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Additions to the Bryophyte Flora of the Republic of Northern Macedonia

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Additions to the Bryophyte Flora of the Republic of Northern Macedonia cryptogamie Bryologie 2019 ● 40 ● 15 Additions to the bryophyte flora of the Republic of Northern Macedonia Beáta PAPP, Jovana PANTOVIC´ & Marko S. SABOVLJEVIC´ DIRECTEUR DE LA PUBLICATION : Bruno David, Président du Muséum national d’Histoire naturelle RÉDACTEURS EN CHEF / EDITORS-IN-CHIEF : Denis LAMY ASSISTANTS DE RÉDACTION / ASSISTANT EDITORS : Marianne SALAÜN ([email protected]) MISE EN PAGE / PAGE LAYOUT : Marianne SALAÜN RÉDACTEURS ASSOCIÉS / ASSOCIATE EDITORS Biologie moléculaire et phylogénie / Molecular biology and phylogeny Bernard GOFFINET Department of Ecology and Evolutionary Biology, University of Connecticut (United States) Mousses d’Europe / European mosses Isabel DRAPER Centro de Investigación en Biodiversidad y Cambio Global (CIBC-UAM), Universidad Autónoma de Madrid (Spain) Francisco LARA GARCÍA Centro de Investigación en Biodiversidad y Cambio Global (CIBC-UAM), Universidad Autónoma de Madrid (Spain) Mousses d’Afrique et d’Antarctique / African and Antarctic mosses Rysiek OCHYRA Laboratory of Bryology, Institute of Botany, Polish Academy of Sciences, Krakow (Pologne) Bryophytes d’Asie / Asian bryophytes Rui-Liang ZHU School of Life Science, East China Normal University, Shanghai (China) Bioindication / Biomonitoring Franck-Olivier DENAYER Faculté des Sciences Pharmaceutiques et Biologiques de Lille, Laboratoire de Botanique et de Cryptogamie, Lille (France) Écologie des bryophytes / Ecology of bryophyte Nagore GARCÍA MEDINA Department of Biology (Botany), and Centro de Investigación en Biodiversidad y Cambio Global (CIBC-UAM), Universidad Autónoma de Madrid (Spain) Cryptogamie, Bryologie est indexé dans / Cryptogamie, Bryologie is indexed in: – Biological Abstracts – Current Contents – Science Citation Index – Publications bibliographiques du CNRS (Pascal). Cryptogamie, Bryologie est distribué en version électronique par / Cryptogamie, Bryologie is distributed electronically by: – BioOne® (http://www.bioone.org) Cryptogamie, Bryologie est une revue en flux continu publiée par les Publications scientifiques du Muséum, Paris Cryptogamie, Bryologie is a fast track journal published by the Museum Science Press, Paris Les Publications scientifiques du Muséum publient aussi / The Museum Science Press also publish: Adansonia, Geodiversitas, Zoosystema, Anthropozoologica, European Journal of Taxonomy, Naturae, Cryptogamie sous-sections Algologie, Mycologie. Diffusion – Publications scientifiques Muséum national d’Histoire naturelle CP 41 – 57 rue Cuvier F-75231 Paris cedex 05 (France) Tél. : 33 (0)1 40 79 48 05 / Fax : 33 (0)1 40 79 38 40 [email protected] / http://sciencepress.mnhn.fr © Publications scientifiques du Muséum national d’Histoire naturelle, Paris, 2019 ISSN (imprimé / print) : 1290-0796 / ISSN (électronique / electronic) : 1776-0992 Additions to the bryophyte flora of the Republic of Northern Macedonia Beáta PAPP Botanical Department, Hungarian Natural History Museum, H-1431, Budapest, Pf. 137 (Hungary) Jovana PANTOVIĆ Marko S. SABOVLJEVIĆ Institute of Botany and Botanical Garden, Faculty of Biology, University of Belgrade, Takovska 43, 11000 Belgrade (Serbia) Submitted on 4 September 2018 | Accepted on 20 March 2019 | Published on 9 October 2019 Papp B., Pantović J. & Sabovljenić M. S. 2019. — Additions to the bryophyte flora of the Republic of Northern Mac- edonia. Cryptogamie, Bryologie 40 (15): 159-166. https://doi.org/10.5252/cryptogamie-bryologie2019v40a15. http:// cryptogamie.com/bryologie/40/15 ABSTRACT The bryophyte flora research in the Republic of Northern Macedonia has been neglected for decades. KEY WORDS Here, we report 18 new records, namely two liverworts and 16 moss species for this territory. For each Chorology, liverworts, new species presented here, the next information is given: collection data, short notes on distribution mosses, in Europe, the Balkan countries and the Mediterranean Region, as well as frequency and IUCN threat new records, categories in the Balkan countries if any. For taxonomically interesting species the peculiarities such as Balkans, Mediterranean, diagnostic morphological characters are explained. With these new records the bryophyte flora of the SE Europe. Republic of Northern Macedonia counts 575 species (one hornwort, 104 liverworts and 470 mosses). RÉSUMÉ Additions à la flore bryologique de la République de Macédoine du Nord. L’étude de la flore bryologique de la République de Macédoine du Nord a longtemps été négligée. MOTS CLÉS Dans cet article, deux hépatiques et 16 mousses ont été nouvellement récoltées sur ce territoire. Chorologie, hépatiques, Pour chacune sont précisées les données de récolte, la distribution en Europe, dans les Balkans et la mousses, Région méditerranéenne ainsi que leur fréquence et leur niveau de menace selon l’UICN dans les nouveaux signalements, Balkans. Pour les espèces taxonomiquement intéressantes les particularités et les caractères morpho- Balkans, Mediterranée, logiques diagnostiques sont ajoutés. Avec ces nouveaux reports la flore bryologique de la République SE Europe. de Macédoine du Nord comprend 575 espèces (une anthocérote, 104 hépatiques et 470 mousses). CRYPTOGAMIE, BRYOLOGIE • 2019 • 40 (15) © Publications scientifiques du Muséum national d’Histoire naturelle, Paris. www.cryptogamie.com/bryologie 159 Papp B. et al. INTRODUCTION Ellis et al. 2014, 2017). Sabovljević et al. (2011) concluded that the Republic of Northern Macedonia, although still very The Republic of Northern Macedonia (previously Former poorly explored bryologically, given the number of reported Yugoslav Republic of Macedonia or the Republic of Macedo- species in relation to the small surface area of the country, nia) is situated in SE Europe, in the central south part of the belongs to one of the richest countries in Europe regarding Balkan Peninsula. It occupies an area of 25 713 km2, 40% of the bryophyte flora diversity. which is forested, 51% is in agriculture use and 9% is non- The aim of present study was to contribute to the knowl- productive; its surface is composed of a complex mosaic of edge of underexplored bryophyte flora of Northern Macedo- various metamorphic, sedimentary and igneous rocks; the relief nia, and here we present a significant contribution i.e. new structure is very diverse and rather interesting, represented by national records. mountains, valleys, ravines, narrow gorges, saddles and other forms, being mountains cover approximately two-thirds of the land; due to specific natural and geographic characteristics, MATERIAL AND METHODS the prevailing climates country are the Mediterranean and the continental (in Spirovska 2003). The studied samples were collected in a part of the Scardo- Mihajlov et al. (2011) reported the biodiversity of Mac- Pindic massive, namely Galičica Mt, where the focus of edonia to be characterized by high heterogeneity and high bryological investigation was in October 2016. levels of relict and endemic species. Gaston & David (1994) The voucher specimens are deposited in the Bryophyte col- and Spirovska (2003) stated that analyses of biodiversity lection of the Hungarian Natural History Museum, Budapest richness for individual countries within the European con- (BP). tinent rank the Republic of Northern Macedonia at the The nomenclature for liverworts and mosses follows Ros very top of the list of countries considered to be one of the et al. (2007, 2013) respectively, except for Fissidens incurvus European Hot Spots. Starke ex Röhl., where we accept the concept of Erzberger The first conspectus of bryophytes for the present territory (2016), and the Ephemerum serratum (Hedw.) Hampe com- of the Republic of Northern Macedonia made by Pavletić plex, for which we follow Ellis & Price (2015). (1955) counted 213 species (no hornworts, 27 liverworts and 186 mosses). Later on, Martinčič (1968) reported 267 species (in total 285 with infraspecific taxa). Spirovska (2003) reported STUDY AREA 349 species (one hornwort, 52 liverworts and 296 mosses) Galičica Mt is a National Park founded in 1958, which among which two were stated to be endemic (namely Mieli- covers 227.5 km², situated in the SW part of Northern choferia paradoxa Herzog and Orthotrichum insidiosum Her- Macedonia between the two biggest lakes namely Ohrid (in zog). At present M. paradoxa is still considered to be endemic the E) and Prespa (in the W) and the Albanian border in and its accepted name is Brachymenium paradoxum (Herzog) the S (Fig. 1). The range extends for 35 km long stretching A.J. Shaw (Shaw 1987), while O. insidiosum is considered to from N to S, its highest peak is Magaro reaching 2254 m be a synonym of the widely distributed O. rupestre Schleich. altitude; it has poor hydrology on mainly calcareous bed- ex Schwaegr (Lewinsky 1993). Cekova (2005) recorded rock; the two above mentioned lakes are a kind of thermal 397 bryophyte species in the Republic of Northern Mac- regulator, which moderates the climate in this region, not edonia (one hornwort, 66 liverworts and 330 mosses). In permitting extremely high or low temperatures, hence this the check-list of SE European liverworts and hornworts, region is under the influence of moderate continental or Sabovljević & Natcheva (2006) reported 69 species (one atypical sub-Mediterranean climate (Custerevska 2016). It hornwort and 68 liverworts), and Sabovljević et al. (2008) is a high-mountain region considered as the center of high reported 345 mosses species for Northern Macedonia. In the floristic diversity of vascular plants (over 1000 species) check list of Mediterranean
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