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Lech Borowiec Department of Biodiversity and Evolutionary Taxonomy University of Wrocław Cytacje - Citations (cytacje w czasopismach indeksowanych zaznaczone na czerwono – citations in journals indexed by ISI marked with red) 1977 1. Materiały do znajomości muchówek z rodziny wpleszczowatych (Diptera, Hippoboscidae) pasożytujących na ptakach w Polsce. Pol. Pismo ent., 47: 37-42. 1. Draber-Mońko A., 2007. Wpleszczowate (Hippoboscidae). Pp. 169-172, w: Fauna Polski – charakterystyka i wykaz gatunków (ed. Bogdanowicz et al.). Muzeum i Instytut Zoologii PAN, Warszawa, 505 str. 2. Zatwarnicki T., 2020. Bibliografia Dipterologiczna Polski (1597-2020). Dipteron, 37(01): 1-361. 1978 2. Nowe dane o susówkach Polski (Coleoptera, Chrysomelidae, Halticinae) Polski ze szczególnym uwzględnieniem Śląska. Pol. Pismo ent., 48: 423-427. 3. Burakowski B., Mroczkowski M., Stefańska J., 1985. Chrząszcze Coleoptera. Buprestoidea, Elateroidea i Cantharoidea. W: Katalog Fauny Polski, XXIII, 10, 401 pp. 4. Doguet S., 1994. Coleopteres Chrysomelidae. Volume 2 Alticinae. In: Faune de France 80, Paris, 694 pp. 5. Warchałowski A., 1996. Ubersicht der westpalaarktischen Arten der Gattung Longitarsus Berthold, 1827 (Coleoptera: Chrysomelidae: Halticinae). Genus, suppl. 1996: 266 pp. 6. Gruev B., Doeberl M., 1997. General distribution of the flea beetles in the Palaearctic subregion (Coleoptera: Chrysomelidae: Alticinae). Scopolia: 37: 1-496. 7. Ścibior R., 2002. Drugie stanowisko Longitarsus fulgens (Foudras, 1860) (Coleoptera: Chrysomelidae: Alticinae) w Polsce. Wiad. ent., 20 (2001): 176. 8. Lopatin I.K., Aleksandrovich O.R., Konstantinov, A.S. 2004. Check list of Leaf-beetle Chrysomelidae (Coleoptera) of the eastern Europe and northern Asia. Mantis, Olsztyn, 343 pp. 9. Kisiel P. i inni, 2015. Świat zwierząt. W: A. Żelaźniewicz (red.), Przyroda Dolnego Śląska. Polska Akademia Nauk, Oddział we Wrocławiu, 321-374 str. 3. Przegląd muchówek z rodziny wpleszczowatych (Diptera, Hippoboscidae) pasożytujących na ptakach w Polsce. Pol. Pismo ent., 48: 565-572. 10. Draber-Mońko A., 2007. Wpleszczowate (Hippoboscidae). Pp. 169-172, w: Fauna Polski – charakterystyka i wykaz gatunków (ed. Bogdanowicz et al.). Muzeum i Instytut Zoologii PAN, Warszawa, 505 str. 11. Zatwarnicki T., 2020. Bibliografia Dipterologiczna Polski (1597-2020). Dipteron, 37(01): 1-361. 1979 4. Chrysomelidae (Coleoptera) new to the Bulgarian fauna. Pol. Pismo ent., 49: 481-483. 12. Gruev B., 1980. Pregled na vidovite ot podsemeistvata Eumolpinae, Chrysomelinae, Hispinae i Cassidinae na sem. Chrysomelidae (Insecta: Coleoptera) ystanovieni po Bylgarsko Tschernomorie. Trav. Scien. Univ. Plovdiv, 18: 83-96. 13. Gruev B., Tomov V., 1984. Coleoptera, Chrysomelidae Part 1. In: Fauna na Bulgaria 13, Sofia, 220 pp. 14. Gruev B., Tomov V., 1986. Coleoptera, Chrysomelidae Part 2. In: Fauna na Bulgaria 16, Sofia, 388 pp. 15. Warchałowski A., 1991. Chrysomelidae – stonkowate, część II. In: Fauna Polski 13, Warszawa, 347 pp. 16. Sassi D., 1994. Le Cassidine appenniniche del Museo Di Soria Naturale di Verona (Coleoptera Chrysomelidae). Boll. Mus. civ. St. nat. Verona, 18: 53-90. 17. Warchałowski A., 1996. Ubersicht der westpalaarktischen Arten der Gattung Longitarsus Berthold, 1827 (Coleoptera: Chrysomelidae: Halticinae). Genus, suppl. 1996: 266 pp. 18. Gruev B., Doeberl M., 1997. General distribution of the flea beetles in the Palaearctic subregion (Coleoptera: Chrysomelidae: Alticinae). Scopolia: 37: 1-496. 19. Gruev B., Tomov V., 1998. Coleoptera: Chrysomelidae. In: Catalogus Faunae Bulgaricae 3: 160 pp. 20. Bordy B., 2000. Coleopteres Chrysomelidae, volume 3 Hispinae et Cassidinae. Faune de France, 85: 250 pp. + 26 pl. 21. Warchałowski A., 2000. Chrysomelidae – stonkowate, część VII. In: Fauna Polski 22, Warszawa, 357 pp. 22. Gruev B., 2004. The Leaf Beetles (Insecta: Coleoptera: Chrysomelidae) of the Stara Planina Ridge and the Fore-Balkan (Bulgaria). Fauna and Zoogeography. Trav. Sci. Univ. Plovdiv, 40: 37-75. 23. Lopatin I.K., Aleksandrovich O.R., Konstantinov, A.S. 2004. Check list of Leaf-beetle Chrysomelidae (Coleoptera) of the eastern Europe and northern Asia. Mantis, Olsztyn, 343 pp. 24. Gruev B., Doeberl M., 2005. General distribution of the flea beetles in the Palaearctic subregion (Coleoptera: Chrysomelidae: Alticinae).Supplement. Pensoft, Sofia-Moscow: 239 pp. 25. Gruev B., Tomov V., 2007. A distributional atlas and catalogue of the Leaf beetles of Bulgaria. Zoocartographia Balcanica, vol. 3, Pensoft, 350 pp. 6. Otiorhynchus lugdunensis Boheman, 1843 (Col. Curculionidae) nowy gatunek ryjkowca dla fauny Polski. Prz. zool., 23: 326-327. 26. Burakowski B., Mroczkowski M., Stefańska J., 1986. Chrząszcze Coleoptera. Cucujoidea część 2. W: Katalog Fauny Polski, XXIII, 13, 278 pp. 1 1980 7. Eine neue paläarktische Gattung der Samenkäfer (Coleoptera, Bruchidae). Pol. Pismo ent., 50: 127-131. 27. Udayagiri S., Wadhi S.R., 1982. A key to world bruchid genera. NBPGR Sci. Monogr. No. 5, New Delhi, 16 pp. 28. Decelle J., Lodos N., 1989. Contribution to the study of legume weevils of Turkey (Coleoptera: Bruchidae). Bull. Annls Soc. r. belge Ent., 125: 163-212. 29. Anton K.-W., Halperin J., Calderon M., 1997. An annotated list of the Bruchidae (Coleoptera) of Israel and adjacent areas. Israel Journ. Entomol., 31: 59-96. 30. Porca M., 2003. The actual stage the knowledge about the damages bruchides. Journ. Central European Agr., 4, 3: 199-208. 31. Yus Ramos R., 2007. Genera de Coleópteros de la Península Ibérica e Islas Baleares: familia Bruchidae 1 (Coleoptera, Chrysomeloidea). Bol. Asoc. Esp. Ent., 31: 65-114. 32. Anton K.-W. 2010. Subfamily Bruchinae Latreille, 1802. In: I. Löbl, A. Smetana (ed.). Catalogue of Palaearctic Coleoptera. Volume 6. Chrysomeloidea. Apollo Books, 339-353 pp. 33. Yus Ramos R., Fancello L., Coello García P., 2010. Contribución al conocimiento de la fauna de brúquidos (Coleoptera: Bruchidae) de la isla de Cerdeña (Italia). Boletín de la Sociedad Entomológica Aragonesa, 47: 209‒221. 34. Yus Ramos R., 2010. Sobre la presencia de Acanthobruchidius spiniger (Baudi, 1886) en la isla de Cerdena (Italia). Redescription del macho (Coleoptera: Bruchidae). Heteropterous Rev. Entomol., 10: 131-138. 8. Strąkowce-Bruchidae. W: Klucze do oznaczania owadów Polski, część XIX, zeszyt 93. PWN, Warszawa-Wrocław, 51 pp. 35. Zampetti F., 1982. Pozizione sistematica di alcuni species appartenenti al. genre Bruchidius Schilsky (Coleoptera, Bruchidae). Boll. Mus. Civ. Stor. Nat. Verona, 8: 383-410. 36. Wendt H., 1984. Zur Kenntnis der Bruchidenfauna Bulgariens (Col. Phytophaga). Dtsch. Entom. Zietschr., 31: 153-167. 37. Schmitt M., 1985. Versuch einer phylogenetisch-systematischen Analyse der Criocerinae (Coleoptera, Chrysomelidae). Zool. Beitr., 29: 35-85. 38. Burakowski B., Mroczkowski M., Stefańska J., 1986. Chrząszcze Coleoptera. Dermestoidea, Bostrychoidea, Cleroidea i Lymexyloidea. W: Katalog Fauny Polski, XXIII, 11, 243 pp. 39. Niezgodziński P., 1986. Zagadnienie występowania i aklimatyzacji strąkowca fasolowego Acanthoscelides obtectus Say (Col., Bruchidae) w Polsce. Opolskie Tow. Przyj. Nauk, zesz. przyr., 24: 51-61. 40. Wendt H., 1986. Beitrage zur Insektenfauna der DDR: Coleoptera-Bruchidae (Chrysomeloidea). I Zur Biologie und Verbreitung. Mitt. Zool. Mus. Berlin, 62: 103-133. 41. Johnson C.D., 1990. Systematics of the seed beetle genus Acanthoscelides (Bruchidae) of northern South America. Trans. Amer. Entomol. Soc., 116: 297-618. 42. Strejcek J., 1990. Brouci celedi Bruchidae, Urodonidae a Anthribidae. In: Zoologicke klice. Academia, Praha, 87 pp. + XXIV pl. 43. Wendt H., 1993. Bruchus ecalcaratus K. Daniel, 1906 – ein synonym zu Bruchus rufimanus Boheman, 1833 (Coleoptera, Chrysomeloidea, Bruchidae). Dtsch. Ent. Z., 40: 161-165. 44. Biondi M., Daccordi M., Regalin R., Zampetti M.F., 1995. Coleoptera Polyphaga XV (Chrysomelidae, Bruchidae). Checklist delle specie della fauna italiana. Calderini, 34 pp. 45. Romero J., Johnson C.D.., 2000. Revision of the genus Zabrotes Horn of Mexico (Coleoptera : Bruchidae : Amblycerinae). Trans. Am. Entomol. Soc., 126 (2): 221-274. 46. Ruta R., Konwerski Sz., Kubisz D., 2005. Uwagi o krajowych strąkowcach (Coleoptera: Bruchidae). Wiad. entomol., 24: 235-241. 47. Yus Ramos R., 2009. Paleoacanthoscelides gilvus (Gyllenhal, 1839) (Coleoptera: Bruchidae) en la fauna ibero-balear. Revision del genero. Heteropterous Rev. Entomol., 9: 111-122. 48. Niedojad K., 2012. Chrząszcze z nadrodziny Chrysomeloidea (Coleoptera) w Sudetach Środkowych. Przyroda Sudetów, 15: 67-84. 49. Strejcek J., 2012. Bruchidae, Urodontidae. Icones Insectorum Europae Centralis, Folia Heyrovskiana, no. 15: 24 pp. 50. Johnson C.D., Southgate B.J., Delobel A., 2004. A revision of the Caryedontini (Coleoptera:Bruchidae: Pachymerinae) of Africa and the Middle East. Mem. Amer. Entomol. Soc., 44: 120 pp. 51. Zuber M., 2021. Hromadný výskyt dvou druhů kriticky ohrožených zrnokazů Bruchidius cinerascens (Gyllenhal, 1833) a Bruchidius pusillus (Germar, 1824) v dolním Pojizeří (Coleoptera: Chrysomelidae: Bruchinae). Elateridarum, 15: 94-96. 9. A new species of Acanthoscelides Schilsky from Bulgaria (Coleoptera, Bruchidae). Pol. Pismo ent., 50: 167-170. 52. Wendt H., 1984. Zur Kenntnis der Bruchidenfauna Bulgariens (Col. Phytophaga). Dtsch. Entom. Zietschr., 31: 153-167. 53. Anton K.-W., 1991. Neuzumeldende Samenkafer-Arten fur Mitteleuropa (Coleoptera: Bruchidae). Mitt. Entom. Gesellsch. Basel, 41: 97-100. 54. Tuda M., Shima K., Johnson C.D., 2001. Establishment of Acanthoscelides pallidipennis (Coleoptera:
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    Blattkäfer (Coleoptera: Megalopodidae, Orsodacnidae et Chryso- melidae excl. Bruchinae) Bestandssituation. Stand: März 2013 Wolfgang Bäse Einführung Exkremente zum Schutz vor Feinden auf dem Rücken. Nur wenige Blattkäfer-Arten sind durch ihre wirt- Zu den Blattkäfern gehören nach Löbl & Smetana schaftliche Bedeutung allgemein bekannt. Hierzu gehö- (2010) drei Familien. So werden die ehemaligen Un- ren der Kartoffelkäfer, der Rübenschildkäfer (Cassida terfamilien Zeugophorinae als Megalopodidae und die nebulosa), Vertreter der Kohlerdflöhe (Phyllotreta spp.) Orsodacninae als Orsodacnidae interpretiert. Die ur- und die Spargel-, Getreide- und Lilienhähnchen (Crio- sprüngliche Familie der Samenkäfer (Bruchidae) zählt ceris spp., Oulema spp. und Lilioceris spp.). Viele Arten jetzt als Unterfamilie (Bruchinae) zu den Chrysomelidae. sind jedoch durch die Zerstörung naturnaher Standorte In dieser Arbeit fehlen die Samenkäfer, da die Datenlage gefährdet. So waren die Schilfkäfer ursprünglich an die momentan als nicht ausreichend angesehen wird. dynamischen Auenbereiche der Bäche und Flüsse gebun- Zu den größten Käferfamilien der Welt gehörend, sind den. Durch Grundwasserabsenkungen, Uferzerstörung die Blattkäfer ohne Berücksichtigung der Samenkäfer in und intensive Freizeitnutzung wurden viele ursprüng- Deutschland mit 510 Arten (Geiser 1998) vertreten. liche Lebensräume zerstört. Weniger spezialisierte Arten Der Habitus der Blattkäfer ist nicht einheitlich. Ne- sind vielfach noch ungefährdet, da sie auf sekundäre ben dem typischen gewölbten bis eiförmigen Habitus, Lebensräume wie Teiche oder Gräben ausweichen kön- wie er vom Kartoffelkäfer (Leptinotarsa decemlineata) nen. Die seltener nachgewiesenen Arten sind oft hoch- bekannt ist, gibt es bockkäferähnliche Formen bei den spezialisiert. So ist Donacia obscura nur in Mooren zu Schilfkäfern (Donaciinae), flachgedrückte Vertreter bei finden, während Macroplea mutica im Binnenland an den Schildkäfern (Cassida spp.), die eher zylindrisch ge- Salzseen gebunden ist.
  • Alien Beetle Species in the Republic of Moldova: a Review of Their Origin and Main Impact

    Alien Beetle Species in the Republic of Moldova: a Review of Their Origin and Main Impact

    Российской академии наук Институт проблем экологии и эволюции им. А.Н. Северцова (ИПЭЭ РАН) Российский Журнал Биологических Инвазий (ISSN – 1996–1499) Основан в январе 2008 г. Выходит 4 раза в год Главный редактор академик РАН Дгебуадзе Юрий Юлианович Заместитель главного редактора д.б.н., Петросян Варос Гарегинович Ответственный секретарь к.б.н., Дергунова Наталья Николаевна Редакционная коллегия к.б.н., В.В. Бобров, д.б.н., Ю.К. Виноградова, д.б.н., А.Ю. Звягинцев, д.б.н., С.С. Ижевский, д.б.н., И.Н. Ильин, д.б.н., Крылов А.В., к.б.н., В.Ю. Масляков, к.б.н., О.В. Морозова, академик РАН, Д.С. Павлов, д.б.н., А.Н. Пельгунов, д.б.н., Н.М. Пронин, к.б.н., Ю.В. Слынько, д.б.н., И.В. Телеш, к.б.н., И.Ю. Фенева, к.б.н., Л.А. Хляп, д.б.н., Шиганова Т.А., д.б.н., Г.Х. Щербина Тематика журнала Теоретические вопросы биологических инвазий (теория, моделирование, результаты наблюдений и экспериментов): инвазионные коридоры, векторы инвазий, адаптации видов– вселенцев, уязвимость аборигенных экосистем, оценка риска инвазий, генетические, экологические, биологические, биогеографические и эволюционные аспекты влияния чужеродных видов на биологическое разнообразие биосистем различных уровней организации. Мониторинг инвазионного процесса (сообщения о нахождении организмов за пределами естественного ареала, динамике расселения, темпах натурализации). Методы, средства накопления, обработки и представления данных прикладных исследований (новые разработки, моделирование, результаты исследований) с применением фактографических и геоинформационных систем. Использование результатов исследований биологических инвазий (методы и новые фундаментальные результаты) при изучении морских, пресноводных и наземных видов, популяций, сообществ и экосистем. Контроль, рациональное использование и борьба с видами вселенцами.
  • Biology of the Bruchidae +6178

    Biology of the Bruchidae +6178

    Ann. Rev. Entomol 1979. 24:449-73 Copyright @ 1979 by Annual Reviews Inc. All rights reserved BIOLOGY OF THE BRUCHIDAE +6178 B. J. Southgate Biology Department, Pest Infestation Control Laboratory, Ministry of Agriculture, Fisheries, and Food, Slough SL3 7HJ, Berks, England INTRODUCTION Species of Bruchidae breed in every continent except Antarctica. The larg­ est number of species live in the tropical regions of Asia, Africa, and Central and South America. Many species have obvious economic importance because they breed on grain legumes and consume valuable proteins that would otherwise be eaten by man. Other species, however, destroy seeds of an immense number of leguminous trees and shrubs, which, though they have no obvious economic value, stem the advance of the deserts into the marginal cultivated areas of the world. When this ecosystem is mismanaged by practices such as over­ grazing, then any organism that restricts the normal regeneration of seed­ lings will, in the long run, affect agriculture adversely. This has been demonstrated recently in some African and Middle Eastern semiarid zones (65). The present interest in the management of arid areas and in the introduc­ Annu. Rev. Entomol. 1979.24:449-473. Downloaded from www.annualreviews.org Access provided by Copyright Clearance Center on 11/01/20. For personal use only. tion of alternative tree species to provide timber, fodder, or shade has stimulated a detailed study of the ecology of some leguminous trees and shrubs that has revealed some deleterious effects of bruchid beetles on the seeds of these plants (42, 43, 59). It has also emphasized the inadequacy of our knowledge of the taxonomy and biology of these beetles.
  • Six New Species of the Genus Agroiconota Spaeth, 1913 (Coleoptera: Chrysomelidae: Cassidinae), with a Key to the Genus

    Six New Species of the Genus Agroiconota Spaeth, 1913 (Coleoptera: Chrysomelidae: Cassidinae), with a Key to the Genus

    ANNALES ZOOLOGICI (W arszawa), 2005, 55(1): 61-74 SIX NEW SPECIES OF THE GENUS AGROICONOTA SPAETH, 1913 (COLEOPTERA: CHRYSOMELIDAE: CASSIDINAE), WITH A KEY TO THE GENUS L e c h B o r o w ie c Zoological Institute, University of Wroclaw, Przybyszewskiego 63/77, 51-148 Wroclaw, Poland; e-mail: [email protected] Abstract. — Agriconota atromaculata (Peru), A. atropunctata (Bolivia), A. carlobrivioi (Bolivia, Brazil, Peru), A. gibbipennis (Brazil), A. paraguayana (Paraguay), and A. sanareen- sis (Venezuela), new to the science, are described. New records of several species, colour photos of all species, and a key to the genus Agroiconota Spaeth, 1913 are given. Key words.— Entomology, taxonomy, new species, new records, Coleoptera, Chrysomelidae, Cassidinae, Agroiconota, Neotropical Region. Introduction Ta x n o n o m y The genus Agroiconota was proposed by Spaeth Agroiconota Spaeth, 1913 (1913) for Cassida tristriata Fabricius, 1792 (type spe­ cies by original designation) and Coptocycla stupidula Agroiconota Spaeth, 1913:142 (type species:Cassida tristriata Fabricius, 1792, by original designation), 1914: 121, Hincks 1952: 341, Seeno Boheman, 1855. In 1936, Spaeth reviewed the genus in and Wilcox 1982: 178, Riley 1986: 106, Borowiec 1999: 322. key form. He keyed 16 species, including 6 new to the science. Next year, Spaeth (1937) described another new The genus Agroiconota Spaeth, 1913 is a m em ber of species in the genus. Blake (1970) described a new species the tribe Cassidini. It is well distinguished from other from Cuba. At last, Buzzi (1996) described Agroiconota genera of the tribe by the following combination of char­ urbanae from Brazil (Mato Grosso) but Borowiec (1998) acters: venter of pronotum without antennal grooves, synonymised it with A.
  • The Cassidinae Beetles of Longnan County (Jiangxi, China): Overview and Community Composition

    The Cassidinae Beetles of Longnan County (Jiangxi, China): Overview and Community Composition

    Biodiversity Data Journal 7: e39053 doi: 10.3897/BDJ.7.e39053 Research Article The Cassidinae beetles of Longnan County (Jiangxi, China): overview and community composition Peng Liu‡, Chengqing Liao‡‡, Jiasheng Xu , Charles L. Staines§, Xiaohua Dai ‡,| ‡ Leafminer Group, School of Life Sciences, Gannan Normal University, Ganzhou, China § Smithsonian Environmental Research Center, Edgewater, United States of America | National Navel-Orange Engineering Research Center, Ganzhou, China Corresponding author: Xiaohua Dai ([email protected]) Academic editor: Flávia Rodrigues Fernandes Received: 13 Aug 2019 | Accepted: 16 Oct 2019 | Published: 18 Oct 2019 Citation: Liu P, Liao C, Xu J, Staines CL, Dai X (2019) The Cassidinae beetles of Longnan County (Jiangxi, China): overview and community composition. Biodiversity Data Journal 7: e39053. https://doi.org/10.3897/BDJ.7.e39053 Abstract There are few reports on the community composition and diversity pattern of the Cassidinae species of China. Compared to the neighbouring provinces of Guangdong, Fujian and Zhejiang, the Cassidinae richness in Jiangxi Province is under-reported. Longnan City, a biodiversity hotspot in Jiangxi Province, was chosen to obtain the first overview of the Cassidinae beetles. The sample coverage curves for the three sample sites reached an asymptote which indicated sampling was sufficient for data analysis. A total of eight tribes, 16 genera, 59 species and 1590 individuals of Cassidinae beetles were collected. Most belonged to the tribe Hispini (1121 individuals; 70.5%), followed by the tribe Cassidini (161 individuals; 10.13%) and the tribe Oncocephalini (159 individuals; 10.0%). The remainder (149 individuals) belonged to five tribes (Gonophorini, Basiprionotini, Callispini, Notosacanthini and Aspidimorphini). The tribes Notosacanthini, Aspidimorphini and Oncocephalini were newly recorded for Jiangxi Province.
  • Download PDF File

    Download PDF File

    Myrmecological News 23 41-59 Vienna, September 2016 Analyzing large-scale and intranidal phenotype distributions in eusocial Hymenoptera – a taxonomic tool to distinguish intraspecific dimorphism from heterospecificity Bernhard SEIFERT Abstract Ant and termite nests are long-term stable, semi-closed systems constantly producing conspecific worker populations of related individuals over many generations. Accordingly, nests of these eusocial insects, as they are found in nature, offer free of cost an analysis situation that has to be generated in other groups of organisms by controlled rearing experiments. A test system based on analyzing intranidal and large-scale phenotype distributions and comparing the observed distributions with predictions for different scenarios of heterospecificity and intraspecific dimorphism is introduced by a case study on ants. The test system, named DIMORPH test, allows a taxonomist to distinguish if discrete character syndromes represent separate species or an intraspecific phenomenon. One of the most important parameters within the test system is the abundance and distribution of phenotypically mixed nest populations. Five biological explanations are possible for ant nests with a mixture of discrete phenotypes: They may represent (1) geneti- cally determined intraspecific morphs, (2) intraspecific modifications induced by environmental factors, (3) the associ- ation of a temporary social parasite with a host species, (4) the association of a permanent social parasite with a host species, and (5) a parabiotic association of two basically independent (self-sustaining) species. The paper explains the biological background of the scenarios (1) to (5) and presents mathematical models and generalizations from empirical data to predict phenotype distributions for each scenario under variable conditions.