7 00 JC 08 av nd C i W 00 56 A. ur C1 JC 46 01 4 en i C 70 M W 00 8 elb e 00 A 01 A. di 7 M 0 05 A. ula 2 045 .ne 4 A ad C 0 W 00 9 pa e C2 A ph .a me 70 M 05 A. ula tra .ne oh dm tu MW 00 27 .pa us M ph ipta e s W 01 A rath ni W0 JC ohi m tus M 0 57 .za ara 8 M 053 000 pta en MW 001 A sb s C1 W0 9 A 43 me os W 531 .ara icu 7 042 .diz A.r no M 02 1 A urs C1 4 A ine ipa s WE 66 .elb icus MW .iph nsis rtii E02 32 A burs C18 MW 003 idam C1 W 002 A.el icus 002 28 A on 7 W 316 burs 69 A .iphi C14 M 00 A.el tis MW9 .iphid dam MW 110 eudac 9203 amo on W00 A.ps i A.tur n C1 M 8009 dussi MW99 cicus 4 AD9 5 A.fir C30 M 526 A C24 0012 rdussii W99135 .turcicu MW 06 A.fi 25 A.turcic s MW990 irdussii C MW us C25 0234 A.f 99565 A.ta MW0 igberti C25 MW9937 nkeri 98285 A.s 2 A.baytopi C MW .artvinensis C25 MW 27 MW99058 A 99344 A.baytopi 1 A.firdussii MW MW0015 98103 A.iphicarmon MW99247 A.firdussii C24 MW99559 A.tankeri MW99413 A.firdussii C25 MW99309 A.baytopi MW9942 hani 2 A.firdussii C25 WE02662 A.rovs MW98162 A rovshani MW .actis C17 W00176 A. s 98284 A. M phigenide MW9 sigberti 8001 A.i des 8097 A.e WE9 phigeni MW980 rnesti C 001 A.i mara MW 79 A.ly 18 DS01 A.dag s 9808 cius 00005 nide MW98 9 A.ly DS oseido ami M 138 cius A.p hahr W98 A.pos 00001 A.s enes MW 189 eid DS 5001 em ni M 994 A.ho on C E8 acha age W 08 pffe 20 W 1 A. enh 14 M 981 A.h ri C1 249 A.t ii C i W9 54 opffe 6 E0 451 hoff offi MW 906 A.po ri C W E02 rsc ch 56 M 99 1 A. se 15 W A.e .ers i C W 50 pu idon 93 A an 57 MW998 1 A tna C2 003 327 uri i C s A 180 .pu mi 1 W 00 sch ian mu D 81 A tna C2 M W us ur hu 85 M 98 83 .po m 5 M kla sch st C s M W 01 A se i C A. us po us ia W 00 2 A .po ido 25 62 kla A. m uc 0 W 0 12 .a se n 02 A. 8 hu la C2 M E 0 9 lti id C1 0 9 35 st .g s 20 W 026 17 A va on 9 MW 25 00 po A u C s M 7 .go ga C 00 W A. 02 le s iu 1 M W 00 75 A. rb ns 19 W M 7 0 ru eu ar 1 9 1 A go u M 4 00 ae ul .d C 0 M W 9 78 . r no 03 E .c r A ii 1 M W 9 46 go bu vi 0 W A ae n C 10 93 5 A rb n C W 9 .c 01 ru ii 1 M W 00 57 A .g un ov 2 M 0 A 1 bi n i C 1 M W 0 1 . or i C 0 04 5 0 iru i C 1 0 7 A d b ov 2 3 0 A. b un ii 1 1 M W 9 3 9 . am un i C 0 W0 3 W 4 . ir n i C 1 1 M W 9 9 3 A a o o 1 M 00 M 4 A b u i C 1 9 0 0 . lti cl v 9 4 7 . ir n i 5 W W 9 0 5 A c va e i W 0 6 A b u i C 7 M E 9 9 . y g s M 0 2 7 . ir n ii 5 M 9 4 9 A p a W 0 4 A b u C 8 s W 0 9 4 4 . s n an ka 0 5 . ir n s C li M W 2 4 8 A m e eu s n M 0 6 A b ru li l is M 9 6 4 . e u s d W 7 . i l s y ll s M W 9 9 9 A c d u M 0 4 2 A b y li h y li W 2 y r o l 0 0 . h l is M 9 8 3 1 A .c a h x i W 0 1 2 yl p h y l 3 i M W 1 5 a e C M A p . l 2 W 9 8 3 A . y n b 2 W 0 7 . h A .p h y d M 8 1 3 c a e a rx 5 0 0 0 A p p C e W W 9 6 . y n u e M 0 6 . A . h a s WE W 9 8 3 3 A s a C W 2 0 p i b a M 9 e e s s 0 0 A 4 A . n e i W E 9 8 1 1 A . n u 1 0 5 7 d a s 9 a n e C C

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64 A molecular phylogeny of Agrodiaetus

7 7 5

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W . f e e e . k e a a 5 . m i e M . M M 3 d e l u 8 T . u 9 . b b r . p p z e 9 2 A 4 A o c 1 W 9 A . h 4 W . . u u e c i 7 W A A e 9 5 t A . c i r 3 3 h 3 A h i s C 1 C 6 A A h h 0 6 u n 1 r 8 0 9 A . . t e 9 4 6 c 0 C e 5 i C s 0 . a C 1 9 1 2 g r u 0 . 7 8 1 2 A 7 M 9 3 1 A . s o 0 p e u 5 1 8 . 4 5 A u b e s M 2 C 6 . 1 s 9 9 t l ar e p 6 2 A a M W A 2 9 a ic u 1 3 K 9 4 2 . e 0 b s W 6 2 5 A e C1 m 9 2 0 n s c h 2 7 4 8 0 5 . i n i W 3 . m 9939 9 2 4 A i s r u 0 . 0 2 4 s J M e r 9 9 s 0 2 0 p 9 5 1 A n a u c 0 V A i a 9 0 9 . n r i u C 0 u a 0 9 2 9 . r 2 i E P W 4 W 2 y W 9 b s M 1 0 . . a s W0 E 9 8 2 2 a l u r c C 0 r r W 8 7 A i 0 0 m e r . W E 3 A . b M W 0 06 2 e y r g t 9 9 8 3 e l u s i 8 0 M W p M 3 W 9 4 . s u g u M r i W W 9 8 A v 0 1 4 o p u M 9 8 e b W 3 n y M 9 2 l u C s 9 m u s W D W 9 1 A . o 2 5 0 1 A r a l i s M 0 n i n 9 9 V g a l M W 9 0 1 e b k 2 M 5 4 A ed u s A W 0 5 8 A . l o n e 9 0 . i ic M W 0 e a a C A 8 . a a o s M 8 9 5 A . r m i M W 0 9 . a u M W u r r an a 0 7 . A c g lc n o n M W 9 9 266 0 51 0 A a u d tr W 9 a r s M .s 7 M 8 1 A r . a e e u n D 0 0 0 1 6 .c h a s 1 9 2 A t a me s d s M W 0 0 1 A c u M W 8 .i a g A E 0 1 A .s m th C M 9 1 7 .t s x e ti o M W 0 3 1 a e W 0 o a e s s W 0 0 0 9 A .h ra a i 9 M W 8 8 r u r M W 0 0 a l d 1 W 9 9 1 5 A u r rx ti a W 0 0 1 A z u a C 95 8 2 L .a le ic e s M 8 8 6 2 . a v i 9 1 2 9 . n n u s M W 9 9 2 3 A .p p ad 95 4 2 L o t s s M D 8 0 1 A a v M 3 L 8 .o s e i A W 9 0 3 .z p s 75 W 6 . L s sm ro s 0 5 7 A a lli C M 0 L co . s s M W 2 2 .z y s W 1 . r sy m a M W 0 01 26 A h lli M 0 0 co y r a r M E 0 2 4 .p y 5 W0 1 34 r do ia r 9 7 A h is 8 M 1 01 L yd n c W 9 3 4 .p yll C 25 O W 0 8 .c o iu a MWW 99 17 A h llis C K M 01 59 L o ni s M 9 52 .p hy sii 24 96 W 0 L .co rid us W 9 4 A .p s C 25 02 0 92 .a r o M W 00 6 A rdu sii C 2 11 L lb id n M 03 8 .fi us sis L.c 16 .al ica on W98 34 A ird en MW ae L bic n M D 00 13 .f in 7 M 9 le .c a s A W 94 7 A rtv 1 W0 96 stis or ns M 9 4 A.a is C 25 M 10 12 si ido W 92 8 act sii C W9 11 L.c mu n M W9 05 A. us s M 82 L.b or s M 99 62 fird cti W9 78 el ido W 81 A. uda M 96 L.b larg n M W9 234 pse sii W0 08 L ella us M 00 A. dus 25 046 .be rgu MW 009 A.fir rti C 3 M 9 L. llarg s D98 51 igbe s C2 W00 bella us A 001 A.s ridate M 116 rgu MW 8285 mith sis M W982 P.ar s W9 3 A. anen W990 20 P gus M 9820 .senn C14 13 N.c .loew MW 621 A damon MW98 oeles ii E02 A.iphi 029 M tina W 00269 idamon MW00 .daphn MW 8 A.iph C17 076 M.d is W0032 izinensis MW002 aphnis M 0539 A.d 20 90 M.marci MW0 .gorbunovi C JC000 da MW00177 A 63 P.icarus 78 A.gorbunovi JC00061 P.andr MW001 onicus MW00129 A.gorbunovi C20 MW01025 P.icarus WE02674 A.gorbunovi JC00042 P.eroides MW00330 A.ps .menelaos eudoxerxes C15 JC00029 P MW00179 A.cyan roides forsteri MW9 eus MW00530 P.e arus 9094 A.cyane 00412 P.ic MW99448 us C17 MW .icarus MW9 A.cyaneu 02006 P las 9006 A s C18 MW P.dory WE024 .firduss 99014 orylas MW 54 A.m ii C30 MW 19 P.d es M 9946 ofidii 010 ersit W99 5 A.ka MW 3 P.th ites MW 353 nduli 0108 thers a M 992 A.alt C25 MW 2 P. fatim W9 40 A ivaga 0030 N. ona AD 905 .alti ns C MW 9301 .cor s M 980 7 A vaga 22 W9 4 N ndu W9 12 .me ns C M 050 ma us MW 81 A.a rhab 21 W0 P.a and ia M 981736 ltiva a C M 47 am D W A.w gan 17 90 P. ornellia S 983 0 A ag s W9 01 P.c rne ia D 00 13 .m ner M 020 8 co el M S0 001 A ara i C W P. orn us W 10 A .se sc 18 M 990335 .c hin s MW 0 01 .pos rta hi MW 82 P rr inu n M 05 A vu C1 9 64 my rh o M W 00 47 .ip eidonlen 6 MW 82 P. r ed ri W 0 44 A hi si 9 0 my .a he n M 03 4 .b ge ides s C W 55 P. P c o M W 00 9 A iru ni 20 M 9 7 6 es m on M W 0 4 3 .b n de 9 53 32 . da e 0 01 09 A. iru ii C s W 9 0 P . am a M W 0 0 A er n 1 M 9 0 39 A .d s 8 M W 0 2 2 . s ii 0 W W 0 3 A da 3 s M W 0 0 6 A ca ch C1 M M 0 61 e C si 9 0 07 7 . e o 1 0 9 46 e n 1 M W 0 1 2 A bi ru ffi JC 9 5 um a e s M W 9 0 0 .b ru le i C W 9 h s i C a 0 M W 9 3 1 A i n u 1 9 . a n is lc 5 M 9 5 4 .b ru ii s 4 M A d a t a 0 W 9 9 6 7 A i n C C W 5 e ro s n C 2 1 M W 91 5 5 A .d ru ii 1 2 M 0 m a e e s C 2 s M W 9 5 a n C 1 0 6 u . c n e M 9 9 9 A .p r i A l m a s C i W W 2 3 i i 1 9 h .a . lc ti e a W 9 9 5 A .t o u C 0 9 0 A s is g s e e M 9 8 0 0 a s s 1 A. 4 A a t l s i M E 0 3 A .t n th 1 W 1 2 n e s u in s a e M W 0 8 1 a M 9 0 1 e c f s 1 M W 0 0 0 0 9 A . n k u 0 3 7 l e . a e s 2 M W 2 tur e m 5 0 1 a c . r n s 3 2 M 0 1 4 3 A . k r 9 2 m . l A i 4 M W 9 6 t e i u C 0 8 A b C 2 M W 0 7 9 u c 9 X a a e 4 i M W 9 A . r s J 0 9 . A . 7 a . r C 4 i M 9 2 6 i r i s M W 9 6 p c i W 0 3 f s C t W W 8 2 A . c C h . b i W 9 3 6 i 5 A A W h us W o r s s W W 8 p i W c 5 C i 0 W 99 0 A . c 8 u i i M t 1 a o 0 W 8 0 2 i A t a 3 9 A i

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MW M Fig. 64. Maximum Parsimony Majority Rule Consensus Tree (ITS-2)

65 Chapter 3

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66 A molecular phylogeny of Agrodiaetus

100 Agrodiaetus 100 MW00076 M.daphnis 70 100 MW98029 M.daphnis MW00290 M.marcida 100 MW00504 N.corona 100 MW99301 N.fatima 99 100 MW02001 P.amandus 100 MW99047 P.amandus 100 MW00302 P.thersites 100 MW01083 P.thersites MW01019 P.dorylas 100 MW99014 P.dorylas 100 MW02006 P.icarus 100 100 JC00029 P.menelaos 100 JC00051 A.menelaos 100 JC00042 P.eroides 100 MW00530 P.eroides forsteri MW00412 P.icarus 100 100 MW01025 P.icarus 100 JC00061 P.andronicus 100 JC00063 P.icarus JC00039 P.escheri MW00326 P.aedon 100 100 MW99537 P.myrrhinus 100 MW99550 P.myrrhinus 100 MW98264 P.cornelia MW98235 P.cornelia 100 100 MW99038 P.cornelia MW99013 N.coelestina MW98270 A.anteros MW01061 A.cramera 100 100 MW99001 A.torulensis 100 MW99097 A.isauricus Aricia 99 JC00055 A.artaxerxes MW01048 A.agestis 100 100 MW02033 A.agestis MW00024 V.alcedo MW00525 C.semiargus 57 100 MW02034 C.semiargus MW98220 P.loewii MW00116 P.argus 58 83 MW00469 L.bellargus 78 100 MW99608 L.bellargus 100 MW98278 L.bellargus 100 MW01011 L.bellargus 100 MW98129 L.ossmar 51 MW98185 L.ossmar 100 100 MW98228 L.syriaca 100 MW99514 L.corydonius Lysandra MW99536 L.corydonius 100 MW01092 L.albicans 0 MW01059 L.albicans 100 MW01034 L.coridon 55 MW01018 L.coridon 55 MW01116 L.coridon 100 MW99612 L.coridon 100 69 OK96022 L.caelestissimus MW00517 V.morgianus MW99134 A.eumedon MW99163 P.pylaon 100 MW99303 K.eurypilus MW99018 A.pyrenaicus MW02025 T.theophrastus Fig. 66. MP Consensus Tree of the combined data set (ITS-2 & COI) excl. Agrodiaetus

67 Chapter 3

100 MW98136 A.w agneri C18 100 MW98139 A.w agneri C19 100 MW98313 A.sertavulensis C20 76 MW98170 A.maraschi C16 100 MW99240 A.altivagans C21 100 AD98012 A.altivagans 100 MW99057 A.merhaba C17 100 MW99353 A.altivagans C22 99 MW99465 A.kanduli C25 100 WE02454 421 A.mofidii WE02621 A.sennanensis 100 MW99094 A.cyaneus C17 75 MW99448 A.cyaneus C18 99 MW00179 A.cyaneus 100 MW00330 A.pseudoxerxes C15 99 WE02674 A.gorbunovi 100 MW00178 A.gorbunovi 100 MW00129 A.gorbunovi C20 MW00177 A.gorbunovi C20 poseidon-group MW98203 A.mithridates C23 90 MW98285 A.sigberti C25 53 MW99006 A.firdussii C30 MW00234 A.firdussii C25 83 MW99058 A.artvinensis C25 55 AD98009 A.pseudactis 53 MW98162 A.actis C17 MW99413 A.firdussii C25 100 75 MW00151 A.firdussii 100 MW99247 A.firdussii C24 MW98097 A.ernesti C18 MW98079 A.lycius 100 MW98189 A.hopfferi C16 53 MW99408 A.hopfferi C15 MW98180 A.poseidon C19 66 MW99501 A.putnami C25 100 MW98138 A.poseidon C20 100 100 MW98154 A.poseidon C21 DS00001 A.poseidonides 100 WE02491 A.achaemenes WE85001 A.shahrami MW00393 A.erschoffii C14 100 AD98036 A.phyllis 100 MW99174 A.phyllis 59 MW00348 A.phyllis C85 100 100 MW00452 A.phyllis C75 74 MW00262 A.klausschuriani C56 65 WE02451 A.tenhageni WE00002 A.glaucias erschoffii-group 99 MW00101 A.darius 61 MW00347 A.posthumus C85 MW00409 A.caeruleus C20 100 MW00444 A.birunii C11 100 MW00547 A.birunii C10 100 MW00072 A.birunii C11 58 MW00102 A.birunii C11 63 MW00267 A.birunii C10 DS01001 A.iphigenides 100 MW98049 A.iphigenia C14 MW99009 A.iphigenia C12 100 MW99135 A.turcicus C25 76 MW99203 A.turcicus C24 100 MW98103 A.iphicarmon 100 51 MW99372 A.baytopi C27 iphigenia-group MW99565 A.tankeri 100 MW99559 A.tankeri MW99309 A.baytopi 100 MW00176 A.rovshani 100 100 WE02662 A.rovshani 100 MW99292 A.pierceae C21 MW99341 A.pierceae C23 100 MW98240 A.theresiae C59 MW98294 A.guezelmavi 100 MW99393 A.antidolus C42 72 100 MW99406 A.antidolus C44 100 MW99286 A.kurdistanicus C55 100 MW00226 A.femininoides C27 100 100 WE02671 A.femininoides C27 100 WE02614 A.morgani C27 WE02591 A.peilei C39 MW98205 A.dama 100 MW00110 A.elbursicus C18 100 MW00316 A.elbursicus C17 100 MW00051 A.elbursicus C17 100 92 100 MW00058 A.elbursicus C18 MW00232 A.elbursicus C18 carmon-group 100 WE02661 A.arasbarani MW00127 A.paulae C17 100 WE02531 A.zarathustra C22 MW00032 A.hamadanensis C22 90 AD98001 A.surakovi 100 MW98009 A.carmon MW98261 A.schuriani C80 100 AD98018 A.ninae 100 MW99508 A.ninae C34 99 MW99479 A.turcicola C20 100 MW99226 A.zapvadi 100 MW99374 A.zapvadi C19 AD98024 A.huberti 88 MW99095 A.huberti C34 100 66 MW99552 A.huberti C33 MW00539 A.dizinensis C17 MW00269 A.iphidamon C14 100 iphidamon-group 100 MW00328 A.iphidamon 58 MW99382 A.demavendi C70 56 WE02677 A.demavendi 100 MW00185 A.demavendi C70 MW00015 A.demavendi C70 JC01014 A.admetus JC00043 A.ripartii 91 JC00045 A.nephohiptamenos 100 MW99407 A.demavendi C60 admetus-group MW99196 A.ripartii C90 100 WE02535 A.lorestanus 92 MW00189 A.demavendi 100 100 MW99105 A.demavendi C66 100 MW01014 A.ripartii C90 69 WE02431 A.khorasanensis 100 MW00064 A.valiabadi MW00498 A.valiabadi C23 JM00001 A.fabressei 100 MW01001 A.ainsae MW01039 A.fabressei 100 MW01053 A.ainsae 100 MW01107 A.fulgens 100 MW99591 A.humedasae C38 71 100 MW99605 A.humedasae JC00040 A.aroaniensis dolus-group MW98172 A.menalcas 100 MW98315 A.alcestis C20 100 MW98212 A.alcestis C21 100 MW00231 A.alcestis C19 100 MW99471 A.dantch.Xmenalcas C50 100 MW99319 A.dantchenkoi C42 100 MW99276 A.dantchenkoi C42 100 MW99164 A.interjectus C31 MW99274 A.dantchenkoi C42 MW99546 A.damon 100 MW99613 A.damon Outgroup Fig. 67. MP Consensus Tree of the combined data set (ITS-2 & COI) of Agrodiaetus

68 A molecular phylogeny of Agrodiaetus

55 Agrodiaetus & Polyommatus (partim) AF170864 Euphilotes bernardino MW01061 A.cramera 100 MW99001 A.torulensis MW99097 A.isauricus 61 87 MW01048 A.montensis 57 MW02033 A.montensis 77 JC00055 A.artaxerxes MW99028 A.agestis 61 MW00020 A.agestis 96 JC00057 A.artaxerxes JC00062 A.agestis MW00024 V.alcedo 100 MW99430 V.alcedo MW00116 P.argus 99 MW99124 P.christophi MW00326 P.aedon MW99538 P.myrrhinus 64 MW99537 P.myrrhinus 100 MW99550 P.myrrhinus 100 MW98264 P.cornelia MW98235 P.cornelia 96 MW99038 P.cornelia MW98228 L.syriaca 99 MW99140 L.corydonius 56 MW99514 L.corydonius 55 MW98185 L.ossmar 94 MW99042 L.corydonius 93 MW99536 L.corydonius MW98129 L.ossmar MW00469 L.bellargus 64 MW98278 L.bellargus 98 99 MW99435 L.bellargus MW99608 L.bellargus MW01011 L.bellargus OK99001 L.gennargenti 62 MW01116 L.coridon OK96022 L.caelestissimus 54 MW01103 L.coridon MW99612 L.coridon 65 MW01092 L.albicans MW01059 L.albicans MW01018 L.coridon MW01034 L.albicans MW00517 V.morgianus MW00525 C.semiargus 100 MW02034 C.semiargus MW98220 P.loewii MW98270 A.anteros MW99013 N.coelestina 59 WE00003 N.diana MW99018 A.pyrenaicus MW99134 A.eumedon MW99163 P.pylaon 50 98 MW99303 K.eurypilus AF170863 Apodemia mormo MW99520 L.thetis MW98230 L.tityrus MW98094 L.thersamon MW02009 L.phlaeas 55 MW00119 L.candens 91 MW99515 L.virgaureae MW00044 L.asabinus MW00017 L.alciphron 100 MW02007 L.alciphron MW00497 F.quercus MW01022 L.boeticus 100 MW02028 L.boeticus 64 MW98265 L.boeticus 50 MW01023 S.pirithous MW99158 S.hyrcanicum 96 MW01027 S.esculi 59 MW99398 S.myrtale MW01120 C.marshalli MW02008 C.argiolus 100 MW99084 C.argiolus MW02021 Z.knysna MW02025 T.theophrastus 56 MW02031 P.abencerragus 97 MW99080 P.vicrama MW98002 C.osiris MW99045 M.arion MW99221 T.endymion MW99425 C.trochylus 0 MW02024 I.feisthamelii JC02001 I.podalirius 100 99 AF170873 Iphiclides podalirius 81 MW01114 I.feisthamelii AF044007 Colias eurytheme AF170860 Coenonympha tullia AF044006 Papilio machaon Fig. 68. Bootstrap Tree (COI) excl. Agrodiaetus and closely related Polyommatus species

69 Chapter 3

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70 A molecular phylogeny of Agrodiaetus

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71 Chapter 3

95 Agrodiaetus MW00530 P.eroides forsteri 90 JC00029 P.menelaos 60 JC00051 P.menelaos 100 JC00042 P.eroides MW00412 P.icarus MW01025 P.icarus 87 JC00061 P.andronicus 59 JC00063 P.icarus JC00039 P.escheri 73 MW00290 M.marcida MW00076 M.daphnis 100 89 MW98029 M.daphnis 100 MW00302 P.thersites MW01083 P.thersites 81 MW01019 P.dorylas 84 99 MW99014 P.dorylas 73 MW00504 N.corona MW99301 N.fatima 56 MW02001 P.amandus 95 MW99047 P.amandus MW02006 P.icarus 95 MW00326 P.aedon 100 MW99537 P.myrrhinus 100 MW99550 P.myrrhinus MW98264 P.cornelia 100 MW98235 P.cornelia 100 MW99038 P.cornelia MW99013 N.coelestina MW98270 A.anteros MW01061 A.cramera 84 100 MW99001 A.torulensis 100 MW99097 A.isauricus Aricia JC00055 A.artaxerxes 67 MW01048 A.agestis 84 86 MW02033 A.agestis MW00024 V.alcedo MW00116 P.argus 55 MW00469 L.bellargus 82 MW99608 L.bellargus 97 MW98278 L.bellargus 0 80 MW01011 L.bellargus 98 MW98129 L.ossmar 96 MW98185 L.ossmar 90 MW98228 L.syriaca 100 MW99514 L.corydonius MW99536 L.corydonius Lysandra 53 MW99612 L.coridon 51 OK96022 L.caelestissimus 99 MW01116 L.coridon MW01092 L.albicans MW01059 L.albicans 52 MW01018 L.coridon MW01034 L.albicans MW00517 V.morgianus MW00525 C.semiargus 100 MW02034 C.semiargus MW98220 P.loew ii MW99018 A.pyrenaicus MW99134 A.eumedon MW99163 P.pylaon 100 MW99303 K.eurypilus MW02025 T.theophrastus

Fig. 71. Bootstrap Tree of the combined data set (ITS-2 & COI) excl. Agrodiaetus

72 A molecular phylogeny of Agrodiaetus

57 MW98136 A.wagneri C18 MW98139 A.wagneri C19 51 MW98170 A.maraschi C16 MW98313 A.sertavulensis C20 MW99240 A.altivagans C21 79 AD98012 A.altivagans 53 MW99057 A.merhaba C17 MW99353 A.altivagans C22 MW99465 A.kanduli C25 MW00330 A.pseudoxerxes C15 WE02674 A.gorbunovi 100 98 MW00178 A.gorbunovi 96 MW00129 A.gorbunovi C20 MW00177 A.gorbunovi C20 MW00179 A.cyaneus MW98203 A.mithridates C23 MW99094 A.cyaneus C17 99 MW99448 A.cyaneus C18 WE02621 A.sennanensis poseidon-group WE02454 421 A.mofidii MW99058 A.artvinensis C25 70 MW98285 A.sigberti C25 MW99006 A.firdussii C30 99 MW98162 A.actis C17 53 MW00234 A.firdussii C25 AD98009 A.pseudactis MW99413 A.firdussii C25 MW00151 A.firdussii 68 MW99247 A.firdussii C24 MW98079 A.lycius MW98097 A.ernesti C18 MW98180 A.poseidon C19 MW99501 A.putnami C25 72 MW98138 A.poseidon C20 70 85 MW98154 A.poseidon C21 MW98189 A.hopfferi C16 67 MW99408 A.hopfferi C15 83 MW99292 A.pierceae C21 MW99341 A.pierceae C23 92 MW98240 A.theresiae C59 MW98294 A.guezelmavi 76 MW99393 A.antidolus C42 58 MW99406 A.antidolus C44 59 MW99286 A.kurdistanicus C55 73 MW00226 A.femininoides C27 95 WE02671 A.femininoides C27 69 WE02614 A.morgani C27 WE02591 A.peilei C39 MW98205 A.dama 87 MW00110 A.elbursicus C18 89 MW00316 A.elbursicus C17 59 MW00051 A.elbursicus C17 74 54 MW00058 A.elbursicus C18 84 MW00232 A.elbursicus C18 carmon-group 66 WE02661 A.arasbarani MW00127 A.paulae C17 93 WE02531 A.zarathustra C22 MW00032 A.hamadanensis C22 63 AD98001 A.surakovi 66 MW98009 A.carmon MW98261 A.schuriani C80 100 MW99226 A.zapvadi MW99374 A.zapvadi C19 80 AD98018 A.ninae 72 MW99508 A.ninae C34 57 MW99479 A.turcicola C20 AD98024 A.huberti MW99095 A.huberti C34 95 51 MW99552 A.huberti C33 99 WE02491 A.achaemenes WE85001 A.shahrami WE02451 A.tenhageni WE00002 A.glaucias MW00393 A.erschoffii C14 MW00262 A.klausschuriani C56 97 AD98036 A.phyllis 57 99 MW99174 A.phyllis MW00348 A.phyllis C85 erschoffii-group 93 MW00452 A.phyllis C75 64 57 MW00101 A.darius MW00347 A.posthumus C85 MW00409 A.caeruleus C20 52 MW00547 A.birunii C10 69 MW00444 A.birunii C11 MW00267 A.birunii C10 74 MW00072 A.birunii C11 MW00102 A.birunii C11 DS00001 A.poseidonides DS01001 A.iphigenides 100 MW00064 A.valiabadi MW00498 A.valiabadi C23 JM00001 A.fabressei 99 MW01001 A.ainsae MW01039 A.fabressei MW01053 A.ainsae 97 91 MW01107 A.fulgens 100 MW99591 A.humedasae C38 80 MW99605 A.humedasae JC00040 A.aroaniensis dolus-group MW98172 A.menalcas 68 MW98315 A.alcestis C20 68 MW98212 A.alcestis C21 MW00231 A.alcestis C19 83 MW99471 A.dantch.Xmenalcas C50 60 MW99319 A.dantchenkoi C42 58 MW99276 A.dantchenkoi C42 63 MW99164 A.interjectus C31 95 MW99274 A.dantchenkoi C42 WE02431 A.khorasanensis MW01014 A.ripartii C90 97 MW00189 A.demavendi 73 MW99105 A.demavendi C66 54 WE02535 A.lorestanus 73 MW99196 A.ripartii C90 MW99407 A.demavendi C60 100 JC00043 A.ripartii admetus-group JC00045 A.nephohiptamenos JC01014 A.admetus WE02677 A.demavendi MW99382 A.demavendi C70 100 MW00015 A.demavendi C70 MW00185 A.demavendi C70 70 MW98103 A.iphicarmon 70 MW99372 A.baytopi C27 MW99565 A.tankeri MW99559 A.tankeri baytopi-species-group 76 MW99309 A.baytopi 89 MW00176 A.rovshani 78 99 WE02662 A.rovshani MW00539 A.dizinensis C17 MW00269 A.iphidamon C14 98 iphidamon-group 100 MW00328 A.iphidamon MW98049 A.iphigenia C14 99 MW99009 A.iphigenia C12 MW99135 A.turcicus C25 100 MW99203 A.turcicus C24 MW99546 A.damon 99 MW99613 A.damon Outgroup Fig. 72. Bootstrap Tree of the combined data set (ITS-2 & COI) of Agrodiaetus

73 Chapter 3

Statistical parsimony networks The following table gives an overview of the statistical parsimony networks calculated from the COI data set. The parsimony connection limit was 11 steps for a parsimony probability of 95%. Such a connection limit would only allow the calculation of networks for populations of the same species or very closely related species. In the case of Agrodiaetus 33 different networks were obtained with this connection limit. On the other hand, calculation time increases more than exponentially with an increasing connection limit which rendered the calculation of a complete network for Agrodiaetus unfeasible. Instead, networks were calculated for the main Agrodiaetus clades which were obtained from the combined COI- and ITS2-analysis (i.e. all clades with more than two sequences). The connection limit was increased until all haplotypes were connected, with the only exception of A. klausschuriani in the erschoffii-group, a taxon which was so distant that it seemed more reasonable to exclude it from the analysis. The number of steps required is stated in Tab. 9.

Tab. 9. Connection limits in the COI network analysis Group sequences excluded taxa haplo- steps fig. types Iphiclides 4 3 14 Fig. 73 Polyommatus-aedon-group 7 6 31 Fig. 74 Aricia-agestis-group 12 11 13 Fig. 75 Lysandra 21 19 18 Fig. 76 Agrodiaetus-admetus-group 27 19 15 Fig. 77 Agrodiaetus-menalcas-group 25 17 16 Fig. 78 Agrodiaetus-carmon-group 44 34 20 Fig. 79 Agrodiaetus-erschoffii-group 24 A. klausschuriani 21 18 Fig. 80 Agrodiaetus-iphigenia-group 16 15 18 Fig. 81 Agrodiaetus-poseidon-group 49 39 13 Fig. 82

The number of haplotypes is always lower than the number of sequences, because some sequences turned out to have the same haplotype (or only differed in ambiguities). In these cases, the graph shows only one representative haplotype and the size of the bounding box is increased according to the frequency of this haplotype. Tab. 10 lists those representative haplotypes together with the codes of the collapsed haplotypes. Usually these collapsed haplotypes belong to the same taxon as the reference specimen, but in some cases they were designated to different taxa which is indicated with an asterisk (*). Identical COI haplotypes were found in the following different taxa. • Iphiclides feisthamelii and Iphiclides podalirius • Agrodiaetus zapvadi and Agrodiaetus turcicola • Agrodiaetus kurdistanicus and Agrodiaetus antidolus • Agrodiaetus peilei and Agrodiaetus morgani • Agrodiaetus fabressei and Agrodiaetus ainsae • Agrodiaetus cyaneus and Agrodiaetus merhaba • Agrodiaetus sertavulensis and Agrodiaetus wagneri This indicates that taxa could be identical genetically or that gene flow persists between them. If the bounding box is a rectangle, this haplotype was calculated to have the highest outgroup probability.

In several cases the networks provide information which could not be obtained from the phylogenetic trees. Extant ancestral haplotypes appear in the Polyommatus-aedon-group

74 A molecular phylogeny of Agrodiaetus

(Polyommatus myrrhinus, Fig. 74), in the Agrodiaetus-menalcas-group (Agrodiaetus dantchenkoi and Agrodiaetus fabressei, Fig. 78), in the Agrodiaetus-carmon-group (Agrodiaetus surakovi, Fig. 79), and in the Agrodiaetus-poseidon-group (Agrodiaetus putnami and A. pseudactis, Fig. 82). Crosslinks between haplotypes of different taxa indicate that gene flow persists between them. This appears to be the case in the species pairs Lysandra corydonius and Lysandra ossmar (Fig. 73), Lysandra coridon and Lysandra albicans (Fig. 76), Agrodiaetus demavendi and Agrodiaetus ripartii (Fig. 77), and Agrodiaetus huberti and Agrodiaetus ninae (Fig. 79).

Tab. 10. Collapsed haplotypes in the COI network analysis Group Reference Taxon specimen codes of collapsed haplotypes specimen * taxon different from the reference taxon code Iphiclides MW01114 I. feisthamelii AF170873* Aricia JC00062 A. agestis MW00020 Lysandra MW00469 L. bellargus MW99608 MW99140 L. corydonius MW99514 P. aedon MW99537 P. myrrhinus MW99550 A. carmon MW00051 A. elbursicus MW00056 MW00058 MW00232 MW99226 A. zapvadi MW99314* MW99374 MW99286 A. kurdistanicus MW99376* MW99393 MW99473* WE02591 A. peilei WE02614* MW00032 A. hamadanensis MW00001 A. dolus JM00001 A. fabressei MW01001* MW01039 MW00064 A. valiabadi MW01039 MW00229 A. alcestis MW00231 MW99274 A. dantchenkoi MW99276 MW99319 MW99320 MW99471 A. iphigenia MW99009 A. iphigenia MW99170 A. erschoffii MW0060 A. birunii MW00060 MW00072 MW00102 MW00476 A. admetus JC00045 A. nephohiptamenos JC00046 MW00183 A. demavendi MW00185 MW00186 MW99068 A. ripartii MW99196 MW99104 A. demavendi MW99141 MW99263 A. ripartii MW99264 MW99382 A. demavendi MW99381 WE02677 A. poseidon MW99006 A. firdussii MW00125 MW00129 A. gorbunovi MW00177 WE02675 MW99413 A. firdussii MW00151 MW99422 MW99448 A. cyaneus MW00179 MW99059* MW99449 MW98313 A. sertavulensis MW98139* MW99061 A. putnami MW99507

Fig. 73. COI-Parsimony-Network of Iphiclides

75 Chapter 3

Fig. 74. COI-Parsimony-Network of the P.aedon-group

Fig. 75. COI-Parsimony-Network of the Aricia agestis-group

76 A molecular phylogeny of Agrodiaetus

Fig. 76. COI-Parsimony-Network of Lysandra

77 Chapter 3

Fig. 77. COI-Parsimony-Network of the Agrodiaetus admetus-group

78 A molecular phylogeny of Agrodiaetus

Fig. 78. COI-Parsimony-Network of the Agrodiaetus-menalcas-group

79 Chapter 3

Fig. 79. COI-Parsimony-Network of the Agrodiaetus carmon-group

80 A molecular phylogeny of Agrodiaetus

Fig. 80. COI-Parsimony-Network of the Agrodiaetus erschoffii-group

81 Chapter 3

Fig. 81. COI-Parsimony-Network of the Agrodiaetus iphigenia-group

82 A molecular phylogeny of Agrodiaetus

Fig. 82. COI-Parsimony-Network of the Agrodiaetus-poseidon-group

83 Chapter 3

Fig. 83. ITS2-Parsimony-Network of Agrodiaetus (grouped) incl. connected outgroups

84 A molecular phylogeny of Agrodiaetus

Fig. 84. ITS2-Parsimony-network of Agrodiaetus (excluding outgroups)

85 Chapter 3

Fig. 85. ITS2-Parsimony-Network of Lysandra

For the ITS-2 dataset the connection limit was 12 steps for a parsimony probability of 95%. The parsimony analysis of all Polyommatiti ITS-2 sequences with this connection limit produces 8 different networks, if gaps are counted as missing characters. Most taxa, including all taxa of subgenus Agrodiaetus are found in one network. If A. surakovi is excluded from the analysis (due to more than 10% missing character information), A. hamadanensis is disconnected from the carmon-group (Fig. 83). This network includes also most other taxa of the genus Polyommatus. Exceptions are the genus Lysandra (Fig. 85) and Polyommatus thersites which form two separate networks. The genus Agrodiaetus is only connected via Agrodiaetus damon to the outgroup, and the shortest distance is to Polyommatus cornelia (4 steps). A. damon is further connected (with a maximum of 4 steps) to the admetus-group (3 steps), to A. valiabadi (4 steps), the other members of the dolus-group (4 steps) and to A. iphidamon (3 steps). A. iphidamon is the stem species of all other groups which closely resemble the groups found in the combined Bayesian analysis with the following exceptions: A. klausschuriani connects to A. hopfferi in the poseidon-group and A. putnami is not connected to the poseidon-group but to A. iphidamon and to A. birunii in the erschoffii-group. A. glaucias forms a group together with A. tenhageni which is directly connected to A. iphidamon and not to the other members of the erschoffii-group. A. phyllis is also split from the erschoffii-group. The complete network of ITS-2 haplotypes is presented in Fig. 84. A list of collapsed haplotypes can be found in Tab. 11.

86 A molecular phylogeny of Agrodiaetus

Tab. 11. Collapsed haplotypes in the ITS-2 network analysis Net- Reference Taxon specimen codes of collapsed haplotypes work specimen * taxon different from the reference taxon code # reference specimen for further collapsed haplotypes bold: also collapsed if gaps are coded as 5th character 1 AD98012 Agrodiaetus altivagans MW99353 1 AD98036 Agrodiaetus phyllis MW00452 MW99174 1 JC00043 Agrodiaetus ripartii JC00045* JC01014* WE02431* 1 MW00051 Agrodiaetus elbursicus MW00110 1 MW00064 Agrodiaetus valiabadi MW00498 1 MW00129 Agrodiaetus gorbunovi MW00177 MW00178 1 MW00176 Agrodiaetus rovshani WE02662 1 MW00189 Agrodiaetus demavendi MW99105 MW99196* 1 MW00226 Agrodiaetus femininoides WE02614* 1 MW00231 Agrodiaetus alcestis MW98212 MW98315 MW99164* MW99274# 1 MW00234 Agrodiaetus firdussii MW98162* MW99058* MW99247 1 MW00328 Agrodiaetus iphidamon MW99372# 1 MW01001 Agrodiaetus ainsae MW01039* MW01053 1 MW98136 Agrodiaetus wagneri MW98139 1 MW98172 Agrodiaetus menalcas MW99471 1 MW99009 Agrodiaetus iphigenia MW99309 MW99565 1 MW99135 Agrodiaetus turcicus MW99203 1 MW99274 Agrodiaetus dantchenkoi MW99276 MW99319 1 MW99372 Agrodiaetus baytopi WE02491* WE85001* 1 MW99546 Agrodiaetus damon MW99613 1 MW99591 Agrodiaetus humedasae MW99605 1 JC00029 Polyommatus menelaos JC00042* JC00051 1 JC00061 Polyommatus andronicus JC00063* 1 MW00412 Polyommatus icarus MW00530* 1 MW99537 Polyommatus myrrhinus MW99550 7 MW00469 Lysandra bellargus MW99608 7 MW01018 Lysandra coridon MW01034* MW01092# 7 MW01092 Lysandra coridon MW01116 8 JC00055 Aricia artaxerxes MW01048*

In the case of gaps counted as 5th character most Agrodiaetus were also connected in one network, only A. antidolus, A. kurdistanicus and A. femininoides together formed a separate network and three other taxa (A. hamadanensis, A. maraschi and A. paulae) remained unconnected.

Discussion

Comparisons with traditional systematics The basal splits in the COI gene tree correspond to the traditional classification of Lycaenidae (as far as these are represented in the Palaearctic region) into three subfamilies (Theclinae, , ) according to ELIOT (1973) or tribes (Theclini, Lycaenini, Polyommatini) according to HESSELBARTH et al. (1995) and support modern views of SCOTT & WRIGHT (1990) and FIEDLER (1991b) that Lycaenini represent the most ancient subdivision of those three taxa. The clades within the tribus Polyommatini also largely follow current subdivisions into subtribes according to HESSELBARTH et al. (1995) and the monophyly of Polyommatiti is supported. Although taxon sampling of outgroups was necessarily coarse, the resolution achieved with the available molecular data was surprisingly good. The only exception appears to be Euphilotes bernardino (Barnes & McDunnough, 1917) which is

87 Chapter 3 sometimes treated as a subspecies of Euphilotes battoides (Behr, 1906). The trees surprisingly indicate that this species should be included in the Polyommatiti (Polyommatus section sensu ELIOT 1973). However, in his original description of the genus Euphilotes, MATTONI (1977) treated this taxon (together with Philotiella) as a relative of Philotes. Species currently assigned to Euphilotes and Philotiella, respectively, had previously been treated under the genus name Philotes, and ever since MATTONI's split all these taxa have been regarded, on the grounds of similarities in male genitalia and wing morphology, as belonging to the Glaucopsyche section sensu ELIOT (1972) (which was later split into two subtribes, Glaucopsychiti and Scolitantiditi, by HESSELBARTH et al. 1995). On the other hand, MATTONI (1977) noted that Euphilotes do differ in a number of points from "true" Philotes, and therefore there is a possibility that traditional morphology-based classification of Euphilotes as members of Scolitantiditi needs to be revised. However, the taxonomic identity of the specimen from which the GenBank sequence AF170864 is derived has not been ascertained so far (F. Sperling pers. comm.), and there remains a slight possibility that a misidentification with the phenotypically very similar Plebeius (= Icaricia) acmon (Westwood & Hewitson, [1852]) has occurred. Thus, unless the identity of this voucher specimen has been cross- checked (which is still in existence, F. Sperling pers. comm.) and in the absence of further supportive data (e.g. presence of an eversible female gonoporus, the best morphological autapomorphy of the true Polyommatiti: HÄUSER 1993), the affiliation of Euphilotes bernardino with the Polyommatiti remains uncertain.

Within Polyommatiti the COI and ITS2 gene trees correspond with each other in the exclusion of Cyaniris from a monophyletic genus Polyommatus but not in the position of Lysandra. While the COI gene tree which places Lysandra within Polyommatus confirms current morphology-based systematics, its basal placement within Polyommatiti in the ITS2 gene tree might be caused by alignment problems in the most distant groups and the high divergence of Lysandra sequences due to an apparent faster evolutionary rate, which might be caused by microsatellites. Disagreement in the COI and ITS2 data sets occur mostly in the less well resolved parts of the tree. The combined tree provides the best resolution within Agrodiaetus which proves that both data sets complement each other. The resulting clades obtained from the MP and Bayesian phylogenetic analysis are largely concordant with each other, but do not correspond very well with current groupings (HESSELBARTH et al. 1995; BÁLINT & JOHNSON 1997; ECKWEILER & HÄUSER 1997). However, these traditional, morphology-based approaches are also far from being congruent. In Tab. 12 the new subdivisions based on molecular phylogenetic data are contrasted with the three most recent conventional classification schemes.

Tab. 12. Agrodiaetus-clades on the basis of molecular data in comparison to previous morphology-based grouping concepts Agrodiaetus-clades n Hesselbarth Bálint & Eckweiler on the basis of molecular data et al. (1995) Johnson & Häuser (1997) (1997) damon-group damon ([Denis & Schiffermüller], 1775) 45 damon damon damon admetus-group admetus (Esper, [1783]) 78-80 admetus admetus admetus ripartii (Freyer, 1830) 90 admetus admetus admetus nephohiptamenos (Brown & Coutsis, 1978) ca.90 admetus admetus admetus demavendi (Pfeiffer, 1938) 66-76 admetus admetus admetus khorasanensis (Carbonell, 2001) 84 admetus lorestanus Eckweiler, 1997 69-74 admetus

88 A molecular phylogeny of Agrodiaetus

Agrodiaetus-clades n Hesselbarth Bálint & Eckweiler on the basis of molecular data et al. (1995) Johnson & Häuser (1997) (1997) dolus-group ainsae (Forster, 1961) 108-110 dolus dolus dolus fulgens (de Sagarra, 1925) 103 dolus fabressei (Oberthür, 1910) 90 admetus admetus admetus humedasae (Toso & Baletto, 1976) 38 admetus admetus admetus aroaniensis (Brown, 1976) 48 admetus admetus admetus menalcas (Freyer, [1837]) 85 dolus dolus dolus alcestis (Zerny, 1932) 19-21 admetus admetus admetus interjectus (de Lesse, 1960) 29-32 admetus admetus admetus dantchenkoi (Lukhtanov & Wiemers, 2003) 40-42 valiabadi (Rose & Schurian, 1977) 23 admetus admetus iphidamon-group iphidamon (Staudinger, 1899) 14 damon damon dizinensis (Schurian, 1982) 17 carmon carmon damon carmon-group ninae (Forster, 1956) 33-37 transcaspicus transcaspicus damon turcicola (Koçak, 1977) 19-20 transcaspicus damon huberti (Carbonell, 1993) 33-37 carmon damon zapvadi (Carbonell, 1993) 18-19 transcaspicus damon elbursicus (Forster, 1956) 16-18 transcaspicus transcaspicus damon paulae (Wiemers & De Prins, 2003) 17 arasbarani (Carbonell & Naderi, 2000) zarathustra Eckweiler, 1997 ca. 22 damon pierceae (Lukhtanov & Dantchenko, 2002) 22 carmon (Herrich-Schäffer, [1851]) 81-82 carmon carmon damon schuriani (Rose, 1978) 81-82 damon surakovi Dantchenko & Lukhtanov, 1994 50 damon sekercioglui (Lukhtanov & Dantchenko, 2002) 50 hamadanensis (de Lesse, 1959) 21-22 carmon dama dama theresiae Schurian, van Oorschot & van den 63 transcaspicus dama dama Brink, 1992 guezelmavi Olivier, Puplesiene, van der Poorten, 42 De Prins & Wiemers, 1999 dama (Staudinger, 1892) 41 poseidon dama dama karindus (Riley, 1921) 66-68 dama peilei Bethune-Baker, 1921 39 dolus dolus femininoides (Eckweiler, 1987) 27 dolus dolus morgani (Le Cerf, 1909) 25-27 dolus dolus dolus kurdistanicus (Forster, 1961) 57-62 dolus dolus dolus antidolus (Rebel, 1901) 40-41 dolus dolus dolus erschoffii-group erschoffii (Lederer, 1869) 13-15 erschoffii erschoffii achaemenes Skala, 2002 shahrami Skala, 2001 128-131 klausschuriani Ten Hagen, 1999 56 tenhageni Schurian & Eckweiler, 1999 phyllis (Christoph, 1877) 78-82 damon dolus damon glaucias (Lederer, 1871) glaucias erschoffii darius Eckweiler & Ten Hagen, 1998 caeruleus (Staudinger, 1871) 20 poseidon transcaspicus damon posthumus (Christoph, 1877) ca. 85 damon damon damon birunii Eckweiler & Ten Hagen, 1998 10-11 iphigenides-group iphigenides (Staudinger, 1886) 65-67 poseidonides iphigenides poseidonides-group dagmara (Grum-Grshimailo, 1888) erschoffii dagmara poseidonides (Staudinger, 1886) 24 poseidonides iphigenides 89 Chapter 3

Agrodiaetus-clades n Hesselbarth Bálint & Eckweiler on the basis of molecular data et al. (1995) Johnson & Häuser (1997) (1997) iphigenia-group iphigenia (Herrich-Schäffer, [1847]) 12-16 damon damon damon turcicus (Koçak, 1977) 24 carmon carmon damon baytopi (de Lesse, 1959) 27 damon damon damon rovshani Dantchenko & Lukhtanov, 1994 damon damon tankeri (de Lesse, 1960) 20-21 damon damon damon iphicarmon Eckweiler & Rose, 1993 29 damon damon damon poseidon-group poseidon (Herrich-Schäffer, [1851]) 19-22 poseidon poseidon damon putnami (Lukhtanov & Dantchenko, 2002) 25-27 poseidon hopfferi (Herrich-Schäffer, [1851]) 15 poseidon poseidon damon lycius (Carbonell, 1996) 21-22 poseidon damon ernesti Eckweiler, 1989 18 damon pseudactis (Forster, 1960) 29 damon firdussii (Forster, 1956) 24-32 actis actis damon actis (Herrich-Schäffer, [1851]) 17 actis actis damon artvinensis (Carbonell, 1997) 21-22 actis actis damon sigberti Olivier, van der Poorten, Puplesiene & 25-29 De Prins, 2000 haigi (Lukhtanov & Dantchenko, 2002) 21-25 mithridates (Staudinger, 1878) 21-27 dolus admetus dolus mofidii (de Lesse, 1963) 34-35 damon dolus damon altivagans (Forster, 1956) 18-23 actis actis damon kanduli (Lukhtanov & Dantchenko, 2002) 25 merhaba De Prins, van der Poorten, Borie, 16-17 carmon carmon damon Oorschot, Riemis & Coenen, 1991 wagneri (Forster, 1956) 16-18 actis actis damon maraschi (Forster, 1956) 16 actis damon sertavulensis (Koçak, 1979) 20 actis damon pseudoxerxes (Forster, 1956) 15-16 carmon carmon damon sennanensis (de Lesse, 1959) 28-30 dolus dolus dolus gorbunovi Dantchenko & Lukhtanov, 1994 19-20 actis damon cyaneus (Staudinger, 1899) 16-20 carmon carmon damon

The ITS-2 network analysis supports the larger clades found in the COI and combined cladograms but also provides evidence that the bad resolution in some closely related species groups and the peripheral position of certain taxa like A. iphidamon and A. iphigenides in the cladograms is due to the survival of stem species. A. damon appears to be directly derived from the ancestor of all Agrodiaetus and A. iphidamon the stem species for the iphigenides-, iphigenia-, erschoffii-, poseidon-, carmon- and poseidonides-clades. The short distances between the stem species and the subnetworks provide evidence for the fast radiation of Agrodiaetus. Phylogenetic relationships within the genus Agrodiaetus are discussed in more detail in Chapter 4.

Congruence between gene trees and species trees in outgroups If relationships between closely related species are discussed the question of congruence between gene trees and species trees arises. If both are identical, species (if defined as reproductively isolated units) should form monophyletic groups in the cladograms. Apparently some exceptions can be found, one of the most striking examples being Polyommatus icarus (Rottemburg, 1775). While populations from Spain to Iran appear as a monophyletic group (including the Greek Polyommatus andronicus Coutsis & Ghavalas, 1995 which has been separated only recently from P. icarus based on disputable evidence), the Moroccan specimen of P. icarus is placed outside this clade. This result came as a surprise

90 A molecular phylogeny of Agrodiaetus because Northwest African populations of P. icarus are thought to represent the same subspecies as in Europe (TENNENT 1996). The COI and ITS-2 p-distances between the Moroccan and Eurasian populations of P. icarus differ to a much higher degree than in other species (Tab. 13), including those with well differentiated subspecies in Northwest Africa (like Polyommatus amandus), and are on the level of well differentiated species. The Moroccan specimen also differs in phenotype from the other icarus (f. celina Austaut), but P. icarus is an extremely variable species throughout its vast, trans-Palaearctic distributional range. Without further material it cannot be decided if Northwest African populations of P. icarus are so divergent from Eurasian ones that they should better be seen as representing a distinct Polyommatus species.

Tab. 13. P-distances between Moroccan and Eurasian populations Species Country comparison p-distance COI p-distance ITS-2 Aricia montensis Morocco-Spain 0.006 0.005 Celastrina argiolus Morocco-Turkey 0.013 Cyaniris semiargus Morocco-Iran 0.012 0.011 Iphiclides feisthamelii Morocco-Spain 0.021 Iphiclides feisthamelii - podalirius Spain - Greece 0.003 Lampides boeticus Morocco-Spain 0.003 0.001 Lampides boeticus Morocco-Turkey 0.002 Lycaena alciphron Morocco-Iran 0.027 Polyommatus amandus Morocco-Turkey 0.037 0.007 Polyommatus icarus Morocco-Spain 0.068 0.025 Polyommatus icarus Morocco-Greece 0.063 0.017 Polyommatus icarus Morocco-Turkey 0.059 Polyommatus icarus Morocco-Iran 0.060 0.015 Polyommatus icarus Spain-Iran 0.012 0.017 Polyommatus andronicus - icarus Greece 0.007 0.000

Another surprise involving a Moroccan taxon was found in the species pair Iphiclides podalirius – feisthamelii (Papilionidae). The latter replaces the former in Northwest Africa and the Iberian Peninsular. Opinions differ whether these two taxa represent different species or just subspecies and no detailed studies are known from the contact zone in the French Pyrenees. The COI haplotype of I. feisthamelii (Duponchel, 1832) from Spain is identical to the I. podalirius (Linnaeus, 1758) sequence AF170873 in GenBank and very similar to Greek I. podalirius, but Moroccan and Spanish I. feisthamelii sequences are very different from each other (Tab. 13, Fig. 73). Although it cannot be ruled out that the GenBank sequence represents I. feisthamelii, because the specimen is from “France” where both taxa occur and the voucher specimen is unavailable (Sperling, pers. comm.), the COI data indicate a high level of differentiation between Northwest African and European populations but not between the two taxa feisthamelii and podalirius.

Further cases where the gene trees do not seem to correspond with taxonomical species catagories are found outside Agrodiaetus in the following species groups: Aricia agestis- artaxerxes, Polyommatus eros-eroides-menelaos, Meleageria daphnis-marcida, Polyommatus aedon-myrrhinus-cornelia, and the genus Lysandra.

Aricia agestis (Denis & Schiffermüller, 1775) and A. artaxerxes (Fabricius, 1793) are very closely related species which are known to interbreed (AAGARD et al. 2002; HOEGH- GULDBERG 1979, 1982). The two Greek specimens of Aricia artaxerxes had a very different

91 Chapter 3

COI haplotype, one of them (JC00057) almost identical to Aricia agestis. This might be due to mtDNA introgression because these specimens were found sympatrical with A. agestis on Mt. Taiyetos.

The Polyommatus eros-eroides complex consists of mostly allopatric populations whose relationships are unclear. P. menelaos Brown, 1996 from Peleponnesos is thought to be closely related to P. eros Ochsenheimer, 1808 by most authors and not to P. eroides Frivaldszky, 1835 which is found in mainland Greece, but the status of P. eroides and P. eros as distinct species is only founded on very slight differences in coloration and in habitat, with P. eros inhabiting mainly higher altitudes above 1800 m. Although the total ranges of both taxa overlap they are not known to occur in sympatry. P. eroides forsteri Pfeiffer, 1938 from Elburs Mts. in Iran is treated as a distinct species by BÁLINT (1993) and CARBONELL (1994b). The taxon yildizae Kocak, 1977 from Kop Dağı Mts. in Turkey was described as a subspecies of P. eroides and treated as such by CARBONELL (1994b) but transferred to P. forsteri by BÁLINT (1993) and to P. eros by HESSELBARTH et al. (1995) even though they noticed that the distribution of this taxon between two subspecies of P. eroides is hard to explain on zoogeographic grounds. Different species boundaries are again suggested by TUZOV et al. (2000a) who combine high altitude populations from Azerbaijan, Iran and Turkey (ssp. moletti Carbonell, 1994) under the name Polyommatus erotulus Nekrutenko, 1985, whereas GORBUNOV (2001) recognizes even five units in this species complex. Available genetic data only support a division between Iranian forsteri on one side and Greek/Turkish eroides/menelaos/yildizae on the other side which agrees best with CARBONELL (1994b). Nominotypical P. eros (from the Alps) have not been investigated though.

Meleageria marcida (Lederer, 1872) is a discoloured variety of Meleageria daphnis (Denis & Schiffermüller, 1775) confined to the Northern slopes of Elburs Mts. in Iran which was raised to species rank by BÁLINT & JOHNSON (1997) although hybrids with intermediate phenotype are known (SCHURIAN 1989b). The discoloration is probably an adaption to the specific climatic conditions (low solar radiation) on the north side of Elburs Mts. (BIRO et al. 2003). The COI sequence of M. marcida from Veresk is very different from the other daphnis populations, but another specimen from Kendevan Pass has a haplotype similar to daphnis. This suggests that M. marcida is well differentiated genetically, but gene flow exists in contact zones (one of which is near Kendevan Pass).

Polyommatus myrrhinus (Staudinger, 1901) from Eastern Anatolia was described as a subspecies of Polyommatus myrrha (Herrich-Schäffer, [1851]) which is found in other parts of Anatolia. HESSELBARTH et al. (1995) ranked it as a subspecies of Polyommatus aedon (Christoph, 1887) from Elburs Mts. (Iran) whereas BÁLINT & JOHNSON (1997) treated it as a distinct species. The large genetic distance in the COI data supports the specific distinctness of myrrhinus and aedon but does not exclude the possibility that P. myrrhinus is conspecific with true P. myrrha which was not analyzed. The COI and ITS2 haplotypes of Polyommatus cornelia (Gerhard, [1850]) are extremely similar to P. myrrhinus. Although both taxa are sometimes united in the subgenus Sublysandra (e.g. BÁLINT & JOHNSON 1997) they are quite different in wing pattern and definitely constitute different species which occur sympatrically in Anatolia without hybridization. The molecular data set implies that these taxa are very young and therefore not well differentiated genetically. It does not agree well with the conclusions of FIEDLER et al. (1994) drawn from the study of life history parameters which suggest a closer relationship of P. cornelia (= P. candalus (Herrich-Schäffer, [1851]) to P. icarus than to the P. myrrha-complex. It should be noted that P. cornelia is a very variable butterfly (HESSELBARTH et al. 1995) and karyological data (DE LESSE 1960a) indicate that it might comprise different species.

92 A molecular phylogeny of Agrodiaetus

The subgenus Lysandra is a group of closely related taxa some of which are known to interbreed with each other (SCHURIAN 1989b & 1989c). Many taxa and allopatric populations differ in chromosome numbers (DE LESSE 1960a, 1969) which range between n=24 in Lysandra syriaca (Tutt, [1910]) and n=92 in populations of Lysandra coridon (Poda, 1761) from the Balkan Peninsula. Because of its plesiomorphic chromosome number, L. syriaca (from Lebanon and South Turkey) is thought (SCHURIAN 1989b) to represent the most basal Lysandra (together with L. punctifera (Oberthür, 1876) from Morocco). The COI data confirm this hypothesis. L. bellargus (Rottemburg, 1775) (n=45) which has an extensive distribution from Spain to Iran is found sympatrically with most other Lysandra taxa. Hybrids with L. coridon have sometimes been found in nature (polonus Zeller, 1847), but F1 hybrids are probably sterile and extensive hybridization experiments with L. coridon, L. hispana and L. ossmar have failed (SCHURIAN, 1989b). The COI and ITS2 data confirm the monophyly of L. bellargus populations from Spain, Italy, Turkey and Iran without indications of gene flow with other Lysandra species. L. ossmar (Gerhard, 1851) and L. corydonius (Herrich-Schäffer, 1852) are thought to be sister species differing in the upperside wing colour (grey vs. blue) which are parapatrically distributed in West and Central Anatolia (L. ossmar) and Eastern Anatolia to Caucasus and Transcaucasus (L. corydonius). Their chromosome numbers are identical (n=84), but L. ossmar has two large chromosomes compared to three in L. corydonius (DE LESSE 1969). Natural hybrids are known from the contact zone in Sivas, Erzincan and Erzurum Province. SCHURIAN (1989b) managed to breed such hybrids until the F3-generation. The ITS2 gene tree conforms to the supposed species tree, but the sequences are very similar and only two positions are parsimony-informative. In the COI gene two different groups of haplotypes can be identified which differ in five positions, but the specimens from Sivas and Erzurum (near/in the contact zone) possess the haplotypes of the opposite species which causes the difference of the COI gene tree from the species tree. Although more material would be needed to clarify this situation, the data indicate extensive mtDNA introgression across the contact zone. (The two specimens from Erzurum have the same haplotype apart from two positions, one of which is identical to L. coridon from Italy in MW99042. This specimen also has missing character information at 16% of all COI nucleotides which causes the odd placement with L. coridon in the Bayesian analysis. The MP analysis seems to be less sensitive to missing character information and correctly places the two corydonius from Erzurum together.) The remaining Lysandra taxa belong to the Lysandra coridon species complex which has been the main focus of LELIEVRE’s (1992) study of allozymes in this group. Lysandra coridon (Poda, 1761) is the most widespread species which is found from Northern Spain (ssp. asturiensis De Sagarra, 1924) through Central Europe (type locality: Graz in Austria) to Greece. Allozyme variation indicates a division between a western and eastern group of populations (SCHMITT & SEITZ 2001). The chromosome numbers increase gradually from n=87 in Southwest Europe (Spain and Italy) to n=92 in Southeast Europe (Macedonia) (DE LESSE 1969). The allopatric taxa caelestissimus Verity, 1921 from Montes Universales (Central Spain) and gennargenti Leigheb, 1987 from Sardinia are thought to be subspecies of L. coridon by most authors but treated as distinct species by KUDRNA (2002). LEIGHEB (1987) suggested a close relationship between gennargenti and caelestissimus due to similarities in wing coloration. Allozyme variation however indicates that gennargenti populations originated from the Italian mainland and differentiation is due to gene drift and inbreeding (MARCHI et al. 1995). Lysandra albicans (Gerhard, 1851) with a chromosome number of n=82 replaces L. coridon in Central and Southern Spain. Hybrids with L. caelestissimus have often been found and even named (caerulescens Tutt) and supposed hybrids with L. coridon asturiensis are known from the contact zone (e.g. near Peñahorada/Burgos; LELIEVRE, 1992). Specimen MW01018 is from the same locality and looks like the specimen figured as possible hybrid in LELIEVRE (1992:

93 Chapter 3 photo 2, fig. 16). The picture is further complicated by the fact that another species, Lysandra hispana (Herrich-Schäffer, 1852) with n=84 chromosomes, occurs in coastal regions from Catalonia to Tuscany. The 2nd generation of this bivolitine species can occur together with the only generation of L. albicans and L. coridon and can not be reliably separated at these locations (SCHURIAN, 1989b). L. coridon and L. hispana have been crossed very successfully until the F3 generation (BEURET 1956-1959). It is possible that the specimens from Sta. Coloma de Queralt (Tarragona) represent this taxon. The ITS2- and COI-sequences from the taxa analyzed in the Lysandra coridon-complex form one monophyletic clade which confirms that they are not well differentiated genetically. Populations of L. albicans are placed at basal positions in the tree and in the network which might indicate that L. albicans is the ancestral taxon, but the material is not sufficient for far-reaching conclusions.

Comparison with allozyme results The only study of allozymes in Agrodiaetus was published by MENSI et al. (1994) who investigated some monomorphic Agrodiaetus species together with Agrodiaetus damon ([Denis & Schiffermüller], 1775). Unfortunately the chromosome number of the specimens used was not determined and therefore the identification of several taxa is doubtful, especially those from Anatolia (A. ripartii, A. demavendi and A. interjectus). A. interjectus is not known from Van Province, but four other similar karyospecies occur there: A. dantchenkoi, A. alcestis, A. demavendi and A. ripartii. Despite these flaws, some interesting comparisons with the DNA results are possible. Allozyme and DNA results correspond in the following points: • A. damon forms a distinct clade from the other investigated Agrodiaetus species. • The monomorphic Agrodiaetus species (A. fabressei, A. humedasae, A. admetus, A. ripartii) are closely related and form one clade together with the dimorphic taxa A. menalcas, A. dolus and A. fulgens. • A. admetus and A. ripartii are very closely related (possibly sister species). The only differences are in the position of A. fabressei and A. humedasae which cluster within the dolus-group in the DNA analysis but occupy an ancestral position in the allozyme studies. Of interest is also the position of two taxa which were not included in the DNA study: • A. dolus (Hübner, 1823) from South France appears closely related to A. fulgens and A. menalcas, with A. dolus vittatus (Oberthür, 1892) from Aveyron more closely related to A. fulgens than to A. dolus dolus (Bouches du Rhône). This result would question the status of A. fulgens as a distinct species. • A. exuberans (Verity, 1926), a very local endemic from Oulx (Torino, Italy) appears to be an ancestral taxon, closely related to A. fabressei and A. humedasae.

Hybridization in Agrodiaetus Despite the presumed close relationships between many Agrodiaetus species, not much is known about hybridization in this subgenus of Polyommatus. In contrast to the related subgenus Lysandra, where extensive hybridization experiments have been conducted (SCHURIAN 1989b), the only available evidence for hybrids in Agrodiaetus so far stems from a few collected specimens which are thought to represent hybrids due to their intermediate phenotype. The following natural hybrids (all of them males) have been recorded: • within Agrodiaetus: o A. ripartii (Freyer, 1830) x damon ([Denis & Schiffermüller], 1775) (SCHURIAN & HOFMANN 1975) o A. ripartii x menalcas (Freyer, [1837]) (SCHURIAN & HOFMANN 1980) o A. antidolus aereus Eckweiler, 1998 x (?) cyaneus (Staudinger, 1899) (TEN HAGEN 2003)

94 A molecular phylogeny of Agrodiaetus

• of Agrodiaetus with other Polyommatus subgenera: o A. damon x Meleageria daphnis ([Denis & Schiffermüller], 1775) (REBEL 1920) o A. damon x Lysandra coridon (Poda, 1761) (REBEL 1930a) o A. damon x Polyommatus icarus (Rottemburg, 1775) (REBEL 1930b) o A. ectabanensis x Meleageria daphnis elamita (Le Cerf, 1913) (TEN HAGEN 2003) o A. turcicus (Koçak, 1977) x Polyommatus icarus (TEN HAGEN 2003) The problem with such records is that the conclusions about the parental species of those natural hybrids (which might also be just aberrations) are very uncertain. In the case of the presumed hybrid ripartii x menalcas, at least one other karyospecies similar to A. ripartii, A. alcestis (Zerny, 1932), probably occurs at the site at Zelve (Nevşehir Prov., Turkey). Obviously, hybrids between phenotypically similar species would usually be overlooked (the same is true for any female hybrid within Agrodiaetus) or recorded as a variation of one of the parental species, and only hybrids between very different phenotypes are likely to be discovered.

During the expeditions to Turkey, Iran, Italy and Spain, only one specimen was found which appeared to be a hybrid because of its intermediate phenotype. This male specimen (MW99471) from Erek Dağı (Van Prov., Turkey) has brown wings with traces of silvery scales which would be expected in a hybrid between a species of the “brown” Agrodiaetus complex and one with silvery males. Possible parental species were two species with silvery males, A. menalcas (Freyer, [1837]) and A. kurdistanicus (Forster, 1961), which were both found at the site, and four karyospecies with brown males which are known to occur in Van Province, A. alcestis, A. dantchenkoi (Lukhtanov & Wiemers, 2003), A. demavendi (Pfeiffer, 1938) and A. ripartii. Mitochondrial DNA is maternally inherited and would therefore indicate the mother species of the presumed hybrid. In this case, the mtDNA COI sequence turned out to be identical to those of two specimens of A. dantchenkoi from nearby Kurubaş Geçidi. In nuclear genes there is a 50% chance that either the paternal or maternal copy is inherited. The ITS-2 sequence was identical to the sequence of A. menalcas including the ‘A’ in ITS-2 position 613 in the aligned data set which was not found in any other Polyommatiti sequence. Thus it can be concluded that specimen MW99471 is a hybrid between a female of A. dantchenkoi which has a chromosome number of n=40-42 and a male of A. menalcas which has the double chromosome number of n=85. The chromosome number of the hybrid specimen is approximately the same as in A. dantchenkoi and this would be expected in an F1 hybrid if a complete pairing between each dantchenkoi chromosome with two menalcas chromosomes is achieved by trivalent formation in meiosis whereas intermediate chromosome numbers usually occur in Fn and backcrosses. Chromosome numbers in hybrids can also be higher than those of their parents if no complete pairing is achieved. A. dantchenkoi and A. menalcas appear to be very closely related and might even be sister species and the occurrence of a hybrid between them is therefore hardly surprising.

Meiotic behaviour of chromosomes in hybrids of two closely related Lepidoptera species with a different karyotype, Antheraea roylei (n=31) and A. pernyi (n=49), was studied by NAGARAJU & JOLLY (1986) who observed the formation of 18 trivalents + 13 bivalents (n=31) in F1 hybrids and two different karyotypes with either 49 bivalents or 9 trivalents + 31 bivalents (n=40) in backcrosses.

A comparison of mtDNA and nuclear sequences did not reveal any other hybrid in Agrodiaetus. Although hybrids between very closely related species (such as those with identical nuclear or mtDNA sequences) can not safely be detected in this way, the existence

95 Chapter 3 of hybrids between distantly related species, especially those from different species groups within the studied material can firmly be excluded.

Radiation of Agrodiaetus and the colonization of Europe Genetic data confirm current opinions (e.g. HESSELBARTH et al. 1995) that Agrodiaetus is a very young radiation originating in the Pliocene, with most intense speciation during the Pleistocene. The biogeographical origin of the subgenus Agrodiaetus remains unsolved, because the sister group within Polyommatus could not be exactly determined and A. damon, a close relative of the ancestor species of Agrodiaetus, has an extremely extensive distribution from the Pyrenees to Mongolia, where it is confined to isolated mountain ranges like most other Agrodiaetus species. The lower altitudinal limits of most Agrodiaetus increase from the West (Spain: 500 m) to the East (Iran: 1500 m), but are usually around 1000 m. Only few species are found in the lowlands or even near sea level (e.g. A. admetus) and some are even confined to very high altitudes (e.g. A. faramarzii Skala, 2001 at 4000 m in the Zagros range of Iran). It can be assumed that A. damon had an almost continuous distribution at the end of the Pleistocene (Dryas 10000 years ago) but populations retreated into the mountains when the climate warmed up and became isolated from each other. Obviously the centre of Agrodiaetus radiation is Eastern Anatolia, Transcaucasia and Iran. Not only do these regions have the highest species diversity, but species of these regions are present in almost all species groups (with the only exception of some Central Asian ones) and their stem species are also found in these areas, most notably A. iphidamon from Elburs Mts. in northern Iran. The orogeny enabled them to evade even severe climate changes by moving up and down the mountains without the need of long range dispersal. Many species are now confined to isolated mountain tops with subalpine or alpine Onobrychis steppe and have a discontinuous distribution, but during at least ten arctic periods of the Pleistocene these populations were able to extend their distribution and to intermix, but were separated again during the interstadials, thus producing complicated speciation patterns.

Europe was only reached by few Agrodiaetus species which belong to the dolus-, admetus- and iphigenia-clade. European endemic (karyo-)species are only found in the dolus-clade and therefore this clade probably represents the first European radiation. The ancestor species is a close relative of the three Anatolian species A. dantchenkoi, A. interjectus and A. alcestis. Relicts of this first colonization are the monomorphic species A. fabressei (Central Spain), A. humedasae (Aosta, Italy), and A. aroaniensis (Greece) which appear to have survived the last glaciation in close proximity to their current occurrence. The Southwest Mediterranean A. dolus, A. ainsae and A. fulgens appear very close genetically to A. fabressei (nuclear and mtDNA) but also to the phenotypically more similar Anatolian A. menalcas (allozymes, v. MENSI et al. 1994), thus their origin remains unclear. The admetus-clade represents a distinct colonization including A. ripartii which is widely distributed in the Mediterranean region (Northern Spain, Southern France, Balkans) through Anatolia and Russia to Altai and the closely related Anatolian A. admetus which adapted especially well to low altitudes and occurs throughout the Balkan peninsula. The only species with blue-coloured males (apart from A. damon) that got a foothold in Europe is the Anatolian A. iphigenia which marginally extended its distribution to Southern Greece during the Pleistocene and is now restricted in Europe to Mt. Chelmos in the Peleponnesos.

96 Systematics of Agrodiaetus

Chapter 4: Systematics of Agrodiaetus based on molecular evidence – a new perspective

The following systematic list attempts to summarize the molecular results to infer the relationships between species of the subgenus Agrodiaetus taking into account the available evidence from karyological, morphological and biogeographic studies. It includes only those taxa which were included in the molecular study.

damon-group This group contains only one species with a most extensive distribution from Spain to Mongolia. It represents the sister of all other Agrodiaetus according to the nuclear DNA data set. damon ([Denis & Schiffermüller], 1775) Despite its vast distribution the genetic variation appears small. Samples from the French Alps and from NE Turkey are genetically very similar. The sister taxon of A. damon remains unclear. iphidamon-group This ancestral group consists of two closely related species confined to the Iranian Elburs Mountains. iphidamon (Staudinger, 1899) This species represents a genotype that is close to the presumed ancestor of all species groups with blue males (iphigenides-, iphigenia-, erschoffii-, carmon-, actis- and poseidonides- group). The nuclear ITS-2 sequences are identical with possible stem species in the iphigenia- group (A. baytopi) and erschoffii-group (A. shahrami & A. achaemenes). dizinensis (Schurian, 1982) This local endemic which is only known from the type locality Dizin (Central Elburs Mts.) appears to be most closely related to A. iphidamon. ECKWEILER & HÄUSER (1997) suggest a close relationship to A. kendevani (Forster, 1956), a taxon which could not be included in this study. iphigenides-group A Central Asian species group of which only one species was sampled. iphigenides (Staudinger, 1886) poseidonides-group This Central Asian group appears very distant from the other groups. Study of more Central Asian taxa is necessary to decide if e.g. it can be united with the iphigenides-group. poseidonides (Staudinger, 1886) In the ITS-2 network analysis this taxon seems to be most closely related to A. iphigenides. dagmara (Grum-Grshimailo, 1888) Genetically this taxon is closely related to A. poseidonides.

97 Chapter 4

iphigenia-group Small Anatolian species group. Possible ancestor species: A. iphidamon. The following four taxa with mostly allopatric distribution appear to be very closely related genetically. baytopi (de Lesse, 1959) The ITS-2 sequence of this East Anatolian species is identical to the suggested ancestor species iphidamon and therefore it could be the stem species of the iphigenia-group. tankeri (de Lesse, 1960) This taxon differs from the previous species in phenotype and karyotype and is distributed parapatrically in NE Turkey while A. baytopi is found in SE Turkey. A sympatric occurrence is known from Tahir Geçidi (Ağrı Prov., Turkey; HESSELBARTH et al. 1995). ITS-2 sequences of both taxa are identical and their mtDNA haplotypes overlap which raises the question if speciation is complete or if gene flow persists. rovshani Dantchenko & Lukhtanov, 1994 This taxon whose karyotype remains unknown occurs in Azarbaijan and Iranian Azarbaijan and is genetically close to A. baytopi and A. tankeri. iphicarmon Eckweiler & Rose, 1993 Genetic data confirm the karyological results that this taxon is not a subspecies of A. iphigenia but instead closely related to A. baytopi. turcicus (Koçak, 1977) This Eastern Anatolian species is also closely related to A. baytopi and A. tankeri but occurs sympatrically with them. iphigenia (Herrich-Schäffer, [1847]) This is the only representative of this group whose range extends to Europe (Peleponnesos, Greece). Populations from different parts of Anatolia and Armenia are genetically very similar and well differentiated from the other members of the iphigenia-group, which are often found sympatrically. admetus-group This group has an extensive distribution from Northern Spain to Kazakhstan and includes only monomorphic brown species with an elevated number of chromosomes (n=65-90). The ancestor species of this group seems to be a close relative of A. damon.

The following three taxa are karyospecies with an allopatric distribution, but indistinguishable on morphological grounds. ripartii (Freyer, 1830) Despite of their similarities in phenotype and karyotype populations of this widespread taxon from Spain, Greece and Turkey do not form a monophyletic group which could be taken as an indication that A. ripartii represents the stem species for the remaining taxa in this group. Alternatively it is possible that speciation is incomplete and gene flow between the karyospecies A. ripartii and A. demavendi persists. khorasanensis (Carbonell, 2001) This taxon from Kopet Dagh (Iran) with a chromosome number intermediate between the previous and the following taxon is genetically close to A. ripartii. demavendi (Pfeiffer, 1938) Genetic and karyological data indicate that this karyospecies might consist of several species with only slightly different chromosome numbers but further investigations which include the precise determination of chromosome numbers are necessary. Dugijan (Azarbaijan-e Sharqi, Iran) turned out to be one interesting location where two different mtDNA haplotypes were found which might indicate the coexistence of two different species. These haplotypes were both found from Eastern Turkey to Northwest Iran. Slight differences in phenotype have led to recent descriptions of new species like A. ahmadi (Carbonell, 2001) and A. urmiaensis

98 Systematics of Agrodiaetus

(Schurian & ten Hagen, 2003), unfortunately without karyological data, and it is uncertain whether the recorded differences can be used to delimit species. lorestanus Eckweiler, 1997 This taxon was described as a subspecies of A. demavendi and its karyotype is probably identical (Carbonell, 2001). Genetic data confirm the close relationship to A. demavendi. nephohiptamenos (Brown & Coutsis, 1978) This taxon from Macedonia might constitute a synonym of A. ripartii unless it can be confirmed that is has a different karyotype. Genetically and phenotypically it appears to be very close to A. ripartii. admetus (Esper, [1783]) This species has an extensive distribution from the Balkans to Western Siberia and occurs sympatrically with most of the former taxa. Genetically it is closely related to A. ripartii and A. demavendi. dolus-group This is a group of taxa with brown or whitish males which is distributed from Southern Spain to Elburs Mts. (Iran) and Lebanon. It corresponds to the menalcas-group which resulted from the Bayesian analysis of COI and ITS-2. The ancestor species of this group seems to be a close relative of A. damon. valiabadi (Rose & Schurian, 1977) This species appears to be the most ancestral in the dolus-group. dolus (Hübner, [1823]) Material of this taxon from Southern France with whitish males could not be included in this study, but the following two taxa are apparently very closely related. ainsae (Forster, 1961) This is the Spanish representative of A. dolus which differs only slightly in chromosome numbers. fulgens (de Sagarra, 1925) DNA sequences of this taxon are identical to A. ainsae which further questions its status as a distinct allopatric species (see Chapter 2). fabressei (Oberthür, 1910) Despite the similar phenotype and karyotype this taxon does not seem to be closely related to ripartii. Instead it appears to be very closely related to A. ainsae to which it is allopatric in distribution (MUNGUIRA et al. 1995). interjectus (de Lesse, 1960) The comparison of nuclear and mtDNA confirms DE LESSE’s opinion (1960b) that this taxon is not a hybrid species between A. alcestis and A. demavendi or A. ripartii despite of its intermediate chromosome number but instead it is genetically very close to A. alcestis. It occurs sympatrically with A. alcestis and A. demavendi. dantchenkoi (Lukhtanov & Wiemers, 2003) This karyospecies is closely related to A. interjectus to which it is allopatric in distribution. alcestis (Zerny, 1932) The two subspecies with slightly different karyotypes, the nominate one from Lebanon and Anatolia and ssp. karacetinae Lukhtanov & Dantchenko, 2002 from Kordestan have similar ITS-2 sequences but different and independently evolved COI haplotypes indicating possible specific distinctness of these two taxa. menalcas (Freyer, [1837]) Although this Anatolian taxon has whitish males like A. dolus/ainsae/fulgens it appears to be more closely related to A. alcestis. aroaniensis (Brown, 1976) This Greek taxon also appears most closely related to A. alcestis. humedasae (Toso & Baletto, 1976) This Italian endemic from Aosta valley is very closely related to A. aroaniensis.

99 Chapter 4 erschoffii-group This is a predominantly Iranian group with one species reaching Central Anatolia. Most members of this group appear closely related to their ancestor species A. iphidamon. shahrami Skala, 2001 This high mountain endemic from the Zagros range appears to be one of the most ancestral because its ITS-2 sequence is identical to A. iphidamon. achaemenes Skala, 2002 A very close relative of A. shahrami with almost identical sequences. It is an allopatric endemic of another high mountain in the Zagros range. phyllis (Christoph, 1877) This species which ranges from Elburs Mts. to Central Anatolia represents a monophyletic group of populations which are well differentiated genetically from all other members of this group. glaucias (Lederer, 1871) The closest relative of this taxon which was described from Elburs Mts. (Iran) appears to be A. birunii. tenhageni Schurian & Eckweiler, 1999 This taxon from Khorasan (Iran) appears to be closely related to the ancestor species A. iphidamon. klausschuriani ten Hagen, 1999 The origin of this species remains unclear. In the ITS-2 network analysis it connects with the actis-group. posthumus (Christoph, 1877) This taxon appears to be restricted to the eastern Elburs Mts. The genetic analysis confirms the specific distictness from A. phyllis which occurs sympatrically at the type locality and has a similar chromosome number but a different karyotype with two very large chromosomes. darius Eckweiler & ten Hagen, 1998 Genetically this taxon from Central Elburs Mts. is most closely related to the allopatric A. posthumus from eastern Elburs. birunii Eckweiler & ten Hagen, 1998 This taxon was described as a subspecies of A. posthumus from central Elburs Mts. (Iran) but according to the genetic analysis, which agrees with the karyological results, this taxon must be considered specifically distinct from A. posthumus. caeruleus (Staudinger, 1871) This is the only Agrodiaetus species in this study which has blue-coloured females. It is distributed from Transcaucasus to Transcaspia and similar taxa also occur in Central Asia. In Elburs Mts. it occurs sympatrically with several taxa in this group (e.g. A. phyllis, A. posthumus, A. erschoffii and A. glaucias) which appear as the closest relatives in this study. erschoffii (Lederer, 1869) This species from Northeast Iran appears to be genetically most distinct from the other species in this group and its closest relatives remain unclear. carmon-group The ancestor species of this heterogenous group which is distributed from Anatolia to Iran appears to be A. iphidamon. carmon (Herrich-Schäffer, [1851]) This Anatolian taxon is the first in a group of four closely related and apparently allopatric taxa, some of which have a different karyotype. Specimens from very distant locations (Antalya Prov. in SW Turkey and Kars Prov. in NE Turkey) appear to be very similar genetically.

100 Systematics of Agrodiaetus schuriani (Rose, 1978) Although the sampled specimen of this taxon from Cappadokia is from a location which is between the two sampled localities of carmon, the sequences are quite distinct. This result indicates that the taxon schuriani is not a synonym of carmon, but further investigations are necessary to verify its status and distribution. A close relation to A. surakovi as indicated by ROSE (2002) seems possible on the basis of the molecular results. surakovi Dantchenko & Lukhtanov, 1994 This taxon represents A. carmon in Armenia and Azarbaijan but has a different karyotype. Genetically it is very similar to A. carmon. sekercioglui (Lukhtanov & Dantchenko, 2002) The COI data confirm that this taxon from SE Turkey is closely related to A. surakovi and was correctly placed as a subspecies of the latter. pierceae (Lukhtanov & Dantchenko, 2002) This taxon is very similar in phenotype to A. huberti (Carbonell, 1993) but has a different karyotype and seems to replace it in Southeast Turkey (Van and Hakkari Prov.). According to LUKHTANOV & DANTCHENKO (2002b) it is genetically (mtDNA) close to A. kendevani (Forster, 1956) and A. zarathustra neglecta (Dantchenko, 2000)2 and not to A. huberti despite the very similar appearance. The author can confirm the genetic distance to A. huberti. According to the COI data set A. pierceae is very close to A. schuriani but according to the ITS2 data set its relationships within the carmon-group are less clear because it appears to be quite distinct to any other members of this group. A. pierceae occurs sympatrically with A. surakovi sekercioglui at Çatak. The following 8 taxa represent a group of closely related taxa, some of which are allopatric to each other and may differ only karyologically. Their relationships are not well understood. Most of its members were included in the transcaspicus-group sensu HESSELBARTH et al. (1995). ninae (Forster, 1956) The first member of this group, which was described as a subspecies of A. transcaspicus but differs from it in karyotype, is distributed from Armenia to Eastern Anatolia turcicola (Koçak, 1977) This taxon which appears to replace A. ninae in Van Province (Turkey) can only be distinguished from the latter by its karyotype and the genetic data confirm that this taxon is very closely related to it. huberti (Carbonell, 1993) Prior to CARBONELL (1993) this taxon which is widely distributed in Northeastern Anatolia and occurs sympatrically with A. ninae was confused with the latter due to its similar appearance and identical karyotype. The nuclear and mitochondrial DNA data sets differ in the position of this taxon. A. huberti shares very similar COI haplotypes with A. ninae, but the ITS-2 sequences are distinct, confirming its specific status. elbursicus (Forster, 1956) Nominotypical material of this taxon from Elburs Mts. (Iran) appears to be very close to A. ninae according to the nuclear DNA data set, but quite distinct from it according to the mtDNA data set. The number of chromosomes is half the number found in A. ninae. zapvadi (Carbonell, 1993) The phenotype and karyotype of this taxon from Van Province (Turkey) is very similar to elbursicus, but genetically it is distinct. According to the mtDNA data set it is most closely related to A. huberti. zarathustra Eckweiler, 1997 This Iranian taxon from Lorestan has a peculiar phenotype and its karyotype is also different from the other members of the ninae-species group. Genetically it is very close to A. elbursicus (mtDNA) and A. zapvadi (ITS-2).

2 This taxon was described from Armenia but the name A. zarathustra neglecta Dantchenko, 2002 appears to be a nomen nudum because it was published without description and without stating any differentiating characters. 101 Chapter 4 arasbarani (Carbonell & Naderi, 2000) This taxon from Azarbaijan-e Sharqi (Iran) is similar to A. zarathustra in phenotype. Genetically it is closely related to A. elbursicus. paulae Wiemers & De Prins, 2003 The phenotype of this new species from Ahar Pass (Iran) is very distinct from any other member of this group and it has previously been confused with A. altivagans (ECKWEILER, pers. comm.). According to the genetic data it is most closely related to A. zarathustra, A. arasbarani, A. elbursicus and A. zapvadi and its karyotype is like those of the latter two taxa. dama (Staudinger, 1892) This very rare Anatolian species is genetically very distant from the other taxa in this group. It appears basal in the ITS-2 network and therefore might represent an ancestral taxon. hamadanensis (de Lesse, 1959) This Iranian species with a peculiar phenotype (males with violett wing upperside) is genetically very distant from other Agrodiaetus but seems to be most closely related to A. dama according to the mtDNA data set. theresiae Schurian, van Oorschot & van den Brink, 1992 The relationships of this local Anatolian endemic remain unclear although the mtDNA data set indicates A. surakovi as closest relative. Possibly this is another ancestral relict species comparable to A. dama. guezelmavi Olivier, Puplesiene, van der Poorten, De Prins & Wiemers, 1999 The genetic data confirm that this local Anatolian endemic is closely related to its sister species A. theresiae from which it differs only by its karyotype. antidolus (Rebel, 1901) This Eastern Anatolian species is the first in a group of 6 genetically closely related taxa. The first four taxa have males with a silvery-brown upperside and are allopatric in distribution. kurdistanicus (Forster, 1961) This taxon which is very similar to antidolus in phenotype but differs in chromosome number is only known from Van Province (Turkey). Genetically these karyotypically different specimens from Van Province do not differ from specimens with the antidolus-karyotype found in Hakkari Province. femininoides (Eckweiler, 1987) The darkest member of this group from Zanjan (NW Iran) again has a different chromosome number but genetically it is close to the former two taxa from Turkey. morgani (Le Cerf, 1909) The phenotype of this taxon from Iranian Kordestan and Lorestan is similar to A. antidolus but its chromosome number is the same as in A. femininoides. Its mtDNA is also similar to the latter. peilei Bethune-Baker, 1921 This rare species whose males have a unique golden brown wing colour occurs sympatrically with the former and the following species. It differs from them in karyotype and has a chromosome number intermediate between these two species. Genetically it is closely related to both of them and its COI haplotype is even identical with A. morgani. karindus (Riley, 1921) This Iranian taxon which occurs sympatrically with the former two species and differs from them in phenotype and karyotype appears to be misplaced in this group at first glance. Unlike the other members of the antidolus-species group its males display a bright blue upperside wing colour, similar to A. dama and therefore it was described as a subspecies of the latter. However the analysis of its mtDNA reveals that its haplotype is almost identical to A. peilei. Striking similarities to this species can also be found in the form of the wings and in the underside wing pattern.

102 Systematics of Agrodiaetus poseidon-group This group comprises mainly Anatolian, Caucasian, and few Iranian elements. Its ancestor species is probably A. iphidamon. hopfferi (Herrich-Schäffer, [1851]) This Anatolian species with conspicuous male phenotype (greyish blue upperside) appears to be the stem species of this group. poseidon (Herrich-Schäffer, [1851]) The genetic data confirm that this Anatolian species is closely related to A. hopfferi. It has a similar phenotype but blue-coloured males and a different karyotype. Its distribution closely resembles that of A. hopfferi but it is absent from Southeast Turkey. In Northeast Turkey it seems to be replaced by the following karyospecies: putnami (Lukhtanov & Dantchenko, 2002) Genetically this taxon from NE Turkey which was separated from A. poseidon because of its higher number of chromosomes seems to be very close to A. poseidon and its COI haplotype is almost identical. In the ITS-2 network however it clusters with the erschoffii-group, but this is due to only two plesiomorphic nucleotide character states shared with A. birunii but which are also found in A. iphidamon. Possibly this taxon is ancestral to A. poseidon. lycius (Carbonell, 1996) The genetic data confirm the view of ECKWEILER & HÄUSER (1997) that this local taxon which occurs near Antalya just off the Southwestern distributional limit of A. poseidon and A. hopfferi and which shares the karyotype of the former is closely related to both species.

The following 8 taxa belong to a group of genetically very closely related taxa and most of them share very similar phenotypes. The reason why the MP- and Bayesian analysis produced unresolved trees is the fact that several populations appear as stem populations of others which can be seen in the network analysis (Fig. 82). This is usually the case in populations of the same species. Although karyological results indicate that several species are involved, there is no indication of genetic differentiation of such karyospecies. actis (Herrich-Schäffer, [1851]) This taxon from Central Anatolia is the first named of the actis-species group and apparently has a low chromosome number (n=17) although topotypical material has not been investigated yet. Genetically however it is not differentiated from the following taxa. firdussii (Forster, 1956) This taxon was described from Elburs Mts. and with n=31-34 its chromosome number seems to be about double the number in actis. Populations from Northwest Iran and Eastern Anatolia which were included in this study have a slightly lower and variable number (see chapter 2) but there is no indication of genetic differentiation between populations or specimens with lower (n=ca.25) and higher (n=28-30) number of chromosomes. pseudactis (Forster, 1960) With n=29 the chromosome number of this taxon from Azarbaijan appears to be the same as in many populations from Iranian Azarbaijan and eastern Anatolia. If populations of A. firdussii from Elburs Mts. should turn out to be specifically distinct, the name pseudactis might apply to populations from Northwest Iran and eastern Anatolia, otherwise the name might be sunk into synonymy with firdusii (as done by HESSELBARTH et al. 1995). A specimen from Armenia which was included in this study appears to represent the stem populations for all the others. This would indicate that the actis-species group originated in Armenia. ernesti Eckweiler, 1989 This local taxon which was described as a subspecies of A. firdussii from mountains near Antalya at the southwestern range limit of this species group is genetically more differentiated than any other taxon in this group. It also differs in phenotype (e.g. lighter blue upperside of the males). The chromosome number (n=18) is very similar to A. actis. HESSELBARTH et al. (1995) synonymized this taxon under A. sertavulensis, although both taxa have a different phenotype. This action is not supported by genetic and karyological data.

103 Chapter 4 artvinensis (Carbonell, 1997) Some populations from Northeast Turkey differ slightly in phenotype and chromosome number (n=21-22). Genetically however, these populations appear almost identical to those from Armenia (pseudactis). sigberti Olivier, van der Poorten, Puplesiene & De Prins, 2000 Some populations from the Taurus Mts. and the Pontic chain with dark phenotype are covered under this name. However, the phenotype of this taxon is very variable and many specimens cannot be separated from typical actis/firdussii/pseudactis. The karyotype is the same as in pseudactis. Genetically these populations do not seem to be differentiated either. Two specimens from the same population appear at slightly different places in the network. Although these specimens exhibit the extremes in phenotype variation found in this taxon, the genetic analysis also does not indicate that two different species are involved. haigi (Lukhtanov & Dantchenko, 2002) Populations from Southeast Turkey appear to have a slightly lower chromosome number (n=21-25) than surrounding populations from Northwest Iran and Northeast Turkey and such specimens were also included in the genetic analysis but they do not appear to be well differentiated genetically. The same probably applies to A. bilgini (Dantchenko & Lukhtanov, 2002) which was described from Torul (Prov. Gümüşhane) and has the same chromosome number as A. haigi.

The remaining taxa form the crown group in Agrodiaetus and appear very closely related genetically. Due to the very similar phenotypes and karyotypes their systematics remains poorly understood. It appears that they are very young evolutionary units in the process of speciation. altivagans (Forster, 1956) The genetic analysis reveals that the material referred to this taxon in this study is heterogenous and might constitute several species, despite similar chromosome numbers (n=21-23). Alternatively it is one genetic unit with a variable phenotype and comprising several different haplotypes. In this case the following four taxa might represent karyologically diverged populations at the periphery of its range which are in the process of speciation and therefore not well differentiated genetically. The specimen from Armenia, where the type locality of A. altivagans is located, seems to occupy an ancestral position in the COI network analysis (Fig. 82) and it appears to be genetically distinct from the Turkish specimens: MW99165 from Erzincan (Erzurum Prov.) and MW99240 from Güzeldere Geçidi (Van Prov.) appear closely related to A. wagneri and A. maraschi whereas MW99353 and MW99357 from Güzeldere Geçidi (Van Prov.) are genetically similar to A. gorbunovi. The phenotype of the latter two specimens differs from typical altivagans and appears more similar to A. wagneri sensu HESSELBARTH et al. (1995) but this taxon has a different karyotype. wagneri (Forster, 1956) This taxon was treated as a subspecies of A. altivagans by DE LESSE (1962a) but elevated to species rank by HESSELBARTH et al. (1995) who claim that both taxa occur sympatrically in eastern Turkey. Their delimitation of this taxon however remains far from clear and the only karyological data available are from DE LESSE (1962a) who checked one specimen from the type locality which had n=16 chromosomes. The specimen included in this study is from Nevşehir which is 300 km east of the type locality in the centre of distribution of wagneri and 400 km west of the closest population of A. altivagans in Erzincan (according to HESSELBARTH et al. 1995). The phenotype of this specimen is typical for A. wagneri and several similar specimens from the same population are figured in HESSELBARTH et al. (1995: Plate 122, fig. 14-17). Its chromosome number is n=18 which is the same as recorded by De Lesse (1962a) from A. altivagans of Erzincan Prov. Genetically this specimen is also very similar to A. altivagans from Erzincan (MW99165). Two further specimens which are similar to A. wagneri sensu HESSELBARTH et al. (1995) from Van Province (MW99353 and MW99357) turned out to have the same chromosome number as A. altivagans (n=21-23) and are therefore placed under this taxon. Genetically these two specimens do not appear closely related to A. wagneri.

104 Systematics of Agrodiaetus maraschi (Forster, 1956) HESSELBARTH et al. (1995) synonymized this taxon from Maraş under A. wagneri, partly because of its identical chromosome number (n=16) which DE LESSE (1962a) reported from Kayseri, 170 km west of its type locality. The genetic analysis includes a specimen from Gürün (Sivas Prov.), 130 km north of the type locality whose phenotype appears to be very similar to the holotype of A. maraschi. Its chromosome number also turned out to be n=16. Genetically it is almost identical to A. altivagans from Erzincan and Van Prov. (specimen MW99240, n=21). sertavulensis (Koçak, 1979) HESSELBARTH et al. (1995) raised this taxon from the Taurus Mts. in Turkey to species level and synonymized A. ernesti with it, one reason for this action being the alleged sympatry of A. sertavulensis with A. wagneri at the type locality. Genetically however this taxon appears to be almost identical to A. wagneri and A. maraschi but distinct from A. ernesti. A. sertavulensis (n=20) seems to represent an allopatric population of the altivagans-complex and the “sympatric” wagneri might be just an intrapopulational variation. gorbunovi Dantchenko & Lukhtanov, 1994 This is another allopatric taxon of the altivagans-complex from Azarbaijan. Genetically specimens from three different populations in Northwest Iran are almost identical with each other and with two A. altivagans from Van Province (Turkey). damocles (Herrich-Schäffer, [1844]) This Russian taxon with a chromosome number of n=24-27 has only recently been recorded from Turkey (Erzincan Prov.) and described as ssp. kanduli (LUKHTANOV & DANTCHENKO, 2002). The holotype is phenotypically very similar to A. wagneri sensu HESSELBARTH et al. (1995). According to the mtDNA data set of LUKHTANOV & DANTCHENKO (2002) this taxon is closely related to A. damocles damocles, A. damocles rossicus, A. damocles krymaeus and A. altivagans. In this study a specimen from Çatak (Van Province) is included which resembles A. damocles kanduli in phenotype and which has the same chromosome number (n=25). Genetically it turned out to be very similar to A. gorbunovi and to A. altivagans from Güzeldere Geçidi (Van Province) with n=22. It seems that karyological results do not coincide well with phenotypic or genetic variation in this complex and further karyological investigations are necessary to clarify species boundaries. At present, the genetic data only indicate the existence of one species (A. damocles) with a variable phenotype and karyotype which comprises the taxa damocles, altivagans, wagneri, maraschi, sertavulensis & gorbunovi. mofidii (de Lesse, 1963) This Northeast Iranian taxon from Kopet Dagh (ssp. mofidii, n=34-35) and Kuh-e-Sorkh (ssp. sorkhensis Eckweiler, 2003, n=45) is genetically very close to A. altivagans to which it is allopatric in distribution. mithridates (Staudinger, 1878) The systematic position of this monomorph Anatolian species with a variable chromosome number of n=21-27 has been uncertain. It was placed into the admetus-group (with monomorph males) by BÁLINT & JOHNSON (1997), but into the dolus-group (with silvery males) by HESSELBARTH et al. (1995) and ECKWEILER & HÄUSER (1997). Genetically it is closely related to A. sennanensis and A. hopfferi according to the nuclear DNA data set, but to A. altivagans and A. maraschi according to the mtDNA data set. The range of A. mithridates appears to be the intersection of the range of A. hopfferi with the range of the A. altivagans- complex. sennanensis (de Lesse, 1959) This northwest Iranian taxon appears to be closely related to the allopatric A. hopfferi (ITS-2) and to the sympatric A. cyaneus (COI). pseudoxerxes (Forster, 1956) The closest relative of this Iranian taxon from Elburs Mts. seems to be A. gorbunovi and A. cyaneus. ECKWEILER & HÄUSER (1997) list it as a subspecies of A. kendevani although De Lesse (1962a) found both taxa sympatrically at Kendevan Pass and treats A. pseudoxerxes as a subspecies of A. altivagans and the close relationship with this species can now be confirmed

105 Chapter 4

genetically. According to the mtDNA data set of Lukhtanov & Dantchenko (2002) A. kendevani is genetically closely related to A. pierceae and A. zarathustra both of which belong to the carmon-group. This result also rules out conspecifity of A. pseudoxerxes with A. kendevani. cyaneus (Staudinger, 1899) This taxon was described from Georgia and is distributed from the Caucasus and Kordestan to Iran. Genetically it appears distinct but closely related to the altivagans-complex. Specimens from Van Province (Turkey) and Marand (Iranian Azarbaijan) which belong to the nominate subspecies are very similar genetically, but the specimen of ssp. damalis (Riley, 1921) from Lorestan in Iran appears to be genetically distinct. merhaba De Prins, van der Poorten, Borie, Oorschot, Riemis & Coenen, 1991 This taxon is similar to A. cyaneus in phenotype and karyotype and only known from northeastern Anatolia (Artvin and Erzurum Prov.). The genetic data confirm its close relationship with A. cyaneus to which it is allopatric in distribution and it might be more appropriate to treat it as a subspecies of A. cyaneus.

106 Evolution of morphological traits in Agrodiaetus

Chapter 5: Evolution of morphological traits in Agrodiaetus

Introduction Although differences in wing pattern and coloration between many Agrodiaetus species are very subtle to anyone who is not familiar with this group of butterflies, they often represent the only means of identification and classification. Yet even experts are often unable to determine single specimens because of considerable infraspecific variation (within and between populations). On the other hand, the subgenus Agrodiaetus includes also very different colour morphs. Even though the majority of species has males with an iridescent blue upperside, like the majority of Lycaenidae, the subgenus Agrodiaetus also includes many species with different upperside coloration, most of them brown, some silvery, whitish or even golden brown. Together with the extent of androconial patches, these colour morphs have even been the main characters to delimit species groups within Agrodiaetus. The iridescent coloration is confined to the males, whereas females of most Agrodiaetus species are dull brown. It therefore appears that the male coloration has evolved through sexual selection. Females of the closely related Polyommatus icarus only mate once and exhibit mate choice (KNÜTTEL & FIEDLER 2001), and it should be noted that butterflies discern colour very differently from the human eye, because they also have colour receptors for the ultraviolet light spectrum. Unfortunately no studies have been conducted to infer the coloration of Agrodiaetus males as it would appear to the eye of the butterfly, and not even the visible colour has been measured and analysed quantitatively. A quantitative analysis of wing pattern characters by means of multivariate techniques is also beyond the scope of this thesis (but hopefully can be conducted in the future). This chapter aims to pinpoint four important morphological characters which have been used for the systematics of Agrodiaetus and evaluate them in a qualitative approach: 1. Presence of a distinct white streak on the hindwing underside 2. Presence of orange submarginal lunules on the hindwing underside 3. Presence of well developed androconial patches on the forewing upperside of males 4. Ground colour of the male upperside The first character is the main character used to delimit the subgenus Agrodiaetus although some species or individuals with the white streak missing are still regarded as members of this subgenus due to other distinctive characters. The remaining characters have been used in combination to characterize species groups within Agrodiaetus.

Material and methods The presence of the first three characters (white streak, orange submarginal lunules and androconial patches) was checked from the wing vouchers. 1211 wing vouchers of Agrodiaetus and 521 wing vouchers of outgroup species which are kept in glass slide mounts were available for the analysis (Appendix 2). Wings were also scanned with a Microtek Scanmaker E6 at 600 dpi resolution in RGB Modus (16.7 million colours) and will be made available online through MorphBank (http://www.morphbank.net/). The ground colour of the forewing upperside was copied from a representative spot in the area of the cell using Adobe Photoshop and pasted onto the nodes of the ITS2-network for visualization.

107 Chapter 5

Results A distinct white streak was found in most Agrodiaetus but not in any outgroup species. Within Agrodiaetus the white streak was found missing in the following species or specimens:

Clade Species Specimens with missing streak admetus-clade Agrodiaetus admetus all males, females streak reduced admetus-clade Agrodiaetus lorestanus WE02535 (=1/2) dolus-clade Agrodiaetus fabressei all dolus-clade Agrodiaetus humedasae all dolus-clade Agrodiaetus aroaniensis JC00040, 041, 047 (=3/4) carmon-clade Agrodiaetus hamadanensis all carmon-clade Agrodiaetus dama all carmon-clade Agrodiaetus karindus all carmon-clade Agrodiaetus peilei all carmon-clade Agrodiaetus femininoides all carmon-clade Agrodiaetus morgani all carmon-clade Agrodiaetus antidolus MW99376-379, 393, 404 (=7/11) erschoffii-clade Agrodiaetus tenhageni all poseidon-clade Agrodiaetus sennanensis all poseidon-clade Agrodiaetus mithridates very weak (1)

Orange submarginal lunules were present in the following species:

Clade Species iphigenides-clade Agrodiaetus iphigenides poseidonides-clade Agrodiaetus poseidonides poseidonides-clade Agrodiaetus dagmara

Androconia are present in all Agrodiaetus but in some species they are organized in hairy androconial patches on the forewing upperside. The extension of these patches however is subject to considerable variation and is difficult to measure. The following list includes only species with extensive androconial patches which are not restricted to the wing venation:

Clade Species Specimens admetus-clade all dolus-clade all carmon-clade Agrodiaetus peilei all carmon-clade Agrodiaetus femininoides partial carmon-clade Agrodiaetus morgani all carmon-clade Agrodiaetus kurdistanicus all carmon-clade Agrodiaetus antidolus all poseidon-clade Agrodiaetus hopfferi partial poseidon-clade Agrodiaetus sennanensis all poseidon-clade Agrodiaetus mithridates all

The ground colour of the reference specimens in the ITS2-network (see Fig. 84) is presented in Fig. 86 and the presence of well-developed androconial patches is indicated with an ‘A’. The species names of the reference specimens can be found in Fig. 84 and Tab. 11.

108 Evolution of morphological traits in Agrodiaetus

Fig. 86. Ground colour and presence of androconia (A) mapped on ITS2-MP-network 109 Chapter 5

Discussion HÄUSER & ECKWEILER (1997) state the presence of a distinct white streak on the hindwing underside which is „sometimes entirely absent but never partly present (as in other groups of Polyommatus)” as the only morphological diagnostic feature for the delimitation of the subgenus Agrodiaetus. (The second diagnostic feature stated is the oligophagy on Onobrychis and Hedysarum.) The molecular results have confirmed the delimitation of Agrodiaetus sensu HÄUSER & ECKWEILER (1997) as a monophyletic group within Polyommatus and also the exclusion of Polyommatus thersites which has a vestigial white streak (like many Polyommatus species) and whose larva also feeds on Onobrychis. (P. thersites is included within Agrodiaetus e.g. by BÁLINT & JOHNSON (1997) and TOLMAN & LEWINGTON (1997).) It can also be confirmed that the loss of the white streak in several Agrodiaetus species occurred independently in different clades and that there is individual variation of this character in some species. According to HESSELBARTH et al. (1995) the streak is mostly absent in A. admetus but can sometimes be present and in females it is often only partly present (contrary to the statement in HÄUSER & ECKWEILER 1997). In A. mithridates the variation of this character is considerable with the white streak present, hardly visible or totally absent. A. demavendi lorestanus Eckweiler, 1997 was described because of the absence of the white streak in 90% of the individuals. HESSELBARTH et al. (1995) discuss the possibility that the white streak represents a diagnostic character to separate the two karyospecies A. antidolus and A. kurdistanicus. In the material used for this thesis all specimens of the karyospecies A. kurdistanicus (from Van Province) had a white streak. In A. antidolus (from Hakkari Province) the white streak was absent in all specimens from Yüksekova and Dilezi Geçidi but present in a part of the specimens from Haruna Geçidi and Dez valley. Specimens of A. antidolus with and without white streak were included in the molecular study which did not reveal any genetic differentiation between these two forms. In summary, there is good evidence to accept the white streak as an autapomorphic character state of Agrodiaetus, which has subsequently been lost multiple times, most notably in the antidolus subclade.

Orange submarginal lunules were only present in the three Central Asian species. (This character can be found in many further Central Asian Agrodiaetus species which could not be included in this study.) The presence of such lunules is a plesiomorphic character which is found in most other species of the genus Polyommatus (but not in Meleageria s.str. and Neolysandra). Therefore these Central Asian species might be thought to represent ancestral taxa but this does not appear to be true from the molecular analysis. However, this possibility can not be entirely ruled out, because Polyommatus species confined to Central Asia were not sampled and the sister group to Agrodiaetus might be found there.

According to ECKWEILER & HÄUSER (1997) the upperside coloration and presence of well- developed androconial patches are the main characters to delimit species groups within Agrodiaetus. Apart from the Central Asian taxa which were separated due to their biogeographical origin (iphigenides- and dagmara-group), despite of differences in upperside coloration and androconial patches, all other blue-coloured Agrodiaetus with sparsely developed androconial patches were grouped together (damon-group) and the taxa with well developed androconial patches were split into three groups, one with only brown-coloured males (admetus-group), one with only blue-coloured males (dama-group) and one with differently (including white-) coloured males (dolus-group). From Fig. 86 it appears that well-developed androconial patches were only found in non-blue-coloured males, a discrepancy to ECKWEILER & HÄUSER (1997) which needs explanation. Some taxa of their dama-group (like A. theresiae) have slightly better developed androconia than other blue Agrodiaetus but these are not comparable to those found in the admetus- and dolus-group. In A. hamadanensis the androconial patches are hardly developed at all. Individual variation can

110 Evolution of morphological traits in Agrodiaetus also explain some differences in judgement between authors. The most extreme variation was observed in A. hopfferi, a species with variable bluish-silvery coloured males which was placed in the damon-group by ECKWEILER & HÄUSER (1997). Some specimens have well- developed androconial patches whereas these are hardly visible in others and absent in the more bluish males from Hakkari Province. On the other hand, two specimens with brown males appear in Fig. 86 which do not have well-developed androconial patches. One of them is A. glaucias, a member of the erschoffii-group, which still has blue basal suffusion on the wing upperside. The same is true for specimen WE02671 of A. femininoides from Ardabil Province whereas A. femininoides specimens from Zanjan Province, which have only slight traces of basal suffusion, all have well-developed androconial patches. To conclude, it appears that androconial patches are especially well developed in species with non-blue-coloured males. This makes sense because a well-developed scent-based mate recognition system is needed in monomorph Agrodiaetus to replace the visual mate recognition system.

A second question concerns the systematic value of these characters. Fig. 86 reveals that the admetus-clade only consists of monomorphic brown species and the dolus-clade only contains species with brown- or white-coloured males, but A. sennanensis and A. mithridates are found within the poseidon- and several other brown Agrodiaetus in the carmon-clade. In the case of A. mithridates it was noted already by HESSELBARTH et al. (1995) that this species can not be closely related to the monomorphic Agrodiaetus of the admetus-group, e.g. because of the different structure of the androconial patches. Differences can also be found in other taxa, e.g. in the antidolus-species group where the androconial patch is limited to the area below the central cell whereas it extends along the costal vein in the admetus- and dolus-clade. A most interesting case consists of the three taxa A. peilei (brown males), A. morgani (silvery males) and A. karindus (blue males) which seem to be very closely related (with almost identical nuclear and mtDNA haplotypes) and occur sympatrically in Lorestan (Iran), but have different karyotypes. Although the underside wing pattern is extremely similar in these three taxa they were even placed into different groups due to the striking differences in upperside wing colour. This case might be an interesting example of a sexually selected character displacement reinforcing premating isolation.

Current data suggest that in Agrodiaetus the loss of the blue iridescent wing colour coupled with the expansion of the androconial patches requires few genetic changes and happened independently several times in the course of the radiation of Agrodiaetus. Therefore the value of these characters to infer the systematics and evolutionary history of Agrodiaetus is limited, and the group of “brown” Agrodiaetus apparently is no monophyletic unit. Discoloration, the loss of the iridescent blue colour is also found in many other genera of Lycaenidae, see BÁLINT & JOHNSON (1997) for an overview, but most species are found within Agrodiaetus and Aricia. The reasons for it are unknown in most cases, but in Meleageria (daphnis) marcida discoloration appears to be an adaptation to local climatic conditions of the northern Elburs with reduced solar radiation (BIRÓ et al. 2003). This could also explain the brown colour of Agrodiaetus valiabadi which occurs in the same area, but it is less plausible for most other brown Agrodiaetus species. Instead, sexual selection might be the driving force in these cases. Further studies coupled with experiments and wing colour measurements which must include the hidden wing pattern in the UV spectrum (like KNÜTTEL & FIEDLER 2001 in Polyommatus icarus) could reveal exciting insights into the evolution of mate recognition systems in Agrodiaetus.

111 Chapter 5

Summary A qualitative analysis of major wing pattern characters in Agrodiaetus revealed that the distinct white streak appears to be an autapomorphy of Agrodiaetus but secondarily can get lost again as an individual variation. The presence of a plesiomorphic wing character in Central Asian Agrodiaetus would suggest an ancestral position but this is not corroborated by the molecular analysis. An obvious strong correlation exists between the loss of blue iridescent coloration of males and an enlargement of their andronconial patches which indicates a switch from a visual sexual recognition system to an olfactorial one. Both character states evolved independently several times in Agrodiaetus and can not be used to define species groups within Agrodiaetus.

112 The role of chromosome evolution in the radiation of Agrodiaetus

Chapter 6: The role of chromosome evolution in the radiation of Agrodiaetus

Introduction Although the incredible variation in chromosome numbers among Agrodiaetus is well-known since the pioneering work of DE LESSE (1960a), the reasons for this pattern are not understood. HESSELBARTH et al. (1995) hypothesize that fission of chromosomes should increase the ability of species to adapt to new environmental conditions, because exchange of chromosomal material during meiosis would be facilitated. Correspondingly, chromosome fusions should lower the recombination potential leading to a more stable genome which would be beneficial for species already adapted to specific, more stable environmental conditions. However, these predictions have never been empirically tested. If these predictions are true, a positive correlation between the number of chromosomes and range size should be expected, because species with very restricted ranges are usually specifically adapted to local conditions, whereas widely distributed species accept broader amplitudes of habitat conditions. This hypothesis will be tested in this chapter.

Another question concerns the role of karyotype evolution in the radiation of Agrodiaetus. Is it the cause for the high number of species in this subgenus? Did it even cause sympatric speciation? In Metazoa sympatric speciation has hardly been proven, but convincing cases are found among recent radiations of freshwater fishes (e.g. MCCUNE & LOVEJOY 1998; RUNDLE et al. 2000; SCHLIEWEN et al. 2001) and it is common in plants where polyploidy often leads to speciation. Putting karyological results into a phylogenetic framework should help to elucidate these questions.

Other interesting aspects of karyotype evolution include biogeographical patterns and phylogenetic signal in chromosome numbers. Do species with high or low chromosome numbers occur in certain biogeographical areas? Is there a direction of change in chromosome numbers? Do numbers change gradually or by stepwise multiplication or division? This chapter aims to find answers to these questions.

Material and methods Chromosome numbers (or their mean in case of a range of numbers) were plotted against the range size of all currently known karyospecies (N=88). A rough score of the range size (RS) was assigned in analogy to KUDRNA (1986), but adapted for the Palaearctic region: 1. Species widespread over the whole (or nearly all of) the Palaearctic region; 2. Species widespread over large parts of the Palaearctic region; 3. Species distributed over one or more smaller parts of the Palaearctic region; 4. Species restricted to one or more territories smaller than the above; 5. Species confined to a small area, such as one mountain range, or a single (known) site. The Spearman rank-difference correlation coefficient was calculated to test for any (positive or negative) correlation between both variables and a Kruskal-Wallis-ANOVA was also conducted to check correlations between range size and chromosome numbers. These tests appear justified because no indication exists that range size is depending on the phylogenetic position.

Two methods were applied to investigate the karyological data in a phylogenetic framework. Firstly, statistical parameters were calculated for each clade obtained from the combined COI- and ITS-2 phylogenetic analysis in order to see if patterns of chromosome evolution differ

113 Chapter 6 between clades. All clades with only one or two taxa were lumped together. The chromosome numbers used were the mean values for each recognized taxon listed in chapter 2 taking into account own as well as literature data. In a second approach, chromosome numbers were mapped onto the ITS-2 phylogenetic network to look for evidence of any direction in karyotype evolution. In this case actual chromosome numbers of the specimens used to construct the network were applied. Only if such data were not available, chromosome numbers were inferred from other sources (including literature data) and marked as such. If chromosome numbers change gradually and a direction of chromosome evolution exists, taxa which are distant from the supposed ancestral haplotype of A. damon should have a more derived karyotype (i.e. highly fissioned or highly fused) than those near the origin of the network.

Results A list of all recognized karyospecies in alphabetical order together with their (mean) haploid chromosome number (n) and range size (RS) is found in Tab. 14 and the number of taxa attributed to each range size category are shown in the following table:

RS 1 2 3 4 5 Taxa 0 2 14 39 33

The correlation between both variables is shown in Fig. 87.

Tab. 14. List of species with (mean) haploid chromosome number and range size Taxon n RS Taxon n RS Taxon n RS Taxon n RS actis 17 4 dantchenkoi 41 5 iphicarmon 29 5 poseidon 21 4 admetus 79 3 deebi 67 5 iphidamon 14 4 poseidonides 24 4 ainsae 109 5 demavendi 71 3 iphigenia 14 3 posthumus 85 5 alcestis 20 3 dizinensis 17 5 iphigenides 66 3 pseudactis 29 4 altivagans 21 4 dolus 124 5 juldusus 67 4 pseudoxerxes 16 4 antidolus 41 4 ectabanensis 18 4 karindus 67 4 putnami 26 5 ardschira 114 4 elbursicus 17 4 khorasanensis 84 4 ripartii 90 2 aroaniensis 48 5 eriwanensis 32 4 klausschuriani 56 5 sennanensis 29 4 artvinensis 22 5 ernesti 18 5 kurdistanicus 60 5 sertavulensis 20 5 aserbeidschanus 23 4 erschoffii 14 3 lycius 22 5 shahrami 130 5 baytopi 27 4 fabressei 90 4 maraschi 16 5 shamil 17 4 birunii 11 5 femininoides 27 4 menalcas 85 3 sigberti 27 4 caeruleus 20 4 firdussii 28 3 merhaba 17 5 surakovi 50 4 carmon 82 3 fulgens 103 5 mithridates 24 3 tankeri 21 5 carmonides 16 4 galloi 66 5 mofidii 35 4 theresiae 63 5 ciscaucasicus 16 4 gorbunovi 20 4 morgani 26 4 transcaspicus 53 4 cyaneus 18 4 guezelmavi 42 5 ninae 35 4 turcicola 20 5 dagestanicus 40 4 hamadanensis 22 4 paulae 17 5 turcicus 24 4 dama 41 5 hopfferi 15 3 peilei 39 5 valiabadi 23 5 damocles 25 3 huberti 35 4 pfeifferi 107 4 wagneri 17 4 damon 45 2 humedasae 38 5 phyllis 80 3 zapvadi 19 5 damone 67 3 interjectus 31 5 pierceae 22 5 zarathustra 22 4

114 The role of chromosome evolution in the radiation of Agrodiaetus

s

r 140

be 120 100 num

e 80 m

o 60 s

o 40 m

o 20 r

h 0 C 2345 Range size

Fig. 87. Correlation between chromosome numbers and range Neither the Spearman Rank Correlation Coefficient (R = 0.053, p>= 0.621, Z = 0.494) nor the Kruskal-Wallis-ANOVA (H=3.74, p=0.2914) returned a significant result for a correlation between both variables.

Fig. 88 displays the variation of chromosome numbers and Tab. 15 lists statistical parameters for each clade, apart from the few species of the ancestral damon-, iphidamon-, iphigenides- and poseidonides-clade which were lumped together (“others”).

Tab. 15. Statistical parameters of chromosome number variation in Agrodiaetus clades admetus dolus iphigenia erschoffii carmon poseidon others Total Number of taxa 6 11 5 7 20 23 5 77 Minimum 66 19 12 10 16 15 14 10 Maximum 90 125 29 131 82 45 67 131 Mean 80.9 64.8 22.9 56.4 38.9 21.8 33.2 40.9 Median 81.5 48.0 24.0 56.0 37.0 20.5 24.0 27.0 Standard deviation 8.5 38.2 5.9 44.6 18.3 5.2 22.0 29.0

30

25

20 admetus-clade dolus-clade Number of taxa 15 iphigenia-clade 10 erschoffii-clade 5 carmon-clade 0 poseidon-clade Tot 10 al Total 20 pose 30 ca ido 40 rmo n-cla 50 er n-c de 60 scho lade 70 iphig ffii-c 80 en lad

0 d i e 90 olus a-cla ad -clad de 10 metu e 110 s-cla Clades 120 de 130 Chromosome number (n)

Fig. 88. Variation of chromosome numbers in different Agrodiaetus clades

115 Chapter 6

Different clades have significantly different chromosome numbers (Tab. 16).

Tab. 16. Kruskal-Wallis-ANOVA: Chromosome number differences between clades.

Chromosome no. by Clade n Rank sum Mean rank admetus-group 4 256.5 64.13 dolus-group 10 520.5 52.05 iphigenia-group 5 136.0 27.20 erschoffii-group 7 281.5 40.21 carmon-group 20 794.5 39.73 poseidon-group 21 481.0 22.90 others 5 158.0 31.60

Kruskal-Wallis statistic 23.33 p 0.0007 (chisqr approximation, corrected for ties)

The ITS-2 MP-network with chromosome numbers mapped onto it is shown in Fig. 89. The taxa names can be looked up in Fig. 84. If chromosome numbers were inferred from other specimens than those used to construct the network, these are shown in brackets {}.

Subnetworks which resemble the clades obtained in the phylogenetic trees are named and marked with a polygon, and exceptions are also labelled. All taxa in the admetus-group and most taxa in the dolus-group have high chromosome numbers whereas low chromosome numbers prevail in the remaining groups. Exceptions are A. posthumus (n=85) in the erschoffii- and A. carmon in the carmon-group (n=81-82). Low chromosome numbers are often found in stem species (e.g. A. birunii (n=10-11) in the erschoffii-group, A. iphigenia (n=13-14) in the iphigenia-clade, A. hopfferi (n=15-16) in the poseidon-group and A. alcestis (n=19-21) in the dolus-group whereas a higher chromosome number appears to be a derived condition. Numbers between n=30 and n=60 are common in the dolus- and especially the carmon-group but hardly found in most other groups. Exceptions are A. damon (n=45) and two taxa in the poseidon-group, A. mofidii sorkhensis, n=45 and A. klausschuriani (n=56), but the phylogenetic position of the latter taxon is uncertain.

Discussion No significant correlation was found between chromosome numbers and range size which questions the assumption that the number of chromosomes influences the ability of a species to adapt to environmental conditions. Circumstantial evidence for differing habitat requirements between taxa with low and taxa with high chromosome numbers is not apparent either. Closely related karyospecies inhabit similar habitats, e.g. A. alcestis (n=19-21) and A. menalcas (n=85) in Anatolia, A. birunii (n=10-11) and A. posthumus (n=85) in central and eastern Elburs Mts. or A. morgani (n=27) and A. karindus (n=66-68) in Lorestan, Iran. Currently it seems more reasonable to assume that karyological differences are not adaptive, but occur more or less accidentally in Agrodiaetus.

116 The role of chromosome evolution in the radiation of Agrodiaetus

Fig. 89. Chromosome numbers mapped onto the ITS2-MP-network

117 Chapter 6

In order to see if there is a directional change of chromosome numbers in the radiation of Agrodiaetus, we shall have a first look at possible sister species of the ancestral A. damon, which has a chromosome number of n=45. Such a number is not found in any close relative of Agrodiaetus (apart from Lysandra bellargus) and appears to be caused by fission of the modal and probably ancestral number of n=22-24 which is found in most Polyommatus species. Species with higher numbers are only found in the derived subgenus Lysandra (reaching from n=24 in the ancestral L. syriaca to n=87-92 in L. coridon) and even higher in a group of closely related species sometimes united into the subgenus Plebicula (LORKOVIĆ 1990): P. dorylas (n=147-151), P. nivescens from Spain (n=190-191) and P. atlanticus from Morocco (n=221-223). Of interest is also the variation found within P. cornelia (n=26-38) which appears closely related to A. damon in the ITS-2 network. The closest relatives of A. damon among Agrodiaetus according to the ITS-2 network have chromosome numbers which are multiples or fractions of n=45: almost triple in A. shahrami (n=130), double in A. ripartii (n=90), half in A. valiabadi (n=23) and a third in A. iphidamon (n=14). Thus it appears that considerable variations in chromosome numbers have appeared early in the radiation of Agrodiaetus. Accordingly, chromosome numbers also vary in each Agrodiaetus clade, although differences can be seen between clades. The admetus-clade has the highest mean number and variation is low. The highest variation is found in the (mainly Iranian) erschoffii- clade which also has the taxa with the lowest (A. birunii, n=10) and highest (A. shahrami, n=130) chromosome number found in Agrodiaetus. The poseidon-clade which includes mainly Anatolian faunal elements has the lowest variation and also the lowest mean in chromosome numbers, most of which vary between n=15 and n=35. Most chromosome number changes in this group (and part of the carmon-group) are due to slight variations of the modal value 24. On the other hand, high chromosome numbers seem to appear suddenly, probably by simultaneous fission of the whole chromosome set which leads to multiple numbers. Phylogenetically they appear to be in a deadlock position. Fig. 89 does not show any clear case where highly fissioned chromosomes were fused again to low numbers.

Interestingly, the most extreme values are not found in the most derived taxa but they appear mainly in ancestral clades. Chromosome numbers in the poseidon-group, which turned out to be the crown group of Agrodiaetus in the molecular phylogenetic analysis and which includes many taxa in the process of speciation, has the lowest variation in chromosome numbers most of which are still around the modal value. This also indicates that most of the drastic changes of chromosome numbers happened early in the radiation of Agrodiaetus and therefore corroborates the taxonomic practice to attribute species status to karyospecies with highly divergent chromosome numbers.

Most closely related karyospecies are allopatric or parapatric in distribution. If karyologically well-differentiated sister species occur in sympatry, they also differ considerably in phenotype: A. alcestis (brown males, n=19-21) and A. menalcas (white males, n=85) in Anatolia, A. hopfferi (silvery males, n=15) and A. poseidon (blue males, n=19-22) in Anatolia, or A. morgani (silvery males, n=27), A. peilei (brown males, n=39) and A. karindus (blue males, n=66-68) in Lorestan, Iran. Apparently the coexistence is only possible if females can differentiate the males of closely related species by their phenotype. Otherwise extensive hybridization between different “karyospecies” would occur and difficulties in the pairing of chromosomes during meiosis would lead to a very low fitness of the offspring. Therefore it can be concluded that there is little evidence for sympatric speciation caused by changes in karyotype. Instead, changes in chromosome number seem to be a common by-product of allopatric speciation and phenotypic differentiation is necessary for sister species to occur in sympatry. However, it is probable that chromosomal changes have been important for the

118 The role of chromosome evolution in the radiation of Agrodiaetus radiation of Agrodiaetus during the Pleistocene. Due to periodically changing climate conditions, altitudinal range shifts of populations may have lead to their isolation from each other during the interstadia. Chromosomal changes which occured in allopatric populations confined to isolated mountain tops did not allow remixing of gene pools when their ranges met again during recurrent periods of cooler climate. These processes could explain the high species diversity in Agrodiaetus, in particular in areas with complex topographic relief such as Anatolia, Transcaucasia or Iran.

Summary A correlation analysis of chromosome numbers and range size did not support the hypothesis that the number of chromosomes has an adaptive value. A comparison of chromosome numbers between evolutionary units revealed different patterns. High numbers dominate in the two clades with exclusively brown or silvery-white males. Fission events have happened early in the radiation of Agrodiaetus and independently within different clades. A high degree of fragmentation appeared to represent an irreversible condition when chromosome numbers were mapped onto a phylogenetic network. No evidence was found that changes of chromosome numbers lead to sympatric speciation, rather karyotype evolution in allopatry could have prevented introgression between genotypes if allopatrically evolved taxa came secondarily into sympatry. In this regard, karyotype differentiation may have played a major role in the radiation of Agrodiaetus during the Pleistocene.

119 Summary

Summary

The subgenus Agrodiaetus which is distributed extensively in the Palaearctic region and especially in Southwest Asia is a species-rich group of blues which are often very similar morphologically but show a large variation in their chromosome numbers (n=10-125). The karyology of 64 taxa was successfully studied and the previously unknown chromosome numbers of 17 taxa were revealed. By means of molecular techniques it was possible to clarify the phylogeny of this group for the first time. A. damon, a species which is extremely widespread in the Palearctic region and has a constant chromosome number of n=45, is the sister-species of all other species of Agrodiaetus that have been tested. Presumably the colonization of Europe took place before the main radiations in the Anatolian-Caucasian- Iranian region. The greatest differences in the number of chromosomes were found in allopatrically distributed sister-species. It was surprising that morphologically extremely similar sympatric species with different numbers of chromosomes were not closely related, but often belonged to totally different clades, whereas some closely related sympatric species differed greatly in wing colours. This leads to the conclusion that changes in the number of chromosomes do not lead to sympatric speciation, but instead appear as a by-product of allopatric speciation and that such young species can only occur in sympatry after a sufficient differentiation in their phenotype to exclude erroneous matings. The comparison of mitochondrial with nuclear DNA sequences has also shown that hybridizations in Agrodiaetus are rare events. Supposedly they do not occur more frequently in Agrodiaetus than in other Lycaenidae. Only one of the specimen tested, which already drew our attention by its intermediate wing coloration, after molecular testing proved to be a hybrid of two nearly related but quite differently coloured species.

Zusammenfassung

Die in der Paläarktis und besonders in Vorderasien weit verbreitete Untergattung Agrodiaetus ist eine sehr artenreiche Gruppe von Bläulingen, welche sich morphologisch oft sehr ähnlich sehen, sich aber durch eine extrem starke Variation ihrer Chromosomenzahlen (n=10-125) auszeichnen. 64 Taxa konnten karyologisch erfolgreich untersucht und die bisher unbekannte Chromosomenzahl von 17 Taxa aufgedeckt werden. Mit Hilfe molekularer Techniken ist es nun erstmals gelungen, die Phylogenie dieser Gruppe weitgehend aufzuklären. Demnach ist A. damon, eine Art mit extrem weiter Verbreitung in der Paläarktis und konstanter Chromosomenzahl (n=45), die Schwesterart aller übrigen untersuchten Agrodiaetus-Arten. Die Besiedlung Europas erfolgte vermutlich schon vor den Hauptradiationen im anatolisch- kaukasich-iranischen Raum. Die größten Unterschiede in den Chromosomenzahlen finden sich bei allopatrisch verbreiteten Schwesterarten. Überraschend war, dass morphologisch extrem ähnliche sympatrisch vorkommende Arten mit unterschiedlichen Chromosomenzahlen nicht näher miteinander verwandt sind, sondern oft sogar zu ganz verschiedenen Verwandtschaftsgruppen gehören, wohingegen sich einige sehr nahe verwandte sympatrisch vorkommende Arten stark in der Flügelfärbung unterscheiden. Daraus lässt sich der Schluss ziehen, dass Veränderungen der Chromosomenzahlen nicht zu sympatrischer Artbildung führen, sondern stattdessen ein Nebenprodukt allopatrischer Artbildung darstellen und dass solche jungen Arten erst dann wieder in Sympatrie auftreten können, wenn sie sich auch phänotypisch ausreichend differenziert haben, um Fehlpaarungen zu vermeiden. Der Vergleich von mitochondrialen mit Kern-DNA-Sequenzen hat auch gezeigt, dass Hybridisationen bei Agrodiaetus seltene Ereignisse darstellen. Sie treten vermutlich nicht viel

120 Acknowledgements häufiger auf als bei anderen Lepidopteren-Gattungen. Nur einer der untersuchten Falter, der bereits durch eine intermediäre Färbung aufgefallen war, erwies sich nach molekularer Untersuchung als Hybrid zwischen zwei nahe verwandten, aber recht unterschiedlich gefärbten Arten.

Acknowledgements

This work would not have been successfully accomplished without the help of many collegues whose support and contributions I gratefully acknowledge:

To my mentor Prof. Dr. Clas M. Naumann (Bonn) I am especially indebted. He initiated this work and I have to thank him for his continuous and generous support at the Zoologisches Forschungsinstitut und Museum Alexander Koenig and his sympathy in difficult situations of life. I was especially pleased to be able to accompagny him on an excursion to the core area of Agrodiaetus, where he also helped with the tedious work of preparations. Without his aid and his outstanding expertise this work would not have been accomplished. Prof. Dr. Konrad Fiedler (Bayreuth) brought in his excellent knowledge on Lycaenidae and thus was instrumental in initiating this project as well as bringing it to a satisfactory end. I am looking forward to future cooperation. Dr. habil. Bernhard Misof (Bonn) I have to thank for taking the task of Korreferat. His excellent supervision of laboratory work was indispensable for sequencing such a great amount of material. Dr. Jurate De Prins (Antwerpen) instructed me in her excellent method of karyological preparations and helped in preparing and analyzing numerous karyological slides. The hospitality of herself and her husband, Willy De Prins, during the many visits to Antwerp was extraordinary, and I am grateful for their friendship and two extremely successful joint expeditions to Anatolia. Dr. Vladimir Lukhtanov (St. Petersburg), the leading expert on Agrodiaetus karyology, demonstrated his new method of two-phase chromosome preparation as a visiting scientist to the Museum Koenig. He supported me with the preparation and analysis of a great number of preparations. I am greatful for his cordial cooperation. Karen Meusemann (Bonn) did a major proportion of chromosome preparations in the course of her diploma thesis on the karyology of Agrodiaetus. Thanks to her unusual efforts it was possible to analyze such a lot of preparations, and our discussions were helpful for both of us. I thank my other student assistents, Esther Meyer, Ruth Rottscheidt, Meike Thomas and Manuela Brenk for their dedicated support with molecular lab work. Manuela also helped in the production of several diagrams. Without this help the work would not have been completed yet. Most helpful was the cooperation of the other members of the Museum’s lab crew. Above all I have to thank Dr. Axel Hille, the first head of lab, for his introduction, Rainer Sonnenberg and Anja Schunke for further tutoring and numerous helps during the whole project and Oliver Niehuis for excellent cooperation in sequencing and the organization of the joint excursion to Morocco.

121 Acknowledgements

Claudia Etzbauer, the soul of our lab, secured excellent working conditions, and with her sense of humour helped in frustrating periods of work. I thank the lepidopterists of the Museum Koenig, Dr. Dieter Stüning, Dr. Wolfgang Speidel and Dr. Harald Fänger for friendly support and numerous hints. Dirk van der Poorten & Alain Olivier (Antwerpen) helped to identify most of the Anatolian material and organized the two most successful joint expeditions to Central and Eastern Turkey. Dr. Wolfgang Eckweiler (Frankfurt), the leading expert of Iranian Agrodiaetus, contributed valuable material, and I am grateful for his help in identifying Iranian material and many inspiring discussions. Dr. Klaus G. Schurian (Sulzbach/Taunus) also helped in the identification of Agrodiaetus from Iran. Thank you for your friendship and interesting discussions. Dr. Sigbert Wagener (Oberhausen) magnanimously lent me his representative Agrodiaetus collection from Anatolia and explained his ideas on the evolution of Agrodiaetus. Dr. Alexandre Dantchenko (Moskow) supported me with material from Armenia. I thank him for interesting discussions during his time as visiting scholar to the Museum Koenig. John Coutsis (Athens) provided important Agrodiaetus-material from Greece and I also thank him for most interesting information on the genital morphology of Agrodiaetus. He also did preparations and excellent genitalia drawings from my material. An evaluation of this material was not possible for this thesis but will be done in a forthcoming publication of John and further joint work. Dr. José Munguira (Madrid) not only provided material from Spain but also helped with information on Spanish localities of Agrodiaetus. With Dr. Otakar Kudrna (Schweinfurt) I had interesting discussions and I thank him for providing two important Lysandra taxa for this project. Dr. Emilio Balletto (Torino) was very helpful when my car broke down during a visit to Torino on the excursion to Italy and I am greatful for his hospitality. Dr. Hendrik-Jan Megens (Leiden) supplied information on PCR-primers and a copy of his excellent thesis. Sven Erlacher (Jena) also helped with information on PCR primers for ND1. Thank you!

My loving wife Paula not only helped in looking after our child and the household while I worked long hours in the lab. She also contributed to this thesis considerably by scanning Agrodiaetus wings and typing a part of the reference list. Thank you very much for your support, even though my scientific work caused time and financial constraints to our family. Dear little Pascal (sorry, you have grown already and tomorrow you will start school) has encouraged me with his cheerfulness, even in difficult periods.

Finally, I am especially grateful to my parents, Wolfgang & Helga Wiemers (Münster) for their continuous support throughout my postgraduate studies. Without their help and encouragement this work could not have been carried out.

122 References

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142 Hilfsmittel & Erklärung

Hilfsmittel

Die Arbeit wurde auf einem PC mit MS Word 2000 verfasst. Die Tabellen wurden in MS Excel 2000 erstellt und für die Bearbeitung von Graphiken wurde MS Photo Editor und Adobe Photoshop Elements verwendet. Statistische Berechnungen erfolgten mit Analyse-It V.1.6.8. Für die molekulargenetische Auswertung kamen folgende Computerprogramme zum Einsatz: BioEdit V.5.0.9, ClustalX V.1.8.1, ForCon V.1.0, Mega V.1.0, V.2.1 & V.3.0 beta, MrBayes V.3.0b4, PAUP V.4.0 beta10, TCS V.1.13 und TreeView V.1.6.6.

Erklärung

Ich versichere, dass ich diese Arbeit selbständig verfasst, keine anderen Quellen und Hilfsmittel als die angegebenen benutzt und die Stellen der Arbeit, die anderen Werken dem Wortlaut oder dem Sinn nach entnommen sind, kenntlich gemacht habe. Die Arbeit hat in gleicher oder ähnlicher Form keiner anderen Prüfungsbehörde vorgelegen.

Bonn, den 28. September 2003

143

144 Appendix 1 Karyological results in Agrodiaetus

Code Genus Species Chromosomes Date Locality Altitude Province Country WE 02491 Agrodiaetus achaemenes many (atypical) 21.07.2002 Gardaneh ye Cheri 2800-3000m Bakhtiari Iran WE 02492 Agrodiaetus achaemenes many (atypical) 21.07.2002 Gardaneh ye Cheri 2800-3000m Bakhtiari Iran MW 98162 Agrodiaetus actis n=ca.17 25.07.1998 Gökpinar 1700 m Sivas Turkey MW 98166 Agrodiaetus actis n=17 25.07.1998 Gökpinar 1700 m Sivas Turkey MW 98057 Agrodiaetus admetus n=80 13.07.1998 Camkuyuzu 1750 m Antalya Turkey MW 98082 Agrodiaetus admetus n=ca.73-80 15.07.1998 Cukurelma 1300 m Antalya Turkey MW 98245 Agrodiaetus admetus n=78-80 29.07.1998 Saimbeyli-Fälle 1200-1500 m Adana Turkey MW 01003 Agrodiaetus ainsae n=100-108 17.07.2001 Ilarduya 550 m Alava Spain MW 01004 Agrodiaetus ainsae n=ca.108 17.07.2001 Ilarduya 550 m Alava Spain MW 01054 Agrodiaetus ainsae n=ca.108 20.07.2001 Sta. Maria 500 m Huesca Spain MW 00229 Agrodiaetus alcestis n=ca.19 16.07.2000 Qazayd Dagh 2300 m Zanjan Iran MW 00231 Agrodiaetus alcestis n=ca.19 16.07.2000 Qazayd Dagh 2300 m Zanjan Iran MW 98212 Agrodiaetus alcestis n=21 28.07.1998 Saimbeyli-Fälle 1500 m Adana Turkey MW 98315 Agrodiaetus alcestis n=20 06.08.1998 Yellibeli Geçidi 1800 m Karaman Turkey MW 99299 Agrodiaetus alcestis n=19 18.07.1999 Çatak 1600-1900 m Van Turkey MW 99380 Agrodiaetus alcestis n=19 20.07.1999 Yüksekova 1800 m Hakkari Turkey MW 99165 Agrodiaetus altivagans n=ca.21 15.07.1999 Çaglayan 1500 m Erzincan Turkey MW 99240 Agrodiaetus altivagans n=21 17.07.1999 Güzeldere Geç. 2500 m Van Turkey MW 99253 Agrodiaetus altivagans n=ca.23 17.07.1999 Güzeldere Geç. 2500 m Van Turkey MW 99338 Agrodiaetus altivagans n=ca.21 19.07.1999 Güzeldere Geç. 2600 m Van Turkey MW 99353 Agrodiaetus altivagans n=21-22 19.07.1999 Güzeldere Geç. 2500 m Van Turkey MW 99376 Agrodiaetus antidolus n>30 20.07.1999 Yüksekova 1800 m Hakkari Turkey MW 99377 Agrodiaetus antidolus n=42 20.07.1999 Yüksekova 1800 m Hakkari Turkey MW 99378 Agrodiaetus antidolus n>40 20.07.1999 Yüksekova 1800 m Hakkari Turkey MW 99389 Agrodiaetus antidolus n=40-42 20.07.1999 Dilezi Geçidi 2100 m Hakkari Turkey MW 99393 Agrodiaetus antidolus n=38-43 21.07.1999 Haruna Geçidi 2000 m Hakkari Turkey MW 99394 Agrodiaetus antidolus n=42 21.07.1999 Haruna Geçidi 2000 m Hakkari Turkey MW 99403 Agrodiaetus antidolus n=ca.40 22.07.1999 Dez Çay 1500 m Hakkari Turkey MW 99406 Agrodiaetus antidolus n=ca.44 22.07.1999 Dez Çay 1500 m Hakkari Turkey MW 99058 Agrodiaetus artvinensis n=20-25 08.07.1999 Kiliçkaya 1350 m Artvin Turkey MW 99064 Agrodiaetus artvinensis n=21-22 08.07.1999 Kiliçkaya 1350 m Artvin Turkey MW 99372 Agrodiaetus baytopi n=27-28 19.07.1999 Güzeldere Geç. 2500 m Van Turkey MW 00060 Agrodiaetus birunii n=11 11.07.2000 Pul-e Zanguleh 2400 m Mazandaran Iran MW 00061 Agrodiaetus birunii 2n=ca.21 11.07.2000 Pul-e Zanguleh 2400 m Mazandaran Iran MW 00062 Agrodiaetus birunii 2n=20-22 11.07.2000 Pul-e Zanguleh 2400 m Mazandaran Iran MW 00068 Agrodiaetus birunii n=11 11.07.2000 Gardaneh-ye Lavashm 2900 m Mazandaran Iran MW 00069 Agrodiaetus birunii 2n=ca.20 11.07.2000 Gardaneh-ye Lavashm 2900 m Mazandaran Iran MW 00070 Agrodiaetus birunii 2n=20-22 11.07.2000 Gardaneh-ye Lavashm 2900 m Mazandaran Iran MW 00071 Agrodiaetus birunii 2n=22 11.07.2000 Gardaneh-ye Lavashm 2900 m Mazandaran Iran MW 00072 Agrodiaetus birunii 2n=20-22 11.07.2000 Gardaneh-ye Lavashm 2900 m Mazandaran Iran MW 00102 Agrodiaetus birunii n=11 12.07.2000 Dizin Pass 3000 m Tehran Iran MW 00103 Agrodiaetus birunii n=11 12.07.2000 Dizin Pass 3000 m Tehran Iran MW 00115 Agrodiaetus birunii n=10-11 12.07.2000 Shemshak 2900 m Tehran Iran MW 00267 Agrodiaetus birunii n=10 18.07.2000 Veresk 1800-1950 m Mazandaran Iran MW 00444 Agrodiaetus birunii n=11 26.07.2000 Polur 2200 m Tehran Iran MW 00445 Agrodiaetus birunii 2n=20 26.07.2000 Polur 2200 m Tehran Iran MW 00450 Agrodiaetus birunii 2n=20-22 26.07.2000 Polur 2200 m Tehran Iran MW 00457 Agrodiaetus birunii 2n=20-22 26.07.2000 Polur 2200 m Tehran Iran MW 00463 Agrodiaetus birunii 2n=20-22 26.07.2000 Polur 2200 m Tehran Iran MW 00476 Agrodiaetus birunii 2n=20-22 28.07.2000 Golestanak 2700-3200 m Mazandaran Iran MW 00480 Agrodiaetus birunii 2n=ca.20 28.07.2000 Golestanak 2700-3200 m Mazandaran Iran MW 00482 Agrodiaetus birunii 2n=ca.20 28.07.2000 Golestanak 2700-3200 m Mazandaran Iran MW 00485 Agrodiaetus birunii 2n=ca.20 28.07.2000 Golestanak 2700-3200 m Mazandaran Iran MW 00487 Agrodiaetus birunii 2n=ca.20 28.07.2000 Golestanak 2700-3200 m Mazandaran Iran MW 00503 Agrodiaetus birunii 2n=20 31.07.2000 Takht-e Suleyman 3000 m Mazandaran Iran MW 00547 Agrodiaetus birunii n=10 04.08.2000 Dizin Pass 3200-3300 m Tehran Iran MW 00335 Agrodiaetus caeruleus n=20 21.07.2000 Shakuh 2600 m Golestan Iran MW 00409 Agrodiaetus caeruleus n=20 23.07.2000 Hajiabad 2150 m Golestan Iran MW 00421 Agrodiaetus caeruleus n=19-20 23.07.2000 Hajiabad 2150 m Golestan Iran MW 99060 Agrodiaetus carmon n>=79 08.07.1999 Kiliçkaya 1350 m Artvin Turkey MW 99094 Agrodiaetus cyaneus n=17 11.07.1999 Kagizman 1400 m Kars Turkey MW 99375 Agrodiaetus cyaneus n=18 20.07.1999 Zernek Brj. 1900 m Van Turkey MW 99448 Agrodiaetus cyaneus n=18 23.07.1999 Zernek Brj. 1900 m Van Turkey MW 99475 Agrodiaetus cyaneus n=18-19 25.07.1999 Erek Dagi 2200 m Van Turkey MW 99274 Agrodiaetus dantchenkoi n=40-42 17.07.1999 Kurubas Geçidi 2200 m Van Turkey MW 99276 Agrodiaetus dantchenkoi n=40-43 17.07.1999 Kurubas Geçidi 2200 m Van Turkey MW 99278 Agrodiaetus dantchenkoi n=40-45 17.07.1999 Kurubas Geçidi 2200 m Van Turkey MW 99319 Agrodiaetus dantchenkoi n=42 18.07.1999 Çatak 2000-2200 m Van Turkey MW 99320 Agrodiaetus dantchenkoi n=40-41 18.07.1999 Çatak 2000-2200 m Van Turkey MW 99471 Agrodiaetus dantch.Xmenal. n=45-50 25.07.1999 Erek Dagi 2200 m Van Turkey MW 00014 Agrodiaetus demavendi n>=68 09.07.2000 Samqabad 1900-2100 m Tehran Iran MW 00015 Agrodiaetus demavendi n=65-75 09.07.2000 Samqabad 1900-2100 m Tehran Iran MW 00183 Agrodiaetus demavendi n=ca.70-80 15.07.2000 Dugijan 2000 m Azarbayjan-e Sharqi Iran MW 00185 Agrodiaetus demavendi n=many 15.07.2000 Dugijan 2000 m Azarbayjan-e Sharqi Iran MW 00186 Agrodiaetus demavendi n=many 15.07.2000 Dugijan 2000 m Azarbayjan-e Sharqi Iran MW 00189 Agrodiaetus demavendi n=many 15.07.2000 Dugijan 2000 m Azarbayjan-e Sharqi Iran MW 99102 Agrodiaetus demavendi n>50 11.07.1999 Akçay 1200 m Kars Turkey MW 99105 Agrodiaetus demavendi n=66 11.07.1999 Akçay 1200 m Kars Turkey

- 145 - Appendix 1 Karyological results in Agrodiaetus

Code Genus Species Chromosomes Date Locality Altitude Province Country MW 99110 Agrodiaetus demavendi n=55-70 11.07.1999 Akçay 1200 m Kars Turkey MW 99142 Agrodiaetus demavendi n=ca.66 13.07.1999 Akçay 1200 m Kars Turkey MW 99148 Agrodiaetus demavendi n=57-65 13.07.1999 Akçay 1200 m Kars Turkey MW 99150 Agrodiaetus demavendi n=64-76 13.07.1999 Akçay 1200 m Kars Turkey MW 99152 Agrodiaetus demavendi n>30 13.07.1999 Akçay 1200 m Kars Turkey MW 99382 Agrodiaetus demavendi n=67-71 20.07.1999 Yüksekova 1800 m Hakkari Turkey MW 99383 Agrodiaetus demavendi n=59 20.07.1999 Yüksekova 1800 m Hakkari Turkey MW 99384 Agrodiaetus demavendi n=67 20.07.1999 Yüksekova 1800 m Hakkari Turkey MW 99386 Agrodiaetus demavendi n=many 20.07.1999 Yüksekova 1800 m Hakkari Turkey MW 99391 Agrodiaetus demavendi n=many 20.07.1999 Dilezi Geçidi 2100 m Hakkari Turkey MW 99392 Agrodiaetus demavendi n=many 20.07.1999 Dilezi Geçidi 2100 m Hakkari Turkey MW 99405 Agrodiaetus demavendi n=many 22.07.1999 Dez Çay 1500 m Hakkari Turkey MW 99407 Agrodiaetus demavendi n>60 22.07.1999 Dez Çay 1500 m Hakkari Turkey MW 00539 Agrodiaetus dizinensis n=17 04.08.2000 Dizin Pass 3200-3300 m Tehran Iran MW 00051 Agrodiaetus elbursicus n=17 11.07.2000 Nesa 2100 m Tehran Iran MW 00052 Agrodiaetus elbursicus 2n>=28 11.07.2000 Nesa 2100 m Tehran Iran MW 00057 Agrodiaetus elbursicus n=ca.17 11.07.2000 Kendevan 2150 m Mazandaran Iran MW 00058 Agrodiaetus elbursicus n=18 11.07.2000 Pul-e Zanguleh 2400 m Mazandaran Iran MW 00059 Agrodiaetus elbursicus n=16 11.07.2000 Pul-e Zanguleh 2400 m Mazandaran Iran MW 00065 Agrodiaetus elbursicus n=17 11.07.2000 Gardaneh-ye Lavashm 2900 m Mazandaran Iran MW 00107 Agrodiaetus elbursicus n=ca.15-20 12.07.2000 Shahrestanak 2000 m Tehran Iran MW 00110 Agrodiaetus elbursicus n=17-18 12.07.2000 Shahrestanak 2000 m Tehran Iran MW 00232 Agrodiaetus elbursicus n=18 16.07.2000 Qazayd Dagh 2300 m Zanjan Iran MW 00316 Agrodiaetus elbursicus n=16-18 19.07.2000 Asadbar 2500 m Tehran Iran MW 98097 Agrodiaetus ernesti n=18 21.07.1998 Dedegöl Geçidi 1700 m Isparta Turkey MW 00393 Agrodiaetus erschoffii n=ca.14 23.07.2000 Hajiabad 2150 m Golestan Iran MW 00406 Agrodiaetus erschoffii n=ca.13-14 23.07.2000 Hajiabad 2150 m Golestan Iran MW 01036 Agrodiaetus fabressei n=many 19.07.2001 Abejar 1100 m Soria Spain MW 01039 Agrodiaetus fabressei n>75 19.07.2001 Abejar 1100 m Soria Spain MW 00226 Agrodiaetus femininoides n=27 16.07.2000 Qazayd Dagh 2300 m Zanjan Iran MW 00227 Agrodiaetus femininoides n=ca.27 16.07.2000 Qazayd Dagh 2300 m Zanjan Iran MW 00228 Agrodiaetus femininoides n=27 16.07.2000 Qazayd Dagh 2300 m Zanjan Iran WE 02671 Agrodiaetus femininoides n=27 31.07.2002 Khalkhal, Gollijeh 1900m Ardabil Iran WE 02672 Agrodiaetus femininoides n=27 31.07.2002 Khalkhal, Gollijeh 1900m Ardabil Iran MW 00216 Agrodiaetus firdussii n=ca.29 16.07.2000 Qazayd Dagh 2300 m Zanjan Iran MW 00217 Agrodiaetus firdussii n=29 16.07.2000 Qazayd Dagh 2300 m Zanjan Iran MW 00218 Agrodiaetus firdussii n=28 16.07.2000 Qazayd Dagh 2300 m Zanjan Iran MW 00234 Agrodiaetus firdussii n>=24 16.07.2000 Qazayd Dagh 2300 m Zanjan Iran MW 99006 Agrodiaetus firdussii n=ca.30 05.07.1999 Çaglayan 1500 m Erzincan Turkey MW 99007 Agrodiaetus firdussii n=28-31 05.07.1999 Çaglayan 1500 m Erzincan Turkey MW 99172 Agrodiaetus firdussii n=32 15.07.1999 Çaglayan 1500 m Erzincan Turkey MW 99241 Agrodiaetus firdussii n=25 17.07.1999 Güzeldere Geç. 2500 m Van Turkey MW 99247 Agrodiaetus firdussii n=ca.24 17.07.1999 Güzeldere Geç. 2500 m Van Turkey MW 99413 Agrodiaetus firdussii n=25 22.07.1999 Dez Çay 1500 m Hakkari Turkey MW 99417 Agrodiaetus firdussii n=26 22.07.1999 Dez Çay 1500 m Hakkari Turkey MW 99422 Agrodiaetus firdussii n=ca.25 22.07.1999 Dez Çay 1500 m Hakkari Turkey MW 99441 Agrodiaetus firdussii n=25 23.07.1999 Dez Çay 1500-1700 m Hakkari Turkey MW 99443 Agrodiaetus firdussii n=27 23.07.1999 Dez Çay 1500-1700 m Hakkari Turkey MW 99444 Agrodiaetus firdussii n=ca.25 23.07.1999 Dez Çay 1500-1700 m Hakkari Turkey MW 99457 Agrodiaetus firdussii n=26 24.07.1999 Çatak 1600-1900 m Van Turkey MW 01106 Agrodiaetus fulgens n=ca.98-103 23.07.2001 Sta. Coloma de Queralt 700 m Tarragona Spain MW 01107 Agrodiaetus fulgens n>90 23.07.2001 Sta. Coloma de Queralt 700 m Tarragona Spain MW 00129 Agrodiaetus gorbunovi n=20 13.07.2000 Ahar Pass 1800-1850 m Azarbayjan-e Sharqi Iran MW 00177 Agrodiaetus gorbunovi n=ca.20 15.07.2000 Dugijan 2000 m Azarbayjan-e Sharqi Iran WE 02675 Agrodiaetus gorbunovi n=19 31.07.2002 Khalkhal, Gollijeh 1900m Ardabil Iran MW 98293 Agrodiaetus guezelmavi n=43 04.08.1998 Taskent 1450 m Konya Turkey MW 98297 Agrodiaetus guezelmavi n=ca.40-43 04.08.1998 Taskent 1450 m Konya Turkey MW 98298 Agrodiaetus guezelmavi n=ca.40 04.08.1998 Taskent 1450 m Konya Turkey MW 00004 Agrodiaetus hamadanensis n=22 09.07.2000 Samqabad 1900-2100 m Tehran Iran MW 00005 Agrodiaetus hamadanensis n=ca.22 09.07.2000 Samqabad 1900-2100 m Tehran Iran MW 00032 Agrodiaetus hamadanensis n=ca.22 10.07.2000 Safedabad 2000 m Tehran Iran WE 02676 Agrodiaetus hamadanensis n=21 31.07.2002 Khalkhal, Gollijeh 1900m Ardabil Iran MW 98189 Agrodiaetus hopfferi n=15-16 27.07.1998 Gündüzbey 1300 m Malatya Turkey MW 98190 Agrodiaetus hopfferi n=15-16 27.07.1998 Gündüzbey 1300 m Malatya Turkey MW 98191 Agrodiaetus hopfferi 2n=30 27.07.1998 Gündüzbey 1300 m Malatya Turkey MW 99408 Agrodiaetus hopfferi n=15 22.07.1999 Dez Çay 1500 m Hakkari Turkey MW 99453 Agrodiaetus hopfferi n=15 24.07.1999 Çatak 1600-1900 m Van Turkey MW 99053 Agrodiaetus huberti n=36 08.07.1999 Kiliçkaya 1350 m Artvin Turkey MW 99095 Agrodiaetus huberti n=33-34 11.07.1999 Kagizman 1400 m Kars Turkey MW 99552 Agrodiaetus huberti n=ca.33 29.07.1999 Kop Geçidi 2350 m Bayburt Turkey MW 99590 Agrodiaetus humedasae 2n=ca.70-80 14.08.1999 Pondel 900 m Aosta Italy MW 99591 Agrodiaetus humedasae n=ca.38 14.08.1999 Pondel 900 m Aosta Italy MW 99595 Agrodiaetus humedasae 2n>66 14.08.1999 Pondel 900 m Aosta Italy MW 99596 Agrodiaetus humedasae 2n=ca.76 14.08.1999 Pondel 900 m Aosta Italy MW 99164 Agrodiaetus interjectus n=31 14.07.1999 Çiftlik 1900 m Erzurum Turkey MW 98101 Agrodiaetus iphicarmon n=ca.30 21.07.1998 Dedegöl Geçidi 1700 m Isparta Turkey MW 98102 Agrodiaetus iphicarmon n=ca.30 21.07.1998 Dedegöl Geçidi 1700 m Isparta Turkey MW 98104 Agrodiaetus iphicarmon n=29 21.07.1998 Dedegöl Geçidi 1700 m Isparta Turkey

- 146 - Appendix 1 Karyological results in Agrodiaetus

Code Genus Species Chromosomes Date Locality Altitude Province Country MW 00269 Agrodiaetus iphidamon n=ca.14 18.07.2000 Veresk 1800-1950 m Mazandaran Iran MW 00274 Agrodiaetus iphidamon n=14 18.07.2000 Veresk 1800-1950 m Mazandaran Iran MW 00287 Agrodiaetus iphidamon n=14 18.07.2000 Veresk 1800-1950 m Mazandaran Iran MW 00338 Agrodiaetus iphidamon n=ca.14 21.07.2000 Shakuh 2600 m Golestan Iran MW 00362 Agrodiaetus iphidamon n=14 21.07.2000 Shakuh 2600 m Golestan Iran MW 00381 Agrodiaetus iphidamon n=14 21.07.2000 Shakuh 3000 m Golestan Iran MW 00424 Agrodiaetus iphidamon n=13-15 23.07.2000 Hajiabad 2150 m Golestan Iran MW 98049 Agrodiaetus iphigenia n=14 12.07.1998 Salur Dagi 1700-1900 m Fethiye Turkey MW 98106 Agrodiaetus iphigenia n=15 21.07.1998 Dedegöl Geçidi 1700 m Isparta Turkey MW 99009 Agrodiaetus iphigenia n=12 05.07.1999 Çaglayan 1500 m Erzincan Turkey MW 99154 Agrodiaetus iphigenia n=11-14 13.07.1999 Kagizman 1400 m Kars Turkey MW 99170 Agrodiaetus iphigenia n=12 15.07.1999 Çaglayan 1500 m Erzincan Turkey MW 99465 Agrodiaetus kanduli n=25 24.07.1999 Çatak 1600-1900 mVan Turkey WE 02611 Agrodiaetus karindus n=68 27.07.2002 40 km SW Saqqez 1800-1900m Kordestan Iran WE 02612 Agrodiaetus karindus n>=66 27.07.2002 40 km SW Saqqez 1800-1900m Kordestan Iran MW 00259 Agrodiaetus klausschuriani n=56-58 18.07.2000 Veresk 1800-1950 m Mazandaran Iran MW 00261 Agrodiaetus klausschuriani n>47 18.07.2000 Veresk 1800-1950 m Mazandaran Iran MW 00262 Agrodiaetus klausschuriani n=56 18.07.2000 Veresk 1800-1950 m Mazandaran Iran MW 00264 Agrodiaetus klausschuriani n=56 18.07.2000 Veresk 1800-1950 m Mazandaran Iran MW 99286 Agrodiaetus kurdistanicus n=54-56 18.07.1999 Çatak 1600-1900 m Van Turkey MW 99287 Agrodiaetus kurdistanicus n=56-68 18.07.1999 Çatak 1600-1900 m Van Turkey MW 99288 Agrodiaetus kurdistanicus n=62 18.07.1999 Çatak 1600-1900 m Van Turkey MW 99473 Agrodiaetus kurdistanicus n>=55 25.07.1999 Erek Dagi 2200 m Van Turkey MW 98069 Agrodiaetus lycius n=21-22 15.07.1998 Cukurelma 1300 m Antalya Turkey MW 98070 Agrodiaetus lycius n=21-22 15.07.1998 Cukurelma 1300 m Antalya Turkey MW 98071 Agrodiaetus lycius n=22 15.07.1998 Cukurelma 1300 m Antalya Turkey MW 98072 Agrodiaetus lycius n=21-22 15.07.1998 Cukurelma 1300 m Antalya Turkey MW 98170 Agrodiaetus maraschi n=16 25.07.1998 Gökpinar 1700 m Sivas Turkey MW 98022 Agrodiaetus menalcas n=85 12.07.1998 Gülübeli Geçidi 1500 m Fethiye Turkey MW 98064 Agrodiaetus menalcas n>75 13.07.1998 Camkuyuzu 1750 m Antalya Turkey MW 99496 Agrodiaetus menalcas n=85 25.07.1999 Erek Dagi 2200 m Van Turkey MW 99057 Agrodiaetus merhaba n=ca.17 08.07.1999 Kiliçkaya 1350 m Artvin Turkey MW 99059 Agrodiaetus merhaba n=15-17 08.07.1999 Kiliçkaya 1350 m Artvin Turkey MW 98203 Agrodiaetus mithridates n=23 27.07.1998 Gündüzbey 1300 m Malatya Turkey WE 02453 Agrodiaetus mofidii n=ca.45 17.07.2002 Kuh-e-Sorkh, Kadkan 2100-2500m Khorasan Iran WE 02454 Agrodiaetus mofidii 2n>80 17.07.2002 Kuh-e-Sorkh, Kadkan 2100-2500m Khorasan Iran WE 02614 Agrodiaetus morgani n=27 27.07.2002 40 km SW Saqqez 1800-1900m Kordestan Iran MW 99508 Agrodiaetus ninae n=34 26.07.1999 Agrı 1800 m Agrı Turkey MW 00126 Agrodiaetus paulae n=17 13.07.2000 Ahar Pass 1800-1850 m Azarbayjan-e Sharqi Iran MW 00127 Agrodiaetus paulae n=17 13.07.2000 Ahar Pass 1800-1850 m Azarbayjan-e Sharqi Iran MW 00155 Agrodiaetus paulae n=ca.18 13.07.2000 Ahar Pass 1800-1850 m Azarbayjan-e Sharqi Iran MW 00157 Agrodiaetus paulae n=ca.17 13.07.2000 Ahar Pass 1800-1850 m Azarbayjan-e Sharqi Iran MW 00158 Agrodiaetus paulae n=ca.17 13.07.2000 Ahar Pass 1800-1850 m Azarbayjan-e Sharqi Iran MW 00161 Agrodiaetus paulae n=17 13.07.2000 Ahar Pass 1800-1850 m Azarbayjan-e Sharqi Iran WE 02591 Agrodiaetus peilei n=39 27.07.2002 Qamchiyan 1800-1900m Kordestan Iran WE 02592 Agrodiaetus peilei n=39 27.07.2002 Qamchiyan 1800-1900m Kordestan Iran WE 02593 Agrodiaetus peilei n=39 27.07.2002 Qamchiyan 1800-1900m Kordestan Iran MW 00236 Agrodiaetus phyllis n=many 16.07.2000 Qazayd Dagh 2300 m Zanjan Iran MW 00307 Agrodiaetus phyllis n=76-78 19.07.2000 Asadbar 2900 m Tehran Iran MW 00348 Agrodiaetus phyllis n=82-86 21.07.2000 Shakuh 2600 m Golestan Iran MW 00355 Agrodiaetus phyllis n=many 21.07.2000 Shakuh 2600 m Golestan Iran MW 00452 Agrodiaetus phyllis n>75 26.07.2000 Polur 2200 m Tehran Iran MW 99292 Agrodiaetus pierceae n=21 18.07.1999 Çatak 1600-1900 m Van Turkey MW 99341 Agrodiaetus pierceae n=22-24 19.07.1999 Güzeldere Geç. 2600 m Van Turkey MW 99416 Agrodiaetus pierceae n=22 22.07.1999 Dez Çay 1500 m Hakkari Turkey MW 98137 Agrodiaetus poseidon n=21 22.07.1998 Zelve 1100 m Nevsehir Turkey MW 98138 Agrodiaetus poseidon n=20-21 22.07.1998 Zelve 1100 m Nevsehir Turkey MW 98141 Agrodiaetus poseidon n=21 22.07.1998 Zelve 1100 m Nevsehir Turkey MW 98144 Agrodiaetus poseidon n=21 22.07.1998 Zelve 1100 m Nevsehir Turkey MW 98145 Agrodiaetus poseidon n=20 22.07.1998 Zelve 1100 m Nevsehir Turkey MW 98154 Agrodiaetus poseidon n=21 22.07.1998 Zelve 1100 m Nevsehir Turkey MW 98178 Agrodiaetus poseidon n=19 25.07.1998 Gökpinar 1700 m Sivas Turkey MW 98180 Agrodiaetus poseidon n=19 25.07.1998 Gökpinar 1700 m Sivas Turkey MW 98183 Agrodiaetus poseidon n=19 25.07.1998 Gökpinar 1700 m Sivas Turkey MW 00347 Agrodiaetus posthumus n=ca.85 21.07.2000 Shakuh 2600 m Golestan Iran MW 00330 Agrodiaetus pseudoxerxes n=15 21.07.2000 Shakuh 2600 m Golestan Iran MW 99061 Agrodiaetus putnami n=ca.25 08.07.1999 Kiliçkaya 1350 m Artvin Turkey MW 99501 Agrodiaetus putnami n=25 26.07.1999 Agrı 1800 m Agrı Turkey MW 01014 Agrodiaetus ripartii n=ca.90 18.07.2001 Ubierna 900 m Burgos Spain MW 01015 Agrodiaetus ripartii n=ca.90 18.07.2001 Ubierna 900 m Burgos Spain MW 98100 Agrodiaetus ripartii n>80 21.07.1998 Dedegöl Geçidi 1700 m Isparta Turkey MW 98215 Agrodiaetus ripartii n=ca.90 28.07.1998 Saimbeyli-Fälle 1500 m Adana Turkey MW 99068 Agrodiaetus ripartii n=ca.90 08.07.1999 Kiliçkaya 1350 m Artvin Turkey MW 99196 Agrodiaetus ripartii n>85 15.07.1999 Çaglayan 1500 m Erzincan Turkey MW 99218 Agrodiaetus ripartii n=many 15.07.1999 Çaglayan 1500 m Erzincan Turkey MW 99219 Agrodiaetus ripartii n=many 15.07.1999 Çaglayan 1500 m Erzincan Turkey MW 99220 Agrodiaetus ripartii n=many 15.07.1999 Çaglayan 1500 m Erzincan Turkey MW 99263 Agrodiaetus ripartii n>71 17.07.1999 Kurubas Geçidi 2200 m Van Turkey

- 147 - Appendix 1 Karyological results in Agrodiaetus

Code Genus Species Chromosomes Date Locality Altitude Province Country MW 99574 Agrodiaetus ripartii n=many 29.07.1999 Kop Geçidi 2350 m Bayburt Turkey MW 98261 Agrodiaetus schuriani n=ca.75-80 30.07.1998 Gezbeli Geçidi 1800 m Kayseri Turkey MW 99289 Agrodiaetus sekercioglui n>=46 18.07.1999 Çatak 1600-1900 m Van Turkey MW 98305 Agrodiaetus sertavulensis n=20 06.08.1998 Yellibeli Geçidi 1800 m Karaman Turkey MW 98313 Agrodiaetus sertavulensis n=ca.20 06.08.1998 Yellibeli Geçidi 1800 m Karaman Turkey MW 98279 Agrodiaetus sigberti n=28 31.07.1998 Ala Daglar 2900 m Kayseri Turkey MW 98285 Agrodiaetus sigberti n=25 31.07.1998 Ala Daglar 2700 m Kayseri Turkey MW 98240 Agrodiaetus theresiae n=59 29.07.1998 Saimbeyli-Fälle 1200-1500 m Adana Turkey MW 98241 Agrodiaetus theresiae n=ca.59 29.07.1998 Saimbeyli-Fälle 1200-1500 m Adana Turkey MW 98243 Agrodiaetus theresiae n=63 29.07.1998 Saimbeyli-Fälle 1200-1500 m Adana Turkey MW 99258 Agrodiaetus turcicola n=20 17.07.1999 Kurubas Geçidi 2200 m Van Turkey MW 99262 Agrodiaetus turcicola n=20 17.07.1999 Kurubas Geçidi 2200 m Van Turkey MW 99272 Agrodiaetus turcicola n=ca.18-22 17.07.1999 Kurubas Geçidi 2200 m Van Turkey MW 99273 Agrodiaetus turcicola n=ca.18-22 17.07.1999 Kurubas Geçidi 2200 m Van Turkey MW 99294 Agrodiaetus turcicola n=22 18.07.1999 Çatak 1600-1900 m Van Turkey MW 99314 Agrodiaetus turcicola n=ca.20 18.07.1999 Çatak 2000-2200 m Van Turkey MW 99321 Agrodiaetus turcicola n=20 18.07.1999 Çatak 2000-2200 m Van Turkey MW 99464 Agrodiaetus turcicola n=20 24.07.1999 Çatak 1600-1900 m Van Turkey MW 99478 Agrodiaetus turcicola n=19 25.07.1999 Erek Dagi 2200 m Van Turkey MW 99479 Agrodiaetus turcicola n=20 25.07.1999 Erek Dagi 2200 m Van Turkey MW 99483 Agrodiaetus turcicola n=19-20 25.07.1999 Erek Dagi 2200 m Van Turkey MW 99135 Agrodiaetus turcicus 2n=ca.50 12.07.1999 Badilli 1800-2000 m Igdir Turkey MW 99189 Agrodiaetus turcicus n=ca.24 15.07.1999 Çaglayan 1500 m Erzincan Turkey MW 99203 Agrodiaetus turcicus n=22-24 15.07.1999 Çaglayan 1500 m Erzincan Turkey MW 00498 Agrodiaetus valiabadi n=23 30.07.2000 5 km S Valiabad 1900 m Mazandaran Iran MW 98136 Agrodiaetus wagneri n=18 22.07.1998 Zelve 1100 m Nevsehir Turkey MW 98139 Agrodiaetus wagneri n=19/21 22.07.1998 Zelve 1100 m Nevsehir Turkey MW 99228 Agrodiaetus zapvadi n=18 17.07.1999 Zernek Brj. 1900 m Van Turkey MW 99229 Agrodiaetus zapvadi n=ca.17 17.07.1999 Zernek Brj. 1900 m Van Turkey MW 99373 Agrodiaetus zapvadi n=18 20.07.1999 Zernek Brj. 1900 m Van Turkey MW 99374 Agrodiaetus zapvadi n=19 20.07.1999 Zernek Brj. 1900 m Van Turkey MW 99476 Agrodiaetus zapvadi n=18-20 25.07.1999 Erek Dagi 2200 m Van Turkey WE 02531 Agrodiaetus zarathustra n=ca.22 25.07.2002 30 km W Dorud 2100m Lorestan Iran WE 02533 Agrodiaetus zarathustra n=ca.22 25.07.2002 30 km W Dorud 2100m Lorestan Iran

Chromosome numbers in bold type are new for this taxon.

- 148 - Appendix 2 List of material

Code Sex Genus Species Wgs Fix Kar Alc DNA Cytb CO1 ND1 ITS2 Fig. Date Locality Area Altitude Province Country AD 98001 ♂ Agrodiaetus surakovi 1 1 1 1 1 I-1 20.07.1998 Gnyshik village Aiotzdzorsky range 1800-2200 m Transcaucasia Armenia AD 98002 ♂ Agrodiaetus surakovi 1 1 20.07.1998 Gnyshik village Aiotzdzorsky range 1800-2200 m Transcaucasia Armenia AD 98003 ♂ Agrodiaetus surakovi 1 1 20.07.1998 Gnyshik village Aiotzdzorsky range 1800-2200 m Transcaucasia Armenia AD 98004 ♂ Agrodiaetus surakovi 1 1 20.07.1998 Gnyshik village Aiotzdzorsky range 1800-2200 m Transcaucasia Armenia AD 98005 ♂ Agrodiaetus surakovi 1 1 20.07.1998 Gnyshik village Aiotzdzorsky range 1800-2200 m Transcaucasia Armenia AD 98006 ♂ Agrodiaetus surakovi 1 1 20.07.1998 Gnyshik village Aiotzdzorsky range 1800-2200 m Transcaucasia Armenia AD 98007 ♂ Agrodiaetus surakovi 1 1 20.07.1998 Gnyshik village Aiotzdzorsky range 1800-2200 m Transcaucasia Armenia AD 98008 ♂ Agrodiaetus surakovi 1 1 20.07.1998 Gnyshik village Aiotzdzorsky range 1800-2200 m Transcaucasia Armenia AD 98009 ♂ Agrodiaetus pseudactis 1 1 1 1 1 I-2 20.07.1998 Gnyshik village Aiotzdzorsky range 1800-2200 m Transcaucasia Armenia AD 98010 ♂ Agrodiaetus pseudactis 1 1 20.07.1998 Gnyshik village Aiotzdzorsky range 1800-2200 m Transcaucasia Armenia AD 98011 ♂ Agrodiaetus pseudactis 1 1 20.07.1998 Gnyshik village Aiotzdzorsky range 1800-2200 m Transcaucasia Armenia AD 98012 ♂ Agrodiaetus altivagans 1 1 1 1 1 20.07.1998 Gnyshik village Aiotzdzorsky range 1800-2200 m Transcaucasia Armenia AD 98013 ♂ Agrodiaetus altivagans 1 1 20.07.1998 Gnyshik village Aiotzdzorsky range 1800-2200 m Transcaucasia Armenia AD 98014 ♂ Agrodiaetus altivagans 1 1 20.07.1998 Gnyshik village Aiotzdzorsky range 1800-2200 m Transcaucasia Armenia AD 98015 ♂ Agrodiaetus altivagans 1 1 20.07.1998 Gnyshik village Aiotzdzorsky range 1800-2200 m Transcaucasia Armenia AD 98016 ♂ Agrodiaetus altivagans 1 1 20.07.1998 Gnyshik village Aiotzdzorsky range 1800-2200 m Transcaucasia Armenia AD 98017 ♂ Agrodiaetus altivagans 1 1 20.07.1998 Gnyshik village Aiotzdzorsky range 1800-2200 m Transcaucasia Armenia AD 98018 ♂ Agrodiaetus ninae 1 1 1 1 1 20.07.1998 Gnyshik village Aiotzdzorsky range 1800-2200 m Transcaucasia Armenia AD 98019 ♂ Agrodiaetus ninae 1 1 20.07.1998 Gnyshik village Aiotzdzorsky range 1800-2200 m Transcaucasia Armenia AD 98020 ♂ Agrodiaetus ninae 1 1 20.07.1998 Gnyshik village Aiotzdzorsky range 1800-2200 m Transcaucasia Armenia AD 98021 ♂ Agrodiaetus ninae 1 1 20.07.1998 Gnyshik village Aiotzdzorsky range 1800-2200 m Transcaucasia Armenia AD 98022 ♂ Agrodiaetus ninae 1 1 20.07.1998 Gnyshik village Aiotzdzorsky range 1800-2200 m Transcaucasia Armenia AD 98023 ♂ Agrodiaetus ninae 1 1 20.07.1998 Gnyshik village Aiotzdzorsky range 1800-2200 m Transcaucasia Armenia AD 98024 ♂ Agrodiaetus huberti 1 1 1 1 1 20.07.1998 Gnyshik village Aiotzdzorsky range 1800-2200 m Transcaucasia Armenia AD 98025 ♂ Agrodiaetus huberti 1 1 20.07.1998 Gnyshik village Aiotzdzorsky range 1800-2200 m Transcaucasia Armenia AD 98026 ♂ Agrodiaetus huberti 1 1 20.07.1998 Gnyshik village Aiotzdzorsky range 1800-2200 m Transcaucasia Armenia AD 98027 ♂ Agrodiaetus huberti 1 1 20.07.1998 Gnyshik village Aiotzdzorsky range 1800-2200 m Transcaucasia Armenia AD 98028 ♂ Agrodiaetus huberti 1 1 20.07.1998 Gnyshik village Aiotzdzorsky range 1800-2200 m Transcaucasia Armenia AD 98029 ♂ Agrodiaetus iphigenia 1 1 1 1 20.07.1998 Gnyshik village Aiotzdzorsky range 1800-2200 m Transcaucasia Armenia AD 98030 ♂ Agrodiaetus iphigenia 1 1 20.07.1998 Gnyshik village Aiotzdzorsky range 1800-2200 m Transcaucasia Armenia AD 98031 ♂ Agrodiaetus iphigenia 1 1 20.07.1998 Gnyshik village Aiotzdzorsky range 1800-2200 m Transcaucasia Armenia AD 98032 ♂ Agrodiaetus iphigenia 1 1 20.07.1998 Gnyshik village Aiotzdzorsky range 1800-2200 m Transcaucasia Armenia AD 98033 ♂ Agrodiaetus iphigenia 1 1 20.07.1998 Gnyshik village Aiotzdzorsky range 1800-2200 m Transcaucasia Armenia AD 98034 ♂ Agrodiaetus iphigenia 1 1 20.07.1998 Gnyshik village Aiotzdzorsky range 1800-2200 m Transcaucasia Armenia AD 98035 ♂ Agrodiaetus iphigenia 1 1 20.07.1998 Gnyshik village Aiotzdzorsky range 1800-2200 m Transcaucasia Armenia AD 98036 ♂ Agrodiaetus phyllis 1 1 1 1 1 20.07.1998 Gnyshik village Aiotzdzorsky range 1800-2200 m Transcaucasia Armenia AD 98037 ♂ Agrodiaetus phyllis 1 1 20.07.1998 Gnyshik village Aiotzdzorsky range 1800-2200 m Transcaucasia Armenia AD 98038 ♂ Agrodiaetus phyllis 1 1 20.07.1998 Gnyshik village Aiotzdzorsky range 1800-2200 m Transcaucasia Armenia AD 98039 ♂ Agrodiaetus phyllis 1 1 20.07.1998 Gnyshik village Aiotzdzorsky range 1800-2200 m Transcaucasia Armenia AD 98040 ♂ Agrodiaetus phyllis 1 1 20.07.1998 Gnyshik village Aiotzdzorsky range 1800-2200 m Transcaucasia Armenia AD 98041 ♂ Agrodiaetus phyllis 1 1 20.07.1998 Gnyshik village Aiotzdzorsky range 1800-2200 m Transcaucasia Armenia AD 98042 ♂ Agrodiaetus phyllis 1 1 20.07.1998 Gnyshik village Aiotzdzorsky range 1800-2200 m Transcaucasia Armenia AD 98043 ♂ Agrodiaetus phyllis 1 1 20.07.1998 Gnyshik village Aiotzdzorsky range 1800-2200 m Transcaucasia Armenia AD 98044 ♂ Agrodiaetus phyllis 1 1 20.07.1998 Gnyshik village Aiotzdzorsky range 1800-2200 m Transcaucasia Armenia AD 98045 ♂ Agrodiaetus phyllis 1 1 20.07.1998 Gnyshik village Aiotzdzorsky range 1800-2200 m Transcaucasia Armenia AD 98046 ♂ Agrodiaetus phyllis 1 1 20.07.1998 Gnyshik village Aiotzdzorsky range 1800-2200 m Transcaucasia Armenia AD 98047 ♂ Agrodiaetus phyllis 1 1 20.07.1998 Gnyshik village Aiotzdzorsky range 1800-2200 m Transcaucasia Armenia AD 98048 ♂ Agrodiaetus phyllis 1 1 20.07.1998 Gnyshik village Aiotzdzorsky range 1800-2200 m Transcaucasia Armenia AD 98049 ♂ Agrodiaetus phyllis 1 1 20.07.1998 Gnyshik village Aiotzdzorsky range 1800-2200 m Transcaucasia Armenia DS 00001 ♂ Agrodiaetus poseidonides 1 1 1 1 I-3 23.06.2000 Safedou Darvaz Mts. 2500 m Tajikistan DS 00002 ♂ Agrodiaetus damone 1 1 I-4 03.07.2000 Kuvandyk South Urals 2000 m Orenburg Russia DS 00003 ♂ Agrodiaetus damocles 1 1 I-5 05.07.2000 Kuvandyk South Urals Orenburg Russia DS 00004 ♂ Agrodiaetus phyllides 1 1 I-6 15.06.2000 Kitabsky national reserve Zeravshansky Mts. 1500-2500 m Uzbekistan DS 00005 ♂ Agrodiaetus dagmara 1 1 1 I-7 23.06.2000 Nikolaevsky Pass Khozratishoh Mts. 2500 m Tajikistan DS 01001 ♂ Agrodiaetus iphigenides 1 1 1 1 I-8 08.06.2001 Kitabsky national reserve Zeravshansky Mts. 1500-2500 m Uzbekistan DS 95001 ♂ Agrodiaetus actinides 1 1 I-9 24.07.1995 Transalaisky Mts. Aram-Kungei Range 3400-4400 m Kirgizia DS 99001 ♂ Agrodiaetus damone 1 1 12.07.1999 Kuray Tadzhilu riv. 1700-1800 m SE Altai Mts Russia DS 99002 ♂ Agrodiaetus transcaspicus 1 1 03.07.1999 Nohur Kopet Dag Turkmenia JC 00029 ♂ Polyommatus menelaos 1 1 1 1 1 16.06.2000 Mt. Taiyetos 1180-1200 m Peloponnisos Greece JC 00038 ♂ Polyommatus menelaos 1 1 16.06.2000 Mt. Taiyetos 1180-1200 m Peloponnisos Greece JC 00039 ♂ Polyommatus escheri 1 1 1 1 1 09.07.2000 Mt. Falakro 1650 m Macedonia Greece JC 00040 ♂ Agrodiaetus aroaniensis 1 1 1 1 1 I-10 04.07.2000 Mt. Helmos 1350 m Peloponnisos Greece JC 00041 ♂ Agrodiaetus aroaniensis 1 1 04.07.2000 Mt. Helmos 1350 m Peloponnisos Greece JC 00042 ♂ Polyommatus eroides 1 1 1 1 1 08.07.2000 Rodopi Mts. 1200 m Macedonia Greece JC 00043 ♂ Agrodiaetus ripartii 1 1 1 1 1 21.06.2000 Mt. Helmos 1350-1500 m Peloponnisos Greece JC 00044 ♂ Agrodiaetus aroaniensis 1 1 04.07.2000 Mt. Helmos 1350 m Peloponnisos Greece JC 00045 ♂ Agrodiaetus nephohiptamenos 1 1 1 1 1 I-11 07.07.2000 Mt. Orvilos 1200-2100 m Macedonia Greece JC 00046 ♂ Agrodiaetus nephohiptamenos 1 1 1 1 07.07.2000 Mt. Orvilos 1200-2100 m Macedonia Greece JC 00047 ♂ Agrodiaetus aroaniensis 1 1 04.07.2000 Mt. Helmos 1350 m Peloponnisos Greece JC 00048 ♂ Agrodiaetus nephohiptamenos 1 1 07.07.2000 Mt. Orvilos 1200-2100 m Macedonia Greece JC 00051 ♂ Polyommatus menelaos 1 1 1 1 1 09.07.2000 Mt. Falakro 1650 m Macedonia Greece JC 00053 ♂ Polyommatus escheri 1 1 09.07.2000 Mt. Falakro 1650 m Macedonia Greece JC 00054 ♂ Agrodiaetus ripartii 1 1 21.06.2000 Mt. Helmos 1350-1500 m Peloponnisos Greece JC 00055 ♂ Aricia artaxerxes 1 1 1 1 1 16.06.2000 Mt. Taiyetos 1180-1200 m Peloponnisos Greece JC 00056 ♂ Aricia artaxerxes 1 1 16.06.2000 Mt. Taiyetos 1180-1200 m Peloponnisos Greece JC 00057 ♂ Aricia artaxerxes 1 1 1 1 1 16.06.2000 Mt. Taiyetos 1180-1200 m Peloponnisos Greece JC 00060 ♂ Polyommatus menelaos 1 1 16.06.2000 Mt. Taiyetos 1180-1200 m Peloponnisos Greece JC 00061 ♂ Polyommatus andronicus 1 1 1 1 1 09.07.2000 Mt. Falakro 1650 m Macedonia Greece JC 00062 ♂ Aricia agestis 1 1 1 1 09.07.2000 Mt. Falakro 1650 m Macedonia Greece JC 00063 ♂ Polyommatus icarus 1 1 1 1 1 09.07.2000 Mt. Falakro 1650 m Macedonia Greece JC 00064 ♂ Polyommatus andronicus 1 1 07.07.2000 Mt. Orvilos 1200-2100 m Macedonia Greece JC 00067 ♂ Polyommatus icarus 1 1 09.07.2000 Mt. Falakro 1650 m Macedonia Greece JC 00070 ♂ Polyommatus andronicus 1 1 09.07.2000 Mt. Falakro 1650 m Macedonia Greece JC 00072 ♂ Polyommatus andronicus 1 1 09.07.2000 Mt. Falakro 1650 m Macedonia Greece JC 00074 ♂ Polyommatus andronicus 1 1 07.07.2000 Mt. Orvilos 1200-2100 m Macedonia Greece JC 01014 Agrodiaetus admetus 1 1 1 1 1 I-12 14.06.2001 Mt. Taiyetos 1200-1300 m Peloponnisos Greece JC 01018 Aricia agestis 1 14.06.2001 Mt. Taiyetos 1200-1300 m Peloponnisos Greece JC 01020 Agrodiaetus admetus 1 14.06.2001 Mt. Taiyetos 1200-1300 m Peloponnisos Greece JC 01027 Coenonympha arcania 1 08.07.2001 Vathirema Rhodopi Mts. 1000 m Macedonia Greece JC 01033 Coenonympha arcania 1 08.07.2001 Vathirema Rhodopi Mts. 1000 m Macedonia Greece JC 01038 Coenonympha arcania 1 08.07.2001 Vathirema Rhodopi Mts. 1000 m Macedonia Greece JC 01039 Coenonympha arcania 1 08.07.2001 Vathirema Rhodopi Mts. 1000 m Macedonia Greece JC 02001 Iphiclides podalirius 1 1 1 06.09.2002 Agia Marina Spetses Island 0 m Greece JC 02002 Pontia daplidice 1 06.09.2002 Agia Marina Spetses Island 0 m Greece JC 02003 Pontia daplidice 1 06.09.2002 Agia Marina Spetses Island 0 m Greece JC 02004 Pontia daplidice 1 06.09.2002 Agia Marina Spetses Island 0 m Greece JC 02005 Pontia daplidice 1 06.09.2002 Agia Marina Spetses Island 0 m Greece JM 00001 ♂ Agrodiaetus fabressei 1 1 1 1 1 I-13 18.07.2000 Mogorrita Tragacete Cuenca Spain JM 00002 ♂ Agrodiaetus fabressei 1 1 18.07.2000 Mogorrita Tragacete Cuenca Spain JM 00003 ♂ Agrodiaetus fabressei 1 1 18.07.2000 Mogorrita Tragacete Cuenca Spain JM 00004 ♂ Agrodiaetus fabressei 1 1 18.07.2000 Mogorrita Tragacete Cuenca Spain MW 00001 ♂ Agrodiaetus hamadanensis 1 C P 1 1 1 09.07.2000 Samqabad nordöstl. Abyek 1900-2100 m Tehran Iran MW 00002 ♂ Agrodiaetus hamadanensis 1 C 1 09.07.2000 Samqabad nordöstl. Abyek 1900-2100 m Tehran Iran MW 00003 ♂ Agrodiaetus hamadanensis 1 C 1 09.07.2000 Samqabad nordöstl. Abyek 1900-2100 m Tehran Iran MW 00004 ♂ Agrodiaetus hamadanensis 1 C P 1 09.07.2000 Samqabad nordöstl. Abyek 1900-2100 m Tehran Iran MW 00005 ♂ Agrodiaetus hamadanensis 1 C P 1 09.07.2000 Samqabad nordöstl. Abyek 1900-2100 m Tehran Iran MW 00006 ♂ Agrodiaetus hamadanensis 1 C 1 09.07.2000 Samqabad nordöstl. Abyek 1900-2100 m Tehran Iran MW 00007 ♂ Agrodiaetus hamadanensis 1 C 1 09.07.2000 Samqabad nordöstl. Abyek 1900-2100 m Tehran Iran MW 00008 ♂ Agrodiaetus hamadanensis 1 C 1 09.07.2000 Samqabad nordöstl. Abyek 1900-2100 m Tehran Iran MW 00009 ♂ Agrodiaetus demavendi 1 C P 1 09.07.2000 Samqabad nordöstl. Abyek 1900-2100 m Tehran Iran MW 00010 ♂ Agrodiaetus demavendi 1 C 1 09.07.2000 Samqabad nordöstl. Abyek 1900-2100 m Tehran Iran MW 00011 ♂ Agrodiaetus demavendi 1 C 1 09.07.2000 Samqabad nordöstl. Abyek 1900-2100 m Tehran Iran MW 00012 ♂ Agrodiaetus demavendi 1 C P 1 09.07.2000 Samqabad nordöstl. Abyek 1900-2100 m Tehran Iran MW 00013 ♂ Agrodiaetus demavendi 1 C P 1 09.07.2000 Samqabad nordöstl. Abyek 1900-2100 m Tehran Iran MW 00014 ♂ Agrodiaetus demavendi 1 C P 1 09.07.2000 Samqabad nordöstl. Abyek 1900-2100 m Tehran Iran

- 149 - Appendix 2 List of material

Code Sex Genus Species Wgs Fix Kar Alc DNA Cytb CO1 ND1 ITS2 Fig. Date Locality Area Altitude Province Country MW 00015 ♂ Agrodiaetus demavendi 1 C P 1 1 1 1 I-14 09.07.2000 Samqabad nordöstl. Abyek 1900-2100 m Tehran Iran MW 00016 ♂ Agrodiaetus demavendi 1 C P 1 09.07.2000 Samqabad nordöstl. Abyek 1900-2100 m Tehran Iran MW 00017 ♂ Lycaena alciphron 1 1 1 1 09.07.2000 Samqabad nordöstl. Abyek 1900-2100 m Tehran Iran MW 00018 ♂ Polyommatus icarus 1 1 09.07.2000 Samqabad nordöstl. Abyek 1900-2100 m Tehran Iran MW 00019 ♂ Polyommatus icarus 1 1 09.07.2000 Samqabad nordöstl. Abyek 1900-2100 m Tehran Iran MW 00020 ♂ Aricia agestis 1 1 1 1 09.07.2000 Samqabad nordöstl. Abyek 1900-2100 m Tehran Iran MW 00021 Lycaena phlaeas 1 1 09.07.2000 Samqabad nordöstl. Abyek 1900-2100 m Tehran Iran MW 00022 ♀ Agrodiaetus hamadanensis 1 1 09.07.2000 Samqabad nordöstl. Abyek 1900-2100 m Tehran Iran MW 00023 ♂ Vacciniina alcedo 1 1 09.07.2000 Samqabad nordöstl. Abyek 1900-2100 m Tehran Iran MW 00024 ♂ Vacciniina alcedo 1 1 1 1 1 09.07.2000 Samqabad nordöstl. Abyek 1900-2100 m Tehran Iran MW 00025 ♂ Vacciniina alcedo 1 1 09.07.2000 Samqabad nordöstl. Abyek 1900-2100 m Tehran Iran MW 00026 ♂ Agrodiaetus hamadanensis 1 1 09.07.2000 Samqabad nordöstl. Abyek 1900-2100 m Tehran Iran MW 00027 ♂ Agrodiaetus hamadanensis 1 1 09.07.2000 Samqabad nordöstl. Abyek 1900-2100 m Tehran Iran MW 00028 ♀ Agrodiaetus hamadanensis 1 1 09.07.2000 Samqabad nordöstl. Abyek 1900-2100 m Tehran Iran MW 00029 ♂ Agrodiaetus hamadanensis 1 1 09.07.2000 Samqabad nordöstl. Abyek 1900-2100 m Tehran Iran MW 00030 ♂ Agrodiaetus demavendi 1 C P 1 09.07.2000 Samqabad nordöstl. Abyek 1900-2100 m Tehran Iran MW 00031 ♂ Agrodiaetus hamadanensis 1 C P 1 10.07.2000 Safedabad westl. Sirud / Karaj 2000 m Tehran Iran MW 00032 ♂ Agrodiaetus hamadanensis 1 C P 1 1 1 1 1 I-15 10.07.2000 Safedabad westl. Sirud / Karaj 2000 m Tehran Iran MW 00033 ♂ Agrodiaetus hamadanensis 1 C P 1 10.07.2000 Safedabad westl. Sirud / Karaj 2000 m Tehran Iran MW 00034 ♀ Agrodiaetus hamadanensis 1 1 10.07.2000 Safedabad westl. Sirud / Karaj 2000 m Tehran Iran MW 00035 ♀ Agrodiaetus hamadanensis 1 1 10.07.2000 Safedabad westl. Sirud / Karaj 2000 m Tehran Iran MW 00036 ♀ Agrodiaetus hamadanensis 1 1 10.07.2000 Safedabad westl. Sirud / Karaj 2000 m Tehran Iran MW 00037 ♀ Agrodiaetus hamadanensis 1 1 10.07.2000 Safedabad westl. Sirud / Karaj 2000 m Tehran Iran MW 00038 ♂ Polyommatus icarus 1 1 10.07.2000 Safedabad westl. Sirud / Karaj 2000 m Tehran Iran MW 00039 ♂ Polyommatus icarus 1 1 10.07.2000 Safedabad westl. Sirud / Karaj 2000 m Tehran Iran MW 00040 ♂ Polyommatus icarus 1 1 10.07.2000 Safedabad westl. Sirud / Karaj 2000 m Tehran Iran MW 00041 ♀ Polyommatus icarus 1 1 10.07.2000 Safedabad westl. Sirud / Karaj 2000 m Tehran Iran MW 00042 ♀ Polyommatus icarus 1 1 10.07.2000 Safedabad westl. Sirud / Karaj 2000 m Tehran Iran MW 00043 ♀ Polyommatus icarus 1 1 10.07.2000 Safedabad westl. Sirud / Karaj 2000 m Tehran Iran MW 00044 ♂ Lycaena asabinus 1 1 1 1 10.07.2000 Safedabad westl. Sirud / Karaj 2000 m Tehran Iran MW 00045 ♂ Lycaena alciphron 1 1 10.07.2000 Safedabad westl. Sirud / Karaj 2000 m Tehran Iran MW 00046 ♂ Lycaena phlaeas 1 1 10.07.2000 Safedabad westl. Sirud / Karaj 2000 m Tehran Iran MW 00047 ♂ Lycaena tityrus 1 1 10.07.2000 Safedabad westl. Sirud / Karaj 2000 m Tehran Iran MW 00048 ♂ Agrodiaetus elbursicus 1 C P 1 11.07.2000 Nesa 1 km N Nesa 2100 m Tehran Iran MW 00049 ♂ Agrodiaetus elbursicus 1 C P 1 11.07.2000 Nesa 1 km N Nesa 2100 m Tehran Iran MW 00050 ♂ Agrodiaetus elbursicus 1 C P 1 11.07.2000 Nesa 1 km N Nesa 2100 m Tehran Iran MW 00051 ♂ Agrodiaetus elbursicus 1 C P 1 1 1 1 11.07.2000 Nesa 1 km N Nesa 2100 m Tehran Iran MW 00052 ♂ Agrodiaetus elbursicus 1 C P 1 11.07.2000 Nesa 1 km N Nesa 2100 m Tehran Iran MW 00053 ♂ Agrodiaetus elbursicus 1 C 1 11.07.2000 Nesa 1 km N Nesa 2100 m Tehran Iran MW 00054 ♂ Agrodiaetus elbursicus 1 C 1 11.07.2000 Nesa 1 km N Nesa 2100 m Tehran Iran MW 00055 ♂ Agrodiaetus elbursicus 1 C P 1 11.07.2000 Nesa 1 km N Nesa 2100 m Tehran Iran MW 00056 ♂ Agrodiaetus elbursicus 1 C P 1 1 1 1 11.07.2000 Kendevan 5 km N Kendevan 2150 m Mazandaran Iran MW 00057 ♂ Agrodiaetus elbursicus 1 C P 1 11.07.2000 Kendevan 5 km N Kendevan 2150 m Mazandaran Iran MW 00058 ♂ Agrodiaetus elbursicus 1 C P 1 1 1 11.07.2000 Pul-e Zanguleh 15 km NO Kendevan 2400 m Mazandaran Iran MW 00059 ♂ Agrodiaetus elbursicus 1 C P 1 1 1 1 I-16 11.07.2000 Pul-e Zanguleh 15 km NO Kendevan 2400 m Mazandaran Iran MW 00060 ♂ Agrodiaetus birunii 1 C P 1 1 1 11.07.2000 Pul-e Zanguleh 15 km NO Kendevan 2400 m Mazandaran Iran MW 00061 ♂ Agrodiaetus birunii 1 C P 1 11.07.2000 Pul-e Zanguleh 15 km NO Kendevan 2400 m Mazandaran Iran MW 00062 ♂ Agrodiaetus birunii 1 C P 1 11.07.2000 Pul-e Zanguleh 15 km NO Kendevan 2400 m Mazandaran Iran MW 00063 ♂ Agrodiaetus birunii 1 C 1 11.07.2000 Pul-e Zanguleh 15 km NO Kendevan 2400 m Mazandaran Iran MW 00064 ♂ Agrodiaetus valiabadi 1 C P 1 1 1 1 11.07.2000 Pul-e Zanguleh 15 km NO Kendevan 2400 m Mazandaran Iran MW 00065 ♂ Agrodiaetus elbursicus 1 C P 1 11.07.2000 Gardaneh-ye Lavashm 20 km NO Kendevan 2900 m Mazandaran Iran MW 00066 ♂ Agrodiaetus elbursicus 1 C P 1 11.07.2000 Gardaneh-ye Lavashm 20 km NO Kendevan 2900 m Mazandaran Iran MW 00067 ♂ Agrodiaetus elbursicus 1 C 1 11.07.2000 Gardaneh-ye Lavashm 20 km NO Kendevan 2900 m Mazandaran Iran MW 00068 ♂ Agrodiaetus birunii 1 C P 1 11.07.2000 Gardaneh-ye Lavashm 20 km NO Kendevan 2900 m Mazandaran Iran MW 00069 ♂ Agrodiaetus birunii 1 C P 1 11.07.2000 Gardaneh-ye Lavashm 20 km NO Kendevan 2900 m Mazandaran Iran MW 00070 ♂ Agrodiaetus birunii 1 C P 1 11.07.2000 Gardaneh-ye Lavashm 20 km NO Kendevan 2900 m Mazandaran Iran MW 00071 ♂ Agrodiaetus birunii 1 C P 1 11.07.2000 Gardaneh-ye Lavashm 20 km NO Kendevan 2900 m Mazandaran Iran MW 00072 ♂ Agrodiaetus birunii 1 C P 1 1 1 1 1 11.07.2000 Gardaneh-ye Lavashm 20 km NO Kendevan 2900 m Mazandaran Iran MW 00073 ♂ Agrodiaetus birunii 1 C P 1 11.07.2000 Gardaneh-ye Lavashm 20 km NO Kendevan 2900 m Mazandaran Iran MW 00074 ♀ Agrodiaetus birunii 1 1 11.07.2000 Gardaneh-ye Lavashm 20 km NO Kendevan 2900 m Mazandaran Iran MW 00075 ♂ Polyommatus icarus 1 1 11.07.2000 Nesa 1 km N Nesa 2100 m Tehran Iran MW 00076 ♂ Meleageria daphnis 1 1 1 1 1 1 11.07.2000 Nesa 1 km N Nesa 2100 m Tehran Iran MW 00077 ♂ Meleageria daphnis 1 1 11.07.2000 Nesa 1 km N Nesa 2100 m Tehran Iran MW 00078 ♂ Meleageria daphnis 1 1 11.07.2000 Nesa 1 km N Nesa 2100 m Tehran Iran MW 00079 ♀ Meleageria daphnis 1 1 11.07.2000 Nesa 1 km N Nesa 2100 m Tehran Iran MW 00080 ♀ Meleageria daphnis 1 1 11.07.2000 Nesa 1 km N Nesa 2100 m Tehran Iran MW 00081 ♀ Meleageria daphnis 1 1 11.07.2000 Nesa 1 km N Nesa 2100 m Tehran Iran MW 00082 ♂ Polyommatus amandus 1 1 11.07.2000 Kendevan 5 km N Kendevan 2150 m Mazandaran Iran MW 00083 ♂ Polyommatus amandus 1 1 11.07.2000 Kendevan 5 km N Kendevan 2150 m Mazandaran Iran MW 00084 ♂ Plebeius pylaon 1 1 11.07.2000 Valiabad nördl. Valiabad 1600 m Mazandaran Iran MW 00085 ♀ Lycaena alciphron 1 1 11.07.2000 Pul-e Zanguleh 15 km NO Kendevan 2400 m Mazandaran Iran MW 00086 ♂ Polyommatus icarus 1 1 11.07.2000 Pul-e Zanguleh 15 km NO Kendevan 2400 m Mazandaran Iran MW 00087 ♂ Meleageria marcida 1 1 1 1 11.07.2000 Pul-e Zanguleh 15 km NO Kendevan 2400 m Mazandaran Iran MW 00088 ♂ Meleageria marcida 1 1 11.07.2000 Pul-e Zanguleh 15 km NO Kendevan 2400 m Mazandaran Iran MW 00089 ♂ Meleageria marcida 1 1 11.07.2000 Pul-e Zanguleh 15 km NO Kendevan 2400 m Mazandaran Iran MW 00090 ♂ Meleageria marcida 1 1 11.07.2000 Pul-e Zanguleh 15 km NO Kendevan 2400 m Mazandaran Iran MW 00091 ♂ Meleageria marcida 1 1 11.07.2000 Pul-e Zanguleh 15 km NO Kendevan 2400 m Mazandaran Iran MW 00092 ♂ Agrodiaetus elbursicus 1 1 11.07.2000 Nesa 1 km N Nesa 2100 m Tehran Iran MW 00093 ♂ Agrodiaetus elbursicus 1 1 11.07.2000 Nesa 1 km N Nesa 2100 m Tehran Iran MW 00094 ♀ Agrodiaetus elbursicus 1 1 11.07.2000 Nesa 1 km N Nesa 2100 m Tehran Iran MW 00095 ♀ Agrodiaetus elbursicus 1 1 11.07.2000 Nesa 1 km N Nesa 2100 m Tehran Iran MW 00096 ♂ Agrodiaetus birunii 1 1 11.07.2000 Pul-e Zanguleh 15 km NO Kendevan 2400 m Mazandaran Iran MW 00097 ♂ Agrodiaetus birunii 1 1 11.07.2000 Pul-e Zanguleh 15 km NO Kendevan 2400 m Mazandaran Iran MW 00098 ♂ Agrodiaetus birunii 1 1 11.07.2000 Pul-e Zanguleh 15 km NO Kendevan 2400 m Mazandaran Iran MW 00099 ♂ Agrodiaetus phyllis 1 1 11.07.2000 Pul-e Zanguleh 15 km NO Kendevan 2400 m Mazandaran Iran MW 00100 ♂ Agrodiaetus birunii 1 1 11.07.2000 Pul-e Zanguleh 15 km NO Kendevan 2400 m Mazandaran Iran MW 00101 ♂ Agrodiaetus darius 1 C P 1 1 1 1 I-17 12.07.2000 Dizin Pass südl. Dizin 3000 m Tehran Iran MW 00102 ♂ Agrodiaetus birunii 1 C P 1 1 1 1 12.07.2000 Dizin Pass südl. Dizin 3000 m Tehran Iran MW 00103 ♂ Agrodiaetus birunii 1 C P 1 12.07.2000 Dizin Pass südl. Dizin 3000 m Tehran Iran MW 00104 ♂ Agrodiaetus birunii 1 C 1 12.07.2000 Dizin Pass südl. Dizin 3000 m Tehran Iran MW 00105 ♂ Agrodiaetus elbursicus 1 C P 1 12.07.2000 Shahrestanak nördl. Tehran 2000 m Tehran Iran MW 00106 ♂ Agrodiaetus elbursicus 1 C 1 12.07.2000 Shahrestanak nördl. Tehran 2000 m Tehran Iran MW 00107 ♂ Agrodiaetus elbursicus 1 C P 1 12.07.2000 Shahrestanak nördl. Tehran 2000 m Tehran Iran MW 00108 ♂ Agrodiaetus elbursicus 1 C 1 12.07.2000 Shahrestanak nördl. Tehran 2000 m Tehran Iran MW 00109 ♂ Agrodiaetus elbursicus 1 C 1 12.07.2000 Shahrestanak nördl. Tehran 2000 m Tehran Iran MW 00110 ♂ Agrodiaetus elbursicus 1 C P 1 1 1 1 12.07.2000 Shahrestanak nördl. Tehran 2000 m Tehran Iran MW 00111 ♂ Agrodiaetus elbursicus 1 C P 1 12.07.2000 Shahrestanak nördl. Tehran 2000 m Tehran Iran MW 00112 ♂ Agrodiaetus elbursicus 1 C P 1 12.07.2000 Gajereh 2200 m Tehran Iran MW 00113 ♂ Agrodiaetus elbursicus 1 C 1 12.07.2000 Gajereh 2200 m Tehran Iran MW 00114 ♂ Agrodiaetus elbursicus 1 C 1 12.07.2000 Gajereh 2200 m Tehran Iran MW 00115 ♂ Agrodiaetus birunii 1 C P 1 12.07.2000 Shemshak südl. Dizin Pass 2900 m Tehran Iran MW 00116 ♂ Plebeius argus 1 1 1 1 1 12.07.2000 Shemshak südl. Dizin Pass 2900 m Tehran Iran MW 00117 ♂ Neolysandra corona 1 1 12.07.2000 Shemshak südl. Dizin Pass 2900 m Tehran Iran MW 00118 ♂ Neolysandra corona 1 1 12.07.2000 Shemshak südl. Dizin Pass 2900 m Tehran Iran MW 00119 ♂ Lycaena candens 1 1 1 1 12.07.2000 Shemshak südl. Dizin Pass 2900 m Tehran Iran MW 00120 ♀ Lycaena candens 1 1 12.07.2000 Shemshak südl. Dizin Pass 2900 m Tehran Iran MW 00121 ♂ Lycaena tityrus 1 1 12.07.2000 Shemshak südl. Dizin Pass 2900 m Tehran Iran MW 00122 ♂ Vacciniina morgianus 1 1 12.07.2000 Dizin Pass südl. Dizin 3000 m Tehran Iran MW 00123 ♂ Neolysandra corona 1 1 12.07.2000 Dizin Pass südl. Dizin 3000 m Tehran Iran MW 00124 ♂ Neolysandra corona 1 1 12.07.2000 Dizin Pass südl. Dizin 3000 m Tehran Iran MW 00125 ♂ Agrodiaetus firdussii 1 C P 1 1 1 13.07.2000 Ahar Pass 20 km SW Ahar 1800-1850 m Azarbayjan-e Sharqi Iran MW 00126 ♂ Agrodiaetus paulae 1 C P 1 13.07.2000 Ahar Pass 20 km SW Ahar 1800-1850 m Azarbayjan-e Sharqi Iran MW 00127 ♂ Agrodiaetus paulae 1 C P 1 1 1 1 I-18 13.07.2000 Ahar Pass 20 km SW Ahar 1800-1850 m Azarbayjan-e Sharqi Iran MW 00128 ♂ Agrodiaetus paulae 1 C P 1 13.07.2000 Ahar Pass 20 km SW Ahar 1800-1850 m Azarbayjan-e Sharqi Iran MW 00129 ♂ Agrodiaetus gorbunovi 1 C P 1 1 1 1 I-19 13.07.2000 Ahar Pass 20 km SW Ahar 1800-1850 m Azarbayjan-e Sharqi Iran

- 150 - Appendix 2 List of material

Code Sex Genus Species Wgs Fix Kar Alc DNA Cytb CO1 ND1 ITS2 Fig. Date Locality Area Altitude Province Country MW 00130 ♂ Agrodiaetus phyllis 1 C P 1 13.07.2000 Ahar Pass 20 km SW Ahar 1800-1850 m Azarbayjan-e Sharqi Iran MW 00131 ♂ Agrodiaetus phyllis 1 C 1 13.07.2000 Ahar Pass 20 km SW Ahar 1800-1850 m Azarbayjan-e Sharqi Iran MW 00132 ♂ Agrodiaetus paulae 1 C P 1 13.07.2000 Ahar Pass 20 km SW Ahar 1800-1850 m Azarbayjan-e Sharqi Iran MW 00133 ♂ Agrodiaetus phyllis 1 C 1 13.07.2000 Ahar Pass 20 km SW Ahar 1800-1850 m Azarbayjan-e Sharqi Iran MW 00134 ♂ Agrodiaetus phyllis 1 C 1 13.07.2000 Ahar Pass 20 km SW Ahar 1800-1850 m Azarbayjan-e Sharqi Iran MW 00135 ♂ Agrodiaetus phyllis 1 C P 1 13.07.2000 Ahar Pass 20 km SW Ahar 1800-1850 m Azarbayjan-e Sharqi Iran MW 00136 ♂ Agrodiaetus phyllis 1 C 1 13.07.2000 Ahar Pass 20 km SW Ahar 1800-1850 m Azarbayjan-e Sharqi Iran MW 00137 ♂ Agrodiaetus phyllis 1 C 1 13.07.2000 Ahar Pass 20 km SW Ahar 1800-1850 m Azarbayjan-e Sharqi Iran MW 00138 ♂ Agrodiaetus phyllis 1 C 1 13.07.2000 Ahar Pass 20 km SW Ahar 1800-1850 m Azarbayjan-e Sharqi Iran MW 00139 ♂ Agrodiaetus phyllis 1 C 1 13.07.2000 Ahar Pass 20 km SW Ahar 1800-1850 m Azarbayjan-e Sharqi Iran MW 00140 ♂ Agrodiaetus phyllis 1 C 1 1 1 13.07.2000 Ahar Pass 20 km SW Ahar 1800-1850 m Azarbayjan-e Sharqi Iran MW 00141 ♂ Agrodiaetus phyllis 1 1 13.07.2000 Ahar Pass 20 km SW Ahar 1800-1850 m Azarbayjan-e Sharqi Iran MW 00142 ♀ Agrodiaetus firdussii 1 1 13.07.2000 Ahar Pass 20 km SW Ahar 1800-1850 m Azarbayjan-e Sharqi Iran MW 00143 ♀ Agrodiaetus firdussii 1 1 13.07.2000 Ahar Pass 20 km SW Ahar 1800-1850 m Azarbayjan-e Sharqi Iran MW 00144 ♀ Agrodiaetus firdussii 1 1 13.07.2000 Ahar Pass 20 km SW Ahar 1800-1850 m Azarbayjan-e Sharqi Iran MW 00145 ♀ Agrodiaetus firdussii 1 1 13.07.2000 Ahar Pass 20 km SW Ahar 1800-1850 m Azarbayjan-e Sharqi Iran MW 00146 ♀ Agrodiaetus alcestis 1 1 13.07.2000 Ahar Pass 20 km SW Ahar 1800-1850 m Azarbayjan-e Sharqi Iran MW 00147 ♀ Agrodiaetus 1 1 13.07.2000 Ahar Pass 20 km SW Ahar 1800-1850 m Azarbayjan-e Sharqi Iran MW 00148 ♀ Agrodiaetus phyllis 1 1 13.07.2000 Ahar Pass 20 km SW Ahar 1800-1850 m Azarbayjan-e Sharqi Iran MW 00149 ♂ Polyommatus icarus 1 1 13.07.2000 Ahar Pass 20 km SW Ahar 1800-1850 m Azarbayjan-e Sharqi Iran MW 00150 ♂ Polyommatus icarus 1 1 13.07.2000 Ahar Pass 20 km SW Ahar 1800-1850 m Azarbayjan-e Sharqi Iran MW 00151 ♂ Agrodiaetus firdussii 1 C P 1 1 1 1 13.07.2000 Ahar Pass 20 km SW Ahar 1800-1850 m Azarbayjan-e Sharqi Iran MW 00152 ♂ Agrodiaetus firdussii 1 C 1 13.07.2000 Ahar Pass 20 km SW Ahar 1800-1850 m Azarbayjan-e Sharqi Iran MW 00153 ♂ Agrodiaetus firdussii 1 C 1 13.07.2000 Ahar Pass 20 km SW Ahar 1800-1850 m Azarbayjan-e Sharqi Iran MW 00154 ♂ Agrodiaetus firdussii 1 C 1 13.07.2000 Ahar Pass 20 km SW Ahar 1800-1850 m Azarbayjan-e Sharqi Iran MW 00155 ♂ Agrodiaetus paulae 1 C P 1 13.07.2000 Ahar Pass 20 km SW Ahar 1800-1850 m Azarbayjan-e Sharqi Iran MW 00156 ♂ Agrodiaetus firdussii 1 C 1 13.07.2000 Ahar Pass 20 km SW Ahar 1800-1850 m Azarbayjan-e Sharqi Iran MW 00157 ♂ Agrodiaetus paulae 1 C P 1 1 1 13.07.2000 Ahar Pass 20 km SW Ahar 1800-1850 m Azarbayjan-e Sharqi Iran MW 00158 ♂ Agrodiaetus paulae 1 C P 1 13.07.2000 Ahar Pass 20 km SW Ahar 1800-1850 m Azarbayjan-e Sharqi Iran MW 00159 ♂ Agrodiaetus paulae 1 C P 1 13.07.2000 Ahar Pass 20 km SW Ahar 1800-1850 m Azarbayjan-e Sharqi Iran MW 00160 ♂ Agrodiaetus paulae 1 C P 1 13.07.2000 Ahar Pass 20 km SW Ahar 1800-1850 m Azarbayjan-e Sharqi Iran MW 00161 ♂ Agrodiaetus paulae 1 C P 1 13.07.2000 Ahar Pass 20 km SW Ahar 1800-1850 m Azarbayjan-e Sharqi Iran MW 00162 ♂ Agrodiaetus paulae 1 C P 1 13.07.2000 Ahar Pass 20 km SW Ahar 1800-1850 m Azarbayjan-e Sharqi Iran MW 00163 ♂ Agrodiaetus phyllis 1 C 1 13.07.2000 Ahar Pass 20 km SW Ahar 1800-1850 m Azarbayjan-e Sharqi Iran MW 00164 ♂ Agrodiaetus phyllis 1 C 1 13.07.2000 Ahar Pass 20 km SW Ahar 1800-1850 m Azarbayjan-e Sharqi Iran MW 00165 ♂ Agrodiaetus phyllis 1 C 1 13.07.2000 Ahar Pass 20 km SW Ahar 1800-1850 m Azarbayjan-e Sharqi Iran MW 00166 ♂ Agrodiaetus phyllis 1 C 1 13.07.2000 Ahar Pass 20 km SW Ahar 1800-1850 m Azarbayjan-e Sharqi Iran MW 00167 ♂ Agrodiaetus phyllis 1 C P 1 13.07.2000 Ahar Pass 20 km SW Ahar 1800-1850 m Azarbayjan-e Sharqi Iran MW 00168 ♂ Agrodiaetus phyllis 1 C P 1 13.07.2000 Ahar Pass 20 km SW Ahar 1800-1850 m Azarbayjan-e Sharqi Iran MW 00169 ♂ Agrodiaetus phyllis 1 C P 1 13.07.2000 Ahar Pass 20 km SW Ahar 1800-1850 m Azarbayjan-e Sharqi Iran MW 00170 ♂ Agrodiaetus alcestis 1 C P 1 13.07.2000 Ahar Pass 20 km SW Ahar 1800-1850 m Azarbayjan-e Sharqi Iran MW 00171 ♂ Plebeius argus 1 C 1 13.07.2000 Ahar Pass 20 km SW Ahar 1800-1850 m Azarbayjan-e Sharqi Iran MW 00172 ♂ Plebeius argus 1 1 13.07.2000 Ahar Pass 20 km SW Ahar 1800-1850 m Azarbayjan-e Sharqi Iran MW 00173 ♂ Polyommatus thersites 1 1 13.07.2000 Ahar Pass 20 km SW Ahar 1800-1850 m Azarbayjan-e Sharqi Iran MW 00174 ♀ Satyrium abdominalis 1 1 13.07.2000 Ahar Pass 20 km SW Ahar 1800-1850 m Azarbayjan-e Sharqi Iran MW 00175 ♂ Satyrium abdominalis 1 1 13.07.2000 Ahar Pass 20 km SW Ahar 1800-1850 m Azarbayjan-e Sharqi Iran MW 00176 ♂ Agrodiaetus rovshani 1 C P 1 1 1 1 I-20 15.07.2000 Dugijan 30 km NO Marand 2000 m Azarbayjan-e Sharqi Iran MW 00177 ♂ Agrodiaetus gorbunovi 1 C P 1 1 1 1 15.07.2000 Dugijan 30 km NO Marand 2000 m Azarbayjan-e Sharqi Iran MW 00178 ♂ Agrodiaetus gorbunovi 1 C P 1 1 1 1 15.07.2000 Dugijan 30 km NO Marand 2000 m Azarbayjan-e Sharqi Iran MW 00179 ♂ Agrodiaetus cyaneus 1 C P 1 1 1 1 I-21 15.07.2000 Dugijan 30 km NO Marand 2000 m Azarbayjan-e Sharqi Iran MW 00180 ♂ Agrodiaetus phyllis 1 C 1 15.07.2000 Dugijan 30 km NO Marand 2000 m Azarbayjan-e Sharqi Iran MW 00181 ♂ Agrodiaetus phyllis 1 C 1 15.07.2000 Dugijan 30 km NO Marand 2000 m Azarbayjan-e Sharqi Iran MW 00182 ♂ Agrodiaetus demavendi 1 C P 1 15.07.2000 Dugijan 30 km NO Marand 2000 m Azarbayjan-e Sharqi Iran MW 00183 ♂ Agrodiaetus demavendi 1 C P 1 1 1 15.07.2000 Dugijan 30 km NO Marand 2000 m Azarbayjan-e Sharqi Iran MW 00184 ♂ Agrodiaetus demavendi 1 C P 1 15.07.2000 Dugijan 30 km NO Marand 2000 m Azarbayjan-e Sharqi Iran MW 00185 ♂ Agrodiaetus demavendi 1 C P 1 1 1 1 15.07.2000 Dugijan 30 km NO Marand 2000 m Azarbayjan-e Sharqi Iran MW 00186 ♂ Agrodiaetus demavendi 1 C P 1 1 1 15.07.2000 Dugijan 30 km NO Marand 2000 m Azarbayjan-e Sharqi Iran MW 00187 ♂ Agrodiaetus demavendi 1 C P 1 15.07.2000 Dugijan 30 km NO Marand 2000 m Azarbayjan-e Sharqi Iran MW 00188 ♂ Agrodiaetus demavendi 1 C P 1 15.07.2000 Dugijan 30 km NO Marand 2000 m Azarbayjan-e Sharqi Iran MW 00189 ♂ Agrodiaetus demavendi 1 C P 1 1 1 1 1 15.07.2000 Dugijan 30 km NO Marand 2000 m Azarbayjan-e Sharqi Iran MW 00190 ♂ Agrodiaetus demavendi 1 C P 1 15.07.2000 Dugijan 30 km NO Marand 2000 m Azarbayjan-e Sharqi Iran MW 00191 ♂ Lysandra bellargus 1 C 1 15.07.2000 Dugijan 30 km NO Marand 2000 m Azarbayjan-e Sharqi Iran MW 00192 ♂ Polyommatus icarus 1 C 1 15.07.2000 Dugijan 30 km NO Marand 2000 m Azarbayjan-e Sharqi Iran MW 00193 ♂ Polyommatus thersites 1 1 15.07.2000 Dugijan 30 km NO Marand 2000 m Azarbayjan-e Sharqi Iran MW 00194 ♂ Polyommatus icarus 1 1 15.07.2000 Dugijan 30 km NO Marand 2000 m Azarbayjan-e Sharqi Iran MW 00195 ♂ Polyommatus icarus 1 1 15.07.2000 Dugijan 30 km NO Marand 2000 m Azarbayjan-e Sharqi Iran MW 00196 ♂ Meleageria daphnis 1 1 15.07.2000 Dugijan 30 km NO Marand 2000 m Azarbayjan-e Sharqi Iran MW 00197 ♂ Meleageria daphnis 1 1 15.07.2000 Dugijan 30 km NO Marand 2000 m Azarbayjan-e Sharqi Iran MW 00198 ♀ Agrodiaetus demavendi 1 1 15.07.2000 Dugijan 30 km NO Marand 2000 m Azarbayjan-e Sharqi Iran MW 00199 ♀ Agrodiaetus demavendi 1 1 15.07.2000 Dugijan 30 km NO Marand 2000 m Azarbayjan-e Sharqi Iran MW 00200 ♂ Agrodiaetus phyllis 1 C P 1 15.07.2000 Dugijan 30 km NO Marand 3000 m Azarbayjan-e Sharqi Iran MW 00201 ♂ Agrodiaetus phyllis 1 C 1 15.07.2000 Dugijan 30 km NO Marand 3000 m Azarbayjan-e Sharqi Iran MW 00202 ♂ Agrodiaetus phyllis 1 C 1 15.07.2000 Dugijan 30 km NO Marand 3000 m Azarbayjan-e Sharqi Iran MW 00203 ♂ Agrodiaetus phyllis 1 C P 1 15.07.2000 Dugijan 30 km NO Marand 3000 m Azarbayjan-e Sharqi Iran MW 00204 ♂ Agrodiaetus phyllis 1 C P 1 15.07.2000 Dugijan 30 km NO Marand 3000 m Azarbayjan-e Sharqi Iran MW 00205 ♀ Agrodiaetus phyllis 1 C 1 15.07.2000 Dugijan 30 km NO Marand 3000 m Azarbayjan-e Sharqi Iran MW 00206 ♂ Polyommatus icarus 1 C 1 15.07.2000 Dugijan 30 km NO Marand 3000 m Azarbayjan-e Sharqi Iran MW 00207 ♂ Polyommatus icarus 1 C P 1 15.07.2000 Dugijan 30 km NO Marand 3000 m Azarbayjan-e Sharqi Iran MW 00208 ♂ Agrodiaetus phyllis 1 1 15.07.2000 Dugijan 30 km NO Marand 3000 m Azarbayjan-e Sharqi Iran MW 00209 ♀ Agrodiaetus phyllis 1 1 15.07.2000 Dugijan 30 km NO Marand 3000 m Azarbayjan-e Sharqi Iran MW 00210 ♀ Agrodiaetus phyllis 1 1 15.07.2000 Dugijan 30 km NO Marand 3000 m Azarbayjan-e Sharqi Iran MW 00211 ♀ Agrodiaetus phyllis 1 1 15.07.2000 Dugijan 30 km NO Marand 3000 m Azarbayjan-e Sharqi Iran MW 00212 ♀ Agrodiaetus phyllis 1 1 15.07.2000 Dugijan 30 km NO Marand 3000 m Azarbayjan-e Sharqi Iran MW 00213 ♂ Lycaena thersamon 1 1 15.07.2000 Dugijan 30 km NO Marand 3000 m Azarbayjan-e Sharqi Iran MW 00214 ♀ Lycaena thersamon 1 1 15.07.2000 Dugijan 30 km NO Marand 3000 m Azarbayjan-e Sharqi Iran MW 00215 ♂ Agrodiaetus firdussii 1 C P 1 16.07.2000 Qazayd Dagh 25 km O Zanjan 2300 m Zanjan Iran MW 00216 ♂ Agrodiaetus firdussii 1 C P 1 16.07.2000 Qazayd Dagh 25 km O Zanjan 2300 m Zanjan Iran MW 00217 ♂ Agrodiaetus firdussii 1 C P 1 16.07.2000 Qazayd Dagh 25 km O Zanjan 2300 m Zanjan Iran MW 00218 ♂ Agrodiaetus firdussii 1 C P 1 16.07.2000 Qazayd Dagh 25 km O Zanjan 2300 m Zanjan Iran MW 00219 ♂ Agrodiaetus firdussii 1 C P 1 16.07.2000 Qazayd Dagh 25 km O Zanjan 2300 m Zanjan Iran MW 00220 ♂ Agrodiaetus phyllis 1 C 1 16.07.2000 Qazayd Dagh 25 km O Zanjan 2300 m Zanjan Iran MW 00221 ♂ Agrodiaetus phyllis 1 C 1 16.07.2000 Qazayd Dagh 25 km O Zanjan 2300 m Zanjan Iran MW 00222 ♂ Agrodiaetus phyllis 1 C 1 16.07.2000 Qazayd Dagh 25 km O Zanjan 2300 m Zanjan Iran MW 00223 ♂ Meleageria daphnis 1 1 16.07.2000 Qazayd Dagh 25 km O Zanjan 2300 m Zanjan Iran MW 00224 ♂ Meleageria daphnis 1 1 16.07.2000 Qazayd Dagh 25 km O Zanjan 2300 m Zanjan Iran MW 00225 ♀ Meleageria daphnis 1 1 16.07.2000 Qazayd Dagh 25 km O Zanjan 2300 m Zanjan Iran MW 00226 ♂ Agrodiaetus femininoides 1 C P 1 1 1 1 I-22 16.07.2000 Qazayd Dagh 25 km O Zanjan 2300 m Zanjan Iran MW 00227 ♂ Agrodiaetus femininoides 1 C P 1 16.07.2000 Qazayd Dagh 25 km O Zanjan 2300 m Zanjan Iran MW 00228 ♂ Agrodiaetus femininoides 1 C P 1 16.07.2000 Qazayd Dagh 25 km O Zanjan 2300 m Zanjan Iran MW 00229 ♂ Agrodiaetus alcestis 1 C P 1 1 1 16.07.2000 Qazayd Dagh 25 km O Zanjan 2300 m Zanjan Iran MW 00230 ♂ Agrodiaetus alcestis 1 C P 1 16.07.2000 Qazayd Dagh 25 km O Zanjan 2300 m Zanjan Iran MW 00231 ♂ Agrodiaetus alcestis 1 C P 1 1 1 1 1 I-23 16.07.2000 Qazayd Dagh 25 km O Zanjan 2300 m Zanjan Iran MW 00232 ♂ Agrodiaetus elbursicus 1 C P 1 1 1 1 16.07.2000 Qazayd Dagh 25 km O Zanjan 2300 m Zanjan Iran MW 00233 ♂ Agrodiaetus firdussii 1 C 1 16.07.2000 Qazayd Dagh 25 km O Zanjan 2300 m Zanjan Iran MW 00234 ♂ Agrodiaetus firdussii 1 C P 1 1 1 1 I-24 16.07.2000 Qazayd Dagh 25 km O Zanjan 2300 m Zanjan Iran MW 00235 ♂ Agrodiaetus firdussii 1 C P 1 16.07.2000 Qazayd Dagh 25 km O Zanjan 2300 m Zanjan Iran MW 00236 ♂ Agrodiaetus phyllis 1 C P 1 16.07.2000 Qazayd Dagh 25 km O Zanjan 2300 m Zanjan Iran MW 00237 ♂ Agrodiaetus phyllis 1 C 1 16.07.2000 Qazayd Dagh 25 km O Zanjan 2300 m Zanjan Iran MW 00238 ♂ Agrodiaetus phyllis 1 C 1 16.07.2000 Qazayd Dagh 25 km O Zanjan 2300 m Zanjan Iran MW 00239 ♂ Agrodiaetus phyllis 1 C 1 16.07.2000 Qazayd Dagh 25 km O Zanjan 2300 m Zanjan Iran MW 00240 ♂ Agrodiaetus phyllis 1 C P 1 16.07.2000 Qazayd Dagh 25 km O Zanjan 2300 m Zanjan Iran MW 00241 ♂ Agrodiaetus phyllis 1 C P 1 16.07.2000 Qazayd Dagh 25 km O Zanjan 2300 m Zanjan Iran MW 00242 ♀ Agrodiaetus femininoides 1 1 16.07.2000 Qazayd Dagh 25 km O Zanjan 2300 m Zanjan Iran MW 00243 ♀ Agrodiaetus femininoides 1 1 16.07.2000 Qazayd Dagh 25 km O Zanjan 2300 m Zanjan Iran MW 00244 ♀ Agrodiaetus alcestis 1 1 16.07.2000 Qazayd Dagh 25 km O Zanjan 2300 m Zanjan Iran

- 151 - Appendix 2 List of material

Code Sex Genus Species Wgs Fix Kar Alc DNA Cytb CO1 ND1 ITS2 Fig. Date Locality Area Altitude Province Country MW 00245 ♂ Agrodiaetus phyllis 1 1 16.07.2000 Qazayd Dagh 25 km O Zanjan 2300 m Zanjan Iran MW 00246 ♂ Agrodiaetus phyllis 1 1 16.07.2000 Qazayd Dagh 25 km O Zanjan 2300 m Zanjan Iran MW 00247 ♂ Agrodiaetus phyllis 1 1 16.07.2000 Qazayd Dagh 25 km O Zanjan 2300 m Zanjan Iran MW 00248 ♀ Agrodiaetus firdussii 1 1 16.07.2000 Qazayd Dagh 25 km O Zanjan 2300 m Zanjan Iran MW 00249 ♀ Agrodiaetus firdussii 1 1 16.07.2000 Qazayd Dagh 25 km O Zanjan 2300 m Zanjan Iran MW 00250 ♂ Polyommatus thersites 1 1 16.07.2000 Qazayd Dagh 25 km O Zanjan 2300 m Zanjan Iran MW 00251 ♂ Polyommatus icarus 1 1 16.07.2000 Qazayd Dagh 25 km O Zanjan 2300 m Zanjan Iran MW 00252 ♂ Polyommatus icarus 1 1 16.07.2000 Qazayd Dagh 25 km O Zanjan 2300 m Zanjan Iran MW 00253 ♂ Plebeius argus 1 1 16.07.2000 Qazayd Dagh 25 km O Zanjan 2300 m Zanjan Iran MW 00254 ♀ Plebeius argus 1 1 16.07.2000 Qazayd Dagh 25 km O Zanjan 2300 m Zanjan Iran MW 00255 ♂ Meleageria daphnis 1 1 16.07.2000 Qazayd Dagh 25 km O Zanjan 2300 m Zanjan Iran MW 00256 ♂ Meleageria daphnis 1 1 16.07.2000 Qazayd Dagh 25 km O Zanjan 2300 m Zanjan Iran MW 00257 ♀ Meleageria daphnis 1 1 16.07.2000 Qazayd Dagh 25 km O Zanjan 2300 m Zanjan Iran MW 00258 ♀ Meleageria daphnis 1 1 16.07.2000 Qazayd Dagh 25 km O Zanjan 2300 m Zanjan Iran MW 00259 ♂ Agrodiaetus klausschuriani 1 C P 1 1 1 18.07.2000 Veresk südl. Veresk 1800-1950 m Mazandaran Iran MW 00260 ♂ Agrodiaetus klausschuriani 1 C P 1 18.07.2000 Veresk südl. Veresk 1800-1950 m Mazandaran Iran MW 00261 ♂ Agrodiaetus klausschuriani 1 C P 1 18.07.2000 Veresk südl. Veresk 1800-1950 m Mazandaran Iran MW 00262 ♂ Agrodiaetus klausschuriani 1 C P 1 1 1 1 I-25 18.07.2000 Veresk südl. Veresk 1800-1950 m Mazandaran Iran MW 00263 ♂ Agrodiaetus klausschuriani 1 C P 1 18.07.2000 Veresk südl. Veresk 1800-1950 m Mazandaran Iran MW 00264 ♂ Agrodiaetus klausschuriani 1 C P 1 18.07.2000 Veresk südl. Veresk 1800-1950 m Mazandaran Iran MW 00265 ♀ Agrodiaetus klausschuriani 1 1 18.07.2000 Veresk südl. Veresk 1800-1950 m Mazandaran Iran MW 00266 ♂ Agrodiaetus birunii 1 C P 1 18.07.2000 Veresk südl. Veresk 1800-1950 m Mazandaran Iran MW 00267 ♂ Agrodiaetus birunii 1 C P 1 1 1 1 18.07.2000 Veresk südl. Veresk 1800-1950 m Mazandaran Iran MW 00268 ♂ Agrodiaetus birunii 1 C P 1 18.07.2000 Veresk südl. Veresk 1800-1950 m Mazandaran Iran MW 00269 ♂ Agrodiaetus iphidamon 1 C P 1 1 1 1 I-26 18.07.2000 Veresk südl. Veresk 1800-1950 m Mazandaran Iran MW 00270 ♂ Agrodiaetus iphidamon 1 C 1 18.07.2000 Veresk südl. Veresk 1800-1950 m Mazandaran Iran MW 00271 ♂ Agrodiaetus iphidamon 1 C 1 18.07.2000 Veresk südl. Veresk 1800-1950 m Mazandaran Iran MW 00272 ♂ Agrodiaetus iphidamon 1 C 1 18.07.2000 Veresk südl. Veresk 1800-1950 m Mazandaran Iran MW 00273 ♂ Agrodiaetus iphidamon 1 C 1 18.07.2000 Veresk südl. Veresk 1800-1950 m Mazandaran Iran MW 00274 ♂ Agrodiaetus iphidamon 1 C P 1 18.07.2000 Veresk südl. Veresk 1800-1950 m Mazandaran Iran MW 00275 ♂ Agrodiaetus iphidamon 1 C 1 18.07.2000 Veresk südl. Veresk 1800-1950 m Mazandaran Iran MW 00276 ♂ Agrodiaetus iphidamon 1 C 1 18.07.2000 Veresk südl. Veresk 1800-1950 m Mazandaran Iran MW 00277 ♂ Agrodiaetus iphidamon 1 1 18.07.2000 Veresk südl. Veresk 1800-1950 m Mazandaran Iran MW 00278 ♂ Agrodiaetus iphidamon 1 1 18.07.2000 Veresk südl. Veresk 1800-1950 m Mazandaran Iran MW 00279 ♂ Agrodiaetus iphidamon 1 C 1 18.07.2000 Veresk südl. Veresk 1800-1950 m Mazandaran Iran MW 00280 ♂ Agrodiaetus iphidamon 1 C 1 18.07.2000 Veresk südl. Veresk 1800-1950 m Mazandaran Iran MW 00281 ♂ Agrodiaetus iphidamon 1 C 1 18.07.2000 Veresk südl. Veresk 1800-1950 m Mazandaran Iran MW 00282 ♂ Agrodiaetus iphidamon 1 C 1 18.07.2000 Veresk südl. Veresk 1800-1950 m Mazandaran Iran MW 00283 ♂ Agrodiaetus iphidamon 1 C 1 18.07.2000 Veresk südl. Veresk 1800-1950 m Mazandaran Iran MW 00284 ♂ Agrodiaetus iphidamon 1 C 1 18.07.2000 Veresk südl. Veresk 1800-1950 m Mazandaran Iran MW 00285 ♂ Agrodiaetus iphidamon 1 C 1 18.07.2000 Veresk südl. Veresk 1800-1950 m Mazandaran Iran MW 00286 ♂ Agrodiaetus iphidamon 1 C 1 18.07.2000 Veresk südl. Veresk 1800-1950 m Mazandaran Iran MW 00287 ♂ Agrodiaetus iphidamon 1 C P 1 18.07.2000 Veresk südl. Veresk 1800-1950 m Mazandaran Iran MW 00288 ♂ Agrodiaetus iphidamon 1 C P 1 18.07.2000 Veresk südl. Veresk 1800-1950 m Mazandaran Iran MW 00289 ♀ Vacciniina alcedo 1 C 1 18.07.2000 Veresk südl. Veresk 1800-1950 m Mazandaran Iran MW 00290 ♂ Meleageria marcida 1 C P 1 1 1 1 18.07.2000 Veresk südl. Veresk 1800-1950 m Mazandaran Iran MW 00291 ♂ Meleageria marcida 1 C 1 18.07.2000 Veresk südl. Veresk 1800-1950 m Mazandaran Iran MW 00292 ♂ Meleageria marcida 1 C 1 18.07.2000 Veresk südl. Veresk 1800-1950 m Mazandaran Iran MW 00293 ♂ Meleageria marcida 1 C P 1 18.07.2000 Veresk südl. Veresk 1800-1950 m Mazandaran Iran MW 00294 ♂ Meleageria marcida 1 C P 1 18.07.2000 Veresk südl. Veresk 1800-1950 m Mazandaran Iran MW 00295 ♂ Meleageria marcida 1 1 18.07.2000 Veresk südl. Veresk 1800-1950 m Mazandaran Iran MW 00296 ♂ Meleageria marcida 1 1 18.07.2000 Veresk südl. Veresk 1800-1950 m Mazandaran Iran MW 00297 ♂ Meleageria marcida 1 1 18.07.2000 Veresk südl. Veresk 1800-1950 m Mazandaran Iran MW 00298 ♂ Meleageria marcida 1 1 18.07.2000 Veresk südl. Veresk 1800-1950 m Mazandaran Iran MW 00299 ♂ Meleageria marcida 1 1 18.07.2000 Veresk südl. Veresk 1800-1950 m Mazandaran Iran MW 00300 ♂ Meleageria marcida 1 1 18.07.2000 Veresk südl. Veresk 1800-1950 m Mazandaran Iran MW 00301 ♀ Lysandra bellargus 1 1 18.07.2000 Veresk südl. Veresk 1800-1950 m Mazandaran Iran MW 00302 ♂ Polyommatus thersites 1 1 1 1 1 1 18.07.2000 Veresk südl. Veresk 1800-1950 m Mazandaran Iran MW 00303 ♂ Polyommatus icarus 1 1 18.07.2000 Veresk südl. Veresk 1800-1950 m Mazandaran Iran MW 00304 ♂ Agrodiaetus phyllis 1 C P 1 19.07.2000 Asadbar westl. Nesa 2900 m Tehran Iran MW 00305 ♂ Agrodiaetus phyllis 1 C 1 19.07.2000 Asadbar westl. Nesa 2900 m Tehran Iran MW 00306 ♂ Agrodiaetus phyllis 1 C 1 19.07.2000 Asadbar westl. Nesa 2900 m Tehran Iran MW 00307 ♂ Agrodiaetus phyllis 1 C P 1 19.07.2000 Asadbar westl. Nesa 2900 m Tehran Iran MW 00308 ♂ Agrodiaetus phyllis 1 C 1 19.07.2000 Asadbar westl. Nesa 2500 m Tehran Iran MW 00309 ♂ Agrodiaetus phyllis 1 C 1 19.07.2000 Asadbar westl. Nesa 2500 m Tehran Iran MW 00310 ♂ Agrodiaetus phyllis 1 C P 1 19.07.2000 Asadbar westl. Nesa 2500 m Tehran Iran MW 00311 ♂ Agrodiaetus phyllis 1 C P 1 19.07.2000 Asadbar westl. Nesa 2500 m Tehran Iran MW 00312 ♂ Agrodiaetus elbursicus 1 C P 1 19.07.2000 Asadbar westl. Nesa 2500 m Tehran Iran MW 00313 ♂ Agrodiaetus elbursicus 1 C 1 19.07.2000 Asadbar westl. Nesa 2500 m Tehran Iran MW 00314 ♂ Agrodiaetus elbursicus 1 C 1 19.07.2000 Asadbar westl. Nesa 2500 m Tehran Iran MW 00315 ♂ Agrodiaetus elbursicus 1 C P 1 19.07.2000 Asadbar westl. Nesa 2500 m Tehran Iran MW 00316 ♂ Agrodiaetus elbursicus 1 C P 1 1 1 1 19.07.2000 Asadbar westl. Nesa 2500 m Tehran Iran MW 00317 ♂ Vacciniina alcedo 1 C 1 19.07.2000 Asadbar westl. Nesa 2500 m Tehran Iran MW 00318 ♂ Plebeius loewii 1 1 19.07.2000 Asadbar westl. Nesa 2500 m Tehran Iran MW 00319 ♀ Polyommatus thersites 1 1 19.07.2000 Asadbar westl. Nesa 2500 m Tehran Iran MW 00320 ♂ Aricia agestis 1 1 19.07.2000 Asadbar westl. Nesa 2500 m Tehran Iran MW 00321 ♂ Meleageria daphnis 1 1 19.07.2000 Asadbar westl. Nesa 2500 m Tehran Iran MW 00322 ♀ Meleageria daphnis 1 1 19.07.2000 Asadbar westl. Nesa 2500 m Tehran Iran MW 00323 ♂ Plebeius loewii 1 1 19.07.2000 Asadbar westl. Nesa 2900 m Tehran Iran MW 00324 ♂ Plebeius loewii 1 1 19.07.2000 Asadbar westl. Nesa 2900 m Tehran Iran MW 00325 ♂ Plebeius loewii 1 1 19.07.2000 Asadbar westl. Nesa 2900 m Tehran Iran MW 00326 ♂ Polyommatus aedon 1 C 1 1 1 1 21.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00327 ♂ Agrodiaetus erschoffii 1 C P 1 1 1 21.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00328 ♂ Agrodiaetus iphidamon 1 C P 1 1 1 1 1 21.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00329 ♂ Agrodiaetus pseudoxerxes 1 C P 1 21.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00330 ♂ Agrodiaetus pseudoxerxes 1 C P 1 1 1 1 I-27 21.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00331 ♂ Agrodiaetus pseudoxerxes 1 C P 1 21.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00332 ♂ Agrodiaetus phyllis 1 C 1 21.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00333 ♂ Agrodiaetus phyllis 1 C 1 21.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00334 ♂ Agrodiaetus phyllis 1 C 1 21.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00335 ♂ Agrodiaetus caeruleus 1 C P 1 1 1 21.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00336 ♂ Agrodiaetus caeruleus 1 C P 1 21.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00337 ♂ Agrodiaetus caeruleus 1 C 1 21.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00338 ♂ Agrodiaetus iphidamon 1 C P 1 21.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00339 ♂ Agrodiaetus caeruleus 1 C 1 21.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00340 ♂ Agrodiaetus caeruleus 1 C 1 21.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00341 ♂ Agrodiaetus iphidamon 1 C 1 21.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00342 ♀ Agrodiaetus caeruleus 1 1 21.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00343 ♀ Agrodiaetus caeruleus 1 1 21.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00344 ♂ Agrodiaetus posthumus 1 C P 1 21.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00345 ♂ Agrodiaetus caeruleus 1 C 1 21.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00346 ♂ Agrodiaetus phyllis 1 C 1 21.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00347 ♂ Agrodiaetus posthumus 1 C P 1 1 1 1 I-28 21.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00348 ♂ Agrodiaetus phyllis 1 C P 1 1 1 1 I-29 21.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00349 ♂ Agrodiaetus iphidamon 1 C 1 21.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00350 ♂ Agrodiaetus iphidamon 1 C 1 21.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00351 ♂ Agrodiaetus phyllis 1 C 1 21.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00352 ♂ Agrodiaetus caeruleus 1 C 1 21.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00353 ♂ Agrodiaetus phyllis 1 C 1 21.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00354 ♂ Agrodiaetus phyllis 1 C 1 21.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00355 ♂ Agrodiaetus phyllis 1 C P 1 21.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00356 ♂ Agrodiaetus iphidamon 1 C 1 21.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00357 ♂ Agrodiaetus caeruleus 1 C P 1 21.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00358 ♂ Agrodiaetus posthumus 1 C P 1 1 1 21.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00359 ♂ Agrodiaetus iphidamon 1 C 1 21.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran

- 152 - Appendix 2 List of material

Code Sex Genus Species Wgs Fix Kar Alc DNA Cytb CO1 ND1 ITS2 Fig. Date Locality Area Altitude Province Country MW 00360 ♂ Agrodiaetus phyllis 1 C 1 21.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00361 ♂ Agrodiaetus phyllis 1 C 1 21.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00362 ♂ Agrodiaetus iphidamon 1 C P 1 21.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00363 ♂ Agrodiaetus iphidamon 1 C 1 21.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00364 ♂ Polyommatus aedon 1 C 1 21.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00365 ♂ Agrodiaetus caeruleus 1 C P 1 21.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00366 ♂ Agrodiaetus caeruleus 1 C 1 21.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00367 ♂ Agrodiaetus phyllis 1 C 1 21.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00368 ♂ Agrodiaetus iphidamon 1 C 1 21.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00369 ♂ Agrodiaetus iphidamon 1 C 1 21.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00370 ♂ Agrodiaetus posthumus 1 C P 1 21.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00371 ♂ Agrodiaetus phyllis 1 C 1 21.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00372 ♂ Agrodiaetus iphidamon 1 C 1 21.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00373 ♂ Agrodiaetus caeruleus 1 C 1 21.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00374 ♂ Agrodiaetus iphidamon 1 C 1 21.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00375 ♂ Agrodiaetus iphidamon 1 C 1 21.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00376 ♂ Agrodiaetus caeruleus 1 C 1 21.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00377 ♂ Agrodiaetus posthumus 1 C P 1 21.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00378 ♂ Agrodiaetus posthumus 1 C P 1 21.07.2000 Shakuh 36°32.418'N-54°25.956 E 3000 m Golestan Iran MW 00379 ♂ Agrodiaetus posthumus 1 C P 1 21.07.2000 Shakuh 36°32.418'N-54°25.956 E 3000 m Golestan Iran MW 00380 ♂ Agrodiaetus phyllis 1 C 1 21.07.2000 Shakuh 36°32.418'N-54°25.956 E 3000 m Golestan Iran MW 00381 ♂ Agrodiaetus iphidamon 1 C P 1 21.07.2000 Shakuh 36°32.418'N-54°25.956 E 3000 m Golestan Iran MW 00382 ♂ Agrodiaetus phyllis 1 C P 1 21.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00383 ♂ Agrodiaetus caeruleus 1 C P 1 21.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00384 ♂ Agrodiaetus erschoffii 1 C P 1 22.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00385 ♂ Agrodiaetus erschoffii 1 C P 1 22.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00386 ♀ Agrodiaetus caeruleus 1 1 22.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00387 ♂ Agrodiaetus iphidamon 1 C 1 23.07.2000 Hajiabad 25 km SSW Gorgan 2150 m Golestan Iran MW 00388 ♂ Agrodiaetus iphidamon 1 C 1 23.07.2000 Hajiabad 25 km SSW Gorgan 2150 m Golestan Iran MW 00389 ♂ Agrodiaetus iphidamon 1 C P 1 23.07.2000 Hajiabad 25 km SSW Gorgan 2150 m Golestan Iran MW 00390 ♂ Agrodiaetus caeruleus 1 C 1 23.07.2000 Hajiabad 25 km SSW Gorgan 2150 m Golestan Iran MW 00391 ♂ Agrodiaetus caeruleus 1 C 1 23.07.2000 Hajiabad 25 km SSW Gorgan 2150 m Golestan Iran MW 00392 ♂ Agrodiaetus iphidamon 1 C 1 23.07.2000 Hajiabad 25 km SSW Gorgan 2150 m Golestan Iran MW 00393 ♂ Agrodiaetus erschoffii 1 C P 1 1 1 1 I-30 23.07.2000 Hajiabad 25 km SSW Gorgan 2150 m Golestan Iran MW 00394 ♂ Agrodiaetus erschoffii 1 C P 1 23.07.2000 Hajiabad 25 km SSW Gorgan 2150 m Golestan Iran MW 00395 ♂ Agrodiaetus erschoffii 1 1 23.07.2000 Hajiabad 25 km SSW Gorgan 2150 m Golestan Iran MW 00396 ♂ Agrodiaetus erschoffii 1 1 23.07.2000 Hajiabad 25 km SSW Gorgan 2150 m Golestan Iran MW 00397 ♂ Agrodiaetus erschoffii 1 1 23.07.2000 Hajiabad 25 km SSW Gorgan 2150 m Golestan Iran MW 00398 ♂ Agrodiaetus erschoffii 1 1 23.07.2000 Hajiabad 25 km SSW Gorgan 2150 m Golestan Iran MW 00399 ♂ Agrodiaetus erschoffii 1 1 23.07.2000 Hajiabad 25 km SSW Gorgan 2150 m Golestan Iran MW 00400 ♀ Agrodiaetus caeruleus 1 1 23.07.2000 Hajiabad 25 km SSW Gorgan 2150 m Golestan Iran MW 00401 ♀ Agrodiaetus caeruleus 1 1 23.07.2000 Hajiabad 25 km SSW Gorgan 2150 m Golestan Iran MW 00402 ♀ Agrodiaetus caeruleus 1 1 23.07.2000 Hajiabad 25 km SSW Gorgan 2150 m Golestan Iran MW 00403 ♂ Agrodiaetus posthumus 1 C P 1 24.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00404 ♂ Agrodiaetus phyllis 1 C 1 23.07.2000 Hajiabad 25 km SSW Gorgan 2150 m Golestan Iran MW 00405 ♂ Agrodiaetus iphidamon 1 C 1 23.07.2000 Hajiabad 25 km SSW Gorgan 2150 m Golestan Iran MW 00406 ♂ Agrodiaetus erschoffii 1 C P 1 23.07.2000 Hajiabad 25 km SSW Gorgan 2150 m Golestan Iran MW 00407 ♂ Agrodiaetus caeruleus 1 C 1 23.07.2000 Hajiabad 25 km SSW Gorgan 2150 m Golestan Iran MW 00408 ♂ Agrodiaetus caeruleus 1 C 1 23.07.2000 Hajiabad 25 km SSW Gorgan 2150 m Golestan Iran MW 00409 ♂ Agrodiaetus caeruleus 1 C P 1 1 1 1 1 I-31 23.07.2000 Hajiabad 25 km SSW Gorgan 2150 m Golestan Iran MW 00410 ♂ Agrodiaetus caeruleus 1 C P 1 23.07.2000 Hajiabad 25 km SSW Gorgan 2150 m Golestan Iran MW 00411 ♂ Agrodiaetus caeruleus 1 C 1 23.07.2000 Hajiabad 25 km SSW Gorgan 2150 m Golestan Iran MW 00412 ♂ Polyommatus icarus 1 C 1 1 1 1 1 23.07.2000 Hajiabad 25 km SSW Gorgan 2150 m Golestan Iran MW 00413 ♂ Polyommatus icarus 1 C 1 23.07.2000 Hajiabad 25 km SSW Gorgan 2150 m Golestan Iran MW 00414 ♂ Agrodiaetus caeruleus 1 C 1 23.07.2000 Hajiabad 25 km SSW Gorgan 2150 m Golestan Iran MW 00415 ♂ Agrodiaetus iphidamon 1 C 1 23.07.2000 Hajiabad 25 km SSW Gorgan 2150 m Golestan Iran MW 00416 ♂ Agrodiaetus iphidamon 1 C 1 23.07.2000 Hajiabad 25 km SSW Gorgan 2150 m Golestan Iran MW 00417 ♂ Agrodiaetus caeruleus 1 C 1 23.07.2000 Hajiabad 25 km SSW Gorgan 2150 m Golestan Iran MW 00418 ♂ Agrodiaetus caeruleus 1 C 1 23.07.2000 Hajiabad 25 km SSW Gorgan 2150 m Golestan Iran MW 00419 ♂ Agrodiaetus posthumus 1 C P 1 24.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00420 ♂ Agrodiaetus iphidamon 1 C 1 23.07.2000 Hajiabad 25 km SSW Gorgan 2150 m Golestan Iran MW 00421 ♂ Agrodiaetus caeruleus 1 C P 1 23.07.2000 Hajiabad 25 km SSW Gorgan 2150 m Golestan Iran MW 00422 ♂ Agrodiaetus iphidamon 1 C 1 23.07.2000 Hajiabad 25 km SSW Gorgan 2150 m Golestan Iran MW 00423 ♂ Agrodiaetus iphidamon 1 C 1 23.07.2000 Hajiabad 25 km SSW Gorgan 2150 m Golestan Iran MW 00424 ♂ Agrodiaetus iphidamon 1 C P 1 1 1 23.07.2000 Hajiabad 25 km SSW Gorgan 2150 m Golestan Iran MW 00425 ♂ Agrodiaetus erschoffii 1 C P 1 23.07.2000 Hajiabad 25 km SSW Gorgan 2150 m Golestan Iran MW 00426 ♀ Agrodiaetus caeruleus 1 C 1 I-32 23.07.2000 Hajiabad 25 km SSW Gorgan 2150 m Golestan Iran MW 00427 ♀ Agrodiaetus caeruleus 1 1 23.07.2000 Hajiabad 25 km SSW Gorgan 2150 m Golestan Iran MW 00428 ♀ Agrodiaetus caeruleus 1 1 23.07.2000 Hajiabad 25 km SSW Gorgan 2150 m Golestan Iran MW 00429 ♀ Agrodiaetus caeruleus 1 1 23.07.2000 Hajiabad 25 km SSW Gorgan 2150 m Golestan Iran MW 00430 ♀ Agrodiaetus caeruleus 1 1 23.07.2000 Hajiabad 25 km SSW Gorgan 2150 m Golestan Iran MW 00431 ♂ Polyommatus aedon 1 C 1 24.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00432 ♂ Agrodiaetus phyllis 1 C 1 24.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00433 ♂ Agrodiaetus caeruleus 1 C 1 24.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00434 ♂ Agrodiaetus posthumus 1 C P 1 24.07.2000 Shakuh 35 km S Gorgan 2900 m Golestan Iran MW 00435 ♂ Polyommatus aedon 1 C 1 25.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00436 ♂ Polyommatus aedon 1 C P 1 25.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00437 ♂ Polyommatus aedon 1 C P 1 25.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00438 ♂ Agrodiaetus posthumus 1 C P 1 25.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00439 ♂ Agrodiaetus phyllis 1 1 25.07.2000 Shakuh 36°32.418'N-54°25.956 E 2600 m Golestan Iran MW 00440 ♂ Agrodiaetus birunii 1 C 1 26.07.2000 Polur südl. Polur 2200 m Tehran Iran MW 00441 ♂ Agrodiaetus birunii 1 C 1 26.07.2000 Polur südl. Polur 2200 m Tehran Iran MW 00442 ♂ Agrodiaetus birunii 1 C 1 26.07.2000 Polur südl. Polur 2200 m Tehran Iran MW 00443 ♂ Agrodiaetus birunii 1 C 1 26.07.2000 Polur südl. Polur 2200 m Tehran Iran MW 00444 ♂ Agrodiaetus birunii 1 C P 1 1 1 1 26.07.2000 Polur südl. Polur 2200 m Tehran Iran MW 00445 ♂ Agrodiaetus birunii 1 C P 1 26.07.2000 Polur südl. Polur 2200 m Tehran Iran MW 00446 ♂ Agrodiaetus birunii 1 C 1 26.07.2000 Polur südl. Polur 2200 m Tehran Iran MW 00447 ♂ Agrodiaetus phyllis 1 C P 1 26.07.2000 Polur südl. Polur 2200 m Tehran Iran MW 00448 ♂ Agrodiaetus darius 1 C P 1 26.07.2000 Polur südl. Polur 2200 m Tehran Iran MW 00449 ♂ Agrodiaetus birunii 1 C 1 26.07.2000 Polur südl. Polur 2200 m Tehran Iran MW 00450 ♂ Agrodiaetus birunii 1 C P 1 26.07.2000 Polur südl. Polur 2200 m Tehran Iran MW 00451 ♂ Agrodiaetus phyllis 1 C 1 26.07.2000 Polur südl. Polur 2200 m Tehran Iran MW 00452 ♂ Agrodiaetus phyllis 1 C P 1 1 1 1 26.07.2000 Polur südl. Polur 2200 m Tehran Iran MW 00453 ♂ Agrodiaetus phyllis 1 C 1 26.07.2000 Polur südl. Polur 2200 m Tehran Iran MW 00454 ♂ Agrodiaetus phyllis 1 C 1 26.07.2000 Polur südl. Polur 2200 m Tehran Iran MW 00455 ♂ Agrodiaetus phyllis 1 C 1 26.07.2000 Polur südl. Polur 2200 m Tehran Iran MW 00456 ♂ Agrodiaetus phyllis 1 C 1 26.07.2000 Polur südl. Polur 2200 m Tehran Iran MW 00457 ♂ Agrodiaetus birunii 1 C P 1 26.07.2000 Polur südl. Polur 2200 m Tehran Iran MW 00458 ♂ Agrodiaetus phyllis 1 C P 1 26.07.2000 Polur südl. Polur 2200 m Tehran Iran MW 00459 ♂ Agrodiaetus phyllis 1 C 1 26.07.2000 Polur südl. Polur 2200 m Tehran Iran MW 00460 ♂ Agrodiaetus phyllis 1 C 1 26.07.2000 Polur südl. Polur 2200 m Tehran Iran MW 00461 ♂ Agrodiaetus phyllis 1 C 1 26.07.2000 Polur südl. Polur 2200 m Tehran Iran MW 00462 ♂ Agrodiaetus phyllis 1 C 1 26.07.2000 Polur südl. Polur 2200 m Tehran Iran MW 00463 ♂ Agrodiaetus birunii 1 C P 1 26.07.2000 Polur südl. Polur 2200 m Tehran Iran MW 00464 ♂ Agrodiaetus phyllis 1 1 26.07.2000 Polur südl. Polur 2200 m Tehran Iran MW 00465 ♂ Agrodiaetus phyllis 1 1 26.07.2000 Polur südl. Polur 2200 m Tehran Iran MW 00466 ♀ Agrodiaetus birunii 1 1 26.07.2000 Polur südl. Polur 2200 m Tehran Iran MW 00467 ♂ Lycaena thetis 1 1 26.07.2000 Polur südl. Polur 2200 m Tehran Iran MW 00468 ♂ Lycaena thetis 1 1 26.07.2000 Polur südl. Polur 2200 m Tehran Iran MW 00469 ♂ Lysandra bellargus 1 1 1 1 1 26.07.2000 Polur südl. Polur 2200 m Tehran Iran MW 00470 ♂ Polyommatus icarus 1 1 26.07.2000 Polur südl. Polur 2200 m Tehran Iran MW 00471 ♂ Polyommatus icarus 1 1 26.07.2000 Polur südl. Polur 2200 m Tehran Iran MW 00472 ♂ Agrodiaetus birunii 1 C P 1 27.07.2000 Pul-e Zanguleh 15 km NO Kendevan 2400 m Mazandaran Iran MW 00473 ♂ Agrodiaetus birunii 1 C 1 27.07.2000 Pul-e Zanguleh 15 km NO Kendevan 2400 m Mazandaran Iran MW 00474 ♂ Polyommatus aedon 1 C 1 28.07.2000 Golestanak Nature Reserve 25 km NO Kendevan 2700-3200 m Mazandaran Iran

- 153 - Appendix 2 List of material

Code Sex Genus Species Wgs Fix Kar Alc DNA Cytb CO1 ND1 ITS2 Fig. Date Locality Area Altitude Province Country MW 00475 ♂ Neolysandra corona 1 C P 1 28.07.2000 Golestanak Nature Reserve 25 km NO Kendevan 2700-3200 m Mazandaran Iran MW 00476 ♂ Agrodiaetus birunii 1 C P 1 1 1 28.07.2000 Golestanak Nature Reserve 25 km NO Kendevan 2700-3200 m Mazandaran Iran MW 00477 ♂ Agrodiaetus iphidamon 1 C 1 28.07.2000 Golestanak Nature Reserve 25 km NO Kendevan 2700-3200 m Mazandaran Iran MW 00478 ♂ Agrodiaetus iphidamon 1 C 1 28.07.2000 Golestanak Nature Reserve 25 km NO Kendevan 2700-3200 m Mazandaran Iran MW 00479 ♂ Agrodiaetus iphidamon 1 C 1 28.07.2000 Golestanak Nature Reserve 25 km NO Kendevan 2700-3200 m Mazandaran Iran MW 00480 ♂ Agrodiaetus birunii 1 C P 1 28.07.2000 Golestanak Nature Reserve 25 km NO Kendevan 2700-3200 m Mazandaran Iran MW 00481 ♂ Agrodiaetus birunii 1 C 1 28.07.2000 Golestanak Nature Reserve 25 km NO Kendevan 2700-3200 m Mazandaran Iran MW 00482 ♂ Agrodiaetus birunii 1 C P 1 28.07.2000 Golestanak Nature Reserve 25 km NO Kendevan 2700-3200 m Mazandaran Iran MW 00483 ♂ Agrodiaetus birunii 1 C P 1 28.07.2000 Golestanak Nature Reserve 25 km NO Kendevan 2700-3200 m Mazandaran Iran MW 00484 ♂ Agrodiaetus birunii 1 C P 1 28.07.2000 Golestanak Nature Reserve 25 km NO Kendevan 2700-3200 m Mazandaran Iran MW 00485 ♂ Agrodiaetus birunii 1 C P 1 28.07.2000 Golestanak Nature Reserve 25 km NO Kendevan 2700-3200 m Mazandaran Iran MW 00486 ♂ Agrodiaetus birunii 1 C P 1 28.07.2000 Golestanak Nature Reserve 25 km NO Kendevan 2700-3200 m Mazandaran Iran MW 00487 ♂ Agrodiaetus birunii 1 C P 1 28.07.2000 Golestanak Nature Reserve 25 km NO Kendevan 2700-3200 m Mazandaran Iran MW 00488 ♂ Agrodiaetus birunii 1 C P 1 28.07.2000 Golestanak Nature Reserve 25 km NO Kendevan 2700-3200 m Mazandaran Iran MW 00489 ♂ Agrodiaetus birunii 1 C P 1 28.07.2000 Golestanak Nature Reserve 25 km NO Kendevan 2700-3200 m Mazandaran Iran MW 00490 ♂ Polyommatus eroides 1 C 1 28.07.2000 Golestanak Nature Reserve 25 km NO Kendevan 2700-3200 m Mazandaran Iran MW 00491 ♂ Polyommatus eroides 1 C 1 28.07.2000 Golestanak Nature Reserve 25 km NO Kendevan 2700-3200 m Mazandaran Iran MW 00492 ♂ Polyommatus eroides 1 1 28.07.2000 Golestanak Nature Reserve 25 km NO Kendevan 2700-3200 m Mazandaran Iran MW 00493 ♂ Polyommatus eroides 1 1 28.07.2000 Golestanak Nature Reserve 25 km NO Kendevan 2700-3200 m Mazandaran Iran MW 00494 ♂ Agrodiaetus birunii 1 1 28.07.2000 Golestanak Nature Reserve 25 km NO Kendevan 2700-3200 m Mazandaran Iran MW 00495 ♀ Agrodiaetus birunii 1 1 28.07.2000 Golestanak Nature Reserve 25 km NO Kendevan 2700-3200 m Mazandaran Iran MW 00496 ♀ Lampides boeticus 1 1 30.07.2000 5 km S Valiabad Valiabad 1900 m Mazandaran Iran MW 00497 ♂ Favonius quercus 1 1 1 1 1 30.07.2000 5 km S Valiabad Valiabad 1900 m Mazandaran Iran MW 00498 ♂ Agrodiaetus valiabadi 1 C P 1 1 1 1 I-33 30.07.2000 5 km S Valiabad Valiabad 1900 m Mazandaran Iran MW 00499 ♂ Agrodiaetus elbursicus 1 C P 1 30.07.2000 Kendevan 5 km N Kendevan 2150 m Mazandaran Iran MW 00500 ♂ Agrodiaetus elbursicus 1 C 1 30.07.2000 Kendevan 5 km N Kendevan 2150 m Mazandaran Iran MW 00501 ♂ Agrodiaetus phyllis 1 C 1 31.07.2000 Takht-e Suleyman südwestl. Marzanabad 3000 m Mazandaran Iran MW 00502 ♂ Agrodiaetus birunii 1 C 1 31.07.2000 Takht-e Suleyman südwestl. Marzanabad 3000 m Mazandaran Iran MW 00503 ♂ Agrodiaetus birunii 1 C P 1 31.07.2000 Takht-e Suleyman südwestl. Marzanabad 3000 m Mazandaran Iran MW 00504 ♂ Neolysandra corona 1 C 1 1 1 1 31.07.2000 Takht-e Suleyman südwestl. Marzanabad 3000 m Mazandaran Iran MW 00505 ♂ Neolysandra corona 1 C 1 31.07.2000 Takht-e Suleyman südwestl. Marzanabad 3000 m Mazandaran Iran MW 00506 ♂ Neolysandra corona 1 C 1 31.07.2000 Takht-e Suleyman südwestl. Marzanabad 3000 m Mazandaran Iran MW 00507 ♂ Neolysandra corona 1 C 1 31.07.2000 Takht-e Suleyman südwestl. Marzanabad 3000 m Mazandaran Iran MW 00508 ♂ Neolysandra corona 1 C 1 31.07.2000 Takht-e Suleyman südwestl. Marzanabad 3000 m Mazandaran Iran MW 00509 ♂ Agrodiaetus birunii 1 1 31.07.2000 Takht-e Suleyman südwestl. Marzanabad 3000 m Mazandaran Iran MW 00510 ♂ Neolysandra corona 1 1 31.07.2000 Takht-e Suleyman südwestl. Marzanabad 3000 m Mazandaran Iran MW 00511 ♀ Neolysandra corona 1 1 31.07.2000 Takht-e Suleyman südwestl. Marzanabad 3000 m Mazandaran Iran MW 00512 ♂ Neolysandra corona 1 C 1 31.07.2000 Takht-e Suleyman südwestl. Marzanabad 3000 m Mazandaran Iran MW 00513 ♂ Neolysandra corona 1 C 1 31.07.2000 Takht-e Suleyman südwestl. Marzanabad 3000 m Mazandaran Iran MW 00514 ♂ Neolysandra corona 1 C 1 31.07.2000 Takht-e Suleyman südwestl. Marzanabad 3000 m Mazandaran Iran MW 00515 ♂ Vacciniina morgianus 1 C 1 31.07.2000 Takht-e Suleyman südwestl. Marzanabad 3000 m Mazandaran Iran MW 00516 ♂ Polyommatus amandus 1 1 31.07.2000 Takht-e Suleyman südwestl. Marzanabad 3000 m Mazandaran Iran MW 00517 ♂ Vacciniina morgianus 1 1 1 1 1 31.07.2000 Takht-e Suleyman südwestl. Marzanabad 3600 m Mazandaran Iran MW 00518 ♂ Vacciniina morgianus 1 1 31.07.2000 Takht-e Suleyman südwestl. Marzanabad 3600 m Mazandaran Iran MW 00519 ♀ Vacciniina morgianus 1 1 31.07.2000 Takht-e Suleyman südwestl. Marzanabad 3600 m Mazandaran Iran MW 00520 ♀ Vacciniina morgianus 1 1 31.07.2000 Takht-e Suleyman südwestl. Marzanabad 3600 m Mazandaran Iran MW 00521 ♂ Vacciniina morgianus 1 1 31.07.2000 Takht-e Suleyman südwestl. Marzanabad 3600 m Mazandaran Iran MW 00522 ♂ Vacciniina morgianus 1 1 31.07.2000 Takht-e Suleyman südwestl. Marzanabad 3600 m Mazandaran Iran MW 00523 ♂ Neolysandra corona 1 C 1 01.08.2000 Takht-e Suleyman südwestl. Marzanabad 3500-3700 m Mazandaran Iran MW 00524 ♂ Neolysandra corona 1 C P 1 01.08.2000 Takht-e Suleyman südwestl. Marzanabad 3500-3700 m Mazandaran Iran MW 00525 ♂ Cyaniris semiargus 1 C 1 1 1 1 01.08.2000 Takht-e Suleyman südwestl. Marzanabad 3500-3700 m Mazandaran Iran MW 00526 ♂ Cyaniris semiargus 1 C 1 01.08.2000 Takht-e Suleyman südwestl. Marzanabad 3500-3700 m Mazandaran Iran MW 00527 ♂ Vacciniina morgianus 1 C 1 01.08.2000 Takht-e Suleyman südwestl. Marzanabad 3500-3700 m Mazandaran Iran MW 00528 ♂ Vacciniina morgianus 1 C 1 01.08.2000 Takht-e Suleyman südwestl. Marzanabad 3500-3700 m Mazandaran Iran MW 00529 ♂ Vacciniina morgianus 1 C 1 01.08.2000 Takht-e Suleyman südwestl. Marzanabad 3500-3700 m Mazandaran Iran MW 00530 ♂ Polyommatus eroides 1 C 1 1 1 1 1 01.08.2000 Takht-e Suleyman südwestl. Marzanabad 3500-3700 m Mazandaran Iran MW 00531 ♂ Polyommatus eroides 1 C 1 01.08.2000 Takht-e Suleyman südwestl. Marzanabad 3500-3700 m Mazandaran Iran MW 00532 ♂ Polyommatus eroides 1 C 1 01.08.2000 Takht-e Suleyman südwestl. Marzanabad 3500-3700 m Mazandaran Iran MW 00533 ♂ Polyommatus eroides 1 C 1 01.08.2000 Takht-e Suleyman südwestl. Marzanabad 3500-3700 m Mazandaran Iran MW 00534 ♂ Polyommatus eroides 1 C P 1 01.08.2000 Takht-e Suleyman südwestl. Marzanabad 3500-3700 m Mazandaran Iran MW 00535 ♀ Polyommatus eroides 1 1 01.08.2000 Takht-e Suleyman südwestl. Marzanabad 3500-3700 m Mazandaran Iran MW 00536 ♀ Cyaniris semiargus 1 1 01.08.2000 Takht-e Suleyman südwestl. Marzanabad 3500-3700 m Mazandaran Iran MW 00537 ♂ Lycaena thersamon 1 1 01.08.2000 Takht-e Suleyman südwestl. Marzanabad 3500-3700 m Mazandaran Iran MW 00538 ♀ Lycaena thersamon 1 1 01.08.2000 Takht-e Suleyman südwestl. Marzanabad 3500-3700 m Mazandaran Iran MW 00539 ♂ Agrodiaetus dizinensis 1 C P 1 1 1 1 I-34 04.08.2000 Dizin Pass südl. Dizin 3200-3300 m Tehran Iran MW 00540 ♂ Agrodiaetus dizinensis 1 C 1 04.08.2000 Dizin Pass südl. Dizin 3200-3300 m Tehran Iran MW 00541 ♂ Agrodiaetus dizinensis 1 C 1 04.08.2000 Dizin Pass südl. Dizin 3200-3300 m Tehran Iran MW 00542 ♂ Agrodiaetus dizinensis 1 C 1 04.08.2000 Dizin Pass südl. Dizin 3200-3300 m Tehran Iran MW 00543 ♂ Agrodiaetus dizinensis 1 C 1 04.08.2000 Dizin Pass südl. Dizin 3200-3300 m Tehran Iran MW 00544 ♂ Agrodiaetus dizinensis 1 C 1 04.08.2000 Dizin Pass südl. Dizin 3200-3300 m Tehran Iran MW 00545 ♂ Agrodiaetus dizinensis 1 C P 1 04.08.2000 Dizin Pass südl. Dizin 3200-3300 m Tehran Iran MW 00546 ♀ Agrodiaetus dizinensis 1 C 1 04.08.2000 Dizin Pass südl. Dizin 3200-3300 m Tehran Iran MW 00547 ♂ Agrodiaetus birunii 1 C P 1 1 1 1 I-35 04.08.2000 Dizin Pass südl. Dizin 3200-3300 m Tehran Iran MW 00548 ♂ Agrodiaetus birunii 1 C P 1 04.08.2000 Dizin Pass südl. Dizin 3200-3300 m Tehran Iran MW 00549 ♂ Agrodiaetus posthumus 1 C 1 04.08.2000 Dizin Pass südl. Dizin 3200-3300 m Tehran Iran MW 00550 ♂ Agrodiaetus posthumus 1 C 1 04.08.2000 Dizin Pass südl. Dizin 3200-3300 m Tehran Iran MW 00551 ♀ Agrodiaetus posthumus 1 1 04.08.2000 Dizin Pass südl. Dizin 3200-3300 m Tehran Iran MW 00552 ♀ Lycaena thersamon 1 1 28.07.2000 Golestanak Nature Reserve 25 km NO Kendevan 2700-3200 m Mazandaran Iran MW 01001 ♂ Agrodiaetus ainsae 1 C P 1 1 1 1 17.07.2001 Ilarduya westl. Eguino 550 m Alava Spain MW 01002 ♂ Agrodiaetus ainsae 1 C 1 17.07.2001 Ilarduya westl. Eguino 550 m Alava Spain MW 01003 ♂ Agrodiaetus ainsae 1 C P 1 17.07.2001 Ilarduya westl. Eguino 550 m Alava Spain MW 01004 ♂ Agrodiaetus ainsae 1 C P 1 17.07.2001 Ilarduya westl. Eguino 550 m Alava Spain MW 01005 ♂ Agrodiaetus ainsae 1 C P 1 17.07.2001 Ilarduya westl. Eguino 550 m Alava Spain MW 01006 ♂ Agrodiaetus ainsae 1 C P 1 17.07.2001 Ilarduya westl. Eguino 550 m Alava Spain MW 01007 ♂ Agrodiaetus ainsae 1 C 1 17.07.2001 Ilarduya westl. Eguino 550 m Alava Spain MW 01008 ♂ Agrodiaetus ainsae 1 C 1 17.07.2001 Ilarduya westl. Eguino 550 m Alava Spain MW 01009 ♂ Polyommatus escheri 1 C P 1 1 1 17.07.2001 Ilarduya westl. Eguino 550 m Alava Spain MW 01010 ♂ Polyommatus escheri 1 C P 1 17.07.2001 Ilarduya westl. Eguino 550 m Alava Spain MW 01011 ♂ Lysandra bellargus 1 C 1 1 1 1 17.07.2001 Ilarduya westl. Eguino 550 m Alava Spain MW 01012 ♂ Polyommatus escheri 1 C 1 17.07.2001 Ilarduya westl. Eguino 550 m Alava Spain MW 01013 ♂ Agrodiaetus ripartii 1 C 1 18.07.2001 Ubierna 20 km N Burgos 900 m Burgos Spain MW 01014 ♂ Agrodiaetus ripartii 1 C P 1 1 1 1 I-36 18.07.2001 Ubierna 20 km N Burgos 900 m Burgos Spain MW 01015 ♂ Agrodiaetus ripartii 1 C P 1 18.07.2001 Ubierna 20 km N Burgos 900 m Burgos Spain MW 01016 ♂ Agrodiaetus ripartii 1 C 1 18.07.2001 Ubierna 20 km N Burgos 900 m Burgos Spain MW 01017 ♂ Agrodiaetus ripartii 1 C P 1 18.07.2001 Ubierna 20 km N Burgos 900 m Burgos Spain MW 01018 ♂ Lysandra coridon 1 C 1 1 1 1 18.07.2001 Ubierna 20 km N Burgos 900 m Burgos Spain MW 01019 ♂ Polyommatus dorylas 1 C P 1 1 1 1 18.07.2001 Ubierna 20 km N Burgos 900 m Burgos Spain MW 01020 ♂ Polyommatus dorylas 1 C 1 18.07.2001 Ubierna 20 km N Burgos 900 m Burgos Spain MW 01021 ♀ Polyommatus dorylas 1 1 18.07.2001 Ubierna 20 km N Burgos 900 m Burgos Spain MW 01022 ♂ Lampides boeticus 1 1 1 1 18.07.2001 Ubierna 20 km N Burgos 900 m Burgos Spain MW 01023 ♂ Leptotes pirithous 1 1 1 1 18.07.2001 Ubierna 20 km N Burgos 900 m Burgos Spain MW 01024 ♂ Aricia agestis 1 1 18.07.2001 Ubierna 20 km N Burgos 900 m Burgos Spain MW 01025 ♀ Polyommatus icarus 1 1 1 1 1 18.07.2001 Ubierna 20 km N Burgos 900 m Burgos Spain MW 01026 ♂ Plebeius argus 1 1 18.07.2001 Ubierna 20 km N Burgos 900 m Burgos Spain MW 01027 ♀ Satyrium esculi 1 1 1 1 18.07.2001 Ubierna 20 km N Burgos 900 m Burgos Spain MW 01028 ♀ Satyrium esculi 1 1 18.07.2001 Ubierna 20 km N Burgos 900 m Burgos Spain MW 01029 ♀ Satyrium esculi 1 1 18.07.2001 Ubierna 20 km N Burgos 900 m Burgos Spain MW 01030 ♂ Satyrium esculi 1 1 18.07.2001 Ubierna 20 km N Burgos 900 m Burgos Spain MW 01031 ♀ Satyrium spini 1 1 18.07.2001 Ubierna 20 km N Burgos 900 m Burgos Spain MW 01032 ♀ Satyrium spini 1 1 18.07.2001 Ubierna 20 km N Burgos 900 m Burgos Spain MW 01033 ♂ Gonepteryx cleopatra 1 1 18.07.2001 Ubierna 20 km N Burgos 900 m Burgos Spain MW 01034 ♂ Lysandra albicans 1 C P 1 1 1 1 19.07.2001 Abejar Sierra de Cabrejas 1100 m Soria Spain MW 01035 ♂ Lysandra albicans 1 C 1 19.07.2001 Abejar Sierra de Cabrejas 1100 m Soria Spain MW 01036 ♂ Agrodiaetus fabressei 1 C P 1 19.07.2001 Abejar Sierra de Cabrejas 1100 m Soria Spain MW 01037 ♂ Agrodiaetus fabressei 1 C P 1 19.07.2001 Abejar Sierra de Cabrejas 1100 m Soria Spain

- 154 - Appendix 2 List of material

Code Sex Genus Species Wgs Fix Kar Alc DNA Cytb CO1 ND1 ITS2 Fig. Date Locality Area Altitude Province Country MW 01038 ♂ Agrodiaetus fabressei 1 C P 1 19.07.2001 Abejar Sierra de Cabrejas 1100 m Soria Spain MW 01039 ♂ Agrodiaetus fabressei 1 C P 1 1 1 1 19.07.2001 Abejar Sierra de Cabrejas 1100 m Soria Spain MW 01040 ♂ Agrodiaetus fabressei 1 C P 1 19.07.2001 Abejar Sierra de Cabrejas 1100 m Soria Spain MW 01041 ♀ Agrodiaetus fabressei 1 1 19.07.2001 Abejar Sierra de Cabrejas 1100 m Soria Spain MW 01042 ♀ Agrodiaetus fabressei 1 1 19.07.2001 Abejar Sierra de Cabrejas 1100 m Soria Spain MW 01043 ♀ Agrodiaetus fabressei 1 1 19.07.2001 Abejar Sierra de Cabrejas 1100 m Soria Spain MW 01044 ♀ Agrodiaetus fabressei 1 1 19.07.2001 Abejar Sierra de Cabrejas 1100 m Soria Spain MW 01045 ♀ Agrodiaetus fabressei 1 1 19.07.2001 Abejar Sierra de Cabrejas 1100 m Soria Spain MW 01046 ♀ Agrodiaetus fabressei 1 1 19.07.2001 Abejar Sierra de Cabrejas 1100 m Soria Spain MW 01047 ♀ Agrodiaetus fabressei 1 1 19.07.2001 Abejar Sierra de Cabrejas 1100 m Soria Spain MW 01048 ♂ Aricia agestis 1 1 1 1 1 19.07.2001 Abejar Sierra de Cabrejas 1100 m Soria Spain MW 01049 ♀ Polyommatus icarus 1 1 19.07.2001 Abejar Sierra de Cabrejas 1100 m Soria Spain MW 01050 ♂ Lycaeides idas 1 1 19.07.2001 Abejar Sierra de Cabrejas 1100 m Soria Spain MW 01051 ♀ Satyrium acaciae 1 1 19.07.2001 de Camaros Sierra de Urbión 1200 m Soria Spain MW 01052 ♂ Agrodiaetus ainsae 1 C P 1 20.07.2001 Sta. Maria Embalse de la Pena 500 m Huesca Spain MW 01053 ♂ Agrodiaetus ainsae 1 C P 1 1 1 1 I-37 20.07.2001 Sta. Maria Embalse de la Pena 500 m Huesca Spain MW 01054 ♂ Agrodiaetus ainsae 1 C P 1 20.07.2001 Sta. Maria Embalse de la Pena 500 m Huesca Spain MW 01055 ♂ Agrodiaetus ainsae 1 C P 1 20.07.2001 Sta. Maria Embalse de la Pena 500 m Huesca Spain MW 01056 ♂ Agrodiaetus ainsae 1 C 1 20.07.2001 Sta. Maria Embalse de la Pena 500 m Huesca Spain MW 01057 ♂ Agrodiaetus ainsae 1 C 1 20.07.2001 Sta. Maria Embalse de la Pena 500 m Huesca Spain MW 01058 ♂ Agrodiaetus ainsae 1 C 1 20.07.2001 Sta. Maria Embalse de la Pena 500 m Huesca Spain MW 01059 ♂ Lysandra albicans 1 C P 1 1 1 1 20.07.2001 Sta. Maria Embalse de la Pena 500 m Huesca Spain MW 01060 ♂ Polyommatus thersites 1 1 20.07.2001 Sta. Maria Embalse de la Pena 500 m Huesca Spain MW 01061 ♂ Aricia cramera 1 1 1 1 1 20.07.2001 Sta. Maria Embalse de la Pena 500 m Huesca Spain MW 01062 ♂ Leptidea sinapis 1 1 20.07.2001 Sta. Maria Embalse de la Pena 500 m Huesca Spain MW 01063 ♂ Coenonympha dorus 1 1 20.07.2001 Sta. Maria Embalse de la Pena 500 m Huesca Spain MW 01064 ♂ Pyronia cecilia 1 1 20.07.2001 Sta. Maria Embalse de la Pena 500 m Huesca Spain MW 01065 ♂ Pyronia tithonus 1 1 20.07.2001 Sta. Maria Embalse de la Pena 500 m Huesca Spain MW 01066 ♀ Pyronia bathseba 1 1 20.07.2001 Sta. Maria Embalse de la Pena 500 m Huesca Spain MW 01067 ♀ Maniola jurtina 1 1 20.07.2001 Sta. Maria Embalse de la Pena 500 m Huesca Spain MW 01068 ♂ Hipparchia semele 1 1 20.07.2001 Sta. Maria Embalse de la Pena 500 m Huesca Spain MW 01069 ♂ Hipparchia fagi 1 1 20.07.2001 Sta. Maria Embalse de la Pena 500 m Huesca Spain MW 01070 ♂ Hipparchia fidia 1 1 20.07.2001 Sta. Maria Embalse de la Pena 500 m Huesca Spain MW 01071 ♂ Hipparchia fidia 1 1 20.07.2001 Sta. Maria Embalse de la Pena 500 m Huesca Spain MW 01072 ♂ Agrodiaetus ripartii 1 C P 1 1 1 21.07.2001 Triste Embalse de la Pena 600 m Huesca Spain MW 01073 ♂ Agrodiaetus ripartii 1 C 1 21.07.2001 Triste Embalse de la Pena 600 m Huesca Spain MW 01074 ♂ Agrodiaetus ripartii 1 C 1 21.07.2001 Triste Embalse de la Pena 600 m Huesca Spain MW 01075 ♂ Agrodiaetus ainsae 1 C 1 21.07.2001 Triste Embalse de la Pena 600 m Huesca Spain MW 01076 ♂ Agrodiaetus ainsae 1 C 1 21.07.2001 Triste Embalse de la Pena 600 m Huesca Spain MW 01077 ♂ Agrodiaetus ainsae 1 C P 1 21.07.2001 Triste Embalse de la Pena 600 m Huesca Spain MW 01078 ♂ Agrodiaetus ainsae 1 C P 1 1 1 21.07.2001 Triste Embalse de la Pena 600 m Huesca Spain MW 01079 ♂ Agrodiaetus ainsae 1 C 1 21.07.2001 Triste Embalse de la Pena 600 m Huesca Spain MW 01080 ♂ Lysandra albicans 1 C 1 21.07.2001 Triste Embalse de la Pena 600 m Huesca Spain MW 01081 ♂ Lysandra albicans 1 C 1 21.07.2001 Triste Embalse de la Pena 600 m Huesca Spain MW 01082 ♂ Polyommatus escheri 1 C 1 21.07.2001 Triste Embalse de la Pena 600 m Huesca Spain MW 01083 ♂ Polyommatus thersites 1 C 1 1 1 1 21.07.2001 Triste Embalse de la Pena 600 m Huesca Spain MW 01084 ♂ Polyommatus thersites 1 C 1 21.07.2001 Triste Embalse de la Pena 600 m Huesca Spain MW 01085 ♂ Polyommatus thersites 1 C 1 21.07.2001 Triste Embalse de la Pena 600 m Huesca Spain MW 01086 ♂ Polyommatus thersites 1 C 1 21.07.2001 Triste Embalse de la Pena 600 m Huesca Spain MW 01087 ♀ Polyommatus thersites 1 1 21.07.2001 Triste Embalse de la Pena 600 m Huesca Spain MW 01088 ♂ Coenonympha arcania 1 1 21.07.2001 Triste Embalse de la Pena 600 m Huesca Spain MW 01089 ♀ Coenonympha dorus 1 1 21.07.2001 Triste Embalse de la Pena 600 m Huesca Spain MW 01090 ♂ Lasiommata megera 1 1 21.07.2001 Triste Embalse de la Pena 600 m Huesca Spain MW 01091 ♂ Melanargia galathea 1 1 21.07.2001 Triste Embalse de la Pena 600 m Huesca Spain MW 01092 ♂ Lysandra albicans 1 C 1 1 1 1 22.07.2001 Boltana Pirineos Centrals 650 m Huesca Spain MW 01093 ♂ Agrodiaetus ainsae 1 C 1 22.07.2001 Boltana Pirineos Centrals 650 m Huesca Spain MW 01094 ♀ Agrodiaetus ainsae 1 1 22.07.2001 Boltana Pirineos Centrals 650 m Huesca Spain MW 01095 ♀ Agrodiaetus ainsae 1 1 22.07.2001 Boltana Pirineos Centrals 650 m Huesca Spain MW 01096 ♀ Agrodiaetus ainsae 1 1 22.07.2001 Boltana Pirineos Centrals 650 m Huesca Spain MW 01097 ♂ Leptotes pirithous 1 1 22.07.2001 Boltana Pirineos Centrals 650 m Huesca Spain MW 01098 ♂ Agrodiaetus ainsae 1 C 1 23.07.2001 Boltana Pirineos Centrals 650 m Huesca Spain MW 01099 ♂ Lysandra albicans 1 C 1 23.07.2001 Boltana Pirineos Centrals 650 m Huesca Spain MW 01100 ♂ Lysandra albicans 1 C 1 23.07.2001 Boltana Pirineos Centrals 650 m Huesca Spain MW 01101 ♂ Papilio machaon 1 1 23.07.2001 Boltana Pirineos Centrals 650 m Huesca Spain MW 01102 ♂ Lysandra coridon 1 C 1 23.07.2001 Sta. Coloma de Queralt 700 m Tarragona Spain MW 01103 ♂ Lysandra coridon 1 C 1 1 1 23.07.2001 Sta. Coloma de Queralt 700 m Tarragona Spain MW 01104 ♂ Agrodiaetus ripartii 1 C P 1 23.07.2001 Sta. Coloma de Queralt 700 m Tarragona Spain MW 01105 ♂ Agrodiaetus ripartii 1 C P 1 1 1 23.07.2001 Sta. Coloma de Queralt 700 m Tarragona Spain MW 01106 ♂ Agrodiaetus fulgens 1 C P 1 23.07.2001 Sta. Coloma de Queralt 700 m Tarragona Spain MW 01107 ♂ Agrodiaetus fulgens 1 C P 1 1 1 1 I-38 23.07.2001 Sta. Coloma de Queralt 700 m Tarragona Spain MW 01108 ♂ Agrodiaetus fulgens 1 C P 1 23.07.2001 Sta. Coloma de Queralt 700 m Tarragona Spain MW 01109 ♂ Agrodiaetus fulgens 1 C P 1 23.07.2001 Sta. Coloma de Queralt 700 m Tarragona Spain MW 01110 ♂ Agrodiaetus fulgens 1 C P 1 23.07.2001 Sta. Coloma de Queralt 700 m Tarragona Spain MW 01111 ♀ Agrodiaetus fulgens 1 1 23.07.2001 Sta. Coloma de Queralt 700 m Tarragona Spain MW 01112 ♀ Agrodiaetus fulgens 1 1 23.07.2001 Sta. Coloma de Queralt 700 m Tarragona Spain MW 01113 ♂ Lasiommata megera 1 1 23.07.2001 Sta. Coloma de Queralt 700 m Tarragona Spain MW 01114 ♀ Iphiclides feisthamelii 1 1 1 1 24.07.2001 Taradell SO Vic 700 m Barcelona Spain MW 01115 ♂ Lysandra coridon 1 1 24.07.2001 Taradell SO Vic 700 m Barcelona Spain MW 01116 ♂ Lysandra coridon 1 1 1 1 1 24.07.2001 Taradell SO Vic 700 m Barcelona Spain MW 01117 ♂ Arethusana arethusa 1 1 24.07.2001 Seva S Vic 600 m Barcelona Spain MW 01118 ♂ Pararge aegeria 1 1 25.07.2001 Lloret de Mar 600 m Girona Spain MW 01119 ♂ Pararge aegeria 1 1 25.07.2001 Lloret de Mar 600 m Girona Spain MW 01120 ♂ Cacyreus marshalli 1 1 1 1 27.07.2001 Maruéjols-les-Gardons 100 m Hérault France MW 01121 ♂ Minois dryas 1 1 27.07.2001 Maruéjols-les-Gardons 100 m Hérault France MW 01401 Zygaena filipendulae 1 1 19.07.2001 Abejar Sierra de Cabrejas 1100 m Soria Spain MW 01402 Zygaena filipendulae 1 1 19.07.2001 Abejar Sierra de Cabrejas 1100 m Soria Spain MW 01403 Zygaena lonicerae 1 1 19.07.2001 Montenegro de Camaros Sierra de Urbión 1200 m Soria Spain MW 01404 Zygaena lonicerae 1 1 19.07.2001 Montenegro de Camaros Sierra de Urbión 1200 m Soria Spain MW 01405 Zygaena lonicerae 1 1 19.07.2001 Montenegro de Camaros Sierra de Urbión 1200 m Soria Spain MW 01406 Zygaena lonicerae 1 1 19.07.2001 Montenegro de Camaros Sierra de Urbión 1200 m Soria Spain MW 01407 Zygaena lonicerae 1 1 19.07.2001 Montenegro de Camaros Sierra de Urbión 1200 m Soria Spain MW 01408 Zygaena filipendulae 1 1 19.07.2001 Montenegro de Camaros Sierra de Urbión 1200 m Soria Spain MW 01409 Zygaena lonicerae 1 1 19.07.2001 Montenegro de Camaros Sierra de Urbión 1200 m Soria Spain MW 01410 Zygaena hilaris 1 1 21.07.2001 Triste Sierra de Urbión 600 m Soria Spain MW 01411 Zygaena occitanica 1 1 21.07.2001 Triste Sierra de Urbión 600 m Soria Spain MW 01412 Zygaena occitanica 1 1 21.07.2001 Triste Sierra de Urbión 600 m Soria Spain MW 01413 Zygaena fausta 1 1 21.07.2001 Triste Sierra de Urbión 600 m Soria Spain MW 01414 Zygaena occitanica 1 1 21.07.2001 Triste Sierra de Urbión 600 m Soria Spain MW 01415 Zygaena filipendulae 1 1 24.07.2001 Taradell SO Vic 600 m Barcelona Spain MW 01416 Zygaena filipendulae 1 1 24.07.2001 Taradell SO Vic 600 m Barcelona Spain MW 02001 ♂ Polyommatus amandus 1 1 1 1 1 09.07.2002 Oukaimeden Hoher Atlas 2300m Marrakech Morocco MW 02002 ♂ Polyommatus amandus 1 1 09.07.2002 Oukaimeden Hoher Atlas 2300m Marrakech Morocco MW 02003 ♂ Polyommatus amandus 1 1 09.07.2002 Oukaimeden Hoher Atlas 2300m Marrakech Morocco MW 02004 ♀ Polyommatus amandus 1 1 09.07.2002 Oukaimeden Hoher Atlas 2300m Marrakech Morocco MW 02005 ♀ Polyommatus amandus 1 1 09.07.2002 Oukaimeden Hoher Atlas 2300m Marrakech Morocco MW 02006 ♂ Polyommatus icarus 1 1 1 1 1 09.07.2002 Oukaimeden Hoher Atlas 2300m Marrakech Morocco MW 02007 ♀ Lycaena alciphron 1 1 1 1 09.07.2002 Oukaimeden Hoher Atlas 2300m Marrakech Morocco MW 02008 ♀ Celastrina argiolus 1 1 1 1 09.07.2002 Oukaimeden Hoher Atlas 2300m Marrakech Morocco MW 02009 ♂ Lycaena phlaeas 1 1 1 1 09.07.2002 Oukaimeden Hoher Atlas 2300m Marrakech Morocco MW 02010 ♂ Pararge aegeria 1 1 09.07.2002 Oukaimeden Hoher Atlas 2300m Marrakech Morocco MW 02011 ♂ Hyponephele maroccana 1 1 09.07.2002 Oukaimeden Hoher Atlas 2300m Marrakech Morocco MW 02012 ♂ Gonepteryx cleopatra 1 1 09.07.2002 Oukaimeden Hoher Atlas 2300m Marrakech Morocco MW 02013 ♀ Argynnis pandora 1 1 09.07.2002 Oukaimeden Hoher Atlas 2300m Marrakech Morocco MW 02014 ♂ Aricia artaxerxes 1 1 09.07.2002 Oukaimeden Hoher Atlas 2400m Marrakech Morocco MW 02015 ♂ Pseudochazara atlantis 1 1 09.07.2002 Oukaimeden Hoher Atlas 2800m Marrakech Morocco

- 155 - Appendix 2 List of material

Code Sex Genus Species Wgs Fix Kar Alc DNA Cytb CO1 ND1 ITS2 Fig. Date Locality Area Altitude Province Country MW 02016 ♂ Pseudochazara atlantis 1 1 09.07.2002 Oukaimeden Hoher Atlas 2800m Marrakech Morocco MW 02017 ♂ Coenonympha vaucheri 1 1 12.07.2002 Tizi-n-Tacheddirt Hoher Atlas 3000m Marrakech Morocco MW 02018 ♀ Pieris segonzaki 1 1 12.07.2002 Tizi-n-Tacheddirt Hoher Atlas 3200m Marrakech Morocco MW 02019 ♂ Pontia daplidice 1 1 12.07.2002 Tizi-n-Tacheddirt Hoher Atlas 3200m Marrakech Morocco MW 02020 ♀ Hipparchia aristaeus 1 1 12.07.2002 Tizi-n-Tacheddirt Hoher Atlas 3200m Marrakech Morocco MW 02021 ♂ Zizeeria knysna 1 1 1 1 11.07.2002 10 km N Tizi-n-Test Hoher Atlas 1700m Marrakech Morocco MW 02022 ♂ Berberia lambessanus 1 1 12.07.2002 Tizi-n-Tacheddirt Hoher Atlas 3000m Marrakech Morocco MW 02023 ♂ Berberia lambessanus 1 1 12.07.2002 Tizi-n-Tacheddirt Hoher Atlas 3000m Marrakech Morocco MW 02024 ♀ Iphiclides feisthamelii 1 1 1 1 11.07.2002 10 km N Tizi-n-Test Hoher Atlas 1700m Marrakech Morocco MW 02025 ♂ Tarucus theophrastus 1 1 1 1 1 14.07.2002 Tourchte Hoher Atlas 1400m Marrakech Morocco MW 02026 ♂ Tarucus theophrastus 1 1 14.07.2002 Tourchte Hoher Atlas 1400m Marrakech Morocco MW 02027 ♂ Polyommatus icarus 1 1 14.07.2002 Tourchte Hoher Atlas 1400m Marrakech Morocco MW 02028 ♀ Lampides boeticus 1 1 1 1 14.07.2002 Tourchte Hoher Atlas 1400m Marrakech Morocco MW 02029 ♀ Hyponephele maroccana 1 1 14.07.2002 Tourchte Hoher Atlas 1400m Marrakech Morocco MW 02030 ♀ Hyponephele maroccana 1 1 14.07.2002 Tourchte Hoher Atlas 1400m Marrakech Morocco MW 02031 ♀ Pseudophilotes abencerragus 1 1 1 1 15.07.2002 Oukaimeden Hoher Atlas 2700m Marrakech Morocco MW 02032 ♀ Pseudophilotes abencerragus 1 1 15.07.2002 Oukaimeden Hoher Atlas 2700m Marrakech Morocco MW 02033 ♂ Aricia artaxerxes 1 1 1 1 1 15.07.2002 Oukaimeden Hoher Atlas 2700m Marrakech Morocco MW 02034 ♂ Cyaniris semiargus 1 1 1 1 1 15.07.2002 Oukaimeden Hoher Atlas 2700m Marrakech Morocco MW 02035 ♀ Polyommatus icarus 1 1 15.07.2002 Oukaimeden Hoher Atlas 2700m Marrakech Morocco MW 02036 tripolinus 1 1 15.07.2002 Oukaimeden Hoher Atlas 2700m Marrakech Morocco MW 02037 ♂ Satyrus ferula 1 1 15.07.2002 Oukaimeden Hoher Atlas 2700m Marrakech Morocco MW 02038 Issoria lathonia 1 1 15.07.2002 Oukaimeden Hoher Atlas 2700m Marrakech Morocco MW 02039 ♂ Gonepteryx cleopatra 1 1 15.07.2002 Oukaimeden Hoher Atlas 2700m Marrakech Morocco MW 02040 ♂ Colias crocea 1 1 15.07.2002 Oukaimeden Hoher Atlas 2700m Marrakech Morocco MW 02041 ♂ Thymelicus acteon 1 1 15.07.2002 Oukaimeden Hoher Atlas 2700m Marrakech Morocco MW 02042 ♂ Pieris brassicae 1 1 15.07.2002 Oukaimeden Hoher Atlas 2700m Marrakech Morocco MW 98001 ♂ Polyommatus icarus 1 C 1 1 11.07.1998 Karabayir südl. Korkuteli 1400 m Antalya Turkey MW 98002 ♂ Cupido osiris 1 C 1 1 1 11.07.1998 Karabayir südl. Korkuteli 1400 m Antalya Turkey MW 98003 ♀ Cupido osiris 1 C 1 11.07.1998 Karabayir südl. Korkuteli 1400 m Antalya Turkey MW 98004 ♀ Plebeius loewii 1 C 1 11.07.1998 Karabayir südl. Korkuteli 1400 m Antalya Turkey MW 98005 ♂ Plebeius loewii 1 C 1 11.07.1998 Karabayir südl. Korkuteli 1400 m Antalya Turkey MW 98006 ♂ Lycaena thersamon 1 C 1 1 1 11.07.1998 Karabayir südl. Korkuteli 1400 m Antalya Turkey MW 98007 ♂ Polyommatus thersites 1 C 1 11.07.1998 Karabayir südl. Korkuteli 1400 m Antalya Turkey MW 98008 ♂ Polyommatus thersites 1 C 1 1 1 11.07.1998 Karabayir südl. Korkuteli 1400 m Antalya Turkey MW 98009 ♂ Agrodiaetus carmon 1 C P 1 1 1 1 I-39 11.07.1998 Karabayir südl. Korkuteli 1400 m Antalya Turkey MW 98010 ♂ Agrodiaetus carmon 1 C P 1 11.07.1998 Karabayir südl. Korkuteli 1400 m Antalya Turkey MW 98011 ♂ Meleageria daphnis 1 C 1 11.07.1998 Karabayir südl. Korkuteli 1400 m Antalya Turkey MW 98012 ♂ Meleageria daphnis 1 C 1 11.07.1998 Karabayir südl. Korkuteli 1400 m Antalya Turkey MW 98013 ♂ Meleageria daphnis 1 C 1 11.07.1998 Karabayir südl. Korkuteli 1400 m Antalya Turkey MW 98014 ♀ Satyrium ilicis 1 C 1 12.07.1998 Gülübeli Geçidi westl. Elmali 1500 m Fethiye Turkey MW 98015 ♂ Plebeius loewii 1 C 1 12.07.1998 Gülübeli Geçidi westl. Elmali 1500 m Fethiye Turkey MW 98016 ♂ Plebeius loewii 1 C 1 12.07.1998 Gülübeli Geçidi westl. Elmali 1500 m Fethiye Turkey MW 98017 ♀ Aricia agestis 1 C 1 12.07.1998 Gülübeli Geçidi westl. Elmali 1500 m Fethiye Turkey MW 98018 ♂ Polyommatus icarus 1 C 1 12.07.1998 Gülübeli Geçidi westl. Elmali 1500 m Fethiye Turkey MW 98019 ♂ Agrodiaetus menalcas 1 C 1 1 12.07.1998 Gülübeli Geçidi westl. Elmali 1500 m Fethiye Turkey MW 98020 ♂ Agrodiaetus menalcas 1 C P 1 1 1 12.07.1998 Gülübeli Geçidi westl. Elmali 1500 m Fethiye Turkey MW 98021 ♂ Agrodiaetus menalcas 1 C 1 12.07.1998 Gülübeli Geçidi westl. Elmali 1500 m Fethiye Turkey MW 98022 ♂ Agrodiaetus menalcas 1 C P 1 12.07.1998 Gülübeli Geçidi westl. Elmali 1500 m Fethiye Turkey MW 98023 ♂ Agrodiaetus menalcas 1 C 1 12.07.1998 Gülübeli Geçidi westl. Elmali 1500 m Fethiye Turkey MW 98024 ♂ Agrodiaetus menalcas 1 C 1 12.07.1998 Gülübeli Geçidi westl. Elmali 1500 m Fethiye Turkey MW 98025 ♂ Agrodiaetus menalcas 1 C 1 12.07.1998 Gülübeli Geçidi westl. Elmali 1500 m Fethiye Turkey MW 98026 ♂ Agrodiaetus admetus 1 C P 1 12.07.1998 Gülübeli Geçidi westl. Elmali 1500 m Fethiye Turkey MW 98027 ♂ Agrodiaetus admetus 1 C 1 12.07.1998 Gülübeli Geçidi westl. Elmali 1500 m Fethiye Turkey MW 98028 ♂ Meleageria daphnis 1 C 1 1 12.07.1998 Gülübeli Geçidi westl. Elmali 1500 m Fethiye Turkey MW 98029 ♂ Meleageria daphnis 1 C 1 1 1 1 12.07.1998 Gülübeli Geçidi westl. Elmali 1500 m Fethiye Turkey MW 98030 ♂ Meleageria daphnis 1 C 1 12.07.1998 Gülübeli Geçidi westl. Elmali 1500 m Fethiye Turkey MW 98031 ♂ Meleageria daphnis 1 C 1 12.07.1998 Gülübeli Geçidi westl. Elmali 1500 m Fethiye Turkey MW 98032 ♂ Meleageria daphnis 1 C 1 12.07.1998 Gülübeli Geçidi westl. Elmali 1500 m Fethiye Turkey MW 98033 ♂ Meleageria daphnis 1 C 1 12.07.1998 Gülübeli Geçidi westl. Elmali 1500 m Fethiye Turkey MW 98034 ♂ Meleageria daphnis 1 C 1 12.07.1998 Gülübeli Geçidi westl. Elmali 1500 m Fethiye Turkey MW 98035 ♂ Meleageria daphnis 1 C 1 12.07.1998 Gülübeli Geçidi westl. Elmali 1500 m Fethiye Turkey MW 98036 ♂ Meleageria daphnis 1 C 1 12.07.1998 Gülübeli Geçidi westl. Elmali 1500 m Fethiye Turkey MW 98037 ♂ Meleageria daphnis 1 C 1 12.07.1998 Gülübeli Geçidi westl. Elmali 1500 m Fethiye Turkey MW 98038 ♂ Meleageria daphnis 1 C 1 12.07.1998 Gülübeli Geçidi westl. Elmali 1500 m Fethiye Turkey MW 98039 ♂ Meleageria daphnis 1 C 1 12.07.1998 Gülübeli Geçidi westl. Elmali 1500 m Fethiye Turkey MW 98040 ♂ Meleageria daphnis 1 C P 1 12.07.1998 Gülübeli Geçidi westl. Elmali 1500 m Fethiye Turkey MW 98041 ♂ Meleageria daphnis 1 C P 1 1 12.07.1998 Gülübeli Geçidi westl. Elmali 1500 m Fethiye Turkey MW 98042 ♂ Agrodiaetus menalcas 1 C 1 12.07.1998 Gülübeli Geçidi westl. Elmali 1500 m Fethiye Turkey MW 98043 ♂ Agrodiaetus menalcas 1 C 1 12.07.1998 Gülübeli Geçidi westl. Elmali 1500 m Fethiye Turkey MW 98044 ♂ Agrodiaetus menalcas 1 C 1 12.07.1998 Gülübeli Geçidi westl. Elmali 1500 m Fethiye Turkey MW 98045 ♂ Agrodiaetus menalcas 1 C 1 12.07.1998 Gülübeli Geçidi westl. Elmali 1500 m Fethiye Turkey MW 98046 ♂ Agrodiaetus menalcas 1 C P 1 12.07.1998 Gülübeli Geçidi westl. Elmali 1500 m Fethiye Turkey MW 98047 ♂ Agrodiaetus menalcas 1 C P 1 1 12.07.1998 Gülübeli Geçidi westl. Elmali 1500 m Fethiye Turkey MW 98048 ♀ Meleageria daphnis 1 C 1 12.07.1998 Salur Dagi westl. Elmali 1700-1900 m Fethiye Turkey MW 98049 ♂ Agrodiaetus iphigenia 1 C P 1 1 1 1 12.07.1998 Salur Dagi westl. Elmali 1700-1900 m Fethiye Turkey MW 98050 ♂ Meleageria daphnis 1 C 1 12.07.1998 Salur Dagi westl. Elmali 1700-1900 m Fethiye Turkey MW 98051 ♂ Meleageria daphnis 1 C 1 12.07.1998 Salur Dagi westl. Elmali 1700-1900 m Fethiye Turkey MW 98052 ♂ Plebeius loewii 1 C 1 12.07.1998 Salur Dagi westl. Elmali 1700-1900 m Fethiye Turkey MW 98053 ♂ Lycaena alciphron 1 C 1 12.07.1998 Salur Dagi westl. Elmali 1700-1900 m Fethiye Turkey MW 98054 ♂ Lycaena alciphron 1 C 1 12.07.1998 Salur Dagi westl. Elmali 1700-1900 m Fethiye Turkey MW 98055 ♂ Agrodiaetus admetus 1 C P 1 1 1 13.07.1998 Camkuyuzu Bey Daglari 1750 m Antalya Turkey MW 98056 ♂ Agrodiaetus admetus 1 C 1 13.07.1998 Camkuyuzu Bey Daglari 1750 m Antalya Turkey MW 98057 ♂ Agrodiaetus admetus 1 C P 1 13.07.1998 Camkuyuzu Bey Daglari 1750 m Antalya Turkey MW 98058 ♂ Agrodiaetus admetus 1 C 1 13.07.1998 Camkuyuzu Bey Daglari 1750 m Antalya Turkey MW 98059 ♂ Agrodiaetus admetus 1 C 1 13.07.1998 Camkuyuzu Bey Daglari 1750 m Antalya Turkey MW 98060 ♂ Agrodiaetus admetus 1 C 1 13.07.1998 Camkuyuzu Bey Daglari 1750 m Antalya Turkey MW 98061 ♀ Agrodiaetus admetus 1 1 13.07.1998 Camkuyuzu Bey Daglari 1750 m Antalya Turkey MW 98062 ♂ Agrodiaetus admetus 1 C P 1 13.07.1998 Camkuyuzu Bey Daglari 1750 m Antalya Turkey MW 98063 ♀ Agrodiaetus admetus 1 1 13.07.1998 Camkuyuzu Bey Daglari 1750 m Antalya Turkey MW 98064 ♂ Agrodiaetus menalcas 1 C P 1 13.07.1998 Camkuyuzu Bey Daglari 1750 m Antalya Turkey MW 98065 ♂ Agrodiaetus menalcas 1 C 1 13.07.1998 Camkuyuzu Bey Daglari 1750 m Antalya Turkey MW 98066 ♂ Agrodiaetus menalcas 1 C 1 13.07.1998 Camkuyuzu Bey Daglari 1750 m Antalya Turkey MW 98067 ♂ Agrodiaetus menalcas 1 1 13.07.1998 Camkuyuzu Bey Daglari 1750 m Antalya Turkey MW 98068 ♀ Agrodiaetus menalcas 1 1 13.07.1998 Camkuyuzu Bey Daglari 1750 m Antalya Turkey MW 98069 ♂ Agrodiaetus lycius 1 C P 1 1 15.07.1998 Cukurelma nördl. Elmali 1300 m Antalya Turkey MW 98070 ♂ Agrodiaetus lycius 1 C P 1 15.07.1998 Cukurelma nördl. Elmali 1300 m Antalya Turkey MW 98071 ♂ Agrodiaetus lycius 1 C P 1 15.07.1998 Cukurelma nördl. Elmali 1300 m Antalya Turkey MW 98072 ♂ Agrodiaetus lycius 1 C P 1 15.07.1998 Cukurelma nördl. Elmali 1300 m Antalya Turkey MW 98073 ♀ Agrodiaetus lycius 1 1 15.07.1998 Cukurelma nördl. Elmali 1300 m Antalya Turkey MW 98074 ♂ Agrodiaetus lycius 1 1 15.07.1998 Cukurelma nördl. Elmali 1300 m Antalya Turkey MW 98075 ♂ Agrodiaetus lycius 1 1 15.07.1998 Cukurelma nördl. Elmali 1300 m Antalya Turkey MW 98076 ♂ Agrodiaetus lycius 1 1 15.07.1998 Cukurelma nördl. Elmali 1300 m Antalya Turkey MW 98077 ♂ Agrodiaetus lycius 1 1 15.07.1998 Cukurelma nördl. Elmali 1300 m Antalya Turkey MW 98078 ♂ Agrodiaetus lycius 1 1 15.07.1998 Cukurelma nördl. Elmali 1300 m Antalya Turkey MW 98079 ♂ Agrodiaetus lycius 1 1 1 1 1 I-40 15.07.1998 Cukurelma nördl. Elmali 1300 m Antalya Turkey MW 98080 ♂ Agrodiaetus menalcas 1 C 1 15.07.1998 Cukurelma nördl. Elmali 1300 m Antalya Turkey MW 98081 ♂ Agrodiaetus menalcas 1 1 15.07.1998 Cukurelma nördl. Elmali 1300 m Antalya Turkey MW 98082 ♂ Agrodiaetus admetus 1 C P 1 15.07.1998 Cukurelma nördl. Elmali 1300 m Antalya Turkey MW 98083 ♀ Agrodiaetus admetus 1 C 1 15.07.1998 Cukurelma nördl. Elmali 1300 m Antalya Turkey MW 98084 ♀ Agrodiaetus admetus 1 1 1 1 15.07.1998 Cukurelma nördl. Elmali 1300 m Antalya Turkey MW 98085 ♀ Agrodiaetus admetus 1 1 15.07.1998 Cukurelma nördl. Elmali 1300 m Antalya Turkey MW 98086 ♂ Agrodiaetus admetus 1 1 15.07.1998 Cukurelma nördl. Elmali 1300 m Antalya Turkey MW 98087 ♀ Agrodiaetus admetus 1 1 15.07.1998 Cukurelma nördl. Elmali 1300 m Antalya Turkey MW 98088 ♂ Agrodiaetus admetus 1 1 15.07.1998 Cukurelma nördl. Elmali 1300 m Antalya Turkey

- 156 - Appendix 2 List of material

Code Sex Genus Species Wgs Fix Kar Alc DNA Cytb CO1 ND1 ITS2 Fig. Date Locality Area Altitude Province Country MW 98089 ♂ Agrodiaetus lycius 1 C P 1 1 1 20.07.1998 Güneykent Barla Dagi 1300 m Isparta Turkey MW 98090 ♂ Agrodiaetus lycius 1 C P 1 20.07.1998 Güneykent Barla Dagi 1300 m Isparta Turkey MW 98091 ♀ Agrodiaetus ripartii 1 C 1 20.07.1998 Güneykent Barla Dagi 1300 m Isparta Turkey MW 98092 ♂ Agrodiaetus alcestis 1 C 1 20.07.1998 Güneykent Barla Dagi 1300 m Isparta Turkey MW 98093 ♀ Plebeius argus 1 C 1 20.07.1998 Güneykent Barla Dagi 1300 m Isparta Turkey MW 98094 ♀ Lycaena thersamon 1 C 1 1 1 20.07.1998 Güneykent Barla Dagi 1300 m Isparta Turkey MW 98095 ♂ Lycaena thersamon 1 C 1 20.07.1998 Güneykent Barla Dagi 1300 m Isparta Turkey MW 98096 ♂ Lycaena thetis 1 C 1 20.07.1998 Güneykent Barla Dagi 1300 m Isparta Turkey MW 98097 ♂ Agrodiaetus ernesti 1 C P 1 1 1 1 I-41 21.07.1998 Dedegöl Geçidi westl. Kurucuova 1700 m Isparta Turkey MW 98098 ♂ Agrodiaetus iphicarmon 1 C P 1 21.07.1998 Dedegöl Geçidi westl. Kurucuova 1700 m Isparta Turkey MW 98099 ♂ Polyommatus dorylas 1 C P 1 21.07.1998 Dedegöl Geçidi westl. Kurucuova 1700 m Isparta Turkey MW 98100 ♂ Agrodiaetus ripartii 1 C P 1 1 21.07.1998 Dedegöl Geçidi westl. Kurucuova 1700 m Isparta Turkey MW 98101 ♂ Agrodiaetus iphicarmon 1 C P 1 21.07.1998 Dedegöl Geçidi westl. Kurucuova 1700 m Isparta Turkey MW 98102 ♂ Agrodiaetus iphicarmon 1 C P 1 1 1 21.07.1998 Dedegöl Geçidi westl. Kurucuova 1700 m Isparta Turkey MW 98103 ♂ Agrodiaetus iphicarmon 1 C P 1 1 1 1 I-42 21.07.1998 Dedegöl Geçidi westl. Kurucuova 1700 m Isparta Turkey MW 98104 ♂ Agrodiaetus iphicarmon 1 C P 1 21.07.1998 Dedegöl Geçidi westl. Kurucuova 1700 m Isparta Turkey MW 98105 ♂ Agrodiaetus iphicarmon 1 1 21.07.1998 Dedegöl Geçidi westl. Kurucuova 1700 m Isparta Turkey MW 98106 ♂ Agrodiaetus iphigenia 1 C P 1 1 1 21.07.1998 Dedegöl Geçidi westl. Kurucuova 1700 m Isparta Turkey MW 98107 ♂ Agrodiaetus iphicarmon 1 C P 1 21.07.1998 Dedegöl Geçidi westl. Kurucuova 1700 m Isparta Turkey MW 98108 ♂ Agrodiaetus ripartii 1 C 1 21.07.1998 Dedegöl Geçidi westl. Kurucuova 1700 m Isparta Turkey MW 98109 ♂ Agrodiaetus ripartii 1 1 21.07.1998 Dedegöl Geçidi westl. Kurucuova 1700 m Isparta Turkey MW 98110 ♂ Agrodiaetus ripartii 1 C P 1 21.07.1998 Dedegöl Geçidi westl. Kurucuova 1700 m Isparta Turkey MW 98111 ♂ Agrodiaetus ripartii 1 1 21.07.1998 Dedegöl Geçidi westl. Kurucuova 1700 m Isparta Turkey MW 98112 ♂ Agrodiaetus ripartii 1 1 21.07.1998 Dedegöl Geçidi westl. Kurucuova 1700 m Isparta Turkey MW 98113 ♂ Agrodiaetus iphicarmon 1 1 21.07.1998 Dedegöl Geçidi westl. Kurucuova 1700 m Isparta Turkey MW 98114 ♂ Agrodiaetus iphicarmon 1 1 21.07.1998 Dedegöl Geçidi westl. Kurucuova 1700 m Isparta Turkey MW 98115 ♂ Agrodiaetus iphicarmon 1 1 21.07.1998 Dedegöl Geçidi westl. Kurucuova 1700 m Isparta Turkey MW 98116 ♂ Agrodiaetus iphicarmon 1 1 21.07.1998 Dedegöl Geçidi westl. Kurucuova 1700 m Isparta Turkey MW 98117 ♂ Agrodiaetus iphicarmon 1 1 21.07.1998 Dedegöl Geçidi westl. Kurucuova 1700 m Isparta Turkey MW 98118 ♂ Agrodiaetus iphicarmon 1 1 21.07.1998 Dedegöl Geçidi westl. Kurucuova 1700 m Isparta Turkey MW 98119 ♂ Agrodiaetus iphicarmon 1 1 21.07.1998 Dedegöl Geçidi westl. Kurucuova 1700 m Isparta Turkey MW 98120 ♂ Agrodiaetus ripartii 1 1 21.07.1998 Dedegöl Geçidi westl. Kurucuova 1700 m Isparta Turkey MW 98121 ♂ Agrodiaetus iphicarmon 1 1 21.07.1998 Dedegöl Geçidi westl. Kurucuova 1700 m Isparta Turkey MW 98122 ♂ Agrodiaetus ernesti 1 1 21.07.1998 Dedegöl Geçidi westl. Kurucuova 1700 m Isparta Turkey MW 98123 ♂ Lycaena thetis 1 1 21.07.1998 Dedegöl Geçidi westl. Kurucuova 1700 m Isparta Turkey MW 98124 ♂ Lycaena virgaureae 1 1 21.07.1998 Dedegöl Geçidi westl. Kurucuova 1700 m Isparta Turkey MW 98125 ♂ Lycaena virgaureae 1 1 21.07.1998 Dedegöl Geçidi westl. Kurucuova 1700 m Isparta Turkey MW 98126 ♀ Lycaena phlaeas 1 1 21.07.1998 Dedegöl Geçidi westl. Kurucuova 1700 m Isparta Turkey MW 98127 ♂ Polyommatus dorylas 1 1 21.07.1998 Dedegöl Geçidi westl. Kurucuova 1700 m Isparta Turkey MW 98128 ♂ Polyommatus dorylas 1 1 1 1 21.07.1998 Dedegöl Geçidi westl. Kurucuova 1700 m Isparta Turkey MW 98129 ♂ Lysandra ossmar 1 1 1 1 1 21.07.1998 Dedegöl Geçidi westl. Kurucuova 1700 m Isparta Turkey MW 98130 ♂ Lysandra ossmar 1 1 21.07.1998 Dedegöl Geçidi westl. Kurucuova 1700 m Isparta Turkey MW 98131 ♂ Meleageria daphnis 1 1 21.07.1998 Dedegöl Geçidi westl. Kurucuova 1700 m Isparta Turkey MW 98132 ♂ Meleageria daphnis 1 1 21.07.1998 Dedegöl Geçidi westl. Kurucuova 1700 m Isparta Turkey MW 98133 ♂ Polyommatus icarus 1 1 1 1 21.07.1998 Dedegöl Geçidi westl. Kurucuova 1700 m Isparta Turkey MW 98134 ♂ Polyommatus icarus 1 1 21.07.1998 Dedegöl Geçidi westl. Kurucuova 1700 m Isparta Turkey MW 98135 ♂ Polyommatus icarus 1 1 1 21.07.1998 Dedegöl Geçidi westl. Kurucuova 1700 m Isparta Turkey MW 98136 ♂ Agrodiaetus wagneri 1 C P 1 1 1 1 I-43 22.07.1998 Zelve nördl. Ürgüp 1100 m Nevsehir Turkey MW 98137 ♂ Agrodiaetus poseidon 1 C P 1 1 22.07.1998 Zelve nördl. Ürgüp 1100 m Nevsehir Turkey MW 98138 ♂ Agrodiaetus poseidon 1 C P 1 1 1 1 22.07.1998 Zelve nördl. Ürgüp 1100 m Nevsehir Turkey MW 98139 ♂ Agrodiaetus wagneri 1 C P 1 1 1 1 22.07.1998 Zelve nördl. Ürgüp 1100 m Nevsehir Turkey MW 98140 ♂ Agrodiaetus poseidon 1 C P 1 22.07.1998 Zelve nördl. Ürgüp 1100 m Nevsehir Turkey MW 98141 ♂ Agrodiaetus poseidon 1 C P 1 22.07.1998 Zelve nördl. Ürgüp 1100 m Nevsehir Turkey MW 98142 ♂ Agrodiaetus poseidon 1 C P 1 22.07.1998 Zelve nördl. Ürgüp 1100 m Nevsehir Turkey MW 98143 ♂ Agrodiaetus poseidon 1 C P 1 22.07.1998 Zelve nördl. Ürgüp 1100 m Nevsehir Turkey MW 98144 ♂ Agrodiaetus poseidon 1 C P 1 22.07.1998 Zelve nördl. Ürgüp 1100 m Nevsehir Turkey MW 98145 ♂ Agrodiaetus poseidon 1 C P 1 22.07.1998 Zelve nördl. Ürgüp 1100 m Nevsehir Turkey MW 98146 ♂ Agrodiaetus poseidon 1 C 1 22.07.1998 Zelve nördl. Ürgüp 1100 m Nevsehir Turkey MW 98147 ♂ Agrodiaetus poseidon 1 C 1 22.07.1998 Zelve nördl. Ürgüp 1100 m Nevsehir Turkey MW 98148 ♂ Agrodiaetus poseidon 1 C 1 22.07.1998 Zelve nördl. Ürgüp 1100 m Nevsehir Turkey MW 98149 ♂ Agrodiaetus wagneri 1 C P 1 1 22.07.1998 Zelve nördl. Ürgüp 1100 m Nevsehir Turkey MW 98150 ♂ Agrodiaetus poseidon 1 C 1 22.07.1998 Zelve nördl. Ürgüp 1100 m Nevsehir Turkey MW 98151 ♀ Agrodiaetus wagneri 1 C 1 22.07.1998 Zelve nördl. Ürgüp 1100 m Nevsehir Turkey MW 98152 ♂ Agrodiaetus poseidon 1 C 1 22.07.1998 Zelve nördl. Ürgüp 1100 m Nevsehir Turkey MW 98153 ♂ Agrodiaetus poseidon 1 C 1 22.07.1998 Zelve nördl. Ürgüp 1100 m Nevsehir Turkey MW 98154 ♂ Agrodiaetus poseidon 1 C P 1 1 1 1 1 I-44 22.07.1998 Zelve nördl. Ürgüp 1100 m Nevsehir Turkey MW 98155 ♂ Lysandra ossmar 1 C P 1 22.07.1998 Zelve nördl. Ürgüp 1100 m Nevsehir Turkey MW 98156 ♂ Agrodiaetus iphigenia 1 C 1 25.07.1998 Gökpinar Gürün 1700 m Sivas Turkey MW 98157 ♂ Agrodiaetus iphigenia 1 C 1 25.07.1998 Gökpinar Gürün 1700 m Sivas Turkey MW 98158 ♂ Agrodiaetus actis 1 1 25.07.1998 Gökpinar Gürün 1700 m Sivas Turkey MW 98159 ♂ Agrodiaetus actis 1 C P 1 25.07.1998 Gökpinar Gürün 1700 m Sivas Turkey MW 98160 ♂ Agrodiaetus actis 1 C P 1 1 25.07.1998 Gökpinar Gürün 1700 m Sivas Turkey MW 98161 ♂ Agrodiaetus poseidon 1 1 25.07.1998 Gökpinar Gürün 1700 m Sivas Turkey MW 98162 ♂ Agrodiaetus actis 1 C P 1 1 1 1 I-45 25.07.1998 Gökpinar Gürün 1700 m Sivas Turkey MW 98163 ♂ Agrodiaetus actis 1 1 1 25.07.1998 Gökpinar Gürün 1700 m Sivas Turkey MW 98164 ♂ Agrodiaetus ripartii 1 C P 1 25.07.1998 Gökpinar Gürün 1700 m Sivas Turkey MW 98165 ♀ Agrodiaetus actis 1 1 25.07.1998 Gökpinar Gürün 1700 m Sivas Turkey MW 98166 ♂ Agrodiaetus actis 1 C P 1 25.07.1998 Gökpinar Gürün 1700 m Sivas Turkey MW 98167 ♂ Agrodiaetus poseidon 1 C 1 25.07.1998 Gökpinar Gürün 1700 m Sivas Turkey MW 98168 ♂ Agrodiaetus actis 1 1 25.07.1998 Gökpinar Gürün 1700 m Sivas Turkey MW 98169 ♂ Agrodiaetus poseidon 1 1 25.07.1998 Gökpinar Gürün 1700 m Sivas Turkey MW 98170 ♂ Agrodiaetus maraschi 1 C P 1 1 1 1 I-46 25.07.1998 Gökpinar Gürün 1700 m Sivas Turkey MW 98171 ♂ Agrodiaetus actis 1 C P 1 25.07.1998 Gökpinar Gürün 1700 m Sivas Turkey MW 98172 ♂ Agrodiaetus menalcas 1 C 1 1 1 1 1 I-47 25.07.1998 Gökpinar Gürün 1700 m Sivas Turkey MW 98173 ♂ Agrodiaetus actis 1 1 25.07.1998 Gökpinar Gürün 1700 m Sivas Turkey MW 98174 ♂ Agrodiaetus maraschi 1 C P 1 25.07.1998 Gökpinar Gürün 1700 m Sivas Turkey MW 98175 ♂ Agrodiaetus poseidon 1 C 1 25.07.1998 Gökpinar Gürün 1700 m Sivas Turkey MW 98176 ♂ Agrodiaetus actis 1 1 25.07.1998 Gökpinar Gürün 1700 m Sivas Turkey MW 98177 ♂ Agrodiaetus menalcas 1 1 25.07.1998 Gökpinar Gürün 1700 m Sivas Turkey MW 98178 ♂ Agrodiaetus poseidon 1 C P 1 25.07.1998 Gökpinar Gürün 1700 m Sivas Turkey MW 98179 ♂ Agrodiaetus poseidon 1 1 25.07.1998 Gökpinar Gürün 1700 m Sivas Turkey MW 98180 ♂ Agrodiaetus poseidon 1 C P 1 1 1 1 25.07.1998 Gökpinar Gürün 1700 m Sivas Turkey MW 98181 ♂ Agrodiaetus poseidon 1 1 25.07.1998 Gökpinar Gürün 1700 m Sivas Turkey MW 98182 ♂ Agrodiaetus poseidon 1 1 1 25.07.1998 Gökpinar Gürün 1700 m Sivas Turkey MW 98183 ♂ Agrodiaetus poseidon 1 C P 1 1 1 25.07.1998 Gökpinar Gürün 1700 m Sivas Turkey MW 98184 ♀ Agrodiaetus actis 1 C 1 25.07.1998 Gökpinar Gürün 1700 m Sivas Turkey MW 98185 ♂ Lysandra ossmar 1 C 1 1 1 1 25.07.1998 Gökpinar Gürün 1700 m Sivas Turkey MW 98186 ♂ Pseudophilotes vicrama 1 C 1 25.07.1998 Gökpinar Gürün 1700 m Sivas Turkey MW 98187 ♂ Lysandra ossmar 1 C 1 25.07.1998 Gökpinar Gürün 1700 m Sivas Turkey MW 98188 ♂ Agrodiaetus hopfferi 1 C 1 1 27.07.1998 Gündüzbey Yesilyurt 1300 m Malatya Turkey MW 98189 ♂ Agrodiaetus hopfferi 1 C P 1 1 1 1 I-48 27.07.1998 Gündüzbey Yesilyurt 1300 m Malatya Turkey MW 98190 ♂ Agrodiaetus hopfferi 1 C P 1 27.07.1998 Gündüzbey Yesilyurt 1300 m Malatya Turkey MW 98191 ♂ Agrodiaetus hopfferi 1 C P 1 27.07.1998 Gündüzbey Yesilyurt 1300 m Malatya Turkey MW 98192 ♂ Agrodiaetus hopfferi 1 C 1 27.07.1998 Gündüzbey Yesilyurt 1300 m Malatya Turkey MW 98193 ♂ Agrodiaetus hopfferi 1 C 1 27.07.1998 Gündüzbey Yesilyurt 1300 m Malatya Turkey MW 98194 ♂ Agrodiaetus hopfferi 1 C 1 27.07.1998 Gündüzbey Yesilyurt 1300 m Malatya Turkey MW 98195 ♂ Agrodiaetus hopfferi 1 C P 1 27.07.1998 Gündüzbey Yesilyurt 1300 m Malatya Turkey MW 98196 ♂ Agrodiaetus hopfferi 1 1 27.07.1998 Gündüzbey Yesilyurt 1300 m Malatya Turkey MW 98197 ♂ Agrodiaetus hopfferi 1 1 27.07.1998 Gündüzbey Yesilyurt 1300 m Malatya Turkey MW 98198 ♂ Agrodiaetus hopfferi 1 1 27.07.1998 Gündüzbey Yesilyurt 1300 m Malatya Turkey MW 98199 ♂ Agrodiaetus hopfferi 1 1 27.07.1998 Gündüzbey Yesilyurt 1300 m Malatya Turkey MW 98200 ♂ Agrodiaetus hopfferi 1 1 27.07.1998 Gündüzbey Yesilyurt 1300 m Malatya Turkey MW 98201 ♂ Agrodiaetus hopfferi 1 1 27.07.1998 Gündüzbey Yesilyurt 1300 m Malatya Turkey MW 98202 ♂ Agrodiaetus hopfferi 1 1 1 27.07.1998 Gündüzbey Yesilyurt 1300 m Malatya Turkey MW 98203 ♂ Agrodiaetus mithridates 1 C P 1 1 1 1 I-49 27.07.1998 Gündüzbey Yesilyurt 1300 m Malatya Turkey

- 157 - Appendix 2 List of material

Code Sex Genus Species Wgs Fix Kar Alc DNA Cytb CO1 ND1 ITS2 Fig. Date Locality Area Altitude Province Country MW 98204 ♀ Agrodiaetus hopfferi 1 C 1 27.07.1998 Gündüzbey Yesilyurt 1300 m Malatya Turkey MW 98205 ♂ Agrodiaetus dama 1 1 1 1 27.07.1998 Gündüzbey Yesilyurt 1300 m Malatya Turkey MW 98206 ♀ Agrodiaetus hopfferi 1 C 1 27.07.1998 Gündüzbey Yesilyurt 1300 m Malatya Turkey MW 98207 ♂ Meleageria daphnis 1 C 1 27.07.1998 Gündüzbey Yesilyurt 1300 m Malatya Turkey MW 98208 ♂ Lysandra bellargus 1 C P 1 27.07.1998 Gündüzbey Yesilyurt 1300 m Malatya Turkey MW 98209 ♂ Agrodiaetus hopfferi 1 1 1 27.07.1998 Gündüzbey Yesilyurt 1300 m Malatya Turkey MW 98210 ♂ Agrodiaetus admetus 1 C 1 28.07.1998 Saimbeyli-Fälle Saimbeyli 1500 m Adana Turkey MW 98211 ♂ Agrodiaetus admetus 1 C 1 28.07.1998 Saimbeyli-Fälle Saimbeyli 1500 m Adana Turkey MW 98212 ♂ Agrodiaetus alcestis 1 C P 1 1 1 1 28.07.1998 Saimbeyli-Fälle Saimbeyli 1500 m Adana Turkey MW 98213 ♂ Agrodiaetus alcestis 1 C 1 1 28.07.1998 Saimbeyli-Fälle Saimbeyli 1500 m Adana Turkey MW 98214 ♂ Agrodiaetus alcestis 1 C 1 28.07.1998 Saimbeyli-Fälle Saimbeyli 1500 m Adana Turkey MW 98215 ♂ Agrodiaetus ripartii 1 C P 1 28.07.1998 Saimbeyli-Fälle Saimbeyli 1500 m Adana Turkey MW 98216 ♂ Agrodiaetus alcestis 1 C 1 28.07.1998 Saimbeyli-Fälle Saimbeyli 1500 m Adana Turkey MW 98217 ♀ Agrodiaetus alcestis 1 C 1 28.07.1998 Saimbeyli-Fälle Saimbeyli 1500 m Adana Turkey MW 98218 ♂ Agrodiaetus alcestis 1 C 1 28.07.1998 Saimbeyli-Fälle Saimbeyli 1500 m Adana Turkey MW 98219 ♂ Plebeius loewii 1 C 1 1 28.07.1998 Saimbeyli-Fälle Saimbeyli 1500 m Adana Turkey MW 98220 ♂ Plebeius loewii 1 1 1 1 1 28.07.1998 Saimbeyli-Fälle Saimbeyli 1500 m Adana Turkey MW 98221 ♀ Plebeius loewii 1 1 28.07.1998 Saimbeyli-Fälle Saimbeyli 1500 m Adana Turkey MW 98222 ♂ Polyommatus cornelia 1 C P 1 28.07.1998 Saimbeyli-Fälle Saimbeyli 1500 m Adana Turkey MW 98223 ♂ Polyommatus cornelia 1 1 28.07.1998 Saimbeyli-Fälle Saimbeyli 1500 m Adana Turkey MW 98224 ♂ Polyommatus cornelia 1 1 28.07.1998 Saimbeyli-Fälle Saimbeyli 1500 m Adana Turkey MW 98225 ♂ Polyommatus cornelia 1 C P 1 1 1 28.07.1998 Saimbeyli-Fälle Saimbeyli 1500 m Adana Turkey MW 98226 ♂ Polyommatus cornelia 1 1 28.07.1998 Saimbeyli-Fälle Saimbeyli 1500 m Adana Turkey MW 98227 ♂ Polyommatus thersites 1 C 1 1 28.07.1998 Saimbeyli-Fälle Saimbeyli 1500 m Adana Turkey MW 98228 ♂ Lysandra syriaca 1 C 1 1 1 1 1 28.07.1998 Saimbeyli-Fälle Saimbeyli 1500 m Adana Turkey MW 98229 ♀ Lycaena tityrus 1 C 1 28.07.1998 Saimbeyli-Fälle Saimbeyli 1500 m Adana Turkey MW 98230 ♂ Lycaena tityrus 1 1 1 1 28.07.1998 Saimbeyli-Fälle Saimbeyli 1500 m Adana Turkey MW 98231 ♀ Lycaena tityrus 1 1 28.07.1998 Saimbeyli-Fälle Saimbeyli 1200 m Adana Turkey MW 98232 ♂ Agrodiaetus alcestis 1 C 1 28.07.1998 Saimbeyli-Fälle Saimbeyli 1200 m Adana Turkey MW 98233 ♂ Agrodiaetus alcestis 1 C 1 28.07.1998 Saimbeyli-Fälle Saimbeyli 1200 m Adana Turkey MW 98234 ♂ Agrodiaetus firdussii 1 C P 1 28.07.1998 Saimbeyli-Fälle Saimbeyli 1200 m Adana Turkey MW 98235 ♂ Polyommatus cornelia 1 C 1 1 1 1 28.07.1998 Saimbeyli-Fälle Saimbeyli 1200 m Adana Turkey MW 98236 ♂ Polyommatus cornelia 1 1 28.07.1998 Saimbeyli-Fälle Saimbeyli 1200 m Adana Turkey MW 98237 ♂ Polyommatus cornelia 1 1 28.07.1998 Saimbeyli-Fälle Saimbeyli 1200 m Adana Turkey MW 98238 ♂ Lysandra syriaca 1 C 1 28.07.1998 Saimbeyli-Fälle Saimbeyli 1200 m Adana Turkey MW 98239 ♂ Celastrina argiolus 1 1 28.07.1998 Saimbeyli-Fälle Saimbeyli 1200 m Adana Turkey MW 98240 ♂ Agrodiaetus theresiae 1 C P 1 1 1 1 I-50 29.07.1998 Saimbeyli-Fälle Saimbeyli 1200-1500 m Adana Turkey MW 98241 ♂ Agrodiaetus theresiae 1 C P 1 29.07.1998 Saimbeyli-Fälle Saimbeyli 1200-1500 m Adana Turkey MW 98242 ♂ Agrodiaetus theresiae 1 C P 1 29.07.1998 Saimbeyli-Fälle Saimbeyli 1200-1500 m Adana Turkey MW 98243 ♂ Agrodiaetus theresiae 1 C P 1 29.07.1998 Saimbeyli-Fälle Saimbeyli 1200-1500 m Adana Turkey MW 98244 ♂ Agrodiaetus alcestis 1 C P 1 1 29.07.1998 Saimbeyli-Fälle Saimbeyli 1200-1500 m Adana Turkey MW 98245 ♂ Agrodiaetus admetus 1 C P 1 29.07.1998 Saimbeyli-Fälle Saimbeyli 1200-1500 m Adana Turkey MW 98246 ♂ Lysandra syriaca 1 C P 1 29.07.1998 Saimbeyli-Fälle Saimbeyli 1200-1500 m Adana Turkey MW 98247 ♂ Polyommatus thersites 1 1 1 29.07.1998 Saimbeyli-Fälle Saimbeyli 1200-1500 m Adana Turkey MW 98248 ♂ Polyommatus icarus 1 1 29.07.1998 Saimbeyli-Fälle Saimbeyli 1200-1500 m Adana Turkey MW 98249 ♂ Polyommatus thersites 1 C 1 29.07.1998 Saimbeyli-Fälle Saimbeyli 1200-1500 m Adana Turkey MW 98250 ♂ Polyommatus icarus 1 1 29.07.1998 Saimbeyli-Fälle Saimbeyli 1200-1500 m Adana Turkey MW 98251 ♂ Polyommatus thersites 1 C P 1 29.07.1998 Saimbeyli-Fälle Saimbeyli 1200-1500 m Adana Turkey MW 98252 ♂ Plebeius loewii 1 1 29.07.1998 Saimbeyli-Fälle Saimbeyli 1200-1500 m Adana Turkey MW 98253 ♂ Plebeius loewii 1 1 1 29.07.1998 Saimbeyli-Fälle Saimbeyli 1200-1500 m Adana Turkey MW 98254 ♂ Polyommatus cornelia 1 1 1 29.07.1998 Saimbeyli-Fälle Saimbeyli 1200-1500 m Adana Turkey MW 98255 ♂ Meleageria daphnis 1 C 1 29.07.1998 Saimbeyli-Fälle Saimbeyli 1200-1500 m Adana Turkey MW 98256 ♂ Meleageria daphnis 1 1 29.07.1998 Saimbeyli-Fälle Saimbeyli 1200-1500 m Adana Turkey MW 98257 ♂ Meleageria daphnis 1 1 29.07.1998 Saimbeyli-Fälle Saimbeyli 1200-1500 m Adana Turkey MW 98258 ♂ Meleageria daphnis 1 1 29.07.1998 Saimbeyli-Fälle Saimbeyli 1200-1500 m Adana Turkey MW 98259 ♂ Agrodiaetus iphigenia 1 C 1 30.07.1998 Gezbeli Geçidi 1800 m Kayseri Turkey MW 98260 ♂ Agrodiaetus firdussii 1 C P 1 30.07.1998 Gezbeli Geçidi 1800 m Kayseri Turkey MW 98261 ♂ Agrodiaetus schuriani 1 C P 1 1 1 1 II-1 30.07.1998 Gezbeli Geçidi 1800 m Kayseri Turkey MW 98262 ♀ Agrodiaetus firdussii 1 C 1 30.07.1998 Gezbeli Geçidi 1800 m Kayseri Turkey MW 98263 ♂ Agrodiaetus ripartii 1 C 1 30.07.1998 Gezbeli Geçidi 1800 m Kayseri Turkey MW 98264 ♂ Polyommatus cornelia 1 1 1 1 1 30.07.1998 Gezbeli Geçidi 1800 m Kayseri Turkey MW 98265 ♂ Lampides boeticus 1 1 1 1 30.07.1998 Gezbeli Geçidi 1800 m Kayseri Turkey MW 98266 ♂ Agrodiaetus iphigenia 1 C 1 30.07.1998 Erciyes Dagi Kayakevi 2000 m Kayseri Turkey MW 98267 ♂ Agrodiaetus iphigenia 1 C 1 30.07.1998 Erciyes Dagi Kayakevi 2000 m Kayseri Turkey MW 98268 ♂ Agrodiaetus iphigenia 1 C 1 30.07.1998 Erciyes Dagi Kayakevi 2000 m Kayseri Turkey MW 98269 ♂ Aricia isauricus 1 1 30.07.1998 Erciyes Dagi Kayakevi 2000 m Kayseri Turkey MW 98270 ♂ Aricia anteros 1 C P 1 1 1 1 30.07.1998 Erciyes Dagi Kayakevi 2000 m Kayseri Turkey MW 98271 ♂ Meleageria daphnis 1 C 1 1 - 30.07.1998 Erciyes Dagi Kayakevi 2000 m Kayseri Turkey MW 98272 ♂ Meleageria daphnis 1 C 1 30.07.1998 Erciyes Dagi Kayakevi 2000 m Kayseri Turkey MW 98273 ♂ Meleageria daphnis 1 C 1 30.07.1998 Erciyes Dagi Kayakevi 2000 m Kayseri Turkey MW 98274 ♂ Meleageria daphnis 1 1 1 1 30.07.1998 Erciyes Dagi Kayakevi 2000 m Kayseri Turkey MW 98275 ♂ Meleageria daphnis 1 1 30.07.1998 Erciyes Dagi Kayakevi 2000 m Kayseri Turkey MW 98276 ♂ Meleageria daphnis 1 1 30.07.1998 Erciyes Dagi Kayakevi 2000 m Kayseri Turkey MW 98277 ♂ Meleageria daphnis 1 1 30.07.1998 Erciyes Dagi Kayakevi 2000 m Kayseri Turkey MW 98278 ♂ Lysandra bellargus 1 1 1 1 1 30.07.1998 Ala Daglar Yahyali 1600 m Kayseri Turkey MW 98279 ♂ Agrodiaetus sigberti 1 C P 1 31.07.1998 Ala Daglar Yahyali 2900 m Kayseri Turkey MW 98280 ♂ Polyommatus icarus 1 1 31.07.1998 Ala Daglar Yahyali 2900 m Kayseri Turkey MW 98281 ♂ Polyommatus icarus 1 1 31.07.1998 Ala Daglar Yahyali 2900 m Kayseri Turkey MW 98282 ♀ Agrodiaetus sigberti 1 C 1 31.07.1998 Ala Daglar Yahyali 2700 m Kayseri Turkey MW 98283 ♂ Agrodiaetus sigberti 1 C P 1 1 31.07.1998 Ala Daglar Yahyali 2700 m Kayseri Turkey MW 98284 ♂ Agrodiaetus sigberti 1 C P 1 1 1 II-2 31.07.1998 Ala Daglar Yahyali 2700 m Kayseri Turkey MW 98285 ♂ Agrodiaetus sigberti 1 C P 1 1 1 1 II-3 31.07.1998 Ala Daglar Yahyali 2700 m Kayseri Turkey MW 98286 ♂ Agrodiaetus sigberti 1 C 1 31.07.1998 Ala Daglar Yahyali 2700 m Kayseri Turkey MW 98287 ♂ Agrodiaetus sigberti 1 C 1 31.07.1998 Ala Daglar Yahyali 2700 m Kayseri Turkey MW 98288 ♂ Agrodiaetus sigberti 1 C 1 31.07.1998 Ala Daglar Yahyali 2700 m Kayseri Turkey MW 98289 ♂ Agrodiaetus iphigenia 1 C 1 31.07.1998 Ala Daglar Yahyali 2700 m Kayseri Turkey MW 98290 ♂ Agrodiaetus sigberti 1 C P 1 31.07.1998 Ala Daglar Yahyali 2700 m Kayseri Turkey MW 98291 ♂ Agrodiaetus sigberti 1 1 31.07.1998 Ala Daglar Yahyali 2700 m Kayseri Turkey MW 98292 ♂ Agrodiaetus sigberti 1 1 31.07.1998 Ala Daglar Yahyali 2700 m Kayseri Turkey MW 98293 ♂ Agrodiaetus guezelmavi 1 C P 1 1 04.08.1998 Taskent 1450 m Konya Turkey MW 98294 ♂ Agrodiaetus guezelmavi 1 C P 1 1 1 1 II-4 04.08.1998 Taskent 1450 m Konya Turkey MW 98295 ♂ Agrodiaetus guezelmavi 1 C P 1 04.08.1998 Taskent 1450 m Konya Turkey MW 98296 ♂ Agrodiaetus guezelmavi 1 C P 1 04.08.1998 Taskent 1450 m Konya Turkey MW 98297 ♂ Agrodiaetus guezelmavi 1 C P 1 04.08.1998 Taskent 1450 m Konya Turkey MW 98298 ♂ Agrodiaetus guezelmavi 1 C P 1 04.08.1998 Taskent 1450 m Konya Turkey MW 98299 ♀ Agrodiaetus guezelmavi 1 C P 1 04.08.1998 Taskent 1450 m Konya Turkey MW 98300 ♂ Polyommatus thersites 1 C 1 04.08.1998 Taskent 1450 m Konya Turkey MW 98301 ♀ Agrodiaetus guezelmavi 1 C 1 05.08.1998 Taskent 1450 m Konya Turkey MW 98302 ♀ Agrodiaetus ripartii 1 C 1 05.08.1998 Taskent 1450 m Konya Turkey MW 98303 ♂ Lysandra bellargus 1 1 05.08.1998 Taskent 1450 m Konya Turkey MW 98304 ♂ Agrodiaetus iphigenia 1 1 06.08.1998 Yellibeli Geçidi Ermenek 1800 m Karaman Turkey MW 98305 ♂ Agrodiaetus sertavulensis 1 C P 1 1 06.08.1998 Yellibeli Geçidi Ermenek 1800 m Karaman Turkey MW 98306 ♂ Agrodiaetus sertavulensis 1 C P 1 06.08.1998 Yellibeli Geçidi Ermenek 1800 m Karaman Turkey MW 98307 ♂ Agrodiaetus sertavulensis 1 C P 1 06.08.1998 Yellibeli Geçidi Ermenek 1800 m Karaman Turkey MW 98308 ♂ Agrodiaetus sertavulensis 1 C P 1 06.08.1998 Yellibeli Geçidi Ermenek 1800 m Karaman Turkey MW 98309 ♂ Agrodiaetus sertavulensis 1 C P 1 06.08.1998 Yellibeli Geçidi Ermenek 1800 m Karaman Turkey MW 98310 ♀ Agrodiaetus sertavulensis 1 C 1 06.08.1998 Yellibeli Geçidi Ermenek 1800 m Karaman Turkey MW 98311 ♂ Agrodiaetus iphigenia 1 C P 1 06.08.1998 Yellibeli Geçidi Ermenek 1800 m Karaman Turkey MW 98312 ♂ Agrodiaetus iphigenia 1 C 1 06.08.1998 Yellibeli Geçidi Ermenek 1800 m Karaman Turkey MW 98313 ♂ Agrodiaetus sertavulensis 1 C P 1 1 1 1 II-5 06.08.1998 Yellibeli Geçidi Ermenek 1800 m Karaman Turkey MW 98314 ♀ Agrodiaetus sertavulensis 1 C 1 06.08.1998 Yellibeli Geçidi Ermenek 1800 m Karaman Turkey MW 98315 ♂ Agrodiaetus alcestis 1 C P 1 1 1 1 06.08.1998 Yellibeli Geçidi Ermenek 1800 m Karaman Turkey MW 98316 ♀ Agrodiaetus alcestis 1 C 1 06.08.1998 Yellibeli Geçidi Ermenek 1800 m Karaman Turkey MW 98317 ♂ Meleageria daphnis 1 1 06.08.1998 Yellibeli Geçidi Ermenek 1800 m Karaman Turkey MW 98318 ♂ Meleageria daphnis 1 1 06.08.1998 Yellibeli Geçidi Ermenek 1800 m Karaman Turkey

- 158 - Appendix 2 List of material

Code Sex Genus Species Wgs Fix Kar Alc DNA Cytb CO1 ND1 ITS2 Fig. Date Locality Area Altitude Province Country MW 98319 ♂ Polyommatus cornelia 1 C P 1 06.08.1998 Yellibeli Geçidi Ermenek 1800 m Karaman Turkey MW 98320 ♀ Satyrium spini 1 1 06.08.1998 Yellibeli Geçidi Ermenek 1800 m Karaman Turkey MW 98321 ♂ Lycaena thersamon 1 1 1 06.08.1998 Yellibeli Geçidi Ermenek 1800 m Karaman Turkey MW 98322 ♀ Lycaena thersamon 1 1 06.08.1998 Yellibeli Geçidi Ermenek 1800 m Karaman Turkey MW 99001 ♂ Aricia torulensis 1 C P 1 1 1 1 04.07.1999 Torul 15 km SO Torul 1100 m Gümüshane Turkey MW 99002 ♂ Polyommatus amandus 1 1 04.07.1999 Tersundagi Geç. Siran 2000 m Gümüshane Turkey MW 99003 ♂ Polyommatus amandus 1 1 04.07.1999 Tersundagi Geç. Siran 2000 m Gümüshane Turkey MW 99004 ♂ Agrodiaetus turcicus 1 C P 1 05.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99005 ♂ Agrodiaetus turcicus 1 C P 1 05.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99006 ♂ Agrodiaetus firdussii 1 C P 1 1 1 1 II-6 05.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99007 ♂ Agrodiaetus firdussii 1 C P 1 05.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99008 ♂ Agrodiaetus firdussii 1 C 1 05.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99009 ♂ Agrodiaetus iphigenia 1 C P 1 1 1 1 II-7 05.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99010 ♂ Agrodiaetus firdussii 1 C 1 05.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99011 ♂ Agrodiaetus firdussii 1 C P 1 05.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99012 ♂ Neolysandra coelestina 1 C 1 05.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99013 ♂ Neolysandra coelestina 1 C P 1 1 1 1 05.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99014 ♂ Polyommatus dorylas 1 C 1 1 1 1 1 05.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99015 ♂ Agrıades pyrenaicus 1 C 1 05.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99016 ♂ Agrodiaetus turcicus 1 C 1 05.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99017 ♂ Agrıades pyrenaicus 1 C P 1 05.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99018 ♂ Agrıades pyrenaicus 1 1 1 1 1 05.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99019 ♂ Agrıades pyrenaicus 1 1 05.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99020 ♂ Agrıades pyrenaicus 1 1 05.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99021 ♂ Agrıades pyrenaicus 1 1 05.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99022 ♂ Agrıades pyrenaicus 1 1 05.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99023 ♂ Turanana endymion 1 1 1 05.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99024 ♂ Turanana endymion 1 1 05.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99025 ♂ Plebeius loewii 1 1 05.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99026 ♂ Plebeius loewii 1 1 05.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99027 ♂ Polyommatus icarus 1 1 05.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99028 ♂ Aricia agestis 1 1 1 1 05.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99029 ♂ Polyommatus icarus 1 1 05.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99030 ♂ Polyommatus icarus 1 1 05.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99031 ♂ Agrodiaetus hopfferi 1 C 1 06.07.1999 Çaglayan 5 km O Çaglayan 1500 m Erzincan Turkey MW 99032 ♀ Agrodiaetus ripartii 1 C 1 06.07.1999 Çaglayan 5 km O Çaglayan 1500 m Erzincan Turkey MW 99033 ♂ Agrodiaetus ripartii 1 C P 1 06.07.1999 Çaglayan 5 km O Çaglayan 1500 m Erzincan Turkey MW 99034 ♀ Agrodiaetus ripartii 1 C 1 06.07.1999 Çaglayan 5 km O Çaglayan 1500 m Erzincan Turkey MW 99035 ♀ Agrodiaetus ripartii 1 C 1 06.07.1999 Çaglayan 5 km O Çaglayan 1500 m Erzincan Turkey MW 99036 ♂ Agrodiaetus admetus 1 C 1 06.07.1999 Çaglayan 5 km O Çaglayan 1500 m Erzincan Turkey MW 99037 ♂ Agrodiaetus admetus 1 C 1 06.07.1999 Çaglayan 5 km O Çaglayan 1500 m Erzincan Turkey MW 99038 ♂ Polyommatus cornelia 1 1 1 1 1 06.07.1999 Çaglayan 5 km O Çaglayan 1500 m Erzincan Turkey MW 99039 ♂ Lysandra bellargus 1 1 06.07.1999 Çaglayan 5 km O Çaglayan 1500 m Erzincan Turkey MW 99040 ♂ Polyommatus myrrhinus 1 1 07.07.1999 Köskköy 25 km N Erzurum 1900 m Erzurum Turkey MW 99041 ♂ Polyommatus myrrhinus 1 1 07.07.1999 Köskköy 25 km N Erzurum 1900 m Erzurum Turkey MW 99042 ♂ Lysandra corydonius 1 1 1 1 07.07.1999 Köskköy 25 km N Erzurum 1900 m Erzurum Turkey MW 99043 ♂ Lysandra corydonius 1 1 07.07.1999 Köskköy 25 km N Erzurum 1900 m Erzurum Turkey MW 99044 ♂ Plebeius loewii 1 1 07.07.1999 Köskköy 25 km N Erzurum 1900 m Erzurum Turkey MW 99045 ♀ Maculinea arion 1 1 1 1 1 07.07.1999 Köskköy 25 km N Erzurum 1900 m Erzurum Turkey MW 99046 ♀ Kretania eurypilus 1 1 07.07.1999 Köskköy 25 km N Erzurum 1900 m Erzurum Turkey MW 99047 ♂ Polyommatus amandus 1 1 1 1 1 07.07.1999 Köskköy 25 km N Erzurum 1900 m Erzurum Turkey MW 99048 ♀ Polyommatus amandus 1 1 07.07.1999 Köskköy 25 km N Erzurum 1900 m Erzurum Turkey MW 99049 ♂ Polyommatus amandus 1 1 07.07.1999 Köskköy 25 km N Erzurum 1900 m Erzurum Turkey MW 99050 ♂ Polyommatus amandus 1 1 07.07.1999 Köskköy 25 km N Erzurum 1900 m Erzurum Turkey MW 99051 ♂ Polyommatus amandus 1 1 07.07.1999 Köskköy 25 km N Erzurum 1900 m Erzurum Turkey MW 99052 ♂ Polyommatus amandus 1 1 07.07.1999 Köskköy 25 km N Erzurum 1900 m Erzurum Turkey MW 99053 ♂ Agrodiaetus huberti 1 C P 1 1 1 08.07.1999 Kiliçkaya Yusufeli 1350 m Artvin Turkey MW 99054 ♂ Agrodiaetus huberti 1 C P 1 08.07.1999 Kiliçkaya Yusufeli 1350 m Artvin Turkey MW 99055 ♂ Agrodiaetus huberti 1 C P 1 08.07.1999 Kiliçkaya Yusufeli 1350 m Artvin Turkey MW 99056 ♂ Agrodiaetus huberti 1 C P 1 08.07.1999 Kiliçkaya Yusufeli 1350 m Artvin Turkey MW 99057 ♂ Agrodiaetus merhaba 1 C P 1 1 1 1 II-8 08.07.1999 Kiliçkaya Yusufeli 1350 m Artvin Turkey MW 99058 ♂ Agrodiaetus artvinensis 1 C P 1 1 1 1 II-9 08.07.1999 Kiliçkaya Yusufeli 1350 m Artvin Turkey MW 99059 ♂ Agrodiaetus merhaba 1 C P 1 1 1 08.07.1999 Kiliçkaya Yusufeli 1350 m Artvin Turkey MW 99060 ♂ Agrodiaetus carmon 1 C P 1 1 1 08.07.1999 Kiliçkaya Yusufeli 1350 m Artvin Turkey MW 99061 ♂ Agrodiaetus putnami 1 C P 1 1 1 08.07.1999 Kiliçkaya Yusufeli 1350 m Artvin Turkey MW 99062 ♂ Agrodiaetus iphigenia 1 C 1 08.07.1999 Kiliçkaya Yusufeli 1350 m Artvin Turkey MW 99063 ♂ Agrodiaetus artvinensis 1 C P 1 08.07.1999 Kiliçkaya Yusufeli 1350 m Artvin Turkey MW 99064 ♂ Agrodiaetus artvinensis 1 C P 1 08.07.1999 Kiliçkaya Yusufeli 1350 m Artvin Turkey MW 99065 ♂ Agrodiaetus artvinensis 1 C P 1 08.07.1999 Kiliçkaya Yusufeli 1350 m Artvin Turkey MW 99066 ♂ Agrodiaetus artvinensis 1 C P 1 08.07.1999 Kiliçkaya Yusufeli 1350 m Artvin Turkey MW 99067 ♂ Agrodiaetus ninae 1 C P 1 08.07.1999 Kiliçkaya Yusufeli 1350 m Artvin Turkey MW 99068 ♂ Agrodiaetus ripartii 1 C P 1 1 1 1 08.07.1999 Kiliçkaya Yusufeli 1350 m Artvin Turkey MW 99069 ♂ Agrodiaetus iphigenia 1 C 1 08.07.1999 Kiliçkaya Yusufeli 1350 m Artvin Turkey MW 99070 ♂ Agrodiaetus ripartii 1 C 1 08.07.1999 Kiliçkaya Yusufeli 1350 m Artvin Turkey MW 99071 ♂ Agrodiaetus ripartii 1 C 1 08.07.1999 Kiliçkaya Yusufeli 1350 m Artvin Turkey MW 99072 ♂ Polyommatus thersites 1 C 1 08.07.1999 Kiliçkaya Yusufeli 1350 m Artvin Turkey MW 99073 ♂ Polyommatus icarus 1 1 08.07.1999 Kiliçkaya Yusufeli 1350 m Artvin Turkey MW 99074 ♀ Polyommatus icarus 1 1 08.07.1999 Kiliçkaya Yusufeli 1350 m Artvin Turkey MW 99075 ♂ Polyommatus icarus 1 1 08.07.1999 Kiliçkaya Yusufeli 1350 m Artvin Turkey MW 99076 ♂ Lysandra corydonius 1 1 08.07.1999 Kiliçkaya Yusufeli 1350 m Artvin Turkey MW 99077 ♂ Plebeius loewii 1 1 08.07.1999 Kiliçkaya Yusufeli 1350 m Artvin Turkey MW 99078 ♂ Meleageria daphnis 1 1 08.07.1999 Kiliçkaya Yusufeli 1350 m Artvin Turkey MW 99079 ♂ Meleageria daphnis 1 1 08.07.1999 Kiliçkaya Yusufeli 1350 m Artvin Turkey MW 99080 ♂ Pseudophilotes vicrama 1 1 1 1 08.07.1999 Kiliçkaya Yusufeli 1350 m Artvin Turkey MW 99081 ♂ Pseudophilotes vicrama 1 1 08.07.1999 Kiliçkaya Yusufeli 1350 m Artvin Turkey MW 99082 ♂ Pseudophilotes vicrama 1 1 08.07.1999 Kiliçkaya Yusufeli 1350 m Artvin Turkey MW 99083 ♂ Celastrina argiolus 1 1 08.07.1999 Kiliçkaya Yusufeli 1350 m Artvin Turkey MW 99084 ♂ Celastrina argiolus 1 1 1 1 08.07.1999 Kiliçkaya Yusufeli 1350 m Artvin Turkey MW 99085 ♂ Celastrina argiolus 1 1 08.07.1999 Kiliçkaya Yusufeli 1350 m Artvin Turkey MW 99086 ♂ Cupido osiris 1 1 08.07.1999 Kiliçkaya Yusufeli 1350 m Artvin Turkey MW 99087 ♂ Lycaena alciphron 1 1 08.07.1999 Kiliçkaya Yusufeli 1350 m Artvin Turkey MW 99088 ♂ Agrodiaetus iphigenia 1 C 1 09.07.1999 Demirkent 20 km O Demirkent 1600 m Artvin Turkey MW 99089 ♂ Agrodiaetus iphigenia 1 C 1 09.07.1999 Demirkent 20 km O Demirkent 1600 m Artvin Turkey MW 99090 ♂ Polyommatus dorylas 1 1 09.07.1999 Demirkent 20 km O Demirkent 1600 m Artvin Turkey MW 99091 ♂ Polyommatus eros 1 1 09.07.1999 Demirkent 20 km O Demirkent 1600 m Artvin Turkey MW 99092 ♂ Polyommatus eros 1 1 09.07.1999 Demirkent 20 km O Demirkent 1600 m Artvin Turkey MW 99093 Satyrium ilicis 1 1 09.07.1999 Demirkent 20 km O Demirkent 1600 m Artvin Turkey MW 99094 ♂ Agrodiaetus cyaneus 1 C P 1 1 1 1 11.07.1999 Kagizman 30 km SW Kagizman 1400 m Kars Turkey MW 99095 ♂ Agrodiaetus huberti 1 C P 1 1 1 1 II-10 11.07.1999 Kagizman 30 km SW Kagizman 1400 m Kars Turkey MW 99096 ♀ Agrodiaetus cyaneus 1 C 1 11.07.1999 Kagizman 30 km SW Kagizman 1400 m Kars Turkey MW 99097 ♂ Aricia isauricus 1 C 1 1 1 1 11.07.1999 Kagizman 30 km SW Kagizman 1400 m Kars Turkey MW 99098 ♂ Aricia isauricus 1 C 1 11.07.1999 Kagizman 30 km SW Kagizman 1400 m Kars Turkey MW 99099 ♂ Aricia isauricus 1 C P 1 11.07.1999 Kagizman 30 km SW Kagizman 1400 m Kars Turkey MW 99100 ♂ Agrodiaetus cyaneus 1 C P 1 11.07.1999 Kagizman 30 km SW Kagizman 1400 m Kars Turkey MW 99101 ♂ Agrodiaetus demavendi 1 1 11.07.1999 Akçay Kagizman 1200 m Kars Turkey MW 99102 ♂ Agrodiaetus demavendi 1 C P 1 1 1 11.07.1999 Akçay Kagizman 1200 m Kars Turkey MW 99103 ♂ Agrodiaetus demavendi 1 C P 1 11.07.1999 Akçay Kagizman 1200 m Kars Turkey MW 99104 ♂ Agrodiaetus demavendi 1 C P 1 1 1 11.07.1999 Akçay Kagizman 1200 m Kars Turkey MW 99105 ♂ Agrodiaetus demavendi 1 C P 1 1 1 1 1 II-11 11.07.1999 Akçay Kagizman 1200 m Kars Turkey MW 99106 ♂ Agrodiaetus demavendi 1 C 1 11.07.1999 Akçay Kagizman 1200 m Kars Turkey MW 99107 ♂ Agrodiaetus demavendi 1 C 1 11.07.1999 Akçay Kagizman 1200 m Kars Turkey MW 99108 ♂ Agrodiaetus demavendi 1 C 1 11.07.1999 Akçay Kagizman 1200 m Kars Turkey MW 99109 ♂ Agrodiaetus demavendi 1 C 1 11.07.1999 Akçay Kagizman 1200 m Kars Turkey MW 99110 ♂ Agrodiaetus demavendi 1 C P 1 11.07.1999 Akçay Kagizman 1200 m Kars Turkey MW 99111 ♂ Agrodiaetus demavendi 1 C 1 11.07.1999 Akçay Kagizman 1200 m Kars Turkey

- 159 - Appendix 2 List of material

Code Sex Genus Species Wgs Fix Kar Alc DNA Cytb CO1 ND1 ITS2 Fig. Date Locality Area Altitude Province Country MW 99112 ♂ Agrodiaetus demavendi 1 C 1 11.07.1999 Akçay Kagizman 1200 m Kars Turkey MW 99113 ♂ Agrodiaetus demavendi 1 C 1 11.07.1999 Akçay Kagizman 1200 m Kars Turkey MW 99114 ♀ Agrodiaetus demavendi 1 1 11.07.1999 Akçay Kagizman 1200 m Kars Turkey MW 99115 ♀ Agrodiaetus demavendi 1 1 11.07.1999 Akçay Kagizman 1200 m Kars Turkey MW 99116 ♀ Agrodiaetus demavendi 1 1 11.07.1999 Akçay Kagizman 1200 m Kars Turkey MW 99117 ♀ Agrodiaetus demavendi 1 1 11.07.1999 Akçay Kagizman 1200 m Kars Turkey MW 99118 ♀ Agrodiaetus demavendi 1 1 11.07.1999 Akçay Kagizman 1200 m Kars Turkey MW 99119 ♀ Agrodiaetus demavendi 1 1 11.07.1999 Akçay Kagizman 1200 m Kars Turkey MW 99120 ♀ Agrodiaetus demavendi 1 1 11.07.1999 Akçay Kagizman 1200 m Kars Turkey MW 99121 ♀ Agrodiaetus demavendi 1 1 11.07.1999 Akçay Kagizman 1200 m Kars Turkey MW 99122 ♀ Agrodiaetus demavendi 1 1 11.07.1999 Akçay Kagizman 1200 m Kars Turkey MW 99123 ♀ Agrodiaetus demavendi 1 1 11.07.1999 Akçay Kagizman 1200 m Kars Turkey MW 99124 ♀ Plebeius christophi 1 1 1 1 11.07.1999 Akçay Kagizman 1200 m Kars Turkey MW 99125 ♂ Cupido osiris 1 1 11.07.1999 Akçay Kagizman 1200 m Kars Turkey MW 99126 ♂ Polyommatus thersites 1 1 11.07.1999 Akçay Kagizman 1200 m Kars Turkey MW 99127 ♂ Polyommatus thersites 1 1 11.07.1999 Akçay Kagizman 1200 m Kars Turkey MW 99128 ♂ Polyommatus thersites 1 1 1 1 11.07.1999 Akçay Kagizman 1200 m Kars Turkey MW 99129 ♂ Polyommatus icarus 1 1 11.07.1999 Akçay Kagizman 1200 m Kars Turkey MW 99130 ♂ Agrodiaetus huberti 1 C P 1 12.07.1999 Badilli Tuzluca 1800-2000 m Igdir Turkey MW 99131 ♂ Agrodiaetus huberti 1 C P 1 12.07.1999 Badilli Tuzluca 1800-2000 m Igdir Turkey MW 99132 ♂ Agrodiaetus huberti 1 C P 1 12.07.1999 Badilli Tuzluca 1800-2000 m Igdir Turkey MW 99133 ♀ Agrodiaetus huberti 1 C 1 12.07.1999 Badilli Tuzluca 1800-2000 m Igdir Turkey MW 99134 ♀ Aricia eumedon 1 1 1 1 1 12.07.1999 Badilli Tuzluca 1800-2000 m Igdir Turkey MW 99135 ♂ Agrodiaetus turcicus 1 C P 1 1 1 1 1 12.07.1999 Badilli Tuzluca 1800-2000 m Igdir Turkey MW 99136 ♂ Agrodiaetus firdussii 1 C P 1 12.07.1999 Badilli Tuzluca 1800-2000 m Igdir Turkey MW 99137 ♂ Agrodiaetus firdussii 1 C 1 12.07.1999 Badilli Tuzluca 1800-2000 m Igdir Turkey MW 99138 ♂ Agrodiaetus baytopi 1 C P 1 12.07.1999 Badilli Tuzluca 1800-2000 m Igdir Turkey MW 99139 ♂ Polyommatus dorylas 1 1 12.07.1999 Badilli Tuzluca 1800-2000 m Igdir Turkey MW 99140 ♂ Lysandra corydonius 1 1 1 1 12.07.1999 Badilli Tuzluca 1800-2000 m Igdir Turkey MW 99141 ♂ Agrodiaetus demavendi 1 C P 1 1 1 13.07.1999 Akçay Kagizman 1200 m Kars Turkey MW 99142 ♂ Agrodiaetus demavendi 1 C P 1 13.07.1999 Akçay Kagizman 1200 m Kars Turkey MW 99143 ♂ Agrodiaetus demavendi 1 C 1 13.07.1999 Akçay Kagizman 1200 m Kars Turkey MW 99144 ♂ Agrodiaetus demavendi 1 C P 1 13.07.1999 Akçay Kagizman 1200 m Kars Turkey MW 99145 ♂ Agrodiaetus demavendi 1 C 1 13.07.1999 Akçay Kagizman 1200 m Kars Turkey MW 99146 ♂ Agrodiaetus demavendi 1 C 1 13.07.1999 Akçay Kagizman 1200 m Kars Turkey MW 99147 ♂ Agrodiaetus demavendi 1 C 1 13.07.1999 Akçay Kagizman 1200 m Kars Turkey MW 99148 ♂ Agrodiaetus demavendi 1 C P 1 13.07.1999 Akçay Kagizman 1200 m Kars Turkey MW 99149 ♂ Agrodiaetus demavendi 1 C 1 13.07.1999 Akçay Kagizman 1200 m Kars Turkey MW 99150 ♂ Agrodiaetus demavendi 1 C P 1 13.07.1999 Akçay Kagizman 1200 m Kars Turkey MW 99151 ♂ Agrodiaetus demavendi 1 C P 1 13.07.1999 Akçay Kagizman 1200 m Kars Turkey MW 99152 ♂ Agrodiaetus demavendi 1 C P 1 13.07.1999 Akçay Kagizman 1200 m Kars Turkey MW 99153 ♂ Agrodiaetus iphigenia 1 C 1 13.07.1999 Kagizman 30 km SW Kagizman 1400 m Kars Turkey MW 99154 ♂ Agrodiaetus iphigenia 1 C P 1 13.07.1999 Kagizman 30 km SW Kagizman 1400 m Kars Turkey MW 99155 ♂ Agrodiaetus huberti 1 C P 1 13.07.1999 Kagizman 30 km SW Kagizman 1400 m Kars Turkey MW 99156 ♂ Agrodiaetus huberti 1 C P 1 13.07.1999 Kagizman 30 km SW Kagizman 1400 m Kars Turkey MW 99157 ♀ Agrodiaetus huberti 1 1 13.07.1999 Kagizman 30 km SW Kagizman 1400 m Kars Turkey MW 99158 ♂ Satyrium hyrcanicum 1 1 1 1 13.07.1999 Kagizman 30 km SW Kagizman 1400 m Kars Turkey MW 99159 ♂ Satyrium hyrcanicum 1 1 13.07.1999 Kagizman 30 km SW Kagizman 1400 m Kars Turkey MW 99160 ♀ Satyrium hyrcanicum 1 1 13.07.1999 Kagizman 30 km SW Kagizman 1400 m Kars Turkey MW 99161 ♂ Agrodiaetus turcicus 1 C P 1 14.07.1999 Köskköy 25 km N Erzurum 1900 m Erzurum Turkey MW 99162 ♂ Agrodiaetus turcicus 1 C 1 14.07.1999 Köskköy 25 km N Erzurum 1900 m Erzurum Turkey MW 99163 ♂ Plebeius pylaon 1 C 1 1 1 1 14.07.1999 Köskköy 25 km N Erzurum 1900 m Erzurum Turkey MW 99164 ♂ Agrodiaetus interjectus 1 C P 1 1 1 1 1 II-12 14.07.1999 Çiftlik 5 km NO Çiftlik 1900 m Erzurum Turkey MW 99165 ♂ Agrodiaetus altivagans 1 C P 1 1 1 15.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99166 ♂ Agrodiaetus firdussii 1 C 1 15.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99167 ♂ Agrodiaetus altivagans 1 C P 1 15.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99168 ♂ Agrodiaetus damon 1 C 1 15.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99169 ♂ Agrodiaetus huberti 1 C P 1 15.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99170 ♂ Agrodiaetus iphigenia 1 C P 1 1 1 15.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99171 ♂ Agrodiaetus firdussii 1 C 1 15.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99172 ♂ Agrodiaetus firdussii 1 C P 1 15.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99173 ♂ Agrodiaetus huberti 1 C P 1 15.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99174 ♂ Agrodiaetus phyllis 1 C P 1 1 1 1 15.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99175 ♂ Agrodiaetus phyllis 1 C 1 15.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99176 ♂ Agrodiaetus iphigenia 1 C 1 15.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99177 ♂ Agrodiaetus turcicus 1 C 1 15.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99178 ♂ Agrodiaetus phyllis 1 C 1 15.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99179 ♂ Agrodiaetus huberti 1 C P 1 15.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99180 ♂ Agrodiaetus phyllis 1 C P 1 15.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99181 ♂ Agrodiaetus iphigenia 1 C 1 15.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99182 ♂ Agrodiaetus damon 1 C 1 15.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99183 ♂ Agrodiaetus phyllis 1 C 1 15.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99184 ♂ Agrodiaetus turcicus 1 C 1 15.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99185 ♂ Agrodiaetus phyllis 1 C 1 15.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99186 ♂ Agrodiaetus firdussii 1 C 1 15.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99187 ♂ Agrodiaetus phyllis 1 C 1 1 1 15.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99188 ♂ Agrodiaetus phyllis 1 C 1 15.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99189 ♂ Agrodiaetus turcicus 1 C P 1 15.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99190 ♂ Agrodiaetus turcicus 1 C 1 15.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99191 ♂ Agrodiaetus phyllis 1 C 1 15.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99192 ♂ Agrodiaetus turcicus 1 C 1 15.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99193 ♂ Agrodiaetus phyllis 1 C 1 15.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99194 ♂ Agrodiaetus iphigenia 1 C P 1 15.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99195 ♂ Agrodiaetus phyllis 1 C 1 15.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99196 ♂ Agrodiaetus ripartii 1 C P 1 1 1 1 II-13 15.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99197 ♂ Agrodiaetus phyllis 1 C 1 15.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99198 ♂ Agrodiaetus phyllis 1 C 1 15.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99199 ♂ Agrodiaetus phyllis 1 C 1 15.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99200 ♂ Agrodiaetus phyllis 1 C 1 15.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99201 ♂ Agrodiaetus phyllis 1 C 1 15.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99202 ♂ Agrodiaetus turcicus 1 C 1 15.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99203 ♂ Agrodiaetus turcicus 1 C P 1 1 1 1 II-14 15.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99204 ♂ Agrodiaetus firdussii 1 C 1 15.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99205 ♂ Agrodiaetus phyllis 1 C P 1 15.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99206 ♂ Agrodiaetus ripartii 1 C 1 15.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99207 ♂ Agrodiaetus ripartii 1 C P 1 15.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99208 ♂ Agrodiaetus ripartii 1 C 1 15.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99209 ♂ Agrodiaetus ripartii 1 C 1 15.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99210 ♂ Agrodiaetus ripartii 1 C 1 15.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99211 ♂ Agrodiaetus ripartii 1 C 1 15.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99212 ♂ Agrodiaetus ripartii 1 C P 1 15.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99213 ♂ Agrodiaetus ripartii 1 C 1 15.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99214 ♂ Agrodiaetus ripartii 1 C 1 15.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99215 ♂ Agrodiaetus ripartii 1 C 1 15.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99216 ♂ Agrodiaetus ripartii 1 C 1 15.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99217 ♂ Agrodiaetus ripartii 1 C P 1 15.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99218 ♂ Agrodiaetus ripartii 1 C P 1 15.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99219 ♂ Agrodiaetus ripartii 1 C P 1 15.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99220 ♂ Agrodiaetus ripartii 1 C P 1 15.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99221 ♂ Turanana endymion 1 1 1 1 15.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99222 ♂ Turanana endymion 1 1 15.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99223 ♂ Neolysandra coelestina 1 1 15.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99224 ♂ Neolysandra coelestina 1 1 15.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99225 ♂ Polyommatus dorylas 1 1 15.07.1999 Çaglayan 5 km SO Çaglayan 1500 m Erzincan Turkey MW 99226 ♂ Agrodiaetus zapvadi 1 C P 1 1 1 1 17.07.1999 Zernek Brj. Güzelsu 1900 m Van Turkey

- 160 - Appendix 2 List of material

Code Sex Genus Species Wgs Fix Kar Alc DNA Cytb CO1 ND1 ITS2 Fig. Date Locality Area Altitude Province Country MW 99227 ♂ Agrodiaetus zapvadi 1 C P 1 17.07.1999 Zernek Brj. Güzelsu 1900 m Van Turkey MW 99228 ♂ Agrodiaetus zapvadi 1 C P 1 17.07.1999 Zernek Brj. Güzelsu 1900 m Van Turkey MW 99229 ♂ Agrodiaetus zapvadi 1 C P 1 17.07.1999 Zernek Brj. Güzelsu 1900 m Van Turkey MW 99230 ♀ Agrodiaetus zapvadi 1 C 1 17.07.1999 Zernek Brj. Güzelsu 1900 m Van Turkey MW 99231 ♀ Agrodiaetus zapvadi 1 C 1 17.07.1999 Zernek Brj. Güzelsu 1900 m Van Turkey MW 99232 ♂ Agrodiaetus cyaneus 1 C P 1 17.07.1999 Zernek Brj. Güzelsu 1900 m Van Turkey MW 99233 ♂ Agrodiaetus cyaneus 1 C P 1 17.07.1999 Zernek Brj. Güzelsu 1900 m Van Turkey MW 99234 ♂ Agrodiaetus cyaneus 1 C P 1 17.07.1999 Zernek Brj. Güzelsu 1900 m Van Turkey MW 99235 ♂ Agrodiaetus turcicola 1 C P 1 17.07.1999 Zernek Brj. Güzelsu 1900 m Van Turkey MW 99236 ♂ Agrodiaetus turcicola 1 C P 1 17.07.1999 Güzeldere Geç. Baskale 2500 m Van Turkey MW 99237 ♂ Agrodiaetus altivagans 1 C P 1 17.07.1999 Güzeldere Geç. Baskale 2500 m Van Turkey MW 99238 ♂ Agrodiaetus firdussii 1 C 1 17.07.1999 Güzeldere Geç. Baskale 2500 m Van Turkey MW 99239 ♂ Agrodiaetus altivagans 1 C 1 17.07.1999 Güzeldere Geç. Baskale 2500 m Van Turkey MW 99240 ♂ Agrodiaetus altivagans 1 C P 1 1 1 1 II-15 17.07.1999 Güzeldere Geç. Baskale 2500 m Van Turkey MW 99241 ♂ Agrodiaetus firdussii 1 C P 1 17.07.1999 Güzeldere Geç. Baskale 2500 m Van Turkey MW 99242 ♂ Agrodiaetus huberti 1 C P 1 17.07.1999 Güzeldere Geç. Baskale 2500 m Van Turkey MW 99243 ♂ Agrodiaetus firdussii 1 C 1 17.07.1999 Güzeldere Geç. Baskale 2500 m Van Turkey MW 99244 ♂ Agrodiaetus phyllis 1 C 1 1 17.07.1999 Güzeldere Geç. Baskale 2500 m Van Turkey MW 99245 ♂ Agrodiaetus phyllis 1 C 1 17.07.1999 Güzeldere Geç. Baskale 2500 m Van Turkey MW 99246 ♂ Agrodiaetus firdussii 1 C 1 17.07.1999 Güzeldere Geç. Baskale 2500 m Van Turkey MW 99247 ♂ Agrodiaetus firdussii 1 C P 1 1 1 1 17.07.1999 Güzeldere Geç. Baskale 2500 m Van Turkey MW 99248 ♂ Agrodiaetus phyllis 1 C 1 17.07.1999 Güzeldere Geç. Baskale 2500 m Van Turkey MW 99249 ♂ Agrodiaetus phyllis 1 C P 1 17.07.1999 Güzeldere Geç. Baskale 2500 m Van Turkey MW 99250 ♂ Agrodiaetus cyaneus 1 C P 1 17.07.1999 Güzeldere Geç. Baskale 2500 m Van Turkey MW 99251 ♂ Agrodiaetus phyllis 1 C 1 17.07.1999 Güzeldere Geç. Baskale 2500 m Van Turkey MW 99252 ♂ Agrodiaetus altivagans 1 C P 1 17.07.1999 Güzeldere Geç. Baskale 2500 m Van Turkey MW 99253 ♂ Agrodiaetus altivagans 1 C P 1 17.07.1999 Güzeldere Geç. Baskale 2500 m Van Turkey MW 99254 ♂ Agrodiaetus turcicola 1 C P 1 17.07.1999 Güzeldere Geç. Baskale 2500 m Van Turkey MW 99255 ♀ Agrodiaetus ? 1 C 1 17.07.1999 Güzeldere Geç. Baskale 2500 m Van Turkey MW 99256 ♀ Agrodiaetus ? 1 C 1 17.07.1999 Güzeldere Geç. Baskale 2500 m Van Turkey MW 99257 ♂ Polyommatus icarus 1 C 1 17.07.1999 Güzeldere Geç. Baskale 2500 m Van Turkey MW 99258 ♂ Agrodiaetus turcicola 1 C P 1 1 1 17.07.1999 Kurubas Geçidi Gürpinar 2200 m Van Turkey MW 99259 ♂ Agrodiaetus turcicola 1 C P 1 17.07.1999 Kurubas Geçidi Gürpinar 2200 m Van Turkey MW 99260 ♂ Agrodiaetus turcicola 1 C 1 17.07.1999 Kurubas Geçidi Gürpinar 2200 m Van Turkey MW 99261 ♂ Agrodiaetus turcicola 1 C 1 17.07.1999 Kurubas Geçidi Gürpinar 2200 m Van Turkey MW 99262 ♂ Agrodiaetus turcicola 1 C P 1 17.07.1999 Kurubas Geçidi Gürpinar 2200 m Van Turkey MW 99263 ♂ Agrodiaetus ripartii 1 C P 1 1 1 17.07.1999 Kurubas Geçidi Gürpinar 2200 m Van Turkey MW 99264 ♂ Agrodiaetus ripartii 1 C P 1 1 1 17.07.1999 Kurubas Geçidi Gürpinar 2200 m Van Turkey MW 99265 ♂ Agrodiaetus ripartii 1 C P 1 17.07.1999 Kurubas Geçidi Gürpinar 2200 m Van Turkey MW 99266 ♂ Agrodiaetus turcicola 1 C 1 17.07.1999 Kurubas Geçidi Gürpinar 2200 m Van Turkey MW 99267 ♂ Agrodiaetus turcicola 1 C 1 17.07.1999 Kurubas Geçidi Gürpinar 2200 m Van Turkey MW 99268 ♂ Agrodiaetus turcicola 1 C 1 17.07.1999 Kurubas Geçidi Gürpinar 2200 m Van Turkey MW 99269 ♂ Agrodiaetus turcicola 1 C 1 17.07.1999 Kurubas Geçidi Gürpinar 2200 m Van Turkey MW 99270 ♂ Agrodiaetus turcicola 1 C 1 17.07.1999 Kurubas Geçidi Gürpinar 2200 m Van Turkey MW 99271 ♂ Agrodiaetus turcicola 1 C 1 17.07.1999 Kurubas Geçidi Gürpinar 2200 m Van Turkey MW 99272 ♂ Agrodiaetus turcicola 1 C P 1 17.07.1999 Kurubas Geçidi Gürpinar 2200 m Van Turkey MW 99273 ♂ Agrodiaetus turcicola 1 C P 1 17.07.1999 Kurubas Geçidi Gürpinar 2200 m Van Turkey MW 99274 ♂ Agrodiaetus dantchenkoi 1 C P 1 1 1 1 II-16 17.07.1999 Kurubas Geçidi Gürpinar 2200 m Van Turkey MW 99275 ♀ Agrodiaetus dantchenkoi 1 C 1 17.07.1999 Kurubas Geçidi Gürpinar 2200 m Van Turkey MW 99276 ♂ Agrodiaetus dantchenkoi 1 C P 1 1 1 1 17.07.1999 Kurubas Geçidi Gürpinar 2200 m Van Turkey MW 99277 ♂ Agrodiaetus dantchenkoi 1 C P 1 17.07.1999 Kurubas Geçidi Gürpinar 2200 m Van Turkey MW 99278 ♂ Agrodiaetus dantchenkoi 1 C P 1 17.07.1999 Kurubas Geçidi Gürpinar 2200 m Van Turkey MW 99279 ♂ Meleageria daphnis 1 C P 1 17.07.1999 Kurubas Geçidi Gürpinar 2200 m Van Turkey MW 99280 ♂ Meleageria daphnis 1 C 1 17.07.1999 Kurubas Geçidi Gürpinar 2200 m Van Turkey MW 99281 ♂ Meleageria daphnis 1 C 1 17.07.1999 Kurubas Geçidi Gürpinar 2200 m Van Turkey MW 99282 ♂ Meleageria daphnis 1 C P 1 17.07.1999 Kurubas Geçidi Gürpinar 2200 m Van Turkey MW 99283 ♂ Kretania eurypilus 1 C P 1 17.07.1999 Kurubas Geçidi Gürpinar 2200 m Van Turkey MW 99284 ♂ Polyommatus thersites 1 C P 1 17.07.1999 Kurubas Geçidi Gürpinar 2200 m Van Turkey MW 99285 ♂ Polyommatus icarus 1 C 1 17.07.1999 Kurubas Geçidi Gürpinar 2200 m Van Turkey MW 99286 ♂ Agrodiaetus kurdistanicus 1 C P 1 1 1 1 II-17 18.07.1999 Çatak 5-18 km N Çatak 1600-1900 m Van Turkey MW 99287 ♂ Agrodiaetus kurdistanicus 1 C P 1 18.07.1999 Çatak 5-18 km N Çatak 1600-1900 m Van Turkey MW 99288 ♂ Agrodiaetus kurdistanicus 1 C P 1 18.07.1999 Çatak 5-18 km N Çatak 1600-1900 m Van Turkey MW 99289 ♂ Agrodiaetus sekercioglui 1 C P 1 1 1 II-18 18.07.1999 Çatak 5-18 km N Çatak 1600-1900 m Van Turkey MW 99290 ♂ Agrodiaetus firdussii 1 C P 1 18.07.1999 Çatak 5-18 km N Çatak 1600-1900 m Van Turkey MW 99291 ♂ Agrodiaetus sekercioglui 1 C P 1 18.07.1999 Çatak 5-18 km N Çatak 1600-1900 m Van Turkey MW 99292 ♂ Agrodiaetus pierceae 1 C P 1 1 1 1 18.07.1999 Çatak 5-18 km N Çatak 1600-1900 m Van Turkey MW 99293 ♂ Agrodiaetus baytopi 1 C P 1 18.07.1999 Çatak 5-18 km N Çatak 1600-1900 m Van Turkey MW 99294 ♂ Agrodiaetus turcicola 1 C P 1 18.07.1999 Çatak 5-18 km N Çatak 1600-1900 m Van Turkey MW 99295 ♂ Agrodiaetus pierceae 1 C P 1 18.07.1999 Çatak 5-18 km N Çatak 1600-1900 m Van Turkey MW 99296 ♂ Agrodiaetus pierceae 1 C P 1 18.07.1999 Çatak 5-18 km N Çatak 1600-1900 m Van Turkey MW 99297 ♂ Agrodiaetus iphigenia 1 C 1 18.07.1999 Çatak 5-18 km N Çatak 1600-1900 m Van Turkey MW 99298 ♂ Agrodiaetus alcestis 1 C P 1 18.07.1999 Çatak 5-18 km N Çatak 1600-1900 m Van Turkey MW 99299 ♂ Agrodiaetus alcestis 1 C P 1 18.07.1999 Çatak 5-18 km N Çatak 1600-1900 m Van Turkey MW 99300 ♂ Vacciniina alcedo 1 C 1 18.07.1999 Çatak 5-18 km N Çatak 1600-1900 m Van Turkey MW 99301 ♂ Neolysandra fatima 1 C 1 1 1 1 18.07.1999 Çatak 5-18 km N Çatak 1600-1900 m Van Turkey MW 99302 ♂ Turanana endymion 1 C 1 18.07.1999 Çatak 5-18 km N Çatak 1600-1900 m Van Turkey MW 99303 ♂ Kretania eurypilus 1 1 1 1 1 18.07.1999 Çatak 5-18 km N Çatak 1600-1900 m Van Turkey MW 99304 ♂ Plebeius loewii 1 C 1 18.07.1999 Çatak 5-18 km N Çatak 1600-1900 m Van Turkey MW 99305 ♀ Pseudophilotes vicrama 1 C 1 18.07.1999 Çatak 5-18 km N Çatak 1600-1900 m Van Turkey MW 99306 ♂ Agrodiaetus iphigenia 1 C 1 18.07.1999 Çatak 25-32 km N Çatak 2000-2200 m Van Turkey MW 99307 ♂ Agrodiaetus baytopi 1 C P 1 18.07.1999 Çatak 25-32 km N Çatak 2000-2200 m Van Turkey MW 99308 ♂ Agrodiaetus zapvadi 1 C P 1 18.07.1999 Çatak 25-32 km N Çatak 2000-2200 m Van Turkey MW 99309 ♂ Agrodiaetus baytopi 1 C- 1 1 1 1 II-19 18.07.1999 Çatak 25-32 km N Çatak 2000-2200 m Van Turkey MW 99310 ♂ Agrodiaetus iphigenia 1 C 1 18.07.1999 Çatak 25-32 km N Çatak 2000-2200 m Van Turkey MW 99311 ♂ Agrodiaetus baytopi 1 C P 1 18.07.1999 Çatak 25-32 km N Çatak 2000-2200 m Van Turkey MW 99312 ♂ Agrodiaetus baytopi 1 C P 1 18.07.1999 Çatak 25-32 km N Çatak 2000-2200 m Van Turkey MW 99313 ♂ Agrodiaetus iphigenia 1 C 1 1 18.07.1999 Çatak 25-32 km N Çatak 2000-2200 m Van Turkey MW 99314 ♂ Agrodiaetus turcicola 1 C P 1 1 1 18.07.1999 Çatak 25-32 km N Çatak 2000-2200 m Van Turkey MW 99315 ♂ Agrodiaetus firdussii 1 C 1 18.07.1999 Çatak 25-32 km N Çatak 2000-2200 m Van Turkey MW 99316 ♂ Agrodiaetus firdussii 1 C 1 18.07.1999 Çatak 25-32 km N Çatak 2000-2200 m Van Turkey MW 99317 ♂ Agrodiaetus turcicola 1 C P 1 18.07.1999 Çatak 25-32 km N Çatak 2000-2200 m Van Turkey MW 99318 ♂ Agrodiaetus turcicola 1 C P 1 18.07.1999 Çatak 25-32 km N Çatak 2000-2200 m Van Turkey MW 99319 ♂ Agrodiaetus dantchenkoi 1 C P 1 1 1 1 18.07.1999 Çatak 25-32 km N Çatak 2000-2200 m Van Turkey MW 99320 ♂ Agrodiaetus dantchenkoi 1 C P 1 1 1 18.07.1999 Çatak 25-32 km N Çatak 2000-2200 m Van Turkey MW 99321 ♂ Agrodiaetus turcicola 1 C P 1 18.07.1999 Çatak 25-32 km N Çatak 2000-2200 m Van Turkey MW 99322 ♂ Agrodiaetus dantchenkoi 1 C P 1 18.07.1999 Çatak 25-32 km N Çatak 2000-2200 m Van Turkey MW 99323 ♂ Neolysandra fatima 1 C P 1 18.07.1999 Çatak 25-32 km N Çatak 2000-2200 m Van Turkey MW 99324 ♂ Agrodiaetus turcicola 1 C 1 17.07.1999 Kurubas Geçidi Gürpinar 2200 m Van Turkey MW 99325 ♂ Agrodiaetus dantchenkoi 1 C P 1 17.07.1999 Kurubas Geçidi Gürpinar 2200 m Van Turkey MW 99326 ♂ Agrodiaetus turcicola 1 1 17.07.1999 Kurubas Geçidi Gürpinar 2200 m Van Turkey MW 99327 ♂ Agrodiaetus turcicola 1 1 17.07.1999 Kurubas Geçidi Gürpinar 2200 m Van Turkey MW 99328 ♂ Agrodiaetus turcicola 1 1 17.07.1999 Kurubas Geçidi Gürpinar 2200 m Van Turkey MW 99329 ♂ Agrodiaetus turcicola 1 1 17.07.1999 Kurubas Geçidi Gürpinar 2200 m Van Turkey MW 99330 ♂ Agrodiaetus turcicola 1 1 17.07.1999 Kurubas Geçidi Gürpinar 2200 m Van Turkey MW 99331 ♂ Agrodiaetus turcicola 1 1 17.07.1999 Kurubas Geçidi Gürpinar 2200 m Van Turkey MW 99332 ♂ Agrodiaetus turcicola 1 1 17.07.1999 Kurubas Geçidi Gürpinar 2200 m Van Turkey MW 99333 ♂ Agrodiaetus turcicola 1 1 17.07.1999 Kurubas Geçidi Gürpinar 2200 m Van Turkey MW 99334 ♂ Agrodiaetus turcicola 1 C 1 19.07.1999 Kurubas Geçidi Gürpinar 2200 m Van Turkey MW 99335 ♂ Agrodiaetus cyaneus 1 C P 1 19.07.1999 Kurubas Geçidi Gürpinar 2200 m Van Turkey MW 99336 ♂ Agrodiaetus altivagans 1 C 1 19.07.1999 Güzeldere Geç. Baskale 2600 m Van Turkey MW 99337 ♂ Agrodiaetus altivagans 1 C P 1 19.07.1999 Güzeldere Geç. Baskale 2600 m Van Turkey MW 99338 ♂ Agrodiaetus altivagans 1 C P 1 19.07.1999 Güzeldere Geç. Baskale 2600 m Van Turkey MW 99339 ♂ Agrodiaetus pierceae 1 C 1 1 19.07.1999 Güzeldere Geç. Baskale 2600 m Van Turkey MW 99340 ♂ Agrodiaetus pierceae 1 C 1 19.07.1999 Güzeldere Geç. Baskale 2600 m Van Turkey MW 99341 ♂ Agrodiaetus pierceae 1 C P 1 1 1 1 II-20 19.07.1999 Güzeldere Geç. Baskale 2600 m Van Turkey

- 161 - Appendix 2 List of material

Code Sex Genus Species Wgs Fix Kar Alc DNA Cytb CO1 ND1 ITS2 Fig. Date Locality Area Altitude Province Country MW 99342 ♂ Agrodiaetus phyllis 1 C 1 19.07.1999 Güzeldere Geç. Baskale 2600 m Van Turkey MW 99343 ♂ Agrodiaetus phyllis 1 C 1 19.07.1999 Güzeldere Geç. Baskale 2600 m Van Turkey MW 99344 ♂ Agrodiaetus baytopi 1 C P 1 1 1 19.07.1999 Güzeldere Geç. Baskale 2600 m Van Turkey MW 99345 ♂ Agrodiaetus baytopi 1 C P 1 19.07.1999 Güzeldere Geç. Baskale 2600 m Van Turkey MW 99346 ♂ Agrodiaetus pierceae 1 C 1 19.07.1999 Güzeldere Geç. Baskale 2600 m Van Turkey MW 99347 ♂ Agrodiaetus pierceae 1 C P 1 19.07.1999 Güzeldere Geç. Baskale 2600 m Van Turkey MW 99348 ♂ Agrodiaetus pierceae 1 C 1 19.07.1999 Güzeldere Geç. Baskale 2600 m Van Turkey MW 99349 ♂ Agrodiaetus iphigenia 1 C 1 19.07.1999 Güzeldere Geç. Baskale 2600 m Van Turkey MW 99350 ♂ Agrodiaetus iphigenia 1 C 1 19.07.1999 Güzeldere Geç. Baskale 2600 m Van Turkey MW 99351 ♂ Agrodiaetus zapvadi 1 C P 1 19.07.1999 Güzeldere Geç. Baskale 2600 m Van Turkey MW 99352 ♂ Agrodiaetus turcicola 1 C 1 19.07.1999 Güzeldere Geç. Baskale 2500 m Van Turkey MW 99353 ♂ Agrodiaetus altivagans 1 C P 1 1 1 1 II-21 19.07.1999 Güzeldere Geç. Baskale 2500 m Van Turkey MW 99354 ♂ Agrodiaetus firdussii 1 C 1 19.07.1999 Güzeldere Geç. Baskale 2500 m Van Turkey MW 99355 ♂ Agrodiaetus pierceae 1 C 1 19.07.1999 Güzeldere Geç. Baskale 2500 m Van Turkey MW 99356 ♂ Agrodiaetus phyllis 1 C P 1 19.07.1999 Güzeldere Geç. Baskale 2500 m Van Turkey MW 99357 ♂ Agrodiaetus altivagans 1 C P 1 1 1 19.07.1999 Güzeldere Geç. Baskale 2500 m Van Turkey MW 99358 ♂ Agrodiaetus pierceae 1 C 1 19.07.1999 Güzeldere Geç. Baskale 2500 m Van Turkey MW 99359 ♂ Agrodiaetus pierceae 1 C 1 19.07.1999 Güzeldere Geç. Baskale 2500 m Van Turkey MW 99360 ♂ Agrodiaetus baytopi 1 C P 1 19.07.1999 Güzeldere Geç. Baskale 2500 m Van Turkey MW 99361 ♂ Agrodiaetus baytopi 1 C P 1 19.07.1999 Güzeldere Geç. Baskale 2500 m Van Turkey MW 99362 ♂ Agrodiaetus pierceae 1 C 1 19.07.1999 Güzeldere Geç. Baskale 2500 m Van Turkey MW 99363 ♂ Agrodiaetus baytopi 1 C P 1 19.07.1999 Güzeldere Geç. Baskale 2500 m Van Turkey MW 99364 ♂ Agrodiaetus baytopi 1 C 1 19.07.1999 Güzeldere Geç. Baskale 2500 m Van Turkey MW 99365 ♂ Agrodiaetus pierceae 1 C 1 19.07.1999 Güzeldere Geç. Baskale 2500 m Van Turkey MW 99366 ♂ Agrodiaetus pierceae 1 C P 1 19.07.1999 Güzeldere Geç. Baskale 2500 m Van Turkey MW 99367 ♂ Agrodiaetus baytopi 1 C 1 19.07.1999 Güzeldere Geç. Baskale 2500 m Van Turkey MW 99368 ♂ Agrodiaetus baytopi 1 C 1 19.07.1999 Güzeldere Geç. Baskale 2500 m Van Turkey MW 99369 ♂ Agrodiaetus baytopi 1 C 1 19.07.1999 Güzeldere Geç. Baskale 2500 m Van Turkey MW 99370 ♂ Agrodiaetus pierceae 1 C P 1 19.07.1999 Güzeldere Geç. Baskale 2500 m Van Turkey MW 99371 ♂ Agrodiaetus pierceae 1 C P 1 19.07.1999 Güzeldere Geç. Baskale 2500 m Van Turkey MW 99372 ♂ Agrodiaetus baytopi 1 C P 1 1 1 1 19.07.1999 Güzeldere Geç. Baskale 2500 m Van Turkey MW 99373 ♂ Agrodiaetus zapvadi 1 C P 1 20.07.1999 Zernek Brj. Güzelsu 1900 m Van Turkey MW 99374 ♂ Agrodiaetus zapvadi 1 C P 1 1 1 1 II-22 20.07.1999 Zernek Brj. Güzelsu 1900 m Van Turkey MW 99375 ♂ Agrodiaetus cyaneus 1 C P 1 20.07.1999 Zernek Brj. Güzelsu 1900 m Van Turkey MW 99376 ♂ Agrodiaetus antidolus 1 C P 1 1 1 20.07.1999 Yüksekova 22 km NW Yüksekova 1800 m Hakkari Turkey MW 99377 ♂ Agrodiaetus antidolus 1 C P 1 20.07.1999 Yüksekova 22 km NW Yüksekova 1800 m Hakkari Turkey MW 99378 ♂ Agrodiaetus antidolus 1 C P 1 20.07.1999 Yüksekova 22 km NW Yüksekova 1800 m Hakkari Turkey MW 99379 ♀ Agrodiaetus antidolus 1 C 1 20.07.1999 Yüksekova 22 km NW Yüksekova 1800 m Hakkari Turkey MW 99380 ♂ Agrodiaetus alcestis 1 C P 1 1 1 20.07.1999 Yüksekova 22 km NW Yüksekova 1800 m Hakkari Turkey MW 99381 ♂ Agrodiaetus demavendi 1 C P 1 1 1 20.07.1999 Yüksekova 22 km NW Yüksekova 1800 m Hakkari Turkey MW 99382 ♂ Agrodiaetus demavendi 1 C P 1 1 1 1 20.07.1999 Yüksekova 22 km NW Yüksekova 1800 m Hakkari Turkey MW 99383 ♂ Agrodiaetus demavendi 1 C P 1 20.07.1999 Yüksekova 22 km NW Yüksekova 1800 m Hakkari Turkey MW 99384 ♂ Agrodiaetus demavendi 1 C P 1 20.07.1999 Yüksekova 22 km NW Yüksekova 1800 m Hakkari Turkey MW 99385 ♀ Agrodiaetus demavendi 1 C 1 20.07.1999 Yüksekova 22 km NW Yüksekova 1800 m Hakkari Turkey MW 99386 ♂ Agrodiaetus demavendi 1 C P 1 20.07.1999 Yüksekova 22 km NW Yüksekova 1800 m Hakkari Turkey MW 99387 ♀ Lycaena alciphron 1 C 1 20.07.1999 Yüksekova 22 km NW Yüksekova 1800 m Hakkari Turkey MW 99388 ♂ Lycaena tityrus 1 C 1 20.07.1999 Yüksekova 22 km NW Yüksekova 1800 m Hakkari Turkey MW 99389 ♂ Agrodiaetus antidolus 1 C P 1 20.07.1999 Dilezi Geçidi nordöstl. Yüksekova 2100 m Hakkari Turkey MW 99390 ♂ Agrodiaetus demavendi 1 C P 1 20.07.1999 Dilezi Geçidi nordöstl. Yüksekova 2100 m Hakkari Turkey MW 99391 ♂ Agrodiaetus demavendi 1 C P 1 20.07.1999 Dilezi Geçidi nordöstl. Yüksekova 2100 m Hakkari Turkey MW 99392 ♂ Agrodiaetus demavendi 1 C P 1 20.07.1999 Dilezi Geçidi nordöstl. Yüksekova 2100 m Hakkari Turkey MW 99393 ♂ Agrodiaetus antidolus 1 C P 1 1 1 1 II-23 21.07.1999 Haruna Geçidi südöstl. Yüksekova 2000 m Hakkari Turkey MW 99394 ♂ Agrodiaetus antidolus 1 C P 1 21.07.1999 Haruna Geçidi südöstl. Yüksekova 2000 m Hakkari Turkey MW 99395 ♀ Agrodiaetus antidolus 1 C 1 21.07.1999 Haruna Geçidi südöstl. Yüksekova 2000 m Hakkari Turkey MW 99396 ♂ Lycaena thersamon 1 C P 1 21.07.1999 Haruna Geçidi südöstl. Yüksekova 2000 m Hakkari Turkey MW 99397 ♀ Polyommatus icarus 1 1 21.07.1999 Haruna Geçidi südöstl. Yüksekova 2000 m Hakkari Turkey MW 99398 ♂ Satyrium myrtale 1 1 1 1 21.07.1999 Haruna Geçidi südöstl. Yüksekova 2000 m Hakkari Turkey MW 99399 ♀ Satyrium myrtale 1 1 1 21.07.1999 Haruna Geçidi südöstl. Yüksekova 2000 m Hakkari Turkey MW 99400 ♀ Satyrium myrtale 1 1 21.07.1999 Haruna Geçidi südöstl. Yüksekova 2000 m Hakkari Turkey MW 99401 ♀ Satyrium myrtale 1 1 21.07.1999 Haruna Geçidi südöstl. Yüksekova 2000 m Hakkari Turkey MW 99402 ♀ Satyrium myrtale 1 1 21.07.1999 Haruna Geçidi südöstl. Yüksekova 2000 m Hakkari Turkey MW 99403 ♂ Agrodiaetus antidolus 1 C P 1 22.07.1999 Dez Çay nordöstl. Hakkari 1500 m Hakkari Turkey MW 99404 ♂ Agrodiaetus antidolus 1 C P 1 22.07.1999 Dez Çay nordöstl. Hakkari 1500 m Hakkari Turkey MW 99405 ♂ Agrodiaetus demavendi 1 C P 1 22.07.1999 Dez Çay nordöstl. Hakkari 1500 m Hakkari Turkey MW 99406 ♂ Agrodiaetus antidolus 1 C P 1 1 1 1 1 II-24 22.07.1999 Dez Çay nordöstl. Hakkari 1500 m Hakkari Turkey MW 99407 ♂ Agrodiaetus demavendi 1 C P 1 1 1 1 22.07.1999 Dez Çay nordöstl. Hakkari 1500 m Hakkari Turkey MW 99408 ♂ Agrodiaetus hopfferi 1 C P 1 1 1 1 II-25 22.07.1999 Dez Çay nordöstl. Hakkari 1500 m Hakkari Turkey MW 99409 ♂ Agrodiaetus hopfferi 1 C- P 1 22.07.1999 Dez Çay nordöstl. Hakkari 1500 m Hakkari Turkey MW 99410 ♂ Agrodiaetus hopfferi 1 C- 1 22.07.1999 Dez Çay nordöstl. Hakkari 1500 m Hakkari Turkey MW 99411 ♂ Agrodiaetus turcicola 1 C P 1 22.07.1999 Dez Çay nordöstl. Hakkari 1500 m Hakkari Turkey MW 99412 ♂ Agrodiaetus phyllis 1 C 1 22.07.1999 Dez Çay nordöstl. Hakkari 1500 m Hakkari Turkey MW 99413 ♂ Agrodiaetus firdussii 1 C P 1 1 1 1 II-26 22.07.1999 Dez Çay nordöstl. Hakkari 1500 m Hakkari Turkey MW 99414 ♂ Agrodiaetus altivagans 1 C P 1 22.07.1999 Dez Çay nordöstl. Hakkari 1500 m Hakkari Turkey MW 99415 ♂ Agrodiaetus cyaneus 1 C P 1 22.07.1999 Dez Çay nordöstl. Hakkari 1500 m Hakkari Turkey MW 99416 ♂ Agrodiaetus pierceae 1 C P 1 22.07.1999 Dez Çay nordöstl. Hakkari 1500 m Hakkari Turkey MW 99417 ♂ Agrodiaetus firdussii 1 C P 1 22.07.1999 Dez Çay nordöstl. Hakkari 1500 m Hakkari Turkey MW 99418 ♂ Agrodiaetus pierceae 1 C P 1 22.07.1999 Dez Çay nordöstl. Hakkari 1500 m Hakkari Turkey MW 99419 ♂ Agrodiaetus pierceae 1 C P 1 22.07.1999 Dez Çay nordöstl. Hakkari 1500 m Hakkari Turkey MW 99420 ♂ Agrodiaetus iphigenia 1 C 1 22.07.1999 Dez Çay nordöstl. Hakkari 1500 m Hakkari Turkey MW 99421 ♂ Agrodiaetus iphigenia 1 C 1 22.07.1999 Dez Çay nordöstl. Hakkari 1500 m Hakkari Turkey MW 99422 ♂ Agrodiaetus firdussii 1 C P 1 1 1 22.07.1999 Dez Çay nordöstl. Hakkari 1500 m Hakkari Turkey MW 99423 ♂ Agrodiaetus iphigenia 1 C 1 22.07.1999 Dez Çay nordöstl. Hakkari 1500 m Hakkari Turkey MW 99424 ♂ Lysandra bellargus 1 C 1 22.07.1999 Dez Çay nordöstl. Hakkari 1500 m Hakkari Turkey MW 99425 ♂ Chilades trochylus 1 1 1 1 22.07.1999 Dez Çay nordöstl. Hakkari 1500 m Hakkari Turkey MW 99426 ♂ Chilades trochylus 1 1 22.07.1999 Dez Çay nordöstl. Hakkari 1500 m Hakkari Turkey MW 99427 ♂ Cyaniris semiargus 1 1 22.07.1999 Dez Çay nordöstl. Hakkari 1500 m Hakkari Turkey MW 99428 ♂ Polyommatus icarus 1 1 22.07.1999 Dez Çay nordöstl. Hakkari 1500 m Hakkari Turkey MW 99429 ♂ Polyommatus icarus 1 1 22.07.1999 Dez Çay nordöstl. Hakkari 1500 m Hakkari Turkey MW 99430 ♂ Vacciniina alcedo 1 1 1 1 22.07.1999 Dez Çay nordöstl. Hakkari 1500 m Hakkari Turkey MW 99431 ♂ Vacciniina alcedo 1 1 22.07.1999 Dez Çay nordöstl. Hakkari 1500 m Hakkari Turkey MW 99432 ♂ Vacciniina alcedo 1 1 22.07.1999 Dez Çay nordöstl. Hakkari 1500 m Hakkari Turkey MW 99433 ♂ Vacciniina alcedo 1 1 22.07.1999 Dez Çay nordöstl. Hakkari 1500 m Hakkari Turkey MW 99434 ♂ Lysandra bellargus 1 1 22.07.1999 Dez Çay nordöstl. Hakkari 1500 m Hakkari Turkey MW 99435 ♂ Lysandra bellargus 1 1 1 1 22.07.1999 Dez Çay nordöstl. Hakkari 1500 m Hakkari Turkey MW 99436 ♂ Polyommatus icarus 1 1 22.07.1999 Dez Çay nordöstl. Hakkari 1500 m Hakkari Turkey MW 99437 ♂ Polyommatus icarus 1 1 22.07.1999 Dez Çay nordöstl. Hakkari 1500 m Hakkari Turkey MW 99438 ♀ Kretania eurypilus 1 1 22.07.1999 Dez Çay nordöstl. Hakkari 1500 m Hakkari Turkey MW 99439 ♀ Lycaena phlaeas 1 1 22.07.1999 Dez Çay nordöstl. Hakkari 1500 m Hakkari Turkey MW 99440 ♂ Agrodiaetus iphigenia 1 C P 1 23.07.1999 Dez Çay nordöstl. Hakkari 1500-1700 m Hakkari Turkey MW 99441 ♂ Agrodiaetus firdussii 1 C P 1 23.07.1999 Dez Çay nordöstl. Hakkari 1500-1700 m Hakkari Turkey MW 99442 ♂ Agrodiaetus phyllis 1 C 1 23.07.1999 Dez Çay nordöstl. Hakkari 1500-1700 m Hakkari Turkey MW 99443 ♂ Agrodiaetus firdussii 1 C P 1 23.07.1999 Dez Çay nordöstl. Hakkari 1500-1700 m Hakkari Turkey MW 99444 ♂ Agrodiaetus firdussii 1 C P 1 23.07.1999 Dez Çay nordöstl. Hakkari 1500-1700 m Hakkari Turkey MW 99445 ♂ Agrodiaetus firdussii 1 C P 1 23.07.1999 Dez Çay nordöstl. Hakkari 1500-1700 m Hakkari Turkey MW 99446 ♂ Lysandra bellargus 1 1 1 23.07.1999 Dez Çay nordöstl. Hakkari 1500-1700 m Hakkari Turkey MW 99447 ♂ Agrodiaetus pierceae 1 C P 1 23.07.1999 Zernek Brj. Güzelsu 1900 m Van Turkey MW 99448 ♂ Agrodiaetus cyaneus 1 C P 1 1 1 1 II-27 23.07.1999 Zernek Brj. Güzelsu 1900 m Van Turkey MW 99449 ♂ Agrodiaetus cyaneus 1 C P 1 1 1 23.07.1999 Zernek Brj. Güzelsu 1900 m Van Turkey MW 99450 ♂ Agrodiaetus turcicola 1 C P 1 23.07.1999 Zernek Brj. Güzelsu 1900 m Van Turkey MW 99451 ♂ Agrodiaetus baytopi 1 C P 1 24.07.1999 Çatak 5-18 km N Çatak 1600-1900 m Van Turkey MW 99452 ♂ Agrodiaetus phyllis 1 C 1 24.07.1999 Çatak 5-18 km N Çatak 1600-1900 m Van Turkey MW 99453 ♂ Agrodiaetus hopfferi 1 C P 1 24.07.1999 Çatak 5-18 km N Çatak 1600-1900 m Van Turkey MW 99454 ♂ Agrodiaetus baytopi 1 C- 1 24.07.1999 Çatak 5-18 km N Çatak 1600-1900 m Van Turkey MW 99455 ♂ Agrodiaetus baytopi 1 C 1 24.07.1999 Çatak 5-18 km N Çatak 1600-1900 m Van Turkey MW 99456 ♂ Agrodiaetus phyllis 1 C 1 24.07.1999 Çatak 5-18 km N Çatak 1600-1900 m Van Turkey

- 162 - Appendix 2 List of material

Code Sex Genus Species Wgs Fix Kar Alc DNA Cytb CO1 ND1 ITS2 Fig. Date Locality Area Altitude Province Country MW 99457 ♂ Agrodiaetus firdussii 1 C P 1 24.07.1999 Çatak 5-18 km N Çatak 1600-1900 m Van Turkey MW 99458 ♂ Agrodiaetus turcicus 1 C P 1 24.07.1999 Çatak 5-18 km N Çatak 1600-1900 m Van Turkey MW 99459 ♂ Agrodiaetus phyllis 1 C 1 24.07.1999 Çatak 5-18 km N Çatak 1600-1900 m Van Turkey MW 99460 ♂ Agrodiaetus baytopi 1 C 1 24.07.1999 Çatak 5-18 km N Çatak 1600-1900 m Van Turkey MW 99461 ♂ Agrodiaetus baytopi 1 C P 1 24.07.1999 Çatak 5-18 km N Çatak 1600-1900 m Van Turkey MW 99462 ♂ Agrodiaetus phyllis 1 C 1 24.07.1999 Çatak 5-18 km N Çatak 1600-1900 m Van Turkey MW 99463 ♂ Agrodiaetus phyllis 1 C P 1 24.07.1999 Çatak 5-18 km N Çatak 1600-1900 m Van Turkey MW 99464 ♂ Agrodiaetus turcicola 1 C P 1 24.07.1999 Çatak 25 km N Çatak 1600-1900 m Van Turkey MW 99465 ♂ Agrodiaetus kanduli 1 C P 1 1 1 1 II-28 24.07.1999 Çatak 25 km N Çatak 1600-1900 m Van Turkey MW 99466 ♂ Agrodiaetus iphigenia 1 C 1 24.07.1999 Çatak 25 km N Çatak 1600-1900 m Van Turkey MW 99467 ♂ Neolysandra fatima 1 1 24.07.1999 Çatak 5-18 km N Çatak 1600-1900 m Van Turkey MW 99468 ♂ Neolysandra fatima 1 1 24.07.1999 Çatak 5-18 km N Çatak 1600-1900 m Van Turkey MW 99469 ♂ Neolysandra fatima 1 1 1 1 24.07.1999 Çatak 5-18 km N Çatak 1600-1900 m Van Turkey MW 99470 ♂ Meleageria daphnis 1 1 24.07.1999 Çatak 5-18 km N Çatak 1600-1900 m Van Turkey MW 99471 ♂ Agrodiaetus dantch.Xmenalc. 1 C P 1 1 1 1 1 II-29 25.07.1999 Erek Dagi Van 2200 m Van Turkey MW 99472 ♂ Agrodiaetus kurdistanicus 1 C P 1 25.07.1999 Erek Dagi Van 2200 m Van Turkey MW 99473 ♂ Agrodiaetus kurdistanicus 1 C P 1 1 1 25.07.1999 Erek Dagi Van 2200 m Van Turkey MW 99474 ♀ Agrodiaetus kurdistanicus 1 C 1 25.07.1999 Erek Dagi Van 2200 m Van Turkey MW 99475 ♂ Agrodiaetus cyaneus 1 C P 1 25.07.1999 Erek Dagi Van 2200 m Van Turkey MW 99476 ♂ Agrodiaetus zapvadi 1 C P 1 1 1 25.07.1999 Erek Dagi Van 2200 m Van Turkey MW 99477 ♂ Agrodiaetus zapvadi 1 C P 1 25.07.1999 Erek Dagi Van 2200 m Van Turkey MW 99478 ♂ Agrodiaetus turcicola 1 C P 1 25.07.1999 Erek Dagi Van 2200 m Van Turkey MW 99479 ♂ Agrodiaetus turcicola 1 C P 1 1 1 1 II-30 25.07.1999 Erek Dagi Van 2200 m Van Turkey MW 99480 ♂ Agrodiaetus turcicola 1 C P 1 25.07.1999 Erek Dagi Van 2200 m Van Turkey MW 99481 ♂ Agrodiaetus turcicola 1 C 1 25.07.1999 Erek Dagi Van 2200 m Van Turkey MW 99482 ♂ Agrodiaetus turcicola 1 C 1 25.07.1999 Erek Dagi Van 2200 m Van Turkey MW 99483 ♂ Agrodiaetus turcicola 1 C P 1 25.07.1999 Erek Dagi Van 2200 m Van Turkey MW 99484 ♂ Agrodiaetus turcicola 1 C 1 25.07.1999 Erek Dagi Van 2200 m Van Turkey MW 99485 ♂ Agrodiaetus turcicola 1 C 1 25.07.1999 Erek Dagi Van 2200 m Van Turkey MW 99486 ♂ Agrodiaetus turcicola 1 C 1 25.07.1999 Erek Dagi Van 2200 m Van Turkey MW 99487 ♂ Agrodiaetus turcicola 1 C 1 25.07.1999 Erek Dagi Van 2200 m Van Turkey MW 99488 ♂ Agrodiaetus turcicola 1 C 1 25.07.1999 Erek Dagi Van 2200 m Van Turkey MW 99489 ♂ Agrodiaetus turcicola 1 C 1 25.07.1999 Erek Dagi Van 2200 m Van Turkey MW 99490 ♂ Agrodiaetus turcicola 1 C 1 25.07.1999 Erek Dagi Van 2200 m Van Turkey MW 99491 ♂ Agrodiaetus turcicola 1 C 1 25.07.1999 Erek Dagi Van 2200 m Van Turkey MW 99492 ♂ Agrodiaetus turcicola 1 C P 1 25.07.1999 Erek Dagi Van 2200 m Van Turkey MW 99493 ♂ Agrodiaetus turcicola 1 C P 1 25.07.1999 Erek Dagi Van 2200 m Van Turkey MW 99494 ♂ Agrodiaetus menalcas 1 C P 1 1 1 II-31 25.07.1999 Erek Dagi Van 2200 m Van Turkey MW 99495 ♂ Agrodiaetus menalcas 1 C P 1 25.07.1999 Erek Dagi Van 2200 m Van Turkey MW 99496 ♂ Agrodiaetus menalcas 1 C P 1 25.07.1999 Erek Dagi Van 2200 m Van Turkey MW 99497 ♂ Agrodiaetus menalcas 1 C 1 25.07.1999 Erek Dagi Van 2200 m Van Turkey MW 99498 ♂ Agrodiaetus menalcas 1 C P 1 25.07.1999 Erek Dagi Van 2200 m Van Turkey MW 99499 ♂ Agrodiaetus menalcas 1 1 25.07.1999 Erek Dagi Van 2200 m Van Turkey MW 99500 ♂ Agrodiaetus menalcas 1 1 25.07.1999 Erek Dagi Van 2200 m Van Turkey MW 99501 ♂ Agrodiaetus putnami 1 C P 1 1 1 1 II-32 26.07.1999 Agrı 15 km N Agrı 1800 m Agrı Turkey MW 99502 ♂ Agrodiaetus huberti 1 C P 1 26.07.1999 Agrı 15 km N Agrı 1800 m Agrı Turkey MW 99503 ♂ Agrodiaetus huberti 1 C P 1 26.07.1999 Agrı 15 km N Agrı 1800 m Agrı Turkey MW 99504 ♂ Agrodiaetus putnami 1 C P 1 26.07.1999 Agrı 15 km N Agrı 1800 m Agrı Turkey MW 99505 ♂ Agrodiaetus putnami 1 C P 1 26.07.1999 Agrı 15 km N Agrı 1800 m Agrı Turkey MW 99506 ♂ Agrodiaetus putnami 1 C P 1 26.07.1999 Agrı 15 km N Agrı 1800 m Agrı Turkey MW 99507 ♂ Agrodiaetus putnami 1 C P 1 26.07.1999 Agrı 15 km N Agrı 1800 m Agrı Turkey MW 99508 ♂ Agrodiaetus ninae 1 C P 1 1 1 1 II-33 26.07.1999 Agrı 15 km N Agrı 1800 m Agrı Turkey MW 99509 ♂ Agrodiaetus ninae 1 C P 1 26.07.1999 Agrı 15 km N Agrı 1800 m Agrı Turkey MW 99510 ♂ Agrodiaetus ninae 1 C P 1 26.07.1999 Agrı 15 km N Agrı 1800 m Agrı Turkey MW 99511 ♂ Agrodiaetus ninae 1 C P 1 26.07.1999 Agrı 15 km N Agrı 1800 m Agrı Turkey MW 99512 ♂ Meleageria daphnis 1 1 26.07.1999 Agrı 15 km N Agrı 1800 m Agrı Turkey MW 99513 ♂ Agrodiaetus huberti 1 C P 1 26.07.1999 Cumaçay 5 km N Cumaçay 2000 m Agrı Turkey MW 99514 ♂ Lysandra corydonius 1 1 1 1 1 26.07.1999 Gaziler 10 km N Gaziler 1800 m Igdir Turkey MW 99515 ♂ Lycaena virgaureae 1 1 1 1 26.07.1999 Gaziler 10 km N Gaziler 1800 m Igdir Turkey MW 99516 ♂ Agrodiaetus wagneri 1 C P 1 27.07.1999 Gaziler 10 km N Gaziler 1800 m Igdir Turkey MW 99517 ♂ Agrodiaetus baytopi 1 C P 1 27.07.1999 Gaziler 10 km N Gaziler 1800 m Igdir Turkey MW 99518 ♂ Lysandra corydonius 1 C 1 27.07.1999 Gaziler 10 km N Gaziler 1800 m Igdir Turkey MW 99519 ♂ Lysandra corydonius 1 C 1 27.07.1999 Gaziler 10 km N Gaziler 1800 m Igdir Turkey MW 99520 ♂ Lycaena thetis 1 1 1 1 28.07.1999 Sac Geçidi 20 km W Eleckirt 2000 m Agrı Turkey MW 99521 ♂ Agrodiaetus damon 1 C 1 28.07.1999 Köskköy 25 km N Erzurum 1900 m Erzurum Turkey MW 99522 ♂ Agrodiaetus damon 1 C 1 28.07.1999 Köskköy 25 km N Erzurum 1900 m Erzurum Turkey MW 99523 ♂ Agrodiaetus damon 1 C P 1 28.07.1999 Köskköy 25 km N Erzurum 1900 m Erzurum Turkey MW 99524 ♂ Agrodiaetus wagneri 1 C P 1 28.07.1999 Köskköy 25 km N Erzurum 1900 m Erzurum Turkey MW 99525 ♂ Agrodiaetus wagneri 1 C P 1 28.07.1999 Köskköy 25 km N Erzurum 1900 m Erzurum Turkey MW 99526 ♂ Agrodiaetus turcicus 1 C P 1 1 1 28.07.1999 Köskköy 25 km N Erzurum 1900 m Erzurum Turkey MW 99527 ♂ Agrodiaetus turcicus 1 C 1 28.07.1999 Köskköy 25 km N Erzurum 1900 m Erzurum Turkey MW 99528 ♂ Agrodiaetus turcicus 1 C 1 28.07.1999 Köskköy 25 km N Erzurum 1900 m Erzurum Turkey MW 99529 ♂ Agrodiaetus turcicus 1 C P 1 28.07.1999 Köskköy 25 km N Erzurum 1900 m Erzurum Turkey MW 99530 ♂ Agrodiaetus damon 1 C 1 28.07.1999 Köskköy 25 km N Erzurum 1900 m Erzurum Turkey MW 99531 ♂ Agrodiaetus damon 1 C P 1 28.07.1999 Köskköy 25 km N Erzurum 1900 m Erzurum Turkey MW 99532 ♂ Agrodiaetus menalcas 1 C 1 28.07.1999 Köskköy 25 km N Erzurum 1900 m Erzurum Turkey MW 99533 ♂ Agrodiaetus alcestis 1 C 1 28.07.1999 Köskköy 25 km N Erzurum 1900 m Erzurum Turkey MW 99534 ♂ Polyommatus myrrhinus 1 C P 1 28.07.1999 Köskköy 25 km N Erzurum 1900 m Erzurum Turkey MW 99535 ♂ Polyommatus myrrhinus 1 C 1 28.07.1999 Köskköy 25 km N Erzurum 1900 m Erzurum Turkey MW 99536 ♂ Lysandra corydonius 1 C P 1 1 1 1 28.07.1999 Köskköy 25 km N Erzurum 1900 m Erzurum Turkey MW 99537 ♂ Polyommatus myrrhinus 1 1 1 1 1 28.07.1999 Köskköy 25 km N Erzurum 1900 m Erzurum Turkey MW 99538 ♂ Polyommatus myrrhinus 1 1 1 1 28.07.1999 Köskköy 25 km N Erzurum 1900 m Erzurum Turkey MW 99539 ♂ Lysandra corydonius 1 1 28.07.1999 Köskköy 25 km N Erzurum 1900 m Erzurum Turkey MW 99540 ♂ Lysandra corydonius 1 1 28.07.1999 Köskköy 25 km N Erzurum 1900 m Erzurum Turkey MW 99541 ♂ Lysandra corydonius 1 1 1 28.07.1999 Köskköy 25 km N Erzurum 1900 m Erzurum Turkey MW 99542 ♂ Polyommatus icarus 1 1 28.07.1999 Köskköy 25 km N Erzurum 1900 m Erzurum Turkey MW 99543 ♂ Polyommatus icarus 1 1 28.07.1999 Köskköy 25 km N Erzurum 1900 m Erzurum Turkey MW 99544 ♂ Agrodiaetus alcestis 1 C P 1 28.07.1999 Köskköy 25 km N Erzurum 1900 m Erzurum Turkey MW 99545 ♂ Agrodiaetus alcestis 1 C P 1 28.07.1999 Köskköy 25 km N Erzurum 1900 m Erzurum Turkey MW 99546 ♂ Agrodiaetus damon 1 1 1 1 1 28.07.1999 Köskköy 25 km N Erzurum 1900 m Erzurum Turkey MW 99547 ♀ Satyrium myrtale 1 1 29.07.1999 Kop Geçidi nördl. Askale 2200 m Erzurum Turkey MW 99548 ♂ Lycaena thetis 1 1 29.07.1999 Kop Geçidi nördl. Askale 2200 m Erzurum Turkey MW 99549 ♂ Plebeius loewii 1 1 29.07.1999 Kop Geçidi nördl. Askale 2200 m Erzurum Turkey MW 99550 ♂ Polyommatus myrrhinus 1 C P 1 1 1 1 1 29.07.1999 Kop Geçidi nördl. Askale 2200 m Erzurum Turkey MW 99551 ♂ Agrodiaetus hopfferi 1 C P 1 29.07.1999 Kop Geçidi nördl. Askale 2200 m Erzurum Turkey MW 99552 ♂ Agrodiaetus huberti 1 C P 1 1 1 1 29.07.1999 Kop Geçidi südl. Maden 2350 m Bayburt Turkey MW 99553 ♂ Agrodiaetus huberti 1 C P 1 29.07.1999 Kop Geçidi südl. Maden 2350 m Bayburt Turkey MW 99554 ♂ Agrodiaetus damon 1 C P 1 29.07.1999 Kop Geçidi südl. Maden 2350 m Bayburt Turkey MW 99555 ♂ Agrodiaetus firdussii 1 C 1 29.07.1999 Kop Geçidi südl. Maden 2350 m Bayburt Turkey MW 99556 ♂ Agrodiaetus damon 1 C 1 29.07.1999 Kop Geçidi südl. Maden 2350 m Bayburt Turkey MW 99557 ♂ Agrodiaetus huberti 1 C P 1 29.07.1999 Kop Geçidi südl. Maden 2350 m Bayburt Turkey MW 99558 ♂ Agrodiaetus huberti 1 C P 1 29.07.1999 Kop Geçidi südl. Maden 2350 m Bayburt Turkey MW 99559 ♂ Agrodiaetus tankeri 1 C P 1 1 1 1 29.07.1999 Kop Geçidi südl. Maden 2350 m Bayburt Turkey MW 99560 ♂ Agrodiaetus phyllis 1 C 1 29.07.1999 Kop Geçidi südl. Maden 2350 m Bayburt Turkey MW 99561 ♂ Agrodiaetus phyllis 1 C 1 29.07.1999 Kop Geçidi südl. Maden 2350 m Bayburt Turkey MW 99562 ♂ Agrodiaetus huberti 1 C P 1 29.07.1999 Kop Geçidi südl. Maden 2350 m Bayburt Turkey MW 99563 ♂ Agrodiaetus tankeri 1 C P 1 29.07.1999 Kop Geçidi südl. Maden 2350 m Bayburt Turkey MW 99564 ♂ Agrodiaetus tankeri 1 C P 1 29.07.1999 Kop Geçidi südl. Maden 2350 m Bayburt Turkey MW 99565 ♂ Agrodiaetus tankeri 1 C P 1 1 1 1 II-34 29.07.1999 Kop Geçidi südl. Maden 2350 m Bayburt Turkey MW 99566 ♂ Agrodiaetus tankeri 1 C P 1 29.07.1999 Kop Geçidi südl. Maden 2350 m Bayburt Turkey MW 99567 ♂ Agrodiaetus damon 1 C P 1 29.07.1999 Kop Geçidi südl. Maden 2350 m Bayburt Turkey MW 99568 ♂ Agrodiaetus damon 1 C 1 29.07.1999 Kop Geçidi südl. Maden 2350 m Bayburt Turkey MW 99569 ♂ Agrodiaetus tankeri 1 C P 1 29.07.1999 Kop Geçidi südl. Maden 2350 m Bayburt Turkey MW 99570 ♂ Agrodiaetus tankeri 1 C P 1 29.07.1999 Kop Geçidi südl. Maden 2350 m Bayburt Turkey MW 99571 ♂ Agrodiaetus tankeri 1 C P 1 29.07.1999 Kop Geçidi südl. Maden 2350 m Bayburt Turkey

- 163 - Appendix 2 List of material

Code Sex Genus Species Wgs Fix Kar Alc DNA Cytb CO1 ND1 ITS2 Fig. Date Locality Area Altitude Province Country MW 99572 ♂ Agrodiaetus phyllis 1 C P 1 29.07.1999 Kop Geçidi südl. Maden 2350 m Bayburt Turkey MW 99573 ♂ Agrodiaetus tankeri 1 C P 1 29.07.1999 Kop Geçidi südl. Maden 2350 m Bayburt Turkey MW 99574 ♂ Agrodiaetus ripartii 1 C P 1 29.07.1999 Kop Geçidi südl. Maden 2350 m Bayburt Turkey MW 99575 ♂ Plebeius pylaon 1 C 1 29.07.1999 Kop Geçidi südl. Maden 2350 m Bayburt Turkey MW 99576 ♂ Polyommatus eros 1 C 1 29.07.1999 Kop Geçidi südl. Maden 2350 m Bayburt Turkey MW 99577 ♂ Polyommatus eros 1 C 1 29.07.1999 Kop Geçidi südl. Maden 2350 m Bayburt Turkey MW 99578 ♂ Polyommatus eros 1 1 1 29.07.1999 Kop Geçidi südl. Maden 2350 m Bayburt Turkey MW 99579 ♂ Polyommatus eros 1 1 1 1 29.07.1999 Kop Geçidi südl. Maden 2350 m Bayburt Turkey MW 99580 ♂ Polyommatus eros 1 1 29.07.1999 Kop Geçidi südl. Maden 2350 m Bayburt Turkey MW 99581 ♂ Polyommatus eros 1 1 29.07.1999 Kop Geçidi südl. Maden 2350 m Bayburt Turkey MW 99582 ♂ Polyommatus eros 1 1 29.07.1999 Kop Geçidi südl. Maden 2350 m Bayburt Turkey MW 99583 ♂ Polyommatus eros 1 1 29.07.1999 Kop Geçidi südl. Maden 2350 m Bayburt Turkey MW 99584 ♂ Agrodiaetus ninae 1 C P 1 30.07.1999 Gölyurt Geçidi Pazaryolu 2350 m Erzurum Turkey MW 99585 ♂ Agrodiaetus huberti 1 C P 1 30.07.1999 Gölyurt Geçidi Pazaryolu 2350 m Erzurum Turkey MW 99586 ♂ Agrodiaetus tankeri 1 C P 1 30.07.1999 Gölyurt Geçidi Pazaryolu 2350 m Erzurum Turkey MW 99587 ♂ Agrodiaetus phyllis 1 C 1 30.07.1999 Gölyurt Geçidi Pazaryolu 2350 m Erzurum Turkey MW 99588 ♂ Agrodiaetus humedasae 1 C P 1 1 14.08.1999 Pondel Val di Cogne 900 m Aosta Italy MW 99589 ♂ Agrodiaetus humedasae 1 C P 1 14.08.1999 Pondel Val di Cogne 900 m Aosta Italy MW 99590 ♂ Agrodiaetus humedasae 1 C P 1 14.08.1999 Pondel Val di Cogne 900 m Aosta Italy MW 99591 ♂ Agrodiaetus humedasae 1 C P 1 1 1 1 II-35 14.08.1999 Pondel Val di Cogne 900 m Aosta Italy MW 99592 ♂ Agrodiaetus humedasae 1 C P 1 14.08.1999 Pondel Val di Cogne 900 m Aosta Italy MW 99593 ♂ Agrodiaetus humedasae 1 C P 1 14.08.1999 Pondel Val di Cogne 900 m Aosta Italy MW 99594 ♂ Agrodiaetus humedasae 1 C P 1 14.08.1999 Pondel Val di Cogne 900 m Aosta Italy MW 99595 ♂ Agrodiaetus humedasae 1 C P 1 14.08.1999 Pondel Val di Cogne 900 m Aosta Italy MW 99596 ♂ Agrodiaetus humedasae 1 C P 1 14.08.1999 Pondel Val di Cogne 900 m Aosta Italy MW 99597 ♂ Agrodiaetus humedasae 1 C P 1 14.08.1999 Pondel Val di Cogne 900 m Aosta Italy MW 99598 ♂ Agrodiaetus humedasae 1 C P 1 14.08.1999 Pondel Val di Cogne 900 m Aosta Italy MW 99599 ♂ Agrodiaetus humedasae 1 C P 1 14.08.1999 Pondel Val di Cogne 900 m Aosta Italy MW 99600 ♀ Agrodiaetus humedasae 1 1 14.08.1999 Pondel Val di Cogne 900 m Aosta Italy MW 99601 ♀ Agrodiaetus humedasae 1 1 14.08.1999 Pondel Val di Cogne 900 m Aosta Italy MW 99602 ♀ Agrodiaetus humedasae 1 1 14.08.1999 Pondel Val di Cogne 900 m Aosta Italy MW 99603 ♀ Agrodiaetus humedasae 1 1 14.08.1999 Pondel Val di Cogne 900 m Aosta Italy MW 99604 ♀ Agrodiaetus humedasae 1 1 14.08.1999 Pondel Val di Cogne 900 m Aosta Italy MW 99605 ♀ Agrodiaetus humedasae 1 1 1 1 1 14.08.1999 Pondel Val di Cogne 900 m Aosta Italy MW 99606 ♂ Agrodiaetus damon 1 1 14.08.1999 Pondel Val di Cogne 900 m Aosta Italy MW 99607 ♀ Agrodiaetus damon 1 1 14.08.1999 Pondel Val di Cogne 900 m Aosta Italy MW 99608 ♂ Lysandra bellargus 1 1 1 1 1 14.08.1999 Pondel Val di Cogne 900 m Aosta Italy MW 99609 ♂ Lysandra coridon 1 1 14.08.1999 Pondel Val di Cogne 900 m Aosta Italy MW 99610 ♂ Lysandra coridon 1 1 14.08.1999 Pondel Val di Cogne 900 m Aosta Italy MW 99611 ♂ Lysandra coridon 1 1 14.08.1999 Pondel Val di Cogne 900 m Aosta Italy MW 99612 ♂ Lysandra coridon 1 1 1 1 1 14.08.1999 Pondel Val di Cogne 900 m Aosta Italy MW 99613 ♂ Agrodiaetus damon 1 1 1 1 1 II-36 17.08.1999 Col de Tende Tende 1850 m Alpes Maritimes France MW 99614 ♂ Agrodiaetus damon 1 1 17.08.1999 Col de Tende Tende 1850 m Alpes Maritimes France MW 99615 ♂ Agrodiaetus damon 1 1 17.08.1999 Col de Tende Tende 1850 m Alpes Maritimes France MW 99616 ♂ Agrodiaetus damon 1 1 17.08.1999 Col de Tende Tende 1850 m Alpes Maritimes France MW 99617 ♀ Agrodiaetus damon 1 1 17.08.1999 Col de Tende Tende 1850 m Alpes Maritimes France MW 99618 ♂ Lysandra coridon 1 1 17.08.1999 Col de Tende Tende 1850 m Alpes Maritimes France OK 96022 ♂ Lysandra caelestissimus 1 1 1 1 30.07.1996 Moscardon Montes Universales 1600 m Teruel Spain OK 99001 ♂ Lysandra gennargenti 1 1 1 29.07.1999 M. Perda Liana Lnusei 1150 m Sardinia Italy WE 00001 ♂ Agrodiaetus tenhageni 1 1 19.05.2000 25 km N Torbat-e-Heydariyeh Kuh-e-Sorkh 1700-1800 m Khorasan Iran WE 00002 ♂ Agrodiaetus glaucias 1 1 1 1 II-37 24.05.2000 Voluyeh südöstl. Sari 1500-1600 m Mazandaran Iran WE 00003 Neolysandra diana 11 02.06.2000 Kotayk, vill. Geghadir Armenia WE 02321 ♂ Agrodiaetus pseudoxerxes 1 1 1 13.07.2002 25 km W Fulad Mahalleh 2300 m Semnan Iran WE 02421 ♂ Agrodiaetus mofidii C P 1 1 15.07.2002 Chaman Bid 1700-2000m Khorasan Iran WE 02431 ♂ Agrodiaetus khorasanensis C P 1 1 1 16.07.2002 5 km SW Firizi 1700-1900m Khorasan Iran WE 02451 ♂ Agrodiaetus tenhageni 1 C P 1 1 1 II-38 17.07.2002 Kuh-e-Sorkh, Kadkan 2100-2500m Khorasan Iran WE 02452 ♂ Agrodiaetus tenhageni 1 C P 17.07.2002 Kuh-e-Sorkh, Kadkan 2100-2500m Khorasan Iran WE 02453 ♂ Agrodiaetus mofidii C P 17.07.2002 Kuh-e-Sorkh, Kadkan 2100-2500m Khorasan Iran WE 02454 ♂ Agrodiaetus mofidii 1 C P 1 1 II-39 17.07.2002 Kuh-e-Sorkh, Kadkan 2100-2500m Khorasan Iran WE 02491 ♂ Agrodiaetus achaemenes C P 1 1 1 21.07.2002 Gardaneh ye Cheri, W Samsami 2800-3000m Bakhtiari Iran WE 02492 ♂ Agrodiaetus achaemenes C P 21.07.2002 Gardaneh ye Cheri, W Samsami 2800-3000m Bakhtiari Iran WE 02531 ♂ Agrodiaetus zarathustra 1 C P 1 1 1 II-40 25.07.2002 Dorud, 30 km W Dorud (Paß) E-Seite 2100m Lorestan Iran WE 02532 ♂ Agrodiaetus zarathustra C P 25.07.2002 Dorud, 30 km W Dorud (Paß) E-Seite 2100m Lorestan Iran WE 02533 ♂ Agrodiaetus zarathustra 1 C P 25.07.2002 Dorud, 30 km W Dorud (Paß) E-Seite 2100m Lorestan Iran WE 02534 ♂ Agrodiaetus lorestanus 1 1 1 25.07.2002 Dorud, 30 km W Dorud (Paß) E-Seite 2100m Lorestan Iran WE 02535 ♂ Agrodiaetus lorestanus 1 1 1 1 II-41 25.07.2002 Dorud, 30 km W Dorud (Paß) E-Seite 2100m Lorestan Iran WE 02536 ♂ Agrodiaetus damalis 1 1 1 II-42 25.07.2002 Dorud, 30 km W Dorud (Paß) E-Seite 2100m Lorestan Iran WE 02591 ♂ Agrodiaetus peilei 1 C P 1 1 1 II-43 27.07.2002 Qamchiyan, 30 km N Chenareh 1800-1900m Kordestan Iran WE 02592 ♂ Agrodiaetus peilei 1 C P 27.07.2002 Qamchiyan, 30 km N Chenareh 1800-1900m Kordestan Iran WE 02593 ♂ Agrodiaetus peilei C P 27.07.2002 Qamchiyan, 30 km N Chenareh 1800-1900m Kordestan Iran WE 02611 ♂ Agrodiaetus karindus 1 C P 27.07.2002 40 km SW Saqqez 1800-1900m Kordestan Iran WE 02612 ♂ Agrodiaetus karindus 1 C P 1 1 II-44 27.07.2002 40 km SW Saqqez 1800-1900m Kordestan Iran WE 02613 ♂ Agrodiaetus karindus C P 27.07.2002 40 km SW Saqqez 1800-1900m Kordestan Iran WE 02614 ♂ Agrodiaetus morgani C P 1 1 1 27.07.2002 40 km SW Saqqez 1800-1900m Kordestan Iran WE 02621 ♂ Agrodiaetus sennanensis 1 1 1 28.07.2002 20 km E Mahabad 1900m Azarbayjan-e-Gharbi Iran WE 02622 ♂ Agrodiaetus sennanensis 28.07.2002 20 km E Mahabad 1900m Azarbayjan-e-Gharbi Iran WE 02661 ♂ Agrodiaetus arasbarani 1 1 1 29.07.2002 Mahmutabad, W 2200-2400m Azarbayjan-e-Sharqi Iran WE 02662 ♂ Agrodiaetus rovshani 1 1 1 29.07.2002 Mahmutabad, W Kaleybar 2200-2400m Azarbayjan-e-Sharqi Iran WE 02671 ♂ Agrodiaetus femininoides 1 C P 1 1 1 II-45 31.07.2002 Khalkhal, Gollijeh 1900m Ardabil Iran WE 02672 ♂ Agrodiaetus femininoides C P 31.07.2002 Khalkhal, Gollijeh 1900m Ardabil Iran WE 02673 ♂ Agrodiaetus femininoides C P 31.07.2002 Khalkhal, Gollijeh 1900m Ardabil Iran WE 02674 ♂ Agrodiaetus gorbunovi 1 C P 1 1 1 31.07.2002 Khalkhal, Gollijeh 1900m Ardabil Iran WE 02675 ♂ Agrodiaetus gorbunovi 1 C P 31.07.2002 Khalkhal, Gollijeh 1900m Ardabil Iran WE 02676 ♂ Agrodiaetus hamadanensis 1 C P 1 1 31.07.2002 Khalkhal, Gollijeh 1900m Ardabil Iran WE 02677 ♂ Agrodiaetus demavendi 1 1 1 31.07.2002 Khalkhal, Gollijeh 1900m Ardabil Iran WE 37001 Agrodiaetus mithridates 1 13.07.1995 Gündüzbey 1000-1100 m Malatya Turkey WE 38601 Agrodiaetus carmon 1 24.07.1995 7-15 km N Saimbeyli 1400-1600 m Adana Turkey WE 85001 ♂ Agrodiaetus shahrami 11123.07.2002 30 km N Chelgerd (Pass) 3000-3200m Bakhtiari Iran WE 92001 Agrodiaetus phyllides 1 20.04.1992 Kuschka 700 m Turkmenistan WE 92002 Agrodiaetus phyllides 1 20.04.1992 Kuschka 700 m Turkmenistan WE 94001 ♂ Agrodiaetus actinides 1111.07.1994 Aram-Kungei valley, Alytyn Dara river 3000 m West Transalai Kirgizia WE 94002 Plebeius cyane 1 15.07.1994 Chatir Kul, SW Naryn, Ak Bashi range Tianshan Kirgizia WE 98001 ♂ Agrodiaetus iphigenides 11 22.07.1998 25 km S Song Kul, Molto Tau 2000m Inner Tianshan Kirgizia WE 99001 Agrodiaetus barmifiruze 1 11.06.1999 Ardakan Yazd Iran

Sum 1895 1145 624 1877 370 6 312 25 202 95

Explanations: Code = individual code number used for all parts of a specimens (wings, karyological fixations & preparations, DNA-extraction, PCR and sequencing; in most cases made of collector initials, collecting year and a 3-digit number) Wgs = wing vouchers scanned Fix = chromosome fixations available (C) Kar = karyological preparations carried out (P) Alc = alcohol material available ('1'), otherwise dried material DNA = DNA extractions carried out Cytb = Cytochrome b sequenced CO1 = Cytochrome Oxidase I sequence ND1 = NADH dehydrogenase subunit 1 sequenced ITS2 = Internal Transcribed Spacer 2 sequenced Fig. = wings figured on plates Date = collecting date Locality =collecting locality (incl. altitude, Province, country

- 164 - Appendix 3: Sequence data Cytochrome Oxidase I (COI)

....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| 5 15 25 35 45 55 65 75 85 95 105 115 125 135 145 155 165 175 AD98001 TAGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATCG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGaACATCC CTAAGAATTT TAATTCGTAT AGAATTGAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA AD98009 NNNNNNNNNN NNNNNNNNNC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTAAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA AD98012 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTAAGA ACTCCTGGAT CTTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA AD98018 TAGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATCG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTGAGA ACTCCCGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA AD98024 TAGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATCG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAAT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTGAGA ACTCCCGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA AD98029 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATCTTT GGAATTTGAG CAGGAATAGT GGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTAAGA ACTCCTGGAT CCTTAATTGG AAATGATCAA ATTTATAATA CTATTGTCAC AGCCCATGCA AD98036 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GGACATTGTA TTTTATTTTT GGGATTTGAG CAGGAATAGT GGGAACATCC CTAAGAaTTT TAATTCGTAT AGAACTGAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTATTAC AGCCCATGCA DS00001 NNNNNNNNNN NNNNNNNNNN NNNNNNNNAT AAAGATATTG GAACGTTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT GGGAACATCC CTAAGAATTT TAATCCGTAT AGAATTAAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTACAAtA CTATTGTTAC AGCCCATGCA DS00005 NNNNNNNNNN NNNNNNNNTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT GGGAACATCT TTAAGAATTT TAATTCGTAT AGAATTAAGA ACTCCAGGAT CTTTAATTGG AGATGATCAA ATTTACAATA CTATTGTTAC AGCCCATGCA DS01001 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATCCGTAT AGAATTGAGA ACTCCTGGAT CCTTAATTGG AGACGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA JC00029 TAGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA CTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCA TTAAGAATTC TAATTCGTAT AGAATTGAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA JC00039 CAGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CGGGAATAGT AGGGACATCT TTAAGAATTT TAATTCGTAT AGAATTAAGA ACTCCTGAAT CcTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA JC00040 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCT CTAAGAATTT TAATTCGTAT GGAATTAAGA ACTCCTGGAT CCTTAATTGG AAATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA JC00042 TAGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATtATA CTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCA TTAAGAATTC TAATTCGTAT AGAATTGAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA JC00043 TCGCGAAAaT GACTTTTTTC CACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGGG CAGGAATAGT AGGaACATCT TTAAGAGTTT TAATTCGTAT GGAATTGAGA ACTCCAGGAT CTTTGATTGG AGATGATCAA ATTTATAATA CCATTGTCAC AGCTCATGCA JC00045 TCGCGAAAAT GACTTTTTTC CACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCT TTAAGAGTTT TAATTCGTAT GGAATTGAGA ACTCCAGGAT CTTTGATTGG AGATGATCAA ATTTATAATA CCATCGTTAC AGCTCATGCA JC00046 NNNNNAAAAT GACTTTTTTC CACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCT TTAAGAGTTT TAATTCGTAT GGAATTGAGA ACTCCAGGAT CTTTGATTGG AGATGATCAA ATTTATAATA CCATCGTTAC AGCTCATGCA JC00051 TAGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA CTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCA TTAAGAATTC TAATTCGTAT AGAATTGAGA ACTCCTGGAT CcTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA JC00055 NNNNNNNNAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTCATTTTT GGTATTTGAG CAGGAATAGT AGGAACATCT TTAAGAATTT TAATTCGTAT GGAATTAAGA ATTCCGGGAT CTTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA JC00057 TAGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGTATTTGGG CAGGAATGGT AGGAACATCT CTAAGAATTT TAATTCGTAT GGAATTAAGA ATTCCAGGAT CTTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA JC00061 TAGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA CTTTATTTTT GGAGTTTGAG CAGGAATAGT AGGAACATCA TTAAGAATTC TAATTCGTAT AGAATTGAGA ACCCCTGGAT CATTAATTGG AGATGATCAA ATTTATAATA CTATCGTTAC AGCTCATGCA JC00062 TAGCGAAAAT GACTTTTTTC NACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGTATTTGGG CAGGAATGGT AGGAACATCT CTAAGAATTT TAATTCGTAT GGAATTAAGA ATTCCAGGAT CTTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA JC00063 TAGCGAAAAT GACTTTTTTC NACAAATCAT AAAGATATTG GAACATTATA CTTTATTTTT GGAGTTTGAG CAGGAATAGT AGGAACATCA TTAAGAATTC TAATTCGTAT AGAATTGAGA ACACCTGGAT CATTAATTGG AGATGATCAA ATTTATAATA CTATCGTTAC AGCTCATGCA JC01014 NNNNNNNNNN NNNNNNNNNN NNCAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGGG CAGGAATAGT AGGAACATCT TTAAGAGTTT TAATTCGTAT AGAATTGAGA ACCCCAGGAT CTTTGATTGG AGATGATCAA ATTTATAATA CCATCGTTAC AGCTCATGCA JC02001 TNGCGAAAAT GACTTTATTC TACAAATCAT AAAGATATTG GAACATTATA TTTCATTTTT GGAATTTGAT CAGGAATAGT AGGAACTTCA TTAAGAATAA TTATTCGAAC TGAATTAGGT AATCCAGGTT CTTTAATTGG AGATGATCAA ATTTATAATA CTATTGTAAC AGCACATGCT JM00001 NNNNNNNNNN NNNNNNNNNN NNNAAATCAt AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCC CTAAGAaTTT TAATTCGTAT GGAATTAAGA ACTCCNGGAT CCTTAATTGG AAATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCG MW00001 TAGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATCG GAACATTATA TTTTATTTTT GGAGTTTGAG CAGGGATAGT AGGAACATCT CTAAGAATTT TAATTCGTAT AGAATTGAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA MW00015 TCGCGAAAAT GACTTTTTTC CACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT GGGAACATCT TTAAGAGTTT TAATTCGTAT GGAATTGAGA ACTCCAGGAT CTTTGATTGG AGATGATCAA ATTTATAATA CCATCGTTAC AGCTCATGCA MW00017 TAGCGAAAAT GAaTTTATTC AACAAATCAT AAAGATATTG GAACTTTATA TTTTATTTTT GGAATTtGAG cTGGAATAGT TGGAACCTCT TTAAGaATTT TAATTCGTCT TGAATtAGGT AcCCCAGGAT CTTTAATtGG TGATGATCaA ATCTATAATA CTATTGtAAC TGCTCATGCT MW00020 TAGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGTATTTGGG CAGGAATGGT AGGAACATCT CTAAGAATTT TAATTCGTAT GGAATTAAGA ATTCCAGGAT CTTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA MW00024 TAGCGAAAAT GACTTTTTTC tACAAATCAT AAAGATATTG GAACATTATA TTTTATCTTT GGAATTTGAG CCGGAATAGT AGGAACATCT TTAAGAATTT TAATTCGTAT AGAACTAAGA ATTCCTAGAT CTTTAATTGG TGATGATCAA ATTTATAATA CTATTGTAAC AGCTCACGCA MW00032 TAGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATCG GAACATTATA TTTTATTTTT GGAGTTTGAG CAGGGATAGT AGGAACATCT CTAAGAATTT TAATTCGTAT AGAATTGAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA MW00044 TAGCGAAAAT GACTTTTTTC AACAAATCAT AAAGATATTG GAACTTTATA TTTTATTTTT GGAATTTGAG CTGGAATAGT TGGAACTTCT TTAAGAATTC TAATTCGTCT TGAATTAGGA ACTCCAGGAT CTTTAATTAG AGATGATCAA ATTTATAACA CTATTGTAAC TGCTCATGCT MW00051 TAGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATCG GAACACTATA TTTTATTTTT GGAGTTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTGAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTCAC AGCTCATGCA MW00056 NNNNNNNNNN NNNNNNNNNN NNNNAATCAT AAAGATATCG GNACACTATA TTTTATTTTT GGAGTTTGAG CAGGAATAGT AGGAACATCC NTAAGAATTT TAATTCGTAT AGAATTGAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTCAC AGCTCATGCA MW00058 NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN AAAGATATCG GAACACTATA TTTTATTTTT GGAGTTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTGAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTCAC AGCTCATGCA MW00059 TAACGAAAAT GACTTTTTTC TACAAATCAT AAAGATATCG GAACACTATA TTTTATTTTT GGAGTTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTGAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTCAC AGCTCATGCA MW00060 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GGACATTATA TTTTATTTTT GGAGTTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTAAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAACA CTATTGTTAC AGCTCATGCA MW00064 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGTATTTGGG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT GGAATTAAGA ACTCCCGGAT CTTTAATTGG AAATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA MW00072 NNNNNNNNNN NNNNNNNNNN NNNNAATCAT AAAGATATTG GGACATTATA TTTTATTTTT GGAGTTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTAAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAACA CTATTGTTAC AGCTCATGCA MW00076 TGGCGAAAAT GACTTTTTTC CACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT GGGAACATCC TTAAGAATTT TGATTCGTAT GGAATTAAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA MW00087 NNNNNNNNNN GACTTTTTTC CACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT GGGAACATCC TTAAGAATTT TGATTCGTAT GGAATTAAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA MW00101 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GGACATTATA TTTTATTTTT GGAGTTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTGAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAACA CTATTGTTAC AGCCCATGCA MW00102 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GGACATTATA TTTTATTTTT GGAGTTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTAAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAACA CTATTGTTAC AGCTCATGCA MW00110 NNNNNNNNNN NNNNNNNNNN NNNNAATCAT AAAGATATCG GAACACTATA TTTTATTTTT GGAGTTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTGAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTCAC AGCTCATGCA MW00116 TAGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CGGGAATAGT AGGAACATCT TTAAGAATCT TAATTCGAAT AGAATTAGGA ACTCCTGGAT CTTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA MW00119 TAGCGAAAAT GACTTTTTTC AACAAATCAT AAAGATATTG GAACTTTATA TTTTATTTTT GGAATTTGAG CTGGAATAAT TGGAACTTCC TTAAGAATTT TAATTCGCCT TGAATTAGGT ACTTCAGGGT CTTTAATtGG AGAtGATCaA ATTTATAATA CTATCGTAAC TGCTCATGCT MW00125 TAGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTAAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA MW00127 TAGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATCG GAACACTATA TTTTATTTTT GGAGTTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTGAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTCAC AGCTCATGCA MW00129 TCGCGAAAAT GACTTTTTTC TNNNAATCAT AAAGATATTG GaaCATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGt AGGAACATCA CTAAGAATTT TAATTCGTAT AGAATTAAGA ACCCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA MW00140 NNNNNNNNNN NNNTTTTTTC TACAAATCAT AAAGATATTG GGACATTGTA TTTTATTTTT GGGATTTGAG CAGGAATAGT GGGAACATCC CTAAGAATTT TAATTCGTAT AGAACTGAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTATTAC AGCTCATGCA MW00151 TAGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT GGGAACATCC CTAAGAATtT TAATTCGTAT AGAATTAAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA MW00157 TAGGGAAAAT GACTTTTTTC TACAAATCAT AAAGATATCG GAACACTATA TTTTATTTTT GGAGTTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTGAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTCAC AGCTCATGCA MW00176 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCC CTTAGAATTT TAATCCGTAT AGAATTAAGA ACTCCTGGAT CTTTGATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA MW00177 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCA CTAAGAATTT TAATTCGTAT AGAATTAAGA ACCCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA MW00178 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCA CTAAGAATTT TAATTCGTAT AGAATTAAGA ACCCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA MW00179 TCGCGAAAAT GACTTTTTTC TACAAACCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCA CTAAGAATTT TAATTCGTAT AGAATTAAGA ACCCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA MW00183 TCGCGAAAAT GACTTTTTTC CNCAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT GGGAACATCT TTAAGAGTTT TAATTCGTAT GGAATTGAGA ACTCCAGGAT CTTTGATTGG AGATGATCAA ATTTATAATA CCATCGTTAC AGCTCATGCA MW00185 TCGNGAAAAN GACTTTTTTC CACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT GGGAACATCT TTAAGAGTTT TAATTCGTAT GGAATTGAGA ACTCCAGGAT CTTTGATTGG AGATGATCAA ATTTATAATA CCATCGTTAC AGCTCATGCA MW00186 TCGCGAAAAT GACTTTTTTC CACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT GGGAACATCT TTAAGAGTtT TAATTCGTAT GGAATTGAGA ACTCCAGGAT CTTTGATTGG AGATGATCAA ATTTATAATA CCATCGTTAC AGCTCATGCA MW00189 NNNNNNNNNN NNNNNNNNNN NNCAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGGG CAGGAATGGT AGGAACATCT TTAAGAGTTT TAATTCGTAT GGAATTGAGA ACTCCAGGAT CTTTAATTGG AGATGATCAA ATTTATAATA CCATTGTTAC AGCTCATGCA MW00226 TAGCGAAAAT GACTTTTTTC NACAAATCaT AAAGATATTG GAACATTATA TTTTATTTTT GGAGTTTGAG CAGGAATAGT AGGAACATCC TTAAGAATTC TAATTCGTAT AGAATTGAGA ACTCCGGGAT CCTTAATTGG AGACGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA MW00229 NNNNNNNNNN NNNNNNNNNN NNNNAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGGGTTTGAG CAGGAATAGT GGGAACATCT CTAAGAATTT TAATTCGTAT GGAATTAAGA ACTCCCGGAT CCTTAATTGG GAATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA MW00231 NGGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGGGTTTGAG CAGGAATAGT GGGAACATCT CTAAGAATTT TAATTCGTAT GGAATTAAGA ACTCCCGGAT CCTTAATTGG GAATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA MW00232 TAGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATCG GAACACTATA TTTTATTTTT GGAGTTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTGAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTCAC AGCTCATGCA MW00234 NNNNNNNNNN NNNNNNNNNN NNNNNNNCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTAAGA ACTCCTGGAT CTTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA MW00259 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GGACATTATA CTTTATTTTT GGAATTTGAG CAGGAATAGT GGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTGAGA ACTCCTGGAT CTTTAATTGG TGATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA MW00262 NNNNNNNNNN NNNTTTTTTC TACAAATCAT AAAGATATTG GGACATTATA CTTTATTTTT GGAATTTGAG CAGGAATAGT GGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTGAGA ACTCCTGGAT CTTTAATTGG TGATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA

165 Appendix 3: Sequence data Cytochrome Oxidase I (COI)

....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| 5 15 25 35 45 55 65 75 85 95 105 115 125 135 145 155 165 175 MW00267 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GGACATTATA TTTTATTTTT GGAGTTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTAAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAACA CTATTGTTAC AGCTCATGCA MW00269 TAGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTGTTTTT GGAATTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTGAGA ATTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA MW00290 TAGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GGACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT GGGAACATCC TTAAGAATTT TAATTCGTAT AGAATTAAGA ACCCCTGGAT TCTTAATTGG AGATGATCAA ATTTACAATA CTATTGTTAC AGCTCATGCA MW00302 TTGCGAAAAT GACTTTTTTC CACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCC TTAAGAATTT TAATTCGTAT AGAATTGAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAACA CTATTGTTAC AGCTCATGCA MW00316 NNNNNNNNNN NNNNNNNNTC TACAAATCAT AAAGATATCG GAACACTATA TTTTATTTTT GGAGTTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTGAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTCAC AGCTCATGCA MW00326 TCGCGAAAAT GATTATATTC AACAAATCAT AAAGACATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCA CTAAGAATTT TAATTCGAAT AGAATTAAGA ACCCCAGGAT CATTAATTGG TGATGACCAA ATTTATAATA CTATTGTCAC AGCTCATGCA MW00327 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GGACATTATA TTTTATTTTT GGTATTTGAG CAGGGATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT GGAATTGAGA ACCCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA MW00328 TAGCGAAAAT GACTTTTTNN NNNNAATCAT AAAGATATTG GAACATTATA TTTTGTTTTT GGAATTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTGAGA ATTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA MW00330 TCGCGAAAAT GACTTTTTTC TACAAACCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCA CTAAGAATTT TAATTCGTAT AGAATTAAGA ACCCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA MW00335 NNNNNNNNNN NNNNNNTTTC TACAAATCAT AAAGATATTG GGACATTATA TTTTATTTTT GGAGTTTGAG CGGGAATAGT GGGAACATCC CTAAGAATCT TAATTCGTAT AGAATTAAGA ACTCCTGGGT CCTTAATCGG AGATGATCAA ATTTATAACA CTATTGTTAC AGCTCATGCA MW00347 NNNNNNNNNT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CGGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTGAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAACA CTATTGTTAC AGCCCATGCA MW00348 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GGACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAACTGAGA ACTCCGGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTATTAC AGCCCATGCA MW00358 NNNNNNNNNN NNNNNNNNNN NNNNAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CGGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTGAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAACA CTATTGTTAC AGCCCATGCA MW00393 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GGACATTATA TTTTATTTTT GGTATTTGAG CAGGGATAGT AGGAACATCC TTAAGAATTT TAATTCGTAT GGAATTGAGA ACCCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA MW00409 TCGCGAAAAT GACTTTTTTC NNNNAATCAT AAAGATATTG GNACATTATA TTTTATTTTT GGAGTTTGAG CGGGAATAGT GGGAACATCC CTAAGAATCT TAATTCGTAT AGAATTAAGA ACTCCTGGGT CCTTAATCGG AGATGATCAA ATTTATAACA CTATTGTTAC AGCTCATGCA MW00412 TAGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA CTTTATTTTT GGAGTTTGAG CAGGAATAGT AGGAACATCA TTAAGAATTC TAATTCGTAT AGAATTGAGA ACCCCTGGAT CATTAATTGG AGATGATCAA ATTTATAATA CTATCGTTAC AGCTCATGCA MW00424 NNNNNNNNNN NNNNNNNNNN NNNNNNNNAT AAAGATATTG GAACATTATA TTTTGTTTTT GGAATTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTGAGA ATTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA MW00444 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GGACATTATA TTTTATTTTT GGAGTTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTAAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAACA CTATTGTTAC AGCTCATGCA MW00452 NNNNNNNNNN NNNNTTTTTC TACAAATCAT AAAGATATTG GGACATTAtA TTTTATTTTt GGAATTTGAG CAGGAATAGT GGGAACAtCC CTAAGAATTT TAATTCGTAT AGAGCTGAGA ACTCCGGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTATTAC AGCCCATGCA MW00469 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA CTTTATTTTT GGAATTTGAG CGGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTAAGA ACCCCTGGAT CTTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA MW00476 NNNNNNNNNN NNNNNNNNNC TACAAATCAT AAAGATATTG GGACATTATA TTTTATTTTT GGAGTTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTAAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAACA CTATTGTTAC AGCTCATGCA MW00497 TAGCGAAAAT GACTTTTTTC CACAAATCAT AAAGATATTG GAACTTTATA TTTTATTTTT NNNNTTTGAG CAGGAATATT AGGAACATCT TTAAGAATTT TAATTCGTAT AGAATTAGGA ACTCCTGGAT CTTTAATTGG TAATGATCAA ATTTATAATA CAATTGTTAC AGCTCATGCA MW00498 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGTATTTGGG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT GGAATTAAGA ACTCCCGGAT CTTTAATTGG AAATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA MW00504 TAGCGAAAAT GACTTTTTTC tACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTAAGA ACTCCTGGAT CTTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA MW00517 TAGCGAAAAT GACTTTTTTC CACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCT TTAAGAATTT TAATTCGAAT AGAATTAAGA ACACCTGGAT CTTTAATTGG TGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA MW00525 TAGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT GGGAACATCT TTAAGAATCT TAATTCGTAT AGAATTAAGA ACTCCTGGAT CTTTAATTGG TGATGATCAA ATTTATAACA CCATTGTAAC AGCTCATGCT MW00530 TAGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA CTTTATTTTT GGAGTTTGAG CAGGAATAGT GGGAACATCA TTAAGAATTC TAATTCGTAT AGAATTGAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA MW00539 TAGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTGAGA ATTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA MW00547 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GGACATTATA TTTTATTTTT GGAGTTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTAAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAACA CTATTGTTAC AGCTCATGCA MW01001 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT GGAATTAAGA ACTCCTGGAT CCTTAATTGG AAATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCG MW01009 NNNNNNNNNT GANTNNNTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CGGGAATAGT AGGAACATCC TTAAGAATTT TAATTCGTAT AGAATTAAGA ACTCCTGAAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA MW01011 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA CTTTATTTTT GGAATTTGAG CGGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTAAGA ACCCCTGGAT CTTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA MW01014 TCGCGAAAAT GACTTTTTTC CACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGGG CAGGAATAGT AGGAACATCT TTAAGAGTTT TAATTCGTAT GGAATTGAGA ACTCCGGGAT CTTTGATTGG AGATGATCAA ATTTATAATA CCATCGTTAC AGCTCATGCA MW01018 TTGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGGATTTGAG CGGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTAAGA ACCCCTGGAT CTTTGATTGG GGATGATCAA ATTTATAACA CTATTGTTAC AGCTCATGCA MW01019 NNNNNNNNNN NNNNNNNNNN NNNAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCC TTAAGAATTT TAATTCGTAT AGAATTGAGA ACTCCTGAAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTCAC AGCTCATGCA MW01022 TAGCGAAAAT GACTTTTTTC AACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATATT GGGAACATCA TTAAGAATTT TAATTCGAAT AGAGTTAGGA ACTCCAGGAT CTTTAATTGG TGATGATCAA ATTTATAATA CTATTGTAAC AGCTCATGCT MW01023 TAGCGAAAAT GACTTTTTTT CACAAATCAT AAAGATATTG GAACTTTATA TTTTATTTTT GGAATTTGAA GAGGAATATT AGGAACTTCA TTAAGAATTT TAATTCGAAT AGAATTAGGA ACACCAGGAT CTTTTATTGG TGATGATCAA ATTTATAATA CAATCGTCAC AGCTCATGCT MW01025 TAGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA CTTTATTTTT GGAATTTGAG CGGGAATAGT AGGAACATCA TTAAGAATTC TAATTCGTAT AGAATTGAGA ACCCCTGGAT CATTGATTGG AGATGATCAA ATTTATAACA CTATCGTTAC AGCTCATGCA MW01027 AAGCGAAAAT GACTTTTTTC TACAAACCAT AAAGATATTG GAACTTTATA TTTTATTTTT GGAATTTGAG CAGGAATACT AGGAACATCT TTAAGAATTC TAATTCGAAT AGAATTAGGA ACACCAGGAT CTTTAATTGG AGATGATCAA ATTTATAATA CCATCGTAAC AGCTCATGCT MW01034 NNNNNNNNNT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CGGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTAAGA ACCCCTGGAT CTTTGATTGG AGATGATCAA ATTTATAACA CTATTGTTAC AGCTCATGCA MW01039 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT GGAATTAAGA ACTCCTGGAT CCTTAATTGG AAATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCG MW01048 NNNNNNNNNN NNCTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTCATTTTT GGtATTTGAG CAGGAATAGT AGGaACATCT TTAAGAaTTT TAATTCGTAT GGAATTAAGA ATTCCAGGAT CTTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA MW01053 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCT CTAAGAATTT TAATTCGTAT GGAATTAAGA ACTCCTGGAT CCTTAATTGG AAATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCG MW01059 TTGGGAAAAN AACTTTTTTN TAAAAATCAT AAAGATATTG GAACNTTATN TTTTATTTTT GAAATTTGAG CNGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTAAGA ACCCNTGGAT CTTTGATTGG AGATGATCAA ATTTATAACA CTATTGTTAC AGCTCATGCA MW01061 NNNNNNNNNN NNNNNNNNNN TACAAATCAT AAAGATaTTG GAACATTATA TTTCATTTTT GGTATTTGAG CAGGAATAGT AGGAACAtCC TTAAGAATTT TAATTCGTAT AGAATTAAGA ACTCCTGGAT CTTTAATTGG AGATGATCAA ATTTATAACA CTATTGTTAC AGCTCATGCA MW01072 TCGCGAAAAT GACTTTTTTC CACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGGG CAGGAATAGT AGGAACATCT TTAAGAGTTT TAATTCGTAT GGAATTGAGA ACTCCGGGAT CTTTGATTGG AGATGATCAA ATTTATAATA CCATCGTTAC AGCTCATGCA MW01078 NNNNNNNNNN NNNNNNNNTC TACAAATCAT ACAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCC CTAAGAGTTT TAATTCGTAT GGAATTAAGA ACTCCTGGAT CCTTAATTGG AAATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCG MW01083 TTGCGAAAAT GACTTTTTTC CaCAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCC TTAAGAATTT TAATTCGTAT AGAATTGAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAACA CTATTGTTAC AGCTCATGCA MW01092 NNGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGGATTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTAAGA ACCCCTGGAT CTTTAATTGG AGATGATCAA ATTTATAACA CTATTGTTAC AGCTCATGCA MW01103 TTGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGGATTTGAG CAGGAATAGT GGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTAAGA ACTCCTGGAT CTTTGATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCACGCA MW01105 NNNCGAAAAT GACTTTTTTC CACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGGG CAGGAATAGT AGGAACATCT TTAAGAGTTT TAATTCGTAT GGAATTGAGA ACTCCAGGAT CTTTAATTGG AGATGATCAA ATTTATAATA CCATCGTTAC AGCTCATGCA MW01107 NNNNNNNNAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT GGGAACATCT CTAAGAATTT TAATTCGTAT GGAATTAAGA ACTCCTGGAT CCTTAATTGG AAATGATCAA ATTTATAACA CTATTGTTAC AGCCCATGCG MW01114 ATGCGAAAAT GACTTTATTC TACAAATCAT AAAGATATTG GAACATTATA TTTCATTTTT GGAATTTGAT CAGGAATAGT AGGAACTTCA TTAAGAATAA TTATTCGAAC TGAATTAGGT AATCCAGGTT CTTTAATTGG AGATGATCAA ATTTATAATA CTATTGTAAC AGCACATGCT MW01116 TTGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGGATTTGAG CGGGAATAGT GGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTAAGA ACCCCTGGAT CTTTGATTGG GGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA MW01120 TAGCGAAAAT GACTTTTTTC AACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATATT AGGAACATCT TTAAGAATCT TAATTCGTAT AGAATTAGGA AATCCCGGAT CTTTAATTGG AGACGATCAA ATTTACAATA CTATTGTTAC AGCTCATGCT MW02001 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCC TTAAGAATTT TAATTCGTAT AGAATTGAGA ATTCCTGGAT TCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA MW02006 TAGCGAAAAT GATTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGGATAGT GGGAACATCC TTAAGAATTC TAATTCGTAT AGAATtAAGA ACTCCTGGAT CTTTAATCGG AGATGATCaA ATTTATAATA CTATTGTtAC AGCTCATGCA MW02007 NAGCGAAAAT GAATTTATTC AACAAATCAT AAAGATATTG GAACTTTATA TTTTATCTTT GGAATTTGAG CTGGAATAGT AGGAACCTCT TTAAGAATTT TAATTCGTCT TGAATTAGGT ACCCCAGGAT CTTTAATTGG TGATGATCAA ATCTATAATA CTATTGTAAC TGCCCATGCT MW02008 TAGCGAAAAT GACTTTTTTC AACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTC GGGATTTGAG CAGGTATATT AGGAACATCA TTAAGAATCT TAATTCGTAT AGAGTTAGGA ACACCGGGAT CATTAATTGG AGATGATCAA ATTTATAATA CTATTGTCAC AGCTCATGCT MW02009 TTGCGAAAAT GACTTTTTTN NACAAATCAT AAAGATATTG GAACTTTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACCTCC TTAAGAATTT TAATTCGTCT TGAATTAGGT ACTCCAGGAT CTTTAATTGG TGATGATCAA ATTTATAATA CTATTGTAAC TGCACATGCC MW02021 NNNNNNNNNT GATTATTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATATT AGGAACATCA TTAAGAATTT TAATTCGAAT AGAATTAGGT ACTCCTGGAT CATTAATTGG TGATGATCAA ATTTATAATA CTATTGTAAC AGCTCATGCT MW02024 NNNNNNNAAT GACTTTATTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAaTTTGAT CAGGAATAGT AGGAACTTCA TTAAGAATAA TTATTCGAAC TGAATTAGGT AATCCAGGTT CTTTAATTGG AGATGATCAA ATTTATAATA CTATTGTAAC AGCACATGCT MW02025 NNNNNNNNNN NACTTTTTTC AACAAATCAT AAAGATATTG GAACATTATA TTTTAtTTTT GGAATTTGGG CAGGAATATT AGGAACATCT TTAAGAATTT TAATTCGTAT AGAATTAGGA ACACCCGGAT CATTAATTGG AGATGATCAA ATTTATAATA CTATTGTAAC AGCTCATGCT MW02028 TAGCGAAAAT GACTTTTTTC AACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATATT GGGAACATCA TTAAGAATTT TAATTCGAAT AGAGTTAGGA ACTCCAGGAT CTTTAATTGG TGATGATCAA ATTTATAATA CTATTGTAAC AGCTCATGCT MW02031 NNNNNNNNNN NACTTTTTTC AACAAATCAT AAaGATATTG GAaCTTTATA TTTTATTTTT GGAATTTGAG caGGAATATT AGGAaCATCT TTAAGaATTT TAATTCGTAT AGAATTAGGA ACACCTGGAT CATTAATTGG AGATGATCaA ATTTATAATA CTATTGTAAC AGCTCATGCT MW02033 TAGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTCATTTTT GGTATTTGAG CAGGAATAGT AGGAACATCT TTAAGAATTT TAATTCGTAT GGAATTAAGA ATTCCAGGAT CTTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA MW02034 TAGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCT TTAAGAATCT TAATTCGTAT AGAATTAAGA ACTCCTGGAT CTTTAATTGG TGATGATCAA ATTTATAATA CCATTGTAAC AGCTCATGCT MW98002 TAGCGAAAAT GAnTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATATT AGGAACATCT TTAAGAATTT TAATTCGAAT AGAATTAGGA ACCCCAGGAT CATTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCT

166 Appendix 3: Sequence data Cytochrome Oxidase I (COI)

....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| 5 15 25 35 45 55 65 75 85 95 105 115 125 135 145 155 165 175 MW98008 TTGCGAAAAT GACTTTTTTC NACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCC TTAAGAATTT TAATTCGTAT AGAATTGAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAACA CTATTGTTAC AGCTCATGCA MW98009 TAGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATCG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCC TTAAGAATTT TAATTCGTAT AGAATTGAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA MW98020 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAGTTTGAG CAGGAATAGT GGGAACATCT CTAAGAATTT TAATTCGTAT GGAATTAAGA ACCCCCGGAT CCTTAATTGG GAATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA MW98029 TTGCGAAAAT GACTTTTTTC CACAAATCAT AAAGATATTG GGACATTATA TTTTATTTTT GGAATTTGAG CAGGAAtAGt AGGAACATCC TTAAGAATTT TGATTCGTAT GGAATTAAGA ACTCCTGGAT TCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA MW98049 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATCTTT GGAATTTGAG CAGGAATAGT GGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTAAGA ACTCCTGGAT CCTTAATTGG AAATGATCAA ATTTATAATA CTATTGTCAC AGCCCATGCA MW98079 TCTCGAAAAT GACTTTTTTC TACaAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT GGGAACATCC CTAAGAATTT TAATTCGTAT GGAATTAAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA MW98084 TCGCGAAAAT GACTTTTTTC CACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGGG CAGGAATAGT AGGAACATCT TTAAGAGTTT TAATTCGTAT AGAATTGAGA ACTCCAGGAT CTTTGATTGG AGATGATCAA ATTTATAATA CCATCGTTAC AGCTCATGCA MW98089 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT GGGAACATCC CTAAGAATTT TAATTCGTAT GGAATTAAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA MW98094 TAGCGAAAAT GACTTTTTTC AACAAATCAT AAAGATATTG GAACTTTATA TTTTATTTTT GGAATTTGAG CTGGAATAAT TGGAACTTCT TTAAGAATTT TAATTCGTCT TGAATTAGGA ACACCAGGAT CTTTAATTGG AAATGATCAA ATTTATAATA CTATTGTTAC TGCTCATGCT MW98097 TAGGGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT GGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTAAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA MW98103 TCGCGAAAAT GACTTTTTTC TACAaATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATCCGTAT AGAATTAAGA ACTCCTGGAT CTTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA MW98106 NNNNNNNNNN NNNNNNNNNN NACAAATCAT AAAGATATTG GAACATTATA TTTTATCTTT GGAATTTGAG CAGGAATAGT GGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTAAGA ACTCCTGGAT CCTTAATTGG AAATGATCAA ATTTATAATA CTATTGTCAC AGCCCATGCA MW98128 TAGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGG AGGAACATCC TTAAGAATTT TAATTCGTAT AGAATTGAGA ACTCCTGAAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTCAC AGCTCATGCA MW98129 TTGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACaTTATA TTTTANTTTT GGAATTTGAG CAGGAATAAT AGGAACATCC TTAAGAATTT TAATTCGTAT AGAATTAAGA ACCCCTGGAT CTTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA MW98133 TAGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA CTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCA TTAAGAATTC TAATTCGTAT AGAATTGAGA ACCCCTGGAT CATTAATTGG AGATGATCAA ATTTATAATA CTATCGTtAC AGCTCATGCA MW98136 TCGCGAAAaT GACTTTTTTC TNCAAATCAT AAAGATATTG GaACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTAAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA MW98138 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAGTTNGAG CAGGAATAGT GGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTAAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA MW98139 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTAAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA MW98154 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAGTTTGAG CAGGAATAGT GGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTAAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA MW98162 TAGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTAAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA MW98170 TCGCGAAAAT GACTTTTTTC TACAAATCaT AAAGATATTG GaACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCC CTAAGaATTT TAATTCGTAT AGAATTAAGA aCTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA MW98172 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT GGGAACATCT CTAAGAATTT TAATTCGTAT GGAATTAAGA ACCCCTGGAT CCTTAATCGG GAATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA MW98180 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGGATTTGAG CAGGAATAGT GGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTAAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA MW98183 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGNNTTTGAG CAGGAATAGT GGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTAAGA ACTCCTGGAT CCTTAATTGG AAATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA MW98185 NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NAATTCGTAT AGAATTAAGA ACCCCTGGAT CTTTAATTGG AGATGATCAA ATTTATAACA CTATTGTTAC AGCTCATGCA MW98189 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAGTTTGAG CAGGAATAGT GGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTAAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA MW98203 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTAAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA MW98205 TAGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATCG GAACATTATA TTTTATTTTT GGAGTTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTGAGA ACTCCTGGAT CTTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA MW98212 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAGTTTGAG CAGGAATAGT GGGAACATCT CTAAGAATTT TAATTCGTAT GGAATTAAGA ACTCCCGGAT CCTTAATTGG GAATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA MW98220 TAGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAAcaTTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCT TTAAGAATCT TAATTCGTAT AGAATTAAGA ACTCCTGGAT CTTTAATTGG tGATGATCAA ATTTATAATA CTATCGTTAC AGCTCATGCA MW98228 TTGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CGGGAATAGT AGGAACATCC TTAAGAATTT TAATTCGTAT AGAATTAAGA ATCCCTGGAT CTTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA MW98230 TAGCGAAAAT GATTTTTTTC AACAAATCAT AAAGATATTG GAANTTTATA TTTTATTTTT GGAATTTGAG CTGGAATAAT TGGAACTTCT TTAAGAATTT TAATTCGTCT TGAATTAGGT ACTCCAGGAT CTTTAATTGG AGATGATCAA ATCTATAATA CTATTGTAAC TGCTCATGCT MW98235 TCGCGAAAAT GATTTTTTTC CACAAATCAT AAAGACATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGATT AGAGCTAAGG ACCCCAGGAT CATTAATTGG TGACGACCAA ATTTATAATA CTATTGTCAC AGCCCATGCA MW98240 CAGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATCG GAACATTATA TTTTATTTTT GGTNTTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTGAGA ACTCCTGGAT CTTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA MW98261 NNNNNNNNNN NNNNNNNNNN TACAAATCAT AAAGATATCG GAACATTATA TTTTATTTTT GGAGTTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTGAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA MW98264 TCGCGAAAAT GATTTTTTTC CACAAATCAT AAAGACATTG GAACATTATA TTTTATTTTT GGGNTTTGAG CAGGAATAGT AGGAACATCA CTAAGAATTT TAATTCGATT AGAGCTAAGA ACCCCAGGAT CATTAATTGG TGATGACCAA ATTTACAATA CTATTGTCAC AGCCCATGCA MW98265 TAGCGAAAAT GACTTTTTTN NNNAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATATT GGGAACATCA TTAAGAATTT TAATTCGAAT AGAGTTAGGA ACTCCAGGAT CTTTAATTGG TGATGATCAA ATTTATAATA CTATTGTAAC AGCTCATGCT MW98270 TAGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA CTTTATTTTT GGTATTTGAG CAGGAATAGT AGGAACATCA TTAAGAATTT TAATTCGTAT AGAATTGAGA ACTCCAGGGT CTTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA MW98274 TTGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCC TTAAGAATTT TGATTCGTAT GGAATTAAGA ACTCCTGGAT TCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA MW98278 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA CTTTATTTTT GGAATTTGAG CGGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTAAGA ACCCCTGGAT CTTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA MW98284 TAGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTAAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA MW98285 NNNNNNNNNN NNCTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTAAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA MW98294 TAGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATCG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTAAGA ACTCCTGGAT CTTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA MW98313 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAaCATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTAAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA MW98315 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAGTTTGAG CAGGAATAGT GGGAACATCT CTAAGAATTT TAATTCGTAT GGAATTAAGA ACTCCCGGAT CCTTAATTGG GAATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA MW99001 TAGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATCG GAACATTATA TTTCATTTTT GGTATTTGAG CAGGAATAGT AGGAACATCT TTAAGAATTT TAATTCGTAT GGAATTAAGA ATTCCAAGAT CTTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA MW99006 TAGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTAAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA MW99009 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATCTTT GGAATTTGAG CAGGAATAGT GGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTAAGA ACTCCTGGAT CCTTAATTGG AAATGATCAA ATTTATAATA CTATTGTCAC AGCCCATGCA MW99013 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAAGTTTATA TTTTAATTTT GGNNTTTGAG CAGGAATAGT AGGAACATCT TTAAGAATTT TAATTCGAAT GgAATTAAGA ACTCCTGGAT CTTTAATTGG AGANGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA MW99014 TAGCGAAAAT GACTTTTTTC cACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCC TTAAGAATTT TAATTCGTAT AGAATTAAGA ACCCCTAAAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTCAC AGCTCATGCA MW99018 TAGCGAAAAT GACTTTTTTC AACAAATCAT AAAAATATTG GAACATTATA CTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCT TTAAGAATTT TAATTCGTAT GGAATTAAGA ACCCCTGGAT CTTTAATCGG AGATGATCAA ATTTATAATA CCATTGTTAC AGCTCATGCA MW99028 NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNCGTAT GGAATTAAGA ATtCCAGGAT CTTTaATTGG AGATGaTCAA ATTTATAATA CTATTGTTAC AGCTCATGCA MW99038 NNNNNAAAAT GATTTTTTTC CACAAATCAT AAAGACATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGATT AGAGCTAAGG ACCCCAGGAT CATTAATTGG TGATGACCAA ATTTATAATA CTATTGTCAC AGCCCATGCA MW99042 TTGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAAcaTTATA TTTTAATTTT GGAATTTGAG CAGGAATAGT AGGAACATCC TTAAGAATTT TAATTCGTAT AGAATTAAGA ACCCCTGGAT CTTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA MW99045 NNNNNAAAAT GACTTTTTTC AACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATATT AGGAATATCT TTAAGAATTT TAATTCGAAT AGAATTAAGA ACTCCAGGAT ATTTAATTGG AAATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCT MW99047 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCT TTAAGAATTT TAATTCGTAT AGAGTTAAGA GTTCCTGGAT CTTTAATTGG AGATGATCAA ATTTATAACT CTATTGTTAC AGCTCATGCA MW99053 TAGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATCG GAACATTATA TTTTATTTTT GGANTTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTGAGA ACTCCCGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCACGCA MW99057 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTAAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA MW99058 TAGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATttGAG CAGGAATAGT GGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTAAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA MW99059 NNNNNNNNNN NNCTTTTTTC TACAAACCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCA CTAAGAATTT TAATTCGTAT AGAATTAAGA ACCCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA MW99060 NNNNNAAAAT GACTTTTTTC TACAAATCAT AAAGATATCG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTGAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA MW99061 NNNNNNNNNN NNNNNNNNNN NNCAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAGTTTGAG CAGGAATAGT GGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTAAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA MW99068 TCGCGAAAAT GACTTTTTTC CACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATGGT AGGAACATCT TTAAGAGTTT TAATTCGTAT GGAATTGAGA ACTCCAGGaT CTTTAATTGG AGATGATCAA ATTTATaATA CCATTGTTAC AGCTCATGCA MW99080 TAGCGAAAAT GACTTTTTTC AACAAATCAT AAAGATATTG GAACTTTATA TTTTATTTTT GGAATTTGAG CAGGAATATT AGGAACATCT TTAAGAATTT TAATTCGTAT AGAATTAGGA ACACCTGGAT CTTTAATTGG AGATGATCAA ATTTATAATA CTATTGTAAC AGCTCATGCT MW99084 TAGCGAAAAT GACTTTTTTC AACAAATCAT AAAGATATTG GAACTTTATA TTTTAATTTT NNNNTTTGAG CAGGTATATT AGGAACATCA TTAAGAATCT TAATTCGTAT AGAGTTAGGA ACACCAGGAT CATTAATTGG AGATGATCAA ATTTATAATA CTATTGTCAC AGCtCATGCT MW99094 TCGCGAAAAT GACTTTTTTC TACAAACCAT AAAGATATTG GaACATTATA TTTTATTTTT GGNATTTGAG CAGGAATAGT AGGAACATCA CTAAGAaTTT TAATTCGTAT AGAATTAAGA ACCCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTtAC AGCCCATGCA MW99095 NNNNNNNNNN NNNNNNNNNN NACAAATCAT AAAGATATCG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTGAGA ACTCCCGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCACGCA MW99097 TAGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATa TTTCATTTTT GGTATTTGAG CAGGAATAGT AGGAACATCT TTAAGAATTT TAATTCGTAT GGAATTAAGA ATTCCAGGAT CTTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA

167 Appendix 3: Sequence data Cytochrome Oxidase I (COI)

....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| 5 15 25 35 45 55 65 75 85 95 105 115 125 135 145 155 165 175 MW99102 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAGTTTGGG CAGGAATGGT AGGAACATCT TTAAGAGTTT TAATTCGTAT GGAATTAAGA ACTCCAGGAT CTTTGATTGG AGATGATCAA ATTTATAATA CCATTGTTAC AGCTCATGCA MW99104 TCGCGAAAAT GACTTTTTTC CACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGANTTTGGG CAGGAATGGT AGGAACATCT TTAAGAGTTT TAATTCGTAT GGAATTGAGA ACTCCAGGAT CTTTAATTGG AGATGATCAA ATTTATAATA CCATTGTTAC AGCTCATGCA MW99105 TCGCGAAAAT GACTTTTTTC CACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGGG CAGGAATGGT AGGAACATCT TTAAGAGTTT TAATTCGTAT GGAATTGAGA ACTCCAGGAT CTTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA MW99124 TAGCGAAAAT AAATTTTTTC NNNNAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCT TTAAGAATCT TAATTCGTAT AGAATTAGGA ACTCCTGGAT CTTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCtCATGCA MW99128 TTGCGAAAAT GACTTTTTTC CACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCT TTAAGAATTT TAATTCGTAT AGAATTGAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAACA CTATTGTtAC AGCTCATGCA MW99134 TAGCGAAAGT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCT TTAAGAATCT TAATTCGTAT GGAATTAAGA ACCCCTGGAT CTTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA MW99135 NNNNNNNNNN NNNNNNNNNC TACAAATAAT AAAGATNTNG GAACANTATA TTTTATTTTT GGAATATGAG CAGGAATAGC AGGAACATCC CTAAGAATTT TAATTCGTAT GGAATTAAGA ACTCCTGGAT CCTTAATTGG AAATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA MW99140 NNNNNAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CGGGAATAGT AGGAACATCT CTAAGAATTT TAATTCGTAT GGAATTAAGA ACCCCTGGAT CTTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA MW99141 TCGCGAAAAT GACTTTTTTC CACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGANTTTGGG CAGGAATGGT AGGAACATCT TTAAGAGTTT TAATTCGTAT GGAATTGAGA ACTCCAGGAT CTTTAATTGG AGATGATCAA ATTTATAATA CCATTGTTAC AGCTCATGCA MW99158 TAGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACTTTATA TTTTATTTTT NNNATTTGAG CAGGAATATT AGGAACATCT TTAAGAaTTC TAATTCGAAT AGAATTAGGA ACCCCCGGAT CTTTGATTGG AGATGATCAA ATTTATAATA CTATTGTAAC AGCTCATGCT MW99163 TAGCGAAAAT GACTTTTTTC CACAAATCAT AAAGATATTG GAACATTATA TTTTATCTTT GGGATTTGAG CAGGAATAGT AGGAACATCT CTAAGAATTT TAATTCGTAT AGAACTAAGA ATTCCCGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA MW99164 TCGCGaAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGANTTTGAG CAGGAATAGT NGGAACATCc CTaaGAATTT TAATTCGTAT GGAATTAAGA ACTCCCGGAT CCTTAATTGG GAATGATCAA ATTTATAACA CTATTGTTAC AGCCCATGCA MW99165 NNNNNNNNNN NNNNNNNNNC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT GGAATTAAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA MW99170 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATCTTT GGAATTTGAG CAGGAATAGT GGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTAAGA ACTCCTGGAT CCTTAATTGG AAATGATCAA ATTTATAATA CTATTGTCAC AGCCCATGCA MW99174 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GGACATTATA TTTtaTTTTT GGGATTTGAG CAGGAATAGT GGGAACATCT CTAAGAATTT TAATTCGTAT AGAACTGAGA ACTCCTGGAT CCTtAATTGG AGATGATCAA ATTTATAATA CTATTATTAC AGCCCATGCA MW99187 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GGACATTATA TTTTATTTTT GGGATTTGAG CAGGAATAGT GGGAACATCC CTAAGAATTT TAATTCGTAT AGAACTGAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTATTAC AGCCCATGCA MW99196 TCGCGAAAAT GACTTTTTTC CACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATGGT AGGAACATCT TTAAGAGTTT TAATTCGTAT GGAATTGAGA ACTCCAGGAT CTTTAATTGG aGATGATCAA ATTTATAATA CCATTGTTAC AGCTCATGCA MW99203 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGC AGGAACATCC CTAAGAATTT TAATTCGTAT GGAATTAAGA ACTCCTGGAT CCTTAATTGG AAATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA MW99221 TAGCGAAAAT GACTTTTTTC NNcAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGTATTTGAG CAGGAATATT AGGAACATCC TTAAGAATTT TAATCCGTAT AGAATTAGGA ACCCCAGGAT CTTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCT MW99226 TAGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATCG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCT CTAAGAATTT TAATTCGTAT AGAATTAAGA ACTCCCGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA MW99240 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGNNTTTGAG CAGGAATAGT NGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTAAGA ACTCCTGGAT CTTTAATTGG AGATGATCAA ATTTATAATA CNATTGTTAC AGCCCATGCA MW99247 TAGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT GGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTAAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA MW99263 TCGCGAAAAT GACTTTTTTC CACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGANTTTGAG CAGGAATGGT AGGAACATCT TTAAGAGTTT TAATTCGTAT GGAATTGAGA ACTCCAGGAT CTTTGATTGG AGATGATCAA ATTTATAATA CCATTGTTAC AGCTCATGCA MW99264 TCGCGAAAAT GACTTTTTtN NACaaATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATGGT AGGAACATCT TTAAGAGTTT TAATTCGTAT GGAATTGAGA ACTCCAGGAT CTTTGATTGG AGATGATCAA ATTTATAATA CCATTGTTAC AGCTCATGCA MW99274 NNNNGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAGTTTGAG CAGGAATAGT GGGAACATCT CTAAGAATTT TAATTCGTAT GGAATTAAGA ACTCCCGGAT CCTTAATTGG GAATGATCAA ATTTATAACA CTATTGTTAC AGCCCATGCA MW99276 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAGTTTGAG CAGGAATAGT GGGAACATCT CTAAGAATTT TAATTCGTAT GGAATTAAGA ACTCCCGGAT CCTTAATTGG GAATGATCAA ATTTATAACA CTATTGTTAC AGCCCATGCA MW99286 TAGCGAAAAT GACTTTTTTC TACAAAtCAT AAAGATATCG GAACATTATA TTTTATTTTT GGAGTTTGAG CAGGAATAGT AGGAACATCC TTAAGAATTC TAATTCGTAT AGAATTAAGA ACTCCGGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA MW99289 TAGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATCG GAACATTATA TTTTATTTTT GGGNTTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTGAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA MW99292 NNNNGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAGTTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTGAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA MW99301 TAGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATGGT AGGAACATCC TTAAGAATTT TAATTCGTAT AGAATTAAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCACGCA MW99303 TAGCGAAAAT GACTTTTTTC CACAAATCAT AAAGATATTG GaACATTATA TTTTATNTTT GGAATTTGAG CAGGAATAGT AGGAACATCT CTAAGAATTT TAATTCGTAT AGAACTAAGA ATTCCTGGAT CTTTAATTGG GGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA MW99309 NNNNNNNNNT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATCCGTAT AGAATTAAGA ACTCCTGGAT CTTTGATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA MW99314 NNNNNAAAAT GACTTTTTTC TACAAATCAT AAAGATATCG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCT CTAAGAATTT TAATTCGTAT AGAATTAAGA ACTCCCGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA MW99319 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAGTTTGAG CAGGAATAGT GGGAACATCT CTAAGAATTT TAATTCGTAT GGAATTAAGA ACTCCCGGAT CCTTAATTGG GAATGATCAA ATTTATAACA CTATTGTTAC AGCCCATGCA MW99320 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAGTTTGAG CAGGAATAGT GGGAACATCT CTAAGAATTT TAATTCGTAT GGaATTAAGA ACTCCCGGAT CCTTAATTGG GAATGATCAA ATTTATAACA CTATTGTTAC AGCCCATGCA MW99341 NNNNNNNNNN NNNNNTTTTC TACTAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAGTTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTGAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTtAC AGcTCATGCA MW99344 TCGCGAAAAT GACTTTTTTC CACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATCCGTAT AGAATTAAGA ACTCCTGGAT CTTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA MW99353 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCA CTAAGAATTT TAATTCGTAT AGAATTAAGA ACCCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA MW99357 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCA CTAAGAATTT TAATTCGTAT AGAATTAAGA ACCCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA MW99372 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATCCGTAT AGAATTAAGA ACTCCTGGAT CTTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA MW99374 TAGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATCG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCT CTAAGAATTT TAATTCGTAT AGAATTAAGA ACTCCCGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA MW99376 TAGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATCG GAACATTATA TTTTATTTTT GGANTTTGAG CAGGAATAGT AGGAACATCC TTAAGAATTC TAATTCGTAT AGAATTAAGA ACTCCGGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA MW99380 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAGTTTGAG CAGGAATAGT GGGAACATCT CTAAGAATTT TAATTCGTAT GGAATTAAGA ACCCCCGGAT CCTTAATTGG GAATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA MW99381 TCGCGAAAAT GACTTTTTTC CACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGANTTTGAG CAGGAATAGT GGGAACATCT TTAAGAGTTT TAATTCGTAT GGAATTGAGA ACTCCAGGAT CTTTGATTGG AGATGATCAA ATTTATAATA CCATCGTTAC AGCTCATGCA MW99382 TCGCGAAAAT GACTTTTTTC CACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT GGGAACATCT TTAAGAGTTT TAATTCGTAT GGAATTGAGA ACTCCAGGAT CTTTGATTGG AGATGATCAA ATTTATAATA CCATCGTTAC AGCTCATGCA MW99393 NNNNNNNNNN NNNNNNNNNC TACAAATCAT AAAGATATCG GAACATTATA TTTTATTTTT GGAGTTTGAG CAGGAATAGT AGGAACATCC TTAAGAATTC TAATTCGTAT AGAATTAAGA ACTCCGGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA MW99398 TAGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACTTTATA TTTTATTTTt NNAATTTGAG CAGGAATATT AGGAACATCT TTAAGAATTC TAATTCGAAT AGAATTGGGA ACCCCTGGAT CTTTAATTGG AGATGATCAA ATTTATAATA CTATTGTAAC AGCTCATGCT MW99406 TAGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATCG GAACATTATA TTTTATTTTT GGAGTTTGAG CAGGAATAGT AGGAACATCC TTAAGAATTC TAATTCGTAT AGAATTAAGA ACTCCGGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA MW99407 TCGCGAAAAT TACTTTTTTN NACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATGGt AGGAACATCT TTAAGAGTTT TAATTCGTAT GGAATTGAGA ACTCCAGGAt CTTTGATTGG AGATGATCAA ATTTATAATa CCATTGTTAC AGCTCATGCA MW99408 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAGTTTGAG CAGGAATAGT GGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTAAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA MW99413 TAGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT GGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTAAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA MW99422 NNNNNNNNNN NNNNNNNNTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT GGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTAAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA MW99425 TAGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCT TTAAGAATTT TAATTCGTAT AGAATTAGGA ACACCTGGAT CTTTAATTGG AGATGATCAA ATTTATAATA CCATTGTAAC TGCTCATGCT MW99430 TAGCGAAAAT GACTTTTTTC TACAaATCAT AAAGATATTG GAACATTATA TTTTATCTTT GGAATTTGAG CCGGAATAGT AGGAACATCT TTAAGAATTT TAATTCGTAT AGAACTAAGA ATTCCTAGAT CTTTAATTGG TGATGATCAA ATTTANAATA CTATTGTAAC AGCTCACGCA MW99435 CGGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA CTTTATTTTT GGAATTTGAG CGGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTAAGA ACCCCTGGAT CTTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA MW99448 TCGCGAAAAT GACTTTTTTC TACAAACCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCA CTAAGAATTT TAATTCGTAT AGAATTAAGA ACCCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CtATTGTTAC AGCCCATGCA MW99449 TCGCGAAAAT GACTTTTTTC TACAAACCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCA CTAAGAATTT TAATTCGTAT AGAATTAAGA ACCCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA MW99465 NNNNNNNNNN GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCA CTAAGAATTT TAATTCGTAT AGAATTAAGA ACCCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA MW99469 TAGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATGGT NGGAACATCC TTAAGAATTT TAATTCGTAT AGAATTNAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCACGCA MW99471 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAGTTTGAG CAGGAATAGT GGGAACATCT CTAAGAATTT TAATTCGTAT GGAATTAAGA ACTCCCGGAT CCTTAATTGG GAATGATCAA ATTTATAACA CTATTGTTAC AGCCCATGCA MW99473 TAGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATCG GAACATTATA TTTTATTTTT GGAGTTTGAG CAGGAATAGT AGGAACATCC TTAAGAATTC TAATTCGTAT AGAATTAAGA ACTCCGGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA MW99476 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATCG GGACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCT CTAAGAATTT TAATTCGTAT AGAATTAAGA ACTCCCGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA MW99479 TAGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATCG GAACATTATA TTTTATTTTT GGANTTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTGAGA ACTCCCGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATCGTTAC AGCTCATGCA MW99494 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAGTTTGAG CAGGAATAGT GGGAACATCT CTAAGAATTT TAATTCGTAT GGAATTAAGA ACCCCAGGAT CCTTAATTGG GAATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA MW99501 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAGTTTGAG CAGGAATAGT GGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTAAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA MW99508 TAGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATCG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTGAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA MW99514 TTGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CGGGAATAGT AGGAACATCT CTAAGAATTT TAATTCGTAT GGAATTAAGA ACCCCTGGAT CTTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA MW99515 TAGCGAAAAT GACTTTTTTA AACAAATCAT AAAGATATTG GAACTTTATA TTTTATTTTT GGAATTTGAG cTGGAATAAT tGGAACTTCT TTAAGAATTT TaATTCGCCT TGAATTAGGT ACTCCAGGAT CTTTAATTGG AGATGATCAA ATCTATAATA CTATTGTAAC TGCTCATGCT

168 Appendix 3: Sequence data Cytochrome Oxidase I (COI)

....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| 5 15 25 35 45 55 65 75 85 95 105 115 125 135 145 155 165 175 MW99520 NNNNNNNNNN GACTTTTTTC AACAAATCAT AAAGATATTG GAACTTTATA TTTTATTTTT GGAATTTGAG CTGGAATAAT TGGAACTTCT CTAAGAATTC TAATTCGTCT TGAGTTAGGA ACTCCAGGAT CTTTAATTGG AGATGATCAA ATTTATAATA CTATTGTAAC AGCTCATGCT MW99526 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGC AGGAACATCC CTAAGAATTT TAATTCGTAT GGAATTAAGA ACTCCTGGAT CCTTAATTGG AAATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA MW99536 NTGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCC TTAAGAATTT TAATTCGTAT AGAATTAAGA ACCCCTGGAT CTTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA MW99537 TCGCGAAAAT GATTTTTTTC CACAAATCAT AAAGACATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCA CTAAGAATTT TAATTCGATT AGAGCTAAGA ACCCCAGGAT CATTAATTGG TGATGACCAA ATTTACAATA CTATTGTCAC AGCCCATGCA MW99538 TCGCGAAAAT GATTTTTTTC CACAAATCAT AAAGACATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCA CTAAGAATTT TAATTCGATT AGAGCTAAGA ACCCCAGGAT CATTAATTGG TGATGACCAA ATTTACAATA CTATTGTCAC AGCCCATGCA MW99546 TAGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAAcATTATA TTTTATTTTT GGAATTTGAG CAGGAATAAT AGGAACATCT TTAAGAATTT TAATTCGTAT AGAATTGAGA ACTCCTGGAT CTTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA MW99550 TCGCGAAAAT GATTTTTTTC CACAAATCAT AAAGACATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCA CTAAGAATTT TAATTCGATT AGAGCTAAGA ACCCCAGGAT CATTAATTGG TGATGACCAA ATTTACAATA CTATTGTCAC AGCCCATGCA MW99552 TAGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATCG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTGAGA ACTCCCGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCACGCA MW99559 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGGATAGT AGGAACATCC CTAAGAATTT TAATCCGTAT AGAATTAAGA ACTCCTGGAT CTTTGATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA MW99565 NNNNGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATCCGTAT AGAATTAAGA ACTCCTGGAT CTTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCCCACGCA MW99579 TAGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA CTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCA TtaAGAATTC TAATTCGTAT AGAATTGAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATa CTATTGTTAC AGCTCATGCA MW99591 NNNNNNNNNN NNNNNNNNNN TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCT CTAAGAATTT TAATTCGTAT GGAATTAAGA ACTCCTGGAT CCTTAATTGG AAACGATCAA ATTTATAACA CTATTGTTAC AGCTCATGCA MW99605 TCGCGAAAAT GACTTTTTTC TACAAATcAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCT CTAAGAATTT TAATTCGTAT GGAATTAAGA ACTCCTGGAT CCTTAATTGG AAACGATCAA ATTTATAACA CTATTGTTAC AGCTCATGCA MW99608 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA CTTTATTTTT GGAATTTGAG CGGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTAAGA ACCCCTGGAT CTTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA MW99612 TTGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAAGATTATA TTTTAATTTT GGAATTTGAG CAGGAATAGT GGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTAAGA ACCCCTGGAT CTTTAATTGG GGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA MW99613 TCGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAAT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTGAGA ACTCCCGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA OK96022 NNNNNNNNNN NNNNNNNNTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CGGGAATAGT GGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTAAGA ACCCCTGGAT CTTTGATTGG GGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA OK99001 NNNNNNNNNN NNNNNNNNNN NNNNNNNNNT AAAGATATTG GAACATTATA TTTTATTTTT GGGATTTGAG CGGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTAAGA ACCCCTGGAT CTTTGATTGG GGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA WE00002 NNNNNNNNNN NNNNNNNNNN NNNNAATCAT AAAGATATTG GGACATTATA TTTTATTTTT GGAATTTGAG CGGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTGAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAACA CTATTGTTAC AGCTCATGCA WE00003 TAGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACTTTATA TTTTATTTTT GGNNTTTGAG CAGGAATAGT AGGAACATCT TTAAGAATTT TAATTCGAAT AGAATTAAGA ACTCCAGGAT CTTTAATTGG GGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA WE02321 NNNNNNNAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCC TTAAGAATTT TAATTCGTAT AGAATTAAGA ACTCCTGGAT CTTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA WE02421 NNNNNNNNNN NNNNNNNNNN NNNNNNNNNT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTAAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA WE02431 NNNNNNNNNN NNNNNNNNNN NNCAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGGG CAGGAATAGT GGGAACATCT TTAAGAATTT TAATTCGTAT GGAATTAAGA ACTCCGGGAT CTTTAATTGG AGATGATCAA ATTTATAATA CCATCGTTAC AGCTCATGCA WE02451 NNNNNNNAAT GACTTTTTTC NNCAAATCAT AAAGATATTG GGACATTATA CTTTATTTTT GGTATTTGAG CAGGAATAGT GGGAACATCT CTAAGAATTT TAATTCGTAT AGAATTGAGA ACTCCTGGAT TCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA WE02491 NNNNNNNNNN NNNNNNNNNC TACAAATCAT AAAGATATTG GAACGTTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTGAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAACA CTATTGTTAC AGCCCATGCA WE02531 NNNNNNNNNN NNNNNTTTTC TACAAATCAT AAAGATATCG GAACACTATA TTTTATTTTT GGAGTTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTGAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTCAC AGCTCATGCA WE02535 NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NAAGATATTG GAACATTATA CTTTATTTTT GGAATTTGGG CAGGAATGGT AGGAACATCT TTAAGAGTTT TAATTCGTAT GGAATTGAGA ACTCCAGGAT CTTTGATTGG AGATGATCAA ATTTATAATA CCATTGTCAC AGCTCATGCA WE02536 NNNNNNNNNN NNNNNNNNNN NNNNAATCAT AAAGATATTG GAACACTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTAAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA WE02591 NNNNNNNNNN NNNNNNNNNN NNNNNNNCAT AAAGATATCG GAACATTATA TTTTATTTTT GGAGTTTGAG CGGGAATAGT GGGAACATCC TTAAGAATTC TAATTCGTAT AGAATTGAGA ACTCCGGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA WE02612 TAGCGAAAAT GACTTTTTTC TACAAATCAT AAAGATATCG GAACATTATA TTTTATTTTT GGAGTTTGAG CGGGAATAGT GGGAACATCC TTAAGAATTC TAATTCGTAT AGAATTGAGA ACTCCGGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA WE02614 NNNNNNNNNN NNNNNNNNNN NNNNNNGCAT AAAGATATCG GAACATTATA TTTTATTTTT GGAGTTTGAG CGGGAATAGT GGGAACATCC TTAAGAATTC TAATTCGTAT AGAATTGAGA ACTCCGGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA WE02621 NNNNNNNNNN NNNNNNNNNN NNNNNNNCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCA CTAAGAATTT TAATTCGTAT AGAATTAAGA ACCCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA WE02661 NNNNNNNNNT GACTTTTTTC TACAAATCAT AAAGATATCG GAACACTATA TTTTATTTTT GGAGTTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTGAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTCAC AGCTCACGCA WE02662 NNNNNNNNNN NNNNNNNNNN NNNAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCC CTTAGAATTT TAATCCGTAT AGAATTAAGA ACTCCTGGAT CTTTGATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA WE02671 NNNNNNNNNN NNNNNTTTTC TACAAATCAT AAAGATATCG GAACATTATA TTTTATTTTT GGAGTTTGAG CAGGAATAGT AGGAACATCC TTAAGAATTC TAATTCGTAT AGAATTAAGA ACTCCAGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA WE02674 NNNNNNNNNN NNNNNNNNNN NNNAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCA CTAAGAATTT TAATTCGTAT AGAATTAAGA ACCCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCCCATGCA WE02676 NNNNNNNNNT GACTTTTTTC TACAAATCAT AAAGATATCG GAACATTATA TTTTATTTTT GGAGTTTGAG CAGGGATAGT AGGAACATCC CTAAGAATTT TAATTCGTAT AGAATTGAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA WE02677 NNNNNNNNNN NNNNNNNNNN NNNNNNNNNT AAAGATATTG GAACATTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT GGGAACATCT TTAAGAGTTT TAATTCGTAT GGAATTGAGA ACTCCAGGAT CTTTGATTGG AGATGATCAA ATTTATAATA CCATCGTTAC AGCTCATGCA WE85001 NGAAAAAATG ACTTTTTTTC TACAAATCAT AAAGATATTG GAACGTTATA TTTTATTTTT GGAATTTGAG CAGGAATAGT AGGAACATCC TAAAGAATTT TAATTCGTAT AGAATTGAGA ACTCCTGGAT CCTTAATTGG AGATGATCAA ATTTATAACA CTATTGTTAC AGCCCATGCA WE98001 NNNNNNNAAT GACTTTTTTC TACAAATCAT AAAGATATTG GAACATTATA TTTTATTTTT GGAGTTTGAG CAGGAATAGT AGGAACATCC CTAAGAATTT TAATCCGTAT AGAATTGAGA ACTCCCGGAT CCTTAATTGG AGACGATCAA ATTTATAATA CTATTGTTAC AGCTCATGCA

....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| 185 195 205 215 225 235 245 255 265 275 285 295 305 315 325 335 345 355 AD98001 TTTATCATAA TTTTTTTTAT AGTTATACCT ATTATAATCG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGAGC ACCTGATATA GCCTTTCCAC GATTAAATAA TATAAGATTC TGATTATTAC CACCATCATT AATACTACTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA AD98009 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTG GTTCCTTTAA TATTAGGAGC ACCTGATATA GCCTTTCCTC GATTAAATAA CATAAGATTT TGATTACTAC CGCCTTCATT AATATTACTA ATTTCCAGAA GAATCGTAGA AAATGGAGCA AD98012 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTG GTTCCTTTAA TATTGGGAGC ACCTGATATA GCCTTTCCTC GATTAAATAA TATAAGATTT TGATTACTAC CGCCTTCACT AATACTACTA ATTTCCAGAA GAATCGTAGA AAATGGAGCA AD98018 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATCG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGGGC ACCTGATATA GCCTTTCCAC GACTAAATAA TATAAGATTC TGATTATTAC CACCATCATT AATACTACTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA AD98024 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATCG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGGGC ACCTGATATA GCCTTTCCAC GACTAAATAA TATAAGATTC TGATTATTAC CGCCATCATT AATACTACTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA AD98029 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATCG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGAGC ACCTGATATA GCCTTTCCCC GATTAAATAA TATAAGATTT TGGTTATTAC CACCATCATT AATACTATTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA AD98036 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGAGC ACCTGATATA GCCTTTCCAC GATTAAATAA TATAAGATTC TGATTACTAC CGCCATCATT AATATTACTA ATTTCCAGAA GAATTCTAGA AAATGGAGCA DS00001 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAATTGATTA GTTCCTTTAA TATTAGGAGC ACCTGATATA GCCTTTCCTC GTTTAAATAA TATAAGATTC TGATTATTGC CACCATCATT AATTCTATTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA DS00005 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAATTGATTA GTTCCTTTAA TATTAGGAGC ACCTGACATA GCCTTTCCTC GATTAAATAA CATAAGATTC TGATTATTAC CACCATCATT AATTCTATTA ATTTCCAGAA GAATTGTAGA AAATGGGGCA DS01001 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GGGGATTTGG TAATTGATTA GTTCCTTTAA TATTGGGAGC ACCTGATATA GCCTTCCCCC GATTAAATAA TATAAGATTC TGATTATTAC CACCATCATT AATACTACTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA JC00029 TTTATTATAA TTTTTTTTAT AGTTATACCC ATTATAATCG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGGGC ACCTGACATA GCCTTCCCTC GATTAAATAA TATAAGATTT TGATTATTAC CTCCATCATT GATGCTACTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA JC00039 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAATTGATTA ATTCCTTTAA TATTAGGGGC ACCTGATATA GCCTTTCCTC GATTAAATAA TATAAGATTC TGATTATTGC CCCCATCCCT AATACTACTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA JC00040 TTTATTATAA TTTTTTTTAT GGTTATACCT ATTATAATTG GAGGATTCGG TAACTGATTA GTTCCCTTAA TATTAGGAGC ACCTGATATA GCCTTTCCAC GATTAAATAA TATGAGATTT TGATTATTAC CACCATCATT AATACTACTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA JC00042 TTTATTATAA TTTTTTTTAT AGTTATACCC ATTATAATCG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGGGC ACCTGACATA GCCTTCCCTC GATTAAATAA TATAAGATTC TGATTATTAC CCCCATCATT GATGCTACTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA JC00043 TTTATTATaA TTTTTTTTAT AGTTATACCT ATTATAATTG GGGGATTTGG TAACTGATTA GTTCCTTTAA TGTTAGGAGC ACCTGATATA GCCTTCCCCC GATTAAATAA TATAAGATTC TGATTACTAC CACCATCCTT AATACTACTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA JC00045 TTTATTATAA TTTTTTTTAT GGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGGGC ACCTGATATA GCCTTCCCCC GATTAAATAA TATAAGATTC TGATTACTAC CACCATCCTT AATACTACTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA JC00046 TTTATTATAA TTTTTTTTAT GGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGGGC ACCTGATATA GCCTTCCCCC GATTAAATAA TATAAGATTC TGATTACTAC CACCATCCTT AATACTACTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA JC00051 TTTATTATAA TTTTTTTTAT AGTTATACCC ATTATAATCG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGGGC ACCTGACATA GCCTTCCCTC GATTAAATAA TATAAGATTT TGATTATTAC CTCCATCATT GATGCTACTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA JC00055 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATCG GAGGATTTGG TAATTGATTA GTTCCTTTAA TATTAGGAGC ACCTGATATA GCTTTCCCCC GAATAAATAA TATAAGATTT TGATTATTAC CCCCATCATT AATATTATTA ATTTCAAGAA GAATTGTAGA AAATGGAGCA JC00057 TTCATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAATTGATTA GTTCCTTTAA TATTAGGAGC ACCTGATATA GCTTTCCCTC GAATAAATAA TATAAGATTT TGATTATTAC CTCCATCATT AATATTATTA ATTTCAAGAA GAATTGTAGA AAATGGAGCA JC00061 TTTATTATAA TTTTTTTTAT AGTTATACCC ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGAGC ACCTGACATA GCCTTCCCTC GATTAAATAA TATAAGATTC TGATTATTAC CTCCATCATT GATTCTACTA ATTTCTAGAA GAATTGTAGA AAATGGGGCA JC00062 TTCATTATAA TTTTTTTTAT AGTTATACCT ATTATaATTG GAGGATTTGG TAATTGATTA GTTCCTTTAA TATTAGGAGC ACCTGATATA GCTTTCCCTC GAATAAATAA TATAAGATTT TGATTATTAC CCCCATCATT AATATTATTA ATTTCAAGAA GAATTGTAGA AAATGGAGCA JC00063 TTTATTATAA TTTTTTTTAT AGTTATACCC ATTATAATTG GAGGGTTTGG TAACTGATTA GTTCCTTTAA TATTAGGAGC ACCTGACATA GCCTTCCCTC GATTAAATAA TATAAGATTC TGATTATTAC CTCCATCATT GATTCTACTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA JC01014 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TACTAGGAGC ACCTGATATA GCCTTCCCCC GATTAAATAA TATAAGATTC TGATTACTAC CACCATCCTT AATACTACTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA JC02001 TTTATTATAA TTTTTTTTAT AGTTATACCA ATTATAATTG GAGGATTTGG TAATTGATTA GTACCTTTAA TATTAGGAGC TCCTGATATA GCATTTCCTC GTATAAATAA TATAAGATTT TGATTATTAC CCCCCTCTCT AATTCTATTA ATTTCAAGAA GAATTGTAGA AAATGGTGCA

169 Appendix 3: Sequence data Cytochrome Oxidase I (COI)

....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| 185 195 205 215 225 235 245 255 265 275 285 295 305 315 325 335 345 355 JM00001 TTTATTATAA TTTTTTTTAT GGTTATACCT ATTATAATCG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGAGC ACCTGATATA GCTTTTCCAC GATTAAATAA CATAAGATTT TGATTATTAC CGCCATCATT AATACTACTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA MW00001 TTTATTATAA TTTTTTTTAT GGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGAGC ACCTGATATA GCCTTTCCAC GATTAAATAA TATAAGATTC TGATTATTAC CGCCATCATT AATACTACTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA MW00015 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TACTAGGAGC ACCTGATATA GCATTCCCCC GATTAAATAA TATAAGAtTC TGATTACTAC CACCATCCTT AATACTACTA ATTTCTAGAA GAATCGTAGA AAATGGAGCA MW00017 TTTAtTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAATTGACTT ATTCCTTTAA TATTAGGAGC TCCTGATATA GCTTTTCCAC GAATAAATAA TATAAGATTT TGATTATTAC CCCCATCTTT ATTATTGTTA ATTTCAAGAA GAATCGTTGA AAATGGAGCA MW00020 TTCATTATAA TTTTTTTTAT AGTTATACCT ATTATaATTG GAGGATTTGG TAATTGATTA GTTCCTTTAA TATTAGGAGC ACCTGATATA GCTTTCCCTC GAATAAATAA TATAAGATTT TGATTATTAC CCCCATCATT AATATTATTA ATTTCAAGAA GAATTGTAGA AAATGGAGCA MW00024 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAATTGACTA ATTCCTTTAA TATTGGGAGC ACCTGATATA GCTTTTCCTC GAATAAATAA TATAAGATTT TGACTATTAC CACCATCATT AATATTATTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA MW00032 TTTATTATAA TTTTTTTTAT GGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGAGC ACCTGATATA GCCTTTCCAC GATTAAATAA TATAAGATTC TGATTATTAC CGCCATCATT AATACTACTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA MW00044 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG CAATTGACTT GtCCCTTTAA TATTAGGAGC ACCTGATATA GCTTTTCCAC GAATAAATAA TATAAGATTT TGATTATTAC CCCCATCTTT ATTATTATTA ATTTCAAGAA GAATTGTAGA AAATGGAGCA MW00051 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATCG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGGGC ACCTGATATA GCCTTTCCAC GATTAAATAA TATAAGATTC TGATTATTAC CACCATCATT AATACTACTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA MW00056 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATCG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGGGC ACCTGATATA GCCTTTCCAC GATTAAATAA TATAAGATTC TGATTATTAC CACCATCATT AATACTACTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA MW00058 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATCG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGGGC ACCTGATATA GCCTTTCCAC GATTAAATAA TATAAGATTC TGATTATTAC CACCATCATT AATACTACTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA MW00059 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATCG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGGGC ACCTGATATA GCCTTTCCAC GATTAAATAA TATAAGATTC TGATTATTAC CACCATCATT AATACTACTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA MW00060 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTCGG TAACTGATTA GTTCCTTTAA TATTAGGGGC ACCTGATATA GCCTTTCCAC GATTAAATAA TATAAGATTC TGATTACTCC CACCATCATT AATACTATTA ATTTCCAGAA GAATTGTAGA AAATGGAGCG MW00064 TTTATTATAA TTTTTTTTAT GGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TGTTAGGAGC ACCTGATATA GCTTTCCCAC GATTAAATAA TATAAGATTT TGATTATTAC CGCCATCATT AATACTACTA ATTTCTAGAA GAATCGTAGA AAATGGAGCA MW00072 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTCGG TAACTGATTA GTTCCTTTAA TATTAGGGGC ACCTGATATA GCCTTTCCAC GATTAAATAA TATAAGATTC TGATTACTCC CACCATCATT AATACTATTA ATTTCCAGAA GAATTGTAGA AAATGGAGCG MW00076 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAATTGATTA GTTCCTTTAA TATTGGGGGC ACCTGATATA GCCTTTCCTC GATTAAATAA CATAAGATTC TGATTATTAC CCCCATCATT AATACTACTA ATTTCAAGAA GAATTGTAGA AAATGGAGCA MW00087 TTTATTAtAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAATTGATTA GTTCCTTTAA TATTGGGGGC ACCTGATATA GCCTTTCCTC GATTAAATAA CATAAGATTC TGATTATTAC CCCCATCATT AATACTACTA ATTTCAAGAA GAATTGTAGA AAATGGAGCA MW00101 TTTATTATAA TTTTTTTTAT GGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGAGC ACCTGATATA GCCTTTCCAC GATTAAATAA TATAAGATTT TGATTACTCC CACCATCATT AATACTATTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA MW00102 TTtATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTCGG TAACTGATTA GTTCCTTTAA TATTAGGGGC ACCTGATATA GCCTTTCCAC GATTAAATAA TATAAGATTC TGATTACTCC CACCATCATT AATACTATTA ATTTCCAGAA GAATTGTAGA AAATGGAGCG MW00110 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATCG GAGGATTTGG NAACTGATTA GTTCCTTTAA TATTAGGGGC ACCTGATATA GCCTTTCCAC GATTAAATAA TATAAGATTC TGATTATTAC CACCATCATT AATACTACTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA MW00116 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAATTGATTA GTTCCTTTAA TATTAGGAGC CCCAGATATA GCTTTCCCTC GAATAAATAA TATAAGATTT TGATTATTAC CCCCATCTTT AATATTATTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA MW00119 TTTATTATAA TTTTTTTTAT AGTAAtACCT ATTATAATTG GAGGATTTGG AAATTGACTT GtACCTTTAA TATTAGGGGC ACCTGATATA GCCTTTCCAC GAATAAATAA TATAAGATTT TGATTGTTAC CTCCATCCTT ATTATTATTA ATTTCAAGAA GAATCGTCGA AAATGGAGCA MW00125 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTG GTTCCTTTAA TATTAGGAGC ACCTGATATA GCCTTTCCTC GATTAAATAA CATAAGATTT TGATTACTAC CGCCTTCATT AATATTACTA ATTTCCAGAA GAATCGTAGA AAATGGAGCA MW00127 TTTATtATaA TTTTTTTTAT AGTTATACCT ATTATAATCG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGGGC ACCTGATATA GCCTTTCCAC GANTAAaTAA TATAAGATTC TGATTATTAC CGCCATCATT AATACTACTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA MW00129 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTG GTTCCTTTAA TATTGGGGGC ACCTGATATA GCCTTTCCTC GATTAAaTAA TATGAGATTT TGATTACTAC CGCCTTCACT AATaCTATTA ATTTCCAGAA GAATCGTAGA AAATGGaGCa MW00140 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGAGC ACCTGATATA GCCTTTCCAC GATTAAATAA TATAAGATTC TGATTACTAC CGCCATCATT AATATTACTA ATTTCCAGAA GAATTCTAGA AAATGGAGCA MW00151 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGAGC ACCTGATATA GCCTTTCCTC GATTAAATAA TATAAGATTT TGATTACTAC CGCCTTCATT AATATTACTA ATTTCCAGAA GAATCGTAGA AAATGGAGCA MW00157 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATCG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGGGC ACCTGATATA GCCTTTCCAC GATTAAATAA TATAAGATTC TGATTATTAC CGCCATCATT AATACTACTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA MW00176 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATCG GAGGATTTGG TAATTGATTA GTTCCTTTAA TATTAGGGGC ACCTGATATA GCTTTCCCCC GATTAAATAA TATAAGATTC TGATTATTAC CCCCATCATT AATACTACTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA MW00177 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTG GTTCCTTTAA TATTGGGGGC ACCTGATATA GCCTTTCCTC GATTAAATAA TATGAGATTT TGATTACTAC CGCCTTCACT AATACTATTA ATTTCCAGAA GAATCGTAGA AAATGGAGCA MW00178 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTG GTTCCTTTAA TATTGGGGGC ACCTGATATA GCCTTTCCTC GATTAAATAA TATGAGATTT TGATTACTAC CGCCTTCACT AATACTATTA ATTTCCAGAA GAATCGTAGA AAATGGAGCA MW00179 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAAtTG GAGGATTTGG TAACTGATTG GTTCCTTTAA TATTGGGGGC ACCTGATATA GCCTTTCCTC GATTAAATAA TATGAGATTT TGATTACTAC CGCCTTCACT AATACTACTA ATTTCCAGAA GAATCGTAGA AAATGGAGCA MW00183 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TACTGGGAGC ACCTGATATA GCATTCCCCC GATTAAATAA TATAAGATTC TGATTACTAC CACCATCCTT AATACTACTA ATTTCTAGAA GAATCGTAGA AAATGGAGCA MW00185 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TACTGGGAGC ACCTGATATA GCATTCCCCC GATTAAATAA TATAAGATTC TGATTACTAC CACCATCCTT AATACTACTA ATTTCTAGAA GAATCGTAGA AAATGGAGCA MW00186 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TACTGGGAGC ACCTGATATA GCATTCCCCC GATTAAATAA TATAAGATTC TGATTACTAC CACCATCCTT AAtACTACTA ATTTCTAGAA GAATCGTAGA AAaTGGAGCA MW00189 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GGGGATTTGG TAATTGATTA GTTCCTTTAA TACTAGGAGC ACCTGATATA GCCTTCCCCC GATTAAATAA TATAAGATTC TGATTGTTAC CACCATCCTT AATACTACTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA MW00226 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG CAACTGATTA GTTCCTTTAA TATTAGGAGC GCCTGATATA GCCTTTCCAC GATTAAATAA TATAAGATTC TGATTATTGC CACCATCATT AATACTACTA ATTTCTAGAA GAGTTGTAGA AAATGGAGCA MW00229 TTTATTATAa TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGGGC ACCTGATATA GCATTTCCAC GATTAAATAA TATAAGATTT TGATTATTAC CGCCATCATT AATACTACTA ATTTCTAGAA GAATCGTAGA AAATGGAGCA MW00231 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGGGC ACCTGATATA GCATTTCCAC GATTAAATAA TATAAGATTT TGATTATTAC CGCCATCATT AATACTACTA ATTTCTAGAA GAATCGTAGA AAATGGAGCA MW00232 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATCG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGGGC ACCTGATATA GCCTTTCCAC GATTAAATAA TATAAGATTC TGATTATTAC CACCATCATT AATACTACTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA MW00234 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTG GTTCCTTTAA TATTAGGAGC ACCTAATATA GCCTTTCCTC GATTAAATAA CATAAGATTT TGATTACTAC CGCCTTCATT AATATTACTA ATTTCCAGAA GAATCGTAGA AAATGGAGCA MW00259 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTG GTTCCTTTAA TACTAGGGGC ACCTGATATA GCTTTTCCAC GATTAAATAA TATAAGATTC TGATTACTTC CACCATCATT AATACTATTA ATTTCCAGAA GAATTGTAGA AAATGGGGCA MW00262 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTG GTTCCTTTAA TACTAGGGGC ACCTGATATA GCTTTTCCAC GATTAAATAA TATAAGATTC TGATTACTTC CACCATCATT AATACTATTA ATTTCCAGAA GAATTGTAGA AAATGGGGCA MW00267 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTCGG TAACTGATTA GTTCCTTTAA TATTAGGGGC ACCTGATATA GCCTTTCCAC GATTAAATAA TATAAGATTC TGATTACTCC CACCATCATT AATACTATTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA MW00269 TTTATTATAA TTTTTTTTAT GGTTATACCT ATTATAATCG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTGGGAGC ACCTGATATA GCCTTTCCAC GATTAAACAA TATAAGATTC TGATTATTGC CACCATCATT AATATTACTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA MW00290 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAATTGATTA GTTCCTTTAA TATTAGGAGC GCCTGATATA GCTTTTCCTC GATTAAaTAA CATAAGATTC TGATTATTGC CTCCGTCATT AATACTATTA ATTTCAAGAA GAATTGTAGA AAATGGAGCG MW00302 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAATTGATTA GTTCCTTTAA TATTAGGGGC ACCTGATATA GCATTTCCTC GATTAAACAA TATAAGATTC TGATTACTAC CCCCATCATT AATACTACTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA MW00316 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATCG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGGGC ACCTGATATA GCCTTTCCAC GATTAAATAA TATAAGATTC TGATTATTAC CACCATCATT AATACTACTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA MW00326 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAATTGACTA GTTCCTTTAA TATTAGGAGC ACCAGATATA GCTTTCCCAC GATTAAATAA TATAAGATTC TGATTACTAC CCCCATCATT AATATTACTA ATTTCAAGAA GAATTGTAGA AAATGGAGCA MW00327 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GNGGATTTGG TAACTGATTA ATTCCTTTAA TATTAGGAGC ACCTGATATA GCCTTTCCGC GATTAAATAA TATAAGATTC TGATTACTAC CCCCATCATT AATACTACTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA MW00328 TTTATTATAA TTTTTTTTAT GGTTATACCT ATTATAATCG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTGGGAGC ACCTGATATA GCCTTTCCAC GATTAAACAA TATAAGATTC TGATTATTGC CACCATCATT AATATTACTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA MW00330 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTG GTTCCTTTAA TATTAGGGGC ACCTGATATA GCCTTTCCTC GATTAAATAA TATGAGATTT TGATTACTAC CGCCTTCACT AATACTACTA ATTTCCAGAA GAATCGTAGA AAATGGAGCA MW00335 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA ATTCCTTTAA TATTAGGAGC ACCTGATATA GCTTTTCCAC GATTAAATAA TATAAGATTC TGATTACTCC CACCATCATT AATACTACTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA MW00347 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGAGC ACCTGATATA GCCTTTCCAC GATTAAATAA TATAAGATTC TGATTACTCC CACCATCATT AATACTATTA ATTTCCAGAA GAATTGTAGA AAATGGAGCG MW00348 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGGGC ACCTGATATA GCCTTTCCAC GATTAAATAA TATAAGATTC TGATTACTAC CACCATCATT AATATTACTA ATTTCCAGAA GAATTCTAGA AAATGGAGCA MW00358 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGAGC ACCTGATATA GCCTTTCCAC GATTAAATAA TATAAGATTC TGATTACTCC CACCATCATT AATACTATTA ATTTCCAGAA GAATTGTAGA AAATGGAGCG MW00393 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA ATTCCTTTAA TATTAGGAGC ACCTGATATA GCCTTTCCGC GATTAAATAA TATAAGATTC TGATTACTAC CCCCATCATT AATACTACTA ATTTCCAGAA GAATTGTAGA AAATGGGGCA MW00409 TTTATTAtAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA ATTCCTTTAA TATTAGGAGC ACCTGATATA GCTTTTCCAC GATTAAATAA TATAAGATTC TGATTACTCC CACCATCATT AATACTACTA ATTTCCAGAA GAATTGTaGA AAATGGAGCA MW00412 TTTATTATAA TTTTTTTTAT AGTTATACCC ATTATAATTG GAGGGTTTGG TAACTGATTA GTTCCTTTAA TATTAGGGGC ACCTGACATA GCCTTCCCTC GATTAAATAA TATAAGATTC TGATTATTAC CTCCATCATT GATTCTACTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA MW00424 TTTATTATAA TTTTTTTTAT GGTTATACCT ATTATAATCG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTGGGAGC ACCTGATATA GCCTTTCCAC GATTAAACAA TATAAGATTC TGATTATTGC CaCCATCATT AATATTACTA aTTTCCAGAA GAATTGTAGA AAATGGAGCA MW00444 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTCGG TAACTGATTA GTTCCTTTAA TATTAGGGGC ACCTGATATA GCCTTTCCAC GATTAAATAA TATAAGATTC TGATTACTCC CACCATCATT AATACTATTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA MW00452 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGAGC ACCTGATATA GCCTTTCCAC GATTAAATAA TATAAGATTC TGATTACTAC CACCATCATT AATATTACTA ATTTCCAGAA GAATCCTAGA AAATGGAGCA MW00469 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTACCTTTAA TATTAGGAGC ACCTGATATA GCATTCCCTC GATTAAATAA TATAAGATTT TGATTATTAC CCCCATCATT AATACTTTTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA MW00476 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTCGG TAACTGATTA GTTCCTTTAA TATTAGGGGC ACCTGATATA GCCTTTCCAC GATTAAATAA TATAAGATTC TGATTACTCC CACCATCATT AATACTATTA ATTTCCAGAA GAATTGTAGA AAATGGAGCG MW00497 TTTATTATAA TTTTTTTTAT AGTTATACCA ATTATAATTG GGGGATTTGG AAATtGATTA GTCCCTTTAA TATTAGGGGC TCCAGATATA GCTTTCCCCC GAATAAATAA TATAAGaTTT TGATTATTAC CTCCCTCATT AATACTTTTA ATTAGAAGTA GAATTGTAGA AAATGGAGCA MW00498 TTTATTATAA TTTTTTTTAT GGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TGTTAGGAGC ACCTGATATA GCTTTCCCAC GATTAAATAA TATAAGATTT TGATTATTAC CGCCATCATT AATACTACTA ATTTCTAGAA GAATCGTAGA AAATGGAGCA MW00504 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATCG GAGGATTTGG TAACTGATTA ATTCCTTTAA TATTAGGGGC ACCTGATATA GCTTTTCCTC GATTAAATAA TATAAGATTC TGATTATTAC CTCCATCATT AATACTATTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA MW00517 TTTATTATAA TTTTTTTTAT GGTTATACCT ATTATAATTG GAGGATTTGG TAATTGATTG GTTCCATTGA TATTAGGAGC ACCTGATATA GCTTTCCCTC GAATAAATAA TATAAGATTT TGATTATTAC CCCCATCATT AATATTATTA ATTTCAAGAA GAATTGTAGA AAATGGAGTA

170 Appendix 3: Sequence data Cytochrome Oxidase I (COI)

....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| 185 195 205 215 225 235 245 255 265 275 285 295 305 315 325 335 345 355 MW00525 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAATTGATTA GTTCCTTTAA TATTAGGAGC ACCTGATATA GCTTTCCCAC GAATAAATAA TATAAGATTT TGATTATTAC CCCCATCATT AATATTATTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA MW00530 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATCG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGGGC ACCTGATATA GCCTTCCCTC GATTAAATAA TATAAGATTC TGATTATTAC CCCCATCATT GATACTACTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA MW00539 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGAGC ACCTGATATA GCCTTTCCAC GATTAAATAA CATAAGATTC TGATTATTGC CGCCATCATT AATATTGCTA ATCTCCAGAA GAATTGTAGA AAATGGAGCA MW00547 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTCGG TAACTGATTA GTTCCTTTAA TATTAGGGGC ACCTGATATA GCCTTTCCAC GATTAAATAA TATAAGATTC TGATTACTCC CACCATCATT AATACTATTA ATTTCCAGAA GAATTGTAGA AAATGGAGCG MW01001 TTTATTATAA TTTTTTTTAT GGTTATACCT ATTATAATCG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGAGC ACCTGATATA GCTTTTCCAC GATTAAATAA CATAAGATTT TGATTATTAC CGCCATCATT AATACTACTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA MW01009 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAATTGATTA ATTCCTTTAA TATTAGGGGC ACCCGATATA GCCTTTCCTC GATTAAATAA TATAAGATTC TGATTATTGC CCCCATCCCT AATACTATTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA MW01011 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GtACCTTTAA TATTAGGAGC ACCTGATATA GCATTCCCTC GATTAAATAA TATAAGATTT TGATTATTAC CCCCATCATT AATACTTTTA ATTTCTAGAA GAATTGTAGA AAATGGAGCG MW01014 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GGGGATTTGG TAACTGATTA GTTCCTTTAA TATTGGGAGC ACCTGATATA GCCTTCCCCC GATTAAATAA TATAAGATTC TGATTACTAC CACCATCCTT AATACTACTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA MW01018 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGGTTTGG TAACTGATTA GtACCTTTAA TATTAGGGGC ACCTGATATA GCATTCCCTC GATTAAATAA TATAAGATTT TGATTATTAC CTCCATCATT AATACTTTTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA MW01019 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTGA TGTTAGGAGC ACCTGATATA GCCTTTCCAC GATTAAATAA TATAAGATTC TGATTATTGC CCCCATCATT AATACTTCTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA MW01022 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GTGGATTTGG AAACTGATTA ATCCCATTAA TATTAGGAGC ACCTGATATA GCTTTCCCAC GAATAAATAA TATAAGATTT TGATTATTAC CTCCGTCATT AATATTACTA ATTTCAAGAA GAATTGTAGA AAATGGAGCA MW01023 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG AAATTGATTA GTGCCTTTAA TATTAGGAGC CCCTGATATA GCTTTTCCAC GAATAAATAA CATAAGATTT TGATTATTAC CTCCATCCCT TATATTATTA ATTTCAAGAA GAATTGTAGA AAATGGAGCA MW01025 TTTATTATAA TTTTTTTTAT AGTTAtACCC ATTATAATCG GAGGGTTTGG TAACTGATTA GTTCCTTTAA TATTAGGAGC ACCTGACATA GCCTTCCCTC GATTAAATAA TATAAGATTC TGATTATTAC CTCCATCATT GATTCTACTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA MW01027 TTTATCATAA TTTTTTTTAT AGTAATACCT ATTATAATTG GAGGATTTGG AAATTGATTA GTGCCATTAA TATTAGGAGC CCCAGATATA GCTTTTCCAC GAATAAATAA TATAAGATTT TGATTATTAC CACCCTCACT AATATTATTA ATTAGAAGAA GAATTGTAGA AAATGGAGCA MW01034 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGGTTTGG TAACTGATTA GTACCTTTAA TATTGGGAGC ACCTGATATA GCATTCCCTC GATTAAATAA TATAAGATTT TGATTATTAC CTCCATCATT AATACTTTTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA MW01039 TTTATTATAA TTTTTTTTAT GGTTATACCT ATTATAATCG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGAGC ACCTGATATA GCTTTTCCAC GATTAAATAA CATAAGATTT TGATTATTAC CGCCATCATT AATACTACTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA MW01048 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATCG GAGGATTTGG TAATTGATTA GTTCCTTTAA TATTGGGAGC ACCTGATATA GCTTTTCCTC GAATAAATAA TATAAGATTT TGATTATTAC CTCCATCATT AATATTATTA ATTTCAAGAA GAATCGTAGA AAATGGAGCA MW01053 TTTATTATAA TTTTTTTTAT GGTTATACCT ATTATAATCG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGAGC ACCTGATATA GCTTTTCCAC GATTAAATAA CATAAGATTT TGATTATTAC CGCCATCATT AATACTACTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA MW01059 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGGTTTGG TAACTGATTA GTACCTTTAA TATTGGGGGC ACCTGATATA GCATTCCCTC GATTAAATAA TATAAGATTT TGATTATTAC CCCCATCATT AATACTTTTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA MW01061 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAATTGATTA ATTCCTTTAA TATTAGGAGC ACCTGATATA GCTTTCCCTC GAATAAATAA TATAAGATTT TGATTATTAC CCCCATCATT AATATTATTA ATTTCAAGAA GAATTGTAGA AAATGGAGCA MW01072 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GGGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGAGC ACCTGATATA GCCTTCCCCC GATTAAATAA TATAAGATTC TGATTACTAC CACCATCCTT AATACTACTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA MW01078 TTTATTATAA TTTTTTTTAT GGTTATACCT ATTATAATCG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGAGC ACCTGATATA GCTTTTCCAC GATTAAATAA CATAAGATTT TGATTATTAC CGCCATCATT AATACTACTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA MW01083 TTTATTATAA TCTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAATTGATTA GTTCCTTTAA TATTAGGGGC ACCTGATATA GCATTTCCTC GATTAAACAA TATAAGATTC TGATTACTAC CTCCATCATT AATACTGCTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA MW01092 TTtATTATaA TTTTTTTTAT AGTTATACCT ATTATAATtG GAGGGtTTGG TAACTGATTA GTACCTTTAA TATTGGGGGC ACCTGATATA GCATTCCCTC GATTAAATAA TATAAGATTT TGATTATTAC CCCCATCATT AATACTTTTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA MW01103 TTTATCATAA TTTTTTTTAT AGTtATACCT ATTATAATTG GAGGGTTTGG TAACTGATTA GtACCTTTAA TATTGGGGGC ACCTGATATA GCATTCCCTC GATTAAATAA TATAAGATTC TGATTATTAC CTCCATCATT AATACTTTTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA MW01105 TTTATCATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGGGC ACCTGATATA GCCTTCCCCC GATTAAATAA TATAAGATTC TGATTACTAC CACCATCCTT AATACTATTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA MW01107 TTTATTATAA TTTTTTTTAT GGTTATACCT ATTATAATCG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGAGC ACCTGATATA GCTTTTCCAC GATTAAATAA CATAAGATTT TGATTATTAC CGCCATCATT AATACTACTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA MW01114 TTTATTATAA TTTTTTTTAT AGTTATACCA ATTATAATtG GAGGATTTGG tAATTGATTA GTACCTTTAA TATTAGGAGC TCCTGATATA GCATTTCCTC GTATAAATAA TATAAGATTT TGATTATTAC CCCCCTCTCT AATTCTATTA ATTTCAAGAA GAATTGTAGA AAATGGTGCA MW01116 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGGTTTGG CAACTGATTA GtACCTTTAA TATTGGGGGC ACCTGATATA GCATTCCCTC GATTAAATAA TATAAGATTT TGATTATTAC CCCCATCATT AATACTTTTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA MW01120 TTTATCATAA TTTTCTTCAT AGTTATACCT ATTATAATTG GAGGATTTGG AAATTGATTA GTTCCACTTA TATTAGGTGC ACCTGATATA GCTTTCCCAC GAATAAATAA TATAAGATTT TGATTATTAC CCCCATCTTT AATATTATTA ATTTCAAGAA GAATTGTAGA AAATGGAGCA MW02001 TTTATTATAA TTTTTTTTAT AGtTATACCT ATTATAATTG GAGGATTTGG TAATTGATTG GTTCCTTTAA TATTAGGGGC ACCTGATATA GCTTTCCCTC GATTAAATAA CATAAGATTT TGATTATTAC CCCCATCATT AATACTACTG ATTTCTAGAA GAATTGTAGA AAATGGAGCA MW02006 TTTATCATAA TTTTTTTTAT AGTTATACCC ATTATAATTG GAGGATTTGG TAATTGATTA GTTCCTTTAA TGTTAGGGGC ACNTGATATA GCCTTTCCTC GATTAAATAA TATAAGATTC TGATTATTAC CCCCATCACT AATACTACTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA MW02007 TTTATCATAA TTTTTTTTAT AGtTATACCT ATTATAATTG GAGGATTTGG TAATTGACTT ATCCCTTTAA TATTAGGAGC CCCTGATATA GCTTTTCCAC GAATAAATAA TATAAGATTT TGATTATTAC CCCCATCCTT ATTATTATTA ATTTCAAGAA GAATCGTTGA AAATGGAGCA MW02008 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG AAATTGATTA ATTCCTTTAA TATTAGGAGC ACCTGATATA GCATTTCCTC GAATAAATAA TATAAGATTT TGATTATTAC CTCCATCTTT GATACTGTTA ATTTCAAGAA GAATTGTAGA AAATGGAGCA MW02009 TTCATTATAA TTTTTTTTAT GGTTATACCT ATTATAATTG GAGGATTTGG TAATTGACTT GTTCCTTTAA TATTAGGAGC ACCTGATATA GCTTTTCCTC GAATAAATAA TATAAGATTT TGATTATTAC CTCCATCTTT ATTATTATTA ATTTCAAGAA GAATTGTTGA AAATGGAGCA MW02021 TTTATTATAA TTTTCTTTAT AGTTATACCT ATTATAATTG GTGGATTTGG AAATTGATTA GTTCCATTAA TATTAGGAGC CCCTGATATA GCATTTCCAC GAATAAATAA TATAAGATTT TGATTATTAC CTCCATCCTT AATATTATTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA MW02024 TTTATTATAA TTTTTTTTAT AGTTATACCA ATTATAATTG GAGGATTTGG TAATTGATTA GTACCTTTAA TATTAGGAGC CCCTGATATA GCATTTCCTC GTATAAATAA TATAAGATTT TGATTATTAC CTCCCTCTTT AATTCTCCTA ATTTCAAGAA GAATTGTAGA AAATGGTGCA MW02025 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG AAATTGACTT GTACCTTTAA TATTAGGAGC ACCTGATATA GCTTTCCCTC GAATAAATAA TATAAGATTT TGATTATTAC CACCATCATT AATATTATTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA MW02028 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GTGGATTTGG AAACTGATTA ATCCCATTAA TATTAGGAGC ACCTGATATA GCTTTCCCAC GAATAAATAA TATAAGATTT TGATTATTAC CTCCGTCATT AATATTGCTA ATTTCAAGAA GAATTGTAGA AAATGGAGCA MW02031 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG AAATTGATTA GtACCTTTAA TATTAGGAGC ACCTGATATA GCATTTCCAC GAATAAATAA TATAAGATTT TGATTATTAC CTCCATCATT AATATTATTA ATTTCAAGTA GAATTGTAGA AAATGGAGCA MW02033 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATCG GAGGATTTGG tAATTGATTA GTTCCTTTAA TATTGGGAGC ACCTGATATA GCTTTTCCCC GAATAAATAA TATAAGATTT TGATTATTAC CCCCATCATT AATATTATTA ATTTCAAGAA GAATTGTAGA AAATGGAGCA MW02034 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAATTGATTA GTCCCTTTAA TATTAGGAGC ACCTGATATA GCTTTCCCAC GAATAAATAA TATAAGATTT TGATTATTAC CCCCATCATT AATATTATTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA MW98002 TTTATTATAA TTTTTTTTAT AGTTATACCA ATTATAATTG GAGGATTTGG AAATTGATTA GTACCATTAA TATTAGGTGC TCCAGATATA GCATTTCCTC GAATAAATAA CATAAGATTT TGATTGTTAC CACCATCATT AATACTATTA ATTTCAAGAA GAATTGTAGA AAATGGAGCA MW98008 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAATTGATTA GTTCCTTTAA TATTAGGGGC ACCTGATATA GCATTTCCTC GATTAAACAA TATAAGATTc TGATTAcTAC CCCCATCATT AATACTACTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA MW98009 TTTATCATAA TTTTTTTTAT AGTTATACCT ATTATAATCG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGAGC ACCTGATATA GCCTTCCCAC GATTAAATAA TATAAGATTC TGATTATTAC CACCATCATT AATACTACTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA MW98020 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTGA TATTAGGAGC ACCTGATATG GCTTTCCCAC GATTAAATAA TATAAGATTT TGATTATTAC CCCCATCATT AATATTACTA ATTTCTAGAA GAATCGTAGA AAATGGAGCA MW98029 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAATTGATTA ATTCCTTTAA TATTNGGAGC ACCTGATATA GCCTTTCCTC GATTAAATAA CATAAGATTC TGATTATTAC CTCCATCATT AATACTACTA ATTTCAAGAA GAATTGTAGA AAATGGAGCA MW98049 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATCG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGAGC ACCTGATATA GCCTTTCCCC GATTAAATAA TATAAGATTT TGATTATTAC CACCATCATT AATACTATTA ATTTCTAGAA GAATTGTAGA AAACGGAGCA MW98079 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTG GTTCCTTTAA TATTAGGNGC ACCTGATATA GCCTTTCCTC GATTAAATAA TATAAGATTT TGATTACTAC CACCTTCATT AATATTACTA ATTTCTAGAA GAATCTTAGA AAATGGAGCA MW98084 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TACTAGGGGC ACCTGATATA GCCTTCCCCC GATTAAATAA TATAAGATTC TGATTACTAC CACCATCCTT AATACTACTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA MW98089 TTTATtATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GGGGGTTTGG TAACTGATTG GTTCCTTTAA TATTAGGGGC ACCTGATATA GCCTTTCCTC GATTAAATAA TATAAGATTT TGATTACTAC CACCTTCaTT AATATTACTA ATTTCTAGAA GAATCGTAGA AAATGGAGCA MW98094 TTTATTATAA TTTTTTTTAT AGTtATACCT ATTATAATTG GAGGATTTGG TAATTGACTT GTtCCTTTAA TATTAGGAGC NCCTGATATA GCTTTTCCAC GAATAAATAA CATAAGATTT TGATTATTAC CNCCATCATT ATTATTATTA ATTTCAAGAA GAATTGTAGA AAATGGAGCA MW98097 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGAGC ACCTGACATA GCCTTTCCTC GATTAAATAA TATAAGATTT TGATTACTAC CGCCTTCATT AATATTACTA ATTTCCAGAA GAATCGTAGA AAATGGAGCA MW98103 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGGGC ACCTGATATA GCTTTCCCCC GATTAAATAA TATAAGATTC TGATTATTAC CCCCATCATT AATACTACTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA MW98106 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATCG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGAGC ACCTGATATA GCCTTTCCCC GATTAAATAA TATAAGATTT TGGTTATTAC CACCATCATT AATACTATTA ATTTCTAGAA GAATTGTAGa AAATGGGGCA MW98128 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACtGATTA GTTCCTTTAA TGTTAGGGGC ACCTGATATA GCCTTTCCAC GATTAAATAA TATAAGATTC tGATTATTAC CGCCATCATT AATACTTCTA ATTTCTAGaA GAATTGTAGA AAATGGAGCA MW98129 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAATTGATTA GTACCTTTAA TATTAGGAGC ACCCGATATA GCATTCCCTC GATTAAATAA TATAAGATTC TGATTATTAC CCCCATCATT AATACTTTTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA MW98133 TTTATTATAA TTTTTTTTAT AGTTATACCC ATTATAATTG GAGGGTTTGG TAACTGATTA GTTCCTTTAA TATTAGGAGC ACCTGACATA GCCTTCCCTC GATTAAATAA TATAAGATTC TGATTATTAC CTCCATCATT GATTCTACTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA MW98136 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTG GTTCCTTTAA TATTGGGGGC ACCTGATATA GCCTTTCCTC GATTAAATAA TATGAGATTT TGATTACTAC CGCCTTCACT AATACTACTA ATTTCCAGAA GAATCGTAGA AAATGGAGCA MW98138 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTG GTTCCTTTAA TATTAGGAGC ACCTGATATA GCCTTTCCTC GATTAAATAA TATAAGATTT TGATTGCTAC CACCTTCATT AATATTACTA ATTTCCAGAA GAATCGTAGA AAATGGAGCA MW98139 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTG GTTCCTTTAA TATTGGGGGC ACCTGATATA GCCTTTCCTC GATTAAATAA TATGAGATTT TGATTACTAC CGCCTTCACT AATACTACTA ATTTCCAGAA GAATCGTAGA AAATGGAGCA MW98154 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTG GTTCCTTTAA TATTAGGAGC ACCTGATATA GCCTTTCCTC GATTAAATAA TATAAGATTT TGATTGCTAC CACCTTCATT AATATTACTA ATTTCCAGAA GAATCGTAGA AAATGGAGCA MW98162 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGAGC aCCTGATATA GCCTTTCCTC GATTAAaTAA TATAAGATTT TGATTACTAC CGCCTTCATT AATATTACTA ATTTCCAGAA GAATCGTAGA AAATGGAGCA MW98170 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTG GTTCCTTTAA TATTGGGGGC ACCTGATATG GCCTTTCCtC GATTAAaTAA TATGAGATTT TGATTACTAC CGCCTTCACT AATACTACTA ATTTCCAGAA GAATCGTAGA AAATGGAGCA MW98172 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTGA TATTAGGAGC ACCTGATATG GCTTTCCCAC GATTAAATAA TATAAGATTT TGATTATTAC CTCCATCATT AATATTACTA ATTTCTAGAA GAATCGTAGA AAATGGAGCA MW98180 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTG GTTCCTTTAA TATTAGGAGC ACCTGATATA GCCTTTCCTC GATTAAATAA TATAAGATTT TGATTGCTAC CACCTTCATT AATATTACTA ATTTCCAGAA GAATCGTAGA AAATGGAGCA MW98183 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTG GTTCCTTTAA TATTAGGAGC ACCTGATATA GCCTTTCCTC GATTAAATAA TATAAGATTT TGATTGCTAC CACCTTCATT AATATTACTA ATTTCCAGAA GAATCGTAGA AAATGGAGCA MW98185 TTTATTATAA TTTTTTTTAT GGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTACCTTTAA TATTAGGGGC ACCTGATATA GCATTCCCTC GATTAAATAA TATAAGATTT TGATTATTAC CCCCATCATT AATACTTTTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA

171 Appendix 3: Sequence data Cytochrome Oxidase I (COI)

....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| 185 195 205 215 225 235 245 255 265 275 285 295 305 315 325 335 345 355 MW98189 TTTATTATAA TTTTTTTTAT AGTTATACCC ATTATAATTG GAGGATTTGG TAACTGATTG GTTCCTTTAA TATTAGGAGC ACCTGATATA GCCTTTCCTC GATTAAATAA TATAAGATTT TGATTACTAC CACCTTCATT AATATTGCTA ATTTCCAGAA GAATCGTAGA AAATGGAGCA MW98203 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTG GTTCCTTTAA TATTGGGGGC ACCTGATATG GCCTTTCCTC GATTAAATAA TATGAGATTT TGATTACTAC CGCCTTCACT AATACTACTA ATTTCCAGAA GAATCGTAGA AAATGGAGCA MW98205 TTTATTATAA TTTTTTTTAT GGTTATACCT ATCATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGAGC ACCTGATATA GCCTTTCCAC GACTAAATAA TATAAGATTT TGATTATTAC CACCATCATT AATACTACTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA MW98212 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGGGC ACCTGATATA GCCTTTCCAC GATTAAATAA TATAaGATTT TGATTATTAC CGCCATCATT AATACTACTA ATTTCTAGAA GAATCGTAGA AAATGGAGCA MW98220 TTTATTATAA TTTTTTTTAT GGTTATACCT ATTATAATTG GAGGATTTGG TAATTGACTA GTTCCTTTAA TATTAGGAGC ACCTGATATA GCTTTTCCTC GAATAAATAA TATAAGATTT TGATTATTAC CACCATCATT AATATTGTTA ATTTCTAGTA GAATTGTAGA AAATGGAGCA MW98228 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGaTTTGG TAACTGATTA GTACCTTTAA TGTTAGGAGC ACCTGATATA GCATTCCCTC GATTAAATAA TATAAGATTT TGATTGTTAC CCCCATCACT AATACTTTTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA MW98230 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG AAATTGACTT ATCCCTTTAA TATTAGGAGC ACCTGATATA GCTTTTCCAC GAATAAATAA TATAAGATTT TGATTATTAC CTCCATCTCT ATTATTATTA ATTTCAAGAA GAATTGTTGA AAATGGAGCA MW98235 TTTATTATAA TTTTTTTCAT AGTTATACCT ATTATAATTG GAGGGTTTGG TAACTGATTA GTTCCCTTAA TATTAGGGGC ACCAGATATA GCTTTCCCCC GATTAAATAA TATAAGATTC TGACTATTAC CCCCATCATT AATATTATTA ATTTCAAGAA GAATTGTAGA AAATGGAGCA MW98240 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATCG GAGGATTTGG TAATTGATTA GTTCCTTTAA TATTAGGGGC ACCTGATATA GCCTTTCCAC GATTAAATAA TATAAGATTC TGATTACTAC CACCATCATT AATACTACTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA MW98261 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGGGC ACCTGATATA GCCTTTCCAC GATTAAATAA TATAAGATTC TGATTATTAC CACCATCATT AATACTATTA ATTTCCAGAA GGATTGTAGA AAATGGAGCA MW98264 TTTATTATAA TTTTTTTCAT AGTTATACCT ATTATAATTG GAGGGTTTGG TAACTGATTA GTCCCCTTAA TATTAGGAGC ACCAGATATA GCTTTCCCCC GATTAAATAA TATAAGATTC TGACTATTAC CCCCATCATT AATATTATTA ATTTCAAGAA GAATTGTAGA AAATGGAGCA MW98265 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GTGGATTTGG AAACTGATTA ATCCCATTAA TATTAGGAGC ACCTGATATA GCTTTCCCAC GAATAAATAA TATAAGATTT TGATTATTAC CTCCGTCATT AATATTACTA ATTTCAAGAA GAATTGTAGA AAATGGAGCA MW98270 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAATTGATTA GTCCCTTTAA TATTAGGAGC ACCTGATATA GCTTTCCCTC GAATAAATAA TATAAGATTT TGATTATTAC CTCCATCATT AATATTATTA ATTTCAAGAA GAATTGTAGA AAATGGAGCA MW98274 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAATTGATTA GTTCCTTTAA TATTAGGGGC ACCTGATATA GCCTTTCCTC GATTAAATAA CATAAGATTC TGATTATTAC CCCCATCATT AATGCTACTA ATTTCAAGAA GAATCGTAGA AAATGGAGCA MW98278 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTACCTTTAA TATTGGGAGC ACCTGATATA GCATTCCCTC GATTAAATAA TATAAGATTT TGATTATTAC CCCCATCATT AATACTTTTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA MW98284 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGAGC ACCTGATATA GCCTTTCCTC GATTAAATAA TATAAGATTT TGATTACTAC CGCCTTCATT AATATTACTA ATTTCCAGAA GAATCGTAGA AAATGGAGCA MW98285 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTG GTTCCTTTAA TATTAGGAGC ACCTGATATA GCCTTTCCTC GATTAAATAA CATAAGATTT TGATTACTAC CGCCTTCATT AATATTACTA ATTTCCAGAA GAATCGTAGA AAATGGAGCA MW98294 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATCG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTGGGGGC ACCTGATATA GCCTTTCCAC GATTAAATAA TATAAGATTC TGATTACTAC CACCATCATT AATACTACTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA MW98313 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTG GTTCCTTTAA TATTGGGGGC ACCTGATATA GCCTTTCCTC GATTAAATAA TATGAGATTT TGATTACTAC CGCCTTCACT AATACTACTA ATTTCCAGAA GAATCGTAGA AAATGGAGCA MW98315 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGaTTTGG TAACTGATTA GTTCCTTTAA TATTAGGAGC ACCTGATATA GCCTTTCCAC GATTAAATAA TATAAGATTT TGATTATTAC CGCCATCATT AATACTACTA ATTTCTAGAA GAATCGTAGA AAATGGAGCA MW99001 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAATTGATTA GTTCCTTTAA TATTAGGAGC ACCTGATATA GCTTTCCCTC GAATAAATAA TATGAGATTT TGATTATTAC CCCCATCATT AATATTATTA ATTTCAAGAA GAATTGTAGA AAATGGAGCA MW99006 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTG GTTCCTTTAA TATTAGGAGC ACCTGATATA GCCTTTCCTC GATTAAATAA CATAAGATTT TGATTACTAC CGCCTTCATT AATATTACTA ATTTCCAGAA GAATCGTAGA AAATGGAGCA MW99009 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATCG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGAGC ACCTGATATA GCCTTTCCTC GATTAAATAA TATAAGATTT TGGTTATTAC CGCCATCATT AATACTATTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA MW99013 TTTATTATAA TTTTTTTCAT GGTTATACCT ATTATAATTG GAGGATTTGG TAATTGATTA GTCCCCTTAA TATTAGGAGC ACCTGATATA GCTTTCCCTC GATTAAATAA TATAAGATTT TGATTATTAC CCCCATCTTT AATATTACTA ATTTCAAGAA GAATTGTAGA AAATGGAGCA MW99014 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TGTTAGGGGC ACCTGATATA GCCTTTCCAC GATTAAATAA TATAAGATTC TGATTATtAC CACCATCATT AATACTTCTA ATTTCTAGGA GAATTGTAGA AAATGGAGCA MW99018 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAATTGATTA GTTCCTTTAA TATTAGGAGC ACCTGATATA GCTTTCCCAC GAATAAATAA TATAAGATTT TGATTATTAC CACCATCACT AATATTATTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA MW99028 TTCATTATAA TTTTTtTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAATTGATTA GTTCCTtTAA TATTAGGAGC ACCTGATATa GCTCTCCCTC GAATaAATAA TATAAGATTT TGAttATTAC CCCCATCATT AaTATTATTA ATTTCAAGAA GAATtGTAGA AAATGGAGCA MW99038 TTTATTATAA TTTTTTTCAT AGTTATACCT ATTATAATTG GAGGGTTTGG TAACTGATTA GTTCCCTTAA TATTGGGGGC ACCAGATATA GCTTTCCCCC GATTAAATAA TATAAGATTC TGACTATTAC CTCCATCATT AATATTATTA ATTTCAAGAA GAATTGTAGA AAATGGAGCA MW99042 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAATTGATTA GTACCTTTAA TATTAGGAGC ACCTGATATA GCATTTCCTC GATTAAATAA TATAAgATTC TGATTATTAC CTCCATCATT AATACTTTTA ATTTCtAGAA GAATTGTAGA AAATGGAGCA MW99045 TTTATTATAA TTTTTTTTAT AGTTATACCA ATTATAATTG GTGGATTTGG AAATTGATTA GTACCTTTAA TATTAGGAGC TCCTGATATA GCTTTTCCAC GAATAAATAA CATAAGATTT TGATTATTAC CACCTTCATT AATATTATTA ATTTCAAGAA GAATTGTAGA AAATGGAGCA MW99047 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG CAACTGATTA GTTCCTTTAA TATTAGGGGC ACCTGATATA GCTTTCCCTC GATTAAATAA CATAAGATTT TGATTATTAC CCCCATCATT AATACTACTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA MW99053 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATCG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGGGC ACCTGATATA GCCTTTCCAC GACTAAATAA TATAAGATTC TGATTATTAC CACCATCATT AATACTACTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA MW99057 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTG GTTCCTTTAA TATTGGGGGC ACCTGATATG GCCTTTCCTC GATTAAATAA TATGAGATTT TGATTACTAC CACCTTCACT AATACTACTA ATTTCCAGAA GAATCGTAGA AAATGGAGCA MW99058 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GaGGATTTGG TAACTGATTG GTTCCTTTAA TATTAGGAGC ACCTGATATA GCCTTTCCTC GATTAAATAA CATAAGATTT TGATTATTAC CGCCTTCATT AATATTACTA ATTTCCAGAA GAATCGTAGA AAATGGAGCA MW99059 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTG GTTCCTTTAA TATTGGGGGC ACCTGATATA GCCTTTCCTC GATTAAATAA TATGAGATTT TGATTACTAC CGCCTTCACT AATACTACTA ATTTCCAGAA GAATCGTAGA AAATGGAgCA MW99060 TTTATCATAA TTTTTTTTAT AGTTATACCT ATTATAATCG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGAGC ACCTGATATA GCCTTTCCAC GATTAAATAA TATAAGATTC TGATTATTAC CACCATCATT AATACTACTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA MW99061 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTG GTTCCTTTAA TATTAGGAGC ACCTGATATA GCCTTTCCTC GATTAAATAA TATAAGATTT TGATTGCTAC CACCTTCATT AATATTACTA ATTTCCAGAA GAATCGTAGA AAATGGAGCA MW99068 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GGGGATTTGG TAATTGATTA GTTCCTTTAA TACTAGGNGC aCCTGATATa gCCTTCCCCC GATTAAATAA TATAAGATTC TGATTACTAC CACCATCCTT AATACTACTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA MW99080 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG AAATTGACTA GTACCTTTAA TATTAGGAGC ACCTGATATA GCATTTCCAC GAATAAATAA CATAAGATTT TGATTATTAC CTCCATCATT AATATTATTA ATTTCAAGTA GAATTGTAGA AAATGGAGCA MW99084 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG AAATTGATTA ATTCCTTTAA TATTAGGAGC ACCTGATATA GCATTtCCTC GAATAAATAA TATAAGATTT TGATTATTAC CTCCATCTTT GATATTGTTA ATTTCAAGAA GAATTGTAGA AAATGGAGCA MW99094 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GGGGNTTTGG TAACTGATTG GTTCCTTTAA TATTGGGGGC ACCTGATATA GCCTTTCCTC GATTAAATAA TATGAGATTT TGATTACTAC CGCCTTCACT AATATTACTA ATTTCCAGAA GAATCGTAGA AAATGGAGCA MW99095 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATCG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGGGC ACCTGATATA GCCTTTCCAC GACTAAATAA TATAAGATTC TGATTATTAC CGCCATCATT AATACTACTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA MW99097 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAATTGATTA GTTCCTTTAA TATTAGGGGC ACCTGATATA GCTTTCCCTC GAATAAATAA TATGAGATTT TGATTATTAC CTCCATCATT AATATTATTA ATTTCAAGAA GAATTGTAGA AAATGGAGCA MW99102 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GGGGATTTGG TAATTGATTA GTTCCTTTAA TACTAGGAGC ACCTGATATA GCCTTCCCCC GATTAAATAA TATAAGATTC TGATTACTAC CGCCATCCTT AATACTACTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA MW99104 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GGGGATTTGG TAATTGATTA GTTCCTTTAA TACTAGGAGC ACCTGATATA GCCTTCCCCC GATTAAATAA TATAAGATTC TGATTGCTAC CACCATCCTT AATACTACTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA MW99105 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GGGGATTTGG TAATTGATTA GTTCCTTTAA TACTAGGAGC ACCTGATATA GCCTTCCCCC GATTAAATAA TATAAGATTC TGATTGCTAC CACCATCCTT AATACTACTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA MW99124 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAATTGATTA GTACCTTTAA TATTAGGAGC TCCAGATATA GCTTTCCCTC GAATAAATAA TATAAGATTT TGATTATTAC CCCCATCTTT AATATTATTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA MW99128 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAATTGATTA GTTCCTTTAA TATTAGGGGC ACCTGATATA GCATTTCCTC GATTAAATAA TATAAGATTC TGATTACTAC CCCCATCATT AATACTACTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA MW99134 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGGTTTGG TAATTGATTA GTTCCTTTAA TATTAGGAGC CCCTGATATA GCCTTCCCTC GAATAAATAA CATAAGATTT TGATTATTAC CCCCATCATT AATATTATTA ATTTCTAGAA GAATTGTAGA AACTGGAGCA MW99135 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGGGC ACCTGATATA GCCTTTCCCC GATTAAATAA TATAAGATTC TGATTATTAC CTCCATCATT AATACTACTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA MW99140 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GGGGATTTGG TAACTGATTA GTACCTTTAA TATTAGGAGC ACCTGATATA GCATTCCCTC GATTAAATAA TATAAGATTT TGATTATTAC CCCCATCATT AATACTTTTA ATTTCTAGAA GAATCGTAGA AAATGGAGCA MW99141 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GGGGATTTGG TAATTGATTA GTTCCTTTAA TACTAGGAGC ACCTGATATA GCCTTCCCCC GATTAAATAA TATAAGATTC TGATTGCTAC CACCATCCTT AATACTACTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA MW99158 TTTATTATAA TTTTCTTTAT AGTAATACCT ATTATAATTG GAGGATTTGG AAATTGATTA GTACCACTAA TATTAGGAGC ACCAGATATA GCTTTTCCTC GAATAAATAA TATAAGATTT TGATTATTAC CTCCTTCATT AATATTATTA ATTTCAAGAA GAATTGTAGA AAATGGAGCA MW99163 TTTATCATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAATTGATTA GTTCCTTTAA TATTAGGAGC ACCTGATATA GCTTTTCCCC GAATAAATAA TATAAGATTT TGATTATTAC CCCCATCATT AATATTATTA GTTTCAAGAA GTATTGTAGA AAATGGAGCA MW99164 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GNGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGGGC ACCTGATATA GCCTTTCCAC GATTAAATAA TATAAGATTT TGATTATTaC CACCATCATT AATACTACTA ATTTCtAGAA GAATCGTAGA AAATGGAGCA MW99165 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTG GTTCCTTTAA TATTGGGGGC ACCTGATATG GCCTTTCCTC GATTAAATAA TATGAGATTT TGATTACTAC CGCCTTCACT AATACTACTA ATTTCCAGAA GAATCGTAGA AAATGGAGCA MW99170 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATCG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGAGC ACCTGATATA GCCTTTCCTC GATTAAATAA TATAAGATTT TGGTTATTAC CGCCATCATT AATACTATTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA MW99174 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGAGC ACCTGATATA GCCTTTCCAC GATTAAATAA TATAAGATTC TGATTACTAC CGCCATCATT AATATTACTA ATTTCCAGAA GAATTCTAGA AAATGGGGCA MW99187 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GNGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGAGC ACCTGATATA GCCTTTCCAC GATTAAATAA TATAAGATTC TGATTACTAC CGCCATCATT AATATTACTA ATTTCCAGAA GAATTCTAGA AAATGGGGCA MW99196 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GGGGATTTGG TAATTGATTA GTTCCTTTAA TACTAGGAGC ACCTGATATA GCCTTCCCCC GATTAAATAA TATAAGATTC TGATTACTAC CACCATCCTT AATACTACTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA MW99203 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTCCCTTTAA TATTAGGGGC ACCTGATATA GCCTTTCCCC GATTAAATAA TATAAGATTC TGATTATTAC CTCCATCATT AATACTACTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA MW99221 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GTGGATTTGG AAATTGACTA GTACCTTTAA TATTAGGAGC TCCTGATATA GCTTTTCCAC GAATAAATAA TATAAGATTT TGATTATTAC CCCCTTCATT AATATTATTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA MW99226 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATCG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGGGC ACCTGATATA GCCTTTCCAC GACTAAATAA TATAAGATTC TGATTATTAC CACCATCATT AATACTACTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA MW99240 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTG GTTCCTTTAA TATTGGGGGC ACCTGATATG GCCTTTCCtC GATTAAATAA TATGAGATTT TGATTACTAC CGCCTTCACT AATACTACTA ATTTCCAGAA GAATCGTAGA AAATGGAGCA MW99247 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGAGC ACCTGATATA GCCTTTCCTC GATTAAATAA TATAAGATTT TGATTACTAC CGCCTTCATT AATATTACTA ATTTCCAGAA GAATCGTAGA AAATGGAGCA MW99263 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GGGGATTTGG TAATTGATTA GTTCCTTTAA TACTAGGAGC ACCTGATATA GCCTTCCCCC GATTAAATAA TATAAGATTC TGATTACTAC CACCATCCTT AATACTACTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA MW99264 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GGGGATTTGG TAATTGATTA GTTCCTTTAA TACTAGGAGC ACCTGATATA GCCTTCCCCC GATTAAATAA TATAAGATTC TGATTACTAC CACCATCCTT AATACTACTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA MW99274 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGGGC ACCTGATATA GCATTTCCAC GATTAAATAA TATAAGATTT TGATTATTAC CACCATCATT AATACTACTA ATTTCTAGAA GAATCGTAGA AAATGGAGCA

172 Appendix 3: Sequence data Cytochrome Oxidase I (COI)

....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| 185 195 205 215 225 235 245 255 265 275 285 295 305 315 325 335 345 355 MW99276 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGGGC ACCTGATATA GCATTTCCAC GATTAAATAA TATAAGATTT TGATTATTAC CACCATCATT AATACTACTA ATTTCTAGAA GAATCGTAGA AAATGGAGCA MW99286 TTTATTATAA TTTTTTTTAT AGTTATACCT ATtATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTGGGGGC GCCTGATATA GCCTTTCCAC GATTAAATAA TATAAGAtTC TGATTATTGC CACCATCATT AATACTACTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA MW99289 TTTATCATAA TTTTTTTTAT AGTTATACCT ATTATAATCG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGAGC ACCTGATATA GCCTTTCCAC GATTAAATAA TATAAGATTC TGATTATTAC CACCATCATT AATACTACTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA MW99292 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGAGC ACCTGATATA GCCTTTCCAC GATTAAATAA TATAAGATTC TGATTATTAC CACCATCATT AATACTACTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA MW99301 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATCG GGGGATTTGG TAATTGATTG ATTCCTTTAA TATTAGGAGC ACCTGATATA GCCTTTCCCC GATTAAATAA TATAAGATTC TGATTATTAC CCCCATCATT AATACTACTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA MW99303 TTTATCATAA TTTTTTTTAT GGTTATACCT ATTATAATTG GAGGATTTGG TAATTGATTG GTTCCTTTAA TATTAGGAGC ACCTGATATA GCTTTCCCCC GAATAAATAA CATAAGATTT TGATTACTAC CACCATCATT AATATTATTA ATTTCTAGAA GTATTGTAGA AAATGGAGCA MW99309 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATCG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGAGC ACCCGATATA GCTTTCCCCC GATTAAATAA TATAAGATTC TGATTATTAC CTCCATCATT AATACTACTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA MW99314 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATCG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGGGC ACCTGATATA GCCTTTCCAC GACTAAATAA TATAAGATTC TGATTATTAC CACCATCATT AATACTACTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA MW99319 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGGGC ACCTGATATA GCATTTCCAC GATTAAATAA TATAAGATTT TGATTATTAC CACCATCATT AATACTACTA ATTTCTAGAA GAATCGTAGA AAATGGAGCA MW99320 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGGGC ACCTGATATA GCATTTCCAC GATTAAATAA TATAaGATTT TGATTATTAC CACCATCATT AATACTACTA ATTTCTAGAA GAATCGTAGA AAATGGAGCA MW99341 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGAGC ACCTGATATA GCCTTTCCAC GATTAAATAA TATAAGATTC TGATTATTaC CACCATCATT AATACTACTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA MW99344 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGAGC ACCTGATATA GCTTTCCCCC GATTAAATAA TATAAGATTC TGATTATTAC CCCCATCATT AATACTATTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA MW99353 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTG GTTCCTTTAA TATTGGGGGC ACCTGATATA GCCTTTCCTC GATTAAATAA TATGAGATTT TGATTACTAC CGCCTTCACT AATACTACTA ATTTCCAGAA GAATCGTAGA AAATGGAGCA MW99357 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTG GTTCCTTTAA TATTGGGGGC ACCTGATATA GCCTTTCCTC GATTAAATAA TATGAGATTT TGATTACTAC CGCCTTCACT AATACTATTA ATTTCCAGAA GAATCGTAGA AAATGGAGCA MW99372 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGAGC ACCTGATATA GCTTTCCCCC GATTAAATAA TATAAGATTC TGATTATTAC CCCCATCATT AATACTATTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA MW99374 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATCG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGGGC ACCTGATATA GCCTTTCCAC GACTAAATAA TATAAGATTC TGATTATTAC CACCATCATT AATACTACTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA MW99376 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTGGGGGC GCCTGATATA GCCTTTCCAC GATTAAATAA TATAAGATTC TGATTATTGC CACCATCATT AATACTACTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA MW99380 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGGGC ACCTGATATA GCATTTCCAC GATTAAATAA TATAAGATTT TGATTATTAC CACCATCATT AATACTACTA ATTTCTAGAA GAATCGTAGA AAATGGAGCA MW99381 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TACTGGGAGC ACCTGATATA GCATTCCCCC GATTAAATAA TATAAGATTC TGATTACTAC CACCATCCTT AATACTACTA ATTTCTAGAA GAATCGTAGA AAATGGAGCA MW99382 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TACTGGGAGC ACCTGATATA GCATTCCCCC GATTAAATAA TATAAGATTC TGATTACTAC CACCATCCTT AATACTACTA ATTTCTAGAA GAATCGTAGA AAATGGAGCA MW99393 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTGGGGGC GCCTGATATA GCCTTTCCAC GATTAAATAA TATAAGATTC TGATTATTGC CACCATCATT AATACTACTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA MW99398 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGGTTTGG AAATTGATTA GTACCATTAA TATTNGGAGC TCCAGATATA GCTTTTCCTC GAATAAATAA TATAAGATTT TGATTATtAC CTCCTTCACT AATATTATTA ATTTCAAGAA GAATCGTAGA AAATGGAGCA MW99406 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTGGGGGC GCCTGATATA GCCTTTCCAC GATTAAATAA TATAAGATTC TGATTATTGC CACCATCATT AATACTACTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA MW99407 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GGGGATTTGG TAATTGATTA GTTCCTTTAA TACTAGGAGC ACCTGATATA GCCTTCCCCC GATTAAATAA TATAAGATTC TGATTACTAC CACCATCCTT AATACTACTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA MW99408 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTG GTTCCTTTAA TATTAGGAGC ACCTGATATA GCCTTTCCTC GATTAAATAA TATAAGATTT TGATTGCTAC CACCTTCATT AATATTACTA ATTTCCAGAA GAATCGTAGA AAATGGAGCA MW99413 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGAGC ACCTGATATA GCCTTTCCTC GATTAAATAA TATAAGATTT TGATTACTAC CGCCTTCATT AATATTACTA ATTTCCAGAA GAATCGTAGA AAATGGAGCa MW99422 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGAGC ACCTGATATA GCCTTTCCTC GATTAAATAA TATAAGATTT TGATTACTAC CGCCTTCATT AATATTACTA ATTTCCAGAA GAATCGTAGA AAATGGAGCA MW99425 TTTATTATAA TTTTCTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG AAATTGATTA GTACCTTTAA TATTGGGAGC CCCTGATATA GCTTTCCCTC GAATAAATAA CATAAGATTT TGATTATTAC CCCCATCATT AATTTTATTA ATTTCAAGAA GAATCGTAGA AAATGGAGCA MW99430 TtTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAATTGACTA ATTCCTTTAA TATTGGGAGC aCCTGATATA GCTTTTCCtC GAATAAATAA TATAAGATTT TGACTATTAC CACCATCATT AATATTATTA ATTTCtAGAA GAATTGTAGA AAATGGAGCA MW99435 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTACCTTTAA TATTAGGAGC ACCTGATATA GCATTCCCTC GATTAAATAA TATAAGATTT TGATTATTAC CCCCATCATT AATACTTTTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA MW99448 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GNGGATTTGG TAACTGATTG GTTCCTTTAA TATTGGGGGC ACCTGATATA GCCTTTCCTC GATTAAATAA TATGAGATTT TGATTACTAC CGCCTTCACT AATACTACTA ATTTCCAGAA GAATCGTAGA AAATGGAGCA MW99449 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTG GTTCCTTTAA TATTGGGGGC ACCTGATATA GCCTTTCCTC GATTAAATAA TATGAGATTT TGATTACTAC CGCCTTCACT AATACTACTA ATTTCCAGAA GAATCGTAGA AAATGGAGCA MW99465 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTG GTTCCTTTAA TATTGGGGGC ACCTGATATA GCCTTTCCTC GATTAAATAA TATGAGATTT TGATTACTAC CGCCTTCACT AATACTATTA ATTTCCAGAA GAATCGTAGA AAATGGAGCA MW99469 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATCG GGGGATTTGG TAATTGATTG ATTCCTTTAA TATTAGGAGC ACCTGATATA GCCTTTCCCC GATTAAATAA TATAAGATTC TGATTATTAC CCCCATCATT AATACTACTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA MW99471 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGGGC ACCTGATATA GCATTTCCAC GATTAAATAA TATAAGATTT TGATTATTAC CACCATCATT AATACTACTA ATTTCTAGAA GAATCGTAGA AAATGGAGCA MW99473 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTGGGGGC GCCTGATATA GCCTTTCCAC GATTAAATAA TATAAGATTC TGATTATTGC CACCATCATT AATACTACTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA MW99476 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATCG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGGGC ACCTGATATA GCCTTTCCAC GACTAAATAA TATAAGATTC TGATTATTAC CACCATCATT AATACTACTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA MW99479 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATCG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGGGC ACCTGATATA GCCTTTCCAC GATTAAATAA TATAAGATTC TGATTATTAC CGCCATCATT AATACTACTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA MW99494 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGaTTTGG TAACTGATTA GTTCCTTTGA TATTAGGAGC ACCTGATATG GCTTTCCCAC GATTAAATAA TATAAGATTT TGATTATTAC CTCCATCATT AATATTACTA ATTTCTAGAA GAATCGTAGA AAATGGAGCA MW99501 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTG GTTCCTTTAA TATTAGGAGC ACCTGATATA GCCTTTCCTC GATTAAATAA TATAAGATTT TGATTGCTAC CACCTTCATT AATATTACTA ATTTCCAGAA GAATCGTAGA AAATGGAGCA MW99508 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATCG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGGGC ACCTGATATA GCCTTTCCAC GACTAAATAA TATAAGATTC TGATTATTAC CGCCATCATT AATACTACTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA MW99514 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GGGGATTTGG TAACTGATTA GTACCTTTAA TATTAGGAGC ACCTGATATA GCATTCCCTC GATTAAATAA TATAAGATTT TGATTATTAC CCCCATCATT AATACTTTTA ATTTCTAGAA GAATCGTAGA AAATGGAGCA MW99515 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG AAATTGACTT GTCCCCTTAA TATTAGGAGC ACCTGATATA GCTTTTCCAC GAATAAATAA TATAAGATTT TGATTATTAC CCCCATCTTT ATTATTATTA ATTTCAAGAA GAATTGTCGA AAATGGAGCA MW99520 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAATTGACTT GTACCTTTAA TATTAGGAGC CCCCGATATA GCTTTTCCAC GAATAAATAA TATAAGATTT TGATTATTAC CTCCATCTTT ATTATTGTTA ATTTCAAGAA GAATTGTAGA AAATGGAGCA MW99526 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGGGC ACCTGATATA GCCTTTCCCC GATTAAATAA TATAAGATTC TGATTATTAC CTCCATCATT AATACTACTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA MW99536 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAATTGATTA GTACCTTTAA TATTAGGAGC ACCTGATATA GCATTTCCTC GATTAAATAA TATAAGATTC TGATTATTAC CTCCATCATT AATACTTTTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA MW99537 TTTATTATAA TTTTTTTCAT AGTTATACCT ATTATAATTG GAGGGTTTGG TAACTGATTA GTTCCCTTAA TATTAGGAGC ACCAGATATA GCTTTCCCCC GATTAAATAA TATAAGATTC TGACTATTAC CCCCATCATT AATATTATTA ATTTCAAGAA GAATTGTAGA AAATGGAGCA MW99538 TTTATTATAA TTTTTTTCAT AGTTATACCT ATTATAATTG GAGGGTTTGG TAACTGATTA GTTCCCTTAA TATTAGGAGC ACCAGATATA GCTTTCCCCC GATTAAATAA TATAAGATTC TGACTATTAC CCCCATCATT AATATTATTA ATTTCAAGAA GAATTGTAGA AAATGGAGCA MW99546 TTTATTATAA TTTTTTTCAT AGTTATACCT ATTATAATCG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGGGC ACCTGATATA GCCTTTCCAC GATTAAATAA TATAAGATTC TGATTATTAC CCCCATCATT AATACTTCTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA MW99550 TTTATTATAA TTTTTTTCAT AGTTATACCT ATTATAATTG GAGGGTTTGG TAACTGATTA GTTCCCTTAA TATTAGGAGC ACCAGATATA GCTTTCCCCC GATTAAATAA TATAAGATTC TGACTATTAC CCCCATCATT AATATTATTA ATTTCAAGAA GAATTGTAGA AAATGGAGCA MW99552 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATCG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGGGC ACCTGATATA GCCTTTCCAC GACTAAATAA TATAAGATTC TGATTATTAC CGCCATCATT AATACTACTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA MW99559 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATCG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTGGGAGC ACCTGATATA GCTTTCCCCC GATTAAATAA TATAAGATTC TGATTATTAC CTCCATCATT AATACTACTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA MW99565 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGAGC ACCTGATATA GCTTTCCCCC GATTAAATAA TATAAGATTC TGATTATTAC CTCCATCATT AATACTACTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA MW99579 TTTATTATAA TTTTTTTTAT AGTTATACCC ATTATAATCG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGGGC ACCTGACATA GCCTTCCCTC GATTAAATAA TATAAGATTC TGATTATTAC CCCCATCATT GATGCTACTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA MW99591 TTTATTATAA TTTTTTTTAT GGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCCTTAA TATTAGGGGC ACCTGATATA GCCTTTCCAC GATTAAATAA TATAAGATTT TGATTATTAC CGCCATCATT AATACTACTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA MW99605 TTTATTATAA TTTTTTTTAT GGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCCTTAA TATTAGGGGC ACCTGATATA GCCTTTCCAC GATTAAATAA TATAAGATTT TGATTATTAC CGCCATCATT AATACTACTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA MW99608 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTACCTTTAA TATTAGGAGC ACCTGATATA GCATTCCCTC GATTAAATAA TATAAGATTT TGATTATTAC CCCCATCATT AATACTTTTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA MW99612 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGGTTTGG TAACTGATTA GTACCTTTAA TATTGGGGGC ACCTGATATA GCATTCCCTC GATTAAATAA TATAAGATTC TGATTATTAC CTCCATCATT AATACTTTTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA MW99613 TTTATTATAA TTTTTTTCAT GGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGGGC ACCTGATATA GCCTTTCCAC GATTAAATAA TATAAGAtTC TGATTATTAC CCCCATCATT AATACTTCTA ATTTCCAGAA GAATTGTAGA AAGTGGAGCA OK96022 TTTATTATAA TTTTTTTTAT GGTTATACCT ATTATAATTG GAGGGTTTGG CAACTGATTA GTACCTTTAA TATTGGGAGC ACCTGATATA GCATTCCCTC GATTAAATAA TATAAGATTT TGATTATTAC CTCCATCATT AATACTTTTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA OK99001 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGGTTTGG TAACTGATTA GTACCTTTAA TATTGGGGGC ACCTGATATA GCATTCCCTC GATTAAATAA TATAAGATTT TGATTATTAC CTCCATCATT AATACTTTTA ATTTCTAGAA GAATTGTAGA AAACGGAGCA WE00002 TTTATCATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGAGC ACCTGATATA GCCTTTCCAC GATTAAATAA TATAAGATTC TGATTACTAC CACCTTCATT AATACTaCTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA WE00003 TTCATTATAA TTTTTTTTAT GGTTATACCT ATTATAATTG GAGGATTTGG TAATTGATTA GTTCCTCTAA TATTAGGGGC ACCTGATATA GCTTTCCCCC GATTAAATAA TATAAGATTT TGATTATTAC CCCCATCTTT AATACTGTTA ATTTCAAGAA GAGTTGTAGA AAATGGAGCA WE02321 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTG GTTCCTTTAA TATTGGGGGC ACCTGATATA GCTTTTCCTC GATTAAATAA TATGAGATTT TGATTACTAC CGCCTTCACT AATACTACTA ATTTCCAGAA GAATCGTAGA AAATGGAGCA WE02421 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTG GTTCCTTTAA TATTAGGGGC ACCTGATATG GCCTTTCCTC GATTAAATAA TATGAGATTT TGATTACTAC CGCCTTCACT AATACTACTA ATTTCCAGAA GAATCGTAGA AAATGGAGCA WE02431 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GGGGATTTGG TAACTGATTA GTTCCTTTAA TACTAGGAGC ACCTGATATA GCCTTCCCCC GATTAAATAA TATAAGATTC TGATTACTAC CACCATCCTT AATACTACTA ATTTCTAGGA GAATTGTAGA AAATGGAGCA WE02451 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAATTGATTA GTTCCTTTAA TATTAGGAGC ACCTGATATA GCCTTTCCAC GATTAAATAA TATAAGATTC TGATTACTAC CCCCATCATT AATATTATTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA WE02491 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGGGC ACCTGATATA GCCTTCCCAC GATTAAATAA TATAAGATTC TGATTACTAC CCCCATCATT AATACTACTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA

173 Appendix 3: Sequence data Cytochrome Oxidase I (COI)

....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| 185 195 205 215 225 235 245 255 265 275 285 295 305 315 325 335 345 355 WE02531 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATCG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGGGC ACCTGATATA GCCTTTCCAC GATTAAATAA TATAAGATTC TGATTATTAC CGCCATCATT AATACTACTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA WE02535 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GGGGATTTGG TAATTGATTA GTTCCTTTAA TACTGGGAGC ACCTGATATA GCCTTCCCTC GATTAAATAA TATAAGATTC TGATTACTTC CACCATCCTT AATACTACTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA WE02536 TTTATTATAA TTTTTTTTAT GGTTATACCT ATCATAATTG GAGGATTTGG TAACTGATTG GTTCCTTTAA TATTGGGAGC ACCTGATATG GCCTTTCCTC GATTAAATAA TATGAGATTT TGATTACTAC CGCCTTCACT AATACTACTA ATTTCCAGAA GAATCGTAGA AAATGGAGCA WE02591 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTGGGGGC ACCTGATATA GCCTTTCCAC GATTAAATAA TATAAGATTC TGATTATTGC CGCCATCATT AATACTACTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA WE02612 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTGGGGGC GCCTGATATA GCCTTTCCAC GATTAAATAA TATAAGATTC TGATTATTGC CGCCATCATT AATACTACTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA WE02614 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTGGGGGC ACCTGATATA GCCTTTCCAC GATTAAATAA TATAAGATTC TGATTATTGC CGCCATCATT AATACTACTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA WE02621 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTG GTTCCTTTAA TATTGGGGGC ACCTGATATA GCCTTTCCTC GATTAAATAA TATGAGATTT TGATTACTAC CACCTTCACT AATACTACTA ATTTCCAGAA GAATCGTAGA AAATGGAGCA WE02661 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATCG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGGGC ACCTGATATA GCCTTTCCAC GATTAAATAA TATAAGATTC TGATTATTAC CGCCATCATT AATACTACTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA WE02662 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATCG GAGGATTTGG TAATTGATTA GTTCCTTTAA TATTAGGGGC ACCTGATATG GCTTTCCCCC GATTAAATAA TATAAGATTC TGATTATTAC CCCCATCATT AATACTACTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA WE02671 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGAGC GCCTGATATA GCCTTTCCAC GATTAAATAA TATAAGATTC TGATTATTGC CACCATCATT AATACTACTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA WE02674 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTG GTTCCTTTAA TATTGGGGGC ACCTGATATA GCCTTTCCTC GATTAAATAA TATGAGATTT TGATTACTAC CGCCTTCACT AATACTATTA ATTTCCAGAA GAATCGTAGA AAATGGAGCA WE02676 TTTATTATAA TTTTTTTTAT GGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGAGC ACCTGATATA GCCTTTCCAC GATTAAATAA TATAAGATTC TGATTATTAC CACCATCATT AATACTACTA ATTTCTAGAA GAATTGTAGA AAATGGAGCA WE02677 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TACTGGGAGC ACCTGATATA GCATTCCCCC GATTAAATAA TATAAGATTC TGATTACTAC CACCATCCTT AATACTACTA ATTTCTAGAA GAATCGTAGA AAATGGAGCA WE85001 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAACTGATTA GTTCCTTTAA TATTAGGGGC ACCTGATATA GCCTTCCCAC GATTAAATAA TATAAGATTC TGATTACTAC CCCCATCATT AATACTACTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA WE98001 TTTATTATAA TTTTTTTTAT AGTTATACCT ATTATAATTG GAGGATTTGG TAATTGATTA GTTCCTTTAA TATTAGGAGC ACCTGATATA GCCTTCCCCC GATTAAATAA TATAAGATTC TGATTATTAC CACCATCATT AATACTATTA ATTTCCAGAA GAATTGTAGA AAATGGAGCA

....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| 365 375 385 395 405 415 425 435 445 455 465 475 485 495 505 515 525 535 AD98001 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC CCTTCACTTA GCAGGAATTT CTTCAATTTT AGGAGCaATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA AD98009 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC CCTTCATTTG GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT CAATATACGG GTAAATAAAT TATCTTTTGA TCAAATATCA AD98012 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAACA TTGCACATAG AGGATCGTCT GTAGATTTAG CAATTTTCTC TCTTCATTTG GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT CAATATACGG GTAAATAACC TATCTTTTGA TCAAATATCA AD98018 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGG GTAAATAATT TATCTTTTGA TCAAATATCA AD98024 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA AD98029 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACACAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCACTTA GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT tAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA AD98036 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACACAG AGGATCATCT GTAGATTTAG CAATTTTCTC CCTTCATTTA GCGGGAATTT CTTCAATTTT AGGGGCAATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAACT TAACCTTTGA TCAAATAtCA DS00001 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTGGATTTAG CAATTTTCTC TCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTCATTA CAACTATTAT TAATATACGG GTAAATAATT TATCTTTTGA TCAAATATCT DS00005 GGGACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAACA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTa CAACTATTAT TAATATACGG GTAAATAATT TATCTTTTGA TCAAATATCA DS01001 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAACATACGG GTAAATAATT TATCATTTGA TCAAATATCA JC00029 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACACAG AGGATCTTCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCTGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA JC00039 GGAACAGGAT GAACAGTTTA TCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTCCATTTA GCTGGAATTT CTTCAATTTT AGGGGCAATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAATT TATCCTTTGA TCAAATACCA JC00040 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCAAATA TTGCACATGG GGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATCAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA JC00042 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACACAG AGGATCTTCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCTGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAATT TATCCTTTGA TCAAATATCA JC00043 GGAACAGGGT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATGG AGGATCATCT GTAGATTTAG CAATTTTCTC ACTTCATTTA GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT CAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCC JC00045 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATGG AGGATCATCT GTAGATTTAG CAATTTTCTC ACTTCATTTA GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACtATTAT CAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCC JC00046 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATGG AGGATCATCT GTAGATTTAG CAATTTTCTC ACTTCATTTA GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT CAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCC JC00051 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACACAG AGGATCTTCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCTGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA JC00055 GGAACAGGAT GAACAGTCTA CCCCCCACTT TCATCTAATA TTGCTCATGG AGGATCATCT GTAGATTTAG CAATTTTTTC TCTTCATTTA GCTGGAATCT CTTCAATTTT AGGAGCTATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAATT TATCGTTTGA CCAAATATCA JC00057 GGAACAGGAT GAACAGTCTA CCCCCCACTT TCATCTAATA TTGCTCATGG AGGATCATCT GTAGATTTAG CAATTTTTTC TCTTCATTTA GCAGGAATCT CTTCAATTTT AGGAGCTATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAATT TATCATTTGA CCAAATATCA JC00061 GGAACAGGAT GAACAGTTTA TCCCCCACTT TCATCTAATA TTGCACACAG AGGATCTTCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCTGGGATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATCAT TAATATACGA GTAAATAATT TATCCTTTGA TCAAATATCA JC00062 GGAACAGGAT GAACAGTCTA CCCCCCACTT TCATCTAATA TTGCTCATGG AGGATCATCT GTAGATTTAG CAATTTTTTC TCTTCATTTA GCAGGAATCT CTTCAATTTT AGGAGCTATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAATT TATCaTTTGA CCAAATATCA JC00063 GGAACAGGAT GAACAGTTTA TCCCCCACTT TCATCTAATA TTGCACACAG AGGATCTTCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCTGGGATTT CTTCAATTTT AGGAGCaATT AATTTTATTA CAACTATCAT TAATAtACGA GTAAATAATT TATCCTTTGA TCAAATATCA JC01014 GGGACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATGG AGGATCATCT GTAGATTTAG CAATTTTCTC ACTTCATTTG GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT CAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCC JC02001 GGAACTGGAT GAACAGTCTA CCCCCCACTA TCATCTAATA TTGCACATAG AGGAAGATCT GTTGATCTTG CTATTTTTTC ACTTCACTTA GCTGGTATTT CATCTATTTT AGGTGCAATT AATTTTATTA CAACAATTAT TAATATACGA ATTAATAATA TATCATTTGA TCAAATACCT JM00001 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCAAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCAGGaATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACCATCAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW00001 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CTACTATTAT TAATATACGG GTAAATAATT TATCTTTTGA TCAAATATCA MW00015 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATGG AGGATCATCT GTAGATTTAG CAATTTTCTC ACTTCATTTA GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT CAATATGCGA GTAAATAATT TATCTTTTGA TCAAATATCT MW00017 GGAACAGGAT GAACAGTTTA CCCCCCaCTT TCATCTAATA TTGCTCATAG AGGATCATCT GTTGATTTAA CTATTTTTTC TTTACATTTA GCAGGTATTT CCTCAATTTT AGGAGCTATT AATTTTATTA CAACTATTAT TAATATACGA ATTAATAATT TATCTTTTGA TCAAATATCT MW00020 GGAACAGGAT GAACAGTCTA CCCCCCACTT TCATCTAATA TTGCTCATGG AGGATCATCT GTAGATTTAG CAATTTTTTC TCTTCATTTA GCAGGAATCT CTTCAATTTT AGGAGCTATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAATT TATCATTTGA CCAAATATCA MW00024 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACACAG AGGGTCATCT GTTGATTTAG CAATTTTTTC TCTTCATTTA GCTGGAATTT CTTCAATTTT AGGAGCTATT AATTTTaTTA CAACTATTAT TAATATGCGA GTAAACAATA TATCTTTTGA TCAAATATCT MW00032 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CTACTATTAT TAATATACGG GTAAATAATT TATCTTTTGA TCAAATATCA MW00044 GGAACAGGAT GAACAGTTTA TCCCCCACTT TCATCTAATA TTGCTCATAG AGGATCTTCT GTTGATTTAG CTATTTTCTC TTTACATTTA GCAGGTATCT CTTCAATTTT AGGAGCTATT AATTTTATTA CAACTATTAT TAATATACGA ATTAATAATT TATCATTTGA TCAAATATCT MW00051 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC GCTTCATTTA GCCGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW00056 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC GCTTCATTTA GCCGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW00058 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC GCTTCATTTA GCCGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW00059 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC GCTTCATTTA GCCGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW00060 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACACAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAACT TATCCTTTGA TCAAATATCA MW00064 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCAAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATCAT TAATAtGCGG GTAAATAATT TATCTTTTGA TCAAATATCA MW00072 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACACAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAACT TATCCTTTGA TCAAATATCA MW00076 GGAACAGGAT GAACGGTTTA CCCCCCGCTT TCATCTAATA TTGCACATAG AGGGTCATCT GTAGACTTAG CAATTTTCTC ACTTCATTTA GCTGGAATTT CTTCAATTTT AGGGGCAATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW00087 GGAACAGGAT GAACGGTTTA CCCCCCGCTT TCATCTAATA TTGCACATAG AGGGTCATCT GTAGACTTAG CAATTTTCTC ACTTCATTTA GCTGGAATTT CTTCAATTNT AGGGGCAATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW00101 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACACAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW00102 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACACAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAACT TATCCTTTGA TCAAATATCA MW00110 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC GCTTCATTTA GCCGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW00116 GGAACAGGAT GAACAGTCTA CCCCCCACTT TCATCTAATA TTGCACACAG AGGATCATCT GTAGATTTAG CAATTTTCTC ACTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCTATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW00119 GGAACAGGAT GAACAGTTTA TCCCCCACTT TCATCTAATA TTGCCCATAG AGGATCGTCT GTTGATTTAG CTATTTTTTC TTTACATTTA GCGGGAATTT CTTCAATTTT AGGAGCTATT AATTTTATTA CAACTATTAT TAATATACGA ATTAATAATT TATCTTTTGA CCAAATATCT MW00125 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC CCTTCATTTA GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT CAATATACGG GTAAATAAAT TATCTTTTGA TCAAATATCA MW00127 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCNCATAG AGGaTCATCT GTAGATTTAG CaATTtTCTC GCTTCATTTA GCCGGAaTTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAATT TATCTTTTga TCAAATATCA MW00129 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAACA TTGCACATAG aGGaTCATCT GTAGATTTaG CAATTTTTTC TCTTCATTTG GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT CAATATACGA GTAAATAACC TATCTTTTGA TCAAATATCA

174 Appendix 3: Sequence data Cytochrome Oxidase I (COI)

....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| 365 375 385 395 405 415 425 435 445 455 465 475 485 495 505 515 525 535 MW00140 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACACAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCGGGAATTT CTTCAATTTT AGGGGCAATT AATTTTATTA CAACTATTAT TAATATACGG GTAAATAACT TAACCTTTGA TCAAATATCA MW00151 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC CCTTCATTTG GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT CAATATACGA GTAAATAAAT TATCTTTTGA TCAAATATCA MW00157 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC GCTTCATTTA GCCGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW00176 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACACAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTCCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGG GTAAATAATT TATCTTTTGA TCAAATATCA MW00177 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAACA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTTTC TCTTCATTTG GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTa CAACTATTAT CAATATACGA GTAAATAACC TATCTTTTGA TCAAATATCA MW00178 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAACA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTTTC TCTTCATTTG GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT CAATATACGA GTAAATAACC TATCTTTTGA TCAAATATCA MW00179 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAACA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTTTC TCTTCATTTG GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT CAATATACGA GTAAATAACC TATCTTTTGA TCAAATATCA MW00183 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATGG AGGATCATCT GTAGATTTAG CAATTTTCTC ACTTCATTTA GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT CAATATGCGA GTAAATAATT TATCTTTTGA TCAAATATCT MW00185 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATGG AGGATCATCT GTAGATTTAG CAATTTTCTC ACTTCATTTA GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT CAATATGCGA GTAAATAATT TATCTTTTGA TCAAATATCT MW00186 GGAACAGGAT GAACAGTTTA CCCNCCAcTT TCATCTAATA TTGCACATGG AGGATCNTCT GTAGANTTAG CaATTTTCTC ACTTCATTTA GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATtaT CAATATGCGA GTAAATAATT TATCTtTTGA TCAAATATCT MW00189 GGAACAGGGT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATGG GGGATCATCT GTAGATTTAG CAATTTTTTC ACTTCATTTA GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT CAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCC MW00226 GGAACAGGGT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC CCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATGCGA GTAAATAATT TATCTTTTGA TCAAATATCA MW00229 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCAAATA TCGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATCAT TAATATACGA GTAAATAATT TATCCTTTGA TCAAATATCA MW00231 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCAAATA TCGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATCAT TAATATACGA GTAAATAATT TATCCTTTGA TCAAATATCA MW00232 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC GCTTCATTTA GCCGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW00234 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC CCTTCATTTA GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT CAATATACGG GTAAATAAAT TATCTTTTGA TCAAATATCA MW00259 GGAACGGGAT GAACAGTTTA TCCCCCACTT TCATCTAATA TTGCGCACAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAACT TATCCTTTGA TCAAATATCA MW00262 GGAACGGGAT GAACAGTTTA TCCCCCACTT TCATCTAATA TTGCGCACAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAACT TATCCTTTGA TCAAATATCA MW00267 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACACAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAACT TATCCTTTGA TCAAATATCA MW00269 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TtGCACATAG AGGATCGTCT GTAGATTTAG CAATTTTCTC CCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW00290 GGAACAGGGT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACACAG AGGATCATCT GTAGACTTAG CAATTTTCTC CCTTCATTTA GCTGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAACATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW00302 GGGACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTCCATTTA GCTGGAATTT CCTcAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGG GTAAATAATT TATCTTTTGA TCAAATATCA MW00316 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC GCTTCATTTA GCCGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW00326 GGGACAGGAT GAACAGTTTA CCCCCCACTT TCATCCAATA TTGCGCATAG AGGATCATCT GTAGATTTAG CAATTTTCTC TTTACATTTA GCTGGAATTT CTTCAATTTT AGGAGCTATT AATTTTATTT CAACTATTAT TAATATACGA GTAAATAATT TATCATTTGA TCAAATATCA MW00327 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCNCACAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTaTtAT TaATATACGG GTAAATAACT TATCCTTTGA TCAAATATCA MW00328 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCGTCT GTAGATTTAG CAATTTTCTC CCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW00330 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAACA TTGCACATAG AGGATCATCT GTAGATTTAG CTATTTTTTC TCTTCATTTG GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT CAATATACGG GTAAATAACC TATCTTTTGA TCAAATATCA MW00335 GGAACAGGAT GAACAGTTTA TCCCCCACTT TCATCTAATA TTGCACACAG AGGATCATCT GTAGACTTAG CAATTTTCTC TCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAACATACGG GTAAATAACT TATCCTTTGA TCAAATATCA MW00347 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACACAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAACT TATCTTTTGA TCAAATATCA MW00348 GGAACGGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACACAG AGGGTCATCT GTGGATTTAG CAATTTTCTC TCTTCATTTA GCGGGAATTT CTTCAATTTT AGGaGCAATT AATTTTATTA CAACTATTAT TAATATACGG GTAAATAACT TAACCTTTGA TCAAATATCA MW00358 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACACAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAACT TATCCTTTGA TCAAATATCA MW00393 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACACAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGG GTAAATAACT TATCCTTTGA TCAAATATCA MW00409 GGAACAGGAT GAACAGTTTA TCCCCCACTT TCATCTAATA TTGCACACAG AGGATCATCT GTAGACTTAG CAATTTTCTC TCTTCATTTA GCAGGAATTT CTTCAATTGT AGGAGCAATT AATTTTATTA CAACTATTAT TAACATACGA GTAAATAACT TATCCTTTGA TCAAATATCA MW00412 GGAACAGGAT GAACAGTTTA TCCCCCACTT TCATCTAATA TTGCACACAG AGGATCTTCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCTGGGATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATCAT TAATATACGA GTAAATAATT TATCCTTTGA TCAAATATCA MW00424 GGaACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCGTCT GTAGATTTAG CAATTTTCTC CCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW00444 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACACAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAACT TATCCTTTGA TCAAATATCA MW00452 GGAACGGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACACAG AGGGTCATCT GTGGATTTAG CAATTTTCTC TCTTCATTTA GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGG GTAAATAACT TAACCTTTGA TCAAATATCA MW00469 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCTCATAG AGGATCATCT GTAGATTTAG CAATTTTCTC ACTTCATTTA GCTGGAATTT CTTCAATTTT AGGAGCTATT AACTTTATTA CAACTATTAT TAATATACGG GTAAATAATT TATCTTTTGA TCAAATATCA MW00476 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACACAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAACT TATCCTTTGA TCAAATATCA MW00497 GGAACAGGAT GAACAGTATA CCCCCCACTT TCATCTAATA tCGCCCATAG AGGAGCTTCT GTTGATTTAG CTATTTTTTC ATTACATTTA GCTGGTATTT CTTCAATTTT AGGAGCTATT AACTTTATTA CAACTATTAT TAATATACGA ATTAGTAATT TATCCTTTGA TCAAATATCA MW00498 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCAAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATCAT TAATATGCGG GTAAATAATT TATCTTTTGA TCAAATATCA MW00504 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC CCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATAtACGA GTAAATAATT TATCCTTTGA TCAAATATCA MW00517 GGAACAGGAT GAACAGTTTA TCCCCCACTT TCATCTAATA TTGCTCATGG AGGATCATCT GTAGATTTAG CAATTTTTTC TCTTCATTTA GCCGGAATTT CTTCAATTTT AGGGGCTATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATaATT TATCTTTTGA TCAAATATCA MW00525 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACACAG AGGATCATCT GTAGATTTAG CAATCTTTTC TCTTCATTTA GCTGGAATTT CTTCAATTCT AGGAGCAATT AATTTTATTA CCACTATTAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCT MW00530 GGAACAGGAT GAACAGTTTA TCCCCCACTT TCATCTAATA TTGCACACAG AGGATCTTCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCTGGAATTT CTTCAATTTT AGGGGCAATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAATT TATCCTTTGA TCAAATATCA MW00539 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC CCTTCACTTA GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CTACTATTAT TAATAtACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW00547 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACACAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGA GTGAATAACT TATCCTTTGA TCAAATATCA MW01001 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCAAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACCATCAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW01009 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTCCATTTA GCTGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT CAACATACGA GTAAATAATT TATCCTTTGA TCAAATACCA MW01011 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCTCATAG AGGATCATCT GTAGATTTAG CAATTTTCTC ACTTCATTTA GCTGGAATTT CTTCAATTTT AGGAGCTATT AACTTTATTA CAACTATTAT TAATATACGG GTAAATAATT TATCTTTTGA TCAAATATCA MW01014 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATGG AGGATCATCT GTAGATTTAG CAATTTTCTC CCTTCATTTA GCGGGAATTT CTTCAATTTT AGGGGCAATT AATTTTATTA CAACTATTAT CAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCC MW01018 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCTCATAG AGGATCATCT GTAGATTTAG CAATTTTCTC ACTTCATTTA GCTGGAATTT CTTCAATTTT AGGAGCTATT AACTTTATTA CAACTATCAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW01019 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCA GTAGATTTAG CAATTTTCTC TCTTCATTTA GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAATT TATCCTTTGA TCAAATATCA MW01022 GGAACAGGAT GAACAGTGTA CCCCCCACTT TCATCTAATA TTGCACATGG AGGTTCATCT GTTGATTTAG CAATTTTTTC ATTACACTTA GCTGGTATTT CTTCAATTTT AGGAGCTATT AATTTTATTA CTACTATTAT TAATATACGA GTAAATAATT TATCATTTGA TCAAATATCA MW01023 GGAACAGGAT GAACAGTGTA TCCCCCACTA TCATCTAATA TTGCTCATGG AGGATCTTCT GTAGATTTAG CTATTTTTTC TTTACATTTA GCTGGAATTT CTTCAATTTT AGGTGCTATT AATTTTATTA CAACTATTAT TAATATACGA GTTAATAATT TATCTTTTGA CCAAATATCT MW01025 GGAACAGGAT GAACAGTTTA TCCCCCACTT TCATCTAATA TTGCACACAG AGGATCTTCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCTGGGATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATCAT TAATATACGA GTAAATAATT TATCCTTTGA TCAAATATCA MW01027 GGAACTGGAT GAACAGTTTA TCCCCCACTT TCATCTAATA TTGCTCATAG AGGATCATCA GTTGATTTAG CCATTTTTTC TCTTCATTTA GCAGGAATTT CATCAATTTT AGGAGCTATT AATTTTATTA CAACTATTAT TAATATACGC ATTAATAATT TATCTTTTGA TCAAATATCA MW01034 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCTCATAG AGGATCATCT GTAGATTTAG CAATTTTCTC ACTTCATTTA GCTGGAATTT CTTCAATTGT AGGAGCTATT AACTTTATTA CAACTATCAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW01039 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCAAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACCATCAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW01048 GGAACAGGAT GAACAGTCTA CCCCCCACTT TCATCTAATA TTGCTCATAG AGGATCATCT GTAGATTTAG CAATTTTTTC TCTTCATTTA GCAGGAaTCT CTTCAATTTT AGGAGCTATT AATTTTATTA CAACTATTAT TAACATACGA GTAAATAATT TATCATTTGA CCAAATATCA MW01053 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCAAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACCATCAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW01059 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCTCATAG AGGATCATCT GTAGATTTAG CAATTTTCTC ACTTCATTTA GCTGGAATTT CTTCAATTTT AGGAGCTATT AACTTTATTA CAACTATCAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW01061 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCTCATGG AGGATCATCT GTAGATTTAG CAATTTTTTC TCTTCATTTA GCTGGAATTT CTTCAATTTT AGGAGCTATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAATT TATCATTTGA CCAAATATCA MW01072 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATGG AGGATCATCT GTAGATTTAG CAATTTTCTC CCTTCATTTA GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT CAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCC MW01078 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCAAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACCATCAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW01083 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCCAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC CCTCCATTTA GCTGGAATTT CCTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW01092 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCTCATAG AGGATCATCT GTAGATTTAG CAATTTTCTC ACTTCATTTA GCTGGAATTT CTTCAATTTT AGGAGCTATT AACTTTATTA CAACTATCAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW01103 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCTCATAG AGGATCATCT GTAGATTTAG CAATTTTCTC ACTTCATTTA GCTGGAATTT CTTCAATTTT AGGAGCTATT AACTTTATTA CAACTATCAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA

175 Appendix 3: Sequence data Cytochrome Oxidase I (COI)

....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| 365 375 385 395 405 415 425 435 445 455 465 475 485 495 505 515 525 535 MW01105 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATGG AGGATCATCT GTAGATTTAG CAATTTTCTC ACTTCATTTA GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT CAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCC MW01107 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCAAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACCATCAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW01114 GGAACTGGAT GAACAGTCTA CCCCCCACTA TCATCTAATA TTGCACATAG AGGAAGATCT GTTGATCTTG CTATTTTTTC ACTTCACTTA GCTGGTATTT CATCTATTTT AGGTGCAATT AATTTTATTA CAACAATTAT TAATATACGA ATTAATAATA TATCATTTGA TCAAATACCT MW01116 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCTCATAG AGGATCATCT GTAGATTTAG CAATTTTCTC ACTTCATTTA GCTGGAATTT CTTCAATTTT AGGAGCTATT AACTTTATTA CAACTATCAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW01120 GGAACAGGAT GAACAGTGTA CCCCCCACTA TCATCTAATA TTGCCCATGG TGGATCATCC GTTGATTTAG CAATTTTTTC TTTACACCTA GCAGGAATTT CATCTATTTT AGGAGCTATT AATTTTATTA CAACTATCAT TAATATACGA GTTAATAATC TTTCATTCGA TCAAATATCT MW02001 GGAACAGGAT GAACAGTTTA TCCCCCACTT TCATCTAATA TTGCACATAG TGGATCATCC GTAGACTTAG CAATTTTCTC ACTTCATTTA GCTGGAATTT CTTCAATTTT AGGAGCTATT AATTTTATTA CAACTATCAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCC MW02006 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TCGCACATAG TGGATCTTCT GTAGATTTAG CGATTTTCTC TCTTCATTTA GCTGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAACATACGA GTAAGTAATT TATCCTTTGA TCAAATATCA MW02007 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCCCATAG AGGGTCATCT GTTGATTTAG CTATTTTTTC ATTACATTTA GCAGGTATTT CATCAATTTT AGGAGCTATT AATTTTATTA CAACTATTAT TAATATACGA ATTAATAATT TATCTTTTGA TCAAATATCT MW02008 GGAACAGGAT GAACAGTGTA CCCCCCACTA TCATCTAATA TTGCACATAG TGGATCATCT GTAGATCTAG CAATTTTCTC ATTACATTTA GCAGGTATTT CATCAATTTT AGGAGCTATT AATTTTATTA CAACTATTAT CAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW02009 GGAACAGGAT GAACAGTTTA TCCCCCACTT TCATCTAATA TTGCTCATAG AGGATCTTCT GTTGATTTAG CTATTTTTTC TTTACATTTA GCAGGTATTT CTTCAATTCT AGGAGCTATT AATTTTATTA CAACTATTAT TAATATACGA ATTAATAATT TATCCTTTGA TCAAATATCA MW02021 GGAACTGGAT GAACTGTGTA CCCCCCaCTA TCATCAAATA TTGCTCATAG AGGATCTTCT GTTGATTTAG CTATTTTTTC TTTACATTTA GCCGGAATTT CTTCAATTTT AGGTGCAATT AATTTTATTA CAACTATTAT TAACATACGT GTAAATAATT TATCTTTTGA CCAAATATCA MW02024 GGAACTGGAT GAACAGTCTA CCCCCCACTA TCATCTAATA TTGCCCATAG AGGAAGATCT GTCGATCTTG CTATTTTCTC ACTTCACTTA GCTGGTATTT CTTCTATTTT AGGTGCAATT AATTTTATTA CAACAATTAT TAATATACGA ATTAATAATA TATCATTTGA TCAAATaCCT MW02025 GGAACAGGAT GAACAGTGTA CCCCCCACTA TCATCAAATA TTGCACATAG AGGATCTTCA GTCGATTTAG CAATTTTCTC TCTTCATTTA GCAGGAATTT CATCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGG GTAAATAATC TATCATTTGA TCAAATATCA MW02028 GGAACAGGAT GAACAGTGTA CCCCCCACTT TCATCTAATA TTGCACATGG AGGTTCATCT GTTGATTTAG CAATTTTTTC ATTACACTTA GCTGGTATTT CTTCAATTTT GGGAGCTATT AATTTTATTA CTACTATTAT TAATATACGA GTAAATAATT TATCATTTGA TCAAATATCA MW02031 GGAACAGGAT GAACAGTGTA CCCCCCACTT TCATCTAATA TTGCTCATAG AGGTTCATCT GTTGATTTAG CAATTTTTTC TCTTCATTTA GCAGGAATTT CATCAATTTT AGGAGCAATT AATTTTATTA CTACAATTAT TAATATACGA GTAAATAATT TATCATTTGA TCAAATATCA MW02033 GGAACAGGAT GAACAGTCTA CCCCCCACTT TCATCTAATA TTGCTCATAG AGGATCATCT GTAGATTTAG CAATTTTTTC TCTTCATTTA GCAGGAATCT CTTCAATTTT AGGAGCTATT AATTTTATTA CAACTATTAT TAACATACGA GTAAATAATT TATCATTTGA CCAAATATCA MW02034 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATGG AGGATCATCT GTAGATTTAG CAATCTTTTC TCTTCATTTA GCTGGAATTT CTTCAATTCT AGGAGCAATT AATTTTATTA CTACTATTAT TAATATACGG GTAAATAATT TATCTTTTGA TCAAATATCT MW98002 GGAACAGGAT GAACAGTGTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTTGATTTAG CAATTTTTTC TTTACACTTA GCAGGAATTT CATCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGA GTTAATAATT TATCATTTGA TCAAATATCA MW98008 GGGACAGGAT GAACAGttTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTCCATTTA GCTGGAATTT CCTCAaTTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGG GTAAATAATT TATCTTTTGA TCAAATATCA MW98009 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC CCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW98020 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCAAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATCAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW98029 GGAACAGGAT GAACGGTTTA CCCCCCACTT TCATCTAATA TTGCACACAG AGGATCATCT GTAGACTTAG CAATTTTCTC ACTTCATTTA GCTGGAATTT CTTCAATTTT AGGGGCAATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW98049 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACACAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW98079 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC CCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTaT caATATACGG GTATATAAAT TATCTTTTGA TCAAATATCA MW98084 GGGACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATGG AGGATCATCT GTAGATTTAG CAATTTTCTC ACTTCATTTG GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT CAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCC MW98089 GGAaCAGGaT GAAcAGTTTA CCCCCCACTT TCATCTAATa TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC CCTTCATTTA GCAGGAATTT CTTCAATTTT AGGGGCAATT AATTTTATTA CAACTATtAT CAATATACGG GTAAATAAAT TATCTTTTGA TCAAATATCA MW98094 GGAACAGGAT GAACAGTTTA TCCCCCACTT TCATCtAATA TTGCTCATAG AGGATCATCT GTGGATTTAG CtATTTTTTC TTTACATTTA GCAGGTATTT CTTCAATTTT AGGAGCTATT AATTTTATCA CAACTATTAT TAATATACGA ATTAATAATT TATCTTTTGA TCAAATATCT MW98097 GGAACAGGAT GAACAGTCTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC CCTTCATTTA GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT CAATATACGG GTAAATAAAT TATCTTTTGA TCAAATATCA MW98103 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC CCCCCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGG GTAAATAATT TATCTTTTGA TCAAATATCA MW98106 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACACAG AGGATCATCT GTTGATTTAG CAATTTTCTC TCTTCATTTA GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW98128 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCA GTAGATTTAG TAATTTTCTC TCTTCATTTG GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAATT TATCCTTTGA TCAAATATCA MW98129 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCTCATAG AGGATCATCT GTAGATTTAG CAATTTTCTC ACTTCATTTA GCTGGCATTT CTTCAATTTT AGGAGCTATT AACTTTATTA CAACTATCAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW98133 GGAACAGGAT GAACAGTTTA TCCCCCACTT TCATCTAATA TTGCACACAG AGGATCTTCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCTGGGATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATCAT TAATATACGA GTAAATAATT TATCCTTTGA TCAAATATCA MW98136 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAACA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTTTC TCTTCATTTG GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CaACTATTAT CAATATACGA GTAAATAACC TATCCTTTGA TCAAATATCA MW98138 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC CCTTCATTTA GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT CAACATACGG GTAAATAAAT TATCTTTTGA TCAAATATCA MW98139 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAACA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTTTC TCTTCATTTG GCAGGAATTT CTTCAATTTT AGGAGCaATT AATTTTATTA CAACTATTAT CAATATACGA GTAAATAACC TATCTTTTGA TCAAATATCA MW98154 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC CCTTCATTTA GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT CAATATACGG GTAAATAAAT TATCTTTTGA TCAAATATCA MW98162 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC CCTTCATTTG GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT CAATATACGG GTAAATAAAT TATCTTTTGA TCAAATATCG MW98170 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAACA TTGCACATAG aGGATCATCT GTAGATTTAG CAATTTTTTC TCTTCATTTG GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT CAATATACGA GTAAATAACC TATCCTTTGA TCAAATATCA MW98172 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCAAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATCAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW98180 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC CCTTCATTTA GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT CAATATACGG GTAAATAAAT TATCTTTTGA TCAAATATCA MW98183 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC CCTTCATTTA GCGGGAATTT CTTCAATTTT AGGAGCaATT AATTTTATTA CAACTATTAT CAATATACGG GTAAATAAAT TATCTTTTGA TCAAATATCA MW98185 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCTCATAG AGGATCATCT GTAGATTTAG CAATTTTCTC ACTTCATTTA GCTGGAATTT CTTCAATTTT AGGAGCTATT AATTTTATTA CAACTATCAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW98189 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC CCTTCATTTG GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT CAATATACGG GTAAATAAAT TATCTTTTGA TCAAATATCA MW98203 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTTTC TCTTCATTTG GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT CAATATACGA GTAAATAACC TATCCTTTGA TCAAATATCA MW98205 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAACATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW98212 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCAAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATCAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW98220 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATGG AGGATCATCT GTAGATTTAG CAATTTTTTC tCTTCATTTA GCTGGAATTT CTTCAATTTT AGGAGCTATT AATTTTATCA CAACAATTAT TAATATACGA GTAAATAGTT TATCTTTTGA TCAAATATCA MW98228 GGAACAGGAT GAACAGTTTA TCCCCCACTT TCATCTAATA TTGCTCATAG AGGATCATCT GTAGATTTAG CAATTTTCTC ACTTCATTTA GCTGGAATTT CTTCAATTTT AGGAGCTATT AACTTTATTA CAACTATCAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW98230 GGAACAGGAT GAACAGTTTA TCCCCCACTT TCATCTAATA TTGCCCATAG AGGATCATCT GTTGATTTAG CTATTTTTTC CTTACATTTA GCAGGTATTT CATCAATTTT AGGAGCTATT AACTTTATTA CAACTATTAT TAATATACGA ATTAATAATT TATCATTTGA TCAAATATCT MW98235 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TCGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC TTTACACTTG GCTGGAATTT CTTCAATTTT AGGAGCTATT AATTTTATTA CAACTATTAT TAACATACGA GTAAATAATT TATCATTTGA TCAAATATCA MW98240 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC CCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGG GTAAATAATT TATCTTTTGA TCAAATATCA MW98261 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC CCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW98264 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TCGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC TTTACACTTG GCTGGAATTT CTTCAATTTT AGGAGCTATT AATTTTATTA CAACTATTAT TAACATACGA GTAAATAATT TATCATTTGA TCAAATATCA MW98265 GGAACAGGAT GAACAGTGTA CCCCCCACTT TCATCTAATA TTGCACATGG AGGTTCATCT GTTGATTTAG CAATTTTTTC ATTACACTTA GCTGGTATTT CTTCAATTTT GGGAGCTATT AATTTTATTA CTACTATTAT TAATATACGA GTAAATAATT TATCATTTGA TCAAATATCA MW98270 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTTTC TCTTCATTTA GCTGGAATTT CTTCAATCTT AGGAGCTATT AATTTTATTA CAACTATCAT TAATATACGA GTAAATAATT TATCTTTTGA CCAAATATCA MW98274 GGAACAGGAT GAACGGTTTA CCCCCCGCTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGACTTAG CAATTTTCTC GCTTCATTTA GCTGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAACATGCGA GTAAATAATT TATCTTTTGA TCAAATATCA MW98278 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCTCATAG AGGATCATCT GTAGATTTAG CAATTTTCTC ACTTCATTTA GCTGGAATTT CTTCAATTTT AGGAGCTATT AACTTTATTA CAACTATTAT TAATATACGG GTAAATAATT TATCTTTTGA TCAAATATCA MW98284 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC CCTTCATTTG GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT CAATATACGG GTAAATAAAT TATCTTTTGA TCAAATATCG MW98285 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC CCTTCATTTA GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT CAATATACGG GTAAATAAAT TATCTTTTGA TCAAATATCA MW98294 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATGG AGGATCATCT GTAGATTTAG CAATTTTCTC CCTTCATTTA GCAGGAATTT CTTCAATTTT AGGGGCAATT AATTTTATTA CAACTATTAT TAATATACGG GTAAATAATT TATCTTTTGA TCAAATATCA MW98313 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAACA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTTTC TCTTCATTTG GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT CAATATACGA GTAAATAACC TATCTTTTGA TCAAATATCA MW98315 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCAAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATCAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW99001 GGAACAGGAT GAACAGTCTA CCCCCCACTT TCATCTAATA TTGCTCATAG AGGATCATCT GTAGATTTAG CAATTTTTTC TCTCCATTTA GCTGGAATTT CTTCAATTTT AGGAGCTATT AATTTTATTA CAACTATTAT TAACATACGG GTAAATAACT TATCTTTTGA CCAAATATCA MW99006 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC CCTTCATTTA GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT CAATATACGG GTAAATAAAT TATCTTTTGA TCAAATATCA MW99009 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACACAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW99013 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCaCATAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCTGGAATTT CTTCAATTTT AGGAGCTATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAAAC TATCCTTTGA TCAAATATCA MW99014 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCA GTAGATTTAG TAATTTTCTC TCTTCATTTG GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAACATACGA GTAAATAATT TATCCTTTGA TCAAATATCA

176 Appendix 3: Sequence data Cytochrome Oxidase I (COI)

....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| 365 375 385 395 405 415 425 435 445 455 465 475 485 495 505 515 525 535 MW99018 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCTCATGG AGGATCATCT GTAGATTTAG CAATTTTTTC TCTTCATCTA GCTGGAATTT CTTCAATTTT AGGAGCTATT AATTTTATTA CAACTATTAT TAATATACGa GTAAATAATT TATCTTTTGa TCAAATaTCa MW99028 GGAACAGGAT GAACAGTCTA CCCCCCACTT TCATCTAATA TtGCTCATGG AGGATCATCT GTAGATTTAG CAATTTTTTC TCTTCATTTA GCAGGAATCT CTTCAATTTT AGGAGCTATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAATT TATCATTTGA CCAAATATCA MW99038 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TCGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC TTTACACTTG GCTGGAATTT CTTCAATTTT AGGAGCTATT AATTTTATTA CAACTATTAT TAACATACGA GTAAATAATT TATCATTTGA TCAAATATCA MW99042 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCtCATAG AGGATCATCT GTAGATTTAG CAATTTTCtC ACTTCATTTA GCTGGCATTT CTTCAATTTT AGGAGCTATT AACTTTATTA CAACTAGCAT TAATATACGA GTaAATAATT TATCTTTTGA TCAAATATCG MW99045 GGAACAGGAT GAACAGTGTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTTGATTTAG CAATTTTTTC TCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCTATT AATTTTATTA CAACAATTAT TAATATAAAA GTAAATAATT TATTTTTTGA TCAAATATCA MW99047 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCTGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGG GTAAATAATT TATCTTTTGA TCAAATATCA MW99053 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGG GTAAATAATT TATCTTTTGA TCAAATATCA MW99057 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAACA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTG GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT CAATATACGA GTAAATAACC TATCTTTTGA TCAAATATCA MW99058 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC CCTTCATTTG GCGGGAATTT CTTcAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT CAATATACGG GTGAATAAAT TATCTTTTGA TCAAATATCA MW99059 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAACA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTTTC TCTTCATTTG GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT CAATATACGA GTAAATAACC TATCTTTTGA TCAAATATCA MW99060 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC CCTTCACTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW99061 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC CCTTCATTTA GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT CAATATACGG GTAAATAAAT TATCTTTTGA TCAAATATCA MW99068 GGaACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATGG AGGATCATCT GTAGATTTAG CAATTTTCTC ACTTCATTTA GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTaT CaATATACGA GTAAATAATT TaTCTTTTGA TCAAATATCC MW99080 GGAACAGGAT GAACAGTGTA CCCCCCACTT TCATCTAATA TTGCCCATAG AGGATCATCT GTTGATTTAG CAATTTTTTC ACTTCATTTA GCAGGAATTT CATCAATTTT AGGAGCAATT AATTTTATTA CTACAATTAT TAATATACGA GTAAATAATT TATCATTTGA TCAAATATCA MW99084 GGAACAGGAT GAACAGTGTA CCCCCCACCA TCATCTAATA TTGCaCATAG TGGAtCATCT GTAGATCTAG CAATTTTCTC ATTACATTTA GCAGGTATTT CATCAATTTT AGGAGCTATT AATTTTATTA CAACTATTAT CAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCC MW99094 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAACA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTTTC TCTTCATTTG GCGGGAATTT CTTCAATTTT AGGNGCAATT AATTTTATTA CAACTATTAT CaATATACGA GTAAATAACC TATCTTTTGA TCAAATATCA MW99095 GGAaCAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW99097 GGAACAGGAT GAACAGTCTA TCCCCCACTT TCATCTAATA TTGCTCATAG AGGATCATCT GTAGATTTAG CAATTTTTTC TCTCCATTTA GCTGGAATTT CTTCAATTTT AGGAGCTATT AATTTTATTA CAACTATTAT TAACATACGA GTAAATAACT TATCTTTTGA CCAAATATCA MW99102 GGAACAGGGT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATGG AGGATCATCT GTAGATTTAG CAATTTTCTC ACTTCATTTA GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT CAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW99104 GGAACAGGGT GAACAGTTTA CCCTCCACTT TCATCTAATA TTGCACATGG GGGATCATCT GTAGATTTAG CAATTTTCTC ACTTCATTTA GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT CAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCC MW99105 GGAACAGGGT GAACAGTTTA CCCTCCACTT TCATCTAATA TTGCACATAG GGGATCATCT GTAGATTTAG CAATTTTCTC ACTTCATTTA GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT CAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCC MW99124 GGAACAGGAT GAACAGTTTA CCCACCACTT TCATCtAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC ACTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCTATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW99128 GGGACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTCCATTTA GCTGGAATTT CCTCAATTTT aGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGG GTAAATAATT TATCTTTTGA TCAAATATCA MW99134 GGAACAGGAT GAACGGTTTA CCCCCCGCTT TCATCAAATA TTGCACATAG AGGATCATCT GTTGATTTAG CAATTTTTTC ACTTCATTTA GCTGGAATTT CTTCTATTTT AGGAGCTATT AACTTTATTA CTACTATTAT TAACATACGA GTNAATAATT TATCTTTTGA TCAATTATCA MW99135 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACACAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTC CAACTATTAT TAATAGACGG GTAAATAATT TATCTTTTGA TCAAATATCA MW99140 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCTCATAG AGGATCATCT GTAGATTTAG CAATTTTCTC ACTTCATTTA GCTGGGATTT CTTCAATTTT AGGAGCTATT AACTTTATTA CAACTATCAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW99141 GGAACAGGGT GAACAGTTTA CCCTCCACTT TCATCTAATA TTGCACATGG GGGATCATCT GTAGATTTAG CAATTTTCTC ACTTCATTTA GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT CAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCC MW99158 GGGACTGGAT GAACAGTTTA TCCCCCACTT TCATCCAATA TTGCCCATAG AGGATCATCA GTTGATTTAG CTATTTTTTC TTTACATTTA GCAGGTATTT CATCAATCTT AGGAGCTATT AATTTTATTA CAACTATTAT TAATATACGA aTTAATAATT TATCTTTTGA TCAAATATCA MW99163 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCAAATA TTGCACATAG AGGATCCTCA GTAGATCTAG CAATTTTTTC TCTTCATCTA GCTGGAATTT CTTCAATTTT AGGAGCTATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCT MW99164 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCAAATA TCGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATcAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW99165 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAACA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTTTC TCTTCATTTG GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT CAATATACGA GTAAATAACC TATCCTTTGA TCAAATATCA MW99170 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACACAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW99174 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACACAG AGGATCATCT GTAGATTTAG CAATTTTCTC CCTTCATTTA GCGGGAATTT CTTCAATTTT AGGGGCAATT AATTTTATTA CAACTATTAT TAATATACGG GTAAATAACT TAACCTTTGA TCAAATATCA MW99187 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACACAG AGGATCATCT GTAGATTTAG CAATTTTCTC CCTTCATTTA GCAGGAATTT CTTCAATTTT NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN MW99196 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATGG AGGATCATCT GTAGATTTAG CAATTTTCTC ACTTCATTTA GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT CAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCC MW99203 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACACAG AGGATCATCT GTAGACTTAG CAATTTTCTC TCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGG GTAAATAATT TATCTTTTGA TCAAATATCA MW99221 GGAACAGGAT GAACAGTGTA CCCCCCACTC TCATCTAATA TTGCTCATAG AGGTTCTTCT GTTGATTTAG CTATTTTTTC TCTTCATTTA GCTGGAATCT CTTCAATTTT AGGAGCTATT AATTTTATTA CAACAATTAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW99226 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGG GTAAATAATT TATCTTTTGA TCAAATATCT MW99240 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAACA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTTTC TCTTCATTTG GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT CAATATACGA GTAAATAACC TATCCTTTGA TCAAATATCA MW99247 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC CCTTCATTTG GCGGGAATTT CTTCAATCTT AGGAGCAATT AATTTTATTA CAACTATTAT CAATATACGA GTAAATAAAT TATCTTTTGA TCAAATATCA MW99263 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATGG AGGATCATCT GTAGATTTAG CAATTTTCTC ACTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT CAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCC MW99264 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATGG AGGATCATCT GTAGATTTAG CAATTTTCTC ACTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT CAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCC MW99274 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCAAATA TCGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATCAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW99276 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCAAATA TCGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATCAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW99286 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC CCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATGCGA GTAAATAATT TATCTTTTGA TCAAATATCA MW99289 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC CCTTCACTTA GCAGGGATTT CTTCAATTTT AGGAGCaATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW99292 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC CCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW99301 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW99303 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCAAATA TTGCACATGG AGGATCCTCA GTAGATTTAG CAATTTTTTC TCTTCATCTA GCTGGAATTT CTTCAATTTT AGGAGCTATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAGTT TATCTTTTGA TCAAATATCT MW99309 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACACAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTCCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGG GTAAATAATT TATCTTTTGA TCAAATATCA MW99314 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGG GTAAATAATT TATCTTTTGA TCAAATATCT MW99319 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCAAATA TCGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATCAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW99320 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCAAATA TCGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATCAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW99341 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC CCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW99344 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACACAG AGGATCATCT GTAGATTTAG CAATTTTCTC ACTCCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGG GTAAATAATT TATCTTTTGA TCAAATATCA MW99353 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAACA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTTTC TCTTCATTTG GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT CAATATACGA GTAAATAACC TATCTTTTGA TCAAATATCA MW99357 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAACA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTTTC TCTTCATTTG GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT CAATATACGA GTAAATAACC TATCTTTTGA TCAAATATCA MW99372 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACACAG AGGATCATCT GTAGATTTAG CAATTTTCTC ACTCCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGG GTAAATAATT TATCTTTTGA TCAAATATCA MW99374 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGG GTAAATAATT TATCTTTTGA TCAAATATCT MW99376 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC CCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATGCGA GTAAATAATT TATCTTTTGA TCAAATATCA MW99380 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCAAATA TCGCACATAG AGGATCGTCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW99381 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATGG AGGATCATCT GTAGATTTAG CAATTTTCTC ACTTCATTTA GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT CAATATGCGA GTAAATAATT TATCTTTTGA TCAAATATCT MW99382 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATGG AGGATCATCT GTAGATTTAG CAATTTTCTC ACTTCATTTA GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT CAATATGCGA GTAAATAATT TATCTTTTGA TCAAATATCT MW99393 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC CCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATGCGA GTAAATAATT TATCTTTTGA TCAAATATCA MW99398 GGAACTGGAT GAACAGTTTA TCCCCCACTT TCATCTAAtA TTGCTCATAG AGGATCATCA GTTGATCTAG CTATTTTTTC TCTTCATTTA GCAGGtATCT CATCAATTTT AGGaGCCATT AATTTTATTA CAACTATTAT NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN MW99406 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC CCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATGCGA GTAAATAATT TATCTTTTGA TCAAATATCA MW99407 GGAACAGGAT GANCaGTTTa CCCCCCaCTT TCATCTAATA TTGCACATGG AGGNTCNTCT GTAGANTTAG CAATTTTCTC ACTTCANTTA GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATtAT CAATATACGA GTAAATAAtT TATCTTTTGA TCAAATATCC MW99408 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC CCTTCATTTA GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT CAATATACGG GTAAATAAAT TATCCTTTGA TCAAATATCA

177 Appendix 3: Sequence data Cytochrome Oxidase I (COI)

....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| 365 375 385 395 405 415 425 435 445 455 465 475 485 495 505 515 525 535 MW99413 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC CCTTCATTTG GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT CAATATACGA GTAAATAAAT TATCTTTTGA TCAAATATCA MW99422 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC CCTTCATTTG GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT CAATATACGA GTAAATAAAT TATCTTTTGA TCAAATATCA MW99425 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCAAATA TTGCTCATGG AGGATCTTCA GTTGATTTAG CAATTTTTTC ATTACACTTA GCTGGTATTT CCTCAATCTT AGGAGCTATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAATT TATCATTTGA TCAAATATCC MW99430 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACACAG AGGGTCATCT GTTGATTTAG CAATTTTTTC TCTTCATTTA GCTGGAATTT CTTCAATTTT AGGAGCTATT AATTTTATTA CAACTATTAT TAATATGCGA GTAAACAATA TATCTTTTGA TCAAATATCT MW99435 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCTCATAG AGGATCATCT GTAGATTTAG CAATTTTCTC ACTTCATTTA GCTGGAATTT CTTCAATTTT AGGAGCTATT AACTTTATTA CAACTATTAT TAATATACGG GTAAATAATT TATCTTTTGA TCAAATATCA MW99448 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAACA TTGCACATAG AGGATCATCT GTaGATTTAG CAATTTTTTC TCTTCATTTG GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT CAATATACGA GTAAATAACC TATCTTTTGA TCAAATATCA MW99449 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAACA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTTTC TCTTCATTTG GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT CAATATACGA GTAAATAACC TATCTTTTGA TCAAATATCA MW99465 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAACA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTTTC TCTTCATTTG GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT CAATATACGA GTAAATAACC TATCTTTTGA TCAAATATCA MW99469 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW99471 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCAAATA TCGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATCAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW99473 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC CCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATGCGA GTAAATAATT TATCTTTTGA TCAAATATCA MW99476 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGG GTAAATAATT TATCTTTTGA TCAAATATCT MW99479 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC GCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGG GTAAATAATT TATCTTTTGA TCAAATATCA MW99494 GGAACAGGAT GAGNAGTTTA CCCCCCACTT TCATCAAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATCtA GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATCAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW99501 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC CCTTCATTTA GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT CAATATACGG GTAAATAAAT TATCTTTTGA TCAAATATCA MW99508 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGG GTAAATAATT TATCTTTTGA TCAAATATCA MW99514 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCTCATAG AGGATCATCT GTAGATTTAG CAATTTTCTC ACTTCATTTA GCTGGGATTT CTTCAATTTT AGGAGCTATT AACTTTATTA CAACTATCAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW99515 GGAACAGGAT GAACAGTTTA TCCCCCACTT TCATCTAATA TGGCTCATAG AGGACCTTCT GTTGATTTAG CTATTTTTTC TTTACATTTA GCAGGTATTT CTTCAATTTT AGGTGCTATT AATTTTATTA CAACTATTAT CAATATACGA ATCAATAATT TATCTTTTGA TCAAATATCT MW99520 GGAACAGGAT GAACAGTTTA TCCCCCACTT TCATCTAATA TTGCCCATAG AGGATCTTCT GTTGATTTAG CTATTTTTTC TTTACATTTA GCAGGTATTT CTTCAATTTT AGGAGCCATT AATTTTATTA CAACTATTAT TAATATACGA ATTAATAACT TATCCTTTGA TCAAATATCC MW99526 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACACAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGG GTAAATAATT TATCTTTTGA TCAAATATCA MW99536 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCTCATAG AGGATCATCT GTAGATTTAG CAATTTTCTC ACTTCATTTA GCTGGCATTT CTTCAATTTT AGGAGCTATT AACTTTATTA CAACTATCAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCG MW99537 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TCGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC TTTACACCTG GCTGGAATTT CTTCAATTtT AGGAGCTATT AATTTTATTA CAACTATTAT TAACATACGA GTAAATAATT TATCATTTGA TCAAATATCA MW99538 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TCGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC TTTACACTTG GCTGGAATTT CTTCAATTTT AGGAGCTATT AATTTTATTA CAACTATTAT TAACATACGA GTAAATAATT TATCATTTGA TCAAATATCA MW99546 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAA CAATTTTCTC ACTTCATTTG GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAACATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW99550 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TCGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC TTTACACCTG GCTGGAATTT CTTCAATTTT AGGAGCTATT AATTTTATTA CAACTATTAT TAACATACGA GTAAATAATT TATCATTTGA TCAAATATCA MW99552 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGG GTAAATAATT TATCTTTTGA TCAAATATCA MW99559 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACACAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTCCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTaTTA CAACTATTAT TAATATACGG GTAAATAATT TATCTTTTGA TCAAATATCA MW99565 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACACAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTCCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGG GTAAATAATT TATCTTTTGA TCAAATATCA MW99579 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACACAG AGGATCTTCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCTGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAATT TATCCTTTGA TCAAATATCA MW99591 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCAAATA TTGCACATGG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATCAT TAATATACGA GTAAATAATT TATCTTTTGA ACAAATATCA MW99605 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCAAATA TTGCACATGG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATCAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW99608 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCTCATAG AGGATCATCT GTAGATTTAG CAATTTTCTC ACTTCATTTA GCTGGAATTT CTTCAATTTT AGGAGCTATT AACTTTATTA CAACTATTAT TAATATACGG GTAAATAATT TATCTTTTGA TCAAATATCA MW99612 GGAACAGGAT GAACAGTTTA TCCCCCACTT TCATCAAATA TTGCCCATAG AGGATCATCT GTAGATTTAG CAATTTTCTC ACTTCAATTA GCTGGAATTT CTTCAATTTT AGGAGCTATT AACTTTATTA CAACTATCAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA MW99613 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAA CAATTTTCTC ACTTCATTTA GCGGGAATTT CTTCAATTTT AGGGGCAATT AATTTTATTA CAACTATTAT TAACATACGA GTAAATAATT TATCTTTTGA TCAAATATCA OK96022 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCTCATAG AGGATCATCT GTAGATTTAG CAATTTTCTC ACTTCATTTA GCTGGAATTT CTTCAATTTT AGGAGCTATT AACTTTATTA CAACTATCAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA OK99001 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCTCATAG AGGATCATCT GTAGATTTAG CAATTTTCTC ACTTCATTTA GCTGGAATTT CTTCAATTTT AGGAGCTATT AACTTTATTA CAACTATCAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA WE00002 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCGCACAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAATT TATCCTTTGA TCAAATATCA WE00003 GGAACAGGAT GAACAGTCTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCACTTA GCTGGAATTT CTTCAATTTT AGGGGCTATT AATTTTATTA CAACTATCAT TAATATACGA GTAAGTAATT TATCATTTGA CCAAATATCA WE02321 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAACA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTTTC TCTTCATTTA GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT CAATATACGA GTAAATAACC TATCTTTTGA TCAAATATCA WE02421 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAACA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTTTC TCTTCATTTG GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT CAATATACGA GTAAATAACC TATCTTTTGA TCAAATATCA WE02431 GGAACAGGGT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATGG AGGATCATCT GTAGATTTAG CAATTTTCTC ACTTCATTTA GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT CAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCC WE02451 GGAACGGGAT GAACAGTTTA CCCCCCACTT TCATCCAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGG GTAAATAAAT TATCCTTTGA TCAAATATCA WE02491 GGAACGGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACACAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGG GTAAATAACT TATCCTTTGA tCAAATATCA WE02531 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC GCTTCATTTA GCCGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA WE02535 GGAACAGGGT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATGG AGGATCATCT GTAGATTTAG CAATTTTCTC ACTTCATTTA GCGGGAATTT CTTCAATTTT AGGGGCAATT AATTTTATTA CAACTATTAT CAATATACGA GTAAATAATT TGTCTTTTGA TCAAATATCC WE02536 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAACA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTG GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT CAATATACGA GTAAATAACC TATCTTTTGA TCAAATATCA WE02591 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC CCTTCATTTA GCGGGAATTT CTTCAATTTT AGGGGCAATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA WE02612 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC CCTTCATTTA GCGGGAATTT CTTCAATTTT AGGGGCAATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA WE02614 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC CCTTCATTTA GCGGGAATTT CTTCAATTTT AGGGGCAATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA WE02621 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAACA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTNTC TCTTCATTTG GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATtA CAACTATTAT CAATATACGA GTAAATAACC TATCTTTTGA TCAAATATCA WE02661 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC GCTTCATTTA GCCGGAATTT CTTCAATTTT AGGGGCAATT AATTTTATTA CAACTATTAT TAATATACGA GTAAATAATT TATCTTTTGA TCAAATATCA WE02662 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACACAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTCCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATACGG GTAAATAATT TATCTTTTGA TCAAATATCA WE02671 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC CCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAATATGCGA GTAAATAATT TATCTTTTGA TCAAATATCA WE02674 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAACA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTTTC TCTTCATTTG GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT CAATATACGA GTAAATAACC TATCTTTTGA TCAAATATCA WE02676 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CTACTATTAT TAATATACGG GTAAATAATT TATCTTTTGA TCAAATATCA WE02677 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATGG AGGATCATCT GTAGATTTAG CAATTTTCTC ACTTCATTTA GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT CAATATGCGA GTAAATAATT TATCTTTTGA TCAAATATCT WE85001 GGAACGGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACACAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCAGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTC CAACTATTAT TAATATACGG GTAAATAACT TATCCTTTGA TCAAATATCA WE98001 GGAACAGGAT GAACAGTTTA CCCCCCACTT TCATCTAATA TTGCACATAG AGGATCATCT GTAGATTTAG CAATTTTCTC TCTTCATTTA GCGGGAATTT CTTCAATTTT AGGAGCAATT AATTTTATTA CAACTATTAT TAACATACGA GTAAATAATT TATCATTTGA TCAAATATCA

....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| 545 555 565 575 585 595 605 615 625 635 645 655 665 675 685 AD98001 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACTATATT ATTAACCGAT CGAAATCTTA ACACCTCATT TTTTGATCCA GCTGGTGGAG GAGATCCAAT TTTATACCAA AD98009 CTATTTATTT GAGCTGTGGG AATTACAGCA TTATTATTAC TTCTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACTGAT CGAAACCTTA ATACCTCATT CTTTGACCCA GCTGGGGGAG GAGATCCAAT TTTATACCAA AD98012 CTATTTATTT GAGCTGTAGG AATTACAGCA TTATTATTAC TTCTATCTTT ACCTGTATTA GCTGGAGCTA TTACCATATT ATTAACTGAT CGAAACCTTA ATACCTCATT CTTTGACCCA GCTGGGGGAG GAGATCCAAT TTTATACCAA AD98018 TTATTTATTT GAGCAGTGGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACTGAT CGAAATCTTA ATACCTCATT CTTTGATCCA GCTGGTGGAG GAGATCCAAT TTTANNNNNN AD98024 TTATTTATTT GAGCAGTGGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACTGAT CGAAATCTTA ACACCTCATT CTTTGATCCA GCTGGTGGAG GAGATCNNNN NNNNNNNNNN AD98029 TTATTTATTT GAGCAGTGGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACCGAT CGAAACCTTA ATACCTCATT CTTTGATCCA GCTGGTGGaG GNNNNNNNNN NNNNNNNNNN AD98036 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACTGAT CGAAACCTTa aTACCTCTTT CTTTGACCCN GCTGGTGGGG GNNNNNNNNN NNNNNNNNNN

178 Appendix 3: Sequence data Cytochrome Oxidase I (COI)

....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| 545 555 565 575 585 595 605 615 625 635 645 655 665 675 685 DS00001 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCTA TTACCATATT ATTAACTGAT CGAAACCTTA ATACCTCATT CTTTGACCCA GCTGGTGGAG GAGATCCAAT TTTATATCAA DS00005 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCTA TTACCATGTT ATTAACTGAT CGAAACCTTA ATACCTCATT CTTTGATCCA GCTGGTGGAG GCGATCCAAT TTTNNNNNNN DS01001 TTATTTATTT GAGCAGTGGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACAGAT CGAAACCTTA ATACCTCATT CTTTGATCCA GCTGGTGGGG GAGATCCATT TTNATATCAA JC00029 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCaA TTACTATATT ATTAACTGAT CGAAACCTCA ATACCTCATT CTTTGATCCA GCTGGAGGGG GAGATCCAAT TTTATATCAA JC00039 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCTA TCACTATATT ATTAACTGAT CGAAATCTTA ATACCTCATT CTTTGATCCT GCAGGAGGAG GAGATCCAAT TTTATATCAA JC00040 TTATTTATTT GAGCAGTGGG AATTACAGCA TTATTATTAC TTTTATCATT GCCTGTATTA GCTGGAGCAA TTACCATATT ATTAACAGAT CGAAATCTTA ATACCTCATT CTTTGATCCA GCTGGTGGGG GAGATCCAAT TTTATATCAA JC00042 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACTATATT ATTAACTGAT CGAAACCTCA ATACCTCATT CTTTGATCCA GCTGGAGGGG GAGATCCAAT TTTATATCAA JC00043 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACTGAT CGAAATCTTA ACACCTCATT CTTTGACCCG GCTGGTGGAG GAGATCCAAT TTTATATcNN JC00045 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACTGAT CGAAATCTTA ACACCTCATT CTTTGACCCG GCTGGTGGAG GAGATCCAAT TTTNNNNNNN JC00046 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACTGAT CGAAATCTTA ACACCTCATT CTTTGACCCG GCTGGTGGAG GAGATCCAAT TTTATTTCNN JC00051 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACTATATT ATTAACTGAT CGAAACCTCA ATACCTCATT CTTTGATCCA GCTGGAGGGG GAGATCCAAT TTTATATCAA JC00055 TTATTTATTT GAGCAGTAGG TATTACCGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACTATATT ATTAACTGAT CGAAATATTA ACACCTCATT TTTTGATCCT GCTGGAGGAG GAGATCCNNN NNNNNNNNNN JC00057 TTATTTATTT GAGCAGTAGG TATTACTGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACTATATT ATTAACTGAT CGAAATATTA aTACCTCATT TTTTGATCCT GCTGGaGGAG GAGATCCTAT TTTNNNNNNN JC00061 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACTATATT ATTAACTGAT CGAAACCTCA ATACCTCATT CTTTGATCCA GCTGGAGGAG GAGATCCAAT TTTATATCAA JC00062 TTATTTATTT GAGCAGTAGG TATTACTGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCaA TTACTATaTT ATTAACTGAT CGAAATATTA ATNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN JC00063 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACTATATT ATTAACGGAT CGAAACCTCA ATACCTCATT CTTTGATCCA GCTGGAGGGG GAGATCCAAT TTTATATCAA JC01014 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACTGAT CGAAATCTTA ACACCTCATT CTTTGACCCG GCTGGTGGAG GAGATCCAAT TTTATATCAA JC02001 TTATTTATTT GAGCTGTTGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCAGTATTA GCTGGAGCTA TTACCATATT ACTTACCGAT CGAAATTTAA ATACTTCATT TTTTGATCCA GCAGGAGGAG GAGATCCTAT TTTATATCAA JM00001 TTATTTATTT GAGCAGTGGG AATTACAGCA TTATTATTAC TTTTATCATT ACCTGTATTA GCTGGAGCaA TTACCATATT ATTAACAGAT CGAAATCTTA aTACCTCATT TTTTGACCCN GCTGGCGGAG GAGATCCAAT TTTANNNNNN MW00001 TTATTTATTT GAGCAGTAGG AATTACGGCA TTATTATTAC TTTTATCCTT ACCTGTATTA GCTGGAGCaA TTACCATATT ATTAACCGAT CGAAACCTTA ACACCTCATT CTTTGATCCG GCTGGTGGGG GAGATCCAAT TTTATACCAN MW00015 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCCGTATTA GCTGGAGCAA TTACCATATT ATTAACCGAT CGAAATCTTA ACACCTCATT CTTTGACCCA GCTGGTGGGG GAGATCCAAT TTTATATCAA MW00017 TTATTTATTT GAGCAGTAGG AATTACAGCT TTATTATTAT TATTATCTTT ACCTGTTTTA GCTGGAGCTA TTACTATATT ATTAACTGAT CGTAATCTAA ATACATCTTT TTTTNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN MW00020 TTATTTATTT GAGCAGTAGG TATTACTGCA TTATNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN MW00024 TTATTTATTT GGGCGGTAGG TATTACTGCT CTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACTGAT CGAAATCTTA ACACTTCATT TTTTGATCCT GCCGGAGGAG GAGATCCAAT TTTATATCAA MW00032 TTATTTATTT GAGCAGTAGG AATTACGGCA TTATTATTAC TTTTATCCTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACCGAT CGAAACCTTA ACACCTCATT CTTTGATCCG GCTGGTGGGG GAGATCCAAT TTTATACCAA MW00044 TTATTTATTT GAGCTGTAGG AATTACAGCT TTATTATTAT TATTATCTTT ACCTGTTTTA GCTGGAGCTA TTACTATATT ATTAACTGAT CGTAATTTAA ATACTTCATT TTTTNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN MW00051 TTATTTATTT GAGCAGTAGG AATTACGGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACCGAT CGAAATCTTA ACACCTCATT CTTTGATCCA GCTGGTGGAG GAGATCCAAT TTTATACCAN MW00056 TTATTTATTT GAGCAGTAGG AATTACGGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACCGAT CGAAATCTTA ACACCTCATT CTTTGATCCA GCTGGTGGAG GAGATCCAAT TTTATACCNN MW00058 TTATTTATTT GAGCAGTAGG AATTACGGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACCGAT CGAAATCTTA ACACCTCATT CTTTGATCCA GCTGGTGGAG GAGATCCAAT TTTATACCAA MW00059 TTATTTATTT GAGCAGTAGG AATTACGGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACCGAT CGAAATCTTA ACACCTCATT CTTTGATCCA GCTGGTGGAG GAGATCCAAT TTTATACCAA MW00060 TTATTTATTT GAGCGGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACTGAT CGAAACCTTA ATACCTCATT CTTTGATCCA GCTGGTGGGG GAGAtCCTAT TTTATATCAA MW00064 TTATTTATtT GGGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCATT ACCTGTATTA GCTGGGGcAA TTACCATATT ATTAACAGAT CGAAATCTTA ATACCTCATT CTTTGACCCG GCTGGCGGAG GAGATCCAAT TTTATNNNNN MW00072 TTATTTATTT GAGCGGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACTGAT CGAAACCTTA ATACCTCATT CTTTGATCCA GCTGGTGGGG GAGATCCTAT TTTATATCAA MW00076 TTATTTGTTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACTATATT ATTAACCGAT CGAAACCTAA ATACCTCATT CTTTGATCCT GCTGGTGGAG GAGATCCCAT TTTATATCAA MW00087 TTATTTGTTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACTATATT ATTAACCGAT CGAAACCTAA ATACCTCATT CTTTGATCCT GCTGGTGGAG GAGATCCAAT TTTATATCAA MW00101 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCAGTATTA GCTGGAGCAA TTACCATATT ATTAACTGAT CGGAACCTTA ATACCTCATT CTTTGACCCG GCTGGTGGAG GAGATCCTAT TTTATATCAA MW00102 TTATTTATTT GAGCGGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTaCCATATT ATTAACTGAT CGAAACCTTA ATACCTCATT CTTTGATCCA GCTGGTGGGG GAGATCCTAT TTTATATCAA MW00110 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACCGAT CGAAATCTTA ACACCTCATT CTTTGATCCA GCTGGTGGAG GAGATCCAAT TTTATACCAA MW00116 TTATTTATTT GAGCGGTCGG AATTACTGCA TTATTATTAC TTTTATCCCT ACCTGTATTA GCTGGAGCAA TTACTATGTT ATTAACAGAT CGGAATCTTA ATANNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN MW00119 TTATTTATTT GAGCTGTAGG AATTACAGCT TTACTATTAT TATTATCTTT ACCTGTTTTA GCTGGAGCTA TTACTATATT ATTAACTGAT CGTAATTTAA ATACTTCTTT TTTTGATCCT GCAGGAGGTG GAGACCCAAT TTTATATCAA MW00125 CTATTTATTT GAGCTGTGGG AATTACAGCA TTATTATTAC TTCTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACTGAT CGAAACCTTA ATACCTCATT CTTTGACCCA GCTGGGGGAG GAGATCCTAT TTTANNNNNN MW00127 TTATTTATTT GAGCAgTAGG AATTAcGGCA TTAtTATTAC TTTTATCTTT AcCTGTaTTA GCTGGAGCAA TTACCATATT ATTAACCGAT CGAAATCTTA ACACCTCATT NTTTGATCCA GCTGGTGGAG GNNNNNNNNN NNNNNNNNNN MW00129 CTATTTATTT GGGCTGTAGG AATTACAGCA TTATTATTAC TTCTATCTTT ACCTGTATTA GCTGGaGCTA TNACCATATT ATTAACCGAT cGAAACCTTA aNACNTCATT CTTTGACCCA GCTGGAGGAG GAGATCCAAT TTTATNNNNN MW00140 TTATTTATTT GAGCGGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACTGAT CGAAACCTTA ATACCTCATT CTTTGACCCA GCTGGTGGGG GAGATCCTAT TTTATATCAA MW00151 CTATTTATTT GAGCTGTGGG AATTACAGCA TTATTATTAC TTCTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACTGAT CGAAACCTTA ATACCTCATT CTTTGACCCA GCTGGGGGAG GAGATCCAAT TTTATACCAA MW00157 TTATTTATTT GAGCAGTAGG AATTACGGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACCGAT CGAAATCTTA ACACCTCATT CTTTGATCCA GCTGGTGGAG GAGATCCAAT TTTATACCAA MW00176 TTATTTATTT GAGCAGTGGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACCGAT CGAAACCTTA ATACCTCATT CTTTGACCCG GCTGGTGGAG GAGATCCAAT TTTATATCAA MW00177 CTATTTATTT GGGCTGTAGG AATTACAGCA TTATTATTAC TTCTATCTTT ACCTGTATTA GCTGGAGCTA TTACCATATT ATTAACCGAT CGAAACCTTA ATACCTCATT CTTTGACCCA GCTGGAGGAG GAGATCCAAT TTTATACCAA MW00178 CTATTTATTT GGGCTGTAGG AATTACAGCA TTATTATTAC TTCTATCTTT ACCTGTATTA GCTGGAGCTA TTACCATATT ATTAACTGAT CGAAACCTTA ATACCTCATT CTTTGACCCA GCTGGAGGAG GAGATCCAAT TTTATACCAA MW00179 CTATTTATTT GGGCTGTAGG AATTACAGCA TTATTATTAC TTCTATCTTT ACCTGTATTA GCTGGAGCTA TTACCATATT ATTAACTGAT CGAAACCTTA ATACCTCATT CTTTGACCCA GCTGGAGGAG GAGATCCAAT TTTATACCAA MW00183 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCCGTATTA GCTGGAGCAA TTACCATATT ATTAACCGAT CGAAATCTTA ACACCTCATT CTTTGACCCA GCTGGTGGGG GAGATCCAAT TTTATATCAA MW00185 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCCGTATTA GCTGGAGCAA TTACCATATT ATTAACCGAT CGAAATCTTA ACACCTCATT CTTTGACCCA GCTGGTGGGG GAGATCCAAT TTTATATCAA MW00186 TTATTTATTT GAGCAGTAGG AATTACAGCA TtATTATtAC TTTTATCTTT ACCCGTATTA GCTGGAGCAA TTACCATATT ATTAACNGAT CGAAATCTTA ACACCTCATT CTTTGACCCA GCTGGTGGGG GAGATCCAAT TTTATATCAA MW00189 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACCGAT CGAAACCTTA ACACCTCGTT CTTTGATCCG GCTGGTGGAG GAGATCCAAT TTTATATCAA MW00226 TTATTTATTT GAGCAGTAGG AATTACGGCA TTATTATTAC TTTTATCTTT ACCTGTaTTA GCTGGAGCAA TTACCATATT ATTAACTGAT CGAAATCTTA ACACCTCATT CTTTGATCCG GCTGGTGGGG GAGATCCAAT TTTATANCAA MW00229 TTATTTATTT GAGCAGTGGG AATTACAGCA TTATTATTAC TTTTATCATT GCCTGTATTA GCTGGAGCAA TTACCATATT ATTAACAGAT CGAAATCTTA ACACCTCATT CTTTGACCCA GCAGGTGGGG GAGATCCAAT TTTATATCAA MW00231 TTATTTATTT GAGCAGTGGG AATTACAGCA TTATTATTAC TTTTATCATT GCCTGTATTA GCTGGAGCAA TTACCATATT ATTAACAGAT CGAAATCTTA ACACCTCATT CTTTGACCCA GCAGGTGGGG GAGATCCAAT TTTATATCAA MW00232 TTATTTATTT GAGCAGTAGG AATTACGGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACCGAT CGAAATCTTA ACACCTCATT CTTTGATCCA GCTGGTGGAG GAGATCCAAT TTTATACCAA MW00234 CTATTTATTT GAGCTGTGGG AATTACAGCA TTATTATTAC TTCTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACTGAT CGAAACCTTA ATACCTCATT CTTTGACCCA GCTGGGGGAG GAGATCCAAT TTTATACCNN MW00259 TTATTTATTT GAGCAGTAGG AATTACAGCA CTATTATTAC TTTTATCTTT ACCTGTATTA GCAGGAGCTA TTACCATATT ATTAACTGAT CGAAACCTTA ATACCTCGTT CTTTGACCCA NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN MW00262 TTATTTATTT GAGCAGTAGG AATTACAGCA CTATTATTAC TTTTATCTTT ACCTGTATTA GCAGGAGCTA TTACCATATT ATTAACTGAT CGAAACCTTA ATACCTCGTT CTTTGACCCA GCTGGCGGGG GGGATCCTAT NNNNNNNNNN MW00267 TTATTTATTT GAGCGGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACTGAT CGAAACCTTA ATACCTCATT CTTTGATCCA GCTGGTGGGG GAGANNNNNN NNNNNNNNNN MW00269 TTATTTATTT GAGCGGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACCGAT CGAAATCTTA ATACCTCATT CTTTGATCCG GCTGGTGGAG GAGATCCAAT TTTATATCAA MW00290 TTATTTGTTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCAGGAGCAA TTACTATACT ATTAACTGAT CGAAACCTAA ATACCNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN MW00302 TTATTTATTT GAGCGGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCTA TTACTATATT ATTaACTGAT CGAAATCTAA ACACCTCATT CTTTGATCCT GCTGGAGGAG GNNNNNNNNN NNNNNNNNNN MW00316 TTATTTATTT GAGCAGTAGG AATTACGGCA TTATTATtAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACCGAT CGAAATCTTA ACACCTCATT CTTTGATCCA GCTGGTGGAG GAGATCCAAT TTTATACCAA MW00326 TTATTTATTT GAGCGGTGGG AATTACAGCA TTATTATTAC TCTTATCTTT ACCTGTATTA GCTGGAGCTA TTACTATATT ATTAACTGAT CGAAATTTAA ACACCTCATT TTTTGACCCT GCTGGAGGAG GAGATCCTAT TTTATATCAA MW00327 TTATTTATTT GAGCAGTaGG AATTACaGCA TTATTGTTAC TTCTGTCTTT ACCtGTATTA GCTGGAGCAA TTACTATATT ATTAACTGAT CGAAACCTTA ATACCTCATT CTTTGATCCC GCTGGTGGAG GAGATCCTAT TTTATATCAA

179 Appendix 3: Sequence data Cytochrome Oxidase I (COI)

....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| 545 555 565 575 585 595 605 615 625 635 645 655 665 675 685 MW00328 TTATTTATTT GAGCGGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACCGAT CGAAATCTTA ATACCTCATT CTTTGATCCG GCTGGTGGAG GAGATCCAAT TTTATATCAA MW00330 CTATTTATTT GGGCTGTAGG AATTACAGCA TTATTATTAC TTCtATCTTT ACCTGTATTA GCTGGaGCTA TTACCATATT ATTAACTGAT CGAAACCTTA ATACCTCATT CTTTGACCCA GCTGGAGGaG GAGATCCAAT TTTATACCAA MW00335 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTGTTA GCTGGAGCAA TCACCATATT ATTAACCGAT CGAAATCTTA ATACCTCATT CTTTGATCCG GCTGGTGGAG GAGATCCTAT TTTATATCAA MW00347 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACTGAT CGAAACCTTA ATACCTCATT CTTTGACCCA GCTGGTGGGG GAGATCCTAT TTTATATCAA MW00348 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACTGAT CGAAACCTTA ATACCTCATT CTTTGACCCA GCTGGTGGGG GAGATCCTAT TCTATACCAa MW00358 TTATTTATTT GAGCAGTAGG AATTACAgCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACTGAT CGAAaCCTTA ATACCTCATT CTTTGACCCA GCTGGTGGGG GAGATCCTAT TTTATANNNN MW00393 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTGTTAC TTCTGTCTTT ACCTGTATTA GCTGGAGCAA TTACTATATT ATTAACTGAT CGAAACCTTA ATACCTCATT CTTTGATCCC GCTGGTGNNN NNNNNNNNNN NNNNNNNNNN MW00409 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTGTTA GCTGGAGCAA TCACCATATT ATTAACCGAT CGAAATCTTA ATACCTCATT CTTTGACCCG GCTGGCGGAG GAGATCCTAT TTTATATCAA MW00412 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTaCTATATT ATTAACGGAT CGAAACCTCA ATACCTCATT CTTTGATCCA GCTGGAGGAG GAGATCCAAT TTTATATCAA MW00424 TTATTTATTT GAGCGGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACCGAT CGAAATCTTA ATACCTCATT CTTTGATCCG GCTGGTGGAG GAGATCCAAT TNNNNNNNNN MW00444 TTATTTATTT GAGCGGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACTGAT CGAAACCTTA ATACCTCATT CTTTGACCCA GCTGGTGGGG GAGATCCTAT TTNNNNNNNN MW00452 TTATTTATTT GAGCGGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCCGTATTA GCTGGAGCAA TTaCCATATT ATTAACTGAt CGAAACCTTA ATACCTCATT CTTTGACCCG GCTGGTGGGG GAGATCCTAT TTNATANCAA MW00469 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAT TATTATCTTT ACCTGTATTA GCTGGAGCTA TTACTATATT ATTAACTGAT CGAAATCTCA ACACTTCATT TTTTGATCCT GCAGGAGGAG GAGATCCAAT CTTATATCAA MW00476 TTATTTATTT GAGCGGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACTGAT CGAAACCTTA ATACCTCATT CTTTGATCCA GCTGGTGGGG GAGATCCTAT TTTATATCAA MW00497 TTATTTATTT GAGCTGTAGG AATTACAGCG TTATTATTAT TATTATCATT ACCAGTATTA GCTGGAGCTA TTACTATATT ATTAACAGAC CGAAATTTAA ATACtTCATT TTTTGACCCA GCAGGTGGAG GAGATCCTAT TCTTTACCAA MW00498 TTATTTATTT GGGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCATT ACCTGTATTA GCTGGGGCAA TTACCATATT ATTAACAGAT CGAAATCTTA ATACCTCATT CTTTGACCCG GCTGGCGGAG GAGATCCAAT TTTATATCAA MW00504 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACTATATT ATTAACTGAT CGAAACCTTA ATACCTCATT TTTTGATCCA GCTGGCGGGG GAGATCCNAT TTTATATCAA MW00517 TTATTTATTT GAGCAGTAGG TATTACAGCT TTACTATTAC TTTTATCATT ACCTGTTTTA GCTGGAGCAA TTACTATATT ATTAACTGAC CGAAATCTTA ATACTTCATT TTTCGATCCT GCTGGAGGAG GAGATCCAAT TTTATATCAN MW00525 TTATTTATTT GAGCAGTAGG AATCACCGCT CTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGGGCAA TTACTATATT ATTAACTGAT CGAAATCTTA ATACTTCATT TTTTGATCCG GCTGGAGGAG GAGATCCAAT CTTANNNNNN MW00530 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTCTATCTTT ACCTGTATTA GCTGGAGCAA TTACTATATT ATTAACTGAT CGAAACCTCA ATACCTCATT CTTTGATCCA GCTGGAGGAG GAGATCCAAT TTTANNNNNN MW00539 TTATTTATTT GAGCGGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCtGGaGCaA TTACCATATT ATTAACCGAT CGAAATCTTA ATACCTCATT CTTTGATCCG GCTGGTGGAG GAGATCCAAT TTTATACCAA MW00547 TTATTTATTT GAGCGGTAGG AATTACNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN MW01001 TTATTTATTT GAGCAGTGGG AATTACAGCA TTATTATTAC TTTTATCATT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACAGAT CGAAATCTTA ATACCTCATT TTTTGACCCC GCTGGCGGAG GAGATCCAAT TTTATATCAA MW01009 TTATTTGTAT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCTA TTACTATATT ATTAACTGAT CGAAACCTTA ATACCTCATT CTTTGATCCT GCAGGAGGAG GAGATCCAAT TTTATATCAC MW01011 TTATTCATTT GAGCAGTAGG AATTACAGCA TTATTATTAT TATTATCTTT ACCTGTATTA GCTGGAGCTA TTACTATATT ATTAACTGAT CGAAATCTCA ACACTTCATT TTTTGATCCT GCAGGAGGAG GAGATCCAAT TTTATATCAA MW01014 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACTGAT CGAAATCTTA ACACCTCATT CTTTGATCCG GCTGGTGGAG GAGATCCAAT TTTATATCAA MW01018 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAT TATTATCTTT ACCTGTATTA GCTGGAGCTA TTACTATATT ATTAACTGAT CGAAATCTTA ACACTTCATT TTTTGATCCT GCTGGAGGAG GAGATCCAAT TCTATACCAA MW01019 TTATTTATTT GAGCGGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACTATATT ATTAACTGAT CGAAACCTTA ATACCTCATT TTTTGATCCT GCTGGAGGAG GAGATCCAAT TTTATATCAA MW01022 TTATTTATTT GAGCTGTTGG TATTACAGCA TTATTATTAT TATTATCTTT ACCTGTTTTA GCGGGAGCTA TTACTATATT ATTAACTGAT CGAAATCTTA ATACTTCTTT TTTTGATCCT GCTGGAGGAG GAGATCCAAT CTTATATCAA MW01023 TTATTTATTT GAGCCGTTGG AATTaCAGCA TTATTACTAT TATTATCTTT ACCTGTATTA GCAGGAGCTA TTACTATATT ATTAACAGAT CGAAATTTAA ATACTTCATT TTTTGACCCT GCCGGAGGAG GAGATCCAAT TTTATATCAA MW01025 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCAGGAGCAA TTACTATATT ATTAACTGAT CGAAACCTCA ATACCTCATT CTTTGATCCA GCTGGAGGAG GAGATCCAAT TTTATATCAA MW01027 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCAGTATTA GCAGGTGCTA TTACTATATT ACTAACTGAT CGAAATTTAA ATACTTCATT TTTTGACCCT GCAGGAGGAG GAGATCCAAT TTTATATCAA MW01034 TTATTTATTT GAGCGGTAGG AATTACAGCA TTATTATTAT TATTATCTTT ACCTGTATTA GCTGGAGCTA TTACTATATT ATTAACTGAT CGAAATCTTA ACACTTCATT NTTTGATCCT GCTGGAGGAG GAGATCCAAN NNNNNNNNNN MW01039 TTATTTATTT GAGCAGTGGG AATTACAGCA TTATTATTAC TTTTATCATT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACAGAT CGAAATCTTA ATACCTCATT TTTTGACCCC GCTGGCGGAG GAGATCCAAT TTTATATCAA MW01048 TTATTTATTT GAGCAGTAGG TATTACCGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGGGCAA TTACTATATT ATTAACTGAT CGAAATATTA ATACCTCATT TTTtGATCCT GcTGGAGGAG GAGATCCNNN NNNNNNNNNN MW01053 TTATTTATTT GAGCAGTGGG AATTACAGCA TTATTATTAC TTTTATCATT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACAGAT CGAAATCTTA ATACCTCATT TTTTGACCCC GCTGGAGGAG GAGATCCAAT TTTATATCAA MW01059 TTATTTATTT GAGCGGTAGG AATTACAGCA TTATTATTAT TATTATCTTT ACCTGTATTA GCTGGAGCTA TTACTATATT ATTAACTGAT CGAAATCTTA ACACTTCATT TTTTGATCCT GCNGGNGGNG GAGANNNNNN NNNNNNNNNN MW01061 TTATTTATTT GAGCTGTAGG TATTACTGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACTATATT ATTAACTGAT CGAAATATTA ATACTTCATT TTTTGATCCT GCTGGAGGAG GAGANCCAAT TTNATATCAA MW01072 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACTGAT CGAAATCTTA ACACCTCATT CTTTGATCCG GCTGGTGGAG GAGATCCAAT TTTATATCAA MW01078 TTATTTATTT GAGCAGTGGG AATTACAGCT TTATTATTAC TTTTATCATT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACAGAT CGAAATCTTA ATACCTCATT TTTTGACCCC GCTGGAGGAG GAGATCCAAT TTTATATCAA MW01083 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCTA TTACTATATT ATTAACTGAT CGAAATCTAA ACACCTCATT CTTTGATCCT GCTGGAGGAG GAGATCCAAT TTTATATCAA MW01092 TTATTTATTT GAGCGGTAGG AATTACAGCA TTATTATTAT TATTATCTTT ACCTGTATTA GCTGGAGCTA TTACTATATT ATTAACTGAT CGAAATCTTA ACACTTCATT TTTTGATCCT GCTGGAGGAG GAGATCCAAT TNTATATCAN MW01103 TTATTTATTT GAGCGGTAGG AATTACAGCA TTATTATTAT TATTATCTTT ACCTGTATTA GCTGGAGCTA TTACTATATT ATTAACTGAT CGAAATCTTA ACACTTCATT TTTTGATCCT GCTGGAGGAG GAGATCCAAT TTTATATCAA MW01105 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACTATATT ATTAACTGAT CGAAATCTTA ACACCTCATT CTTTGACCCG GCTGGNGGAG GAGATCCAAT TTTATATCAA MW01107 TTATTTATTT GAGCAGTGGG AATTACAGCA TTATTATTAC TTTTATCATT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACAGAT CGAAATCTTA ATACCTCATT TTTTGACCCC GCTGGAGGAG GAGATCCAAT TTTATATCAA MW01114 TTATTTATTT GAGCTGTTGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCAGTATTA GCTGGAGCTA TTACCATATT ACTTACTGAT CGAAATTTAA ATACTTCATT TTTTGATCCA GCAGGAGGAG GAGATCCTAT TTTATATCAA MW01116 TTATTTATTT GAGCGGTAGG AATTACAGCA TTGTTATTAT TATTATCTTT ACCTGTATTA GCTGGAGCTA TTACTATATT ATTAACTGAT CGAAATCTTA ACACTTCATT TTTTGATCCT GCTGGAGGAG GAGATCCAAT TTTATATCAA MW01120 TTATTTATTT GAGCTGTTGG TATCACAGCA TTATTACTAC TTTTATCTCT TCCAGTTTTA GCTGGGGCTA TTACTATACT ACTAACCGAT CGAAATTTAA ACACCTCCTT TTTTGACCCT GCAGGGGGAG GTGATCCAAT TTTATATCAA MW02001 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACTATATT ACTAACTGAT CGAAACCTTA ATACCTCATT TTTTGATCCA GCTGGTGGGG GGGATCCAAT TTTATATCNN MW02006 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTCTATCTTT ACCGGTATTG GCTGGAGCAA TTACTATATT ACTAACAGAT CGAAACCTTA ATACATCATT CTTTGATCCT GCTGGAGGAG GAGATCCAAT TTTATACCAA MW02007 TTATTTATCT GAGCAGTAGG AATTACAGCT CTATTATTAC TACTATCTTT ACCTGTTTTA GCTGGAGCTA TTACTATATT ATTAACTGAT CGTAATCTAA ATACATCTTT TTTTGATCCT GCAGGAGGAG GTGACCCAAT TTTATATCAA MW02008 TTATTTATTT GAGCAGTAGG AATTACAGCA CTATTACTTC TTCTTTCTTT ACCTGTTTTA GCTGGTGCTA TTACTATATT ATTAACTGAT CGAAATCTTA ATACTTCATT TTTTGACCCT GCTGGAGGAG GAGACCCAAT TTTATATCAN MW02009 TTATTTATTT GAGCTGTAGG AATTACAGCT TTATTATTAT TATTATCTTT ACCTGTTTTA GCTGGAGCTA TTACAATATT ATTAACTGAC CGTAATTTAA ATACATCNTT TTTTGACCCT GCAGGAGGAN NNNNNNNNNN NNNNNNNNNN MW02021 CTATTTATTT GAGCTGTTGG TATTACAGCT TTATTACTTC TTTTATCTTT ACCTGTTTTA GCAGGGGCTA TTACTATATT ATTAACAGAT CGAAATTTAA ATACCTCATT TTTTGATCCA GCTGGAGGAG GTGATCCAAT TTTATATCAA MW02024 TTATTTATTT GAGCTGTTGG AATTACAGCA TTATTATTAT TATTATCCTT ACCAGTATTA GCTGGAGCTA TTACCATATT ACTTACGGAT CGAAATTTAA ATACTTCATT TTTTGATCCA GCAGGAGGAG GAGATCCTAT TTTATATCAA MW02025 TTATTTATCT GAGCTGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCAGTATTA GCTGGTGCTA tTACCATATT ATTAACAGAT CGAAATCTTA ATACATCATT TTTTGATCCA GCAGGAGGAG GAGATCCAAT TTTATATCAA MW02028 TTATTTATTT GAGCTGTTGG TATTACAGCA TTATTATTAT TATTATCTTT ACCTGTTTTA GCGGGAGCTA TTACTATATT ATTAACTGAT CGAAATCTTA ATACTTCTTT TTTTGATCCT GCTGGAGGAG GAGATCCAAT CTTATATCAA MW02031 TTATTTATTT GAGCAGTAGG TATTACAGCA TTATTATTAT TATTATCTTT ACCTGTTTTA GCGGGTGCTA TTACTATATT ACTAACAGAT CGAAATCTTA ATACTTCTTT NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN MW02033 TTATTTATTT GAGCAGTAGG TATTACCGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACTATATT ATTAACTGAT CGAAATATTA ATACCTCATT TTTTGATCCT GCTGGAGGAG GAGATCCAAT TTTATATCAA MW02034 TTATTTATTT GAGCAGTAGG AATCACCGCT CTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGGGCAA TTACTATATT ATTAACCGAT CGAAATCTTA ATACTTCATT TTTTGATCCG GCTGGAGGAG GAGATCCAAT CTTATATCAA MW98002 TTATTTATTT GAGCTGTAGG AATTaCAGCA TTATTATTAT TGTTATCATT aCCAGTTTTA GCTGGAGCTA TTACAATATT ATTAACTGAT CGAAATTTAA ATACTTCATT TTTTGATCCA GCTGGAGGAG GAGATCCAAT TTTATATCAA MW98008 TTATTTATTT GAGCGGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTAtTA GCTGGNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN MW98009 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACTATATT ATTAACCGAT CGAAATCTTA ACACCTCATT TTTTGATCCA GCTGGTGGAG GAGATCCAAT TTTATACCAA MW98020 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCATT GCCTGTATTA GCTGGAGCAA TTACTATATT ATTAACAGAT CGAAATCTTA ATACCTCATT CTTTGATCCG GCTGGTGGGG GAGATCCAAT NNNNNNNNNN MW98029 TTATTTGTTT GAGCGGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACTATATT ATTAACCGAT CGAAACCTAA ATACCTCATT CTTCGATCCT GCTGGTGGAG GAGATCCAAT TNNNNNNNNN MW98049 TTATTTATTT GAGCAGTGGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACCGAT CGAAACCTTA ATACCTCATT CTTTGATCCA GCTGGTGGAG GAGATCCAAT TTTATACCAA MW98079 CTATTTATTT GAGCTGTAGG AATTACTGCA TTATTATTAC TTCTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACTGAT CGAAACCTTA ATACTTCCTT TTTTGACCCA GCTGGGGGAG GNNNNNNNNN NNNNNNNNNN MW98084 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACCGAT CGAAATCTTA aCACCTCNTT CTTTGACCCG GCTGGTGGAG GaGATCCAAT TTTATATCAA

180 Appendix 3: Sequence data Cytochrome Oxidase I (COI)

....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| 545 555 565 575 585 595 605 615 625 635 645 655 665 675 685 MW98089 CTATTTATTT GAGCTGTAGG AATTACTGCA TTATTATTAC TTCTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACTGAT CGAAACCTTA ATNCTTCCTT NTTTGACCCA GCTGGGGGAG GAGATCCAAT TTTATATCAA MW98094 TTATTTATTT GAGCTGTAGG AATTACAGCT TTATTATTAT TATtATCTTT ACCTGTTTTA GCTGGAGCTA TTACTATATT ATTAACTGAT NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN MW98097 TTATTTATTT GAGCTGTAGG AATTACAGCA TTATTATTAC TTCTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACTGAT CGAAACCTTA ACACCTCATT CTTTGACCCA GCTGGGGGAG GAGATCCAAT TTTATACCAA MW98103 TTATTTATTT GAGCGGTAGG AATCACAGCA TTATTATTAC TTTTACCTTt ACCTGTATTA GCTGGAGCAA TTACCATATT aTTAACCGAT CGAaACCTTA ATACCTCATT CTTTGACCCA GCTGGTGGAG GAGATCCNNN NNNNNNNNNN MW98106 TTATTTATTT GAGCAGTGGG AATTACAGCA TTATTATtAC TTTTATCTTt ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACCGaT CGAAACCTTA ATACCTCATT CTTTGACCCA GCTGGTGGAG GGGATCCAAT TTTATACCAA MW98128 TTATTTATTT GAGCGGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCTA TTACTATATT ACTAACTGAT CGAAATCTTA ATACCTCATT TTTTGATCCT GCTGGAGGAG GAGATCCAAT TTTATATCAA MW98129 TTATTTATTT GAGCGGTAGG AATTACAGCA TTATTATTAT TATTATCTTT ACCTGTATTA GCTGGAGCTA TTACTATATT ATTGACTGAT CGAAACCTTA ACACTTCATT TTTTGATCCT GCTGGAGGAG GaGATCCAAT TTTNNNNNNN MW98133 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACTATATT ATTAACGGAT CGAAACCTCA ATACCTCATT CTTTGATCCA GCTGGAGGAG GAGATCNNNN NNNNNNNNNN MW98136 TTATTTATTT GGGCTGTAGG AATTACAGCA TTATTATTAC TTCTATCTTT ACCTGTATTA GCTGGAGCTA TTACCATATT ATTAACTGAT CGAAACCTTA ATACCTCATT CTTTGACCCA GCTGGAGGAG GAGATCCAAT TTTATACCAA MW98138 CTATTTGTTT GAGCTGTAGG AATTACAGCA TTATTATTAC TTCTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACTGAT CGAAACCTTA ATACCTCGTT CTTTGACCCA GCTGGGGGAG GAGATCCANN NNNNNNNNNN MW98139 TTATTTATTT GGGCTGTAGG AATTACAGCA TTATTATTAC TTCTATCTTT ACCTGTATTA GCTGGAGCTA TTACCATATT ATTAACTGAT CGAAACCTTA ATACCTCATT CTTTGACCCA GCTGGAGGAG GAGATCCAAT TTTATACCAN MW98154 CTATTTGTTT GAGCCGTAGG AATTACAGCA TTATTATTAC TTCTATCTTT ACCtGTATTA GCAGGAGCAA TTACCATATT ATTAACTGAT CGAAACCTTA ATACCTCATT CTTTGACCCA GCTGGGGGAG GAGATCCAAT TTTATANNNN MW98162 CTATTTATTT GAGCTGTGGG AATTACAGCA TTATTATTAC TTCTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACTGAT CGAAACCTTA ATACCTCATT CTTTGACCCA GCTGGGGGAG GAGANNNNNN NNNNNNNNNN MW98170 TTATTTATTT GGGCTGTAGG AATTACAGCA TTATTATTAC TTCTATCTTT ACCTGTATTA GCTGGAGCTA TTACCATATT ATTGACTGAT CGAAACCTTA ATACCTCATT CTTTGACCCA GCTGGAGGAG GAGATCCAAT TTTATACCAA MW98172 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCATT GCCTGTATTA GCTGGAGCAA TTACTATATT ATTAACAGAT CGAAATCTTA ATACCTCATT CTTTGATCCG GCTGGTGGGG GAGATCCAAT TTTATATCAA MW98180 CTATTTATTT GAGCTGTAGG AATTACAGCA TTATTATTAC TTCTATCTTT ACCTGTATTA GCAGGAGCAA TTACCATATT ATTAACTGAT CGAAACCTTA ATACCTCATT CTTTGACCCA GCTGGGGGAG GAGATCCAAT TTTATATCAA MW98183 CTATTTATTT GAGCTGTAGG AATTACAGCA TTATTATTAC TTCTATCTTT ACCTGTATTA GCAGGAGCAA TTACCATATT ATTAACTGAT CGAAACCTTA ATACCTCATT CTTTGACCCA GCTGGGGGNN NNNNNNNNNN NNNNNNNNNN MW98185 TTATTTATTT GAGCGGTAGG AATTACAGCA TTGTTATTAT TACTATCTTT ACCTGTATTA GCTGGAGCTA TTACTATATT ATTAACTGAT CGAAATCTTA ACACTTCATT TTTTGATCCT GCTGGAGGAG GAGATCCAAT TTTATATCAA MW98189 CTATTTGTTT GAGCTGTGGG AATTACAGCA TTATTATTAC TTCTATCTTT ACCTGTATTA GCTGGGGCAA TTACCATATT ATTAACTGAT CGAAACCTTA ATACCTCATT CTTTGACCCA GCTGGNGGGG GAGATCCAAT TTTANNNNNN MW98203 TTATTTATTT GGGCTGTGGG AATTACAGCA TTATTATTAC TTCTATCTTT ACCTGTATTA GCTGGAGCTA TTACCATATT ACTAACTGAT CGAAACCTTA ATACCTCATT CTTTGACCCA GCTGGAGGAG GAGATCCAAT TTTATACCAA MW98205 TTATTTATTT GAGCAGTGGG AATTACGGCA TTATTATTAC TTTTATCTTT GCCTGTATTA GCGGGAGCAA TTACCATATT ATTAACCGAT CGAAATCTTA ACACCTCATT CTTTGATCCA GCTGGTGGAG GAGATCCANT TTTATATCAA MW98212 TTATTTATTT GAGCAGTGGG AATTACAGCA TTATTATTAC TTTTATCATT ACCTGTaTTA GCTGGAGCaA TTACCATATT ATTAACAGAT CGAAATCTTA ACACCTCATT CTTTGATCCA GCAGGTGGAG GAGATCCAAT TTTANNNNNN MW98220 TTATTTATTT GAGCAGTTGG TATTACTGCT TTATTATTAC TTTTATCTTT ACCTGTATTA GCAGGAGCAA TTACTATATT ATTAACAGAT CGAAATCtTA ATACTTCATT TTTTGATCCA GCTGGAGGAG GTGATCCAAT TTTATATCAN MW98228 CTATTTATTT GAGCAGTAGG AATTACAGCA TTATTGTTAC TATTATCATT ACCTGTATTA GCTGGAGCTA TTACTATATT ATTAACTGAT CGAAACCTTA ATACTTCATT TTTTGATCCT GCTGGAGGAG GGGATCCNNN NNNNNNNNNN MW98230 TTATTTATTT GAGCTGTAGG AATTACAGCT TTATTATTAT TATTATCTTT ACCTGTTTTA GCTGGAGCTA TTACCATATT ATTAACTGAT CGTAATTTAA ATACTTCTTT TTTTGATCCT GCAGGAGGTG GAGATCCAAT TTTATACCAA MW98235 TTATTCATCT GAGCGGTAGG AATTACAGCA TTATTATTAC TCTTATCTTT ACCTGTATTA GCTGGAGCTA TTACTATATT ATTAACTGAT CGAAATTTAA ATACCTCATT TTTTGATCCT GCTGGAGGGG GAGATCCTAT TTTATATCAA MW98240 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACAGAT CGAAATCTTA ATACCTCATT CTTTGACCCN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN MW98261 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACTGAT CGAAATCTTA ACACCTCATT CTTTGATCCA GCTGGTGGGG GAGATCCAAT TTTATACCAN MW98264 TTATTCATCT GAGCGGTAGG AATTACAGCA TTATTATTAC TCTTATCTTT ACCTGTATTA GCTGGAGCTA TTACTATATT ATTAACTGAT CGAAATTTAA ATACCTCATT TTTTGATCCT GCTGGAGGAG GAGATCCTAT TTTATATCAA MW98265 TTATTTATTT GAGCTGTTGG TATTACAGCA TTATTATTAT TATTATCTTT ACCTGTTTTA GCGGGAGCTA TTACTATATT ATTAACTGAT CGNAAtCTTA ATACTTCNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN MW98270 TTATTTATTT GAGCTGTAGG TATTACTGCA TTATTACTAC TTCTATCTTT ACCTGTATTA GCTGGAGCTA TCACTATATT ATTAACAGAT CGTAATCTTA ATACCTCATT TTTTGACCCT GCTGGAGGAG GAGATCCAAT TTTATATCAA MW98274 TTATTTGTTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACTATATT ATTAACCGAT CGAAACCTAA ATACCTCATT CTTTGATCCT GCTGGTGGGG GGGATCCAAT TTTATATCAN MW98278 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAT TATTATCTTT ACCTGTATTA GCTGGAGCTA TTACTATATT ATTAACTGAT CGAAATCTCA ACACTTCATT TTTTGATCCT GCAGGAGGAG GAGATCCAAT CTTATATCAA MW98284 CTATTTATTT GAGCTGTAGG AATTACAGCA TTATTATTAC TTCTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACTGAT CGAAACCTTA ATACCTCATT CTTTGACCCA GCTGGGGGAG GAGATCCAAT NNNNNNNNNN MW98285 CTATTTATTT GAGCTGTGGG AATTaCAGCA TTATTATTAC TTCTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACTGAT CGAAACCTTA ATACCTCATT CTTTGACCCA GCTGGGGGAG GAGATCCANN NNNNNNNNNN MW98294 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGcAA TTACCATATT ATTAACCGaT CGAAATCTTA ACACCTCATT CTTTGACCCA GNNNNNNNNN NNNNNNNNNN NNNNNNNNNN MW98313 TTATTTATTT GGGCTGTAGG AATTACAGCA TTATTATTAC TTCTATCTTT ACCTGTATTA GCTGGAGCTA TTACCATATT ATTAACTGAT CGAAACCTTa ATACCTCATT CTTTGACCCA GCTGGAGGAG GAGATCCAAN NNNNNNNNNN MW98315 TTATTTATTT GAGCAGTGGG AATTACAGCA TTATTATTAC TTTTATCATT GCCTGTATTA GCTGGAGCAA TTACCATATT ATTAACAGAT CGAAATCTTA ACACCTCATT CTTTGATCCA GCAGGNNNNN NNNNNNNNNN NNNNNNNNNN MW99001 CTATTTATTT GAGCAGTAGG TATTACTGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCAGGAGCAA TTACTATATT ATTAACAGAT CGAAATATTA ATACCTCATT TTTTGATCCT GCTGGAGGAG GAGATCCAAT CTTATATCAA MW99006 CTATTTATTT GAGCTGTGGG AATTACAGCA TTATTATTAC TTCTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACTGAT CGAAACCTTA NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN MW99009 TTATTTATTT GAGCAGTGGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACCGAT CGAAACCTTA ATACCTCATT CTTTGATCCA GCTGGTGGAG GAGATCNNNN NNNNNNNNNN MW99013 TTATTTATTT GAGCAGTGGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCTA TTACTATATT ATTAACTGAT CGAGATCTTA ATACCTCATT TTTTGATCCt GCTGGTGGTG GAGNNNNNNN NNNNNNNNNN MW99014 TTATTTATTT GAGCGGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACcTGTATTA GCTGGAGCTA TTACTATATT ACTAACTGAT CGAAATCTTA ATACCTCATT TTTTGATCCT GCTGGAGGAG GAGATCCAAT TTTATATCAA MW99018 TTaTTTaTTT GAGCaGTaGG AATTaCTGcA TTaTTaTTaC TTTTaTCTTT ACCTGTATTA GCTGGaGCAA TTACTATATT ATTAACTGAC CGAAACCTAA ATACTTCATT CTTTGaTCCA GCTGGAGGAG GAGANCCNNN NNNNNNNNNN MW99028 TTATTTATTT GAGCAGTAGG TATTACTGCA TTATTATTAC TTTTATCTTT AcCTGTATTA GCTGGAGCAA TTACTATATT ATTAACTGAT CGAAATATTA ATACCTCATT TTTTGATCCT GCTGGAGGAG GAGATCCTAT TTTATATCAA MW99038 TTATTCATCT GAGCGGTAGG AATTACAGCA TTATTATTAC TCTTATCTTT ACCTGTATTA GCTGGAGCTA TTACTATATT ATTAACTGAT CGAAATTTAA ATACCTCATT TTTTGATCCC GCTGGAGGGG GAGATCCTAT TTTATATCAA MW99042 TTATTTATTT GAGCGGTAGG AATTACAGCA TTATTATNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN MW99045 TTATTTATTT GAGCAGTAGG TATTACAGCA TTATTATTAT TATTATCTTT ACCTGTTTTA GCTGGTGCTA TTACTATATT ATTAACAGAT CGAAACCTAA ATACTTCATT TTTTGATCCT GCTGGAGGAG GAGATCCAAT TTTATATCAA MW99047 CTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACTATATT ATTAACTGAT CGANNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN MW99053 TTATTTATTT GAGCGGTGGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACTGAT CGAAATCTTA ACACCTCATT CTTTGATCCA GCNNNNNNNN NNNNNNNNNN NNNNNNNNNN MW99057 CTATTTATTT GAGCTGTAGG AATTACAGCA TTATTATTAC TTCTATCTTT ACCTGTATTA GCTGGAGCTA TTACCATATT ATTAACTGAT CGAAACCTTA ATACCTCATT CTTTGACCCA GcTGGAGGAG GAGATCCAAT TTTATACCAa MW99058 CTATTTATTT GAGCTGTGGG AATTACAGCA TTATTATTAC TTCTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACTGAT CGAAACCTTA ATACCTCATT CTTTGACCCA GCTGGGGGAG GAGATCCAAT TTTATACCAA MW99059 CTATTTATTT GGGCTGTAGG AATTACAGCA TTATTATTAC TTCTATCTTT ACCTGTATTA GCTGGAGCTA TTACCATATT ATTAACTGAT CGAAACCTTA ATACCTCATT CTTTGACCCA GCTGGAGGAG GAGATCCAAT TTTANNNNNN MW99060 TTATTTATCT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACTATATT ATTAACCGAT CGAAATCTTA ACACCTCATT TTTTGATCCA GCTGGTGGAG GAGATCCAAT TTTNNNNNNN MW99061 CTATTTATTT GAGCTGTAGG AATTACAGCA TTATTATTAC TTCTATCTTT ACCTGTATTA GCAGGAGCAA TTACCATATT ATTAACTGAT CGAAACCTTA ATACCTCATT CTTTGACCCA GCTGGGGGAG GAGATCCAAT TTTNNNNNNN MW99068 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGaGCAA TTACCATATT ATTAACCGAT CGAAATCTTA aNACNTCATT CTTTGATCCG GcTGGTGGAG GAGATCCAAT TTTATANNNN MW99080 TTATTTATTT GAGCAGTAGG TATTACAGCA TTATTATTAT TACTATCTTT ACCTGTTTTA GCAGGTGCTA TTACTATATT ATTAACAGAT CGAAATCTTA ATACCTCTTT CTTTGATCCT GCTGGAGGAG GAGATCCAAT TCTATATCAA MW99084 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATtACTTC TTCTTTCTTT ACCTGTTTTA GCTGGTGCTA TTACTATANN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN MW99094 CTATTTATTT GGGCTGTAGG AATTACAGCA TtATTATTAC TTCTATCTTT ACCTGTATTA GCTGGAGCTA TTACCATATT ATTAACTGAT CGAAACCTTA ATACCTCATT CTTTGACCCA GCTGGAGGAG GAGATCCAAT TTTATACCNN MW99095 TTATTTATTT GAGCAGTGGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACTGAT CGAAATCTTA ACACCTCATT CTTTGATCCA GCTGGTGGAG GAGATCCAAT TTTATATCAA MW99097 CTATTTATTT GAGCAGTAGG TATTACTGCA TTATTATTAC tTTTATCTTT ACCTGTATTA GCAGGAGCAA TTACTATATT ATTAACAGAT CGAAATATTA ATACCTCATT TTTTGATCCT GCTGGAGGAG GAGATCCAAT CTTATATCAA MW99102 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACCGAT CGAAATCTTA ACACCTCATT CTTTGATCCG GCTGGTGGAG GAGATCCAAT TTTATATCAA MW99104 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACTATATT ATTAACCGAT CGAAACCTTA ACACCTCGTT CTTTGATCCG GCTGGTGGAG GAGATCCAAN NNNNNNNNNN MW99105 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACTATATT ATTAACCGAT CGAAACCTTA ACACCTCGTT CTTTGANCCG GCTGGTNNNN NNNNNNNNNN NNNNNNNNNN MW99124 TTATTTATTT GAGCAGTCGG AATTACTGCA TTACTATTAC TTTTATCTCT ACCAGTATTA GCCGGAGCAA TTACTATATT ATTAACTGAT CGAAATCTTA ATACTTCATT TTTTGATCCA GCTGGAGGAG GAGATCCAAT TTTATACNNN MW99128 TTATTTATTT GAGCGGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCTA TTACTATATT ATTAACTGAT CGAAATCTAA ACACCTCATT CTTTGATCCT GCTGGAGGAG GAGATCCAAT TTTATACCAA MW99134 TTATTTaTTT GAGCAGTAGG TATTACTGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCGGGTGCAA TTACTATATT ATTAACTGAT CGAAACCTTA ATACTTCGTT TTTTGATCCG GCTGGAGGAG GGGATCCAAT TTTATATCAA MW99135 TTATTTATTT GAGCGGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGGGCAA TTACCATATT ATTAACCGAT CGAAACCTTA ATACCTCATT CTTTGATCCA GCTGGNNNNN NNNNNNNNNN NNNNNNNNNN

181 Appendix 3: Sequence data Cytochrome Oxidase I (COI)

....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| 545 555 565 575 585 595 605 615 625 635 645 655 665 675 685 MW99140 TTATTTATTT GAGCGGTAGG AATTACAGCA TTGTTATTAT TACTATCTTT ACCTGTATTA GCTGGAGCTA TTACTATATT ATTAACTGAT CGAAATCTTA ACACTTCATT TTTTGATCCT GCTGGAGGAG GAGATCCAAT TTTATATCAA MW99141 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACTATATT ATTAACCGAT CGAAACCTTA ACACCTCGTT CTTTGATCCG GCTGGTGGAG GAGATCCAAT TTTATATCAA MW99158 TTATTTATTT GAGCAGTAGG AATTACAGCa TTATTATTAC TTTTATCTTT ACCTGTATTA GCAGGTGCTA TCACTATATT ATTAACTGAT CGAAATTTAA ATACTTCATT TTTTGACCCT GCAGGAGGAG GAGATCCAAT TTTATATCAA MW99163 TTATTTATTT GAGCAGTAGG TATTACTGCT TTGTTATTAC TTTTATCTTT ACCTGTCTTA GCTGGAGCAA TTACTATATT ATTAACAGAT CGAAATCTTA ATACTTCATT TTTTGATCCT GCTGGAGGAG GAGATCCAAT TTTATATCAN MW99164 TTATTTATTT GAGCAGTGGG AATTACcGCA TTATTATTAC TTTTATCATT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACAGAT CGAAATCTTA ACACCTCATT CTTTGATCCA GCTGNNNNNN NNNNNNNNNN NNNNNNNNNN MW99165 TTATTTATTT GGGCTGTAGG AATTACAGCA TTATTATTAC TTCTATCTTT ACCTGTATTA GCTGGAGCTA TTACCATATT ATTAACTGAT CGAAACCTTA ATACCTCATT CTTTGACCCN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN MW99170 TTATTTATTT GAGCAGTGGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACCGAT CGAAACCTTA ATACCTCATT CTTTGATCCA GCTGGTGGAG GAGATCCAAT TTTANNNNNN MW99174 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACTGAT CGAAACCTTA ATACCTCATT CTTTGACCCA GCTGGTGGGG GAGATCCTAT TTTATATCAn MW99187 NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN MW99196 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACCGAT CGAAATCTTA ACACTTCATT CTTTGATCCG GCTGGTGGAN NNNNNNNNNN NNNNNNNNNN MW99203 TTATTTATTT GAGCGGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACCGAT CGAAACCTTA ATACCTCNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN MW99221 TTATTTATTT GAGCTGTAGG TATTACTGCA CTATTATTAT GATTATCTTT ACCAGTTTTA GCTGGAGCTA TTACTATATt ATTAACAGAT CGAGATCTTA ATACTTCATT TTTTGACCCT GCTGGAGGAG GAGATCCAAT TTTATATCAA MW99226 TTATTTATTT GAGCGGTGGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACTGAT CGAAATCTTA ACACCTCATT CTTTGATCCA GCTGGTGGAG GAGATCCAAT TTTATATCAN MW99240 TTATTTATTT GGGCTGTAGG AATTACAGCA TTATTATTAC TTCTATCTTT ACCTGTATTA GCTGGAGCTA TTACCATATT ATTAACTGAT CGAAACCTTA ATACCTCATT CTTTGACCCN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN MW99247 CTATTTATTT GAGCTGTGGG AATTACAGCA TTATTATTAC TTCTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACTGAT CGAAACCTTA ATACCTCATT CTTTGACCCA GCTGGGGGNN NNNNNNNNNN NNNNNNNNNN MW99263 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACCGAT CGAAATCTTA ACACCTCATT CTTTGATCCG GCTGGTGGAG GAGATCCAAT TTNNNNNNNN MW99264 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACCGAt CGAAATCTTA ACACCTCATT CTTTGATCCG GCTGGTGGAG GNNNNNNNNN NNNNNNNNNN MW99274 TTATTTATTT GAGCAGTGGG AATTACAGCA TTATTATTAC TTTTATCATT GCCTGTATTA GCTGGAGCAA TTACCATATT ATTAACAGAT CGAAATCTTA ACACCTCATT CTTTGATCCA GCAGGTGGGG GAGATCCAAT TTTATATCAA MW99276 TTATTTATTT GAGCAGTGGG AATTACAGCA TTATTATTAC TTTTATCATT GCCTGTATTA GCTGGAGCAA TTACCATATT ATTAACAGAT CGAAATCTTA ACACCTCATT CTTTGATCCA GCAGGTGGGG GAGATCCAAT TTTNNNNNNN MW99286 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACCGAT CGAAATCTTA ACACCTCATT CTTTGATCCA GCTGGNNNNN NNNNNNNNNN NNNNNNNNNN MW99289 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCaA TTACTATATT ATtaaCCGAT CGAGATCTTA ACACCTCATT TTTTGATCCA GCTGGTGGNN NNNNNNNNNN NNNNNNNNNN MW99292 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACTGAT CGAAATCTTA ATACCTCATT CTTTGATCCA GCTGGTGGGG GAGATCCAAT TTTATACCAA MW99301 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACTATATT ATTAACTGAT CGAAACCTTA ATACCTCGTT TTTTGACCCA GCTGGCGGGG GAGATCCAAT TTTATACCAA MW99303 TTATTTATCT GAGCAGTAGG TATTACTGCT TTATTGTTAC TTTTATCTTT ACCTGTTTTA GCTGGAGCAA TTACTATATT ATTAACAGAT CGAAATCTTA ATACTTCATT TTTTGATCCT GCTGGAGGAG GAGANCCAAT TTTATATCAN MW99309 TTATTTATTT GAGCAGTGGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACCGAT CGAAACCTTA ATACCTCATT CTTTGATCCG GCTGGTGGAG GAGATCCAAT TTTATATCAA MW99314 TTATTTATTT GAGCGGTGGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACTGAT CGAAATCTTA ACACCTCATT CTTTGATCCA GCTGGTGGAG GAGATCCAAT TTTATATCAA MW99319 TTATTTATTT GAGCAGTGGG AATTACAGCA TTATTATTAC TTTTATCATT GCCTGTATTA GCTGGAGCAA TTACCATATT ATTAACAGAT CGAAATCTTA ACACCTCATT CTTTGATCCA GCAGGTGGGG GAGATCCAAT TTTATATCAA MW99320 TTATTTATTT GAGCAGTGGG AATTACNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN MW99341 TTATTTATTT GAGCAGTAGG AATTACGGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACTGAT CGAAATCTTA ATACCTCATT CTTTGATCCA GCTGGTGGGG GAGATCCAAT TTTATACCAA MW99344 TTATTTATTT GAGCGGtAGG AATCACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACTGAT CGAAACCTTA ATACCTCATT CTTTGACCCA GCTGGTGGAG GAGANNNNNN NNNNNNNNNN MW99353 CTATTTATTT GGGCTGTAGG AATTACAGCA TTATTATTAC TTCTATCTTT ACCTGTATTA GCTGGAGCTA TTACCATATT ATTAACTGAT CGAAACCTTA ATACCTCATT CTTTGACCCA GCTGGGGGAG GAGATCCAAT TTTATACCAN MW99357 CTATTTATTT GGGCTGTAGG AATTACAGCA TTATTATTAC TTCTATCTTT ACCTGTATTA GCTGGAGCTA TTACCATATT ATTAACTGAT CGAAACCTTA ATACCTCATT CTTTGACCCA GCTGGGGGAG GAGATCCAAT TTTATACCAN MW99372 TTATTTATTT GAGCGGTAGG AATCACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACCGAT CGAAACCTTA ATACCTCATT CTTTGACCCA GCTGGTGGGG GAGATCCAAT NNNNNNNNNN MW99374 TTATTTATTT GAGCGGTGGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACTGAT CGAAATCTTA ACACCTCATT CTTTGATCCA GCTGGTGGAG GAGATCCAAT TTTANNNNNN MW99376 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACCGAT CGAAATCTTA ACACCTCATT CTTTGATCCA GCTGGTGGAG GAGATCCAAT TTTATACCAA MW99380 TTATTTATTT GAGCAGTGGG AATTACAGCA TTATTATTAC TTTTATCATT GCCTGTATTA GCTGGAGCAA TTACCATATT ATTAACAGAT CGAAATCTTA ACACCTCATT CTTTGACCCA GCAGGTGGGG GAGATCCAAT TTTATATCAA MW99381 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACCGAT CGAAATCTTA ACACCTCATT CTTTGACCCA GCTGGtGGGG GAGATCCAAT TTTNNNNNNN MW99382 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACCGAT CGAAATCTTA ACACCTCATT CTTTGACCCA GCTGGTGGGG GAGATCCAAT TTTATATCAA MW99393 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACCGAT CGAAATCTTA ACACCTCATT CTTTGATCCA GCTGGTGGAG GAGATCCAAT TTTATACCAA MW99398 NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN MW99406 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACTATATT ATTAACCGAT CGAAATCTTA ACACCTCATT CTTTGATCCA GCTGGTGGAG GAGATCCAAT TTTATACCAN MW99407 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACCGAT CGAAATCTTA ANACNTCATT CTTTGATCCG GCTGGTGGAG GAGATCCAAT TTTATANNNN MW99408 CTATTTATTT GAGCTGTAGG AATTACAGCA TTATTATTAC TTCTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACTGaT CGAAACCTCA ATACCTCATT CTtTGACCCA GCTGGGGGGG GGGATCCAAT NNNNNNNNNN MW99413 CTATTTATTT GAGCTGTGGG AATTACAGCA TTATTATTAC TTCTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACTGAT CGAAACCTTA ATACCTCATT CTTTGACCCA GCTGGGGGAG GAGATCCAAT TTTATACCAA MW99422 CTATTTATTT GAGCTGTGGG AATTACAGCA TTATTATTAC TTCTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACTGAT CGAAACCTTA ATACCTCATT CTTTGACCCA GCTGGGGGAG GAGATCCAAT TTTATACCAA MW99425 TTATTTATTT GAGCTGTAGG AATTACTGCT TTATTATTAC TTTTATCTTT ACCTGTATTA GCAGGAGCTA TTACTATATT ATTAACTGAT CGTAATCTTA ATACATCATT TTTTGACCCT GCTGGAGGAG GAGATCCAAT TTTATATCAA MW99430 TTATTTATTT GGGCGGTAGG TATTACTGCT CTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACTGAT CGAAATCTTA ACACTTCATT TTTTGATCCT GCCGGAGGAG GAGATCCAAT TTNNNNNNNN MW99435 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAT TATTATCTTT ACCTGTATTA GCTGGAGCTA TTACTATATT ATTAACTGAT CGAAATCTCA ACACTTCATT TTTTGATCCT GCAGGAGGAG GAGATCCAAT CTTATATCAA MW99448 CTATTTATTT GGGCTGTAGG AATTACAGCA TTATTATTAC TTCTATCTTT ACCTGTATTA GCTGGAGCTA TTACCATATT ATTAACTGAT CGAAACCTTA ATACCTCATT CTTTGACCCA GCTGGAGGAG GAGATCCAAT TTTATACCAA MW99449 CTATTTATTT GGGCTGTAGG AATTACAGCA TTATTATTAC TTCTATCTTT ACCTGTATTA GCTGGAGCTA TTACCATATT ATTAACTGAT CGAAACcTTA ATACCTCATT CTTTGACCCA GCTGGAGGAG GAGATCCAAT TTTATACCAA MW99465 CTATTTATTT GGGCTGTAGG AATTACAGCA TTATTATTAC TTCTATCTTT ACCCGTATTA GCTGGAGCTA TTACCATATT ATTAACTGAT CGAAACCTTA ATACCTCATT CTTTGACCCA GCTGGAGGAG GAGATCCAAT TTTATACCAA MW99469 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACTATATT ATTAACTGAT CGAAACCTTA ATACCTCGTT TTTTGACCCA GCTGGCGGGG GAGATCCAAT TTTATNNNNN MW99471 TTATTTATTT GAGCAGTGGG AATTACAGCA TTATTATTAC TTTTATCATT GCCTGTATTA GCTGGAGCAA TTACCATATT ATTAACAGAT CGAAATCTTA ACACCTCATT CTTTGATCCA GCAGGTGGGG NNNNNNNNNN NNNNNNNNNN MW99473 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACCGAT CGAAATCTTA ACACCTCATT CTTTGATCCA GCTGGTGGAG GAGATCCAAT TTTATACCAA MW99476 TTATTTATTT GAGCGGTGGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACTGAT CGAAATCTTA ACACCTCATT CTTTGATCCA GCTGGTGGAG GAGATCCAAT TTTATATCAA MW99479 TTATTTATTT GAGCAGTGGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACTGAT CGAAATCTTA ACACCTCATT CTTTGATCCA GCTGGTGGAG GAGATCCANN NNNNNNNNNN MW99494 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCATT GCCTGTATTA GCTGGAGCAA TTACTATATT ATTAACAGAT CGAAATCTTA ATACCTCATT CTTTGATCCG GCTGGTGGAG GAGATCCNNN NNNNNNNNNN MW99501 CTATTTATTT GAGCTGTAGG AATTACAGCA TTATTATTAC TTCTATCTTT ACCTGTATTA GCAGGAGCAA TTACCATATT ATTAACTGAT CGAAACCTTA ATACCTCATT CTTTGACCCA GCTGGGGGAG GAGATCCAAT TTTATATCAA MW99508 TTATTTATTT GAGCAGTGGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACTGAT CGAAATCTTA ACACCTCATT CTTTGACCCA GCTGGTGGAG GAGATCCAAT TTTATATCAA MW99514 TTATTTATTT GAGCGGTAGG AATTACAGCA TTGTTATTAT TACTATCTTT ACCTGTATTA GCTGGAGCTA TTACTATATT ATTAACTGAT CGAAATCtTA ACACtTCATT TTTTGATCCT GCNGGAGGAG GAGATCCAAT TTTANNNNNN MW99515 TTATTTATTT GAGCTGTAGG AATTACAGCT TTACTATTAC TATTATCTTT ACCTGTTTTA GCTGGGGCTA TTACTATATT ATTAACTGAT CGTAATTTAA ATACTTCTTT TTTTGATCCT GCAGGAGGTG GAGATCCAAT TTTATATCAA MW99520 TTATTTATTT GAGCTGTAGG AATTACAGCT TTATTACTAT TACTATCTTT ACCTGTTTTA GCTGGAGCTA TTACTATATT ATTAACTGAT CGTAATTTAA AtACTTCTTT TTTTGATCCT GCAGGTGGTG GAGATCCNAT TTTATATCAA MW99526 TTATTTATTT GAGCGGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGGGCAA TTACCATATT ATTAACCGAT CGAAACCTTA ATACCTCATT CTTTGATCCA GCTGGTGGAG GAGATCNNNN NNNNNNNNNN MW99536 TTATTTATTT GAGCGGTAGG AATTACAGCA TTATTATTAT TATTATCTTT ACCTGTATTA GCTGGAGCTA TTACTATATT ATTAACTGAT CGAAACCTTA ACACTTCATT TTTtGATCCT GCTGGAGGAG GAGATCCAAT TTTATATCAA MW99537 TTATTCATCT GAGCGGTAGG AATTACAGCA TTATTATTAC TCTTATCTTT ACCTGTATTA GCtGGAGCTA TTACTATATT ATTAACTGAT CGAAATTTAA ATACCTCATT TTTTGATCCT GCTGGAGGGG GAGATCCTAT TTTATATCAA MW99538 TTATTCATCT GAGCGGTAGG AATTACAGCA TTATTATTAC TCTTATCTTT ACCTGTATTA GCTGGAGCTA TTACTATATT ATTAACTGAT CGAAATTTAA ATACCTCATT TTTTGATCCT GCTGGAGGGG GAGATCCTAT TTTATATCAA MW99546 TTATTTATTT GAGCTGTAGG AATTACAGCA TTATTATTAC TTTTATCCTT ACCCGTATTA GCTGGTGCAA TTACCATATT ATTAACTGAT CGAAATCTTA ACACCTCATT CTTTGATCCG GCTGGAGGAG GaGATCCAAT TTTATATCAA MW99550 TTATTCATCT GAGCGGTAGG AATTACAGCA TTATTATTAC TCTTATCTTT ACCTGTATTA GCTGGAGCTA TTACTATATT ATTAACTGAT CGAAATTTAA ATACCTCATT TTTTGATCCT GCTGGAGGGG GAGATCCTAT TTTATATCAA

182 Appendix 3: Sequence data Cytochrome Oxidase I (COI)

....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| 545 555 565 575 585 595 605 615 625 635 645 655 665 675 685 MW99552 TTATTTATTT GAGCGGTGGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACTGAT CGAAATCTTA ACACCTCATT CTTTGATCCA GCTGGTGGAG GAGATCCAAT TTTATATCAA MW99559 TTATTTATTT GAGCAGTGGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACCGAT CGAAaTCTTA ATACCTCATT CTTTGATCCG GCTGGTGGAG GAGATCCAaT TTTATATNNN MW99565 TTATTTATTT GAGCGGTAGG AATCACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCAGGAGCAA TTACCATATT ATTAACCGAT CGAAACCTTA ATACCTCATT CTTTGATCCA GCTGGTGGAG GAGATCCAAT TTTATACCAA MW99579 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACTATATT ATTAACTGAT CGAAACCTCA ATACCTCATT CtTTGATCCA GCTGGAGGAG GAGATCCAAT TTTATATCAN MW99591 TTATTTATTT GAGCAGTGGG AATTACAGCA TTATTATTAC TTTTATCATT GCCCGTATTA GCTGGGGCAA TTACCATATT ATTAACAGAT CGAAATCTTA ATACCTCATT CTTTGACCCG GCTGGTGGAG GAGATCCAAT TTTATATCAA MW99605 TTATTTATTT GAGCAGTGGG AATTACAGCA TTATTATTAC TTTTATCATT GCCCGTATTA GCTGGGGCAA TTACCATATT ATTAACAGAT CGAAATCTTA ATACCTCATT CTTTGACCCG GCTGGTGGAG GAGATCNNNN NNNNNNNNNN MW99608 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAT TATTATCTTT ACCTGTATTA GCTGGAGCTA TTACTATATT ATTAACTGAT CGAAATCTCA ACACTTCATT TTTTGATCCT GCAGGAGGAG GAGATCCAAT CTTATATCAA MW99612 TTATTTATTT GAGCGGTAGG AATTACAGCA TTATTATTAT TATTATCTTT ACCTGTATTA GCTGGAGCTA TTACTATaTT ATTAACTGAT CGAAATCTTA ACACTTCATT TTTTGATCCT GCTGGAGGAG GAGATCNNNN NNNNNNNNNN MW99613 TTATTTATTT GAGCTGTGGG AATTACAGCA TTATTATTAC TTTTATCCTT ACCCGTATTA GCTGGTGCAA TTACCATATT ATTAACTGAT CGAAATCTTA ACACCTCATT CTTTGATCCG GCTGGANNNN NNNNNNNNNN NNNNNNNNNN OK96022 TTATTTATTT GAGCGGTAGG AATTACAGCA TTGTTATTAT TATTATCTTT ACCTGTATTA GCTGGAGCTA TTACTATATT ACTAACTGAT CGAAATCTTA ACACTTCATT TTTTGATCCT GCTGGAGGAG GAGATCCAAT TTTATATCAA OK99001 TTATTTATTT GAGCGGTAGG AATTACAGCA TTATTATTAT TATTATCTTT ACCTGTATTA GCTGGAGCTA TTACTATATT ATTAACTGAT CGAAATCTTA ACACTTCATT TTTTGATCCT GCTGGAGGAG GAGATCCAAT TTTATATNNN WE00002 TTATTTATTT GAGCAGTAGG AATtACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTGACAGAT CGAAACCTTA ATACTTCATT CTTTGATCCG GCTGGTGGGG GAGATCCTAT TCNNNNNNNN WE00003 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTCT GCCTGTATTA GCTGGAGCTA TTACTATATT ATTAACTGAT CGAAATCTTA ATACCTCATT NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN WE02321 CTATTTATTT GGGCTGTAGG AATTACAGCA TTATTATTAC TTCTATCTTT ACCAGTATTA GCTGGAGCTA TTACCATATT ATTAACTGAT CGAAACCTTA ATACCTCATT CTTTGACCCA GCTGGGGGAG GAGATCCAAT TTTATACCAA WE02421 CTATTTATTT GGGCTGTAGG AATTACAGCA TTATTATTAC TTCTATCTTT ACCTGTATTA GCTGGAGCTA TTACCATATT ATTAACTGAT CGAAACCTTA ATACCTCATT CTTTGACCCA GCTGGAGGAG GAGATCCAAT TTCAATNNNN WE02431 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACCGAT CGAAATCTTA ATACCTCATT CTTTGACCCG GCTGGTGGAG GAGATCCAAT TTTATATCAN WE02451 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACTGAT CGAAACCTTA ATACTTCATT CTTTGATCCC GCTGGTGGAG GGGATCCTAT TTTATATCAA WE02491 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTGTTA GCTGGAGCAA TTACCATATT ATTAACTGAT CGAAACCTTA ATACCTCATT CTTTGATCCA GCTGGAGGAG GAGATCCTAT TTTNNNNNNN WE02531 TTATTTATTT GAGCAGTAGG AATTACGGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACCGAC CGAAATCTTA ACACCTCATT CTTTGATCCA GCTGGTGGAG GAGATCCAAT TTTATACCNN WE02535 TTATTTATTT GGGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACCGAT CGAAACCTTA ACACCTCATT CTTTGATCCG GCTGGTGGAG GAGATCCAAT TTNNNNNNNN WE02536 CTATTTATTT GGGCTGTAGG AATTACAGCA TTATTATTAC TTCTATCTTT ACCTGTATTA GCTGGAGCTA TTACCATATT ATTAACTGAT CGAAACCTTA ATACCTCATT CTTTGACCCA GCTGGGGGGG GAGATCCAAT TNNNNNNNNN WE02591 TTATTTATTT GAGCAGTGGG AATTACGGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACCGAT CGAAATCTTA ATACCTCATT TTTTGATCCA GCTGGTGGAG GAGATCCAAT TTTATACNNN WE02612 TTATTTATTT GAGCAGTGGG AATTACGGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACCGAT CGAAATCTTA ATACCTCATT TTTTGATCCA GCTGGTGGAG GAGATCCAAT TTTATACCAA WE02614 TTATTTATTT GAGCAGTGGG AATTACGGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACCGAT CGAAATCTTA ATACCTCATT TTTTGATCCA GCTGGTGGAG GAGATCCAAT TTNNNNNNNN WE02621 CTATTTATTT GGGCTGTAGG AATTACAGCA TTATTATTAC TTCTATCTTT ACCTGTATTA GCTGGAGCTA TTACCATATT ATTAACTGAT CGAAACCTTA ATACCTCATT CTTTGACCCA GCTGGAGGAG GAGATCCAAT TTTNNNNNNN WE02661 TTATTTATTT GAGCAGTAGG AATTACGGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACCGAT CGAAATCTTA ACACCTCATT CTTTGATCCA GCTGGTGGAG GAGATCCAAN NNNNNNNNNN WE02662 TTATTTATTT GAGCAGTGGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACCGAT CGAAACCTTA ATACCTCATT CTTTGACCCG GCTGGTGGAG GAGATCCAAT TTTATATCAA WE02671 TTATTTATTT GAGCGGTAGG AATTACGGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACCGAT CGAAATCTTA ACACCTCATT CTTTGATCCA GCTGGTGGAG GAGATCCAAT TTTATACCAA WE02674 CTATTTATTT GGGCTGTAGG AATTACAGCA TTATTATTAC TTCTATCTTT ACCTGTATTA GCTGGAGCTA TTACCATATT ATTAACCGAT CGAAACCTTA ATACCTCATT CTTTGACCCA GCTGGAGGAG GAGATCCAAT TTTATACCAA WE02676 TTATTTATTT GAGCAGTGGG AATTACGGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACCGAT CGAAACCTTA ACACCTCATT CTTTGATCCG GCTGGTGGGG GAGATCCAAT TTTATACCAA WE02677 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACCGAT CGAAATCTTA ACACCTCATT CTTTGACCCA GCTGGTGGGG GAGATCCAAT TTNNNNNNNN WE85001 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTGTTA GCTGGAGCAA TTACCATATT ATTAACTGAT CGAAACCTTA ATACCTCATT CTTTGATCCA GCTGGAGGAG GAGATCNNNN NNNNNNNNNN WE98001 TTATTTATTT GAGCAGTAGG AATTACAGCA TTATTATTAC TTTTATCTTT ACCTGTATTA GCTGGAGCAA TTACCATATT ATTAACAGAT CGAAACCTTA ATACCTCATT CTTTGACCCA GCTGGTGGAG GAGATCCAAT TTTACATCAA

Cytochrome b (Cytb)

....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| 5 15 25 35 45 55 65 75 85 95 105 115 125 135 MW98006 ATTATTACAG GATTATTTTT AACTATATAT TATTCTGCTA ATATTAATTT AGCTTTTTAT AGTGTTAATT ACATTTGTCG AGATGTTAAT TATGGTTGAT TAATTCGAAC TCTTCATGCC AATGGAGCAT CTTTTTTTTT MW98055 ATTATCACAG GATTATTTTT GACTATGTAC TACTCAGCTA ATATTAATTT AGCATTTTTT AGAGTAAATT ATATTTGTCG AGATGTAAAT TATGGATGAT TAATTCGAAC TTTACATGCT AATGGGGCTT CATTTTTTTT MW98102 ATTATCACAG GATTATTTTT GACTATATAT TATTCAGCTA ATATTAATTT AGCATTTTTT AGAGTAAATT ATATTTGTCG AGATGTAAAT TACGGATGAT TAATTCGAAC TTTACATGCT AATGGAGCTT CTTTTTTTTT MW98225 ATTTTTACAG GATTATTTTT AACTATATAT TACTCAGCTA ATATTAATTT AGCATTTTTC AGAGTAAATT ATATCTGCCG AGATGTAAAT TATGGATGAC TAATTCGAAC TTTACATGCC AATGGtGCTT CATTTTtCTT MW99258 ATTATCACAG GATTATTTTT AACTATGTAC TATTCAGCTA ATATTAATTT AGCATTTTTT AGAGTAAATT ACATTTGTCG AGATGTAAAT ?ATGGATGAT TAATTCGAAC TTTACACGCT Aa?GGAGCTT CATTTTTTTT

....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|.. 145 155 165 175 185 195 205 215 225 235 245 255 265 275 MW98006 TATTTGtATT TtCACtCATA TtGGACgAGG AATTTATtAt gAATCATtC- --AATTtAAA ACTTACAtGA AtAATTGGTG tAATtATTTT ATTTATATTA ATAGCAACAg CATTtATAGG ATATGTCCTA CCTGGNN MW98055 TGTTTGTATT TATACTCATA TTGGACGAGG AATTTATTAT GAATCCTTT- --AACCTAAA AATAACATGA ATAATCGGAG TATTGATTTT ATTTATATTG ATAGCAACAG CTTTtATAGG AtATGTTTTA CCTTNNN MW98102 TGTTTGTATT TATATTCATA TtGGACGAGG AATTTATTAT GAATCCTTT- --AACTtAAA AAtAACATGA ATAATCGGAG TATTAATTTt ATTtATATtA ATAGCAACAG CTTTTATAGG AtAtgTTCTN CCCTNNN MW98225 TATTTGCATT TATATTCATA TtGGACGAGG AATTTATCAT GAATCATTCA TTAATTTAAA ATtAACATGA ATAATTGGAG TGTtAAtCtt AtTtAtAtTA AtAGCAACaG CTTTTATAGG ATATGTTTTA CCTTGAG MW99258 TGTTTGTATT TATATTCATA TTGGACGAGG AATTTATTAT GAATCCTTT- --AaCTtAAA AATAACATGA AtAATCGGAG TATTAATTTT ATTTATaTTA ATAGCAACAG CTTTTAtGGG ATACGTTTTA CCTTNNN

183 Appendix 3: Sequence data NADH dehydrogenase subunit 1 (ND1)

....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| 5 15 25 35 45 55 65 75 85 95 105 115 JC00057 TTAGCTTTTT TAACATTAAT AGAACGAAAA GTTTTAAGAT ATATACAAAT TCGTAAAGGA CCTAATAAAG TTGGGTTTTT AGGAATATTA CAACCTTTTT CTGATGGTAT TAAATTATTT MW00032 TTAGCTTTTT TAACATTAAT AGAACGAAAA GTTTTGAGAT ATATACAAAT TCGTAAAGGA CCAAATAAGG TTGGATTTTT AGGAGTATTA CAACCTTTTT CTGATGGTAT TAAATTATTT MW00056 TTAGCTTTTT TAACATTAAT AGAACGAAAA GTTTTAAGAT ACATACAAAT TCGTAAAGGA CCAAATAAGG TTGGATTtTT AGGGATATTA CAACCTTTTT CTGATGGTAT TAAATtATTT MW00072 TTAGCTTTTT TAACACTAAT ASAACGAAAA GTTTTGAGAT ATATTCAAAT TCGTAAAGGG CCAAATAAGG TTGGATTTTT AGGGATATTA CAACCTTTTT CAGACGGTAT TAAATTATTT MW00076 TTAGCTTTTT TAACATTAAT AGAACGAAAA GTTTTAAGAT ATATACAAAT TCGTAAAGGG CCAAATAAGG TTGGATTTTT GGGAATATTA CAACCTTTTT CAGATGGTAT TAAATTATTT MW00189 TTGGCTTTTT TGACACTAAT AGAACGAAAA GTTTTAAGAT ACATGCAAAT TCGTAAAGGT CCAAATAAGG TTGGATTTTT AGGGTTGTTA CAACCTTTTT CGGATGGTGT TAAATTATTT MW00231 TTAGCTTTTT TGACATTAAT AGAACGAAAA GTTTTAAGAT ACATACAAAT TCGTAAAGGA CCGAATAAGG TTGGGTTTTT AGGAATATTA CAACCTTTTT CGGATGGTAT TAAATTATTT MW00302 TTAGCTTTTT TAACATTAAT AGAACGAAAA GTTTTAAGAT ATATGCAAAT TCGTAAGGGA CCAAATAAGG TTGGATTTTT AGGGATATTA CAACCTTTTT CTGATGGTAT TAAATTATTT MW00328 TTAGCTTTTT TAACATTAAT AGAACGAAAA GTTTTGAGAT ATATACAAAT TCGTAAAGGA CCAAATAAGG TTGGATTTTT AGGGATATTA CAACCTTTTT CTGACGGTGT TAAATTATTT MW00409 TTAGCTtTTT TAACATTAAT AGAACGAAAA GTTTTAAGAT ATATACAAAT TCGTAAAGGG CCAAATAAGG TTGGATTTTT AGGAATATTA CAACCTTTTT CAGATGGTGT TAAATTATTT MW00412 TTAGCTTTTT TAACATTAAT AGAACGAAAA GTTTTAAGAT ATATACAAAT TCGTAAAGGG CCAAATAAGG TTGGATTTTT GGGTATATTA CAGCCTTTTT CAGACGGTGT TAAATTGTTC MW00497 TTAGCATTTT TAACTTTATT AGAACGAAAG GTTTTAAGAT ATATACAAAT TCGAAAAGGT CCTAATAAAA TAGGGTTTAT AGGAATGTTA CAGCCTTTTT CTGATGGAAT TAAGTTATTT MW00530 TTAGCTTTTT TAACATTAAT AGAACGAAAA GTTTTAAGAT ATATACAAAT TCGTAAAGGG CCAAATAAGG TTGGATTTTT AGGTATATTA CAGCCTTTTT CAGATGGTGT TAAATTGTTC MW98154 TTAGCTTTTT TAACATTAAT AGAACGAAAA GTTTTGAGAT ATATGCAGAT TCGTAAAGGG CCGAATAAGG TTGGATTTTT AGGAATATTA CAACCTTTTT CAGATGGTAT TAAGTTATTT MW98172 TTAGCTTTTT TGACATTAAT AGAACGAAAA GTTTTAAGAT ACATACAAAT TCGTAAAGGG CCAAATAAGG TTGGATTTTT AGGAATATTA CAACCTTTTT CGGATGGTAT TAAATTATTT MW98228 CTACCTTTTT TGACATTAAT GGAGCGGAAG GTTTTAAGAT ATATACAAAT TCGTAAAGGT CCTAATAAGG TTGGATTTTT AGGGATATTA CAACCTTTTT CAGATGGTAT TAAGTTATTT MW99014 TTaGCTTTTT TAACATTAAT AGAACGAAAA GTTTTAAGAT ACATACAAAT TCGTAAAGGA CCAAATAAAG TTGGATTTTT AGGAATATTA CAACCTTTTT CAGATGGTGT TAAATTATTT MW99045 TTAGCTtTTT TAACATTAAT AGAACGAAAA GTTTTAAGAT ATATACAAAT TCGTAAAGGA CCTAATAAAA TTGGATTTTT AGGTATATTA CAACCATTTT CTGATGGTAT TAAATTATTC MW99068 TTGGCCTTTT TAACATTAAT AGAACGAAAA GTTTTAAGAT ACATGCAAAT TCGCAAAGGT CCAAATAAGG TTGGATTTTT AGGGTTATTA CAACCTTTTT CGGATGGTAT TAAATTATTT MW99105 TTGGCCTTTT TGACACTAAT AGAACGAAAA GTTTTAAGAT ACATGCAAAT TCGTAAAGGT CCAAATAAGG TTGGATTTTT AGGGTTATTA CAACCTTTTT CGGATGGTGT TAAATTATTT MW99135 TTAGCTTTTT TAACATTAAT AGAACGAAAA GTTTTGAGAT ATATACAAAT TCGAAAAGGG CCAAATAAGG TTGGGTTTTT AGGAATATTA CAACCTTTTT CTGACGGTAT TAAATTATTT MW99164 TTAGCTttTT TGACATTAAT AGAACGAAAA GTTTTAAGAT ATATGCAAAT TCGTAAAGGA CCAAATAAGG TTGGGTTTTT AGGAATATTA CAACCTTTTT CGGATGGTAT TAAATTATTT MW99406 TTAGCTTTTT TAACATTAAT AGAACGAAAA GTTTTAAGAT ATATACAAAT TCGTAAAGGA CCAAATAAGG TTGGGTTTTT AGGGATATTG CAACCTTTTT CTGATGGTAT TAAATTATTT MW99471 TTAGCTTTTT TGACATTAAT AGAACGAAAA GTTTTAAGAT ACATACAAAT TCGTAAAGGA CCAAATAAGG TTGGGTTtTT AGGGATATTA CAACCTTTTT CGGATGGTAT TAAATTATTT MW99550 TTAGCTTTTT TAACGTTAAT AGAACGAAAG GTTTTAAGAT ATATACAAAT TCGTAAGGGG CCTAATAAGG TTGGGTTTTT AGGAATATTA CAACCTTTTT CAGATGGTGT AAAACTATTT

....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| 125 135 145 155 165 175 185 195 205 215 225 JC00057 ACTAAAGAAA TGATTTATTT AAATTCATCA AATTATATAT TTTATTATTT GTCTCCTATT ATAGGTTTTA TTTTATCTTT AATAATTTGA ATATTAATTC CTTATTATTT MW00032 ACTAAAGAGA TAATTTATTT AAATTCTTCA AATTATATAT TTTATTATTT ATCTCCTATT GTAGGTTTTA TTTTATCTTT AGTGATTTGA ATATTAATTC CTTATTATTT MW00056 ACTAAAGAGA TAATTTATTT AAATTTTTCA AATTATATAT TTTATTATTT GTCCCCTATT ATAGGTTTTA TTTTATCTTT AGTGATTTGA ATGTTAATTC CTTATTATTT MW00072 ACTAAAGAAA TAATTTATTT AAATTCTTCA AATTATATGT TTTATTATTT ATCTCCTATT ATAGGTTTTA TTTTATCTTT AGTGGTTTGA ATATTAATTC CTTATTATTT MW00076 ACTAAAGAAA TAATTTATTT AAATTCTTCA AATTATATAT TTTATTATTT ATCTCCTATT ATAGGTTTTA TTTTATCTTT GGTAATTTGA ATGTTAATTC CTTATTGTTT MW00189 ACTAAAGAAA TAATTTATTT AAATTCTTCA AATTATATGT TTTATTATTT ATCCCCTATT ATAGGTTTTA TTTTATCTTT AGTGATTTGA ATATTAATTC CTTATTATTT MW00231 ACTAAAGAAA TAATTTATCT AAATTCTTCA AATTATATGT TTTATTATTT ATCTCCTATT ATAGGTTTTA TTTTATCTTT GGTGATTTGA ATATTAATTC CTTATTATTT MW00302 ACTAAAGAAA TAATTTATTT AAATTCTTCA AATTATATGT TTTATTATTT ATCTCCTATT ATAGGTTTTA TTTTATCTTT AGTGATTTGA ATGTTAATTC CTTATTATTT MW00328 ACTAAAGAAA TAATTTATTT AAATTCTTCA AATTATATAT TTTATTATTT GTCTCCTATT ATAGGTTTTA TTTTATCTTT AGTGATTTGA ATGTTAATTC CTTATTATTT MW00409 ACTAAAGAGA TAATTTATTT AAATTCTTCT AATTATATAT TTTATTATTT AtCCCCTATT ATAGGTTTtA TTTtATCTTT aGTAATTTGA ATATTAATTC CTtATTATTT MW00412 ACTAAAGAAA TAATTTATTT AAATTCTTCA AATTATATAT TTTATTATTT TTCTCCTATT ATAGGTTTTA TTTTATCTTT AGTGATTTGA ACATTAATTC CTTATTATTT MW00497 ACAAAAGAAA TAATTTATTT AAATTCTTCT AATTATTTAT TTTATTATTT AGCTCCAGTT ATTGGATTTA TTTTATCTTT AATGGTTTGA ATATTAATTC CTTACTATTT MW00530 ACTAAAGAAA TAATTTATTT AAATTCTTCA AATTATATAT TTTATTATTT TTCTCCTATT ATAGGTTTTA TTTTATCTTT AGTGATTTGA ATATTAATTC CTTATTATTT MW98154 ACTAAAGAAA TAATTTATTT AAATTCTTCA AATTATATAT TTTATTATTT GTCTCCTATT ATAGGTTTTA TTTTATCTTT AGTGATTTGA ATATTAATTC CTTATTATTT MW98172 ACTAAAGAAA TAATTTATCT AAATTCTTCA AATTATATGT TTTATTATTT ATCTCCTATT ATAGGTTTTA TTTTATCTTT GGTGATTTGA ATATTAATTC CTTATTATTT MW98228 ACAAAAGAAA TAATTTATTT AAATTCTTCA AATTATATAT TTTATTATTT ATCTCCTATT ATAGGATTTA TTTTATCTTT AGTAATTTGA ATGTTAATTC CTTATTATTT MW99014 ACTAAAGAAA TAATTTATTT AAATTCTTCA AATTATATAT TTTATTATTT ATCTCCTATT GTAGGTTTTA TTTTATCTTT AGTGGTTTGA ATATTAATTC CTTATTATTT MW99045 ACAAAAGAAA TAATTTATTT AAATTCATCA AATTATTTAT TTTATTATTT ATCTCCAATT ATTGGTTTTA TATTATCTTT AATATTATGA ATATTAATTC CATATTATTT MW99068 ACTAAGGAAA TAATTTATTT AAATTCTTCA AATTATATGT TTTATTATTT ATCTCCTATT ATAGGTTTTA TTTTATCTTT AGTGATTTGA ATATTAATTC CTTATTATTT MW99105 ACTAAAGAAA TAATTTATTT AAATTCTTCA AATTATATGT TTTATTATTT ATCCCCTATT ATAGGTTTTA TTTTATCTTT AGTGATTTGA ATATTAATTC CTTATTATTT MW99135 TCTAAAGAAA TAATTTATTT AAATTCTTCA AATTATATAT TTTATTATTT ATCTCCTATT ATAGGTTTTA TTCTATCTTT GGTAATTTGA GTGTTAATTC CTTATTATTT MW99164 ACTAAAGAAA TAATTTATCT AAATTCTTCA AATTATATGT TTTATTATTT ATCTCCTATT ATAGGTTTTA TTTTATCTTT GGTGATTTGA ATATTAATTC CTTATTATTT MW99406 ACTAAAGAGA TAATTTATTT AAATTCTTCA AACTATATAT TTTATTATTT GTCCCCTATT ATAGGTTTTA TTTTATCTTT AGTGATTTGA ATGTTAATTC CTTATTATTT MW99471 ACTAAAGAAA TAATTTATCT AAATTCTTCA AATTATATGT TTTACTATTT ATCTCCTATT ATAGGTTTTA TTTTATCTTT GGTGATTTGA ATATTAATTC CTTATTATTT MW99550 ACTAAAGAGA TAATTTATTT AAATTTTTCA AATTATATAT TTTATTATTT ATCTCCTATT ATAGGTTTTA TTTTATCCTT GGTGATTTGA ATATTAATTC CTTATTATTT

184 Appendix 3: Sequence data Internal transcribed spacer 2 (ITS-2)

....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| 5 15 25 35 45 55 65 75 85 95 105 115 125 135 145 155 AD98001 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ AD98009 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ AD98012 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ AD98018 NNNNNGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~C~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ AD98024 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTYT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTYGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ AD98036 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ DS00001 TCATAGTGTG AANTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~CT~ ~~~~~~~~~~ ~~~~~~~~~~ DS01001 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ JC00029 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ JC00039 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ JC00040 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ JC00042 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ JC00043 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ JC00045 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ JC00051 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ JC00055 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~~~~ ~~~~~~~~~~ ~~~~~~~~~~ JC00061 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ JC00063 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ JC01014 NNNNNNNGTG AACTGCAGGA CACATKTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ JM00001 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW00015 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW00024 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAATT CATATGCCAC ACTGTTCGTC ~T~~~~~~~~ ~~~~~~~~~~ ~~~~~~~~~~ MW00032 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW00051 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~C~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW00058 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~C~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW00064 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW00072 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW00076 NNNTAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW00101 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW00102 NNATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW00110 NNATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~C~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW00116 NNATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC GTC~~~~GTC T~~~~~~~~~ ~~~~~~~~~~ MW00127 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW00129 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW00151 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW00176 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW00177 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW00178 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW00179 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GC~ ~~~~~~~~~~ ~~~~~~~~~~ MW00185 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW00189 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW00226 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~C~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW00231 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW00232 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~C~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW00234 NNNNNNNNNG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW00262 NNNTAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW00267 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW00269 T~ATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW00290 T~ATASTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW00302 T~ATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ATTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW00316 NNNNNNTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~C~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW00326 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW00328 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW00330 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW00347 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW00348 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW00393 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW00409 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATGTGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW00412 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW00444 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW00452 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW00469 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATACGCCAC ACTGTTCGTC GTC~~~~GTC GTCGTCT~~~ ~~~~~~~~~~ MW00498 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW00504 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW00517 NNNTAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC GT~~~~~AT~ ~~~~~~~~~~ ~~~~~~~~~~ MW00525 TYATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT AATATGCCAC ACTGTTCGTC ~T~~~~~~~~ ~~~~~~~~~~ ~~~~~~~~~~ MW00530 NNNNNNNNTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW00539 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~

185 Appendix 3: Sequence data Internal transcribed spacer 2 (ITS-2)

....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| 5 15 25 35 45 55 65 75 85 95 105 115 125 135 145 155 MW00547 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW01001 NNNNNNNGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW01011 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATACGCCAC ACTGTTCGTC GTC~~~~GTC GTCGTC~T~~ ~~~~~~~~~~ MW01014 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW01018 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATACGCCAC ACTGTTCGTC GTC~~~~GTC GTCGTC~T~~ ~~~~~~~~~~ MW01019 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ATTGTTCGTG ~CCGACTGT~ ~~~~~~~~~~ ~~~~~~~~~~ MW01025 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW01034 TC~TAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATACGCCAC ACTGTTCGTC GTC~~~~GTC GTCGTCGTC~ T~~~~~~~~~ MW01039 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW01048 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~~~~ ~~~~~~~~~~ ~~~~~~~~~~ MW01053 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW01059 NNNNNNTGCG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATACGCCAC ACTGTTCGTC GTC~~~~GTC GTCGTC~T~~ ~~~~~~~~~~ MW01061 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~~~~ ~~~~~~~~~~ ~~~~~~~~~~ MW01083 NNNNNNNNNN AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ATTGTTCGTC ~T~GAC~GTT ~~~~~~~~~~ ~~~~~~~~~~ MW01092 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATACGCCAC ACTGTTCGTC GTC~~~~GTC GTCGTC~T~~ ~~~~~~~~~~ MW01107 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW01116 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATACGCCAC ACTGTTCGTC GTC~~~~GTC GTCGTC~T~~ ~~~~~~~~~~ MW02001 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW02006 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW02025 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAC AATATGCCAC ACTGTTCGTC GTC~~~~~~~ ~~~~~~~~~~ ~~~~~~~~~~ MW02033 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~~~~ ~~~~~~~~~~ ~~~~~~~~~~ MW02034 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT AATATGCCAC ACTGTTCGTC ~TC~~~~~~~ ~~~~~~~~~~ ~~~~~~~~~~ MW98009 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW98029 NNNTAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW98049 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW98079 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATGTGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW98097 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGCC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW98103 TCATAGTGTA AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGG~CCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW98129 NNNNNNNGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATACGCCAC ACTGTTCGTC GTC~~~~GTC GTCGTCGTCG TCGTCGTCTC MW98136 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW98138 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW98139 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW98154 NNNTAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW98162 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW98170 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW98172 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW98180 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW98185 T~ATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATACGCCAC ACTGTTCGTC GTC~~~~GTC GTCGTCGTC~ TC~~~~~~~~ MW98189 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW98203 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW98205 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~C~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW98212 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW98220 TC~TAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~~~~~~~GTC T~~~~~~~~~ ~~~~~~~~~~ MW98228 TC~TAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATACGCCAC ACTGTTCGTC GTC~~~~GTC GTCGTCGTCG TCGTCGTCTC MW98235 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW98240 NNNNNNNNNG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~C~~~~~GC~ ~~~~~~~~~~ ~~~~~~~~~~ MW98261 NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNACATT TCSAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTMGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW98264 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTTT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW98270 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~~~~ ~~~~~~~~~~ ~~~~~~~~~~ MW98278 NNATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATACGCCAC ACTGTTCGTC GTC~~~~GTC GTCGTCN~~~ ~~~~~~~~~~ MW98285 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW98294 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~C~~~~~GC~ ~~~~~~~~~~ ~~~~~~~~~~ MW98313 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW98315 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW99001 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATGTGCCAC ACTGTTCGTC ~T~~~~~~~~ ~~~~~~~~~~ ~~~~~~~~~~ MW99006 T~ATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW99009 NC~TAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW99013 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~~~~ ~~~~~~~~~~ ~~~~~~~~~~ MW99014 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ATTGTTCGTG ~C~~~C~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW99018 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATGTGCCAC ACTGTTCGTC ~T~~~~~~~~ ~~~~~~~~~~ ~~~~~~~~~~ MW99038 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW99047 TMATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW99057 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW99058 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW99094 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW99095 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~C~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW99097 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~~~~ ~~~~~~~~~~ ~~~~~~~~~~ MW99105 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~

186 Appendix 3: Sequence data Internal transcribed spacer 2 (ITS-2)

....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| 5 15 25 35 45 55 65 75 85 95 105 115 125 135 145 155 MW99134 T~ATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~~~~ ~~~~~~~~~~ ~~~~~~~~~~ MW99135 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW99163 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~~~~ ~~~~~~~~~~ ~~~~~~~~~~ MW99164 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW99174 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW99196 TC~TAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW99203 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW99226 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW99240 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW99247 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW99274 NNNNNNNNNN AACTGCAGGA CACATTWGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW99276 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW99286 TCATAGNGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~C~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW99292 T~TTAGTGGG AACTGCAGGA CACATTTGAA CATCGACATT TCSAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~C~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW99301 T~ATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW99303 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TG~GTAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~~~~ ~~~~~~~~~~ ~~~~~~~~~~ MW99309 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW99319 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW99341 NNNNNNNNNN NNNNNNNNNA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~C~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW99353 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW99372 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW99374 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW99382 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW99393 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~C~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW99406 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~C~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW99407 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW99408 NNNNNNNNNG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW99413 TC~TAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW99448 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW99465 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW99471 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW99479 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~C~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW99501 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW99508 TC~TAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~C~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW99514 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATACGCCAC ACTGTTCGTC GTC~~~~GTC GTCGTCGTCG TC~~~~~~TC MW99536 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATACGCCAC ACTGTTCGTC GTC~~~~GTC GTCGTCGTCG TCT~~~~~~~ MW99537 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW99546 CTATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW99550 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW99552 TC~TAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW99559 NNNNNNNNNN NNNNNNNNNN NNNNNNNNNA CATCTACATT NCNAACGCNC ATTGCGGTCC GTGGAGAAAC ATCCANGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW99565 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW99591 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW99605 TCATAGTGTG AACTG~AGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ MW99608 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATACGCCAC ACTGTTCGTC GTC~~~~GTC GTCGTCT~~~ ~~~~~~~~~~ MW99612 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTTT GAGGGCCGGC TGTATAAAAT CATACGCCAC ACTGTTNGTN GTN~~~~GTG GTCGTC~T~~ ~~~~~~~~~~ MW99613 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ OK96022 NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTTT GAGGGCNGGC TGTATAAAAT CATACGCCAC ACTGTTGGTN GTN~~~~GTG GTCGTC~T~~ ~~~~~~~~~~ WE00002 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ WE02431 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ WE02451 T~ATAGTGTG MACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ WE02454 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ WE02491 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ WE02531 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ WE02534 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ WE02535 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ WE02591 NNNNNNTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~C~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ WE02614 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~C~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ WE02621 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ WE02661 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~C~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ WE02662 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ WE02671 T~ATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~C~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ WE02674 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ WE02677 TCATAGTGTG AACTGCAGAA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ WE85001 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAAT CATATGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~ WE94001 TCATAGTGTG AACTGCAGGA CACATTTGAA CATCGACATT TCGAACGCAC ATTGCGGTCC GTGGAGAAAC ATCCAGGACC ACTCCTGTCT GAGGGCCGGC TGTATAAAGT CATGTGCCAC ACTGTTCGTC ~T~~~~~GT~ ~~~~~~~~~~ ~~~~~~~~~~

187 Appendix 3: Sequence data Internal transcribed spacer 2 (ITS-2)

....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| 165 175 185 195 205 215 225 235 245 255 265 275 285 295 305 315 AD98001 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~ARA~ ~CTTA~~~TA AD98009 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGC~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA AD98012 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TGTGTG~~~~ TGGG~~TGCG ~~~~~~~TCC CTGCC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA AD98018 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATGAAGA~ ~CGTAT~~~A AD98024 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTGCC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTCAAAT~ ~~ATG~AGA~ ~CTTG~~~TA AD98036 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA DS00001 ~~~~~~~~~~ ~~~~~~~KCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA DS01001 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGCG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA JC00029 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GACGGT~GT ~~~~~~~~~~ CGCGCTTGTG TT~~~~~~~~ TGGG~~TACG TCACG~~TCC CTGCC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTCAAAT~ ~~ATG~AGA~ ~CATA~~~AA JC00039 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGT~~TACG T~AC~~~TGC ATACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATG~~~AA JC00040 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TGTG~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA JC00042 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GACGGT~GT ~~~~~~~~~~ CGCGCTTGTG TT~~~~~~~~ TGGG~~TACG TCACG~~TCC CTGCC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTCAAAT~ ~~ATG~AGA~ ~CATA~~~AA JC00043 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGGGGTGCG ~~~~~~~TCC CYACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA JC00045 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGGGGTGCG ~~~~~~~TCC CCACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA JC00051 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GACGGT~GT ~~~~~~~~~~ CGCGCTTGTG TT~~~~~~~~ TGGG~~TACG TCACG~~TCC CTGCC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTCAAAT~ ~~ATG~AGA~ ~CATA~~~AA JC00055 ~~~~~~~~~~ ~~~~~~CACG ACGAACAATT GACGGTTCCG ~GGCGGC~GT ~~~~~~~~~~ CGCGCGTGTG ~~~~~~~~~~ TGGG~~TTCG ~~~~~~~CCC ATGCC~~GCG C~~~CTCAC~ G~TGCGGTCC ~GTTYAAAA~ ~~ATG~AGA~ ~CATA~~~AA JC00061 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GACGGT~GT ~~~~~~~~~~ CGCGCTTGTG TT~~~~~~~~ TGGG~~TACG TCACG~~TCC CTGCC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTCAAAT~ ~~ATG~AGA~ ~CATA~~~AA JC00063 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GACGGT~GT ~~~~~~~~~~ CGCGCTTGTG TT~~~~~~~~ TGGG~~TACG TCACG~~TCC CTGCC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTCAAAT~ ~~ATG~AGA~ ~CATA~~~AA JC01014 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TGTG~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA JM00001 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~AA MW00015 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW00024 ~~~~~~~~~~ ~~~~~~CACG ACGAACAATT GACGGTTCCG ~GGAGGC~GT ~~~~~~~~CG CGCGCGTGCG ~~~~~~~~~~ TGGG~~CTCG ~~~~~~~TCC CCGCC~~GCG C~~~CACCC~ G~TGCGGTCC ~GTTCAAAA~ ~~ATG~ATA~ ~CATA~~~AA MW00032 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG GG~~~~~~~~ TGCG~~T~CC ~~~~~~~CCC CCACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CGTA~~~TA MW00051 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG CGACGGT~GT ~~~~~~~~~~ CGCGCTCGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CGTA~~~TA MW00058 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CGTA~~~TA MW00064 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW00072 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~CGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW00076 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TGTG~~~~~~ TGGG~~TACG ~~~~~~~TCC CTACC~~GCG C~~~CTCC~~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~AA MW00101 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TTCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~CGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW00102 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~CGCGGTCC ~GTTTAAAT~ ~~ATG~AAAA ~CATA~~~GA MW00110 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG CGACGGT~GT ~~~~~~~~~~ CGCGCTCGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CGTAT~~~A MW00116 ~~~~~~~~~~ ~~~CACGGCG ~CGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~AGCG CGCG~~TGTG ~~~~~~~~~~ TGGG~~TTAG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTCAAAA~ ~~ATG~AAA~ ~CATA~~~~A MW00127 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGG~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CCACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTCAAAT~ ~~ATG~AGA~ ~CGTGTTATA MW00129 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TGTG~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW00151 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~CGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW00176 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW00177 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TGTG~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW00178 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TGTG~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW00179 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TGTG~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW00185 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW00189 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTCGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW00226 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGGG~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~ATTA~~~TA MW00231 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TGTG~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW00232 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~ATA~ ~CGTA~~~TA MW00234 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW00262 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW00267 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~CGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW00269 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TGTG~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW00290 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TGTG~~~~~~ TGGG~~TACG ~~~~~~~TCC CTAMY~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AKA~ ~CATA~~~AA MW00302 ~~~~~~~~~~ ~~~~GGGACG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ AGCGCTTGTG TT~~~~~~~~ TGGATATACG ~~~~~TATCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTCAAAT~ ~~ATG~AGA~ ~CACA~~~AA MW00316 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG CGACGGT~GT ~~~~~~~~~~ CGCGCTCGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CGTAT~~~A MW00326 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TGTG~~~~~~ TGGG~~TACC ~~~~~~~TCC CTATC~~GCG C~~~CACCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~AA MW00328 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TGTG~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW00330 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TGTG~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW00347 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TTCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~CGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW00348 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCAG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~SA MW00393 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CCGCC~~GCG C~~~GTCCC~ G~CGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW00409 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~GTCCC~ G~CGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW00412 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GACGGT~GT ~~~~~~~~~~ CGCGCTTGTG TT~~~~~~~~ TGGG~~TACG TCACG~~TCC CTGCC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTCAAATA TTATG~AGA~ ~CATA~~~AA MW00444 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~CGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW00452 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW00469 ~~~~~~~~~~ ~~~CACGACG TCGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~AGAGCG CGCGCGTGTG TGTG~~~~~~ CGGG~~TTCG ~~~~~~~TCC GCACGC~GCG CCGCCTCCC~ G~TGCGGTCC ~GTTCAAAA~ ~~AT~~AGA~ ~CATA~~~AA MW00498 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW00504 ~~~~~~~~~~ ~~~~~~~GCG ACGGACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TACG ~~~~~~~TCC CTGCC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~AA MW00517 ~~~~~~~~~~ ~CGC~~~~CG ACGAACAATT GACGGTTCCG ~GGCGGC~GT ~~~~~~~~~~ CGCGCGTGTG ~~~~~~~~~~ TGGG~~TTCG ~~~~~~~TCC CTACC~~GCG C~~~GTCCC~ G~TGCGGTCC ~GTTTAAAA~ ~~ATG~AGA~ ~CATA~~~AA MW00525 ~~~~~~~~~~ ~~~~~~GCCG ACGAACAATT GACGGTTCCG ~GGCGGC~GT ~~~~~~~~CG CGCGCGTG~~ ~~~~~~~~~~ TGGG~~TTCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAA~ ~~ATG~AGA~ ~CATA~~~AA MW00530 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GACGGT~GT ~~~~~~~~~~ CGCGCTTGTG TT~~~~~~~~ TGGG~~TACG TCACG~~TCC CTGCC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTCAAATA TTATG~AGA~ ~CATA~~~AA MW00539 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TGTG~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTACC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA

188 Appendix 3: Sequence data Internal transcribed spacer 2 (ITS-2)

....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| 165 175 185 195 205 215 225 235 245 255 265 275 285 295 305 315 MW00547 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTNCC~ G~CGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW01001 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~AA MW01011 ~~~~~~~~~~ ~~~CACGACG TCGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~AGAGCG CGCGCGTGTG TGTG~~~~~~ CGGG~~TTCG ~~~~~~~TCC GCACGC~GCG CCGCCTCCC~ G~TGCGGTCC ~GTTCAAAA~ ~~AT~~AGA~ ~CATA~~~AA MW01014 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW01018 ~~~~~~CACG GCGCACGGCG TCGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~AGAGCG CGCGCGTGTG TGTG~~~~~~ CGGG~~TTCG ~~~~~~~TCC GCACGC~GCG CCGCCTCCC~ G~TGCGGTCC ~GTTCAAAA~ ~~AT~~AGA~ ~CATA~~~AA MW01019 ~~~~~~~~~~ ~~~~~~~GGG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TT~~~~~~~~ TGGGTGTACG T~ACGTATCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTCAAAT~ ~~ATG~AGA~ ~CATA~~~AA MW01025 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GACGGT~GT ~~~~~~~~~~ CGCGCTTGTG TT~~~~~~~~ TGGG~~TACG TCACK~~TCC CTGCC~~KCG C~~~CTCCC~ G~TGCGGTCC ~GTTCAAAT~ ~~ATG~AGA~ ~CATA~~~AA MW01034 ~~~~~~CACG GCGCACGGCG TCGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~AGAGCG CGCGCGTGTG TGTG~~~~~~ CGGG~~TTCG ~~~~~~~TCC GCACGC~GCG CCGCCTCCC~ G~TGCGGTCC ~GTTCAAAA~ ~~AT~~AGA~ ~CATA~~~AA MW01039 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~AA MW01048 ~~~~~~~~~~ ~~~~~~CACG ACGAACAATT GACGGTTCCG ~GGCGGC~GT ~~~~~~~~~~ CGCGCGTGTG ~~~~~~~~~~ TGGG~~TTCG ~~~~~~~CCC ATGCC~~GCG C~~~CTCAC~ G~TGCGGTCC ~GTTTAAAA~ ~~ATG~AGA~ ~CATA~~~AA MW01053 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~AA MW01059 ~~~~~~CACG GCGCACGGCG TCGAACAATT GACG~TTCCG ~GGCGGT~GT ~~~~AGAGCG CGCGCGTGTG TGTG~~~~~~ CGGG~~TTCG ~~~~~~~TCC GCACGC~GCG CCGCCTCCC~ G~TGCGGTCC ~GTTCAAAA~ ~~AT~~AGA~ ~CATA~~~AA MW01061 ~~~~~~~~~~ ~~~~~~AACG ACGAACAATT GACGGTTCCG ~GGCGGC~GT ~~~~~~~~~~ CGCGCGTGTG ~~~~~~~~~~ TGGG~~TTCG ~~~~~~~CCC ANGCC~~GCG C~~~CTCAC~ G~TGCGGTCC ~GTTTAAAA~ ~~ATG~AGA~ ~CATA~~~TA MW01083 ~~~~~~~~~~ ~~~~~CGACG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TT~~~~~~~~ TGGATATGCG ~~~~~TATCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTCAAAT~ ~~ATG~AGA~ ~CACA~~~AA MW01092 ~~~~~~CACG GCGCACGGCG TCGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~AGAGCG CGCGCGTGTG TGTG~~~~~~ CGGG~~TTCG ~~~~~~~TCC GCACGC~GCG CCGCCTCCC~ G~TGCGGTCC ~GTTCAAAA~ ~~AT~~AGA~ ~CATA~~~AA MW01107 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~AA MW01116 ~~~~~~CACG GCGCACGGCG TCGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~AGAGCG CGCGCGTGTG TGTG~~~~~~ CGGG~~TTCG ~~~~~~~TCC GCACGC~GCG CCGCCTCCC~ G~TGCGGTCC ~GTTCAAAA~ ~~AT~~AGA~ ~CATA~~~AA MW02001 ~~~~~~~~~~ ~~~~~~~GCG ACGGACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGT~~TACG ~~~~~~~TAC CTGCC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~AA MW02006 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TT~~~~~~~~ TGGG~~TACG ~~~~~~~TCC CTGCC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTCAAAT~ ~~ATG~AGA~ ~CATA~~~AA MW02025 ~~~~~~~~~~ ~~~~~~~~TG TCGAACAATT GACGGTTCCT ~GAC~~~~GT ~~~~~~~~~G CACG~~~~~~ ~~~~~~~~~~ TGGG~~CGCG ~~~~~~~CGC G~A~~GCGCG CCTCGTCTC~ G~AACGGTCC ~GTTCAAAA~ ~~ATT~~~~~ ~C~~~~~~AA MW02033 ~~~~~~~~~~ ~~~~~~CACG ACGAACAATT GACGGTTCCG ~GGCGGC~GT ~~~~~~~~~~ CGCGCGTGTG ~~~~~~~~~~ TGGG~~TTCG ~~~~~~~CCC ATGCC~~GCG C~~~CTCAC~ G~TGCGGTCC ~GTTTAAAA~ ~~ATG~AGA~ ~CATA~~~TA MW02034 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGC~GT ~~~~~~~~CG CGCGCGTG~~ ~~~~~~~~~~ TGGG~~TTCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAA~ ~~ATG~AGA~ ~CATA~~~AA MW98009 ~~~~~~~~~~ ~~~~~~~GCG ACGGACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTGCC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CTTA~~~TA MW98029 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TGTG~~~~~~ TGGG~~TACG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~AA MW98049 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW98079 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTCGTG TG~~~~~~~~ TGGGG~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW98097 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW98103 ~~~~~~~~~~ ~~~~~~~GCG ACKAACGATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW98129 ACGGCACACG GCGCAGGGCG TCGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~CGAGCG CGCGCGCGTG TGTG~~~~~~ CGGG~~TTCG ~~~~~~~TCC G~~~~~~~~~ ~~~~~~~~~~ ~~TGCGGTCC ~GTTCAAAA~ ~~AT~~AGA~ ~CATA~~~AA MW98136 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TGTGCG~~~~ TGGG~~TGCG ~~~~~~~TCC CTGCC~~GCG C~~~CTCCCC G~TGCGGTCC CGTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW98138 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW98139 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TGTGCG~~~~ TGGG~~TGCG ~~~~~~~TCC CTGCC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW98154 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATTA~~GA MW98162 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW98170 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TGTGTG~~~~ TGGG~~TGCG ~~~~~~~TCC CTGCC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATATA~TA MW98172 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TGTG~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~KA MW98180 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW98185 ACGGCGCACG GCGCACGGCG TCGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~CGAGCG CGCGCGTGTG TGTG~~~~~~ CGGG~~TTCG ~~~~~~~TCC G~~~~~~~~~ ~~~~~~~~~~ ~~TGCGGTCC ~GTTCAAAA~ ~~AT~~AGA~ ~CATA~~~AA MW98189 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACTGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTCGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW98203 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TGTG~~~~~~ TGGG~~TGCG ~~~~~~~CCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTCAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW98205 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCCTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CTTA~~~TA MW98212 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TGTG~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW98220 ~~~~~~~~~~ ~~~CACGGCG GCGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~CG CGCG~~TGCG TGTG~~~~~~ TGGG~~TTCG ~~~~~~~TCC CTGCCGCGCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAA~ ~~ATG~AAA~ ~CATA~~~AA MW98228 ACGGCGCACG GCGCACGGCG TCGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~CGAGCG CGCGCGTGTG TGTG~~~~~~ CGGG~~TTCG ~~~~~~~TCC GCACGC~GCG CCGCCTCCC~ G~TGCGGTCC ~GTTCAAAA~ ~~AT~~AGA~ ~CATA~~~GA MW98235 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ ~~~~~~TA~~ ~~~~~~~~~~ ~~~~~~~~~~ ~~~~~~CCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~AA MW98240 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CGTA~~~TG MW98261 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC CGTTTAAAT~ ~~ATG~AGAA CTTTG~~~TA MW98264 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ YSCGCTTGTG TG~~~~~~~~ ~~~~~~TA~~ ~~~~~~~~~~ ~~~~~~~~~~ ~~~~~~CCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~AA MW98270 ~~~~~~~~~~ ~~~~~~CGCG GCGAACAACT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCGTGTG ~~~~~~~~~~ TGGG~~TTCG ~~~~~~~TCC CTGCC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAA~ ~~ATG~AGA~ ~CATA~~~AA MW98278 ~~~~~~~~~~ ~~~NNNNNNN NNNNNNNNNN NACGGTTCCG ~GGCGGT~GT ~~~~AGAGCG CGMGCGTGTG TGTA~~~~~~ CGGG~~TTCG ~~~~~~~TCC GCACGC~GCG CCGCCTCCC~ G~TGCGGTCC ~GTTCAAAA~ ~~AT~~AGA~ ~CATA~~~AA MW98285 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TGTG~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW98294 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CGTA~~~TA MW98313 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TGTGTG~~~~ TGGG~~TGCG ~~~~~~~TCC CTGCC~CGCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATR~AGA~ ~CACATA~KA MW98315 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TGTG~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW99001 ~~~~~~~~~~ ~~~~~~CGCG ACGAACAATT GACGGTTCCG ~GGCGGC~GT ~~~~~~~~~~ CGCGCGTGTG ~~~~~~~~~~ TGGG~~TTCG ~~~~~~~CCC CTGCC~~GCG C~~~CTCGC~ G~TGCGGTCC ~GTTTAAAA~ ~~ATG~AGA~ ~CATA~~~AA MW99006 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TGTGTG~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW99009 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW99013 ~~~~~~~~~~ ~~~~~~AGCG ACGAACAATT GACTGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TACG ~~~~~~~TCC CTACCGCGCG C~~~CTACC~ G~TGCGGTCA ~GTTTAAATA TTGTG~AGA~ ~CATA~~~AA MW99014 ~~~~~~~~~~ ~~~~~~~GGG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TT~~~~~~~~ TGGGTGTACG ~~~~~TATCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTCAAAT~ ~~ATG~AGA~ ~CATA~~~AA MW99018 ~~~~~~~~~~ ~~~~~~AACG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCGTGCG ~~~~~~~~~~ GGGG~~TTCG ~~~~~~~TCC CTCCC~~GCG C~~~TTCCC~ G~TGCGGTCC ~GTTTAAAA~ ~~ATG~AGA~ ~CATA~~~AA MW99038 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ ~~~~~~TA~~ ~~~~~~~~~~ ~~~~~~~~~~ ~~~~~~CCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~AA MW99047 ~~~~~~~~~~ ~~~~~~~GCG ACGGACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TGC~~~~~~~ AGGT~~TACG ~~~~~~~TAC CTGCC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~AA MW99057 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TGTGTG~~~~ TGGG~~TGCG ~~~~~~~TGC CTGCC~~KCG C~~~CTCCC~ G~TGCGGKCC ~GTKTAAAT~ ~~ATG~ANA~ ~CATA~~~SA MW99058 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW99094 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TGTGTGTGTG TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW99095 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTGCC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTCAAAT~ ~~ATG~AGA~ ~CTTG~~~TA MW99097 ~~~~~~~~~~ ~~~~~~CACG ACGAACAATT GACGGTTCCG ~GGCGGC~GT ~~~~~~~~~~ CGCGCGTGTG ~~~~~~~~~~ TGGG~~TTCG ~~~~~~~CCC CTGCC~~GCG C~~~CTCGC~ G~TGCGGTCC ~GTTTAAAA~ ~~ATG~AGA~ ~CATA~~~AA MW99105 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTCGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA

189 Appendix 3: Sequence data Internal transcribed spacer 2 (ITS-2)

....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| 165 175 185 195 205 215 225 235 245 255 265 275 285 295 305 315 MW99134 ~~~~~~~~~~ ~~~~~~GACG ACGAACAATT GACGGTTCCG ~GRCGGT~GC ~~~~~~AGCG CGCGCGCGNN ~~~~~~~~~~ NNNN~~NNNN ~~~~~~~NNN NNNNN~~NNN N~~~NNNNN~ N~NNNNNNNN ~NNNNNNNN~ ~~NNN~NNN~ ~NNNN~~~NN MW99135 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~TG ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW99163 ~~~~~~~~~~ ~~~~~~CACG ACGAACAATT GACGGCTCCG ~GGCGGTTGT ~~~~~~~~~~ CGCGCGTGTG ~~~~~~~~~~ TGGG~~TTCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAA~ ~~ATG~AGA~ ~CATA~~~AA MW99164 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW99174 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW99196 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTCGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW99203 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~TG ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW99226 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGG~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CCACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTCAAAT~ ~~ATG~AGA~ ~CGTA~~~TA MW99240 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TGTGTG~~~~ TGGG~~TGCG ~~~~~~~TCC CTGCC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW99247 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW99274 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW99276 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW99286 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGGG~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CTTA~~~TA MW99292 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTGCC~~GSC C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CTTATAATA MW99301 ~~~~~~~~~~ ~~~~~~~GCG ACGGACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TGTG~~~~~~ TGGG~~TACR ~~~~~~~YCC CTGYC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~AA MW99303 ~~~~~~~~~~ ~~~~~~CACG ACGAACAATT GACGGCTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCGTGTG ~~~~~~~~~~ TGGG~~TTCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAA~ ~~ATG~AGA~ ~CATA~~~AA MW99309 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACKGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW99319 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW99341 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTGCC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~ANAC TCATAA~~TA MW99353 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TGTGTG~~~~ TGGG~~TGCG ~~~~~~~TCC CTGCC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW99372 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW99374 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGG~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CCACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTCAAAT~ ~~ATG~AGA~ ~CGTA~~~TA MW99382 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW99393 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGGG~TGCG ~~~~~~~TCC YTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~ATTA~~~TA MW99406 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGGG~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~ATTA~~~TA MW99407 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTGAAT~ ~~ATG~AGA~ ~CATA~~~GA MW99408 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW99413 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW99448 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TGTGTG~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW99465 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TGTGTG~~~~ TGGG~~TGCG ~~~~~~~TCC CTGCC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW99471 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TGTG~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW99479 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CGTA~~~TA MW99501 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW99508 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CGTA~~~TA MW99514 ACGGCGCACG GCGCACAGCG TYTAACAATT GACGGTTCCG ~GCCRGT~GT AYAGMGCGCG CGCGCGTGTG TGTG~~~~~~ CGGG~~TTCG ~~~~~~~TCC GCACGC~GCG CCGCCTCCC~ G~TGCGGTCC ~GTTCAAAA~ ~~AT~~AGA~ ~CATA~~~AA MW99536 ~~~~~~CACG GCGCACGGCG TCGAACAATT GACGGTTCCG ~GCCGGT~GT ~~~~AGAGCG CGCGCGCGTG TGTGTG~~~~ CGGG~~TTCG ~~~~~~~TCC GCACGC~GCG CCGCCTCCC~ G~TGCGGTCC ~GTTCAAAA~ ~~AT~~AGA~ ~CATA~~~AA MW99537 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ ~~~~~~TA~~ ~~~~~~~~~~ ~~~~~~~~~~ ~~~~~~CCC~ GGTGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~AA MW99546 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TACG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW99550 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ ~~~~~~TA~~ ~~~~~~~~~~ ~~~~~~~~~~ ~~~~~~CCC~ GGTGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~AA MW99552 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTGCC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTCAAAT~ ~~ATG~AGA~ ~CTTG~~~TA MW99559 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW99565 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW99591 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TGTG~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW99605 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT RACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TGTG~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA MW99608 ~~~~~~~~~~ ~~~CACGACG TCGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~AGAGCG CGCGCGTGTG TGTG~~~~~~ CGGG~~TTCG ~~~~~~~TCC GCACGCGGCG CCGCCTCCC~ G~TGCGGTCC ~GTTCAAAA~ ~~AT~~AGA~ ~CATA~~~AA MW99612 ~~~~~~CACG GCGCACGGCG TCGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~AGAGCG CGCGCGTGTG TGTG~~~~~~ CGGG~~TTCG ~~~~~~~TCC GCACGC~GCG CCGCCTCCC~ G~TGCGGTCC ~GTTCAAAA~ ~~AT~~AGA~ ~CATA~~~AA MW99613 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TACG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA OK96022 ~~~~~~CACG GCGCACGGCG TCGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~AGAGCG CGCGCGTGTG TGTG~~~~~~ CGGG~~TTCG ~~~~~~~TCC GCACGC~GCG CCGCCTCCC~ G~TGCGGTCC ~GTTCAAAA~ ~~AT~~AGA~ ~CATA~~~AA WE00002 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA WE02431 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA WE02451 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA WE02454 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TGTGTGTG~~ TGGG~~TGCG ~~~~~~~TCC CTGCC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA WE02491 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA WE02531 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGG~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CCACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTCAAAT~ ~~ATG~AGA~ ~CGTA~~~TA WE02534 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~G TGCGCACGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC YTACC~~GCG C~~~YTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA WE02535 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTCGTG TG~~~~~~~~ TGGG~~CGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA WE02591 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAAYT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CTTA~~~TA WE02614 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGGG~TGCG ~~~~~~~TCC YTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~ATTA~~~TA WE02621 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TGTG~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CAYA~~~GA WE02661 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CGTA~~~TA WE02662 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA WE02671 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGGG~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~ATTA~~~TA WE02674 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TGTG~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA WE02677 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA WE85001 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA WE94001 ~~~~~~~~~~ ~~~~~~~GCG ACGAACAATT GACGGTTCCG ~GGCGGT~GT ~~~~~~~~~~ CGCGCTTGTG TG~~~~~~~~ TGGG~~TGCG ~~~~~~~TCC CTACC~~GCG C~~~CTCCC~ G~TGCGGTCC ~GTTTAAAT~ ~~ATG~AGA~ ~CATA~~~GA

190 Appendix 3: Sequence data Internal transcribed spacer 2 (ITS-2)

....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| 325 335 345 355 365 375 385 395 405 415 425 435 445 455 465 475 AD98001 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAG~~~~~~~ ~TTTG~~~CG AGAGCGATTA AD98009 T~GT~~~~~~ ~AATA~~~TG A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGACGGC GACGCGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA AD98012 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGCGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAG~~~~~~~ ~TTTG~~~CG AGAGCGATTA AD98018 T~GT~~~~~~ ~AATATTATA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTAG~~~GA MCAGNNNNNN AD98024 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TYGTGT~~~~ ~GGTGTG~~~ ~~~~TGC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA AD98036 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA DS00001 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA DS01001 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA JC00029 T~G~~~~~~~ TAATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA JC00039 T~G~~~~~~~ TAATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAG AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA JC00040 T~GT~~~~~~ TAATA~~~TG A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAG AAA~~~~~~~ ~TTTA~~~CG AGAGCGATTA JC00042 T~G~~~~~~~ TAATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA JC00043 T~GT~~~~~~ TAATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTMGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA JC00045 T~GT~~~~~~ TAATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA JC00051 T~G~~~~~~~ TAATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA JC00055 T~G~~~~~~~ TAATA~~~TA A~~~~~~~CG ~TTA~~CGTG TTGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCAC CGACGCTCCG TGAAGGTAAA AAA~~~~~~~ ~TTTG~~~CG AGAGTGATTA JC00061 T~G~~~~~~~ TAATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGC~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA JC00063 T~G~~~~~~~ TAATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGC~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA JC01014 T~GT~~~~~~ TAATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA JM00001 T~GT~~~~~~ TAATA~~~TG A~~~~~~~CG ~TCA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTA~~~CG AGAGCGATTA MW00015 T~GT~~~~~~ TAATA~~~TA A~~~~~~~CG ~TTG~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW00024 T~GTTATTAA TAATA~~~TG A~~~~~~~CG ~TTA~~AGTG TTGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCAC CGACGCTCCG TGAAGGTGAA AAG~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW00032 C~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTG~~CGTG TCGTGT~~~~ ~GGTCTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAG~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW00051 T~GT~~~~~~ ~AATATTATA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAAA~~~~~~ ~TTTG~~~CG AGAGCGATTA MW00058 T~GT~~~~~~ ~AATATTATA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW00064 T~GT~~~~~~ TAATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AGA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW00072 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW00076 T~T~~~~~~~ TAATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~GTAC~~~ GGTAGCGCGC TCGCGTCGGC GACRTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW00101 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW00102 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW00110 T~GT~~~~~~ ~AATATTATA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~NN NNNNNNNNNN MW00116 T~A~~~~~~~ AAANG~~~TA ATG~T~AACG CTTC~ACGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGAC GRCGTGCGCC GCT~CCCCAC CGACGCTCCG TGAAGGTGAG AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW00127 T~GT~~~~~~ ~AATA~~~TG A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTGGCGCGC TCGCGTYGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG ASAGCGATTA MW00129 T~GT~~~~~~ ~AATA~~~TG A~~~~~~~CG ~TTA~~CGTG TCGTGC~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGCGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW00151 T~GT~~~~~~ ~AATA~~~TG A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGACGGC GACGCGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW00176 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW00177 T~GT~~~~~~ ~AATA~~~TG A~~~~~~~CG ~TTA~~CGTG TCGTGC~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGCGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW00178 T~G~~~~~~~ TAATA~~~TG A~~~~~~~CG ~TTA~~CGTG TCGTGC~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGCGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW00179 T~GT~~~~~~ GATTA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGCGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW00185 T~GT~~~~~~ TAATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW00189 T~GT~~~~~~ TAATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW00226 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TACCAC GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW00231 T~GT~~~~~~ TAATA~~~TG A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTA~~~CG AGAGCGATTA MW00232 T~G~~~~~~~ TAATATTATA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW00234 T~GT~~~~~~ ~AATA~~~TG A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGACGGC GACGCGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW00262 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGCGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW00267 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW00269 T~GT~~~~~~ ~AATAT~ATA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG NGAAGGTNNN NNN~~~~~~~ ~NNNN~~~NN NNNNNNNNNN MW00290 T~T~~~~~~~ TAATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW00302 T~G~~~~~~~ TTATTT~~TA AC~~~~~~CG ~TTN~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW00316 T~GT~~~~~~ ~RATATTATA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~NN NNNNNNNNNN MW00326 T~G~~~~~~~ TAATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGT TCGCGTCGGC GACGTGCGCC GCT~CCCAGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW00328 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW00330 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~CCGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGCGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW00347 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAG AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW00348 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC SACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW00393 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW00409 T~GTT~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCCCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW00412 T~G~~~~~~~ TAATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW00444 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW00452 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW00469 T~G~~~~~~~ TAATA~~~TA A~~~~~~~CG ~TTA~~CGTG TTGTGT~~~~ ~GGTGTG~~~ TGTGTAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCAC CGACGCTCCG TGAAGGTGAA AAG~~~~~~~ TTTTG~~~CG AGAGCGATTA MW00498 T~GT~~~~~~ TAATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AGA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW00504 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTKAA AAA~~~~~~~ ~TTTG~~~CG AGARCGATTA MW00517 T~G~~~~~~~ CAATA~~~TA A~~~~~~~CG ~TTA~~CGTG TTGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCAC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW00525 T~G~~~~~~~ TAATA~~~TA A~~~~~~~CG ~TTA~~CGTG TTGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCAC CGACGCTCCG TGAACGTGAA AAA~~~~~~~ ~CTTG~~~CG AGAGCGATTA MW00530 T~G~~~~~~~ TAATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW00539 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA

191 Appendix 3: Sequence data Internal transcribed spacer 2 (ITS-2)

....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| 325 335 345 355 365 375 385 395 405 415 425 435 445 455 465 475 MW00547 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW01001 T~GT~~~~~~ TAATA~~~TG A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTC~~ ~~~AGGTNNN NNN~~~~~~~ ~NNNN~~~NN NNNNNNNNNN MW01011 T~G~~~~~~~ TAATA~~~TA A~~~~~~~CG ~TTA~~CGTG TTGTGT~~~~ ~GGTGTG~~~ TGTGTAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCAC CGACGCTCCG TGAAGGTGAA AAG~~~~~~~ TTTTG~~~CG AGAGCGATTA MW01014 T~GT~~~~~~ TAATA~~~TA A~~~~~~~CG ~TTA~~AGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW01018 T~A~~~~~~~ TAATA~~~TA A~~~~~~~CG ~TTA~~CGTG TTGTGT~~~~ ~GGTGTGTG~ TGTGTAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCAC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW01019 T~G~~~~~~~ TAATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGGA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW01025 T~G~~~~~~~ TAATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGC~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTACGC GACGTGCGCC GCT~CCCCGC CKACGCTCCG NGAAGGTGAA AAA~~~~~~~ ~TNNN~~~NN NNNNNNNNNN MW01034 T~A~~~~~~~ TAATA~~~TA A~~~~~~~CG ~TTA~~CGTG TTGTGT~~~~ ~GGTGTG~~~ TGTGTAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCAC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW01039 T~GT~~~~~~ TAATA~~~TG A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTA~~~CG AGAGCGATTA MW01048 T~G~~~~~~~ TAATA~~~TA A~~~~~~~CG ~TTA~~CGTG TTGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCAC CGACGCTCCG TGAAGGTAAA AAA~~~~~~~ ~TTTG~~~CG AGAGTGATTA MW01053 T~GT~~~~~~ TAATA~~~TG A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTA~~~CG AGAGCGATTA MW01059 T~A~~~~~~~ TAATA~~~TA A~~~~~~~CG ~TTA~~CGTG TTGTGT~~~~ ~GGTGTG~~~ TGTGTAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCAC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW01061 T~G~~~~~~~ TAATA~~~TA A~~~~~~~CG ~TTA~~CGTG TTGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCAC CGACGCTCCG TGAAGGTAAA AAA~~~~~~~ ~TTTG~~~CG AGAGTGATTA MW01083 T~G~~~~~~~ TAATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW01092 T~A~~~~~~~ TAATA~~~TA A~~~~~~~CG ~TTA~~CGTG TTGTGT~~~~ ~GGTGTG~~~ TGTGTAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCAC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW01107 T~GT~~~~~~ TAATA~~~TG A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTA~~~CG AGAGCGATTA MW01116 T~A~~~~~~~ TAATA~~~TA A~~~~~~~CG ~TTA~~CGTG TTGTGT~~~~ ~GGTGTG~~~ TGTGTAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCAC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW02001 T~G~~~~~~~ TAATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW02006 T~G~~~~~~~ TAATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW02025 T~GT~~~~~~ TA~~~~~~~~ ~~~~~~~~~~ ~~~~~~~~~~ ~~~~~T~~~~ ~GGTGTGTG~ ~~~~~~C~~~ ~CGAGCGCGC TCTCGTTAGC GACGTGCGCC CCG~CC~~GC CGACGCTCCG TGAAGTTGTG ACTATAACAC TTGTG~~~CG AGAGCGATTA MW02033 T~G~~~~~~~ TAATA~~~TA A~~~~~~~CG ~TTA~~CGTG TTGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCAC CGACGCTCCG TGAAGGTAAA AAA~~~~~~~ ~TTTG~~~CG AGAGTGATTA MW02034 T~G~~~~~~T TAATA~~~TA A~~~~~~~CG ~TTA~~CGTG TTGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCAC CGACGCTCCG TGAAGGTGAA AAAC~~~~~~ ~TTTGAGACG AGAGCGATTA MW98009 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAG~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW98029 T~T~~~~~~~ TAATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~GTAC~~~ GGTAGCSCGC TCGCGTCGGC GACGTGCGCC GYT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW98049 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW98079 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGCGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW98097 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGCGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW98103 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW98129 T~A~~~~~~~ TAATA~~~TA A~~~~~~~CG ~TTA~~CGTG TTGTGT~~~~ ~GGTGTGTG~ TGTGTAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCTCAC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW98136 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGCGCGCC GCT~CCCCGC CGACGCTCCG TGARGGTGAA AAG~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW98138 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGCGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW98139 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TRC~~~ GGTAGCGCGC TCGCGTCGGC GACGCGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAG~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW98154 ATGT~~~~~~ TAATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGCGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW98162 T~GT~~~~~~ ~AATA~~~TG A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGACGGC GACGCGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW98170 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTSGGA SACGCGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAG~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW98172 T~GT~~~~~~ TAATA~~~TG A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC SGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTA~~~CG AGAGCGATTA MW98180 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGCGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW98185 T~A~~~~~~~ TAATA~~~TA A~~~~~~~CG ~TTA~~CGTG TTGTGT~~~~ ~GGTGTGTG~ TGTGTAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCAC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW98189 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGCGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW98203 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGCGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGGA AAA~~~~~~~ ~TTTG~~~CG WGAGCGATTA MW98205 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TACCGC GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAAAAA~~~~ ~TTTG~~~CG AGAGCGATTA MW98212 T~GT~~~~~~ TAATA~~~TG A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTA~~~CG AGAGCGATTA MW98220 TTGT~~~~~~ TAATA~~~TG A~~~~~~~CG ~TTA~~CGTG TTGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCAC CGACGCTCCG TGAAGGTAAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW98228 T~A~~~~~~~ TGATA~~~TA A~~~~~~~CG ~TTA~~CGTG TTGTGT~~~~ ~GGTGTG~~~ TGTGTAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCAC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW98235 T~G~~~~~~~ TAATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGT TCGCGTCGGC GACGTGCGCC GCT~CCCAGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW98240 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW98261 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGCAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAG~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW98264 T~G~~~~~~~ TAATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGT TCGCGTCGGC GACGTGCGCC GCT~CCCAGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW98270 T~T~~~~~~~ TAATA~~~TA A~~~~~~~CG ~TTACACGTG TTGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCAC CGACGCTCCG TGAAGGTAAA AAA~~~~~~~ ~TTTG~~~CG AGAGTGATTA MW98278 T~G~~~~~~~ TAATA~~~TA A~~~~~~~CG ~TTA~~CGTG TTGTGT~~~~ ~GGTGTG~~~ TGTGTAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCAC CGACGCTCCG TGAAGGTGAA AAG~~~~~~~ TTTTG~~~CG AGAGCGATTA MW98285 T~GT~~~~~~ ~AATA~~~TG A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGCGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW98294 T~GT~~~~~~ ~AATG~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AGA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW98313 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGM GACGCGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAG~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW98315 T~GTT~~~~~ ~AATA~~~TG A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTA~~~CG AGAGCGATTA MW99001 TTG~~~~~~~ TAATA~~~TA A~~~~~~~CG ~TTA~~CGTG TTGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCAC CGACGCTCCG TGAAGGTAAA AAA~~~~~~~ ~TTTG~~~CG AGAGTGATTA MW99006 T~GT~~~~~~ ~AATA~~~TG A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGCGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW99009 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW99013 T~G~~~~~TG TAATA~~~TA A~~~~~~~CG ~TTA~~CGTG TTGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGT TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTAAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW99014 T~G~~~~~~~ TAATA~~~TA G~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW99018 T~G~~~~TAA TAATA~~~TA A~~~~~~~CG ~TTA~~CGTG TTGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCAC CGACGCTCCG TGAAGGTAAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW99038 T~G~~~~~~~ TAATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGT TCGCGTCGGC GACGTGCGCC GCT~CCCAGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW99047 T~G~~~~~~~ TAATA~~~TA A~~~~~~CGT ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW99057 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CK ~TTA~~CGTG YC~TGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGSKCGC TCGCNTTNGT GACGCRCGCC GCT~CCCYGC CGACGCTCCG TGAAGGTGAA AAG~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW99058 T~GT~~~~~~ ~AATA~~~TG A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGACGGC GACGCGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW99094 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGCGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW99095 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TGC~~~ GGTAGCGCGC TCGCGTCGGC GACGCGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW99097 TTG~~~~~~~ TAATA~~~TA A~~~~~~~CG ~TTA~~CGTG TTGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCAC CGACGCTCCG TGAAGGTAAA AAA~~~~~~~ ~TTTG~~~CG AGAGTGATTA MW99105 T~GT~~~~~~ TAATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA

192 Appendix 3: Sequence data Internal transcribed spacer 2 (ITS-2)

....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| 325 335 345 355 365 375 385 395 405 415 425 435 445 455 465 475 MW99134 T~T~~~~~~~ TAATA~~~TA A~~~~~~~CG ~TTA~~CGTG TTGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGMGTCGAC RACGTGCGCC GCT~CCCCAC CGACGMTCCG TGAAGGTAAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW99135 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW99163 T~G~~~~~~~ TAATA~~~TA AMCKTWAACG ~TTA~~CGTG TTGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCCACCCCAC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW99164 T~GT~~~~~~ TAATA~~~TG A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTA~~~CG AGAGCGATTA MW99174 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW99196 T~GT~~~~~~ TAATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW99203 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW99226 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGTCGTG TGGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTMGGN GACGTGCGCC GCT~CCCCGC MGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCKATTA MW99240 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGCGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAG~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW99247 T~GT~~~~~~ ~AATA~~~TG A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGACGGC GACGCGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW99274 T~GT~~~~~~ TAATA~~~TG A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCKSC KACGTGCRCC GCT~CCCCGC MGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TKTA~~~CG AGAGCGATTA MW99276 T~GT~~~~~~ TAATA~~~TG A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTA~~~CG AGAGCGATTA MW99286 T~GT~~~~~~ ~AATA~~~T~ ~~~~~~~~~G ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TACCAC GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW99292 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCNN NNNNNNNGAA AAA~~~~~~~ ~NNNN~~~NN NNNNNNNNNN MW99301 T~G~~~~~~~ TAATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW99303 T~G~~~~~~~ TAATA~~~TA A~~~~~~~CG ~TTA~~CGTG TTGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCCACCCCAC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW99309 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW99319 T~GT~~~~~~ TAATA~~~TG A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTA~~~CG AGAGCGATTA MW99341 T~GT~~~~~~ ~AATA~TATA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW99353 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGCGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAG~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW99372 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW99374 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCCTCGGC WACGTGCGCC GCT~CCCCTC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW99382 T~GT~~~~~~ TAATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW99393 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGC~~~~ ~GGTGTG~~~ ~~~~TACCGC GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW99406 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TACCGC GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW99407 T~GT~~~~~~ TAATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AGA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW99408 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGCGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW99413 T~GT~~~~~~ ~AATA~~~TG A~~~~~~~CN ~TTN~~CNTG TCTGGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGACGGC GACGCGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW99448 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGCGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW99465 T~G~~~~~~~ TAATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGCGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAG~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW99471 T~GTT~~~~~ ~AATA~~~TG A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTA~~~CG AGAGCGATTA MW99479 T~GT~~~~~~ ~AATATTATA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW99501 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTACCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW99508 T~GT~~~~~~ ~AATATTATA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW99514 T~A~~~~~~~ TAATA~~~TA A~~~~~~~CG ~TTA~~CGTG TTGTGT~~~~ ~GGTGTG~~~ TGTGTAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCAC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW99536 T~A~~~~~~~ TAATA~~~TA A~~~~~~~CG ~TTA~~CGTG TTGTGT~~~~ ~GGTGTG~~~ TGTGTAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCAC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW99537 T~G~~~~~~~ TAATAATATA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGT TCGCGTCGGC GACGTGCGCC GCT~CCCAGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW99546 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW99550 T~G~~~~~~~ TAATAATATA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGT TCGCGTCGGC GACGTGCGCC GCT~CCCAGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW99552 TTGT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TGC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW99559 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW99565 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW99591 T~GT~~~~~~ TAATA~~~TG A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAG AAA~~~~~~~ ~TTTA~~~CG AGAGCGATTA MW99605 T~GT~~~~~~ TAATA~~~TG A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAG AAA~~~~~~~ ~TTTA~~~CG AGAGCGATTA MW99608 T~G~~~~~~~ TAATA~~~TA A~~~~~~~CG ~TTA~~CGTG TTGTGT~~~~ ~GGTGTG~~~ TGTGTAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCAC CGACGCTCCG TGAAGGTGAA AAG~~~~~~~ TTTTG~~~CG AGAGCGATTA MW99612 T~A~~~~~~~ TAATA~~~TA A~~~~~~~CG ~TTA~~CGTG TTGTGT~~~~ ~GGTGTG~~~ TGTGTAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCAC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA MW99613 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA OK96022 T~A~~~~~~~ TAATA~~~TA A~~~~~~~CG ~TTA~~CGTG TTGTGT~~~~ ~GGTGTG~~~ TGTGTAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCAC CGACGCTCCG TGAAGGTGAG AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA WE00002 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA WE02431 T~GT~~~~~~ TAATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA WE02451 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG T~AAGGTAAA AAT~~~~~~~ ~TTTG~~~CG AGAGCNATTA WE02454 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGCGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGGA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA WE02491 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACKCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA WE02531 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TGC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA WE02534 T~GT~~~~~~ TAATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGM AACGTGCNCC GYT~CCCCGC CGACGYTCCG T~AAGGNKAA AAA~~~~~~~ ~TTTG~~~CR AGAGCRATTA WE02535 T~GT~~~~~~ TAATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA WE02591 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TACCAC GGGAACNCGC TCGCGTNGGC GACGCGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA WE02614 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TYA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TACCAC GGWAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTYCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA WE02621 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGCGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGGA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA WE02661 T~GT~~~~~~ ~GATATTATA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA WE02662 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA WE02671 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TACCAC GGTAACCCGC TCGCGTCGGC GACGTGCGCC GNT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA WE02674 T~GT~~~~~~ ~AATA~~~TG A~~~~~~~CG ~TTA~~CGTG TCGTGC~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGCGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA WE02677 T~GT~~~~~~ TAATA~~~TA A~~~~~~~CG ~TTR~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA WE85001 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA WE94001 T~GT~~~~~~ ~AATA~~~TA A~~~~~~~CG ~TTA~~CGTG TCGTGT~~~~ ~GGTGTG~~~ ~~~~TAC~~~ GGTAGCGCGC TCGCGTCGGC GACGTGCGCC GCT~CCCCGC CGACGCTCCG TGAAGGTGAA AAA~~~~~~~ ~TTTG~~~CG AGAGCGATTA

193 Appendix 3: Sequence data Internal transcribed spacer 2 (ITS-2)

....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| 485 495 505 515 525 535 545 555 565 575 585 595 605 615 625 635 AD98001 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGC ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG AAT~~~~AAA A~~NN~~~NN NNNN~~~~NN N~NNNNNNNN N~~~NNNN~~ AD98009 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ AD98012 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CGCGTTTCTT TT~~~~~~~~ ~~~~~~AACG AA~~~~AGAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~A AD98018 NNNNNNNN~~ NNNNNNNNNN NNNN~~NN~~ ~NN~NNNN~N N~NNNN~NNN ~NNNNNNNN~ ~NNNNNNNN~ NNNNN~~~NN NNNNTTTTTT T~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ AD98024 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CKAA~C T~GTGT~CGT ~CKGCGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT GT~~~~~~~~ ~~~~~~AACG AA~~~~~AAA G~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ AD98036 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ DS00001 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT TTT~~~~~~~ ~~~~~~GACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC A~AAAACACT G~~~TTAT~~ DS01001 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CKAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTCTTTT T~~~~~~~~~ ~~~~~~GACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTATTA JC00029 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~TGTA CACTCTTTT~ CCT~~~~~~~ ~~~~~~AACG AA~~~~GAAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ JC00039 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT T~~~~~~~~~ ~~~~~~AACG AT~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ JC00040 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ JC00042 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~TGTA CACTCTTTT~ CCT~~~~~~~ ~~~~~~AACG AA~~~~GAAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ JC00043 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CRAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ JC00045 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ JC00051 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCGGTGTA CACTCTTTT~ CCT~~~~~~~ ~~~~~~AACG AA~~~~GAAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ JC00055 TCGGTGTG~~ CGGGGCGTGC ACGC~~AC~~ ~GT~CGAA~C T~GTGT~CGT ~CKGCGCGC~ ~GTGATGCG~ CTGCG~~~TA ~~CGTT~~~~ ~~~~~~~~~~ ~~~~~~~~CG AA~~~CGAAA A~~CA~~~CC GCCG~~~~TC G~AAAACACT G~~~TTAT~~ JC00061 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~TGTA CACTCTTTTT CCT~~~~~~~ ~~~~~~AACG AA~~~~GAAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ JC00063 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~TGTA CACTCTTTTT CCT~~~~~~~ ~~~~~~AACG AA~~~~GAAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ JC01014 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CKAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~TGTTAT~~ JM00001 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW00015 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTT~ C~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW00024 TCGGTGAG~~ CGGGGCGTGN NNNN~~NN~~ ~NN~NNNN~N N~NNNN~NNN ~NNNNNNNN~ ~NNNNNNNN~ NNNNN~~~NN ~~NNNN~~~~ ~~~~~~~~~~ ~~~~TGTACG AGAGT~~~~~ A~~CA~~~CC ACCG~~~~TC G~AAAACGCT G~~~TTAT~~ MW00032 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT T~~~~~~~~~ ~~~~~~AACG AA~~~~AAGA A~~CA~~~CC ACCG~~~~TC G~GAAACACT G~~~TTAT~~ MW00051 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGCACGCG~ CTGCG~~~TA CACGTTTTTT T~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW00058 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGCACGCG~ CTGCG~~~TA CACGTTTTTT T~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW00064 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW00072 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT T~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW00076 TCGGCGAG~~ CGGGGCGAGC GCGC~~AC~~ ~KT~CTAA~C T~GTGT~CGT ~CAACKCKC~ ~AAGGCGCG~ CTGCG~TGTA CACGTTTTT~ ~~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW00101 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT T~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW00102 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GGGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT T~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW00110 NNNNNNNN~~ NNNNNNNNNN NNNN~~NN~~ ~NN~NNNN~N N~NNNN~NNN ~NNNNNNNN~ ~NNNNNNNN~ NNNNN~~~NN NNNNNNNNNN ~~~~~~~~~~ ~~~~~~NNNN AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW00116 TCGGTGAG~~ CGGGGCGTGC ACGCC~AG~~ ~GT~CGAA~C TTGCGT~CGT ~CGACGCGC~ ~AGAGTGCG~ CTGCG~CGTA C~~GTTT~~~ GCGT~TTAAC ~~~TGTTT~A AA~~~CG~CG ~~~CA~~~CC GCCG~~~~TC G~AAAACGCT G~~~TTAT~~ MW00127 TCGGCCAR~~ CGGGGCGTGC ACSC~~KC~~ ~GT~HSWA~C T~GTGT~YGM ~CGACSCGC~ ~ARAACGCG~ CTGSG~~K~A CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG AA~~~~AAAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW00129 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGRACGCG~ CTGCG~~~TA CACGTTTCTT CTTTT~~~~~ ~~~~~~AAMG AA~~~~~AAA A~~CA~~~CC ACCGTCTGTC C~GAAACACT G~~~TTAT~~ MW00151 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW00176 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~TC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW00177 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGGACGCG~ CTGCG~~~TA CACGTTTCTT CTTTT~~~~~ ~~~~~~AAAG AA~~~~~AAA A~~CA~~~CC ACCGTCTGTC C~GAAACACT G~~~TTAT~~ MW00178 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGGACGCG~ CTGCG~~~TA CACGTTTCTT CTTTT~~~~~ ~~~~~~AAAG AA~~~~AAAA ~~~CA~~~CC ACCGTCTGTC C~GAAACACT G~~~TTAT~~ MW00179 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT CA~~~~~~~~ ~~~~~TAACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~GAAACACT G~~~TTAT~~ MW00185 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~CA CACGTTTTT~ C~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW00189 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CRACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT TT~~~~~~~~ ~~~~~~AAMG AA~~~~AAAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW00226 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGGGC~ ~AGAACGCG~ CTGCG~~~TG CA~~~~~~~~ ~~~~~~~~~~ ~~~~~~~~~~ ~~~~~AAAAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW00231 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CKACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW00232 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGCACGCG~ CTGCG~~~TA CACGTTTTTT T~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT GTTGTATT~~ MW00234 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CKACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC K~AAAACACT G~~~TTAT~~ MW00262 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT TT~~~~~~~~ ~~~~~~AACG AA~~~~AGAA A~~CG~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW00267 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT T~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW00269 NNNNNNNN~~ NNNNNNNNNN NNNN~~NN~~ ~NN~NNNN~N N~NNNN~NNN ~NNNNNNNN~ ~NNNNNNNN~ NNNNN~~AAA CMAGTTTTTT ~~~~~~~~~~ ~~~~~~AAMG AA~~~AAAAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW00290 TCGGCGAG~~ CGGGGCGAGC GCGC~~AC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~TGTA CRCGTTTTT~ ~~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW00302 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT TTTTTTTTTT G~CAAAAAGG AA~~~AAAGA A~~CA~CACC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW00316 NNNNNNNN~~ NNNNNNNNNN NNNN~~NN~~ ~NN~NNNN~N N~NNNN~NNN ~NNNNNNNN~ ~NNNNNNNN~ NNNNN~~~NN NNNNNNNNNN ~~~~~~~~~~ ~~~~~~NNNN NN~~~~~NNN N~~NN~~~NN NNNN~~~~~C S~AAAA~~~~ ~~~~~~~~~~ MW00326 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG GA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW00328 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG AA~~~~AAAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW00330 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTCTT TT~~~~~~~~ ~~~~~~AACG AA~~~~AAAA A~~CA~~~CC ACAGTCTGTC C~GAAACACT G~~~TTAT~~ MW00347 TCGGCGAG~G CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C Y~KTKT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT T~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW00348 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW00393 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTCTT T~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW00409 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTCTT T~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW00412 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~TGTA CACTCTTTTT CCT~~~~~~~ ~~~~~~AACG AA~~~~GAAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW00444 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT T~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW00452 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW00469 TCGGTGAG~~ CGGGGCGTGC ACGC~~AC~~ ~GT~CGAA~C TTGTGT~CGT ~CGACGCGCG CAGAATGCG~ CTGCG~~~TA CTCGTT~~~~ ~~~~~~~~~C GTTCGTTGCG AA~~~CGCCG ~~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~GTTAT~~ MW00498 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW00504 TCGGCGAG~~ CGGGGNNNNN NNNN~~NN~~ ~NN~NGAA~C T~GTGT~TGT ~YGACGCGC~ ~ARAACGCG~ CTGGGK~~~A CACGTTTTTT AATG~~~AA~ ~~~~~~AAAA AA~AAAGAAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW00517 TCGGTGAG~~ CGGGGCGTGC ACGC~~AC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAATGCG~ CTGCG~~~TA ~~CGTTTTT~ ~~~~~~~~~~ ~~~~~~AAC~ AA~~~~~AAA ~~~CAACACC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW00525 TCGGTGAG~~ AGGGGCGTGC ACGC~~AC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAATGCG~ CTGCG~~~TA ~~CGTTTTTT ~~~~~~~~~~ ~~~~~~~~~~ ~~~~~~~~~A A~~CA~~~CC ACCG~~~~TC GAAAAACACA G~~~TTAT~~ MW00530 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~TGTA CACTCTTTTT CCT~~~~~~~ ~~~~~~AACG AA~~~~GAAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW00539 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG GA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~

194 Appendix 3: Sequence data Internal transcribed spacer 2 (ITS-2)

....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| 485 495 505 515 525 535 545 555 565 575 585 595 605 615 625 635 MW00547 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~ANAACGCN~ CTGCG~~~TA CACGTTTTTT T~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW01001 NNNNNNNN~~ NNNNNNNNNN NNNN~~NN~~ ~NN~NNNN~N N~NNNN~NNN ~NNNNNNNN~ ~NNNNNNNN~ NNNNN~~~NN NNNNNTTTTW ~~~~~~~~~~ ~~~~~~AASR AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW01011 TCGGTGAG~~ CGGGGCGTGC GCGC~~AC~~ ~GT~CGAA~C TTGTGT~CGT ~CGACGCGCG CAGAATGCG~ CTGCG~~~TA CTCGTT~~~~ ~~~~~~~~~C GTTCGTTGCG AA~~~CGCCG ~~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~GTTAT~~ MW01014 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG AA~~~AAAAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW01018 TCGGTGAG~~ CGGGGCGTGC ACGC~~AC~~ ~GT~CGAA~C TTGTGT~CGT ~CGACGCGCG CAGAATGCG~ CTGCG~~~TA CTCGTT~~~~ ~~~~~~~~~C GTTCGTTGCG AA~~~CGCCG ~~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~GTTAT~~ MW01019 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CAAGTTTTTT TCT~~~~~~~ ~~~~~AAACG AA~~~~~AAA A~~CA~CACC ACCG~~~~TC G~AAAGCACT G~~~TTAT~~ MW01025 NNNNNNNN~~ NNNNNNNNNN NNNN~~NN~~ ~NN~NNNN~N N~NNNN~NNN ~NNNNNNNN~ ~NGAACGCG~ TTGGG~TGWA CATTTTTTTT CCT~~~~~~~ ~~~~~~AAMR AA~~~~GAMA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW01034 TCGGTGAG~~ CGGGGCGTGC ACGC~~AC~~ ~GT~CGAA~C TTGTGT~CGT ~CGACGCGCG CAGAATGCG~ CTGCG~~~TA CTCGTT~~~~ ~~~~~~~~~C GTTCGTTGCG AA~~~CGCCG ~~~CA~~~CC RCCG~~~~TC G~AAAACACT G~~GTTAT~~ MW01039 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW01048 TCGGTGTG~~ CGGGGCGTGC ACGC~~AC~~ ~GT~CGAA~C T~GTGT~CGT ~CGGCGCGC~ ~GTGATGCG~ CTGCG~~~TA ~~CGTT~~~~ ~~~~~~~~~~ ~~~~~~~~CG AA~~~CGAAA A~~CA~~~CC GCCG~~~~TC G~AAAACACT G~~~TTAT~~ MW01053 TCGGCGAG~~ CGGGGCGTGC ACGC~~KC~~ ~NN~NNNN~C T~GTGT~CKT ~MTACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG AA~~~~AAAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW01059 TCGGTGAG~~ CGGGGCGTGC ACGC~~AC~~ ~GT~CGAA~C TTGTGT~CGT ~CGACGCGCG CAGAATGCG~ CTGCG~~~TA CTCGTT~~~~ ~~~~~~~~~C GTTCGTTGCG AA~~~CGCCG ~~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~GTTAT~~ MW01061 TCGGTGTG~~ CGGGGCGTGC ACGC~~AC~~ ~GT~CGAA~C T~GTGT~CGT ~CGGCGCGC~ ~GTGATGCG~ CTGCG~~~TA ~~CGTT~~~~ ~~~~~~~~~~ ~~~~~~~~CG AA~~~CGAAA A~~CA~~~CC GCCG~~~~TC G~AAAACACT G~~~TTAT~~ MW01083 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CAAGTTTTTT TTTTTTT~~~ G~CAAAAAGG AA~~~AGAGA A~~CA~CGCC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW01092 TCGGTGAG~~ CGGGGCGTGC ACGC~~AC~~ ~GT~CGAA~C TTGTGT~CGT ~CGACGCGCG CAGAATGCG~ CTGCG~~~TA CTCGTT~~~~ ~~~~~~~~~C GTTCGTTGCG AA~~~CGCCG ~~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~GTTAT~~ MW01107 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG GA~~~~AAAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW01116 TCGGTGAG~~ CGGGGCGTGC ACGC~~AC~~ ~GT~CGAA~C TTGTGT~CGT ~CGACGCGCG CAGAATGCG~ CTGCG~~~TA CTCGTT~~~~ ~~~~~~~~~C GTTCGTTGCG AA~~~CGCCG ~~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~GTTAT~~ MW02001 TCGGCGAG~~ CGGGGCGTGC ACGC~~AC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~TGTA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG AA~AAAAAAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW02006 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT TCT~~~~~~~ ~~~~~~AACG AA~~~~AAAA A~~CAACACC ACCG~~~~TC G~AAAACACT G~~~TTGT~~ MW02025 TCGGT~TA~~ AGGG~CGGGY GTGC~~AC~~ ~~T~CGAA~C TT~TGT~CGT ~CTA~GCGTT GAGAATGCG~ CGAGG~~~TG CTCTCCAT~~ ~~~~~~~~~~ ~~~~~~~~~~ ~~~~~~~~~~ ~~~~~~~~~~ ~~~~~~~~~~ ~~~~~~~~~~ ~~~~~~~~~~ MW02033 TCGGTGTG~~ CGGGGCGTGC ACGC~~AC~~ ~GT~CGAA~C T~GTGT~CGT ~CGGCGCGC~ ~GTGATGCG~ CTGCG~~~TA ~~CGTT~~~~ ~~~~~~~~~~ ~~~~~~~~CG AA~~~CGAAA A~~CA~~~CC GCCG~~~~TC G~AAAACACT G~~~TTAT~~ MW02034 TCGGTGAG~~ AGGGGCGTGC ACGC~~AC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAATGCG~ CTGCG~~~TA ~~CGTTTTTT ~~~~~~~~~~ ~~~~~~~~~~ ~~~~~~~~~A A~~CA~~~CC GCCG~~~~TC GAAAAACACA G~~~TTAT~~ MW98009 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGGACGCG~ CTGCG~~~TA CACGTTTTTT T~~~~~~~~~ ~~~~~~AACG AATAAAAAAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW98029 TCGGCRAG~~ CGGGGCGAGC GCGC~~AC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGGGCGCG~ CTGCG~TGTA CACGTTTTT~ ~~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW98049 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~GGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~TC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW98079 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~ARAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW98097 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW98103 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~TC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW98129 TCGGTGAG~~ CGGGGCGTGC ACGC~~AC~~ ~GT~CGAA~C TTGTGT~TGT ~CAACGCGCG CAGAATGCG~ CTGCG~~~TA CTCGTT~~~~ ~~~~~~~~~C GTTCGTTGCG AA~~~CGCCG ~~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~GTTAT~~ MW98136 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CGCGTTTCTT TT~~~~~~~~ ~~~~~~AACG AA~~~~AGAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~A MW98138 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG AG~~~~~AAA A~~CA~~~CC ACCT~~~~TC G~AAAACACT G~~~TTAT~~ MW98139 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CGCGTTTCTT TT~~~~~~~~ ~~~~~~AACG AA~~~~AGAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~A MW98154 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG AG~~~~~AAA A~~CA~~~CC ACCT~~~~TC G~AAAACACT G~~~TTAT~~ MW98162 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW98170 TCGGMGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~MAAA~C T~GTGT~CGT ~~~ACACGC~ ~~~~~~~~~~ ~~~~~~~~~~ ~~~~~~~~~T ~~~~~~~~~~ ~~~~~~AACG AA~~~~AGAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~A MW98172 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CASGTTTTTT ~~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW98180 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG AG~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW98185 TCGGTGAG~~ CGGGGCGTGC ACGC~~AC~~ ~GT~CGAA~C TTGTGT~CGT ~CGACGCGCG CAGAATGCG~ CTGCG~~~TA CTCGTT~~~~ ~~~~~~~~~C GTTCGTTGCG AA~~~CGCCG ~~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~GTTAT~~ MW98189 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~ANAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG AA~AAAAAAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW98203 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~A T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW98205 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTAT TTTTTCCTTT TTTT~~AACG AA~~~AAAAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW98212 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW98220 TCGGTGAG~~ CGGGGCGTGC ACGCGCAC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAATGCG~ CTGCG~~~TA ~ATNNNNNN~ ~~~~~~~~~~ ~~~~~~NNNA AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TCGT~~ MW98228 TCGGTGAG~~ CGGGGCGTGC ACGC~~AC~~ ~GT~CGAA~C TTGTGT~CGT ~CGACGCGCG CAGAATGCG~ CTGCG~~~TA CTCGTT~~~~ ~~~~~~~~~C GTTCGTTGCG AA~~~CGCCG ~~~CA~~~CC ACCG~~~~TC G~AAAACACC G~~GTTAT~~ MW98235 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~~~~~ AA~CGGAAAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW98240 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CKAA~C T~GTGT~CGT ~CGACGCGC~ ~AGRACGCG~ CTGCG~~~TA CACGTTTTTT TTTTTT~~~~ ~~~~~AAACG AAACGAAAAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW98261 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG AATAAAAAAA A~~CA~~~CC ACMG~~~~TC G~AAAACACT G~~~TTAT~~ MW98264 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG GA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW98270 TCGGTGTG~~ CGGGGCGTGC ACGC~~AC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAATGCG~ CTGCG~~~TA ~~CGTTC~~~ ~~~~~~~~~~ ~~~~~AAAC~ ~A~~~~GAAA ~~~CA~CACC ACCG~~~~TC G~AAAACACG G~~~TTAT~~ MW98278 TCGGTGAG~~ CGGGGCGTGC ACGC~~AC~~ ~GT~CGAA~C TTGTGT~CGT ~CAACGCGCG CAGAATGCG~ CTGCG~~~TA CTCGTT~~~~ ~~~~~~~~~C GTTCGTTGCG AA~~~CGCCG ~~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~GTTAT~~ MW98285 TCGGMGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~ANAA~C T~GTGT~CGT ~MSACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTATTT ~~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW98294 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT TTTTTTTTTT TTTT~WAACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW98313 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CGCGTTTCTT TT~~~~~~~~ ~~~~~~AACG AA~~~~AGAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~A MW98315 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW99001 TCGGTGTG~~ CGGGGCGTGC GCGC~~AC~~ ~GT~CGAA~C T~GTGT~CGT ~CGGCGCGC~ ~GTGATGCG~ CTGCG~~~TA ~~CGTT~~~~ ~~~~~~~~~~ ~~~~~~~~CG AA~~~CGAAA A~~CA~~~CC GCCG~~~~TC G~AAAACGCT G~~~TTAT~~ MW99006 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGK ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW99009 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~TC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW99013 TCGGCGAG~~ CGGGGCGTGC ACGC~~AC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAATGCG~ CTTCG~~~TA ~~CGTTT~~~ ~~~~~~~~~~ ~~~~~~AACA AAA~~C~~AA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTATAT MW99014 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CAAGTTTTTT CTTT~~~~~~ ~~~~~AAACG AA~~~~~AAA A~~CA~CACC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW99018 TCGGTGAG~~ CGGGGCGTGC ACGC~~AC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAATGCG~ CTGCG~~~TA ~~CGTTT~~~ ~~~~~~~~~~ ~~~~~~~~~A AA~~~CAAAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW99038 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CKAA~C T~GTGT~CGT ~CGACGCGC~ ~ASAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~~~~~ AA~CGGAAAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TKAT~~ MW99047 TCGGCGAG~~ CGGGGCGTGC ACGC~~AC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~TGTA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG AA~~AAGAAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW99057 TMKGYCAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CYAA~C T~GTGT~STT ~CNACTCAC~ ~ACAACGCG~ CTGCG~~~TA CGCGTTTNTT TT~~~~~~~~ ~~~~~~WWCK AA~~~~AGAA A~~CA~~~CC CCCG~~~~TC G~AAAACACT G~~~TTAT~A MW99058 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW99094 TCGGCGAG~~ CGGGGCG~GC ACGC~~NN~~ ~NN~NNNN~N NNNNNN~NNN ~NNNNNNNN~ ~NNNNNNNN~ NNNNN~~~NN NNNNTTTTTT TT~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~GAGACACT G~~~TTAT~~ MW99095 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~GTAACG AA~~~~~AAA G~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW99097 TCGGTGTG~~ CGGGGCGTGC ACGC~~AC~~ ~GT~CGAA~C T~GTGT~CGT ~CGGCGCGC~ ~GTGATGCG~ CTGCG~~~TA ~~CGTT~~~~ ~~~~~~~~~~ ~~~~~~~~CG AA~~~CGAAA A~~CA~~~CC GCCG~~~~TC G~AAAACACT G~~~TTAT~~ MW99105 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT T~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~

195 Appendix 3: Sequence data Internal transcribed spacer 2 (ITS-2)

....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| 485 495 505 515 525 535 545 555 565 575 585 595 605 615 625 635 MW99134 TCGGTGAG~~ CGGGGCGTGC ACGC~~AC~~ ~GT~CGAA~C T~GTGT~CGT ~CGGCGCG~G CAGAGTGCG~ CTGCG~~~TA ~~CGTGTTTT TGTTTATT~~ ~~~~~~AAC~ AA~~~~~AAA ~~~CAACACC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW99135 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT TT~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~TC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW99163 TCGGTGAG~~ CGGGGCG~GC ACGC~~AC~~ ~NN~NNNN~N N~NNNN~NNN ~NNNNNNNN~ ~NNNNNNNN~ NNNNN~~~NN NNNNNNNNN~ ~~~~~~~~~~ ~~~~~~NNNN NN~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~T~TTAT~~ MW99164 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW99174 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW99196 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GTTCGAAAC T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT T~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW99203 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT T~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~TC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW99226 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGC ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW99240 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~TGAA~N T~GTGT~CGT CAGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CGCGTTTCTT TT~~~~~~~~ ~~~~~~AACG AA~~~~AGAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~A MW99247 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CKACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW99274 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAW~C T~GTGT~CGT ~NNACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG AA~~~~~AAM A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW99276 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW99286 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TG CA~~~~~~~~ ~~~~~~~~~~ ~~~~~~~~~~ ~~~~~AAAAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW99292 NNNNNNNN~~ NNNNNNNNNN NNNN~~NN~~ ~NN~NNNN~N NNNNNN~NNN ~NNNNNNNN~ ~NNNNNNNN~ NNNNN~~~NN NNNNNNNNNN ~~~~~~~~~~ ~~~~~~NNNN NN~~~~~NNC A~~CC~~~CC MCYG~~~~TM G~AACACTST G~~~TTAT~~ MW99301 TCGGCGAG~~ CGGGGCGTGC ACGC~~AC~~ ~GT~CGAA~C K~GTGT~CGG ~SGAAGCGCA ~AGAACGCG~ CTGCG~~~TA CACGTTATTT TNTG~GGAA~ ~~~~~~AATA AA~~~~AAAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW99303 TCGGTGAG~~ CGGGGCGTGC ACGC~~AC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAATGCG~ CTGCG~~~TA ~~CGTTTCTT ~~~~~~~~~~ ~~~~~~AACG ~~~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~T~TTAT~~ MW99309 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG AA~~~~~AAW A~~CA~~~TC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW99319 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACRCT G~~~TTAT~~ MW99341 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGGACGCG~ CTGCG~~~TA CACGTTCTTT ~~~~~~~~~~ ~~~~~~AACG AA~~~~AAAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTRT~~ MW99353 TCGGCGAG~~ CGGGGCGTGC RCGC~~GC~~ ~GT~CGAA~C TTGTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CGCGTTTCTT TT~~~~~~~~ ~~~~~~AACG AA~~~~AGAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~A MW99372 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C K~GTGT~CGK ~SKACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACSTTTYTT ~~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW99374 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGC ~CGACGCGC~ ~AGAACGCG~ CTGSG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CM~~~CC CCCG~~~~TC G~AAAACACT G~~~TTAT~~ MW99382 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTT~ C~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW99393 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CKAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TG CA~~~~~~~~ ~~~~~~~~~~ ~~~~~~~~~~ ~~~~~~AAAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW99406 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CKAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TG CA~~~~~~~~ ~~~~~~~~~~ ~~~~~~~~~~ ~~~~~~AAAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW99407 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW99408 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG AA~~~AAAAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW99413 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT T~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW99448 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT TCG~T~~~~~ ~~~~~~AACT AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~GAGACACT G~~~TTAT~~ MW99465 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CGCGTTTCTT TT~~~~~~~~ ~~~~~~AACG AAA~~~GAAA ~~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~A MW99471 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW99479 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGCACGCG~ CTGCG~~~TA CGCGTTTTTT T~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC K~AAAACACT G~~~TTAT~~ MW99501 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG AG~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW99508 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGCACGCG~ CTGCG~~~TA CACGTTTTTT T~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW99514 TCGGTGAG~~ CGGGGCGTGC ACGC~~AC~~ ~GT~CGAA~C TTGTGT~CGT ~CGACGCGCG CAGAATGCG~ CTGCG~~~TA CTCGTTGTT~ ~~~~~~~~~C GTTCGTTGCG AA~~~CGCCG ~~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~GTTAT~~ MW99536 TCGGTGAG~~ CGGGGCGTGC ACGC~~AC~~ ~GT~CGAA~C TTGTGT~CGT ~CGACGC~CG CAGAATGCG~ CTGCG~~~TA CTCGTTGTT~ ~~~~~~~~~C GTTCGTTGCG AA~~~CGCCG ~~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~GTTAT~~ MW99537 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG GA~~~~~AAA A~~CACCACC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW99546 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT T~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW99550 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG GA~~~~~AAA A~~CACCACC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW99552 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~GTAACG AA~~~~~AAA G~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW99559 TCKGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CCAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~TC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW99565 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~TC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW99591 TCGGCGAG~~ CGGGGCGTGC GCGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW99605 TCGGCGAG~~ CGGGGCGTGC GCGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CKACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ MW99608 TCGGTGAG~~ CGGGGCGTGC ACGC~~AC~~ ~GT~CGAA~C TTGTGTTCGT TCGACGCGCG CAGAATGCG~ CTGCG~~~TA CTCGTT~~~~ ~~~~~~~~~C GTTCGTTGCG AA~~~CGCCG ~~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~GTTAT~~ MW99612 TCGGTGAG~~ CGGGGCGTGC ACGC~~AC~~ ~GT~CGAA~C TTGTGT~CGT ~CGACGCGCG CAGAATGCG~ CTGCG~~~TA CTCGTT~~~~ ~~~~~~~~~C GTTCGTTGCG AA~~~CGCCG ~~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~GTTAT~~ MW99613 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT T~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ OK96022 TCGGTGAG~~ CGGGGCGTGC ACGC~~AC~~ ~GT~CGAA~C TTGTGT~CGT ~CGACGCGCG CAGAATGCG~ CTGCG~~~TA CTCGTT~~~~ ~~~~~~~~~C GTTCGTTGCG AA~~~CGCCG ~~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~GTTAT~~ WE00002 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGGCGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT TTT~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ WE02431 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~TGTTAT~~ WE02451 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGGCGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT T~~~~~~~~~ ~~~~~~AGCG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ WE02454 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG AA~~~~~TAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ WE02491 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ WE02531 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGC ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ WE02534 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CRAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTYTTT T~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ WE02535 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ WE02591 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTT~ ~~~~~~~~~~ ~~~~~CAACG AAG~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ WE02614 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGRCGGGC~ ~AGAACGCG~ YTGCG~~~TG CA~~~~~~~~ ~~~~~~~~~~ ~~~~~~~~~~ ~~~~~AAAAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ WE02621 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ WE02661 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGCACGCG~ CTGCG~~~TA CACGTTTTTT T~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ WE02662 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~TC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ WE02671 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGSGC~ ~AGAACGCG~ CTGCG~~~TG CA~~~~~~~~ ~~~~~~~~~~ ~~~~~~~~~~ ~~~~~~AAAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ WE02674 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTCTT CTTTT~~~~~ ~~~~~~AAAG AA~~~~~AAA A~~CA~~~CC ACCGTCTGTC C~GAAACACT G~~~TTAT~~ WE02677 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~YA CACGTTTTT~ C~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ WE85001 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGT~CGT ~CGACGCGC~ ~AGAACGCG~ CTGCG~~~TA CACGTTTTTT ~~~~~~~~~~ ~~~~~~AACG AA~~~~~AAA A~~CA~~~CC ACCG~~~~TC G~AAAACACT G~~~TTAT~~ WE94001 TCGGCGAG~~ CGGGGCGTGC ACGC~~GC~~ ~GT~CGAA~C T~GTGTTCGT ~CGACGCGC~ TAGANNNNN~ NNNNN~~~NN NNNNNNNNNN ~~~~~~~~~~ ~~~~~~NNNN NN~~~~~NNN N~~NN~~~NN NNNN~~~~NN N~NNNNNNNN N~~~NNNN~~

196 Appendix 3: Sequence data Internal transcribed spacer 2 (ITS-2)

....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|.. 645 655 665 675 685 695 705 715 725 735 745 755 765 775 785 795 AD98001 ~NNN~~~~NN NNNNNN~~NN N~NN~~~NNN N~~~~~NN~N NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNN~~~N NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNN AD98009 ~ATT~~~~AT TAATTG~~AA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA AD98012 TATT~~~~AT CAATTG~~AA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA AD98018 ~ATT~~~~AT CAATTGTGAA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA AD98024 ~ATT~~~~AT CAATTG~~AA TATT~~~ATA A~~~~~AC~A GTAGGCGGAC TCGACKTCCR AAGGGCGCGT TTTCAACCCC GACGTG~~~C CSATNGATTT CGGTCGCGGA GGCGGAGTCG TTGCNCTNNN NNNNNNNNNN NNNNNNNNNN NNNNNNN AD98036 ~ATT~~~~AT CAATTG~~AA C~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCTGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA DS00001 ~ATT~~~~GT TAATTG~~AA T~TA~TAATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGTGCGCGT CGTCNACCCC CACNTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCC TTGCGCTTAC GGATATCGCG TCTGCCTCAA TTTTTTA DS01001 TATT~~~~AT TAATTG~~AA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA JC00029 ~ATT~~~~AT ~~~TTG~~AA A~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTA~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA JC00039 ~ATT~~~~AT ~~~TTG~~AA T~TA~~~ATG A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTA~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA JC00040 ~ATT~~~~AT TAATTG~~AA TATA~~~ATA A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA JC00042 ~ATT~~~~AT ~~~TTG~~AA A~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTA~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA JC00043 ~ATT~~~~AT TAATTG~~AA T~TA~~~ATA A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATCT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA JC00045 ~ATT~~~~AT TAATTG~~AA T~TA~~~ATA A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATCT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA JC00051 ~ATT~~~~AT ~~~TTG~~AA A~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTA~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTN JC00055 ~ATT~~~~AT ~~~TTG~~AA T~TA~~~ATG A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGCA~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA ATTTTTA JC00061 ~ATT~~~~AT ~~~TTG~~AA A~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTA~~~C CGTTTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA JC00063 ~ATT~~~~AT ~~~TTG~~AA A~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTA~~~C CGTTTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA JC01014 ~ATT~~~~AT TAATTG~~AA T~YA~~~ATA A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATCT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTT~A JM00001 ~ATT~~~~AT TAATTG~~AA T~TA~~~ATA A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW00015 ~ATT~~~~AT TAATTG~~AA T~TA~~~ATA A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATCT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW00024 ~ATT~~~~AT ~~~TTG~~AA T~TG~~~AAT CA~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTA~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA ATTTTTA MW00032 ~ATT~~~~AT CAATTG~~AA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW00051 ~ATT~~~~AT CAATCG~~AA T~TG~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW00058 ~ATT~~~~AT CAATCG~~AA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW00064 ~ATT~~~~AT TAATCG~~AA T~TA~~~ATA A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW00072 ~ATT~~~~AT TAATTG~~AA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW00076 ~ATT~~~~AT ~~~TTGAGAA T~TA~~~ATA GAA~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTA~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTNNNNNNN NNNNNNN MW00101 ~ATT~~~~AT TAATTG~~AA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCNN NNNNNNN MW00102 ~ATT~~~~AT TGATTG~~AA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW00110 ~ATT~~~~AT CAATCG~~AA T~TR~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTN MW00116 ~ATT~~~~AT ~~~TTG~~AA T~TA~~~ATG GA~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTA~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA ATWTATA MW00127 ~ATT~~~~AT CAATTG~~AA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATCGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW00129 ~RTT~~~~GT TAATTG~~AA T~TA~~~ATT G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW00151 ~ATT~~~~AT TAATTG~~AA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW00176 ~AATA~~TAT TAATTG~~AA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW00177 ~ATT~~~~GT TAATTG~~AA T~TA~~~ATT G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW00178 ~ATT~~~~GT TAATTG~~AA T~TA~~~ATT G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW00179 ~ATT~~~~AT TAATTG~~AA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW00185 ~ATT~~~~AT TAATTG~~AA T~TA~~~ATA A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATCT CGGTCGCGGA GGCGGAGTC~ TTGCGCTGAC NGATATCGCG TCTGCCTCAA TTTTTTA MW00189 ~ATT~~~~AT TAATTG~~AA T~TA~~~ATA A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATCT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW00226 ~ATT~~~~AT CGATTG~~AA T~TG~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCTGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW00231 ~ATT~~~~AT TAATTG~~AA T~TA~~~ATA A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW00232 ~ATT~~~~AT CAATTG~~AA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW00234 ~ATT~~~~AT TAATTG~~AA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW00262 ~ATTATTTAT TAATTA~~AA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAN NNNNNNN MW00267 ~ATT~~~~AT TGATTG~~AA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW00269 ~ATT~~~~AT TAATTG~~AA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTN MW00290 ~ATT~~~~AT ~~~TTGAGAA T~TA~~~ATA GAA~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTA~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAW ATTTTTA MW00302 ~ATT~~~~AT ~~~TCG~~AA T~CA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTA~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCMTCAA TTNTATN MW00316 ~~~~~~~~~T CAATCG~~AA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAN NNNNNNN MW00326 ~ATT~~~~AT ~~~TTGA~AA TG~A~~~ATA A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTA~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW00328 ~ATT~~~~AT TAATTG~~AA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW00330 ~ATT~~~~GT TAATTG~~AA T~TA~~~ATT G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCNN NNNNNNN MW00347 ~ATT~~~~AT TAATTG~~AA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW00348 ~ATT~~~~AT CAATTG~~AA C~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CTGTCGCGGA GGCTGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW00393 ~ATT~~~~AT TAATTG~~AA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTCN MW00409 ~ATT~~~~AT TAATTG~~AA C~TA~~~ATA GAG~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW00412 ~ATT~~~~AT ~~~TTG~~AA A~TG~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTA~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW00444 ~ATT~~~~AT TAATTG~~AA T~CA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW00452 ~ATT~~~~AT CAATTG~~AA C~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC KACGTG~~~C CGATTGATTT CGGTCGCGGA GGCTGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW00469 ~GTT~~~~AT ~~~ACG~~AA T~TA~~~ACG A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTA~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA ATTTTTA MW00498 ~ATT~~~~AT TAATCG~~AA T~TA~~~ATA A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW00504 ~ATT~~~~AT ~~~TCG~~AA T~TA~~~ATA A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTA~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTNN MW00517 ~ATT~~~~GT TTGTTG~~AA T~TA~~~ATG A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTA~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA ATTTTTA MW00525 ~ATT~~~~AT ~~~TCG~~AA T~TG~~~ATA A~~~ACA~GA GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTA~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA ATTTTTA MW00530 ~ATT~~~~AT ~~~TTG~~AA A~TG~~~ATA G~~~~~ACAA GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTA~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTT~A MW00539 ~ATT~~~~AT TAATTG~~AA T~TA~~~ATA T~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTN

197 Appendix 3: Sequence data Internal transcribed spacer 2 (ITS-2)

....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|.. 645 655 665 675 685 695 705 715 725 735 745 755 765 775 785 795 MW00547 ~GTT~~~~AT TAATTG~~AA T~TA~~~ACA N~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACNTG~~~C CNATNGATTT CGGTCNCGGA GGCGGANTCN TTGCNCTGCN NGNNNTCGNG TNTGCCTNAA TTTTTTN MW01001 ~ATT~~~~AT TAATTG~~AA T~TA~~~ATA A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTNN MW01011 ~GTT~~~~AT ~~~ACG~~AA T~TA~~~ACG A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTA~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGNNNNNN NNNNNNN MW01014 ~ATT~~~~AT TAATTG~~AA T~TA~~~ATG A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATCT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW01018 ~GTT~~~~AT ~~~ACG~~AA T~TA~~~ACG A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTA~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA ATTTTTA MW01019 ~ATT~~~~AT ~~~TTG~~AA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTA~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW01025 ~ATT~~~~AT ~~~TTG~~AA A~YA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTA~~~C CGTTTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW01034 ~GTT~~~~AT ~~~ACG~~AA T~TA~~~ACG A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTA~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA ATTTTTA MW01039 ~ATT~~~~AT TAATTG~~AA T~TA~~~ATA A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW01048 ~ATT~~~~AT ~~~TTG~~AA T~TA~~~ATG A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGCA~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA ATTTTTA MW01053 ~ATT~~~~AT TAATTG~~AA T~TA~~~ATA A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTGTTTA MW01059 ~GTT~~~~AT ~~~ACG~~AA T~TA~~~ACG A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTA~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA ATTTTTA MW01061 ~ATT~~~~AT ~~~TTG~~AA T~TA~~~ATG A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGCA~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA ATTTTTA MW01083 ~ATT~~~~AT ~~~TCG~~AA T~CA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGCA~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTCN MW01092 ~GTT~~~~AT ~~~ACG~~AA T~TA~~~ACG A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTA~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA ATTTTTA MW01107 ~ATT~~~~AT TAATTG~~AA T~TA~~~ATA A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG THTGCCTCAA TTTTT~A MW01116 ~GTT~~~~AT ~~~ACG~~AA T~TA~~~ACG A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTA~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA ATTTTTA MW02001 ~ATT~~~~AT ~~~TTG~~AA T~TA~~~ATA A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTA~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW02006 ~ATT~~~~AT ~~~TTG~~AA T~CA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTA~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW02025 ~~~~~~~~~~ ~~~~~~~~~~ ~~~~~~~~~~ ~~~~~~~C~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CG~TTCA~CG CGGTT~CGGA GGCGTAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA ATTTTTA MW02033 ~ATT~~~~GT ~~~TTG~~AA T~TA~~~ACG A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGCA~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA ATTTTTA MW02034 ~ATT~~~~AT ~~~TCG~~AA T~TA~~~ATA A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTA~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA ATTTTTA MW98009 ~ACT~~~~AT CAAATG~~AA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW98029 ~ATT~~~~AT ~~~TTGAGAA T~TA~~~ATA GAA~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTA~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA ATTTTTA MW98049 ~ATT~~~~AT TAATTG~~AA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW98079 ~ATT~~~~AT TAATTG~~AA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW98097 ~ATT~~~~AT TAATTG~~AA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW98103 ~ATT~~~~AT TAATTG~~AA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW98129 ~GTT~~~~AT ~~~ACG~~GA T~TA~~~ACG A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTA~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATNNNNNN NNNNNNNNNN NNNNNNN MW98136 TATT~~~~AT CAATTG~~RA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW98138 ~ATC~~~~AT TAATTG~~AA TTCA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW98139 TATT~~~~AT CAATTG~~AA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW98154 ~ATC~~~~AT TAATTG~~AA TTCA~~~ATA A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTNNNNN MW98162 ~ATT~~~~AT TAATTG~~AA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW98170 TATT~~~~AT CAATTG~~AA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW98172 ~ATT~~~~AT TAATTG~~AA T~TA~~~ATA A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGGTTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW98180 ~ATC~~~~AT TAATTG~~AA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW98185 ~GTT~~~~AT ~~~ACG~~GA T~TA~~~ACG A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTA~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA ATTTTTA MW98189 ~ATT~~~~AT TAATTG~~AA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW98203 ~ATT~~~~AT TAATTG~~AA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW98205 ~ATT~~~~AT CAATTG~~AA T~TG~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW98212 ~ATT~~~~AT TAATTG~~AA T~TA~~~ATA A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW98220 ~ATT~~~~AT ~~~TCG~~AA T~TA~~~ATA G~~~~~AA~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA ATTTTGN MW98228 ~GTT~~~~AT ~~~ACG~~AA T~TA~~~ACG A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTA~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA ATTTTTN MW98235 ~ATT~~~~AT ~~~TTGA~AA TTCA~~~ATA A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTA~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW98240 ~ATT~~~~AT CAATTG~~AA T~TG~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW98261 ~ACT~~~~AT CAAATG~~AA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW98264 ~ATT~~~~AT ~~~TTGA~AA TTCA~~~ATA A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTA~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW98270 ~ATT~~~~AT ~~~TTG~~AA T~TA~~~ATG A~~~~~ACAA GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTA~~~C CGATTGACAT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA ATTTTTA MW98278 ~GTT~~~~AT ~~~ACG~~AA T~TA~~~ACG A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTA~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA ATTTTTA MW98285 ~ATT~~~~AT TAATTG~~AA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTYTA MW98294 ~ATT~~~~AT CAATTG~~AA T~TG~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW98313 TGTT~~~~AT CAATTG~~AA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW98315 ~ATTT~~~AT TAATTG~~AA T~TA~~~ATA A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA NNNNNNN MW99001 ~ATT~~~~AT ~~~TTG~~AA T~TA~~~ATG A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGCA~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA ATTTTTA MW99006 ~ATT~~~~AT TAATTG~~AA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW99009 ~ATT~~~~AT TAATTG~~AA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTN MW99013 TATT~~~~AT ~~~TTG~~AA T~CA~~~ATA A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTA~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA ATTTTTA MW99014 ~ATT~~~~AT ~~~TTG~~AA T~TA~~~ACA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTA~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW99018 ~~~~~~~~~~ ~~~~~~~~AA T~TA~~~ATA TA~CACAC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA ATTTTTA MW99038 ~ATT~~~~AT ~~~TTGA~AA TTCA~~~ATA A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTA~~~C CGATTGATTT CTGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCKCG TCTGCCTCAA TTTTTTA MW99047 ~ATT~~~~AT ~~~TTG~~AA T~TA~~~ATA A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTA~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW99057 TATT~~~~AT CAATTG~~AA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW99058 ~ATT~~~~AT TAATTG~~AA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW99094 ~ATCAA~TAT ~AATTG~~AA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW99095 ~ATT~~~~AT CAATTG~~AA T~TG~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TNNNNNN MW99097 ~ATT~~~~AT ~~~TTG~~AA T~TA~~~ATG A~~~~~AA~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGCA~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG YTGCGCTGAC GGATATCGCG TCTGCCTCAA ATTTTGA MW99105 ~ATT~~~~AT TAATTG~~AA T~TA~~~ATA A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATCT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA

198 Appendix 3: Sequence data Internal transcribed spacer 2 (ITS-2)

....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|....| ....|.. 645 655 665 675 685 695 705 715 725 735 745 755 765 775 785 795 MW99134 ~ATT~~~~AT ~~~TTG~~AA T~TG~~~ATA TAA~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTA~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA ATTTTTA MW99135 ~ATT~~~~AT TAATTG~~AA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW99163 ~ATT~~~~AT ~~~TTG~~AA T~TA~~~ATA AN~~~~ACCA GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTA~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA NNNNNNN MW99164 TATT~~~~AT TAATTG~~AA T~TA~~~ATA A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW99174 ~ATT~~~~AT CAATTG~~AA C~TAATAATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCTGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW99196 ~ATT~~~~AT TAATTG~~AA T~TA~~~ATA A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATCT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTN MW99203 ~ATT~~~~AT TAATTG~~AA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTYTA MW99226 ~ATT~~~~AT CGATTG~~AA T~TAAT~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATCGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW99240 TATT~~~~AT CAATTG~~AA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAN NNNNNNN MW99247 ~ATT~~~~AT TAATTG~~AA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW99274 ~ATT~~~~AT TAATTG~~AA T~TA~~~ATA A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCKCGGA GGCGGAGTCG TTGCGSTGAC GGATATCGCG TCKGCCTCAA TTKTYTA MW99276 ~ATT~~~~AT TAATTG~~AA T~TA~~~ATA A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW99286 ~ATT~~~~AT CAATTG~~AA T~TG~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCTGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW99292 ~ATT~~~~AT CGWTTR~~AA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTNNN MW99301 ~ATT~~~~AT ~~~TTG~~AA T~TA~~~ATA A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTA~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW99303 ~ATT~~~~AT ~~~TTG~~AA T~TA~~~ATA ATAA~~ACCA GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTA~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA ATTTTTA MW99309 ~ATTA~TTAT TAATTG~~AA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW99319 ~ATT~~~~AT TAATTG~~AA T~TA~~~ATA A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTKTYTA MW99341 ~ATT~~~~AT CGATTG~~AA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW99353 TATT~~~~AT CAATTG~~AA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW99372 ~ATT~~~~AT TAATTG~~AA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW99374 ~ATT~~~~AT CGATTG~~AA T~TAAT~ATA G~~~~~MC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATCGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATSGCG TYKGCCTCAA TTKTYCA MW99382 ~ATT~~~~AT TAATTG~~AA T~TA~~~ATA A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACTCC GACNTG~~~C CGATTGATCT CGGTCGCGGA GGCGGAGTCG TTGCGCTNAC GGATATCGCG TCTGCCTCAA TTTTTTA MW99393 ~ATT~~~~AT CAATTG~~AA T~TG~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCTGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTKTTTA MW99406 ~ATT~~~~AT CAATTG~~AA T~TG~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCTGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCMTSAA TYYTTTA MW99407 ~ATT~~~~AT TAACTG~~AA T~TA~~~ATA A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATCT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW99408 ~ATT~~~~AT TAATTG~~AA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW99413 ~ATT~~~~AT TAATTG~~AA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA NNNNNNN MW99448 ~ATCAA~TAT ~AATTG~~AA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW99465 TATT~~~~AT CAATTG~~AA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW99471 ~ATT~~~~AT TAATTG~~AA T~TA~~~ATA A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGGTTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTNN MW99479 ~ATT~~~~AT CAATTG~~AA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW99501 ~ATT~~~~AT TAATTG~~AA TTCA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW99508 ~ATT~~~~AT CAATTGTGAA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCCGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW99514 ~GTT~~~~GT ~~~ACG~~AA T~TA~~~ACG A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTA~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA ATTTTKA MW99536 ~GTT~~~~AT ~~~ACG~~AA T~TA~~~ACG A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTA~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA ATTTTTA MW99537 ~ATT~~~~AT ~~~TTGA~AA T~TA~~~ATA A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTACTAC CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW99546 ~ATT~~~~AT TAATTG~~AA T~TA~~~ATA A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTA~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW99550 ~ATT~~~~AT ~~~TTGA~AA T~TA~~~ATA A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTACTAC CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW99552 ~ATT~~~~AT CAATTG~~AA T~TG~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW99559 ~ATT~~~~AT TAATTG~~AA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGNA GGCGGAGTCG NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNN MW99565 ~ATT~~~~AT TAATTG~~AA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW99591 ~ATT~~~~AT TAATTG~~AA TATA~~~ATA A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW99605 ~ATT~~~~AT TAATTG~~AA TATA~~~ATA A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA MW99608 ~GTT~~~~AT ~~~ACG~~AA T~TA~~~ACG A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTA~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA ATTTTTA MW99612 ~GTT~~~~AT ~~~ACG~~AA T~TA~~~ACG A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTA~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA ATTTT~A MW99613 ~ATT~~~~AT TAATTG~~AA T~TA~~~ATA A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTA~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA OK96022 ~GTT~~~~AT ~~~ACG~~AA T~TA~~~ACG A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACNCC CACCTA~~~C CGATTGATTT CNGTCGCGGA GGCGGAGTCG TTGCGCTGAC NGATATCGCG TCTGCCTCAA ATTTTTA WE00002 ~ATT~~~~AT AAATTA~~AA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA WE02431 ~ATT~~~~AT TAATTG~~AA T~TA~~~ATA A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATCT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA WE02451 ~ATT~~~~AT TAATTA~~AA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA WE02454 ~ATT~~~~AT TAATTG~~AA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA WE02491 ~ATT~~~~AT TAATTG~~AA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA WE02531 ~ATT~~~~AT CAATCG~~AA T~TA~~~ACA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA WE02534 ~ATT~~~~AT TAATYG~~AA T~TA~~~ATA A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATCT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TNNNNNN WE02535 ~ATT~~~~AT TAATTG~~AA T~TA~~~ATA A~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATCT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA WE02591 ~ATT~~~~AT CAATTG~~AW T~TG~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA WE02614 ~ATT~~~~AT CGATTG~~AA T~TG~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCKGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA WE02621 ~ATT~~~~AT TAATTG~~AA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA WE02661 ~ATT~~~~AT CAATTGTGAA T~TG~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA WE02662 ~AATA~~TAT TAATTG~~AA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA WE02671 ~ATT~~~~AT CRATTG~~AA T~TG~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCTGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA WE02674 ~RTT~~~~GT TAATTG~~AA T~TA~~~ATT G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA WE02677 ~ATT~~~~AT TAATTG~~AA T~TA~~~ATA A~~~~~AC~A GTANGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATCT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA ATTTT~A WE85001 ~ATT~~~~AT TAATTG~~AA T~TA~~~ATA G~~~~~AC~A GTAGGCGGAC TCGACGTCCG AAGGGCGCGT CGTCGACGCC GACGTG~~~C CGATTGATTT CGGTCGCGGA GGCGGAGTCG TTGCGCTGAC GGATATCGCG TCTGCCTCAA TTTTTTA WE94001 ~NNN~~~~NN NNNNNN~~NN N~NN~~~NNN N~~~~~NN~N NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNN~~~N NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNNNNN NNNNNNN

199 Legend Plate 1 & 2. Agrodiaetus-wings

Plate 1 Plate 2 Fig. Code Species Fig. Code Species 1 AD 98001 M Agrodiaetus surakovi 1 MW 98261 M Agrodiaetus schuriani 2 AD 98009 M Agrodiaetus pseudactis 2 MW 98284 M Agrodiaetus sigberti 3 DS 00001 M Agrodiaetus poseidonides 3 MW 98285 M Agrodiaetus sigberti 4 DS 00002 M Agrodiaetus damone 4 MW 98294 M Agrodiaetus guezelmavi 5 DS 00003 M Agrodiaetus damocles 5 MW 98313 M Agrodiaetus sertavulensis 6 DS 00004 M Agrodiaetus phyllides 6 MW 99006 M Agrodiaetus firdussii 7 DS 00005 M Agrodiaetus dagmara 7 MW 99009 M Agrodiaetus iphigenia 8 DS 01001 M Agrodiaetus iphigenides 8 MW 99057 M Agrodiaetus merhaba 9 DS 95001 M Agrodiaetus actinides 9 MW 99058 M Agrodiaetus artvinensis 10 JC 00040 M Agrodiaetus aroaniensis 10 MW 99095 M Agrodiaetus huberti 11 JC 00045 M Agrodiaetus nephohiptamenos 11 MW 99105 M Agrodiaetus demavendi 12 JC 01014 M Agrodiaetus admetus 12 MW 99164 M Agrodiaetus interjectus 13 JM 00001 M Agrodiaetus fabressei 13 MW 99196 M Agrodiaetus ripartii 14 MW 00015 M Agrodiaetus demavendi 14 MW 99203 M Agrodiaetus turcicus 15 MW 00032 M Agrodiaetus hamadanensis 15 MW 99240 M Agrodiaetus altivagans 16 MW 00059 M Agrodiaetus elbursicus 16 MW 99274 M Agrodiaetus dantchenkoi 17 MW 00101 M Agrodiaetus darius 17 MW 99286 M Agrodiaetus kurdistanicus 18 MW 00127 M Agrodiaetus paulae 18 MW 99289 M Agrodiaetus sekercioglui 19 MW 00129 M Agrodiaetus gorbunovi 19 MW 99309 M Agrodiaetus baytopi 20 MW 00176 M Agrodiaetus rovshani 20 MW 99341 M Agrodiaetus pierceae 21 MW 00179 M Agrodiaetus cyaneus 21 MW 99353 M Agrodiaetus altivagans 22 MW 00226 M Agrodiaetus femininoides 22 MW 99374 M Agrodiaetus zapvadi 23 MW 00231 M Agrodiaetus alcestis 23 MW 99393 M Agrodiaetus antidolus 24 MW 00234 M Agrodiaetus firdussii 24 MW 99406 M Agrodiaetus antidolus 25 MW 00262 M Agrodiaetus klausschuriani 25 MW 99408 M Agrodiaetus hopfferi 26 MW 00269 M Agrodiaetus iphidamon 26 MW 99413 M Agrodiaetus firdussii 27 MW 00330 M Agrodiaetus pseudoxerxes 27 MW 99448 M Agrodiaetus cyaneus 28 MW 00347 M Agrodiaetus posthumus 28 MW 99465 M Agrodiaetus kanduli 29 MW 00348 M Agrodiaetus phyllis 29 MW 99471 M Agrodiaetus dantchenkoiXmenalcas 30 MW 00393 M Agrodiaetus erschoffii 30 MW 99479 M Agrodiaetus turcicola 31 MW 00409 M Agrodiaetus caeruleus 31 MW 99494 M Agrodiaetus menalcas 32 MW 00426 F Agrodiaetus caeruleus 32 MW 99501 M Agrodiaetus putnami 33 MW 00498 M Agrodiaetus valiabadi 33 MW 99508 M Agrodiaetus ninae 34 MW 00539 M Agrodiaetus dizinensis 34 MW 99565 M Agrodiaetus tankeri 35 MW 00547 M Agrodiaetus birunii 35 MW 99591 M Agrodiaetus humedasae 36 MW 01014 M Agrodiaetus ripartii 36 MW 99613 M Agrodiaetus damon 37 MW 01053 M Agrodiaetus ainsae 37 WE 00002 M Agrodiaetus glaucias 38 MW 01107 M Agrodiaetus fulgens 38 WE 02451 M Agrodiaetus tenhageni 39 MW 98009 M Agrodiaetus carmon 39 WE 02454 M Agrodiaetus mofidii 40 MW 98079 M Agrodiaetus lycius 40 WE 02531 M Agrodiaetus zarathustra 41 MW 98097 M Agrodiaetus ernesti 41 WE 02535 M Agrodiaetus lorestanus 42 MW 98103 M Agrodiaetus iphicarmon 42 WE 02536 M Agrodiaetus damalis 43 MW 98136 M Agrodiaetus wagneri 43 WE 02591 M Agrodiaetus peilei 44 MW 98154 M Agrodiaetus poseidon 44 WE 02612 M Agrodiaetus karindus 45 MW 98162 M Agrodiaetus actis 45 WE 02671 M Agrodiaetus femininoides 46 MW 98170 M Agrodiaetus maraschi 1a 1c 2 3 4 5 47 MW 98172 M Agrodiaetus menalcas 1b 1d 48 MW 98189 M Agrodiaetus hopfferi 6 7 8 9 10 49 MW 98203 M Agrodiaetus mithridates : : : : : 50 MW 98240 M Agrodiaetus theresiae a & b: upperside; c & d: underside; a & c: forewing; b & d:hindwing

201 Index Plate 1 & 2. Agrodiaetus-wings

Index A-H Index I-Z Species Plate Fig. Code Sex Species Plate Fig. Code Sex actinides 1 9 DS 95001 M interjectus 2 12 MW 99164 M actis 1 45 MW 98162 M iphicarmon 1 42 MW 98103 M admetus 1 12 JC 01014 M iphidamon 1 26 MW 00269 M ainsae 1 37 MW 01053 M iphigenia 27 MW 99009 M alcestis 1 23 MW 00231 M iphigenides 18 DS 01001 M altivagans 2 15 MW 99240 M kanduli 2 28 MW 99465 M altivagans 2 21 MW 99353 M karindus 2 44 WE 02612 M antidolus 2 23 MW 99393 M klausschuriani 1 25 MW 00262 M antidolus 2 24 MW 99406 M kurdistanicus 2 17 MW 99286 M aroaniensis 1 10 JC 00040 M lorestanus 2 41 WE 02535 M artvinensis 2 9 MW 99058 M lycius 1 40 MW 98079 M baytopi 2 19 MW 99309 M maraschi 1 46 MW 98170 M birunii 1 35 MW 00547 M menalcas 1 47 MW 98172 M caeruleus 1 31 MW 00409 M menalcas 2 31 MW 99494 M caeruleus 1 32 MW 00426 F merhaba 28 MW 99057 M carmon 1 39 MW 98009 M mithridates 1 49 MW 98203 M cyaneus 1 21 MW 00179 M mofidii 2 39 WE 02454 M cyaneus 2 27 MW 99448 M nephohiptamenos 1 11 JC 00045 M dagmara 1 7 DS 00005 M ninae 2 33 MW 99508 M damalis 2 42 WE 02536 M paulae 1 18 MW 00127 M damocles 1 5 DS 00003 M peilei 2 43 WE 02591 M damon 2 36 MW 99613 M phyllides 16 DS 00004 M damone 1 4 DS 00002 M phyllis 1 29 MW 00348 M dantch.Xmenal. 2 29 MW 99471 M pierceae 2 20 MW 99341 M dantchenkoi 2 16 MW 99274 M poseidon 1 44 MW 98154 M darius 1 17 MW 00101 M poseidonides 13 DS 00001 M demavendi 1 14 MW 00015 M posthumus 1 28 MW 00347 M demavendi 2 11 MW 99105 M pseudactis 12 AD 98009 M dizinensis 1 34 MW 00539 M pseudoxerxes 1 27 MW 00330 M elbursicus 1 16 MW 00059 M putnami 2 32 MW 99501 M ernesti 1 41 MW 98097 M ripartii 1 36 MW 01014 M erschoffii 1 30 MW 00393 M ripartii 2 13 MW 99196 M fabressei 1 13 JM 00001 M rovshani 1 20 MW 00176 M femininoides 1 22 MW 00226 M schuriani 21 MW 98261 M femininoides 2 45 WE 02671 M sekercioglui 2 18 MW 99289 M firdussii 1 24 MW 00234 M sertavulensis 25 MW 98313 M firdussii 2 6 MW 99006 M sigberti 22 MW 98284 M firdussii 2 26 MW 99413 M sigberti 23 MW 98285 M fulgens 1 38 MW 01107 M surakovi 11 AD 98001 M glaucias 2 37 WE 00002 M tankeri 2 34 MW 99565 M gorbunovi 1 19 MW 00129 M tenhageni 2 38 WE 02451 M guezelmavi 2 4 MW 98294 M theresiae 1 50 MW 98240 M hamadanensis 1 15 MW 00032 M turcicola 2 30 MW 99479 M hopfferi 1 48 MW 98189 M turcicus 2 14 MW 99203 M hopfferi 2 25 MW 99408 M valiabadi 1 33 MW 00498 M huberti 2 10 MW 99095 M wagneri 1 43 MW 98136 M humedasae 2 35 MW 99591 M zapvadi 2 22 MW 99374 M zarathustra 2 40 WE 02531 M

202 Plate 1. Agrodiaetus-wings

Plate 2. Agrodiaetus-wings

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