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Revista Chilena De Entomología Rev. Chilena Ent. 1989, 17: 5-60 ESTADO ACTUAL DEL CONOCIMIENTO DE LOS COLEÓPTEROS DE CHILE (INSECTA: COLEÓPTERA) Mario Elgueta D.^ y Gerardo Arriagada S.^ RESUMEN A base de una recopilación bibliográfica y conocimiento del grupo, se entrega una visión actualizada del estado de investigación de los Coleóptera de Chile. Se incluye un esquema de clasificación de las familias consideradas válidas y se tratan por separado todas aquellas representadas en Chile; para cada una de ellas se da número de género y especies, incluyendo antecedentes biológicos conocidos. En el caso de familias con pocos representantes, se ofrece un listado taxonómico, siempre que no existan catálogos recientes o se haya señalado expresamente los géneros y número de especies que los constituyen. Georyssidae, Eucinetidae y Limnichidae, los Curculionidae Neopachytychius squamosus Hustache y Naupactus verecundus Hustache y el Silphidae Oxelytrum anticola (Guérin), son citados por primera vez para Chile. Se mencionan los siguientes taxa omitidos en el catálogo de Blackwelder (1944-1957), indicando entre paréntesis el grupo en el cual son tratados en este aporte; se propone además una nueva sinonimia: ABSTRACT Based upon bibliographical review and practical knowledge of this group, an actualized view of the investigation state of chilean Coleóptera is given. A classification of the valid families is included. AU families that are present in Chile are separately analized, and biological data, genera and species numbers are detailed. For small families without recent revisión or checklist, a taxonomic list is given. Georyssidae, Eucinetidae and Limnichidae, the weevils Neopachytychius squamosus Hustache and Naupactus verecundus Hustache, and the carrion beetle Oxelytrum anticola (Guérin) existent in Chile are cited for the first time. Fourteen taxa omitted in Blackwelder's Checklist ( 1 944- 1 957) are mentioned, between parenthesis the group in which these taxa are cited, and a new synonymy is proposed: Brachidia gallerucoides Germain, 1855 (Melyridae) Philophlaeus Germain, 1855 (Biphyllidae - Languriidae) Melanophthalma seminigra Belon: Philippi, 1887 (Lathridiidae) Endophloeus angustatus Solier, 1851 (Colydiidae) Cis chilensis Germain, 1855 (Ciidae) Promecheilus Solier, 1 85 1 (Oedemeridae) Promecheilus variegatus Solier, 1851 (Oedemeridae) Nephrosis Fairmaire &: Germain, 1863 (Oedemeridae) Nephrosis sulcicollis Fairmaire &: Germain, 1863 (Oedemeridae) Oligorrhina Fairmaire &: Germain, 1863 (Oedemeridae) Oligorrhina ruficollis Fairmaire &: Germain, 1863 (Oedemeridae) Nimales Fairmaire & Germain, 1861 (Oedemeridae) Nimales longicornis Fairmaire 8c Germain, 1861 (Oedemeridae) Pseudomeloe machadoi Germain, 1903 (Meloidae) Enneboeus baeckstroemi Pie, 1924 = Archeocrypticvs chilensis Kaszab, 1969 nov. syn. (Archeocrypticidae). 'Sección Entomología, Museo Nacional de Historia Natural, Casilla 787, Santiago-Chile. ^Sociedad Chilena de Entomología, Casilla 21 132, Santiago-Chile. (Recibido: 1 de septiembre de 1988. Aceptado: 20 de septiembre de 1988). Rev. Chilena Ent. 17, 1989 CONTENIDO guos {i.e.: Bréthes, 1925; Fairmaire 8c Ger- main, 1861, 1863; Gemminger & Harold, Pág- 1870; Germain, 1855, 1903; Philippi, 1887; Introducción 6 Solier, 1851). Se ha utilizado, además, como Lista sistemática de las familias del Or- fuente de información adicional, la colección den Coleóptera 7 de Coleóptera del Museo Nacional de Historia Análisis de las familias representadas en Natural (Santiago-Chile), en la cual se encuen- Chile 9 tra el material estudiado e identificado por Recuento del número de géneros y espe- Philibert Germain. cies por familias presentes en Chile . 38 Agradecimientos 39 Se presenta en primer lugar una clasifica- ción Literatura citada 39 actualizada de Coleóptera, aceptada con algunas índice alfabético 53 modificaciones y en forma más o me- nos universal, hasta el nivel de familia; se INTRODUCCIÓN indica con un asterisco las familias para las cuales se conocen representantes en Chile. Di- Para los interesados en el estudio de los co- cha clasificación está basada en los aportes de leópteros de Chile, una obra clásica de consul- Crowson (1955, 1964, 1967, 1970, 1971, 1972, ta es el catálogo de los coleópteros de México, 1973b, 1975, 1981, 1984), Lawrence (1982), América Central, Indias Occidentales y Sud- Lawrence & Newton (1982), Newton (1985), américa (Blackwelder, 1944-1957); sin embar- Sen Gupta 8c Crowson (1966, 1969a, 1969b, go, durante los transcurridos más de 30 años 1973) y Watt (1967a, 1967b, 1974a, 1974b, desde su publicación, se han efectuado nume- 1987), apoyándose también en las contribu- rosas revisiones que involucran descripciones ciones de Abdullah (1969a, 1971), Britton de nuevos taxa, sinonimias transferencias y (1974, 1979), Pal 8c Lawrence (1986) y Scholtz como, asimismo, estudios que han permitido etal. (1987). precisar más claramente los límites de catego- Luego cada familia con representantes en rías supragenéricas. Chile es tratada por separado, en orden siste- No todas estas contribuciones son conoci- mático, señalando antecedentes biológicos dis- das o están a disposición de los investigadores ponibles y un detalle de los taxa conocidos en nacionales; es por ello que se ha estimado de nuestro territorio; en varios casos se entregan interés dar a conocer la información recopila- antecedentes inéditos y se mencionan aquellos da por el personal de la Sección Entomología taxa omitidos en catálogos previos. Se incluye del Museo Nacional de Historia Natural, a en esta parte, la información relativa a Co- través de años de paciente búsqueda. leóptera fósil y Strepsiptera; aun cuando en En esta oportunidad se intenta entregar este último caso no existe un consenso general una visión global del conocimiento actual de para considerarlo como coleópteros (Crow- los coleópteros de Chile, señalando las fami- son, 1955, 1981; Lawrence 8c Newton, 1982; lias representadas, el número de géneros y Riek, 1974, 1979), evidentemente se trata de especies para cada una una lista sistemática y un grupo relacionado. de los taxa de aquellas familias poco numero- En tercer lugar se hace un listado de espe- sas, que no tengan revisiones o catálogos re- cies de aquellas familias poco representadas cientes, o para las cuales no se haya hecho una en nuestra fauna. Por último, se entrega un relación de los géneros que la componen y del recuento del número de géneros especies de número de especies de éstos. y cada las familias representadas en La ejecución de este trabajo está basado en una de Chile. la contribución de Blackwelder (1944-1957) y la información publicada con posterioridad; En la Literatura citada se incluyen las publi- sin embargo, y debido al hecho de haberse caciones posteriores a la contribución de detectado omisiones de taxa propios de Chile, Blackwelder ( 1 944- 1 957) y aquellas anteriores citados o descritos con anterioridad a la publi- que fueron omitidas en este catálogo, en las cación de dicho trabajo, se ha debido revisar cuales se citan o describen taxa presentes en las publicaciones originales y catálogos anti- Chile. Parte de la información recopilada ha . Elgueta y Arriagada: Conocimiento actual Coleóptera de Chile sido extraída de Zoological Record's (años tracts(años 1967, 1968, 1970, 1972, 1973 (sólo 1947 a 1971; excluyendo 1967, 1968 y 1970), volumen 56), 1982 a 1988; hasta vol. 88, nú- Abstraéis of Entomology (años 1970 a 1982; mero 1 ); en las respectivas citas se indica cuan- excluyendo 1972 y 1973) y Biological Abs- do el trabajo no ha sido visto. LISTA SISTEMÁTICA DE LAS FAMILIAS DEL ORDEN COLEÓPTERA (ENTRE PARÉNTESIS SINÓNIMOS Y/O TAXA INCLUIDOS) El asterisco indica las familias representadas en Chile Suborden ARCHOSTEMATA 28. Agyrtidae Superfamilia CUPEDOIDEA 29. Leiodidae (Anisotomidae, Camiaridae, 1. Ommatidae (Tetraphaleridae, Omma- Catopidae, Cholevidae, Colonidae, Em- didae) pelidae, Leptinidae, Leptodiridae, Lio- 2. Cupedidae* didae)* 3. Micromalthidae 30. Scydmaenidae* Suborden ADEPHAGA 31. Micropeplidae Superfamilia CARABOIDEA 32. Dasyceridae 4. Rhysodidae 33. Silphidae* 34. 5. Paussidae (incluye Ozaeninae)* Staphylinidae (Brathinidae, Scaphi- 6. Cicindelidae* diidae)* 7. Carabidae* 35. Pselaphidae (Clavigeridae)* 8. Trachypachidae* Superfamilia EUCINETOIDEA 9. Haliplidae* 36. Clambidae (Calyptomeridae) 10. Amphizoidae 37. Eucinetidae* 1 1 Hygrobiidae 38. Scirtidae (Helodidae, Cyphonidae)* 12. Noteridae Superfamilia DASCILLOIDEA 13. Dytiscidae* 39. Dascillidae* 14. Gyrinidae* 40. Karumiidae (Zarudniolidae)* Suborden MYXOPHAGA 41. Rhipiceridae (Sandalidae)* Superfamilia SPHAERIOIDEA Superfamilia SCARABAEOIDEA 15. Lepiceridae (Cyathoceridae) 42. Lucanidae* 16. Torridincolidae 43. Trogidae* 17. Hydroscaphidae 44. Ceratocanthidae (Acanthoceridae)* 18. Sphaeriidae 45. Passalidae Suborden POLYPHAGA 46. Pleocomidae Superfamilia HYDROPHILOIDEA 47. Ochodaeidae 19. Hydraenidae (Limnebiidae)* 48. Hybosoridae* 20. Spercheidae 49. Glaphyridae* 21. Hydrochidae* 50. Scarabaeidae (Geotrupidae)* 22. Georyssidae* 51. Diphyllostomatidae 23. Hydrophilidae* 52. Glaresidae Superfamilia HISTEROIDEA Superfamilia BYRRHOIDEA 24. Sphaeritidae 53. Byrrhidae* 25. Synteliidae Superfamilia DRYOPOIDEA 26. Histeridae (Niponiidae)* 54. Eulichadidae Superfamilia STAPHYLINOIDEA 55. Ptilodactylidae 27. Ptiliidae (Cephaloplectidae, Limulo- 56. Chelonariidae didae)* 57. Psephenidae (incluye Eubriinae)* Rev. Chilena Ent. 17, 1989 58. Elminthidae (Elmidae, Helminthidae)* Superfamilia LYMEXYLOIDEA 59. Lutrochidae 99. Lymexylidae* 60. Dryopidae (Chiloeidae)*
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