Göteborgs naturhistoriska Museum Årstryck 2020: 33–48 33

Ted von Proschwitz

Faunistical news from the Göteborg Natural History Museum 2019 – snails, and mussels

In this paper the progress of scientific Proschwitz 2002, 2003, 2004, 2005, 2006, malacological work and projects in the Göte- 2007a, 2008, 2009, 2010a, 2011a, 2012, 2013, borg Natural History Museum during 2019 2014a, 2015, 2016, 2017, 2018a, 2019a). (cf. von Proschwitz 2019a), as well as some of the most interesting new records of land and Large freshwater mussels freshwater molluscs from Sweden 2019, and Also in 2019 large materials of freshwater in some cases from earlier years, are reported mussels have been sent in to the museum, and commented. among them should be mentioned: Rather many samples, predominantly from sev- Malacological projects 2019 eral provinces in Southern Sweden (leg: P. To get further information of the projects Ingvarsson, PI Fly-Vatten – och Fiskevård, and their development, see the corresponding Laholm); several large materials from locali- sections in the latest Faunistical news from ties in the counties of Kronoberg, Kalmar, the Göteborg Natural History Museum (von Jönköping and Östergötlands län (leg: C.-J. Månsson, C-J Natur, Torsås); Materials from monitoring of the freshwater pearl mussel, Vinjett: incarnatus. Photo: J. Roth, Margaritifera margaritifera (Linnaeus), from Linköping. the county of Norrbottens län (P. Olofsson); 34 Ted von Proschwitz

Further mussel materials from Dalarnas län The database based on this material was ear- (A. Johansson); Some minor materials, mainly lier referred to as Markfaunadatabasen, but as from Central and Eastern Sweden, have been it also comprises large materials of land and handed over to GNM by S. Lundberg, Vaaka freshwater molluscs which do not belong to Naturkonsult, Stockholm; smaller materials, the “strictly limited” Faunistic invertebrate from several parts of Sweden, have also been research programme (material collected by H. sent to GNM by other collectors (mainly in Lohmander and H. W. Waldén), it will hereaf- connection with determination by ter be referred to as the Göteborgs Naturhis- TvP). toriska Museums (GNM) databas för svenska land- och sötvattensmollusker [The Gothen- The work on the material from the burg Natural History Museum Database for ‘Faunistic invertebrate research Swedish Land and Freshwater Molluscs]. It is program’ and its use in new also in this version that all new additions are conservation and monitoring projects made. During 2019 Eva Andreasson has con- The Göteborg Natural History Museum is tinued to digitalize the mollusc material of A. supported financially through the Museum Nilsson (about 3/4 ready), Jenny Eikestam support programme (Museistödet) from the has continued to digitalize T. von Prosch- Swedish Species Information Centre (Art- witz’ material (about ½ the material ready – Databanken) since 2014, concerning digi- all samples before July 2004 digitalized), and talisation of the huge material from the Fau- T. von Proschwitz is digitalizing and, at the nistic invertebrate research program (Mark- same time doing a taxonomical check of sam- faunainventeringen) (cf. von Proschwitz & ples, of unregistered material of other collec- Andersson 1997, von Proschwitz 1997a). tors, already in the main collection. During

Collector Number of Number of Number of Status mollusc sites mollusc habitats species records of molluscs H. Lohmander 8 401 9 399 67 681 Complete (land-molluscs) H. W. Waldén 15 735 19 303 170 685 Complete (land-molluscs) A. Lörner 1 407 1 592 15 011 Complete I. Wäreborn 170 180 1 357 Complete T. von Proschwitz 2 971 3 848 32 555 In progress A. Nilsson 276 276 2 848 In progress Others 689 970 4 512 In progress Total number 29 649 35 568 294 649

Table 1. The land and freshwater mollusc material in the database of GNM at the end of 2019. (cf. von Pro- schwitz 2018a for the state one year earlier). A vast majority of the numbers are land living species. Note that one site may include more than one habitat. There are also rather large materials of centipedes in the base (1 334 localities; 11 626 species records). Including these the database comprises > 30 000 sites. Faunistical news from the Göteborg Natural History Museum 2019 – snails, slugs and mussels 35

2019 the progressing data laundering of the Man-spread molluscs material already in the data base was finished The project man-spread molluscs has con- (J. Eikestam & T. von Proschwitz). As a con- tinued as in previous years (cf. von Pro- sequence earlier number had to be changed in schwitz 1999a, 2000, 2001, 2002, 2003, 2004, some cases, due to limitations, double regis- 2005, 2006, 2007a, 2008, 2009, 2010a, 2011a, tering etc. – all this has now been corrected. 2012, 2013, 2014a, 2015, 2016, 2017, 2018a, Large land and freshwater mollusc mate- 2019a). rial in GNM, still not digitalized or ordered The number of -samples sent in by the into the main collection are those of J. Bern- public for determination was 33 (compare ström, B. Sundler, F. A. Svalander and G. the very low number 5(!) in the extremely Svensson. Also the materials of limnic mol- dry year 2018), and was in 2019 close to the lusca collected by H. Lohmander and H. W. average (30) in the years before (cf. von Pro- Waldén are intended to be entered into the schwitz 2018a, 2019a). 2019 could be said to data bank. be an average year both concerning climate The material from the Faunistic inverte- (precipitation) and number of slugs (Arion brate research program is continuously used vulgaris). In most places no marked reduc- in several monitoring and reinvestigation tion in the number of slugs could be seen projects, in which the Göteborg Natural His- after the drought 2019 compared to the years tory Museum is co-operating, on consultant before, which indicates a rapid recover of the basis, with other institutions and authorities. populations – in accordance with the large Such projects, dealing with investigation of reproductive potential of this extremely inva- calcareous fens with previously known or sive species. May 2019 was fairly rich in rain, potential occurrences of rare and red-listed which undoubtedly facilitated for the species whorl-snails (Vertigo spp.) are in progress as it overwinters as juveniles – there is usually with the county of Gävleborgs län (provinces a strong correlation between the amount of Gästrikland and Hälsingland). The monitor- precipitation during May and the number of ing of the effects of the tunnel construction A. vulgaris in the following summer-autumn through the ridge of Hallandsås on the land (cf. discussion in von Proschwitz 2016, 2017, mollusc fauna has also been performed on a 2018a, 2019a). yearly schedule (von Proschwitz 2019b). As every year there has been a large interest from the media. TvP was interviewed 2 times Work on the scientific collection by TV, 6 by radio and about 20 by newspa- of land and freshwater molluscs pers/magazines.The information on slugs on in GNM the home pages of the museum (www.gnm. A large number of samples of land snails, se title: Snigelakuten) is also continuously mainly collected by the late Henrik W. updated. Waldén, have to be ordered into the main col- lection, and this work was started in 2019. International co-operation projects About 2/3 of this material, however, still are The polymorphism in colonizing populations to be checked and sorted in – the intention is of Cepaea nemoralis in Sweden is studied in to finish this work in 2020-2021. co-operation with Robert Cameron, Shef- 36 Ted von Proschwitz field University, UK. A study of the present all zoogeographic literature and species lists, situation on the Island of Gotland was pub- as well as in museum collections, the unit lished during the year (Cameron & von Pro- provinces (landskap) is used as it is unchange- schwitz 2019). able and stable. Co-operation with Gerhard & Margrit Falkner and Ira Richling, Naturkundemu- Vertigo geyeri Lindholm, 1925 seum Stuttgart is also in progress in several projects, especially should be mentioned 0,5 mm nomenclatorial and taxonomical problems concerning several groups, especially Euro- pean freshwater mussels, slugs of the genus , and man spread snails occurring in tropical hothouses in Sweden. During the year, the European Red list of land molluscs was published (Neubert et al. 2019). It is the result of the co-operation of a large group of European malacologists, over several years, organized by WWF and the Natural History Museum of Bern, Switzer- land.

Interesting species records Photo: J. Roth, Linköping. Apart from material collected in projects by This fastidious and rare species, occurring GNM (TvP) and reports and material from in calcareous fens, is placed in category NT co-operation partners also some records in (Near Threatened) in the Swedish red-list Artportalen (Swedish Species Information (ArtDatabanken 2015). During 2018-2019 Centre) are included and commented. As aimed search for the species was made in many of the reports there are from non-spe- the county of Gävleborgs län (provinces of cialists the data has to be treated cautiously Gästrikland and Hälsingland) on request of (before validated). This becomes evident as the province administration. Rather many some of the attached photos clearly show a localities are already known from the county different species than the reported. (cf. von Proschwitz 2007b), although geo- Systematics and nomenclature in the sec- graphically they are very restricted. The larg- tion below follows Falkner, Bank & von Pro- est parts of the province are oligothrophic, schwitz (2001), with some adjustment of the and hence unsuitable for V. geyeri, only in the order of families and sub-family taxa follow- area around Gävle in the south of Gästrikland ing Bouchet et al. (2017). In this part the geo- and in the parish Los in Hälsingland, suitable graphical designation province is used for the fen habitats are present. In the Los area 4 new old division of Sweden (= landskap), whereas occurrences were found, and in the calcare- the designation county is used for the modern ous area south of Gävle more than 10 new division into administrative units (= län). In occurrences (all leg. TvP). Faunistical news from the Göteborg Natural History Museum 2019 – snails, slugs and mussels 37

Discus rotundatus (O. F. Müller, 1774) taly by self-fertilization, facilitates the possi- The distribution pattern of D. rotundatus in ble foundation of new populations, but under Sweden is pronounced southwestern-sub- present climatological conditions it could not oceanic. In eastern Sweden the continuous establish out-door populations in Sweden. northern distribution limit runs though the central parts of the province of Östergöt- Boettgerilla pallens Simroth, 1912 land and the southern part of the province of A few new records for this man-spread species Närke (cf. distribution map in Waldén 2007). were reported 2019 in Artportalen (Swedish There are some anthropochorous occur- Species Information Centre): parish Magle- rences in the Lake Mälaren-Stockholm area hem, at Magelhem and Fästan (leg: U. Gärden- (cf. Waldén 1955), but in the northern part fors) October 2019 (In Artportalen, Swedish of the province of Uppland-southern part of species information centre) and in the garden the province of Gästrikland only three pre- market centre Karl Fredrik på Eklaholm, vious localities are known. In 2019 a further Eklaholm, Hejdesten, parish Löderup, Skåne locality was found at the church of Skutskär (leg: J. Roth), September 2019 – all sites situ- in the north-westernmost corner of Uppland ated in the province of Skåne (S. Sweden). It (leg: TvP). was also found in Bemnäs, parish Lycke in the province of Bohuslän (W. Sweden) (leg: Rumina decollata (Linnaeus, 1758) C. Hansson), June 2019 (In Artportalen, An empty juvenile shell was found in the Swedish species information centre), and in garden market centre Karl Fredrik på Ekla- Östansjö, parish of Viby, province of Närke holm, Eklaholm, Hejdesten, parish Löderup, (C. Sweden) – new province record (leg: J. Skåne (leg: J. Roth September 2019). The spe- Roth), November. For details on the spread cies is new for the province of Skåne, even if it and previous records in Sweden see Svens- was found in a market garden at Hemmeslöv son, Rudd & von Proschwitz (1996) and von (county of Skåne but province of Halland, Proschwitz (1994a, 1999a, 2000, 2005, 2008, SW Sweden) in 2018 (cf. von Proschwitz 2009, 2010a, 2011a, 2012, 2013, 2014, 2015, 2019a). At Eklaholm also M. vermiculata, T. 2016, 2017, 2018a, 2019a). B. pallens is now pisana and C. acuta were found, cf. these spe- known to occur in the provinces of Skåne, cies below). R. decollata has earlier been intro- Halland, Småland, Östergötland, Västergöt- duced with vegetables and plants to different land, Bohuslän, Dalsland, Närke, Söder- parts of Sweden, and has also established manland, Uppland, Värmland and Hälsing- a population in the greenhouse of a market land. Due to its subterraneous way of living garden – see references to earlier records in B. pallens is most probably under-recorded. von Proschwitz (1993a, 2007a, 2019a) and further references cited in that paper. Records invadens Reise, Hutchinson, are known from the provinces of Skåne, Hal- Schunack & Schlitt, 2011 [=Deroceras land and Dalsland – in the latter province as panormitanum auctt. non (Lessona & Pollonera, 1882)]. an establish greenhouse population. Further introductions in the coming years are to be A new record for the species 2019 was made expected. Its ability to reproduce uniparen- in the garden market centre Karl Fredrik 38 Ted von Proschwitz

Ambigolimax valentianus (A. Férussac, på Eklaholm, Eklaholm, Hejdesten, parish 1822) Löderup, Skåne (leg: J. Roth), September 2019 – D. invadens was common over the A new record for the species was made in the whole area. There is a rapid, ongoing spread garden market centre Karl Fredrik på Ekla- and establishment of out-door populations holm, Eklaholm, Hejdesten, parish Löderup, of this anthropochorous species in South Skåne (leg: J. Roth), September 2019 – it was and Middle Sweden (cf. von Proschwitz common out-doors (>10 specimens found). 2017, 2018a, 2019a). Information of previous A. valetianus has been known as a green-house records and the dispersal is found in von Pro- species since the 1920’s (Waldén 1960). How- schwitz (1999a, 2000, 2001, 2002, 2003, 2004, ever, in the latter two decades there have been 2005, 2006, 2007a, 2008, 2009, 2011a, 2012, more and more outdoor records. Pro­bably 2013, 2014, 2015, 2016, 2017, 2018a, 2019a). the increasingly warmer climate has made Outdoor occurrences are now known from it possible for the species to survive winter the provinces of Skåne, Halland, Småland, outdoors, at least in the southernmost part Öland, Gotland, Västergötland, Östergöt- – most of the outdoor records are from the land, Bohuslän, Dalsland, Uppland, Värm- province of Skåne. (cf. von Proschwitz 2017, land and Hälsingland. see also D. invadens above). A. valentianus species has been found in the following prov- Deroceras sturanyi (Simroth, 1894) inces (underlined = also out-door records; This, somewhat enigmatic, and rare slug not underlined = only greenhouses): Skåne, is with high probability an old anthropo- Blekinge, Halland, Småland, Östergötland, chorous species in Sweden (cf. discussions in Västergötland, Bohuslän, Dalsland, Upp- Lohmander 1959, Proschwitz 2005, 2019a). land, Västmanland, Dalarna, Gästrikland, In the Swedish area, in which it was earlier Hälsingland, Västerbotten, Norrbotten, Pite (1960’s and 70’s) rather common it seems to lappmark. Further information on records of have disappeared from many of its former A. valentianus in Sweden is given by Waldén localities (cf. von Proschwitz 2019a), whereas (1960) and von Proschwitz (2001, 2007a, an ongoing spread by the help of man seems 2009, 2010a, 2011a, 2013, 2017). to be ongoing in other areas. In 2019 it was found in the garden market centre Karl Fre- Arion rufus (Linnaeus, 1758) drik på Eklaholm, Eklaholm, Hejdesten, Specimens of A. rufus were sent in from parish Löderup, Skåne (leg: J. Roth), Sep- the village Broby, parish Ö. Broby, in the tember 2019 (sympatrically with D. invadens, north-eastern part of the province of Skåne cf. above). So far D. sturanyi has been found (S. Sweden). The species has some further (scattered, often isolated records) in the fol- occurrences in this part of the province. The lowing provinces: Skåne, Blekinge, Småland, origin of these are not known, but it may be Gotland, Östergötland, Västergötland, Upp- an intentional introduction (such are known land, Västmanland and Hälsingland. from e.g. Öland and the Karlskoga-area in Värmland (von Proschwitz 1996c, von Pro- schwitz & Andersen 2010), or unintentional anthropochorous spread with plants or earth. Faunistical news from the Göteborg Natural History Museum 2019 – snails, slugs and mussels 39

Further information on the occurrence and out specimens as proof, the occurrence of C. spread of A. rufus in Sweden can be found in nemoralis in the areas of Karlstad in question von Proschwitz (1993b, 1996a, 1996b, 1996c, cannot be accepted. 1997a, 1997b, 2001a, 2002, 2003, 2004, 2005, 2006, 2007a, 2008, 2009a, 2010a, 2011a, 2014, Cepaea nemoralis (Linnaeus, 1758) 2015, 2017, 2018a) and von Proschwitz & 1 cm Andersen (2010).

Cepaea hortensis (O. F. Müller, 1774) 1 cm

Photo: J. Roth, Linköping. A population of the species was found in the residential area Hälsingeberg in the city of Falun, province of Dalarna (C. Sweden) (leg: S. Möller-Skog, November 2019) (60˚35,626̍ Photo: J. Roth, Linköping. N, 15˚42,471̍ E). This is the so far northern- In Artportalen (the website which is open for most known record on the inland of Sweden. the public to report species’ records) of the In the coastal area of the Baltic Sea it has, Swedish Species Information Centre (ArtDa- however, been found far more to the north, tabanken), there are several reports of Cepaea in the city Härnösand in the coastal area of nemoralis (Linnaeus, 1758) in the years 2008- the province of Ångermanland (62˚38,087' 2009 from the area Gräsdalen-Gruvlyckan in N, 17˚57,323' E) (von Proschwitz 2014). C. the city of Karlstad, S part of Värmland (C. nemoralis has been spreading rapidly in South Sweden). In 2019 TvP visited the area, but and Middle Sweden in the last decades, and could find nothing but colour- and band- the species is now also spreading northwards. ing morphs of the closely related C. horten- C. nemoralis is highly polymorphic concern- sis. Among others yellow morphs with only ing colour- and banding-morphs. See also the the third band (00300), dark to light brown above mentioned co-operation project with morphs such as f. castanea and f. baudonia R. Cameron on polymorphism in colonizing were rather common. The latter morph with populations. So far two studies of the popula- a light brown-reddish lip (var. roseolabiata). tion morf-composition in Sweden have been There is a strong suspicion, that such speci- published: The Göteborg area (W. Sweden) mens have been mistaken for C. nemoralis, (Cameron et al. 2014) and the Island of in which these colour and band morphs are Gotland (Baltic Sea) (Cameron & von Pro- common, and which almost always has a schwitz 2019). Further data is collected every brown lip – usually, however, darker than the year, and further publications are in prepara- lip in var. roseolabiata in C. hortensis. With- tion. 40 Ted von Proschwitz

Massylaea vermiculata (O. F. Müller, known established (introduced) populations 1774) [=Eobania vermiculata (O. F. are from Oldenburg in NW Germany (Wiese Müller, 1774)] 2016) and Belgium (Ronsmans & van den Neucker 2016).

1 cm Cornu aspersum (O. F. Müller, 1774)

1 cm

Photo: J. Roth, Linköping.

A living specimen of this species was found in the garden market centre Karl Fredrik på Ekla- Photo: J. Roth, Linköping. holm, Eklaholm, Hejdesten, parish Löderup, Skåne (leg: J. Roth), September 2019 [New Empty shells of the species were found in for the province of Skåne, earlier found in the a garden in Utby, Herrestad parish, near province of Halland, which belongs to the Uddevalla, province of Bohuslän (W Sweden) county of Skåne] (see also R. decollata above, (leg: J. Svedholm) August 2019 (In Artpor- and T. pisana and C. acuta below). Earlier talen, Swedish species information centre). Swedish records have been made in a garden Records of C. aspersum have been reported, in Askim, S of Göteborg [1998], a Turkish in an increasing number, during the latest groceries shop in Göteborg [2004] (both two decades. In many cases it has been intro- province of Västergötland,W. Sweden). It has duced with plants or vegetables. The species also been found in a flower shop in Uppsala has also been able to establish overwintering (imported with a citrus plant) [2016] (prov- populations in several places in the south- ince of Uppland, E. Sweden), and outside a ern and southwestern parts of Sweden. For plant nursery in Hemmeslöv, (province of further information on the Swedish popula- Halland, but county Skåne, SW. Sweden) tions and introductions see von Proschwitz (for further details on these records see von (1997c, 1999a, 2008, 2014, 2015, 2016, 2017, Proschwitz 1999a, 2005, 2016, 2018a). Fur- 2018a, 2019a) and von Proschwitz & Svens- ther import, with vegetables and plants, of son (1998). So far the species has been found this, originally Mediterranean species, is to in the provinces of Skåne, Halland, Småland, be expected in coming years, but despite that Västergötland, Bohuslän, Södermanland, the continuing warmer climate, establish- [Uppland] and [Norrbotten]. Provinces in ment of overwintering populations outdoors [ ] = only empty shells or found in green- in Sweden still seems improbable. Nearest groceries. Faunistical news from the Göteborg Natural History Museum 2019 – snails, slugs and mussels 41

Theba pisana (O. F. Müller, 1774) seem improbable, that C. acuta could estab- A juvenile empty shell of this species was lish populations in the coastal zone of the found in the garden market centre Karl Fre- southernmost part of Sweden (the province driks på Eklaholm, Eklaholm, Hejdesten, of Skåne), especially in the climatologically parish Löderup, Skåne (leg: J. Roth), Sep- favourable south-eastern part (Österlen), and tember 2019 (see also R. decolata and M. perhaps also on the Baltic islands of Öland vermiculata above, and C. acuta below). It and Gotland – if the climate develops in the has also been recorded in a shop in Alvesta in direction of even warmer. the province of Småland (E. Sweden) (leg: M. Sjödal), January 2019 (In Artportalen, Swed- Xeroplecta intersecta (Poiret, 1801) ish Species Information Centre). There are [=Candidula intersecta (Poiret, 1801)] earlier records of T. pisana from a vegetable Four records from 2019 are reported in Art- store in Kungsbacka, N part of the province portalen (Swedish species information centre) of Halland (W. Sweden) (von Proschwitz all from the province of Skåne (S. Sweden). 2007a) and a market garden in Hemmes- Two from the city of Malmö: Valenciagatan löv, (province of Halland, but county of (D. Nyström) and Oljehamnen (leg. M. Sjö- Skåne, SW Sweden) (von Proschwitz 2018a). dahl et al.) – both new areas for the species, Import of shells with garden seed has also one from Landskrona, Alicegården (leg. H. been reported (Nilsson 1967). As this, origi- Olausson), and one from Rinkaby fiering nally Mediterranean, species has established range, parish Åhus (E part of the province of populations in many places in the atlantic Skåne) (leg. M. Sjödahl) – earlier records are part of W. Europe, as far north as the Island known from both areas. The species inhabits of Helgoland, Germany (Wiese 2016), it may open grasslands and meadows and is spread- be a further possible colonizer in the coastal ing steadily in some parts of S. Sweden. So far areas of the province of Skåne. the species has been found in the provinces of Skåne, Gotland, Västergötland and Bohus- Cochlicella acuta (O. F. Müller, 1774) län (Fåhraeus 1982, von Proschwitz & Sven- A living specimen of C. acuta was found in the sson 1998, von Proschwitz 1999a, 2002, 2003, garden market centre Karl Fredriks på Ekla- 2006, 2016, 2017, 2018a, Sörensson 1999, and holm, Eklaholm, Hejdesten, parish Löderup, further information in Artportalen). Skåne (leg: J. Roth), September 2019 (se also R. decolata, and T. pisana above). The first Candidula unifasciata (Poiret, 1801) Swedish record of this species was made in The species was found in a gravel pit south of a market garden in Hemmeslöv in the south- Stånga, parish Stånga, Island of Gotland (leg. ernmost part of the province of Halland, D. Nyström et al.) September 2019. About (county of Skåne) in 2018 (von Proschwitz 30 sites are known from Gotland. The spe- 2019a). The species has expanded northwards cies is spreading rapidly in South and Middle in the antlantic part of W. Europe in the latter Sweden, partly by the commercial trade with decades and occurs in the coastal areas of Sedum-carpets – cf. the occurrence, often the British Isles, Belgium and the Nether- together with X. obvia, in the centre of lands (cf. Welter-Schultes 2012). It does not traffic roundabouts (von Proschwitz 2016, 42 Ted von Proschwitz

2017, 2018a). It also inhabits dry roadsides and calcareous, man influenced dry habi- tats. Records of the species are known from Skåne, Småland, Öland, Gotland, Östergöt- land, Västergötland, Bohuslän, Uppland and Jämtland (Fåhraeus 1982, Svensson & von Proschwitz 1997, von Proschwitz & Sven- sson 1998, von Proschwitz (1999a, 2000, 2002a, 2003, 2004, 2005, 2007a, 2008, 2009, 2015, 2016, 2017, 2018a, 2018b, 2019a, and further information in Artportalen).

Xerolenta obvia (Menke, 1828) 5 mm The species has been reported from a traffic Cernuella virgata. Photo: J. Roth, Linköping. roundabout at Tyresövägen/Siklöjevägen on Lidingö, Stockholm area, province of Upp- is the second record of the species in Sweden. land (leg. J. Roth) September 2019, and at the It has previously been found in a garden in runic stone field at Gammelbyn, Gävle, prov- Flottsund, Uppland (East Sweden) in 2011 ince of Gästrikland (Eastern Middle Sweden) (Neckheim 2011, von Proschwitz 2013). As (leg. M. Stenmark), April 2019– new prov- Theba pisana and Cochlicella acuta, C. virgata ince record. Like C. unifaciata, X. obvia is a is a Mediterranean species, which by the help “traffic roundabout species”, which is spread of man has spread northward along the coasts with Sedum-carpets. Several such records of France, Belgium, Holland, Germany (Kiel) (and established populations) have been and the British Isles (Kerney 1999, Walter- detected in South and Middle Sweden in later Schultes, 2012 – and distribution maps in years. Records of the species are known from these). It is perhaps a further possible colo- Skåne, Småland, Öland, Gotland, Östergöt- nizer in the coastlands of the province of land, Bohuslän, Södermanland, Uppland and Skåne. Jämtland (von Proschwitz 2001, 2002, 2004, 2006, 2007a, 2007c, 2008, 2014, 2017, 2018a, (O. F. Müller, 2018b, 2019a, and further information in 1774) Artportalen). In 2019 M. incarnatus was found WNW of Österskog in parish Glömminge on the Cernuella virgata (Da Costa, 1778) Island of Öland (leg. J. Roth), 13th Septem- An empty shell of the species was found in ber. Several living specimens were collected. the market garden in Hemmeslöv in the This is a remarkable and zoogeographically southernmost part of the province of Hal- very interesting record as it is the first from land, but belonging to the county of Skåne Öland. The land mollusc fauna of Öland is (leg: J. Roth), June 2018 (cf. H. minuscula, R. very well studied through the intense collec- decollata, C. acuta, C. barbara, T. pisana, and tion on the whole island in the years 1924, C. aspersum in von Proschwitz 2019a). This 1928, 1937, 1939 and 1941 by H. Lohmander Faunistical news from the Göteborg Natural History Museum 2019 – snails, slugs and mussels 43

– during this period 943 localities were sur- 2007, although that map is not complete). veyed. Malacological collecting on Öland has It is common in woodlands (predominantly been performed also in later years by T. von beech forests) in the provinces of Skåne and Proschwitz (von Proschwitz 1999b, 2007c). Blekinge in the south, with gaps only in the Totally there are land molluscs from more agriculturally dominated areas in the south than 1050 localities in the Göteborg Natural and in the spruce dominated zone in the History Museum database. Hence it becomes north. It is also absent in the easternmost part clear that M. incarnatus is an extremely rare of Blekinge. North of the ridge Hallandsås in and local species on Öland. H. Lohmander west and in the province border area between collected in the area around Österskog in the provinces of Blekinge and Småland in the 1941 but not at exactly the same spot as Roth east it becomes much rarer. There is a belt of 2019. The habitat is semi-open deciduous scattered occurrences northwards through woodland, which borders to a small road. the central and eastern parts, although not in Probably the occurrence is indigenous, even the coastal area, of the province of Småland if introduction by man may not be com- (Lohmander 1953, von Proschwitz 1998). pletely excluded. Lohmander (1955), clas- This belt reaches the area of Lake Sommen, sifies M. incarnatus as anthropophobic, but and the east coast of Lake Vättern at Stava, this is contradicted by occurrences in man- parish Ödeshög in the province of Östergöt- influenced sites in the province of Skåne, and land. also in Dalsland (von Proschwitz 1994b) and In W. Sweden there are a few scattered on rural ground, close to the churchyard, at occurrences in the province of Halland north Gårdby in the province of Småland (von Pro- of Hallandsås (Waldén 1969) and in the val- schwitz 2016). leys of rivers Viskan and Ätran in the south- The distribution picture of M. incarnatus ern part of the province of Västergötland (von in Sweden is complex (cf. the map in Waldén Proschwitz 2010b). In central Västergötland the species was recently found, quite isolated, at Dammbo, parish Dimbo (von Proschwitz 2011b, 2012). M. incarnatus is also rather common and spread in an isolated area in the woodlands of Tiveden on the border between the provinces of Västergötland and Närke (Lohmander 1945, 1955, von Proschwitz 2013). The Dammbo record is interesting, as it constitutes a bridge between the occurrences in the river valleys in southern Västergötland and the area in Tiveden. The westernmost outposts of the species in Sweden are three isolated localities in the province of Dalsland 5 mm (von Proschwitz 1994b). The occurrence at Monachoides incarnatus. Photo: J. Roth, Linköping. Österskog on Öland is a remarkable exten- See also vignette on p.33! sion of the Swedish distribution eastwards. 44 Ted von Proschwitz

The total distribution of M. incarnatus is cen- old land mollusc records have been given by tral-eastern European, it does not reach west Jonas Roth (Linköping). Furthermore he has of eastern France and not far east of Poland; supplied me with data of records from other in the south it reaches Bulgaria (cf. distribu- collectors and observers. J. Roth also allowed tion map in Welter-Schultes 2012). In the me to use several of his excellent photos of Baltic States the species is very rare – three snails in this article. Thanks to Peter Nielsen localities in Latvia (Rudzite et al. 2010) con- at the Göteborg Natural History Museum stitute its north-eastern border. It has so far for important work with many different parts not been found in Finland, the information of the project ”Man spread molluscs”. Karin on this in Falkner et al. (2001) is not correct. Ebbesdotter (Göteborg Natural History Museum) should be thanked for perform- Mytilopis leucophaeata (Conrad, 1831) ing the time consuming work with sorting [=Congeria cochleata (Nyst, 1835)]. out the snails from the sifting samples in the The first Swedish records of this invasive conservation and monitoring projects; Göran brackish water mussel were made in 2011, Andersson for remarks on the manuscript; close to the outlets of cooling water from the and Elisabeth Hagström for improving the nuclear power plant of Forsmark, province of language. Uppland (Florin et al. 2013). A further record from Sågarbofjärden, the Baltic cost of Upp- Sammanfattning land, west of Forsmark, was reported by von I rapporten presenteras vetenskapligt arbete Proschwitz (2017, 2018c). In Artportalen inom terrester och limnisk malakologi (Swedish Species Information Centre) two (snäckor, sniglar och musslor) och olika pro- new records of M. lecophaeata from localities jekt där GNM (genom T. von Proschwitz) är east of Forsmark are reported: Höggrundet, engagerat: Stormusslor i sötvatten, Arbetet parish Börstil and Enholmen, parish Gräsö med markfaunamaterialet och användning av (both leg. M. Strand), August 2018. Further det i olika inventerings-, naturskydds- och spread along the Baltic coast of Sweden is to miljöövervakningsprojekt, Kulturspridda be expected (cf. von Proschwitz 2018c). landmollusker (främst spansk skogssnigel), Arbetet med samlingen av land- och sötvat- Acknowledgements tensmollusker på GNM, Internationellt sam- I want to express my appreciation and thanks arbete om nomenklatur och taxonomi hos to all, who have contributed with material vissa grupper av land- och sötvattensmollus- and information to this article. Concern- ker, liksom den nya europeiska rödlistan för ing the freshwater mussels especially should landmollusker. be: Stefan Lundberg (Vaaka Naturkonsult, Dessutom presenteras intressanta fynd Bromma); Per Ingvarsson (PI Fly Vatten- och av land- och sötvattensmollusker, dels från Fiskevård, Laholm); Carl-Johan Månsson Naturhistoriska museets insamlingsverksam- (The Rural Economy and Agricultural Socie- het, dels från andra institutioner, forskare eller ties, E. Sweden); Niklas Wengström (Sport- privatpersoner som står i kontakt med GNM. fiskarna, Göteborg). Much important infor- Speciellt märks nya fynd och uppdateringar mation on localities and material of new and av status för följande arter (understrykningar Faunistical news from the Göteborg Natural History Museum 2019 – snails, slugs and mussels 45

= nya landskaps/länsfynd): Kalkkärrsgryn- T. & Ho r s á k , M. 2014. Cepaea nemoralis snäcka [V. geyeri] (Gästrikland, Hälsingland); (L.) in Göteborg, S.W. Sweden: variation in a Fläckdisksnäcka [D. rotundatus] (Uppland); recent urban invader. — Folia Malacologica 22 (3): 169-182. Bruten sylsnäcka [R. decollata] (Skåne); Ca m e r o n , R. A. D. & v o n Pr o s c h w i t z , T. 2019. Masksnigel [B. pallens] (Skåne, Bohuslän, Cepaea nemoralis (L.) on Gotland, Sweden: Närke); Vagabondsnigel [observera bytet av Spread and variation. — Folia Malacologica 27 svenskt namn från det tidigare växthussni- (4): 307-313. Få h r a e u s , G. 1982. På snäckjakt i gotländska gel (enligt beslut av ArtDatabanken) – arten marker. — Fauna och Flora 77 (6): 291-300. är i dag väletablerad på friland på flera håll i Fa l k n e r , G., Ba n k , R. & v o n Pr o s c h w i t z , T. södra Sverige] [D. invadens] (Skåne); Ham- 2001. Check-list of the non-marine Molluscan Species-group taxa of the States of Northern, marsnigel [D. sturanyi] (Skåne); Valentinsni- Atlantic and Central Europe (CLECOM Area gel [A. valentianus] (Skåne); Röd skogs- I) and their distribution. — Heldia 4(1/2): snigel [A. rufus] (Skåne); Trädgårdssnäcka 1-76. [C. hortensis] (Värmland); Parksnäcka [C. Fl o r i n , A. B., Mo, K., Sv e n s s o n , F., Sc h a g e r - s t r ö m , E., Ka u t s k y , L. & Be r g s t r ö m , L. 2013. nemoralis] (Värmland – korrektion av felak- First record of Conrad’s false mussel, Mytilop- tiga angivelser, Dalarna); Nudelsnäcka [M. sis leucophaeata (Conrad, 1831) in the southern vermiculata] (Skåne); Fläckig vinbergssnäcka Bothnian Sea, Sweden, near a nuclear power plant. — Bioinvasion Records 2 (4): 303-309. [C. aspersum] (Bohuslän); Dynsnäcka [T. Ke r n e y , M. P. 1999. Atlas of the Land and Fresh- pisana] (Skåne, Småland); Smal spetssnäcka water Molluscs of Britain and Ireland. — Col- [C. acuta] (Skåne); Större kalksnäcka [obser- chester (Harley Books). 264 pp. vera bytet av svenskt namn från det tidigare Lo h m a n d e r , H. 1953. Zoologiska avdelningen, berättelse för 1944. — Göteborgs Musei Års- kalksnäcka (enligt beslut av ArtDatabanken) tryck 1945: 11-20. [X. intersecta] (Skåne); Mindre kalksnäcka Lo h m a n d e r , H. 1953. Faunistiskt fältarbete 1951 [observera bytet av svenskt namn från det och 1952 (södra Småland östra och meller- sta delen). — Göteborgs Musei Årstryck 1953: tidigare hedsnäcka (enligt beslut av ArtDa- 31-83. tabanken) [C. unifasciata] (Gotland); Syd- Lo h m a n d e r , H. 1955. Faunistiskt fältarbete 1954 hedsnäcka [X. obvia] (Södermanland, Gäst- (utmed Götalands nordgräns, östra hälften). — Göteborgs Naturhistoriska Museum, Årstryck rikland); Medelhavshedsnäcka [C. virgata] 1955: 30-97. (Halland); Bokskogssnäcka [M. incarnatus] Lo h m a n d e r , H. 1959. Faunistiskt fältarbete i (Öland); Trekantig brackvattenmussla [M. västra och norra Jylland 1954-57. Landmollus- leucophaeata] (Uppland). kerna. — Göteborgs Naturhistoriska Museum, Årstryck 1959: 33-104. Ne c k h e i m , C. M. 2011. Landslakken verzamelen References tijdens een kort bezoek aan Zweden in 2011. Ar t Da t a b a n k e n 2015. Rödlistade arter i Sverige — De Kreukel 2011: 1-5. 2015. — ArtDatabanken SLU, Uppsala. 209 Ne u b e r t , E., Se dd o n , M.B., Al l e n , D.J., Ar r é - pp. b o l a , J., Ba c k e l ja u , T., Ba l a s h o v , I., Ba n k , Bo u c h e t , P., Ro c r o i , J.-P., Ha u s d o r f , B., Ka i m , R., Ca m e r o n , R., d e Fr i a s Ma r t i n s , A.M., De A., Ka n o , Y., Nü t z e l , A., Pa r k h a e v , P., Ma t t i a , W., De d o v , I., Du da , M., Fa l k n e r , Sc h r ö d l , M. & St r o n g , E. E. 2017. Revised G., Fa l k n e r , M., Fe h é r , Z., Ga r g o m i n y , O., Classification, Nomenclator and Typification Ge o r g i e v , D., Gi u s t i , F., Gó m e z Mo l i n e r , of Gastropod and Monoplacophoran Families. B.J., Gr o h , K., Ib á ñ e z , M., Kapp e s , H., Ma n - — Malacologia 6(1/2): 1-526. g a n e l l i , G., Ma r t í n e z -Or t í , A., Na r d i , G., Ca m e r o n , R. A. D., Co x , R. J., v o n Pr o s c h w i t z , Ne i b e r , M.T., Pá l l -Ge r g e l y , B., Pa r m a k e l i s , 46 Ted von Proschwitz

A., Pr i é , V., Re i s c h ü t z , A., Re i s c h ü t z , P.L., v o n Pr o s c h w i t z , T. 1997b. Arion lusitanicus Ro w s o n , B., Rü e t s c h i , J., Sl ap n i k , R., So n , Ma b i l l e and A. rufus (L.) in Sweden: A com- M., Št a m o l , V., Te i x e i r a , D., Tr i a n t i s , K., parison of occurrence, spread and naturali- Va r d i n o ya n n i s , K., v o n Pr o s c h w i t z , T. and zation of two alien slug species. — Heldia 4 Wa l t h e r , F. 2019. European Red List of Ter- Sonderheft 5: 137-138. restrial Molluscs. — IUCN: Cambridge, UK v o n Pr o s c h w i t z , T. 1997c. Fläckig vinbergssnäcka and Brussels, Belgium. 23 pp. https://portals. [Cornu aspersum (O. F. Mü l l e r )] funnen i iucn.org/library/node/48439 Sverige. — Göteborgs Naturhistoriska Museum, Ni l s s o n , A. 1967. Mollusc shells as impurity in Årstryck 1997: 23-27. garden-seeds. — Agri Hortique Genetica 25: v o n Pr o s c h w i t z , T. 1998. Landlevande mollusker 145-160. i Kronobergs län – Förslag till utnyttjande av v o n Pr o s c h w i t z , T. 1993a. On the spread and data från Göteborgs Naturhistoriska Museums development of the anthropochorous ele- markfaunainventering i miljöövervakningsstu- ment in the land-snail fauna of the province dier. – Sammanställningar av lokaler för rödlis- of Dalsland (SW Sweden). — Mitteilungen tade och sällsynta arter, samt lokaler med höga der deutschen malakozoologischen Gesellschaft naturvärden. — Länsstyrelsen i Kronobergs län 50/51: 15-31. - Natur- och kulturmiljöenheten. Meddelande v o n Pr o s c h w i t z , T. 1993b. Den röda skogssnigeln 1998: 22. 113 pp. [With English summary: – Arion rufus (L.) – i Sverige - historia, sprid- Land in the province of Kronobergs ning, utbredning och biologi. — Göteborgs län (SE Sweden) - Use of data from the ’Fau- Naturhistoriska Museum, Årstryck 1993: 31-41. nistic research program’ of the Göteborg Nat- v o n Pr o s c h w i t z , T. 1994a. Masksnigeln – Boett- ural History Museum in environmental moni- gerilla pallens Si m r o t h - en egendomlig kau- toring projects and identification of localities kasisk snigelart i spridning med människan i with high values in natural conservation.] Sverige. — Göteborgs Naturhistoriska Museum, v o n Pr o s c h w i t z , T. 1999a. Faunistiskt nytt 1998 Årstryck 1994: 45-54. – Snäckor, sniglar och musslor. — Göteborgs v o n Pr o s c h w i t z , T. 1994b. Zoogeographical Naturhistoriska Museum, Årstryck 1999: 27-44. Studies on the Land Mollusca of the Province v o n Pr o s c h w i t z , T. 1999b. Landlevande mol- of Dalsland (SW. Sweden). — Acta Regiae Soci- lusker i Kalmar län II. Öland – Förslag till etatis Scientiarum et Litterarum Gothoburgensis utnyttjande av data från Göteborgs Naturhis- Zoologica 15. 152 pp. toriska Museums markfaunainventering i mil- v o n Pr o s c h w i t z , T. 1996a. Röda skogssniglar. — jöövervakningsstudier. – Sammanställningar [p. 96]. In: Gu s t a f s s o n , L. & Ah l é n , I. (eds): av lokaler för rödlistade och sällsynta arter, Sveriges Nationalatlas: Växter och djur. Föränd- samt lokaler med höga naturvärden. — Läns- ringar i floran och faunan. styrelsen Kalmar län informerar. Meddelande v o n Pr o s c h w i t z , T. 1996b. Utbredning och sprid- 1999:13. 143 pp. [With English summary: Land ning av spansk skogssnigel [Arion lusitanicus mollusca in the province of Kalmar län (SE Ma b i l l e ] och röd skogssnigel [Arion rufus Sweden). Part 2. The island of Öland. - Use (L.)] – en översikt av utvecklingen i Sverige. of data from the ’Faunistic research program’ — Göteborgs Naturhistoriska Museum, Års- of the Göteborg Natural History Museum in tryck 1996: 27-45. [With English summary: environmental monitoring projects and identi- Arion lusitanicus Ma b i l l e and A. rufus (L.) in fication of localities with high values in nature Sweden, a review of the occurrence, spread and conservation.] naturalization.] v o n Pr o s c h w i t z , T. 2000. Faunistiskt nytt 1999 v o n Pr o s c h w i t z , T. 1996c. De stora röda sniglarna – Snäckor, sniglar och musslor. — Göteborgs på Öland. — Öländsk bygd Åkerbo Härad. Naturhistoriska Museum, Årstryck 2000: 21-40. Åkerbo Hembygdskrets Årsbok 1996: 56-61. v o n Pr o s c h w i t z , T. 2001. Faunistiskt nytt 2000 v o n Pr o s c h w i t z , T. 1997a. Der Aufbau einer – Snäckor, sniglar och musslor inklusive nationalen Computer-Datenbank für land- något om afrikansk konsnäcka Afropunctum lebende Mollusken im Naturhistorischen seminium (Mo r e l e t ) och större vallsnäcka Museum Göteborg, Schweden. — Mitteilun- Monacha cantiana (Mo n t a g u ) – två för Sve- gen der deutschen malakozoologischen Gesell- rige nya, människospridda landsnäckor. — schaft 60: 27-33. Göteborgs Naturhistoriska Museum, Årstryck Faunistical news from the Göteborg Natural History Museum 2019 – snails, slugs and mussels 47

2001: 19-36. Cepaea nemoralis (Li n n a e u s ) und Helix poma- v o n Pr o s c h w i t z , T. 2002. Faunistiskt nytt 2001 tia Li n n a e u s . — Heldia 5 (3): 75-79. [With – Snäckor, sniglar och musslor. — Göteborgs English summary, p. 78]. Naturhistoriska Museum, Årstryck 2002: 29-46. v o n Pr o s c h w i t z , T. 2008. Faunistical news from v o n Pr o s c h w i t z , T. 2003. Faunistiskt nytt 2002 the Göteborg Natural History Museum 2007 – Snäckor, sniglar och musslor. — Göteborgs – snails, slugs and mussels – with some notes Naturhistoriska Museum, Årstryck 2003: 25-42. on Vertigo ultimathule v o n Pr o s c h w i t z – a v o n Pr o s c h w i t z , T. 2004. Faunistiskt nytt 2003 landsnail species from northernmost Sweden – Snäckor, sniglar och musslor. — Göteborgs new to science. — Göteborgs Naturhistoriska Naturhistoriska Museum, Årstryck 2004: 23-36. Museum, Årstryck 2008: 51-72. v o n Pr o s c h w i t z , T. 2005. Faunistiskt nytt 2004 v o n Pr o s c h w i t z , T. 2009. Faunistical news from – Snäckor, sniglar och musslor inklusive något the Natural History Museum, Göteborg om kinesisk skivsnäcka Gyraulus chinensis 2008 – snails, slugs and mussels – with some (Du n k e r ) och amerikansk tropiksylsnäcka notes on the slug Limacus flavus (Li n n a e u s ) Subulina octona (Br u g u i è r e ) – två för Sverige – refound in Sweden, and Balea heydeni v o n nya, människospridda snäckarter. — Göteborgs Ma l t z a n – a species new to Sweden. Naturhistoriska Museum, Årstryck 2005: 35-61. — Göteborgs Naturhistoriska Museum, Årstryck v o n Pr o s c h w i t z , T. 2006. Faunistiskt nytt 2005 – 2009: 47-68. Snäckor, sniglar och musslor – samt något om v o n Pr o s c h w i t z , T. 2010a. Faunistical news from östlig snytesnäcka Bithynia transsilvanica (E. the Natural History Museum, Göteborg 2009 A. Bi e l z ) – återfunnen i Sverige och kinesisk – snails, slugs and mussels – with some notes dammussla Sinanodonta woodiana (Le a ) – en on Pupilla pratensis (Cl e s s i n ) a land snail spe- för Sverige ny sötvattensmussla. — Göteborgs cies new to Sweden. — Göteborgs Naturhistor- Naturhistoriska Museum, Årstryck 2006: 39-70. iska Museum, Årstryck 2010: 41-62. v o n Pr o s c h w i t z , T. 2007a. Faunistiskt nytt 2006 – v o n Pr o s c h w i t z , T. 2010b. Landlevande mol- Snäckor, sniglar och musslor – samt något om lusker i naturreservaten Rya Åsar och Älmås, ribbpunktsnäcka Paralaoma servilis (Sh u t t l e - (Borås stad, Västra Götalands län 2010). — w o r t h ) – en för Sverige ny, människospridd Meddelanden från Göteborgs Naturhistoriska landsnäcka. — Göteborgs Naturhistoriska Museum 26: 1-40. Museum, Årstryck 2007: 47- 69. v o n Pr o s c h w i t z , T. 2011a. Faunistical news from v o n Pr o s c h w i t z , T. 2007b. Snäckor i rikkärr i the Göteborg Natural History Museum 2010 Gävleborgs län. – Större agatsnäcka – Coch- – snails, slugs and mussels – with some notes licopa nitens (M. von Gallenstein), Kalkkärrs- on Gyraulus stroemi (We s t e r l u n d ) – a fresh- grynsnäcka – Vertigo geyeri Lindholm, Smal- water snail species new to Sweden. — Göte- grynsnäcka – Vertigo angustior Jeffreys och borgs Naturhistoriska Museum, Årstryck 2011: Tandsnäcka – Perforatella bidentata (Gmelin) 39-60. i Gävleborgs län: Återinventering av äldre v o n P r o s c h w i t z , T. 2011b. Inventering av sällsynta lokaler, undersökning av nya objekt samt grynsnäckor i Västra Götalands län 2007-2009. skötselrekommendationer för aktuella loka- — Länsstyrelsen i Västra Götalands län, Rap- ler 2006. — Länsstyrelsen Gävleborg. Rapport port 2011: 61. 235 pp. [With English summary, 2007:14. 43 pp. [English summary, pp. 37-39: pp. 188-190: Rare whorl-snails – Vertigo spp. Cochlicopa nitens (M. von Gallenstein), Vertigo in the province of Västra Götalands län (SW. geyeri Lindholm, Vertigo angustior Jeffreys and Sweden) 2007-2009.] Perforatella bidentata (Gmelin) in the province v o n Pr o s c h w i t z , T. 2012. Faunistical news from of Gävleborgs län (E. Sweden) 2006 – Rein- the Göteborg Natural History Museum 2011 vestigation of old sites and surveys of newe – snails, slugs and mussels. — Göteborgs potential localities, including comments on Naturhistoriska Museum Årstryck 2012: 41-60. threats and recommendations of conservation v o n Pr o s c h w i t z , T. 2013. Faunistical news from measures. the Göteborg Natural History Museum 2012 v o n Pr o s c h w i t z , T. 2007c. Fünf für die Ostsein- – snails, slugs and mussels. — Göteborgs sel Öland neue Landschneckenarten: Ver- Naturhistoriska Museum, Årstryck 2013: 45-54. tigo liljeborgi (We s t e r l u n d ), Xerolenta obvia v o n Pr o s c h w i t z , T. 2014. Faunistical news from (Me n k e ), Candidula unifasciata (Po i r e t ), the Göteborg Natural History Museum 2013 48 Ted von Proschwitz

– snails, slugs and mussels – with som notes on tidsmätningar projekt Hallandsås 2018. Tra- Sphaerium nucleus (S. St u d e r ) – a freshwater fikverket. mussel species new to Sweden. — Göteborgs v o n Pr o s c h w i t z , T. & An d e r s e n , A. 2010. Rød- Naturhistoriska Museum, Årstryck 2014: 39-52. skogsnegl Arion rufus i Sverige og Norge. — v o n Pr o s c h w i t z , T. 2015. Faunistical news from Fauna (Oslo) 63 (1): 2-7. the Göteborg Natural History Museum 2014 v o n Pr o s c h w i t z , T. & An d e r s s o n , G. 1997. Data- – snails, slugs and mussels. — Göteborgs bas för landlevande mollusker, tusenfotingar Naturhistoriska Museum, Årstryck 2015: 31-46. och gråsuggor på Göteborgs Naturhisto- v o n Pr o s c h w i t z , T. 2016. Faunistical news from riska Museum. — Göteborgs Naturhistoriska the Göteborg Natural History Museum 2015 Museum, Årstryck 1997: 29-36. – snails, slugs and mussels – with some notes v o n Pr o s c h w i t z , T. & Sv e n s s o n , U. 1998. Fau- on Leptinaria unilamellata (d’Orbigny) – a nistiskt nytt från Göteborgs Naturhistoriska land snail species new to Sweden. — Göteborgs Museum 1997. — Naturhistoriska Museum, Naturhistoriska Museum, Årstryck 2016: 33-51. Årstryck 1998: 15-20. v o n Pr o s c h w i t z , T. 2017. Faunistical news from Ro n s m a n s , J. & v a n d e n Ne u c k e r , T. (2016): A the Göteborg Natural History Museum 2016 persistent population of the chocolate-band – snails, slugs and mussels – with some notes snail Eobania vermiculata (: Heli- on Rangia cuneata (G. B. Sowerby I) and Myti- cidae) in Belgium. — Belgian Journal of Zool- lopsis leucophaeata (Conrad) – two invasive ogy 146 (1): 66-68. brackish water mussel species new to Sweden. Ru d z i t e , M., Dr e i j e r s , E., Oz o l i n a -Mo l l , L., — Göteborgs Naturhistoriska Museum, Årstryck Pa r e l e , E., Pi l a t e , D., Ru d z i t i s , M. & St a - 2017: 33-52. l a z s , A. 2010. Latvijas gliemji: Sugu noteicejs. v o n Pr o s c h w i t z , T. 2018a. Faunistical news from A guide to the Molluscs of Latvia. — 252 pp. the Göteborg Natural History Museum 2017 Riga (LU Akademiskais apgads). – snails, slugs and mussels – with some notes Sö r e n s s o n , M. 1999. Kalksnäckan Candidula on two imported slugs: Ariolimax columbianus intersecta (Po i r .) – en expanderande stor­ (Gould) and Prophysaon foliolatum (Gould) snäcka på hedmarker. — FaZett 12(1): 13-16. and an introduced slug species Arion subfuscus Sv e n s s o n , U., Ru dd , P. & v o n Pr o s c h w i t z , T. (Draparnaud) s.s. new to Sweden. — Göte- 1996. Faunistiskt nytt från Göteborgs Natur- borgs Naturhistoriska Museum, Årstryck 2018: historiska Museum 1995. — Göteborgs Natur- 29-45. historiska Museum, Årstryck 1996: 17-20. v o n Pr o s c h w i t z , T. 2018b. Malakologische Neu- Wa l d é n , H. W. 1955. The land Gastropoda of the igkeiten in der Land- und Süsswasserfauna vicinity of Stockholm. — Arkiv för Zoologi 2 Schwedens. — In: Wi n k . J. (ed.): Zusammen- (7) Nr 21: 391-448 + Taf. I. fassung der Tagungsbeiträge anlässig der 56. Wa l d é n , H. W. 1960. Om ett par för Sverige nya, Frühjahrstagung der DMG vom 2. Bis 5. Juni anthropochora Landmollusker, Limax valen- 2017 in Warberg. — Mitteilungen der deutschen tianus Férrusac och Deroceras caruanae (Pol- malakozoologischen Gesellschaft 98: 32-33. lonera), jämte några andra, kulturbundna arter. v o n Pr o s c h w i t z , T. 2018c. Two invasive brack- — Göteborgs kungliga Vetenskaps och Vitterhets- ish water mussel species in Sweden: Rangia Samhälles Handlingar (6), Serie B 8 (8): 1-48. cuneata (G. B. Sowerby I) and Mytilopsis leu- Wa l d é n , H. W. 1969. En översikt över Hallands cophaeata (Conrad). — Journal of Conchology landmolluskfauna. — Fauna och flora 64 (2): 43 (1): 111-113. 49-96. v o n Pr o s c h w i t z , T. 2019a. Faunistical news from Wa l d é n , H. W. 2007. Svensk landmolluskatlas. — the Göteborg Natural History Museum 2018 – 271 pp. Stenungsund (Naturcentrum AB). snails, slugs and mussels – with some remarks We l t e r -Sc h u l t e s , F. W. 2012. European non-ma- on Cochlicella acuta (O. F. Müller) and Coch- rine molluscs, a guide for species identifica- licella barbara (Linnaeus) two imported land- tion. — pp. A1-A3, 1-679, Q1-Q78. Göttin- snail species new to Sweden. — Göteborgs gen (Planet Poster Editions). Naturhistoriska Museum, Årstryck 2019: 33-47. Wi e s e , V. 2016. Die Landschnecken Deutschlands. v o n Pr o s c h w i t z , T. 2019b. Inventering av mol- Finden – Erkennen – Bestimmen. 2. Auflage. lusker Hallandsås 2018. — 18 pp. – In: Prövo- — 352 pp. Wiebelsheim (Quelle & Meyer).