Вестник Серия Экологическая Bulletin Ecology Series

ISSN 1563-034Х • ИНДЕКС 75880; 25880

ӘЛ-ФАРАБИ атындағы КАЗАХСКИЙ НАЦИОНАЛЬНЫЙ AL-FARABI KAZAKH ҚАЗАҚ ҰЛТТЫҚ УНИВЕРСИТЕТІ УНИВЕРСИТЕТ имени АЛЬ-ФАРАБИ NATIONAL UNIVERSITY

ХАБАРШЫ ЭКОЛОГИЯ СЕРИЯСЫ ВЕСТНИК СЕРИЯ ЭКОЛОГИЧЕСКАЯ BULLETIN ECOLOGY SERIES

1(33) 2012 ISSN 1563-034 75880 25880

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QR 3 1 (33) 2012 H 3 . W SY -.

22.04.1992 . TJL 766. , J I. R. H.U. R - ! - ( 80- ! !) ………… 8 25.11.99. 956-L …………..……………………………….... 12 ______ .. : U ! ! ^ X.R., ..., . ! ………………….. 12 ( ) R.. U ! - .: 377-33-80, 377-33-34 + 1200, R S.V., ..., . ! ! (. ) ……………………………………………………….. 16 .: 292-70-26 + 2128 Kazuyuki Inubushi K I.., ..., . Revolution of life from ocean to land, ( ) interacting with global environment ……………………………… 20 .: 377-33-34+12-15 F W.W., ..., ., F M.I., ..., ., …………..…………………………………... 21 G F.G., ..., ., I .. L . K- K F.G., ..., ., F.R., ..., ., ! ! – ! R X.J., ..., ………………………………………………………… 21 R .G., ..., ., S R.., ..., ., S W.X., ..., ., W 1. - W H.I., ..., ., X L.V., ..., ., …………….…………………………..….… 28 1.1 N………………………………………………………… 28 I.R. = Ŀ ı ! ..... 28 3 1 (33) 2012 . .., Q.. G 3 5664 Y ! U 20.12.2011. ! ! - 90110 1/8. G ! 3 1. ………………………..…………………………………... 30 U !. T 20 .. .R. W I X 500 . M 3 240 \ !. (Barbatula Toni) G ….. 32 « » .., .. = Ŀ -. (Abramis Brama) ı …………………. 35 050040, .F, .-, I.., R .., R .. 71, SY. T ! U! I « ». Triplophysa Strauchii .Y F ...... 37 2 Y . . 1 (33). 2012

ĭ .., U R.L. Ľ = Ŀ Ŀ ı Ľ ı ...... 42 .I., T .., .. G Ŀ Ŀ (Stizostedion Lucioperca) ı ! Ŀ ...... 45 J . ., . . ! - ! F ( F L) ……………………………………………………………….. 48 JĽ' .ļ., .. F ı ...... 50 JĽ' .ļ., .. W! Ŀ ı ...... 52 J I. R. T ! RWTU ! G-F ……………… 54 .. V W Z U! ...... 59 L J.., .L. L ı ! Ľ ...... 63 L .R., .., .. H! V Ergasilus Sieboldi G …………………… 66 M Q.., Z.. G ! G (Huso Huso L.) ! ! ………………………………………………………………………………………. 70 T.R., I.., .. T- ……………………………………………………………………………………... 73 T.R., I.., J Q.. (Cyprinus Carpio) - ! …………………….. 76 I.. V F ! FT «G» ...... 77 .. T Y- ! W Alosa Kessleri Kessleri (Grimm, 1887)………………………………………………………………………………… 79 T.. R Y (Alburnus Alburnus) " ! G 81

Z.., R .. W . ^ ……………………………………………………………………………… 85 Z.., R .., J ., J .. ! . L …………………………………………………………………………… 88 . . U _ Esox Lucius L., 1758 . . W ………………………………………………………………………... 91 Q.. c ! ! - ……………………………………………………………………………... 94 Q Q.U., J .. T! W ………………………………………………………………………… 96 R . T !! FWUcT, ! ………………………. 99 R .., .U. R!! ! F (Artemia Sp. ) W ...... 102 R .. T !! V T - ! ! ………………………………………………………………. 103 R .., .. !! T V H SY. . 1 (33). 2012 3

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1.2 I...... 156 .R., .. M ! …………………….. 156 .., I .., R.. I W ...... 159 J .. T G H 2005-2009 ……………………………………………………………………………….. 162 .. M Y- H ……………………… 165 I.., M.. H G , F …………. 168 .. S ! W …………………………………………………………………...... 172 M . M., R.. !! W- (V ) …….. 175 " Q . W! F W T 2009-2011 ……. 179 .I. V 4 Y . . 1 (33). 2012

M- …………………………………………………. 182 .Z., R.. H! W- H W ! ………………………………………………………… 185 Q T.., U J.., R .., I.., R .. T V ………………………………………………………………………………………. 188 R L.T. W - 28 F 2011 . ………………………………………………………… 192 R .R. S W M W ! ………………. 194 .. M –R W ………………………... 197 .. R ……………………………………………………………………………………... 201 .., T.., J.., M.. W R! …………………………….. 204 .., R .., R .. W! W ! (Y «S») 2009 . ……………………. 208 Q.. V W ! ! ………………………………………………………………………………... 212 Q.. c ! ! ! ! F! …………………………………….. 216

W 2. - ………………. 220

J Q.., .R., .., U.., .I., M., M.L., R J.. W ! - W! ……………………………………………… 220 Z.., M.., .I., .. F!! ! (mn, co, ni) ! ! . ………………………… 223 .., M.., I.., Z.. ! ! ! . ……………………………………….. 225 . ļ. Ŀı Ŀı Ľĳ ı Ľ ...... 227 .R., .., .., .., M.I., I .., R J.., .., .. G -G U V ...... 230

W 3. G …………………………………………………………………………… 234 .., .M. T ...... 234 R.., .R., .. T ! -F ISUU …………………………………………... 238 .., .., .., R. H . \ ………….. 241 .., R .R., .., .., J . T Limonium Michelsonii Lincz. ! ……………………… 246 JĽ' .ļ., Z.., I.. «Ŀ» ı -ĳ ĳ ĳ H SY. . 1 (33). 2012 5

ı Ŀ ...... 251 U.. R (Heteroptera) U U «Ŀ Ŀ» ……………………………………………………………………… 254 U.. X ! (Heteroptera) ...... 257 M.., .I., ] .L., I.. G Poaceae Barnhart. L F …………………………………………. 260 .Q. U! ……………………………………………………………………………………... 263 I.. c (Culicidae) W X!-^! ……………………………………………………………………………………. 266 R .., .., .M., .. Inula Ľ ĳı !, ! ...... 268 Rĵ .. Xĳı – ...... 271 R .R., .., .., J . T !! ! Limonium Michelsonii Lincz...... 272 R.., .I., M.., .. W W – G R………………………………………….. 279 .. R (Insecta, Coleoptera) I S U U «F-c» ………………… 281

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H U R (03.01.1932-03.05.2001) – – , , ! WWV. U ! ! ! , ! 1949-1954 . 40 . U 30 ! , – , – , ! ! (, , ) . H U . K ! WSI , ! ! ! . V! H.U.R . U 1934. F-F, , ! , ! . H U ! ! . K ! ! M F. U , : 10 , ! . H 1954-1957. H.U.R ! FS WWV, ! R H , . H 1961. , R . V ! . U H U , ! FS WWV, ! F . H ! ! " , ! . Y, ! ! ! . H 1962. H.U.R ! IY ! Alma Mater, ! - . T !! , ! ! " . X. U , " . H 1965. . S. M. . H. U. R ". H R ! ! . . V !. , 1960- H.U.R , " ! , ! , . W " F- W, G-, , , . c . Y. H ! « », ! 1973. ! !. H , - , ! , , ; , ! ; ! ! ! , ! , " . H SY. . 1 (33). 2012 9

S H U . T . H , 20 . H : !, ! ! !, !, , ! !, , ! . ^! ! ! H.U.R ! , !. c , . , , , . Y-! ! H! U , , ! ! . S, 1966. ! " ! . . G , ! . H 1971. H.U.R , !! " , ! " 2001 . H 1976. . H U , , ! , . c !! (1979-1983.), ! (1983-1986.). X ! IY H U, , . H 1974 . H.U.R U - " , ! " . H 20 ! ! I W WWWV, ZSKWT «] ». X, ! . - ! " ! !. ! ! , ! ! ! . V ! ! " ! , ! ! . T . , " . M !, ! RIY, cRcL FS WWWV, FS WWV, !, U R WWWV J 1 HJS . H 1980- H! U «V », ! !. , 5 . «V » (X.1.-1986; X.2.-1987; X.3.-1988; X.4.-1989; X.5.-1992) – H.U.R. H . T , 85 24 . U , ! . H! - , . T !. T ! . H «V » ! . U " . S , , 10 Y . . 1 (33). 2012

. H 1994 . V 12- . S H U : «U F-A (1988), « WWV» (1989), «! » (2- ., 1991; 3- ., 1996), «L G» (1998), ! ! V . G (1992), R V – (1993). X H.U.R ! . H 1960-1990- ! !, , . U" !, , - . 1980- 2,5 . . W 1980- H.U.R ! G- . T ! , ! , ! , , ! ( Y-!) , !. «J», 1987-1990 . ! IY, FS - ! ! " . . V ! , G, FS WWV ! XcT ! ! . T ! H U , " – , , !. U ! , ! ! , , . T H.U.R , , ! , ! . X, 1990-1991 . , XcT -! IcW; 1992-1994 . R ; 1994-1998 . !! «» ; 1996-1998 . W^F - ; 2000-2001 . !! . " . H H U 100 , 13 , WYFV SV V . U H! U ! G, ! ! ! , ! , . H H U 140 , . T , , , ! , , ! , !, , . T ! " W F , , !! ! 1991 . F-F. G , H.U.R W . F-F, U. T W ! R WWWV ! HIGT, U " W R R WWWV, FS WWWV, H SY. . 1 (33). 2012 11

U FS WWV . J Y IY, FS V, !. M H.U.R ! «M ! WWV» (1985), U H W WWV, U U HIGT, R WWV, U , «H ». H H. U. R ! IF V, V, R V, «U » VFS V , «H ». , H! U, !, , , ! ! . R !, , , , 1970- , ! !. H , H.R, !, ! ! . c . J ! R . T ! H! U, , . T , « », . T - . J . Q H U !, : , . M! , H U , . R , !. U ! ! !. S «H» IFM-21 , . H , ! . S : ? H . S ! - ! . H U . H !, !, ! ! . M , . T ! , , , !! . , 1- !, , . S ! ! , . U! ! ! ! H. U. R, !, ! , ! . W! ! – ! , , , ! , .

12 Y . . 1 (33). 2012

YJ 597.745 .. UTQR T TMTTM TZ TMRTL U - !, e-mail: [email protected]

, . W ! ! . c «T » [1], ! V . W, I F TTS ! , !! 2010 R !. T !! , ! 20% , ! [T! .-T! .]. W , !!! ! (, , !) ( , , ..), , [5]. U , , , , !. T! ! ! XX 4 . 1900 90-100 . . V !! : , , , , , . V (! !, !! , ), X [T! .]. V ! ! 60- . H 1966 R « RWTU» («Red Data Book») 79 . H ! « RWTU» ! !. X 1977 193 , 1986 - 287, 1990 - 713, 1996 734 . H ! ! , ! !, «!» , . H , ! . H , . , ! . H-, ! !! ! , , !! . H-, ( ! ) ! . . U ! : !; !; ! . , ! , ( (Acipenseridae), (Salmonidae), (Clupeidae)) . X ( (Huso huso), , (Acipenser gueldenstaedtii)) R W T U «S!! !» [6]. W! ! Mnemiopsis leidyi (Clupeonella). F ! XX ! . U !, . W!, ! – - (Pleuronrctes flesus) (Atherina boyeri). J! ! F ! ! ! [7]. X F-W ( , , ) «U , , » (-U), ! ! ! ! , ! (! ) 18 V U [8]. H SY. . 1 (33). 2012 13

T ! ! , ! , . F . Y! ! , !, . J ! ! V U, ! . , ! , . S ! ! V !!! , ! , ! !! ! , ! !!! , ! !, , ! TTUX .. U , !!! !, !! ! , . S , ! ! . H ! V ! ! R (IUCN) [9], , ! !. H ! ! U! U V: - 357 25 !! 2007. «T ! », ! Y, , . F (R) , , , ! .M. - 3 1106 28 !! 2008 . «T V 2009-2011 » U ! , ! , !! !. U ! ! ! M V 9 ! 2004 3 593-II «T , » ( ! ! ! 21.01.2010 .), ! ! ! , .2. 20-1. : «W ! ! V ! , ! ». , M ! , , ! ! . H , , ( !, ! ! ), , ! , . (SV). H U! U, , ! ! . 45 11 , ! . S !! ! , . T , , , Y- . X ! , ! , ! ! U U. S ! , , , , !, ! , , ! . U , ! ! ! , ! ! ! ! !, , . T ! ! ! . T ! ! . J! ! !! ! [10, 11], ! [12], [13], [14], 14 Y . . 1 (33). 2012

!! [0 Viable populations for conservation// Soulé M.E. (ed.) – Cambridge. Cambridge University Press. - 1987. - 236 p.]. T ! !! - -. U- !, ! - ! . U- , , !, ! ! - . U! ! - , [15]. W! , ! ! !, , ! [16]. U ! , ! ! , . T ! ! [17]. H U U V 10 !! 2000 N 1692 «T ! ! ! V 2030 » , « 104 23 (22,1%), ! , ». U ! «» - ! , , .. [18, 19]. X !!! R , ! ! ! (Brachymystax savinovi) (Thymallus arcticus) [20]. M ! , !, , , ! , , . ! !, . S, ! ! .X U!, ! ! ( ! ). F ! JS- ! R !. V , . W ! , !!! ! !. T ! ! !. W, ! !, ! , , , , , , !, . H Y- . H ! ! ! , ! ! ! , ! ! , . F ! , ! , , !, . S ! ! , , ! ! , ! , ! ! , !. T « !» « » , ! (, ! ..) , ! ! ! , ! ! , !! . U - ! !!! ! . !! !, . W , ! ! ! Y, , !! , . H SY. . 1 (33). 2012 15

T ! «» ! . X ! , . X ! ! H. K ! ! - . T, , - ! , ! , ! ! . T ! ! (, ). , ! , , . U , ! FT, ! !, F . U ! !, , ! , - !!. X ! , , ! -G R F. W , , , !. U ! , ! !, !! , , , , , . J! R- ! V 2011-2025 ., U ! 309-1-8/14 14 ! 2011 ( 11) R ! V. J! ! - ! , ! !, !, ! VRW ! !, \ . J! - SV. S ! V 2004-2015 , , «H ! ! ! ! !! !, ! ! ! ! » [21]. Q 1. Convention on Biological Diversity. – Rio de Janeiro, 1992. – 05 June. 2. Boris Worm, Ray Hilborn et al. Rebuilding Global Fisheries // Science. -2009.- Vol. 325.- P. 578–585. 3. W! . - 2008. – V: FT, 2009. – 196 . 4. Y . X FT .- V: FT, 2008. - 3 5. - 125 . 5. Tonn WM. Climate change and fish communities: A conceptual framework// Transactions Amer. Fisher. Soc. - 1990. - Vol. 119. - P. 337-352. 6. www.redlistiucn.org 7. R H.F., G R.X., K.H., K M., I F.., F W.L. ! // V! ! V : ! !. – F: G, 2005. – W. 6-64. 8. U U V 21 ! 2007 3 521 ( ! ! 24.05.2011 .) T ! , , . 9. IUCN Red List Categories and Criteria: Version 3.1. IUCN Species Survival Commission. - IUCN, Gland, Switzerland and Cambridge, UK, 2001. – 30 pp. 10. MacArthur R.H. Geographical Ecology - New York: Harper and Row, 1972.- 356 p. 11. Richter-Dyn N., Goel N.S. On the extinction of a colonizing species //Theor. Population biology. – 1972. - Vol. 3.- P. 406-423. 12. Leigh E.G. The average lifetime of a population in a varying environment // J. Theor. Biology. - 1981. - Vol. 90.- P. 213- 239. 13. Soulé M.E. Thresholds for survival: maintaining fitness and evolutionary potential // In: Soulé M.E., Wilcox B.A. (eds.) Conservation Biology: an Evolutionary-Ecological Perspective – Sunderland, Mass. Sinauer Associates, 1980.- P. 151-169. 14. Shaffer M.L. Minimum population sizes for species conservation // Bioscience – 1981.- Vol. 31.- P.131-134. 15. R R.H. R ! : c !. – R.: S,1986. – 298 . 16. Frissell C.A., Liss W.J., Gresswell R.E., Nawa R.K., Ebersole L. A resource in crisis: changing the measure of salmon management// Pacific salmon and their ecosystems: status and future options (Stouder D.J., Bisson P.A., NaimannR.J., eds.) - New York: Chapman and Hall, 1997. - P. 411-446. 16 Y . . 1 (33). 2012

17. Reeves G.H., Hall J.D., Roelofs T.D., Hickman T.L., Baker C.O. Rehabilitating and modifying stream habitats // Amer. Fisheries Society Spec. Pub. – 1991.- Vol.19.- P. 519-558. 18. U J.W., V Z.W., ^ R.., ^ S.. V WWWV ! « » // H. . - 1985.- X. 25, H. 1.- W. 16-25. 19. U J.W., W .F., W Q.., F W.W. V . V. - R.: H. , 1994.- 334 . 20. V ! V : SV / SV. – F, 2007. – 34 . 21. ! V 2004-2015 . T Y U V 3 ! 2003 N 1241. *** G' ''' %' , , ' % ' ' ' '''. *** In given article problems of preservation of a biodiversity of a fish fauna are designated, security actions are analysed and the complex of necessary measures for preservation rare and vanishing species of fishes is resulted.

YJ 502.743+597 R.. U] TUT T TMTTM TJTR TTT TR UTJ TO , e-mail: [email protected]

, , , Q . T ! ! !!! ! (TTUX). W 90- TTUX R !! ! , ! Brachymystax lenok . R [1]. T !! R ! !, ! 1985 [2]. M ! TTUX, . S ! ! . S ! F [3], !! . \ ! ! , ! ! TTUX. S, TTUX . H !, , ! , – ! (Hucho taimen), !! TTUX: ! ! ! - [4]. W , , ! TTUX ! ! - ! . J! , X, . R !.

45 40 35 30 1990 25 % 20 2010 15 10 5 0 3456789101112 , 1. . 1990 2010 , ! – ! 2010 – 6 , ! , 7 X , 1990 12 – . U! ! ! - ! . S ! H SY. . 1 (33). 2012 17

!! , ! R, ! ! ! , , . S ! , , – Cyprinus carpio , Abramis brama, !! , , ! , , !!! ! ! . X, ! , ! ! , . T! – ! ! , , . !, ? , ! ! , ! ! !! ! ! [5]. S, !, [6] !! , ! !. H ! ! ! , .. , , . H ! !. H !, ! ! ! V R, « ! !». X , !! ! ! , ! , ! ! !!, ! . W, ! !, , - ! ! , . U , !! ! , ! !! ! : . ! ! ! . X , . U ! ! « !», , ! . Y !! ! ! . R, 1997 . UNESCO «] ». W ( 2) ! , ! ! !, ! . Y!, (ISUU) , , ! ! !, . S !, 2011 ! ISUU – «», G!, « Ŀ», -, «G», , ! , ! , ! ! . W , ISUU, – . , . R . (. ]), - ISUU, , – ! Leuciscus idus , Barbatula toni . , Thymallus arcticus R . Y ! ! , ! !! ! . S, , ! ! . H ISUU ! , , . U ! ! ! , ! !. H-, ! - ! , ! , . T , . 18 Y . . 1 (33). 2012

, . R ] (ISUU «G») ! ! – Perca fluviatilis Sander lucioperca , Abramis brama . H , , !, – 11-15 . H , !!! ! () ! : , ! , – , .

2. R R H-, () ! ( - ! () ) ISUU ! ! . \ ! , ! - . X, Esox lucius, ! . (ISUU «») [7] , , , ! . U , , ! «». U Coregonidae ! – . M, . ^ . . H , ! . H ! , !! ! ! – . G (ISUU «G»), . ^ ( ) (ISUU «»), . U ( ISUU) . S ! ! , ! ! () [8,9], ! , ! !!. F ! ! ! . X, . Posthodiplostomum cuticola !! Abramis brama Rutilus rutilus. H ! - – ! , , ! 100 %. T !!! . H , . S G ] X (ISUU «G») 2011 ! , !, !!! ! . U ! ! . X , ! () ISUU ! ! !. ! , ISUU ! H SY. . 1 (33). 2012 19

! () , ! ! !. S U ! ! ISUU, , ! ! ! ! ; ! ; ! . W ! ! ! () . , TTUX M (ISUU «»), ! TTUX. W ! ! ! . , !! ! , - ! [6]. Y TTUX Q U – ! ! , . , Q ! ! [10] , !! . W , ! , , , !! . S ! , , Q , ! TTUX . - ! . X , ! R , ISUU ! ! !! ! ! () . S!! ! , ! ! ! TTUX. S , TTUX ! ! ! , , , , , ! ! ! . , , ! . ! HYM , TTUX. Q 1. R H.U. T! // V . F: I. - 1992. X. 5. – W. 408-411. 2. G R.X. R//X R . Y- – 2009. X.1, . 1. – W. 212-218. 3. W H.V., X W.V. V F-W //X F . F: R. – 2004. X.1. –W. 175-191. 4. G R.X., T.., K.H. W ! ! F-W! //G, I F! : !, , . R R . ] 1. 22-26 !! 2008, I- F: VT IFIY, 2008. W. 17-23. 5. G R.X. ! R, ! , !//X R . Y- – 2009. X.1, .2. – W. 90-101. 6. M V T ! !. F, F, 7 ! 2006 3 175-III MV 7. I F.., J K.. T //T . V 1: T ( 1964-1998 .) F – 2008. – W. 24-26. 8. U . Q! //http://fish.kulichki.net/?p=262 9. Q! //http://berdfish.com.ua/modules.php?name=News&file=view&news_id=181 10. H Q U ! ! V . R . F-2005. – W. 54. *** K X ' % %', ' % %' . .

20 Y . . 1 (33). 2012

*** In work ways of the decision of the problem questions connected with the organization of fishery, by carrying out land improvements, working out of plans for development of a fish economy and carrying out of researches under programs of Annals of the nature in reservoirs of especially protected natural territories are discussed.

YJ 575.8 Kazuyuki INUBUSHI REVOLUTION OF LIFE FROM OCEAN TO LAND, INTERACTING WITH GLOBAL ENVIRONMENT Graduate School of Horticulture, Chiba University, Matsudo, Chiba 271-8510, Japan, corresponding author: T +81-47-308-8816, F +81-47-308-8720, [email protected]

Revolution of life from ocean to land, interacting with global environment Origin of life is depended much on the geological history of Earth. According to several literatures, formation of Earth assumed about 4.6 billion years ago (BYA). Primitive Ocean started to absorb atmospheric CO2 which created lime stone and reduced global warming. First life activity is estimated to begin about 3.8 BYA as the evidence retained as13C abundance in sedimental rock. Microfossil was also found in sedimentary stone formed in 3.5-3.2 BYA. Magnetic field strengthened protection of life development against cosmic rays in 2.8-2.7 BYA so that oxygenic photosynthesis became active in ocean surface, then cyanobacterial activity became also higher which remains as living fossil, stromatolite. Cyanobacterial can generate O2, fix CO2 and N2. Bigger cells of cyanobacteria, named ‘heterocysts’, can fix N2, by using carbon sources which were derived from other cells by photosynthesis. Banded iron formed in 2.5-2.0 BYA by oxidation of ferrous iron in sea water and precipitation to the bottom of ocean. Eukaryotes appeared from prokaryotes, cyanobacteria became chloroplast of algae and other organisms to create more O2, and also anaerobic microorganisms became aerobic to adapt more O2 in the atmosphere. Such genetic developments are not only horizontal but also vertical or ring wise. Ozone layer was created from excess O2 in stratospheric zone, and then sea level went down, so that life was landing about 500 million years ago. Geological history and revolution of life indicate adaptation of organisms including soil microorganisms to more oxidative and drier conditions. Fossil soils or Paleosols were developed from sediment about 3.2 BYA after microfossil was found. According to USDA Soil Taxonomy, Entisol formed first and Inceptisol followed. After banded iron formed, Laterite appeared. “Green clay” was also created under anaerobic conditions, although it might be extinguished when O2 accumulated in the air. During these changes, final product of organic matter was also shifted from methane (CH4) to carbon dioxide (CO2). In present aquatic sediment, however, deeper layer is still anaerobic and ferrous oxide is reduced to ferric iron. In paddy soil, subsurface layer is also anaerobic under reduced conditions, almost no O2, forming CH4 and rich of ferric iron with greenish grey color, which would be similar as “green clay”. After life landed, more diverse soils were created such as Alfisol and Ultisol and microbial evolutions took places to adapt development of terrestrial ecosystem. Blue- green algae, or cyanobacteria, in marine ecosystem started to adapt high salinity then moving on soil surface, where was much under drier conditions. Cyanobacteria can grow symbiotically with Lichens, Bryophyta, and Pteridophyta. Cyanobacteria in present can still grow on salt-affected arid soils over the world with special salt-registrant mechanisms, and act as moderators of sever soil environments against large temperature fluctuations and soil evaporations. After forest formed in Carboniferous (about 360-290 million years ago), organic matter was accumulated more on soil surface, which may enhance heterotrophic and rhizospheric microorganisms. In Tertiary, dry inland and grassland area developed in the huge continents and grass vegetation formed thick root mat and mychorhizal symbiosis between plant root and soil fungi was also developed. Global climate is changing rapidly nowadays due to intensive human activities, and dry lands are also expanding recently especially inner continents. Mychorhizal symbiosis and other plant-microbe systems with endophytic microorganisms have benefits for plant nutrients such as P, Fe, Zn, Cu and water uptakes by plants and increase resistance to root pathogens. *** X''' , . X'''' L' . *** V , . U M.

H SY. . 1 (33). 2012 21

YJ 597.554(574) .. I LMb U TM M. IJM\ – ZTR UTUQ\TTO JU\ XTT «SV», FT «F!», F,

T P - , , . S , , , (), 40 . P , . T . R , – , . T 50- . H ! ! : , ! ! . H ! ! ! () . X ! !, ! [22], . .S.V! [40] 140 Cyprinidae ( 39 Cyprinidae Leuciscinae) : , , , . T ! . S !! ! ! . V . U ! ! , . W ! . R , ! Y. M 20 ! Y, G, Q . I ! , ( 26%) , ! 8-12% V W W V – . T, , 2,0% [28,29,30]. I ! -, 1946-1955 ., !. R , , , . H Q W V , , ! ! . U . J V W. X «» ! , 8-12%, 100%. , ! [24]. I : ! [7], ! - [8], ! [34] . K Y (, X Y) !! 20- [51]. X ! ! , ! [13]. 22 Y . . 1 (33). 2012

U! ! ! ! !: ! , . H ! ! ! 90- . I ! : 14 !! , 12 – !. J! – « » : – !, ! – ! [19]. U : 1,4 – 2,2 , 1,7 – 6,3 !. T! ! ! , ! [20]. U! !! , ! , . J ! ! ! , ! ! ! !, ! [53]. K! ! ! [14]. I ! . R , ! [15, 41] ! !, [23,37-39,52]. U . X, ! G ! 0,25 – 7,0% [25]. X ! ! (5 . 355 . ! 540 . !) ! .T. [1]. S !!! : 160 ( ) [18]. H WWWV Q.W.G [4] , , , , , , , , , , , , , ! , ! !. R . X, W! 2,0% [46]. H ! [55], . X, ! . V ( J!) , , , , , [57]. I ! ! , , . W [56] ! , . S..S [31] J 70-80% ! . T : «… ! , ! ». K ! [33,42,49] , ! !, ! !. U! , . S.U.W [42] ! 1946 . . F-J ( . R ) . ( .G) ! . T ! ! ! ! , ó (37) (26), – . I.H. S [33] . W (. F !) – . W ! ! . V.I.U [39] Y ! 99 . H - 20 [26], . R! ! . H . ] (!) , ! , ! H SY. . 1 (33). 2012 23

, [36]. I! . ] Q.W.G [3], I.H.S [32] U.F.J! [11]. .F.U [36] 17- !, , ! ; ( ) – . W ! . X ! : I.H.S [32] ! 33%, I.R.J [12] 1958-1967. – 100%. X X. .F.X [48] , «! , -, ! , !, , , ! «», ». U ! . X ] W- J, ! ( ) R -. H !! Balitoridae G ( 36 ) . U, ! (! !, ) [2] . W ( 40%) ! !, [58]. M W ! ! ! . W M, L-L, G X 70-80% ! , - ! [50]. I , : 8000 /, 140 (. G, 1982 ), ! 9,0 / (. G). T ! «» , ! ! . I! , – . ! ! ! . I ! ! ! ó ! ! (20-22). T ! ! – , ó (42-44). U ! – . c !, ! ! , ! . U ! ! ! ! [10]. \ ! ! : 100% . !- ( ) : ! , !, [45]. S : 1,8-2,5 , . H : 2272 . 1914 . ( , 50 ) 29099 . 19823 . 151-200 . H , (: --), - - : – 28- 32., 20 ., ! 40-47 ( – 40,6) ., – 12-20 ( – 15). c 15 50 3+. W, - (0,1%) . T ! ! !. K . .G [54] ! . S 1844 . F ! , , , ! . S ! , ! , ! . K : ! , , , , ! I.H.S [33] ! . W , . U.I.G [5], ! ! . Y -H !, , « ! ! ! ». U! 24 Y . . 1 (33). 2012

( ), ! . J 1949 . G F . X [9]. V ( ) ! 2% ! , ! ( 1+ 6+), () . Y , !!, ! ! . H . U Microcystis . S ! . U [27]. R ! . J H.W. [16, 17] , ! . ..X [47] Y (! 3,1-7,3% (n=1202 ) !! ! (28-30°). ] ! ! , ! !. G.I. [21] -Y 45,5% ! , ! , , ! . , ! ! , . F ! ! !! : ! , . U !! ! , ! ! . I ! ! 34% , - – 20% ( . G), 34%, ! ! 50% – 4,38%, ! – 2,90%. I 0,4-0,5% . S , ! - , 50%, 2,5%, ! 10% (n=200). I - , ! , , ! , . ^ ! 60- ! . H , , ! . X, , ! , ! , ! , ! . (1955-1988.) ! ! ! 27,3% , ! ! , ! , , !. S X L , . G! W F . H . X ! , ( 1974 .) – !. S ! . H ( 9,4 /) . L ! ! !. T. G! W, ( 16,7 /) 1965 . ! . M ! ! !. ] ! – ! . H . R, 1964 . ! , ! ! !, !. V 21,8 44,6 ( 27,0) /. H ! ! ! ( ) !!! ! . V 5,1 70,0 (!! – 33,5) /. H (, , !, , , , , ) 5,0 131,0 (!! 78,0) /. I . H SY. . 1 (33). 2012 25

H . V , ! ! , ! ! 17% (n=600) 1976 . U . H . W , -, ! ! ! 1964 1968 . U ! 1968-1969 . , ! ! 12% (n=100). W ! ! ! ! ! ! . G, ! . W ! ! 34% (11858 .) , . ! . H . X ( 1/) . ] ! . V ! ! 105 /, 9-10 – 108-125 /, 90 / . H . Y F !. ] ! ! 132 /, 37 /. U ! , 44 ! (76%) ! !. ! 25%, ! . V 6,0 110,0 /, 10 34,0 /. T. L H ! (F - !) ! ( ! ) 19 ! 9,0 /. . X! . G! W. M 200 / ! ! ! ! . W! . K ! , ( ), [44]. J! ! ! ( R ). H 2004 (n=210): – 72,9%, – 20,9%, – 6,2% [35]. ! ! ! ! ! ! ! ! !. V 5,0 40,0 (- 14,0) /. H . V ! ! 25 . V ! (1955.) ! 72,0 /, 40 – 34,8 / (!). S – (40% ). H R !. V! «» , – !: ! 64,5 100 ( 85,1)%. ! ! ! ! ! . H , ! !, !. S, !, ! (. R). Y !, ! ! ! !. U !, , ( ) . X, !, ! (W). MQM] H! ! ! ! ! ! , . I- ( ) ! ! - ( ) ! . H , ! ! !! ( ) - ! !. T !, ! ! ! ! – !. H ! ! ! !, !! ! -, ! – -. Q ! ! ! ! (), ! 26 Y . . 1 (33). 2012

50- . T : () , ! ! . T! ! 50- , ! ! ! ! ! . I ! ! , ! , . , ! . U , - ! ! !. S ! ! ! : ! ! W . T! ! ! ( !, ) !. I! , , . T, ! ( ) !. S! ! ! !, ! ! !!! ! ! ! ! ! ! . Q 1. F F.M., Z.S. T Coregonus lavaretus pidschian (Gmelin) ! Coregonus peled (Gmelin) ! T. //X . Y. . FS WWWV. W: 1970.-.72.-W. 52-55. 2. G I.. R! ! (Balitoridae; Cypriniformes; Osteichthyes), G . //F . . . . F: 2010.-20. 3. G Q.W. V. V. III. H.2 R.Q.: 1914. -W.614. 4. G Q.W. V WWWV . - FS WWWV. .2. R.Q.: 1949.- W.821-831. 5. G U.I. T! ! Y. //V , R.: 1925 .IV 31-2. 6. H G.H., R F.U., ^ S.I. W Hypophthalmichthys molitrix Aristichthys nobilis (Cyprinidae). //M. . R.: 1979. . 58. H.2. .- W. 190-196 7. H G. I ! – . //V . R. 1972. 36.- W.9. 8. I M.F. I! ! . //G! U. R.Q.: 1962.- W. 77-79. 9. I F.. . //W - FS WWV. F-F: 1956. H.1. - W.261-269. 10. I F.., W H.F. (Carassius) . // Tethys Aqua Zoological Research, Almaty: 2002. vol. 2. – P.33-37. 11. J! U.F. V ] ! . //X . : 1936. . III, H.1. - W. 67. 12. J I.R. V Barbus Cuvier, 1817 – Y. //V , . 3. (). F-F: S. -1988.- W.24. 13. K F.V., Z.F. T . //W Y ! ISTV. Q.: 1979. H.10.- W. 42-47. 14. F.. X! . //«U! » R.: 1975.- 431. 15. . U M 1875 . // X WU T . WU.: 1878. . VIII. W. 78-109. 16. H.W. H! ! , . // J FS WWWV R.: 1945. . 47. 37 - W.521-526. 17. H.W. W! WWWV. // ! HSTV. Q.: 1948 . 26. H.2.- W. 145-170. 18. Z.H. R ! Abramis brama (L) Rutilus rutilus L (Cyprinidae: Leuciscinae) //F . . G.:- 2009.- 21. 19. Z.R., W H.U., ^ F.. R! (Cyprinus carpio Linne x Carassius carassius (Linne)) W . //c . «G».- F, 1995.:- W. 38-43. 20. Z.R., J H.G. U ! W . // c . «G». F: 1995.- W. 31-38. 21. G.I. R Y . //V -Y . R .-.. U: 2001. W. 34-36. 22. W.I. M ! . «S». R: 1968.-279 . 23. M.R., F! F.M. R . // ! FS FWWV, W. . H.5. –G: 1969.-W. 64-68. 24. Q S.F. W Coregonus ( ). // W . R: 1958. H.2. - W.127-129. 25. R c. U!, !. R. R.: 1974.- 460 . 26. R W.H., W.H., F.H., Q.H. W Carassius auratus x C. carassius (Cyprinidae) ! J W J. // J. S. FS Yii. i: 2009. 36. - W. 191-197. 27. R H.U., J I.R. F . //V , .3. (). «S». F-F: 1988.-W. 247- 248. 28. S S.H. T Y. // V !. 1962. 34. - W. 8-11. 29. S S.H. H! . // X Y.. ISTV. W: 1964. . 6. - W. 19-60. 30. S S.H. R . // X Y.. W SV. W: 1966. . 7. - W. 185-196. H SY. . 1 (33). 2012 27

31. S S.. R , ! ! . // M. R.: 1948. 34. - W.343-353. 32. S I.H. V ! . ]. // K . ! FS WWWV, R.: 1931. . 32. - W. 255 33. S I.H. V F !. I ! . // R WWWV, R . S! !. M. R.: 1940. H.1 (XIV).- W.154. 34. S F.J. I . // F! WWWV. R WWWV, 10-15 ! 1963. .. F-F: 1963.- W.276-278. 35. T (F..I , K..J). V IV – F . G! W-T. «Xi». F: 2011.- W. 62-65 36. U .F. R . ]. (Barbus, Cyprinidae, Pisces) // ! FS WWV, W! .. : 1963. .5. H.2. - W.47-56. 37. U Z.F. T Cyprinidae. //X Y.. WSV. W: 1971. 313. - W. 103-109 38. U V.I. I! , . // U . X: 1962.- W. 221-223. 39. U V.I. T Abramis brama (L) Rutilus rutilus lacustris (Pallas) . Y. // H R.: 1964. . 4. H. 3. - W. 463-465 40. V! .S. I! Cyprinidae. //H R.: 1979. .19. H.6 (119). - W.1025-1042 41. W Q. V V. R.: 1911. 3- . - 1062 . 42. W S.U. I ! . //V . «S». F-F: 1966. H. 5. - W.352-353. 43. W H.U., W X.H. H (Cyprinus caprio Linne x Carassius carassius(Linne)) . // c . «G». F: 1995.- W. 44-46. 44. W H.F., ^ F.., I H.., V F.F., F S.M., F ^.R. H! W-T. //G. ! W F . F: 1986.- W. 293- 295. 45. W H.F., I F.. T ! ! ! . // : !, , !. R , ! 70- SY . F-. F: 2004.- W.200-202. 46. W X.. Abramis brama (L) Blicca bjoerkna (L) W! ! . // H . R.: 1975.- .15. .6 (95). - W. 1120-1123. 47. X .. F ! . // M. . R.: 1966. X.45. H.12..- W. 1826-1834. 48. X .F. T X . //Y. M. .-. I.Y. : 1955. H. 5. - W. 145-147. 49. ^ I.. V F-J. W! . // X M FS WWWV Q.: 1950. .9. - W. 38-43. 50. ^ F.. R (Coregonidae) . //V . X.5. F!, . F-F: 1992.- W. 316-329. 51. _ .I., Q.S. T . // J FS WWWV R.: 1954. . 95. 33. - W. 685-688 52. K.S. I! ! ] (M! W) Rutilus rutilus ! Leuciscus idus L. //H R.: 2003. . 43. 31. - W. 110-117. 53. H.S., W Z.H. I! ! . // J FS WWWV. R.: 1998. . 358. H.5. - W. 716-718. 54. Baer K. Über doppeleibige Missgeburten ober organische Verdoppelungen in Wirbeltieren. // Memoires de L’Academie imperiale des sciences de Saint Petersbourg. Sixieme serie. Sciences mathematiques, physiques et naturelles. Tome VI. 1845. 55. Demandt M.H., Beergek S. Identification of cyprinid hybrids by using geometric morphometrics and microsatellites. // I.Appl. Ichthyol 2009. 36. - C. 695-701. 56. Makino Sajiro, Ojima Jashio, Matsui Joshiichi. Some cytological features of sterility in the carp-funa hybrids. // Annot. Zool. Japon. 1955. V.28. 31. -P.12-16. 57. Rĕdulesku Ion, Vasiliu Natalia. Notĕ asupra hibridului Cyprinus carpio L. x Carassius carassius L. din Ghiolul Rosca. // Bul. Inst. cercetĕri piscicole. 1956. T. 15. 31. p. 67-71. 58. Wheeler A. On the population of roach (Rutilus rutilus), rudd (Scardinius erythrophthalmus) and they hybrid in Esthwaite Water, with notes on the distinctions between them. //I. Fish. Biol. 1976. V.9. - P. 391-400. *** J ' ' ' – ''' ' '''' . X '' ' ''''' '' '. R'' ''' . *** Introduction of carps and cazans into lakes with predominance of the crucian carps presence, is foster to increasing of fish capacity of water body. Crossing between crucian carps and invading species (immigrants) do not occur.

28 Y . . 1 (33). 2012

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1.1 N[XNQIN TL 597 I.R. =Q ľM= R UĨO TORJĨ QĨİ TQTI R M=I= MJI= LĨJO « - » L^W [email protected]

R '' ' %' . - Ľ ı . W = Ŀ . W ^, Q, K, W-X, ı Ľ .. Ŀ !. Wı Ŀ ı Ŀ . = Ŀı 130 2 - . = Ŀ X-F . 1970 . . M J R ļJ=RQ= = Ŀ 2011 . - ! Ľ = Ŀ – ! , Ľ, Ľ ı ĳ ĳ. W ! ĳ ! 15 18- , ! ! 9 12 – (1-). ! ı !, ! Ľ ! [1]. F Ŀ 30-120 , Ŀ 40-60 ĳ. M ļLQ= Lļ TQJ U = Ŀ ı 30 ĳ , ı Ľ . Ľ ı ı ı – .

ı : 1- W Ŀ 8F- G Ŀı 2- Ŀı 9- ^ı 3- 9- ! 10- Ŀı 4- Ŀ ( ) 10F- Ŀ Ŀı 5- Ŀ () 11- K Ŀı 6- 5- ! 12- ı 7- Ŀı 12F- X Ŀı 7F- 7- ! 13- Ŀı 8- ı 14- ==R- 15-Ŀ ı 1-. ĩ ĭ ĩ ĳ'

X (Abramis brama) Ľ F ı ı Ľ - . L ı Ľ ı ı [2]. H SY. . 1 (33). 2012 29

L . Wı Ľ ı ı . G , Ŀ, Ľ . W ı 1970-73 . 51,5 ı Ľĳ . X ı 40% . Wı 70% . 2011 . 567,1 . ĳ ı 25,4 . 2011 . Ľ ı Ŀ , Ŀı Ŀ , Ŀı ĳ () Ŀ ĳ. T Ŀ Ľı ( Ľ ). Wı Ŀ Ľ ı – , Ŀ . X 4 , Ŀ 5-6 . F (V - 4 , W - 6 ). Y Ŀ (!). L , Ŀ, ĳ Ŀ . X 1,5-3 ı Ŀ Ŀ (, Ŀ Ľ .), - . Y Ľ ı ! ı , ı ! . Y 17-19°W [3]. X , Ľ . . ı , ı Ŀ 794,5 ı ı. M 2011 . ı FLX 143,8 ı . Ŀ ı , 2,0, 1,8 ı. F ĳ ı ı, ! Ŀ 1:1 . G 2011 . Ŀ ı 1 13 13 . F 25,3-29,0 6-7 46,1% - (2- ) [1]. 140

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2 . ĩ ĩ ı ı ĭ ĭĵı (2007-2011 .) X !!ı ı Ľ Ŀ , . 2011 . Ľ !!ı 13 , ı 15 , !! . Xı ! Ŀı Ŀ 1 Ŀ. 1- = Ŀ' ĩ ĭ ĩ ı Ŀ ĭ ĭ Ŀ'' ( Ŀ') FLX, L l, Q, q, ı. - ı. - ı n 2006 25,4 421,4 391,0 2,0 1,6 195,5 413 2007 24,0 336,5 295,3 1,9 1,7 - 208 2008 26,0 379,0 339,2 1,9 1,7 122,4 665 2009 25,7 372,5 345,1 2,0 1,8 136,2 535 2010 24,3 348,0 305,0 2,0 1,7 140,8 760 2011 25,4 392,0 338,0 2,0 1,8 143,8 629 30 Y . . 1 (33). 2012

2007-2008 . 2009-2011 . , ı, Ŀ . ı ! Ŀ ĳ 2 [3]. 2- = Ŀ' ĩ ĭ ĩ ı ĭ ĳ ĭ Ŀ'' () ı. FLX, W l, Q, - ı 25,4 392,0 2,0 19,8-453,6 F ı 25,6 415,0 - 17,6-167,4 ^ 23,2 256,0 2,0 - W! Ŀ 24,3 445,0 - 112,0-345,8

2- Ŀ W! Ŀ, F ı, ^ ! Ŀ . W = Ŀ !!ı ı , ı, Ŀ . !!ı Ľ Ŀ. ļJ 1 T ! / , TJY ( ) , G-F . V G- . U //T SV/ SV – F, 2011. – W. 66-67. 2 V . 3 . - F-F: «Ĩ», 1992 . 128-158 . 3 G F. F., R H. U., W S. W ! // G . 3 9. - F., 1975. – 71 . *** P .N. U . G , . *** In article the biology and a modern condition of the widespread and numerous kind of fishes - bream the Kapshagai reservoir and the river Ili. Trade kind. Biological indicators satisfactory, it is not marked decrease or deterioration tendencies.

YJ 664.951.014:543(574.5:615.9) .. , Q.. Tb TRQ\ LQ RQQT UIOTIT TJTQ_ XTT « - !» FT «F!», . F,

N P . , . H ! !! !! ! . U ! !! !! ! , ! ! ! ! ! . W !! ! ! ! , !, ! , !! , , ! , . S , .. ! ! ! [1-4]. T ! ! ! ! . S !!! , ! . X . ! , , UJ. ! , , , , R. G. F, X, ., ! [5-7]. H SY. . 1 (33). 2012 31

W , ! ! S !. M 20- !. RQ RTJ QJTO H - 2011 . ! , ! , , !, ! . S 30 . , . F ! ! [8]. MQb QJTO TLJ H 1 ! ! ! ! . H 2011 . 20 / . Y ! 2011 . ! 2003 2004 ., ! ! !. V ! ! , , . X 1 2003, 2004 2011 ., / I H Zn Cd Pb Cu Q 5,07 0,10 0,77 0,47 2003 W 4,56 0,08 0,68 0,41 L 5,63 0,10 0,56 0,56 Q 3,71 0,11 0,70 0,56 2004 W 3,12 0,10 0,65 0,34 L 5,90 0,08 0,52 0,50 Q 20,0 0,06 0,49 0,11 2011 W 15,0 0,07 0,49 0,69 W 11,0 0,24 0,10 0,57

] ! . T ! , ! ! . X, ! ! ! ! 7+ 12+. R ! ! ! ! ! 7+ 9+, ! 7+, ! 3+ 7+, ! ! ! . S ! : 41 / ! (12+), – 28 / 21 /. R , ! ! . - !. U ! ! , ! , ! , .. , ! , . R ! ! ! . H 2 ! (S) . U! ! ! . S 425,5, ! . V ! ! ! , ! . V , ! ! . T ! . , ! ! . S ! ! ! : 2003 2004 . ! ! , 2011 ., ! Zn> Pb > Cu >Cd S ! ! ! ! , 2011 .: ! Zn> Cu > Pb >Cd ! Zn> Cu > Cd > Pb 32 Y . . 1 (33). 2012

U ! ! 2011. ! ( ), !!! ! ! ! . X 2 2011 . T Zn Cd Pb Cu Q 425,5 20,7 96,1 2,4 W 319,1 24,1 96,1 15,3 W 234,0 82,7 19,6 12,7 W!! ! , /³ 0,047 0,0029 0,0051 0,045 S ! ! ! . T , . ! ! !, ! ! - . W , - ! (RJY) 10 /, ! 0,2 /, – 1,0 / 40 / [9], , !, RJY. Q 1. F S.F. ! ! . Y // R == R. -. . «] L». F, -2004. - W. 40-42. 2. F S.F., QX.., S W.X., G.. T ! !! ! . G // I! !. - 2003. - 31. - W. 99-114. 3. F S.F. T ! ! F // I! !. - 2005. - 34. - W. 168-177. 4. G G.., S H.. X! . G // U ! . G ! . G, -1992.-W.21-22. 5. F S.F., F ^.F. T ! ! . // c! . F:G, -2008.-W.177-184. 6. F S.F., X Q.X., H X.. R ! . // R XII R. -. . «F! – / , W R», F, -2009.-X.1.-W. 394-396. 7. S ! 2010 . F, -2011.- 241. 8. RY 08 – 47/008 «R , , !, ». X, -2002. 9. I ! . W WU 3 4.01.01.03 (Y. 11.06.2003 . 3 447), F, 2006., 322 . *** %'' ' ' ' . R ''', '' % %ii' . *** Level of accumulation of some heavy metals in fabrics of food fishes of the Kapshagai water basin is investigated. The tendency of appreciable strengthening of biomigratory activity of zinc and copper, and also inadequacy cumulation toxic depending on age of fishes and area of their dwelling is revealed.

YJ 597.554.3 (282.25) .R. R TIT ITQb\ (BARBATULA TONI) M JQ TIT O F XTT « - !», -mail: [email protected]

: (, , ) P. W Barbatula toni (Dybowski,1869) – , ! . c . V M H- . S W , , M . ! , ! ! ! . U! ! 174 , ! ! . . RQ RTJ QJTO R 10 ! 2011 , N 47032.49, E 85001.51. H , 20 , 32 ! . J! H SY. . 1 (33). 2012 33

! ! . G (, ) ! .. U 1. H !! S.. ] 2. G Fish. R .. U. U 26 5 ! ( 1). V - (.. U, I.. Q) 3. U (XL). MQb QJTO TLJ W Y , T . K F, ^ W, W . H ! 4. W Barbatula toni (Dybowski,1869) ! G Balitoridae (Swainson, 1839). U ! Nemacheilus 5. R . T ! -!, !. X ! . G ! . I . Y 3 ( ). U!! . . R : D II 7; F II 6; P I 11-12, 11; V I 7-8 7, 7,4. 38 . U : ! 21,5 23,9 % 22,5 %. H 12 %. Y ! ! ; ! – 14,66 %, ! – 6,94 %. H ! – 51,74%, – 37,32%, ! – 22,62 %. X 1 R U U R m .. W Min max T! 94 3,12 13,96 14,86 81 115 ! 82,4 2,46 11,01 13,37 72 99 J 7 0,00 0,00 0,00 7 7 F 6 0,00 0,00 0,00 6 6 V 11 0,12 0,55 4,80 11 12 V 7,4 0,12 0,55 7,40 7 8 % 22,5 0,21 0,92 4,12 21,5 23,9 12 0,34 1,53 12,75 10,5 13,8 ! 14,66 0,32 1,45 9,89 13,5 16,8 ! 6,94 0,16 0,72 10,37 6,2 7,8 ! 51,74 0,25 1,11 2,14 50,3 52,9 ! 37,32 0,23 1,05 2,82 36,4 38,5 ! 22,62 0,39 1,72 7,63 20,2 24,9 ! J 9,32 0,25 1,10 11,87 7,6 10,5 ! J 11,96 0,42 1,87 15,65 10,1 14,9 ! F 6,62 0,16 0,74 11,12 5,8 7,4 ! F 10,66 0,40 1,81 16,99 9 13,7 V 15,10,55 2,46 16,31 12,6 17,9 V 11,520,19 0,83 7,22 10,4 12,4 ! V V 33,240,47 2,08 6,27 31,3 36,2 ! V F 17,260,32 1,41 8,18 16,1 19,7 % 40,080,40 1,80 4,49 36,9 41,3 11,940,38 1,72 14,44 10,2 13,9 26,820,44 1,96 7,29 24,9 29,7 43,20,39 1,74 4,02 40,5 44,9 43,60,61 2,74 6,28 38,8 45,6 34 Y . . 1 (33). 2012

J ! (9,32 %) (11,96 %). T . I , – 15,1 % , – 11,52 % . V! , , – 33,24 % 17,26 %. J ! – 40,8 %. I , 11,94 % . J ! 24,9 % 29,7 %, ! 26,82 % . M – 43,2 %. T , 43,6 % . G . ! ! 3,3 9,9 9 , 6,83 6,08 . H - ! ! 1 5 . T ! , 6-7 ( 2). X 2 T H J, R, -, % ! (-) !! (-) !! . 1 3,3 3,3 2 2 1 4,0 2 3,3-6,4 5,68 2-8 5,4 5 20,0 3 6,5-7,3 6,94 4-8 6,0 14 56,0 4 7,5-8,2 7,8 6-8 7,0 3 12,0 5 9,6-9,9 9,25 8-10 9,0 2 8,0 : 3,3-9,9 6,83 2-10 6,08 25 100 U ! . U ! 3 , 6-7 . ! , ! , 1:2,33.U !, ! ; ! ! IV IV-V. U 3398 4315 , !! 3789 . U ., ! : ! ! 22,5 %, ! ! – 28,7 %, ! ! – 48,8 %. W – . U! , 6. , !, ! , 2,12. H !. H ! . S , ! . W !!! , ! ! . X !, ! , !! ! ! W. Q 1. U .. V . – R.: U! . – 1966. – 376 . 2. ] S.. R ! . – R: W! , 1952. 3. Q I.. G!. – R.: H! . – 1980. – 296 . 4. R H.U., J I.R. . V . X. 4. – F-F: S. – 1989. – W. 58-63. 5. J I.R., R S.^., R .H. F V . ! SFS V – 2010. – 34 (280). –W.18-28. 6. S.. V . – R: S. – 1972. – W. 274-275 *** R P'' ''' '' ' % ''' (, , % ' ) %'' '. *** In this article are given the data about the Siberian stone loach that dwell in the Kenderlik River for the refinement of the systematic status of the earlier not studied points of the area: the morph metrical characteristic and the biological rates such as length, weight, fatness and sex structure of the population.

H SY. . 1 (33). 2012 35

YJ :597 .. , .. =Q ľM=JI= (ABRAMIS BRAMA) QĨİ M=I= LĨJO - , F,

X ''' ', '' ' ''. X - ''', , % '' . F '' 2-14 '. T '' 7,8,9,10 (67%). X 2010 . 48,8-465,3 . X Z 2,0, P – 1,7. G – = ı Ŀ Ľ ĳ : (Schizothorax argentatus), (Noemacheilus barbatulus), Ŀ (Schizothorax argentatus pseudaksaiensis) Ľ (Perca schrenki). ı = Ŀ ı 28 ĳ [1]. Tı Ľ (Acipenser Nudiventris), F ! (Barbus brachycephalus), G ! (Phoxinus phoxinus poljakowi), L ! (Phoxinus phoxinus brachyurus) Ľ G (Perca schrenki) ı . Ľ ı ı 11- ĳ , ı Ľ . G ı Ľ ı , Ľ ı Ŀ. T Ľ ı . Gı ı , ı ! . Lı ı = Ŀ Ŀı !! Ľ . RQ Lļ ļJ=RQ M 2008 - 2010 = Ŀ Ŀ, ĳ . G! 117 , 10% Ŀı. G !ı Ľ ĳ [2]. R Ŀı Ľ «Excel» . M ļLQ= Lļ TQJ QQ X U- . , F ı ĳ . 1949 G – = , M Ŀ, K Ŀ, G . Ŀ [3]. = Ŀ ı Ŀ ĳ. Tı 45 [3]. Gı ( 1). Gı ı Ŀ 24,1-27 , ı 23 %, 19,2 % . F, ı 39,1-42 , ı 3 %, 1,9 % Ŀ. L ı 2,44 [4]. M ı 2-14 1 Ŀ. T 14 ı Ŀ , 1418 , 2 ı , 22,6 . X 20 Ŀ ĳ . G, Ŀ ı 12-15 [5]. F ı 2-14 . Tı 2 Ŀ 7, 8, 9, 10 , ı 67 % . F ı 13 (0,7 %) 14 (0,5 % ) . 1- = Ŀ'' ĩı ĵĩı ' W, % G V , VV WW VV WW VW 7,0-9 2 1 3 1,9 3 9,1-12 6 3 9,3 5,8 9 12,1- 15 4 2 6,2 3,8 6 15,1-18 3 1 4,6 1,9 4 18,1-21 10 5 15,4 9,6 15 21,1-24 1 6 1,5 11,5 7 24,1-27 15 10 23 19,2 25 27,1-30 8 9 12,3 17,3 17 30,1-33 6 4 9,3 7,7 10 33,1-36 5 6 7,7 11,5 11 36 Y . . 1 (33). 2012

36,1-39 3 4 4,6 7,7 7 39,1-42 2 1 3 1,9 3 N 65 52 100 100 117 W , ı ı 2008 - 2010 . (3 ). 2008 . 4 25%, 5 44,7% , 2009 . 5 . 24,2%, 6 . 25,5% . F, 2010 . 5 . 25%, 6 . 30,7% Ľ 7 . 23,9% , 1 2 . F ı Ŀ Ŀ . 2 = Ŀ ı Ŀ. Xı 2010 . 48,8 – 465,3 ı . 1600 14 0,5 1400 13 0,7

1200 12 2,3 11 3,8 1000 10 12

800 9 11,7 8 17,7 600 7 25,6

T ĩ, ĩ, T 400 L, L, 6 5,8 200 5 5,8 4 5,1 0 2 3 4 5 6 7 8 9 10 11 12 13 14 3 3,8 % L, 2 5,3

1-. = Ŀ'' ı ĩ 2-. = Ŀ'' ĩı ĭ ĭĵ. X !! ı Ŀ . R, 1971 . 2:1, 1981-1982 . 1,5:1 Ŀ, 1972 Ľ 1979 . 1,7:1, 1973 Ľ 1975 . 2,4:1 Ŀ [6]. F, 1:1 ( 3). X ı 117 ı 56 - (48%), 54 – (46%) Ľ 7 – (6%) . Xı 2010 ı 1 : 1,1 ı . 2- = Ŀ'' ĩı ĵĭĩ FLX(F ), ı M F T 2008 40,4-373,5 122,4± 2009 61,6-316,2 136,2± 2010 48,8-465,3 169,6± T ı – ı . = Ŀ ı ı - 2,0, - 1,7. L ı ı . P 3 ı ĭ ĭı Ŀ ĭ L W V 2008 1 1,2 2009 1,1 1 2010 1 1,1

30 2008 2009 % 2010 20

0 123456789 L,

3-. = Ŀ'' ĩı ĭ ĭĵı Ŀ ĭ

H SY. . 1 (33). 2012 37

TJ Ľ ı Ľ Ŀ. X !!ı , - Ŀ, Ľ . = Ŀ ĳ ı , , Ľ ĳ ! Ŀı ĳ . ļJ 1. ! – !! ! ! ! . TX]KX. T SV: VFMJKQ ! . 2009. – W. 28 - 29. 2. U .. V . R.: U! U. 1966. -376 . 3. R H.Q. : J. . . F – F, 1973. -404 . 4. \ .U. G! G: F. . . . . X, 1975. - 18 . 5. L . X. 4. R «U», 1983. – 502 . 6. G H.R., J W.. R // H . 1970. X. 10. H. 4. W. 608-621. *** N . , - , , . 2 14 . 67 % 7,8,9,10 . U 2010 . 48,8 - 465,3 . . Z 2,0, P -1,7. *** The investigation of temporary conditions of abramis brama is based on the biological analysis. In the result of the investigation it is revealed that changes in size, weight, age, sexual interrelations and reproduction of the population of abramis brama are not abserved. Age of caches fish were 2-14. There are fish which age were 4, 8, 9, 10 more tnan others. Fecundity of bream in 2010 amounted to thousands of eggs 48,8-465,3. The average fatness of fish for bream Fulton is 2.0, according to Clark 1.7

YJ 597.551.2:575.2 1I.. , 2.. R, 1.. R MR RTZTQTI] UTMQO UTIT I] TRIPLOPHYSA STRAUCHII M .Qb QR T RTR U 1PSY .-Z, F, P, 2Y RI, R, P

U Triplophysa strauchii (Kessler, 1874) 1986-1987 2000-2008 .. .Y F. F . N .Y F. Y ( Balitoridae, Cobitidae) H.U.R ! ! . V! «V », ! V. X H U ! G ! ! ! [1]. R !! ! ! ! ! ! ! ! ! ! . V, ! , !! ! , ! [2,3]. H , ! ! ! !! , ! ! !! ! . H ! !! ! !! «! !», . V! ! !! ! - ! [4,5]. !! ! ! [6]. X , !, ! !! , ! ! . H ! 1980- ! G . S .F ! 450 . [7]. V Y F !!! , .F . K ! 100 , 5 3/. W «W» 1971 , , , G F 1982-86 , ! . W 1990- , ! (!, , , ! ..). T ! [8]. 38 Y . . 1 (33). 2012

\ ! !! ! - ! Triplophysa strauchii (Kessler, 1874), !!. RQ RTJ QJTO V ! .Y F 1986-1987 2000 – 2008 .. .F, . G ! [9,10]. F ! 5 22 . U ! , , !!, !, ; , , , , ! !, ; , , , . H (I..G), ! [11]. ! 1987 ., (.H.R), . V ! (. 1). ] !! [12] b Ms=Mo(Ls/Lo) , Ms - ! ; Mo - ! ; Ls - ; Lo - !; b - ! ! lgMo lgLo ! , !! ! (Intercept) ! . W [13]. J! ! V ANOVA, 2 “ !” CD [14] “!” - d 1,2 [15]. U! ( , , GLM , ) [16,17], ! “NTSYSp” ! 2.02, “Statistica” ! 6.0 “SPSS” ! 18.0. MQb QJTO TLJ H ! ! .Y F, XXI , 1980- XX : ! 15 XXI . 9 . 1 , ! ! . R 2001 , ! 2008 1980- . W 2000-2003 , , , , . U GLM (. 2) ! ! . W ! ! , 1987 . H 2000-2008 . . U ! 1986 2000-2008 . (. 3). , ! ! [18]. S 1 , 1986 2000-2008 . !! 1-3 . T , 2000 2008 . , ! . c , . T 1986 2000 . ! !, ! ! . S 2. X 1 R . N M ±m min max M ±m min max M ±m min max L l_m Q 1986 15 94.05 4.24 70.20 138.30 80.57 3.63 59.00 117.50 13.34 2.53 2.31 39.80 1987 23 100.96 2.33 83.00 125.00 86.85 1.94 70.00 107.00 9.44 0.58 5.10 17.00 2000 3 96.33 6.57 87.00 109.00 79.33 5.24 71.00 89.00 7.09 2.00 4.71 11.06 2001 15 66.13 1.41 61.00 80.00 55.13 1.03 50.00 66.00 2.23 0.14 1.54 3.41 H SY. . 1 (33). 2012 39

2003 14 79.79 2.21 66.00 94.00 66.36 1.91 54.00 80.00 3.96 0.34 1.88 6.65 2006 52 90.75 2.55 51.00 124.00 75.52 2.19 42.00 103.00 7.43 0.53 1.42 16.76 2007 23 103.41 3.20 71.00 126.40 86.08 2.63 58.60 102.70 9.70 0.84 2.57 15.45 2008 37 112.69 2.32 71.50 140.00 93.42 1.89 62.00 114.60 14.17 0.81 6.56 24.10 q_m Fulton Clark 1986 15 10.06 2.45 2.36 30.00 2.18 0.16 1.01 2.86 1.61 0.13 0.78 1.90 1987 23 8.22 0.55 4.90 13.40 1.42 0.03 1.17 1.69 1.20 0.04 0.88 1.47 2000 3 5.31 1.57 3.36 8.41 1.35 0.12 1.16 1.57 1.00 0.10 0.88 1.19 2001 15 1.66 0.10 1.17 2.62 1.31 0.03 1.10 1.53 0.97 0.02 0.83 1.12 2003 14 3.06 0.25 1.50 5.12 1.31 0.04 1.07 1.53 1.02 0.03 0.81 1.20 2006 52 7.59 0.47 2.94 12.92 1.59 0.02 1.00 1.94 1.14 0.02 0.82 1.38 2007 23 8.11 0.70 2.12 12.88 1.42 0.04 0.80 1.73 1.14 0.02 0.93 1.35 2008 37 10.08 0.71 3.66 19.72 1.65 0.03 1.39 2.40 1.17 0.03 0.92 1.54 Ds As P 1986 15 7.60 0.47 7 8 5.30 0.53 5 7 15.80 0.68 15 17 1987 23 7.91 0.42 7 9 5.96 0.77 5 7 11.39 0.89 10 13 2000 3 7.00 1.00 6 8 5.00 0.00 5 5 13.67 0.58 13 14 2001 15 7.53 0.52 7 8 5.00 0.00 5 5 12.27 0.80 11 14 2003 14 7.71 0.47 7 8 5.00 0.39 4 6 13.00 1.41 11 15 2006 29 7.95 0.43 7 9 5.03 0.19 5 6 12.59 0.87 11 14 2007 21 8.05 0.22 8 9 5.29 0.46 5 6 12.19 1.63 8 15 2008 33 8.05 0.47 7 9 5.18 0.46 4 6 13.06 0.83 11 15 VVert N M ±m min max M ±m min max N 1986 15 7.73 0.46 7.0 8.0 40.90 2.18 37.00 44.00 10 1987 23 8.35 0.78 7.0 10.0 35.83 0.71 35.00 37.00 18 2000 3 7.00 0.00 7.0 7.0 35.33 0.58 35.00 36.00 3 2001 15 7.00 0.27 6.5 7.5 35.20 1.15 33.00 37.00 15 2003 14 7.50 0.52 7.0 8.0 35.86 1.61 33.00 39.00 14 2006 29 6.79 0.49 6.0 8.0 35.14 0.92 33.00 37.00 29 2007 21 7.10 0.44 6.0 8.0 35.38 1.16 34.00 38.00 21 2008 33 7.33 0.48 7.0 8.0 35.27 0.94 33.00 37.00 33 min, max, M, ±m – , ; N – !; L – ! , ; l_m – ! , ; Q – ! , ; q_m – , ; Fulton – ; Clark – ; Ds – ; As - ; P, V - ; Vert. – ; X 2 R Test Value F Effect df Error df V Intercept Wilks 0.039577 630.9529 5 130.0000 0.000000 l_m Wilks 0.964492 0.9572 5 130.0000 0.446660 I Wilks 0.087359 12.3221 35 549.2904 0.000000 X 3 . 2 U 1986 ., n=15 . 2000-2008 .., n=144 . d 1,2 CD P Ds 7 8 7.6 0.43 6 10 7.9 0.35 0.13 0.28 0.05 As 5 6.5 5.3 0.44 4 6 5.1 0.19 0.34 0.24 >0.05 P 15 17 15.8 0.53 8 16 12.6 0.96 8.68 1.68 0.001 V 7 8 7.7 0.39 6 8 7.1 0.34 0.22 0.66 0.001 Vert 37 44 40.9 1.72 33 39 35.3 0.87 8.57 1.70 0.001 T! 1. U ! “ !” 40 Y . . 1 (33). 2012

K 1986-1987 . . X ! . U! ! (Eugenvalue=8.39; Total variance=38.18%) ! 1986-1987 2000-2008 . R ! , , ( 51%). H! ! (Eugenvalue=3.22; Total variance=14.64%) ! 2000- 2008 . R ! (19.54%), ! (16.40%) (aD – 11.85%, pD – 9.29%). X! ! (Eugenvalue=1.90; Total variance=8.63%) ! 1986 1987 . R ! , !!, , !! ( 51%). J , ! (P00.05). U 87.2% ! ( , ! ! !! ! ). X ! 1987 ! 100%, 1986 – 93.3%, , 2007 . X ! ! 2000-2008 ! 80% 94%. ! 2003 , 57% (8 ) , 2000 (1 ), 2001 (1 ), 2006 (4 ) . Triplophysa strauchii from Ulken Almaty river count ed t rait s 79 112 72 143 131 113 88

107

5m 55 65 30 105 115 76 61 14m 14012311810129 81673 97 5 9 84 68 10m 9m 1179286 95281421331161037864631113813713413012282777366605810 135999480 20 15m11m7m 2m27 1590 87 13610070561212431129967562856971 6 139 10219 1272314413232 93 98 141120119 91 125 14 22 89 13m 12m 59

8m 25 3m 1m 1141 8 83 6m 4m 4 5718171316747 121 26 126 21 24

1. : 1986 . “m” ! (roots), ! ! . U ! (V00.005), , ! 1986-1987 2000-2008 . R , , , ! , , ! ! . - ! ! . X ! 1986-1987 100%, 2000-2008 90%, 7 179 . H SY. . 1 (33). 2012 41

2. V ! ! , ! .Y F. , ! ! , ! ! ! [19]. Z.W.V [20] , ! !.

3. ! - , , !, - . Y ! , , ! (fast-start response “C-start”). U . F ! ! [21-28]. U!! ! ! ! ! ! . V , , , !! [29-31]. H !, [32]. V , .F ! , ! , [33]. X , ! ! .Y F !!! ! !. Q 1. R H.U. W Cobitidae – H // V – F-F: S, 1989. – X.4. – W.5-69. 2. Q.F. V !! // R – Q. 1987. - W.71-73. 3. V Z.W., U T.F., W T.U. . ! ! - R.: S. - 1982. - 248 . 4. Schluter D. Ecological speciation in postglacial fishes// Philos. Trans. R.Soc. London – 1996. - Ser.B 351. - P.807-814. 5. Schluter D. Ecology and the origin of species// Trends in Ecology and Evolution. - 2001. - Vol.16. - No 7. - P.372-380. 6. Motta Ph.J., Norton S.E., Luczkovich J.J. Perspectives of ecomorphology of bony fishes// Environmental biology of fishes. - 1995. - V.44. - P.11-20. 42 Y . . 1 (33). 2012

7. W H.W. U ! U!// S- ! ! U!: R . 3 ! – F- F: S. - 1990. - W. 145-147. 8. F F.F. H F ! // R ! -G «G-2000»: X !. – F: - , V . - 2000. - W. 6-8. 9. U .. V . R.: U! . - 1966. - 376 . 10. Holcik J. General introduction to fishes. 2. Determination criteria // The freshwater Fishes of Europe.- Aula-Verlag Wiesbaden. - 1989. - Vol.1. Vart 2. - V. 38-58. 11. R R.H., Q G.F., R F.S. T ! ! ! , // H . – 2005. - X.45. - 33. - W. 331-341. 12. Elliott N.G., Haskard K., Kozlov J.A. Morphometric analysis of orange roughy (Hoplostethus atlanticus) off the continental slope of southern Australia //Journal of Fish Biology. - 1995. – V. 46 – V. 202-220. 13. Q I.. G! – R.: H! . - 1990. - 352 . 14. R c. U . - R.: R, 1971. - 454 c. 15. F H.Q., V Z.W. Coregonus lavaretus (L.) // H . 1977. - X.17. - H. 5. - W. 862-878. 16. Sneath P.H.A., Sokal R.R. Numerical Taxonomy – Freeman, San Francisco, - 1973. - 573 p. 17. Press W. H., Flannery B. P., Teukolsky S. A., Vetterling W. T. Numerical recipes – Cambridge, New York, 1986. - 818 p. 18. Webb P.W, LaLiberte G.D., Schrank A.J. Does body and fin form affect the maneuverability of fish traversing vertical and horizontal slits? // Environmental Biology of Fishes – 1996. – V.46. - Pp.7-14. 19. W .F., R H.F., I R.F. J Salmo mykiss () // H – 1988. - X.28. - H.2. - W. 213-221. 20. V Z.W. c! – R.: S, - 1980. 301 . 21. Blake R.W. Fish Locomotion. - Cambridge University Press, Cambridge. - 1983. 22. Domenici P., Blake R.W. The kinematics and performance of fish fast-start swimming// Journal of Experimental Biology - 1997. – 200. - P.1165-1178. 23. Harper D.C., Blake R.W. Fast-start performance of rainbow trout Salmo gairdneri and northern pike Esox lucius. // Journal of Experimental Biology - 1990. – 150. - P.321-342. 24. Langerhans R.B. Predictability of phenotypic differentiation across flow regimes in fishes. // Integrative and Comparative Biology – 2008. – 48. – P. 750-768. 25. Langerhans R.B. Trade-off between steady and unsteady swimming underlies predator-driven divergence in Gambusia affinis. // Journal of Evolutionary Biology - 2009. doi:10.1111/j.1420-9101.2009.01716.x. 26. Langerhans R.B., Layman C.A., Shokrollahi A.M. & DeWitt T.J. Predator-driven phenotypic diversification in Gambusia affinis // Evolution - 2004. - 58: - P. 2305-2318. 27. Webb P.W. Avoidance responses of fathead minnow to strikes by four teleost predators// Journal of Comparative Physiology - 1982. - 147A. – P. 371-378. 28. Webb P.W. Body form, locomotion, and foraging in aquatic vertebrates// American Zoologist - 1984. – 24 – P. 107-120. 29. Q ! F.., J K.F. H! , // H – 1961. – X.1. – H.3 (20). – W.497-509. 30. X .. H! // H - 1968. – X.8. - H.3(50). – W.425- 439. 31. Levin B.A. Drastic shift in the number of lateral line scales in the common roach Rutilis rutilus as a result of heterochronies: experimental data// Journal of applied ichthyology - 2010. - V.26. - Pp.303-306. 32. Balon E.K., Leim K.F. Prelude to ecomorphology of fishes// Environmental biology of fishes. - 1995. - V.44. - P. 7-8. 33. R S.^., G I.., I.W., .. W ! ! G .// G, I F! : !, , : R 2 . 20-24 !! 2010 . I-F – I-F: VT ITKHUT «I- F , - 2010. - W.63-67. *** F '' 1986-1987 % 2000-2008 . ' Triplophysa strauchii (Kessler, 1874) ''' '. U % ''' ' '''. *** A comparative analysis of the state of morphometrical characters of the spotted stone loach Triplophysa strauchii (Kessler, 1874) was made with samples caught in 1986-1987 and 2000-2008 in the Ulken Almaty river. Multivariate analyses show big differences in the investigated plastic and counted characters between fishes caught in the XX-th and XXI-st centuries. The revealed change in the fishes profile with time is a result of negative human impact to the ecosystem of the Ulken Almaty river.

TL 597 .. ĭ, R.L. U UĨO TOR Lļ =Q ľM=JI= ľ QĨİ M=I= LĨJO Lļ Tİ T =RJ= UOJQ LTQJ « - » L^W E-mail: [email protected]

' . . M ' % '' ''. T 10-15 Ŀ , Ľ, Ŀ . T , !! Ľ ı ı ĳı Ľ . F ı Ŀ ı . H SY. . 1 (33). 2012 43

Ŀ , ! ĳ . G ĳ Ŀ K . W ı . Ŀ , Ŀ , 3% . W Ľ ı ĳ . M J R ļJ=RQ= M 2008-2011 . . Ľ ! Ľ = Ŀ ! . Ŀ ! Ľ Ŀ , Ľ Ŀ ĳ Ŀ 18 100 . F , Ŀ Ľ Ľ . L Ľ ı Ľ ı ! . X! 6 , 1, ı Ŀ 3 , ı Ŀ 5 ĳ 10 ĳ . F ı , Ŀ , Ŀ ! ! Ŀı [1]. M ļLQ= Lļ TQJ U Ŀ (Stizostedion lucioperca) G-= 1957-1959 . . = Ŀ Ŀ IcW Ŀ !!ı ĳ Ŀı Ŀ 1270 . Ŀ ı . ı Ŀ ı . Kı ! Ŀ ı . Y ı ı ı , !!ı Ŀ ı [2]. Ŀ . Kı 1975 . T 1990 Ŀ. 2005-2007 . ĳ . Wı Ŀ ı ı 60-80 ( 1). F,

220 200 180 160 140 120 100 80 60 40 20 0 1973 1975 1977 1979 1981 1983 1985 1987 1989 1991 1993 1995 1997 1999 2001 2003 2005 2007 2009 2011* L – ĩ ĭ Ŀ' ı Ŀĭ (2011*- 10 ĭ Ŀ') Ŀ !!ı – 25,0 40,0 4-5 . K, Ľ 8-9 ĳ , , 3 Ŀ [3]. Ŀı 4 5-6 ĳ. Y , ı 3- ĳ Ľ ı !. Ŀı ! , ı ı . S Ŀı . K Ŀ () Ľ , = Ŀ ! , ! . G ı Ŀ, ! . Ŀı Ľ 39,2- 427,3 ı , 156,2 ı , Ŀ Ŀ Ŀ (1-).

44 Y . . 1 (33). 2012

1- 2010-2011 . Ŀ'ı ' ĵĭĩ I , 1 . FLX, ı L N - - - T 2010 . 4 2 25-30 27,5 1520-2684 2102 45,6-67,1 56,3 6 1 78 78 1586 1586 12,4 12,4 7 5 37-177 105 1250-1627 1449,8 46,2-245,1 153,1 8 1 246 246 1760 1760 432,9 432,9 9 1 225 225 1587 1587 404,7 404,7 G 10 - 116 - 1639 - 184,1 2011 . 4 2 26-30 28 1307-1522 1415 39,2-39,6 39,4 5 1 28 28 1453 1453 40,6 40,6 6 6 37-123 91 1250-1627 1449,8 62,4-245,1 103,9 7 3 54-188 139 1207-1536 1431 50,2-288,7 204,5 9 2 247-434 340 1320-1634 1477 403,4-427,3 415,4 G 14 - 124 - 1639 - 156,2

Ŀı ı . R, 2008 Ŀ ĳ ı 1:1,26- ı . 2009 . ı 1:1,8 , 2010-2011 . Lı 1:1,2 Ŀ . Ŀı Ŀ !, ı. G ı Ľ ! Ŀ 15 - (2-). 2- ĩ ĭ ĩ Ŀ'ı ĵĭ Ŀ' (' , ), Ŀ 2011. l l l I! L 1 2 3 M±m T M±m M±m T 2010 1+ 12-12,2 12,1 14,5-15 14,8 - - 2009 2+ 8,4-12,5 10,4 15,8-20,5 18,1 19-22 20 2008 3+ 7,2-13,2 10,3 13,2-20 17,0 18,8-26,5 23,5 2007 4+ 8,2-12-8 9,4 13,3-20,4 16,3 19,2-27,3 22,3 2006 5+ 7,6-12,5 10,3 13,3-20,8 17,4 18,9-30 23,6 2004 7+ 10,4 10,4 17,4 17,4 24,3 24,3 2004-2010 R 7,2-13,2 10,0 13,2-20,8 16,9 18,8-30,0 23,0 T 10,0 10,0 6,9 6,9 6,1 6,1 l l l l l 4 5 6 7 8 N M±m M±m M±m M±m ------2 ------4 20-29,9 25,6 ------23 23,6-34,5 27,6 26-36 29 - - - - - 28 24,2-37,5 29,8 30,3-45 35,9 32,4-46,3 37,2 - - - 17 30,0 30,0 36,0 36,0 42,2 42,2 48,5 48,5 49,2 1 23,6-37,5 28,5 30,3-45 35,9 42,2 42,2 48,5 48,5 49,2 75 5,5 5,5 7,4 7,4 6,3 6,3 6,3 6,3 0,7 75 Ŀ ı ı , 5+,7+ Ŀ. Ŀı ! Ŀı Ŀ , !, ı Ŀ Ŀ (3-). W-, Ŀı (3,5-4 ) ĳ (, , Ŀıı) ı . Rı ĳ !!ı Ľ ĳ Ľ !! ı ĳ . H SY. . 1 (33). 2012 45

3- = Ŀ' ĩ ĭ ĩ Ŀ'ı Ŀ ĭ ĭ Ŀ'' ( Ŀ') L l, Q, q, ı. ı. FLX, ı N - - 2008 32,2 510,0 482,0 1,1 1,0 - 132 2009 35,3 577,2 546,0 1,1 1,0 - 153 2010 33,1 545,0 483,1 1,2 1,0 184,1 226 2011 31,7 484,0 449,0 1,2 1,1 147,7 465 L, Ŀ ĳı . Ĳ Ľ ı ı Ľ - Ŀ . W Ľ ı , Ŀ ĳ Ľ . X , . ı Ŀ !! . W, Ŀ !! ĳ Ľ ı ĳ . W-, Ľ ı . , ı 1 15 , Ŀ ı ! ı 05 Ľ 20 . W Ŀ Ľ Ŀ Ľ ĳ , - Ŀ . ļJ 1. U .. V . – R.: U! , 1966. 2. G-FĿ Ľ ı ı Ľ ı Ŀ , ı Ŀ (VKLGFR) ! Ľ Ľ Ŀ 2013 . GĿ: : ĨML /G^ĨM. –F, 2011.- 56 . 3. G-FĿ Ľ ı ı Ľ ı Ŀ , ı Ŀ (VKLGFR) ! Ľ Ľ Ŀ 2012 . GĿ: : ĨML /G^ĨM. –F, 2010.- 54 . *** , . [ , P . X . *** In article results research of a pike perch which for today is required with great demand in market economy are yielded. The current state and rational uses, and also ways of preservation of a pike perch to the Kapshagai reservoir is characterized. Also recommendations about change of term of an interdiction to fishes are made during spawning.

YJ 597 .I. , .. T, .. Q ľQ=JI= ľ (STIZOSTEDION LUCIOPERCA) İ ITQTIQ ľ== - , -mail: [email protected]

L %' G ''' '' ' ' ' '. P' '' '' . T % ' '' ' . G ' ' . Ľ Ŀ ! . W Ŀ Ľ Ŀ . W Ľ Ŀ ı ĳ . Ŀ . ı T-, K , G-= Ľ FĿ Ŀ ĳ ı . G G Ŀ Ŀ 13 [1]. Ŀ . T Ŀı Ľ . T Ľ ı Ŀ Ŀ ĳ. Ľ Ŀ . K Ŀı , Ŀ !! Ľ ĳ. 46 Y . . 1 (33). 2012

W ı Ŀ ! ı ĳ ĳ. 2002 G Ŀ ı . 2002 G Ľ VI-! Ŀ Ŀ, Ŀı Ŀ Ŀ ı Ľ ĳ, ! , Q, F!Ŀ Ŀ ı ! Ľ . VI Ľ VII ! ı ! Ŀ, , ! (^^) Ľ . G Ľ ! ı Ŀ 2001 Ŀ 2 Ŀ, ^ G ı Ľ (^^)- ĳ. I! ĳ G Ŀı Ŀ ı Ŀ Ŀ, ı ^ G , , Ľ .. ı [2]. T ! ĳ Ľ . T ı ı : Ŀ ı ! Ŀ . G ı ı ĳ . G Ľ , ı ĳ Ŀ, Ŀ . Gı ! , ı Ľ Ŀ . G ı ! ĳ ı Ŀ . F Ľ ĳ Gı ı Ŀ !ı . T Ľ Ľ , ! ! [3]. RQJ R ļJ=RQ M 2011 Ŀ G Ŀı 6 .Ŀ ı . R 4%- ! ĳ . R Ŀı Ľĳ Ľ . F Ŀ ı Ŀ ĳ. W Ľ !. J ĳ , ı 5-7 , ! Ľ ĳ. G - Ľ R ! !. I – !, , Ľ ĳ. I , ! Ŀ [4]. M ļLQ= R QQQ Gı, ! Ľ !ı Ŀ () . S. I Ŀ ĳ ĳ Ľ , Ľ Ŀ . G Ŀ Ľ [5]. Ŀ ! Ľ : Ŀ ı ĳ , Ľ ĳ Ŀ . Xĳ Ŀ Ŀ, ı . Xĳı ı . Iı , ı ! ı Ľ ! ! Ŀ. M Ŀı ı : Ľ ĳı ĳ( 1).

1- Ŀ . ı ĭ, ĳı' ĳ' '. . Ĳ 400 F ĳ ĳ , ! , ĳ . F ĳ !ı ! Ľ . F ĳı ĳ H SY. . 1 (33). 2012 47

ı ! , ! . F ! ĳı ĳ ı ! ĳ . ĳı ĳ ı . ı ĳı Ŀ Ľ . ĳ ! ! . \ Ľ Ľ Ľ Ŀ (! Ŀ, II). G ĳ ĳ Ľ . I ĳ Ľ ! . ı ı . ĳı Ľ ı Ŀ Ŀ, Ŀ Ľ ı [6].

2-Ŀ . U ĳ '' Ľ' Ŀ' ĳ'. . Ĳ 400 Ŀı . G !ı Ŀ Ľ ! ĳ ! . ^ Ľ ! ! ( 2).

3. Ŀ . Rĭ Ŀ'' ĭ . ' . R. Ĳ 400 Ĳ ı ı , Ľ - . R ĳ ı ! ı . . . R Ŀ . ľ ( 3). . W ı. L ı ĳ , . F ĳ ı ĳ . L Ľ Ŀ Ŀı ı ĳ. W ĳ ! ı ĳ . I !ı , ı ! Ŀ . Gı ĳ , ı. F Ŀ ! Ŀ . F Ľ ĳı ĳ , Ŀ ı Ŀ . V! ! , ! !ı Ŀ Ľ ı . T ! ı ı Ŀ Ŀ ĳ. T 48 Y . . 1 (33). 2012

Ŀ ! Ŀ ı . ļJ 1. S Z.J. G! ! F !. F.Q; I STV; 1973.-21 . 2. U Q.U., ^ J.W., Q X.. I . G . // V! ! V : ! !. W . – F: G, 2005. – 367-370 . 3. .. R // H. ,1968.X.8. H.12 4. G. V. R! .1954. 5. S.S. I. V! . //F , . Q. 1962. -77-80 . 6. L .R. R! (Bufo viridis) ! (Rana ridibunda) . –J......-V. F., 2008. -35-37 . *** , . G, . , . Y , . *** Reclaiming processes were observed in the liver of the pike perch were caught in the western part of lake Balkhash. Kernels of hepatocytes were in a division stage that is characteristic for liver of the pike perch were caught in the spring period. In some hepatocytes kernels were big polyploid. Regeneration process was active at all fishes.

YJ 597 . . J, . . TZ UU TTMOTIT UTQbMT RQ IT TJTRT QTQbTIT OT ( UR TJTQ_ LT TM LQTQb) XTT « - !»

I F F , , , . . N – RQ RTJ QJTO R ! ! ! 2011 . F . W !! ! ! 16 80 ! !. F !! ! [1,2.] S! !! "V ". (U ! ! fishbase.org) MQb QJTO TLJ H F 1,5-2 . . H !! 1134 !. U 90% . W ! . T! ! 6 , !! – 1,6 , ! – 3 . H " , !! : [3], " 90- ! . ^ ( 1). X 1 ", , 2011 .

S . L /. F

W. - Carassius auratus auratus + + F - Pseudorsbora parva + - W. T" G – Perca schrenki + - Y ! 2,02 /. H F ! 123 . !. H 1 4 . H 1+ -2+ , ! ! 95%, 8,8 10,3 23 37 ( 2). U - 21,0 304 . W 1:5,25. Y 3,2 3,4,, , 3,3 , .

H SY. . 1 (33). 2012 49

X 2 T , 2011 . H J, R, Y -, . % - !! - !! - !! 1 8,0-9,5 8,8 18-25 23 2,4-3,9 3,4 58 47,2 2 9,0-13,0 10,3 20-70 37 2,7-3,8 3,3 59 48,0 3 16,0-17,5 16,8 136-162 150 3,0-3,3 3,2 3 2,4 4 20,0-21,0 20,6 254-304 284 3,2-3,3 3,2 3 2,4 8,0-21,0 10,0 18-304 39 2,4-3,9 3,3 123 100 U ! ! ! ! ! . U ! ! 0,57 ./3 ! ! ( 3). X 3 , ./3 H H N ! !, ./3 \. F 24 0,57 . L 7 0,17 F . L 25 0,60 T L ! 10 - X, 1085 !. Z ! ! . T , . T , ! ! 22 ., 10,7 . T ! . H 3 – , [3,4] ( 1). H 18 . ! 49 . !. Y ! 0,16 1,02 /, 0,60 /. G ! ! 4. H ! ! ! 2- 7- . 3-4 – 65,3%. U - ! 10,5 23,0 16 176 . 1,3 1,7, 1,4. X 4 T . L, 2011 . H J, R, Y -, . % - !! - !! - !! T 2 10,5-11,5 11,1 16-26 19 1,1-1,7 1,3 9 18,4 3 11,5-14,5 12,4 20-38 27 1,0-1,9 1,4 18 36,7 4 12,0-17,5 15,0 24-90 53 1,2-1,8 1,4 14 28,6 5 18,0-22,0 19,6 106-170 136 1,5-2,0 1,7 6 12,2 6 19,5 19,5 124 124 1,7 1,7 1 2,0 7 23,0 23,0 176 176 1,4 1,4 1 2,0 10,5-23,0 14,2 16-176 51 1,0-2,0 1,4 49 100 2 11,5 11,5 44 44 2,9 2,9 1 5,6 3 12,0-15,5 14,3 48-100 83 2,2-3,3 2,7 16 88,9 4 14,0 14,0 86 86 3,1 3,1 1 5,6 11,5-15,5 14,1 44-100 81 2,2-3,3 2,8 18 100,0 T ! 2+-4+ . U! ! 15,5 100 , ! 11,5 44 , . ! 2,2 – 3,3, – 2,8 ( 4). H ! ! 0,17 ./3 ", ! 0,60 ./3. U , -, ". 50 Y . . 1 (33). 2012

V !! 5. X 5 T " U T L H F U, 29,5 6 W. , 10,7 1,6 V ! ! , S !! !, ! , H 0+, 0+, , . 15-20 15-20 H, % 70 70 U ./ . 150 50 H ", . 4425 900100 V , ! ! . T ! ! . X F, !. I . L, ! ! . Q 1. U .. V . – R: U! 1966. 376 . 2. ! F.. T . - R, Q! ! . 1981. 208 . 3. G F.F., R H.F., R H.U. / "V " .3. F-F, 1988. - 128 . 4. G F.F., R H.F., R H.U. / "V " .4 F-F. 1989. - 103 . *** G F' ' , '' ''' % ''. '- . *** The fish fauna structure, the main biological indicators and productivity of young fishes in high-mountainous reservoirs of Alakol`s region is resulted. The actions on fish economy using are recommended.

YJ 576.8 .ļ. JĽ', .. == Q İ=M= QJ MJ= LQM UM= - . F, , [email protected]

G'' 2010 ' P'' F '' ^ X ' . M ' 175 (% ' 30 , 25 ) . T: , , , , ', % , M %' ' % ' ' ', 13 ' . '' ' ' D. extensus ' 73,3 % . G ĳ ı ı ı . L ı ı , . T , , Ŀ Ŀ. Ĩ ĳ , Ľ ı , , ı ĳ . Xĳ ı Ľ ı . T ĳ Ŀ- . F ı Ľĳ ! ı Ľ ! !ı Ŀ ĳı Ľ !. F ĳ Ŀ : , Ŀ Ŀ, ı Ŀ. G , ı ı Ŀ Ľ . Gı ı ! ! ! ı . Gı Ŀ. L Ľ . H SY. . 1 (33). 2012 51

L , !, ı, Ľ . L ı , ı ĳ Ŀ Ľ (ı ı Ŀ, ı Ŀ Ŀ, () ı Ľ .., ı , , ı Ľ .. Ľ [1]. F ı ! Ŀ ı 30 HSTV (ISTV) ! H.F. J Ľ G « F !» ı Ŀ [2]. 1951 1970 F!ı F ı ! ı FS Y WWV ı , Ľ W.T Tı «U Y» [3] ! Ŀ. W.T. T, V. X Ľ ı F ı ı 12 ĳ . T: Dactylogyrus auriculatus (Nordmann,1832,) D. alatus Linstow, 1878, D. solidus Achmerow, 1948, D. zandti Bychowsky, 1933, D. vastator Nybelin, 1924, D. minor Wagener, 1857, D. parvus Wagener, 1909, D. capoetobramae Kusmenko, 1945, Dactylogyrus sp., Pseudacolpenteron pavlovski Bychowsky et Gussev, 1955, Gyrodactylus parvicopula Bychowsky, 1933, Gyrodactylus sp. 1970 F ıı . 1936 (J Ľ Q, 1937) Ľ 1953, 1961 . . T ı Nitzschia sturionis 100% - , , Ŀ- [6]. T F ıı ! Ľ Ľ Ľ ĳ ı - Ľ ĳ ĳ Ŀ Ŀ. M J Lļ ļJ== 2010 ı Ŀ F ıı 3 Ŀ, ^ Ľ Xĳ Ľ ı , , , , , , Ŀ, Ľ , . ı ĳ 30 25 . R . K. G-U Ľ [4], , ı ĳ «T WWWV» [5] ı . M ļLQ= Lļ TQJ QQ M Ľ ı Ľ ĳ, 13 ĳ (1-).

1- '' ı'' ' Ľ'' ı ĭ ' ' U Q! Dactylogyrus falcatus X D. tuba F D. wunderi X /-/ D. zandti X D. crucifer X D. sphyrna X D. rarissimus X /-/ D. extensus W D. minutus W /-/ Ancyrocephalus paradoxus Ŀ /-/ Gyrodactylus medius W /-/ Dipozoon paradoxum , Paradiplozoon pavlovskii F /-/

1- Ŀ, Ľ ı 6 ı ı 13 ĳ , ı : Dactylogyrus 9 ĳ; Ancyrocephalus – 1 ĳ; Gyrodactylus – 1; Dipozoon – 1; Paradiplozoon - 1. R ı Ŀ , Ľ ı 4 ĳ , - 3, - 2, Ŀ - 1. 52 Y . . 1 (33). 2012

2- '' ı'' ĭ ''ı ĭ Ŀ'' M U c (%) () Q! Dactylogyrus wunderi 70,0 8-44 D. falcatus 63,3 4-150 -//- D. zandti 23,3 2-50 -//- Dipozoon paradoxum 46,6 2-14 -//- D. extensus 73,3 10-64 -//- D. minutus 56,6 2-30 -//- Gyrodactylus medius 30,0 4-20 -//- D. crucifer 50,0 4-50 -//- D. rarissimus 46,6 8-30 -//- D. sphyrna 10,0 2-10 -//- Dipozoon paradoxum 6,6 4 -//- Ŀ Ancyrocephalus paradoxus 33,3 2-16 -//- Gı ! Ŀ Ľ ĳ ı Ľ ı -ı !! ĳ. RĽ D. extensus 73,3%, D. wunderi 70%, D. crucifer 50%, , Ŀ D. tuba Ľ A. paradoxus c Ŀ 20,0 33,3%. R ı ı Dipozoon paradoxum c - 46,6%, – 2-14, c - 6,6%, – 4 . F ı – Ŀ . (2-). M ı Ŀ , ! ! ĳı Ŀ Ŀ. M 30 D. extensus 10-64 , D. wunderi 8-44 , D. crucifer 4-50 . T Ŀ ı ĳ Ŀ. Ĵ F ıı ı . W ı ! ĳ - Ŀ Ŀ. ļJ 1. I S. F., W Z. F., H H. S., I U. U., K K. G., Z Q. S. !. - R.: R, 2003. 2. J H.F., G G.K. F !. // U. . MS FSWWWV. R. – Q, . FS WWWV, - 1934. – X. IY. - W.163-275. 3. T W.T. U Y. – X.: . FS Y WWV, - 1971. - 580 . 4. G! – U! .K. U ( ). - Q.: S, 1985. - 117 . 5. T WWWV /T. . F.H. I. Q.: S, 1985. - X. II. - 424 . 6. T W.T. H F !. – «» Y WWV, 1975. – W.107-109. *** S 2010 . F ^ X . 175 ., ( 30 ., – 25 .). Y : , , , , 13 . N 13 D. extensus, 73,3 %. *** Our studies in the spring and summer of 2010 the Aral Sea ihtioparazitologichesky material collected from the point of Shagalaly Tastubek up around the perimeter. A total of 175 specimens taken a survey., Fish (five species to 30 copies. And chub - 25 copies.). Six species of fish: flounder, bream, carp, chub, perch and roach were found 13 species of monogenetic trematodes. Of the 13 species of parasites, a relatively high degree of invasion observed in carp D. extensus, extent of infestation 73.3%.

YJ 576 .ļ. JĽ', .. J ľM=JI= Qİ \TJ UM=R MRJQ - ĴY. F,

'' ' %'' %'' ' '. '' – Cyprinius carpio, – Abramis brama, – Rutilus rutilus, – Aspius aspius . U 6 ' , Khawia H SY. . 1 (33). 2012 53 sinensis, Ligula intestinalis, Bothriocephalus gowkongensis, Proteocephalus torulosus, Caryophyllaeus fimbriceps, Paradilepis scolecina. G '' ' ' '' . G '' ' ' , . I – ı , , , , ı . ı Ŀ, ı ı Ŀ, ı , ı Ŀ Ľ !! Ľ ĳ [1]. Wı ı ĳ , ! . T ĳ ĳ, ı ı , !, !, , ! . Gı ı Ŀ . Wı W! Ŀ . W! Ŀ T F! ı Ŀı . T F ı Ŀ. ľ ^ ľ X!-^ Ŀı: S ! Ŀı . W! Ŀ Ŀ ı, ! , , ľ Ľ XĽ ı Ŀ ı [2]. W! Ŀ ı F F., J H.F., G G.K., T W.T. Ľ . [3-6]. M RQ Lļ ļJ== 2011 ı ĳ ı W! Ŀ Ľ Ľ Ľ Ŀ, Ŀ - Ľ Ľ ĳ 15 , ! – Cyprinius carpio, – Abramis brama, – Rutilus rutilus, – Aspius aspius . ! ! Ľ ĳ [7]. ļLQ= Lļ TQJ QQ M ı 6 ĳ : Caryophyllaeus fimbriceps, Ligula intestinalis, Khawia sinensis, Bothriocephalus gowkongensis, Proteocephalus torulosus, Paradilepis scolecina (1-). 1- ĵĭ ĭı ı ' ĭ 3 G Caryoph Ligula Khawia Bothriocep Proteoceph Paradilep W M ĳ yllaeus intestinali sinensis halus alus is fimbrice s gowkongen torulosus scolecina ps sis 1 W 2 8 4 - - 2 16 50% (Cyprinius carpio) 2 X 1 - - 4 3 - 8 25% (Abramis brama) 3 X - 7 - - - - 7 21,9% (Rutilus rutilus) 4 F 1 - - - - - 1 3,2% (Aspius aspius) G 4 15 4 4 3 2 32 100

1. ĵĭ ĭı ' ĭ 54 Y . . 1 (33). 2012

1– Ŀ Caryophyllaeus fimbriceps W! Ŀ – 4 : – 2, – 1, – 1 . S Caryophyllaeus fimbriceps Ŀ . X . Caryophyllaeus fimbriceps ı . G ĳ . 6 ! . Gı Ľ, ı . R Ŀ. Ligula intestinalis ı – 15 . T, – 8, – 7 . S ı ı , . Ligula intestinalis ı . K , ĳ . Khawia sinensis – 4 . T . Khawia sinensis ı . J ĳ . X ĳ , ! . Bothriocephalus gowkongensis – 4 . Wı . . Bothriocephalus gowkongensis ı . G ĳ Ľ . Jı ı Ľ ı ! Ŀ . G ĳ Ľ. Proteocephalus torulosus – 3 . X . . Proteocephalus torulosus ı . G 4 , ı ĳ . Paradilepis scolecina – 2 . X . . G 6 ĳ 32 , ı – 16 , 50%. Caryophyllaeus fimbriceps – 2, Ligula intestinalis – 8, Khawia sinensis – 4, Paradilepis scolecina – 2. Kı Ŀ Ligula intestinalis. X ı – 8 , 25%. Caryophyllaeus fimbriceps – 1, Bothriocephalus gowkongensis – 4, Proteocephalus torulosus – 3 . Kı Ŀ Bothriocephalus gowkongensis. X ı – 7 , 21,9%. X 7 Ligula intestinalis . F ı – 1 , 3,2%. X 1 Caryophyllaeus fimbriceps . M Ľ 1 – Ŀ ı ı Ŀ , Ŀ – 50%. X ı Ŀ – 25% , ı Ŀ – 21,9%, ı , Ŀ – 3,2%. ļJ 1. H.. H!, Q.R. R, F.H. U! G . – R.: « U! », 1979. – W 30 2. ! .VIII 137-138. 3. F F.. U ! . W-J. X - . FS WWV, XVI. F-F, 1962. 4. J H. G G. F !. U!. MS FS WWWV. . IV. Q., 1934. 5. T W.T. Y. «H FS YWWV», 1963, 34. 6. T W.T. W-J. «G ! W F », F-F: «S», 1966. 7. R F.U. R ! . – , 1950. – W 9-14 *** N -J . G – Cyprinius carpio, – Abramis brama, – Rutilus rutilus – Aspius aspius. 6 Cestoda. Khawia sinensis, Ligula intestinalis, Bothriocephalus gowkongensis, Proteocephalus torulosus, Caryophyllaeus fimbriceps, Paradilepis scolecina. N , . *** Some commercial fishes like carp – Cyprinius carpio, bream – Abramis brama, roach – Rutilus rutilus and asp – Aspius aspius bellowed to Cyprinidae family were investigated in the Syrdarya River. 6 species of parasites bellowed to Cestoda were observed: Khawia sinensis, Ligula intestinalis, Bothriocephalus gowkongensis, Proteocephalus torulosus, Caryophyllaeus fimbriceps, Paradilepis scolecina. Carp has had the biggest number of parasites and asp has had the lowest one.

YJ 597.5 I. R. J TU UR TR ITO RTU JQ TO I UR TZ Q-QTQbTIT O S VIU SY . -, e-mail: [email protected]

J , RTU . , G-F . H SY. . 1 (33). 2012 55

H ! , !! !, RWTU 1960- !!. S ! 1980- ! ! ! . H , ! . S, 2000 . W RWTU ! H 3.1 [1] , ! !! 2001 ., , ! " !. U H 3.1 : - – Extinct (EX) - – Extinct in the Wild (EW) - U ! – Critically Endangered (CR) - S!! – Endangered (EN) - Y! () – Vulnerable (VU) CR, EN, VU ! (Threatened) - G ! – Near Threatened (NT) - H – Least Concern (LC) - S – Data Deficient (DD) - S – Not Evaluated (NE). T H 3.1 !!! , ! (CR, EN, VU). J! ! : F – ! ! , H - ! (H1) (H2), , , !, W – ! , , D – ! , ( 50 . – CR, 250 . – EN, 1000 . – VU) K – , ! ! , ! . U !!, ! . H, , ! ! V [2] V [3] . F V Y [4] ! , , . W ! . X, , , , ! - , , - ! [5, 6]. W . X, ! V RWTU [7]. G , ! . T - ! . U ! !, , F H, !, ! RWTU. H , F1 – ! – : « , ! , ! ». F1d – – : « ». H – ! ! – : « 10 80% - ! CR, 10 50% - ! EN, 10 20% - ! VU». C, D, E . U ! ! ! . 56 Y . . 1 (33). 2012

! ! . H ! ! (TJY). S . J ! (! (NE) (! DD) - ! ! ! !. H G-F ! 40 , [8, 9]. H V (V) 4 : , ( ), ! ( «» ) (-! !!). , 6 F [10]: ! , W , - (.1). H V F (FT) ! (1 - , 2- , 3-, 4-) ! . X FT ! H ! . T! !, , ( ! LC). X 1. -

33 H U- R- - / 1. Acipenser nudiventris Lovetsky - + 1 V 2. Parasalmo mykiss (Walbaum) - + 3. Abbottina rivularis (Basilewsky) – + 4. Abramis brama (Linnaeus) – + 5. Aristichthys nobilis (Rich.) – + 6. Aspius aspius (Linnaeus) – + 7. Barbus brachycephalus Kessl.- + 2 V 8. Carassius auratus (L.) – - + 9. Ctenopharygodon idella (Val.) – + 10. Cyprinus carpio Linnaeus – + 11. Diptychus dybowskii Kessler – + 12. Diptychus maculates Steind. – + 13. Hemiculter leucisculus (Bas.) – + 14. Hypophthalmichthys molitrix (Val.)- + 15. Leuciscus leuciscus (Linnaeus) – + 16. Megalobrama terminalis (Rich.)- + 17. Phoxinus phoxinus (L.) – ! + 18. Ph. Brachyurus Berg – ! + 3 FT 19. Lagowskiella poljakowi (Kessl.)-! + 3 FT 20. Pseudorasbora parva (Tem. t Schl.) - + 21. Rhodeus sericeus (Pallas) – + 22. Rutilus rutilus (Linnaeus) – + 23. Schizothorax argentatus Kessl.- ! ) Sch. arg. argentatus Kessl.- / ! + ??FT ) Sch. arg. pseudaksaiensis Herz.-/ ! + 1 FT «» + 1 V 24. Tinca tinca (Linnaeus) – + 25. Ictiobus cyprinellus (Val.) – + 26. Nemacheilus sewerzowi G. Nik. – W + 2 FT 27. Triplophyza stolizkai (Steind.)- + 28. Tr. dorsalis (Kessler) – + 29. Tr. strauchi (Kessler) – ! + 30. Tr. labiata (Kessler) – + 4 FT 31. Misgurnus mochoity (Dyb.)- + 32. Silurus glanis Linnaeus – + H SY. . 1 (33). 2012 57

33. Oryzias latipes (Tem. et Schl.) – + 34. Gambusia affinis (Baird et Gir.) – ! + 35. Perca schrenki Kessler – (G-! !!) + 2 V 36. Sander lucioperca (Linnaeus) – + 37. Sander volgensis (Gmelin) – + 38. Micropercops cinctus (Dabry de Th.)- + 39. Rhinogobius cheni (Nichols)- + 40. Channa argus (Cantor) – + S ! ! , G-F ! RWTU H 3.1 ! . – EN: A2acde; B1b (i,iii, v). U! ( F2) – S , , ! 50% 10 3 , , , !: – ! , – , ! / , d – ! , – ! , , , , . I ( H1) – T ! ! , 5 000 ., ! !: – ! ! 5 , b – , , : (i) – !, (iii) – , ! / , (v) – . – EN: A2acde; B1ab (i, iii, v); C 2b. U! ( F2) ( H1) ! . I – VU: A2 ace; B1b (i, iv). U! ( F2) - S , , ! 30% 10 3 , , !: – ! , – ..., – ! ... I ( H1) – T ! ! , 20 000 ., !: – ! !, b – , , : (i) – !, (iv) – !. I – VU: A2 ace; B1 ab (i, iv). V , !. R U ( ! FT) – CR: A2 ace; B1 ab; D. «» (! V) – CR: A2ace; B1abc; D. F2 - S , , ! 80% 10 3 , , Q ! Q , () () ! F ( , .). H1 – T ! ! , 100 . , , ! () (): – ! !, b – , , : (i) – !, (ii) – , (iii) – / !, (iv) – , (v) – ; - ! : (i) – !, (ii) – , (iii) – !, (iv) – . D – U! ! 50 . ! !! . U (! FT) – VU: A2 ace. 58 Y . . 1 (33). 2012

F2 - S , , ! 30% 10 3 , , Q ! Q , !: – ! , – , ! / , – ! , , , , . K , ! !! V , ! . H « » (EN) 2 ; 1 b. I - EN: A2 ace; B1. F2 - S , , ! 50% 10 3 , , Q ! Q , !: – ! , – , ! / !, – ! , , , . H1 – T ! ! , 5 000 . , , ! () () – . T (-! !!) – EN: A1 ace; B1ab. F1 – S , , ! 70% 10 , , ! , , ( ) !: – ! , – .., – ! … H1 – T ! ! , 5000 . , , !: – ! !, b – , , (i) – !, (ii) – , (iii) – , ! / , (iv) – !. K , , , , - «W » (DD). S, ! «H » (LC). Q 1. IUCN Red List Categories: Version 3.1. IUCN, Gland, Switzerland and Cambridge, UK. - 2001. - 32 p. 2. ! V (). - R. – 2001. - 864 . 3. ! V . X. 1.:L. ] 1: U. - F: «S-U» - 2008. - 316 . 4. ! V Y. X. 2 – L. – X - 2003. - 357 . 5. W.Q., U J.W., W! Q.W., V H.H., R Q.S. W! ! RWTU // M - R., 1998. – X. 77, 3 10. - W. 1093-1102. 6. U J.W., W Q.., W .F. ! R W ,1996. V! // H . - R., 1998.- X. 38, 3 1. - W. 159-160. 7. J I.R., R H.F., R H.U., R .H. U ! // W «S RWTU !»: F, SY . -. : c.-2000.- 8. J I.R., R S.^., R .H. F V . W 1.// ! SFS V, . . . - 2010 . - 3 3 (279). - W. 36-49. 9. J I.R., R S.^., R .H. F V . W 2.// ! SFS V, . . . - 2010 .- 3 4 (280). - W. 18-28. 10. ! F . L. - F. - 2006. - 520 . *** ' ', %' IUCN ' ''' %' '. T ' G – F' ', ' ' % '. *** Providing the information on the quantitative criteria, recommended by the IUCN for the categories of the endangered taxons when estimating their status in the protected areas. Giving the estimation of the native species of fishes, as well as of Barbel and Bastard sturgeon of the Balkhash – Alakol basin, based on the aforementioned criteria and categories.

H SY. . 1 (33). 2012 59

YJ 591.0.599.735.5 .. TQb TR Qb TR MLTIT UQb -, F,

J U (N-G ) 2995 . , , , , , , , , , , , , . S . [ ( 25 ), . (23 ). – . V U , , . Q (Silurus glanis L,.1758) !, , . H !, !!! (Ondatra zibethicus L., 1776) - [1-5]. S ! H.W.U [3], 50- . F-J 49 , 70 , 5- (10,2%) V. V [5], 100 (Esox lucius L.,1758) , , . H , K..W [4] , ! ! . S ! 70- ! [6-8] . Y ! . , , . G. H ! ! !! ! , 1957 . (Sander lucioperca L.,1758) . G 23 . [9-10]. c ! ! ! . G! ! ! ! !, ! . U . RQ RTJ QJTO R 1989-1990 . (- ) G , , 2005-2009 . (-!) - (, ) ! . . H 2995 , - ( «G», XTT «-») - !. W , ! - . H 1989 . , FS WWV Z.W.Q, 2007-2009 . SY - F.G. L ( «V !»). F . MQb QJTO TLJ H 1989 .() 854 , 153 (18%). H ! ! (, , ..), ! 1,2% , - 6,5% (.1.).

X1 ] - ] R!, W (, %) H , % R,1989 . 854 153 (18) 10 (6,5) 1,2 F, 1990 . 969 428 (44,2) 8 (1,9) 0,8 R, 1990 . 613 59 (9,6) 4 (6,8) 0,6 , 1990 . 420 69 (16,5) 0 0 F-, 2005 . 68 16 (23,5) 2 (13,3) 2,9 T!-!, 2006 . 25 4 (16,0) 1 (25,0) 4,0 , 2007 . 4 0 0 0 60 Y . . 1 (33). 2012

F-!, 2008 . 31 7 (22,6) 1 (11,3) 3,2 - , 2009 . 5 1 (20,0) 0 0 T!, 2009 . 6 3 (50,0) 0 0 , , . H 1990 . (- ) 2002 , 556 (27,8%). H , 1989-1990 . ( «G») 2856 , - 709 (24,7%), ( 1 3 4- ) – 22 (0,7% 3,1 – ). X , 1989 . 85 , 1990 . – 121, – 153 ( 2 130 . c 1973-1979 . 2 . X , S..Q [6] 300 Z U . G ! H.W.Q [8]. U (1,2; 0,8; 0,6; 0) (n=2856) , ( , IY, H.U.R, , , F.F.G – ), (!, ) , ! ! , , , . R - (G W., G ., I Q., ! M! I. F., Q F., R W., U H., X F., ] G., ^ c. .) , , ( ), , . c ! (G F. – «G»). H 2005-2009 . ( , XTT «-») 139 , 28 (20,1%), .. – 4 (2,9%). G ! (! ) ! , ! , , , !!. H !, ! ! ! ! . , ! ! , !, ! [7]. ! (. .1), !. S !, , ! . H-, ! ! , ; -, , ! . S! 70- ! ! !, , ! . H , ! ! , . H , 1989-1990 2005-2009 . (!, , ), ( ), , , , , (. 2). , ! ! . T, !, . W , ! ! (1989;1990.) , ! Z U!, ! , ! (Fulica atra L.,1758). M 7 ! ! Z U! 50 , 3 10 . J! V ! (: .2). ! ! !! , ! . H 20- ! 1989 . 8070 , , . U , . U 1 3 . T – , , ! . . I 30 80 , – 2 - 4 . J 70 - 90 . W , !, 5 , !. T !, ! – . Y !. T ! . 10 ! 1989 . ( 75 ), ! X . . M, 19 ! 1989 . ( 65 , 80 115 ), . 6 ! ( 65) . 8 ! 1989 . , 165 , - . 18 H SY. . 1 (33). 2012 61

!! . T !! , !!. H - . U !! 1989 . – . !, !! . U 20- !, ! . !, !, , – . T! . H , . , , , , – . X 2 - I , , 1989 1990 1990 1990 2005 2006 2007 2008 2009 2009 n= 2995 V IV V VI IV-V X-XI VII VIII-IX VI- X VII n-854 n-969 n-613 n-420 n-68 n-25 n-4 n-31 n-5 n-6 I.R 10 8 4 0 1 1 0 1 0 0 1) 10 8 4 - 1 1 - 1 - - II. U 3 17 3 0 0 0 0 0 0 0 1) ! 1 ------2) - 1 - 1 ------3) - - 1 ------4) 1 1 1 ------5) - - 1 ------6) - 1 ------7) - 14 ------III. 2 0 0 0 0 0 0 0 0 0 1) ! 1 ------2) ! ( 1 ------IY. 1 10 0 3 1 0 0 0 0 0 1)! ! 1 10 0 3 1 0 0 0 0 0 Y. 79 344 52 61 13 3 0 6 1 5 1) 17 38 5 8 2 - - - - 1 2) 2 ------3) 5 21 ------4) - 45 ------5) 24 51 6 9 5 1 - 3 - - 6) ! 2 - 3 7 - 1 - 2 1 1 7) 27 64 10 11 3 - - - - 1 8) 7 29 7 13 1 1 - - - 1 9) 2 5 1 1 ------10) ------1 - - 11) - 91 20 12 - - - - - 1 YI. 2 9 0 3 0 0 0 0 0 0 1)- 2 8 - 2 ------2) - 1 - 1 ------YII. 5 2 1 2 0 0 0 0 0 1 1) - 2 - 1 - - - - - 1* 2) 5 - 1 1 ------YIII. () 47 25 0 0 2 0 0 0 0 0 IX.J ( , , , , ! 4 13 ------ ..) 62 Y . . 1 (33). 2012

J!, ! ! ! ! - .W ! ! . U , (21 ! 17 . 03 .) !! , , ! , ! , , ! . W . U ! !. U! ! (Rana ridibunda Pallas,1771). K ! 1 2 4,55±0,55 !, ! 2736±330 , ! 0,96±0,31 576±30, ! – 0,41±0,08 216 !, ! . H , ! 12,7 . X , , ! 10 . Y ! , . H , , ! ! - (Astacus astacus). S, ! 2009 . 6 , 3- . H 3 : ! (Carassius auratus), (Cyprinus carpio) (Sander lucioperca), – , (Rutilus rutilus) 3- . MQM] S ( , !, , 2995 , ) !, , ! ! .X - 70-75% ! ! ! . , , ! , , – . U , . X , Z U! ( ! ) , ! , ! ! . X , , , !, . Q 1.W F.F. T. –F-F, 1948. – 250. 2.Q S.U. F! WWWV. –R.,1957. -531. 3. U H.W. F! F-J: F. . . . S. –R.,1951. -19. 4.W K.. T . F-F, 1963.-231 . 5.V V. T ! ! F-J: U- . X,1968.-W.3-173. 6.Q S.. G! ! , . G: F. . . . . – Q, 1975. -19. 7. Q S.., H S.G. U! . G// L.: «V !». -1975,311. –W.22-29. 8. Q H.W. H! ! . //G ! W F . –,1981. –W.120-121. 9. W R.U. F! G//F! WWWV. - ,1972. –W.77- 79. 10. U .F. , G//F! WWWV. –, 1972. –W.65-67. *** T' G '''' (=-G ') ' '' ' 1989-1990 % 2005- 2009 2995 , '. F , '', , , , ', , , ', %' ' '' . P . ' ( 25 ) '' '. L % (23 ) . '' ''' ' – ' ' . L T' G '''' '' '' , '' , '' . *** Nutrition of 2995 exemplars of wels Silurus glanis from water ecosystems of the Southern part of the Balkash Lake was investigated. It was found up that here wels ate mostly food fishes like carp, bream, roach, sander, asp, grass carp, goldfish as well as noncommercial fishes like topmouth gudgeon and gray stone lach. Cannibalism was observed too. Although remains of musk-rats were found in 25 wels nutritions only, harm from wels to musk-rat Ondatra zibethicus was rather significant,. Remains of water birds were found too (in 23 fishes). Acclimatizaed here crayfish was often observed in stomaches of wels recently. People should evaluate imact of wels on water ecosystems of the Southern part of the Balkhash lake not only through concrete number of eaten animals and birds, but trough impossibilities of others to live in running waters so suitable for many animals and birds.

H SY. . 1 (33). 2012 63

YJ 597 J.. L1, .L. 2 LRQ TQJĨ == OJJJİ M=I= IJTQTIQ LĨJO Lļ TZ LOQ J 1 - , F 2X !- , X

'' '' '' '' % ' . L V '' % '' '' ' ' % '' - , , % .. ''' '' '' % . R, L ı Ŀ Ŀ Ľ ı . T ı Ŀ - Ŀ Ŀı Ŀ F, X Ľ ^ Ŀ Ľ ı ĳ ı, . R, «GĿ», «FĿ» Ŀ F, , Ĵ, ı Ľ «XĿ» ^ Ŀı . G Ŀ ! Ŀ ı ı . T Ŀ 1034,7 11 , 11 Ŀ .W 22 . G , ĳ 5660 . Tı Ľ ı Ľ ı ı Ŀ . R, ı ı ! , , Ŀ Ľ , ĳ . , ! , Ŀ ı Ŀ . Ľ GĿ Ŀ ĳ 1995 Ŀ 700 Ľ FĿ Ŀ Ľ 100 , ı 150–200 .W - , ı ! , Ľ , Ľ, Ŀ ĳ . M J Lļ ļJ=RQ ľ Ľ Ŀ Ŀ ı - 2010 ĳ. M ı !, ı ! , . M ı ! ı Google Earth Free ! ı Ŀ ĳ Ľĳ Ŀ, Ŀ ! Ľ . R 2010 . ! ıı Ľ , Ľ . Ĩ- ĳ Ŀ 30-55 , 25-50 . W ı Ľ 2-4 . G ı Ŀ Ľ 12 . ! , Ŀı, ı ĳ U Ľ ] [1, 2] . L ı 8 ı ! - Ľ 32 - . Tı 8, R – 3, V – 6, L – 2, W – 2, G - 4, L – 4, ^ – 3 . M ļLQ= Lļ TQJ QQ I! , Ŀ , , ı ı Ŀı 60-80 , Ŀ Ľ . 2010 Ŀ, L ı ı , , ı , . W ı ı 3-5 , 8 - ( ). M Ŀı ı F Ŀ , 2,2 Ľ 0,9 , Ŀ 195 . Kı J – 0,05 . S, ı . Rı . T ı ı . R, Ŀ ı ı Ŀ , ı ĳ Ŀ . M 64 Y . . 1 (33). 2012

ı ĳ ı Ŀ .F , R -RĿı, ,G H, S .L L, G, J. ^ ĿĿ, W ^Ŀ, F .G, , Ŀ, - - ĳ ı ı Ŀ Ŀ . 2010 ı F,X, ^ Ŀ Ŀ ĳ ı Ŀ ! . R- Ŀ ı Ŀ Ŀ ı ĳ . F Ľ ı ! ı Ľ, , . XĿ Ľ Ŀ , Ŀ , ı Ŀ . S , - - (ı, Ŀ- Ŀ Ľ ..) ĳ ı ı Ŀ Ŀ . L ı ı ĳ Ľ ı ! ĳ Ŀ Ŀ Ľ Ŀ Ŀ . ! ı Ľ 16 ĳ : , Ŀı, Ŀ, , , , , Ŀıı, , , , , , , , Ŀ. XĿ 1- , Ŀ ı Ŀ ĳ ĳ . 1- L ĩ '' ı Ľ ĭ 3 W Ĵ K, F Xı, W ı ı , , Ŀ

1 « , Ŀı, , 3 0,06 18 6 3 X Ŀ X» , , 2 «ı-2» , Ŀı, 1,7 0,6 102 5 2,4 X Ŀ , , 3 «» Ŀ, , 0,7 0,25 17,5 5 3 F, Ŀıı, , , ı 4 «» , , 71 3 1,3 X Ŀ Ŀıı 5 «ĿĽ» , Ŀı, 0,18 0,05 0,9 4 3 G, , ı 6 «H» , , Ŀı 1,2 0,7 84 6 4 G , , 7 «Q- Ŀı, 1,7 0,1 17 8 4,5 , Ŀ » 8 «F ĳ» , Ŀı, 3 0,04 12 4 1,5 G, , ı 9 «» Ŀı, , 0,5 0,3 15 4 0,6 G 10 «^» Ŀı, , 0,47 0,3 14,1 3 1,5 G 11 «ı» Ŀı, , 0,5 0,2 10 5 2,5 ^ Ŀ , , , 12 «» , Ŀı, 0,8 0,6 48 5 1,7 G Ŀ, 13 «W» , Ŀı, , 1,2 0,6 72 5 3 H SY. . 1 (33). 2012 65

14 «K , Ŀı, 0,8 0,3 24 10 6 K » 15 « , Ŀı, 1 0,38 38 10 5 X, , ı» ı 16 «» , Ŀı, 1,5 0,5 75 15 5 Ŀ 17 «F » , Ŀı 0,4 0,15 6 3 1,7 G, ı, 18 «» , , , 0,36 0,2 7,2 5 2 X, Ŀı 19 «» , , 0,5 0,12 6 4 1 G, , Ŀıı, Ŀı 20 «» , Ŀı 0,4 0,15 6 8 3 , Ŀ 21 «Ĳ» , Ŀı, 0,6 0,15 9 4 0,3 , 22 «ĿĿ» , Ŀı, 1,5 0,26 46,8 3,0 0,8 G 23 «H» , Ŀı, 1,6 0,35 56 2,5 1,0 X, , , Ŀ 24 « , Ŀı, 0,72 0,05 3,6 1,85 1,05 L » 25 «L , Ŀı, 0,4 0,03 1,2 4,0 2,7 G » , 26 «G» , Ŀı, , 0,6 0,3 18 4,0 2,7 X, , Ŀ, Ŀ ı 27 « , Ŀı 0,025 0,02 0,05 1,0 0,07 L J» 28 «RĿı» , Ŀı, 0,4 0,2 8 1,5 0,7 G,, , ı 29 «» Ŀı, 0,15 0,05 0,75 1,2 0,5 L 30 «X» , Ŀı, 0,5 0,06 3 2,0 1,0 L , 31 «» , Ŀı 0,4 0,15 6,0 1,6 0,6 K Ŀ, ı 32 «F» , , Ŀı, 2,2 0,9 198 2,0 0,5 X ľ , Ŀ - « X», «H», «ı», «ı-2», «»,«W», «», «H», «X», «F», «G» . W , Ŀı 2 ĳ – «K », «F », «», « J», «RĿı», «», «» . F Ŀ 32 Ŀ ı Ŀı . , 32 . W – , Ŀıı, Ľ .T 2-3 Ŀ . ı ĳ , Ŀ Ŀ ı ı ĳı Ľ ı Ŀ . Wı ı 80 Ľ, Ŀı, Ŀ . X, , Ŀıı, , , Ŀ ! ĳı Ŀ.M Ŀ ĳ ĳ ĳ, Ŀ, Ľ ĳ. G ĳ, Ŀ ĳ Ŀ ĳ. G ı ĿĽ ı Ŀ . G - ĳ , Ľ , Ŀ ĳ ĳ, 66 Y . . 1 (33). 2012

Ŀ , , ı - ĳ ı Ŀ ( ı Ŀ Ŀ), , Ŀ Ŀ ( Ŀ ĳ) . , ı ı Ľ ı Ŀ ! . G ! ! Ŀ, ı . L ! : ı Ŀ ; Ŀ Ľ ı ; ĳ Ŀ ı ı , ĳ ı , Ŀ; Ľ ı . , Ľ Ŀ Ľ ı ĳ ; ! ı ! ĳ ı , - - Ŀ . S, ĳ Ľ ( ĳ, , Ŀ Ľ ..), ! - - ĳ . ļJ 1. U .. V . – R.: U! ,1966. – 306 . 2. ] S.. R ! .-R.: W! , 1952 – 55 . *** 2010 . U L , . *** The paper presents the results of 2010 year of investigations. Data on ihtiofauna species diversity of small reservoirs of Zhambyl region are given. Moreover their morphological characteristics are described.

YJ 597:576.895 .R. L, .. , .. Q UM]TIT ] ERGASILUS SIEBOLDI UTJO UT\Q UO ]T . -

Ergasilus sieboldi 4 . J , . U, , , (20°-25°) , . Y, , : G (Gobiidae) 37 [1, 2], 19 , !! 9 18 – . H W 70% !! ! [3]. G - , 20-25 ! 100 . H F ! !, , !! ! ! , !. G ! !. Y!, ! !! ! , , ! . U! , : ! ! ! ! [4]. H - , , ! ! [5]. H SY. . 1 (33). 2012 67

G !! , , ! !! . H ! , 2010 . RQ RTJ QJTO R ! , ! ! 2010 . R 4 % , ! [6]. H 190 ! , !! 4 : Neogobius fluviatilis (–), Neogobius melanostomus (–!), Neogobius caspius ( ) Mesogobius gymotrachelus ( ). MQb QJTO TLJ G 120 ! , , 53 ! (N. fluviatilis), 27 – ! (N. melanostomus), 15- (M. gymotrachelus) 25 – (N. caspius). H 70 , : – 47, ! – 10, – 7 – 6 !. J! , , . X, 2,4 ; ! – 3,3 ; – 2,3; – 1,8 . H 1:1, ! 1:2 , 1:1, 1:1,5. U , , . T ! ! , ! ! Ergasilus sieboldi, ! Copepoda. E. sieboldi !!! , , [7]. V ! !. J! !! 20 25ºW [8]. F , , . . 2 !, . C K. sieboldi 0,5 1,5 , ! ( 1 F). S , , , ( 1. G). J !! ! ! [9]. W ! ! , 30 - 40 !. , ! 140 ! , !!! , , ! 1,5 . H ! 5-10 ! .

F G 1. () () Kgasilus sieboldi I , ! ! ( 2). 68 Y . . 1 (33). 2012

FG 2. L , K. sieboldi. T . T . . – 200; - 100 W , , !, , !! , ( 3).

F G 3. L , K. sieboldi. L . T , , , T . . 200 U , ( 4 F), ! . U ! , ! ! ( 4 G), ! , ! ! [10]. W , , , , ! , . ! ! . Q , . , ! ! , ! !. U , , , ! ( 5 F). 87 86% II , 14% III . H 36 50% II 50% III . U . U ! ( 5 G). X 33 60,7% ! 2 39,3% III-IV . T , 70% ! III 29% II H SY. . 1 (33). 2012 69

FG 4. - L K. sieboldi . – . T . . 100

. FG 5. - II . \ , . – II – III . T . . - 400; - 100 X , ! , , ! . R! . Q 1. K.S. T V ! – R.: Q! ! , 1981. – 166. 2. J.G. V G! ! ! W Z !// H X 36 – FSS.: S, 1991. – W. 223-231 3. J.G. V, Q.R.W G. : . R.: S,1989. – W.190 - 193 4. .W., K T.H., R c.., J W.I., L S.H. ! !! // I, ! !. – 2001. - 3 3-4. W. 21-27 5. R Q.H. T - , ! ! // . . IV H . . . – F, 2001. – ]. 1. – W.32-33 6. V G. R! . R.: - . - 1953. 718 . 7. S.F. I S.F, W Z.F . ! – R.: R, 2003. – 448. 8. J H.F. U ! - R.: I ! – , 1932. – 151. 9. J F.H. M! - R.: H! , 1981. – 600. 10. H., ! T. V - R.: U,1959. – 29. *** R 4 '' ' '' %' . T ' % ''' '' '. G ' ' , ' % %''' ' (20\-25\) , '. U '' ', %: %'' , - .

70 Y . . 1 (33). 2012

*** In article influence of parasitic invasion Ergasilus sieboldi on reproductive potential of 4 kinds Gobiidae’s fishes were considered. The short characteristic of biology, morphology and life cycle of this parasite were given. It was shown that the greatest contamination of Gobiidae’s fishes were observed during the summer period when the water temperature optimal (20 °-25 °) for reproduction and development Ergasilus sieboldi by the autumn contamination of fishes parasites decreases. It was established that males Gobiidae’s fishes for whom the care of posterity is characteristic are more subject to an attack of parasites: they protect the laying up for emerging larvae and consequently are inactive

YJ 639.2.03 Q..M Z.. Q I U_ UTT UTUQ\ UOTO QI (HUSO HUSO L.) M UTQRTO TIT TTQ MUT XTT

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( ); vm , v f – ttt tt ! t. H! (1) v , v mtt ft ! (M, 2006, M 2011):

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Bt – ! t; Q – !, ! 1 .

. ; Wt - . v , ! (1) mtt !, (M,W- ,1983;M,1986,2006, M, , 2011): v 1 Ft k (T k t k ); (7), F, k, T – . mt M! F, k, T ! (19) .. ^ (1935),(M,1986,2006) - : l qt k ; (8) W pt ; (9) W l ; (10) l, W – t; q,p, – , t 1 l 1; k,,! – , (M,1986,2006). M! (8)-(10) ! (G,1964), «Mirosoft Excel 2003». F, T k ! (12) q,k,C,! - ! (M,1986, 2006, , 2007, M, , 2011): 1 l 1 1 vmp M p p L 2l p k k k L k F ; (11) vmp 1 e ; (12) M ; (13) t p ( ) ; (14) T ; (15) T ( ) ; 2k p q t p t p t p q q q (16)

: vmp – !; l p ,t p – , !! 50% !; M p - !; L – ! ! k !, L 2l p ; T – , !! (18); X – . M! (8)-(10) (7) 1. X 1. M (Huso huso L.). , q k p C l p M! 57.52 0.4523 0.0011 3.29 677.9 1.4881 200.0 , L t T M v T k A p p mp M! 400.0 15.72 72.8 0.0946 0.090 6.954 0.0753

Cr I.I.H (1958): Cr 1.25(Rc Pw Pq ); (17)

: Rc - ! ; Pw -

; Pq - .

I Rc , Pw

Pq , ! – K 2 KK 0.810 : R 1.95W ; (18) (19) (20) Pt Pq (21) (22) c Pw W2 W1; Pq 0.07Wt ; K ; PP ; 2 Rc Pw Pq Pw W , ! , !

R0.5 = 1 . , 2.

72 Y . . 1 (33). 2012

X 2 , 1 . . ( – 50,0%). H- J- R- - - . ]- G- c- U I- T- K 2 KK , , , . . . - . -. - , t, l W, . . . . , , Cr . , , R . v f c , P Pq vm . w . . 0,1 20,3 0,022 0,632 - 1000,0 0,022 0,571 0,519 0,0192 1,387 0,49 2,58 1,1 60,1 0,781 0,467 - 368,2 0,288 0,709 0,357 0,0279 1,367 0,35 3,56 2,1 80,5 2,045 0,372 - 196,1 0,401 0,715 0,256 0,0318 1,254 0,29 4,35 3,1 96,0 3,651 0,305 - 123,2 0,450 0,691 0,197 0,0335 1,151 0,25 5,00 4,1 108,9 5,534 0,255 - 85,57 0,474 0,663 0,159 0,0345 1,070 0,23 5,55 5,1 120,2 7,658 0,216 - 63,74 0,488 0,640 0,133 0,0351 1,010 0,21 6,01 6,1 130,3 10,00 0,186 - 49,95 0,499 0,622 0,115 0,0357 0,965 0,19 6,41 7,1 139,6 12,53 0,161 - 40,68 0,510 0,608 0,102 0,0364 0,932 0,18 6,76 8,1 148,2 15,24 0,142 - 34,13 0,520 0,598 0,091 0,0370 0,909 0,18 7,07 9,1 156,2 18,13 0,126 - 29,30 0,531 0,591 0,084 0,0378 0,891 0,17 7,35 10,1 163,7 21,17 0,114 0,017 25,61 0,542 0,581 0,076 0,0381 0,869 0,16 7,59 11,1 170,9 24,36 0,105 0,033 22,27 0,542 0,561 0,069 0,0378 0,835 0,16 7,81 12,1 177,6 27,70 0,098 0,050 19,19 0,532 0,531 0,062 0,0366 0,787 0,16 8,00 13,1 184,1 31,17 0,093 0,067 16,36 0,510 0,493 0,054 0,0347 0,728 0,15 8,18 14,1 190,4 34,78 0,091 0,083 13,74 0,478 0,448 0,047 0,0322 0,659 0,15 8,33 15,1 196,4 38,51 0,090 0,100 11,35 0,437 0,397 0,040 0,0291 0,583 0,15 8,48 16,1 202,1 42,37 0,090 0,117 9,200 0,390 0,344 0,033 0,0256 0,503 0,15 8,61 17,1 207,7 46,34 0,092 0,133 7,298 0,338 0,290 0,026 0,0219 0,422 0,14 8,73 18,1 213,1 50,43 0,095 0,150 5,653 0,285 0,237 0,021 0,0182 0,345 0,14 8,83 19,1 218,4 54,63 0,100 0,167 4,267 0,233 0,188 0,016 0,0147 0,274 0,14 8,93 20,1 223,5 58,94 0,105 0,167 3,131 0,185 0,146 0,012 0,0116 0,212 0,14 9,02 21,1 228,5 63,36 0,111 0,167 2,281 0,144 0,112 0,009 0,0090 0,163 0,14 9,10 22,1 233,3 67,88 0,118 0,167 1,647 0,112 0,085 0,007 0,0069 0,123 0,14 9,17 23,1 238,0 72,50 0,127 0,167 1,177 0,085 0,064 0,005 0,0053 0,092 0,14 9,24 24,1 242,6 77,22 0,135 0,167 0,832 0,064 0,047 0,003 0,0039 0,068 0,13 9,30 25,1 247,1 82,04 0,145 0,167 0,581 0,048 0,034 0,002 0,0029 0,050 0,13 9,36 26,1 251,5 86,95 0,155 0,167 0,400 0,035 0,025 0,002 0,0021 0,036 0,13 9,41 27,1 255,8 91,95 0,166 0,167 0,271 0,025 0,017 0,001 0,0015 0,025 0,13 9,46 28,1 260,1 97,04 0,177 0,167 0,181 0,018 0,012 0,001 0,0010 0,017 0,13 9,50 29,1 264,2 102,2 0,189 0,167 0,119 0,012 0,008 0,001 0,0007 0,012 0,13 9,54 30,1 268,3 107,5 0,202 0,167 0,076 0,008 0,005 0,000 0,0005 0,008 0,13 9,58 31,1 272,3 112,9 0,215 0,167 0,048 0,005 0,004 0,000 0,0003 0,005 0,13 9,61 32,1 276,2 118,3 0,228 0,167 0,030 0,004 0,002 0,000 0,0002 0,003 0,13 9,64 33,1 280,1 123,8 0,242 0,167 0,018 0,002 0,001 0,000 0,0001 0,002 0,13 9,67 34,1 283,8 129,4 0,257 0,167 0,011 0,001 0,001 0,000 0,0001 0,001 0,13 9,69 35,1 287,6 135,1 0,272 0,167 0,006 0,001 0,001 0,000 0,0000 0,001 0,13 9,72 36,1 291,3 140,9 0,287 0,167 0,003 0,000 0,000 0,000 0,0000 0,000 0,13 9,74 37,1 294,9 146,7 0,302 0,167 0,002 0,000 0,000 0,000 0,0000 0,000 0,13 9,76 38,1 298,4 152,7 0,318 0,167 0,001 0,000 0,000 0,000 0,0000 0,000 0,13 9,77 39,1 302,0 158,7 0,334 0,167 0,001 0,000 0,000 0,000 0,0000 0,000 0,13 9,79 40,1 305,4 164,7 0,351 0,167 0,000 0,000 0,000 0,000 0,0000 0,000 0,13 9,80 H - - - - 2136,6 9,220 11,05 2,499 0,664 17,76 0,22 5,62 V , ! ! t p 15.72 , ! - T 72.8 . S vmp 0.09 (9,0%) ! ! ( 1). W ! ! . S (!) ! 10 . T 1 . ! ! !! 2136.6 . . – 9,22 . ( 2), .. ! ! (2,136 . .) ! (1,0 . ) 2,13 . U (50%- ) ! ! ! 0,1-15,1 – 7,23 ., 78,4% . Y! !! 2136.6 . . 9,22 ., !! 1 . . , ! 17,76 . , 1,926 ! .

! 17,76 . Rc ! ! 11,045 . (62,2%), – , H SY. . 1 (33). 2012 73

– 2,449 (14,07%), – 0,664 . (3,74%), ! ! 3,55 . 20% ( 2). c !

! K 2 = 0,17 17,0%. ! =8,24 , ! 8,24 . H ! ! . G !! , !! 50%- ! !. W ! ! ! . Y ! ( ). I !! , ! ! ! (10 ). c ! ! . W ! ! . W !. U ! ! , . c ! ! , . Q 1. F F.. ! M! // . F. WWWV. - Q: S, 1986. - 230 . 2. G S.. G! // X. HSVT., X.LII, 1964. W.183-258. 3. H I.I. T ! . - Q: S, 1979. - 271 . 4. J! U.F. V // V . -1934.-34.-W. 27-29. 5. J! U.F. T // . ISTV. -1948. - X. 29. - W. 56-64. 6. M F.H. J . - R.: U. -, 1976.-312 . 7. M Q.F., W H.F. Y ! ( ! . K) // V !. - 1983. - 33.- W. 36-37. 8. M Q.F. R , // W. ISTV. -1986. - H. 243. - W. 14-21. 9. M Q.F. G ! . – F…. . ., F, 2006. 58 . 10. M Q.F., .H. U Huso huso (L) // S ( ). F, - «».-2011. – W. 135-151. 11. .H. G ^ . F…. . ., F, 2007. 24 . 12. R I.Q. c! !, J // . ISTV.- 1975.-X. 101.-246 . 13. R I.Q. S ! ! // W. ISTV. - 1984. - H. 222. - W. 3. *** G ''' '' ' '' ' % '' '' ' %' . L'' ''' ' ' % ' %' . *** Study of nutritional requirements of the fish is necessary when solving a number of theoretical and applied assignments associated with artificial regeneration of fish resources, assessment of damage incurred during the technogenic impact or directed formation of internal water basins ichthyofauna. Conducted surveys will allow to approach the issue of volume assessment of artificial reproduction ensuring regeneration and further formation of the beluga reserves in compliance with products of used fodder resources.

YJ: 639.3.05 T.R. , I.. , .. TMT-TT TTJT M . -, F,

– . V – P . X " . H , ! ! ! ! ! ! ". Y! ! (14, 6 ) ! ! ! , 272,0 . . 74 Y . . 1 (33). 2012

H !! " , ! ! 50 %. X, 2005 " 41,9 . , ! 60-70 XX ! 110 .. H . X, 80- XX , ! 10,0 . . H ! ! ! . X, !! , ! ! [1]. X, 2009 ! ! 90,0 . . , – 55,1 . [2]. c , R . H . V , ! ", ! ! ! 5 . [3]. H ! , , ", !! ! ! . G " – - . T ! , . T ! " !! ! ! . U ! ! ! ! . U , !! ! 75-100 /. ] , ! . W!! 20-25 /. J . H 1980 – 1988 . 11 – ! . V ! 48-52 /. U ! " !! 1985-1986 . Y 1986 . " 1500 , 180 , – 120 /. H 1989 - ! " ! " . T! " 10,0 . . T – 900 ., – 90,4 / ( 1). X 1 ", 3 S U, H , V, " / 1 -T! 620 248 400 (.J, , - F) 2 X! (. 200 18 90 , L) 3 J! (. 90 2 22 W) 4 ! (. 900 160 187 G, . R) 500 20 40 5 U! (. 600 23 38 W, ) 6 F! (. 4100 73 18 G, Y,L) 7 Z – ! 1420 170 120 (G H )

8 H – ! 150 112 740 (G! SH)

H SY. . 1 (33). 2012 75

9 ! 66 41 621 ( ) 10 W – ! 1310 24 18 (. U, U!, M) : 9950 900.0 90,4 F !! " ! , ". " H – – T ! ! ! ! ! ! , 400 746 /. U , ! " – . H ! ! ! . X " 150-200 , .. , , ! " ! . H ( 2), " ! , " . X, " – . U ! – , , . () – 20 100 %. X 2 1988 H H, H 23 223 Q 97 L 38 M 18 370,11 U 40,0 W 4,6 W 3 W 10 X 34 _ 36 X ", ! ! , , ! . S! 1990- ! ! . M . R !! !, ". H ! ! " ! ! . X , ! ! . T, ! " ! ! 15 . U ! ! ! ! !! ". Q 1. G I.F., ^ c.K. G . – R., \SXcV, 1981. W.42. 2. R I.I. T – F-F, IY, 1987, .69. 3. ^ F.. X ". – F-F, SSX, 1988, .48 4. O W.c. Y .-R.:F, 1985, .158. *** R '- ''' % '' . '- ''' . P'' ' ' '' . *** Development history, modern conditions, perspectives of development in Kazakhstan of lake commercial industry are considered. Commercial fish industry is under way on small basins on halfintensive technology with important fish species

76 Y . . 1 (33). 2012

YJ:639.3 T.R. , I.. , Q.. J UTMTJQO U (CYPRINUS CARPIO) UIOTIT TT – TTIT TMO . -

P – , - . J . S . – ! (SH) 1973 . V . SH, !, ! (, , , , ! ! ). 700 . H ! , , - ! , ! , ! ! . U !, ! . ! . G , – 100 . . H ! ! . U ! ! , ! ! ! ! !. H ! ! ! ! !!! . Q. T ! ! 30% !. H ! ! . K !! ! ( ) ! ! . \ !!! ! !! !, ! . S ! !! ! . T () ! 2011 U ! ! , ( 1). X 1 . U J W!! V! T! H , , , , % , .. ./ 13.04.11 I 10 5,69 308,3 54,2 32,0 20.04.11 II 7 5,3 214,3 40,4 66,7 22.04.11 III 13 4,8 211,5 44,1 74,4 24.04.11 IV 9 6,1 314,8 51,6 77,8 26.04.11 V 9 5,1 328,7 64,5 65,7 27.04.11 VI 9 4,8 250,0 52,1 84,4 30.04.11 VII 12 5,2 395,8 76,1 65,3 01.05.11 VIII 12 4,6 229,2 49,8 72,7 03.05.11 IX 4,5 : 85 5,12 250,3 48,1 60,1 S! ! ! 13 ! 2011 ., 15-170W. T 2011 ! ! «S -4». V , . , ! 211,5-395,8 .. H ! ! 250,3 . , 50%. T! ! ! – 48, 1 . . 1 . H SY. . 1 (33). 2012 77

H ! 32, 0 84 ,4%. H 60,1 %, !. U , , ! SH. R ! , , . X! ! ! ! . ! , , . H 15 %. 50%, ! . U . W . U . X , . Q 1. U Z.F., H H.F. V – R.:R,2004, .456. 2. W – . X.1,2-R.: F. 1988, .576. 3. \ V.R. M ! . –R.: . HSUT , 1980, .31. *** R – '' % – ''' '' . X ''. &' %' %'' '' '' % '' ''. *** The descriptions of Kapshagai industry, modern conditions of female band are considered. Estimate of female crucian carp to receipt posterity. On basis of analysis reproduction recommend by bumping female band on this industry.

YJ 597+639.3 I.. MTTM ]QTb ]LTJ QRTR UJTTR TMO T «» . F-

F FT «GSX». 2006- 2008 . , " P . , , , , , . U ! " , ! . V ! 5- «V » [1]. H ! !! ! ! ! . \ ! !! ! , ! F !. RQ RTJ QJTO F ! 800 !. H ! !! .R! F-F . F . . ! - . U ! ! ! !. R ! : , 2006-2008 . ! : ( ), ( ) ( ). T [2]. X ! ! [3]. MQb TLJ ! ( Cyprinus carpio, Hypophthalmichthys molitrix Ctenopharyngodon idella) ( Carassius gibelio, Sander lucioperca, Perca schrenkii ). 78 Y . . 1 (33). 2012

H 7 , !! “”: (Pseudorasbora parva), (Hemiculter leusisculus), ! (Abbottina rivularis), (Micropercops cinctus), ! (Oryzias latipes), (Rutilus rutilus) (Rhinogobius similis). H , ! ! . T ! ! ! ( 1).

X 1 2006 . 2007 . 2008 . H T H T H T ! , , , H 2 78 4 66 3 34 Q 7 126 7 62 7 38 T 7 272 7 23 7 77 M H 3 44 2 70 3 30 T 2 28 2 118 2 48 R Q 2 18 2 47 4 35 T ------H 3 12 2 95 2 38 H 4 25 3 85 2 120 Q 7 208 5 570 5 385 T 7 1120 7 938 7 1350 I H 4 12 4 36 2 14 Q 3 48 7 108 7 26 T 5 64 7 190 7 48 H 2-4 , , , ! . ( 50 80%). H!, ! , ! ! ! , !! . H . H 2-3 : , . c ! , ! : ! ! ! 1-1,5 ! - ! !. M ! ! !, !! - !!. K !!! , . U . U ! 650000 25-30 . 1 . H , ! ! ! ! !! ! !. H , , !, ! ! . . H 7 . G ! , ! ! ! !. W , !! 1 4-5 . H, ! , ! ! ! . ! ! (.2).

H SY. . 1 (33). 2012 79

X 2 T «» ( ). 3 H ! / I S, H, S=4,5 S=1,0 S=10,5 % % % 1 F 42 50 102 38,1 68 36,2 2 H - 6 2,2 3 1,6 3 V! 12 14,2 53 19,8 15 8,0 4 22 26,2 26 9,7 12 6,4 5 R - 14 5,2 5 2,7 6 U - 8 3,0 13 6,9 7 8 9,5 46 17,2 48 25,5 8 U! - 5 1,9 8 4,3 9 W - 8 3,0 16 8,5 H: 84 100,0 268 100,0 188 100,0 2, !!! : , 50%, 38,1% 36,2% . ] 9,5% 25,5 %. J! : 26,2%, 9,7% 6,4% . R !! ! (2,2% 1,6%), (5,2 2,7), (3,0 6,9%) - . ! Triplophysa strauchii T.dorsalis. MQM] H F ! , !! ! !: , , ; – ! ; – , , ! , , , . S ! , ! . Q 1. R H.U., J I.R. ! //V , 5 . 1992, .6-44. 2. G F.F., X W.V. - , F, 1999. 347 . 3. G! S.I., S F.R. V !.R.: X R.2004. 389 . *** 2006-2008 ' %' '' %' ' ' . ^ , , ' , , ' '. *** Investigation of fish diversity in the Almaty fish farm “BENT” was provided in 2006-2008. Noncommercial fish species like topmouth gudgeon, amur false gudgeon, common sawbelly, ricefish, beautiful sleeper and amur goby fish were revealed. Fish diversity and abundance depend on ponds exploitation regimen.

YJ 597 .. T TTQ QT-UOTO UTUQ\ QbJ ALOSA KESSLERI KESSLERI (GRIMM, 1887) M- XTT « - !», -mail: [email protected]

- -. R . Y . 2009-2011 . . 2009 . 3+. 4+ 5+. . H SV 2009-2011 , «! - !! ! , ! ! », - , , -, 3 . U 2009 . 3+, 80 Y . . 1 (33). 2012

4+ 5+. H 2010 . 4+, 2011 . 5+. M ! 3 ! 2 !, ! . U! - – ! ! - Alosa. H ! . S . H . Y [1]. S ! -! !! ! !. M 1983 2004 , - Y, ! ! . X ! S U, - « () , !» [2].

RQ RTJ QJTO W !, - ! , ! . H ! ! ! ! , ! . J! ! . J! !, , ! ! ! . V , !. G 2-3 ! ! . 50 ! ! !, ! ! . MQb QJTO TLJ T! !, – . J . 1. , Y -. U , , , . X 1 U , 2009-2011 , ., – . I W 1 , - . %, . %, . %, 1 1 1 2009 14 25.2 16 28.8 5 9.0 2010 17 28.9 18 30.6 7 11.9 2011 11 25.3 15 25.5 4 6.8 W - ! 35-40 % 27-38 % . X ! ! . S . 2 - 2009–2011 . K ! !, ! . V 3+ . W, ! . V 4+ ! 2009 ., ! 9.1 %. H 2010 . 38.6 %, 2011 . – 32.9 %. V 5+ 2009-2010 (4.2 6.5 %, ). M 2011 . – 23.7 %. , , 4+ 5+ ! 2009 . X 2 J - - . MT, 2009–2011 . I H , 3 4 5 !! !! % !! !! % !! !! % . , . , . , , - , - , - 2009 30.8 386 86.7 34.2 514 9.1 37.2 675 4.2 2010 30.7 379 54.9 33.7 484 38. 6 36.4 651 6.5 2011 30.9 381 43.4 33.6 497 32.9 37.8 679 23.7 H SY. . 1 (33). 2012 81

! , 1895 . 100 ! . T 1890- [3]. . H - , 1984 2000 . ! 0.806 4.310 . , – 1.7 .. J ! 3.34 31.3 ., 5.3 .. (H! H.H., 2000). c ! ! . H ! , ! , !. H . Y , ! ! !. U ! ! Y , ! ! - , !! ! . S (! ) ! ! , ! ! ! ! . MQM] H ! ! ! , ! (, , , , !), ! , , , V (, , ) [4], ! - ! ! ! . H ! - . T , ! ! ! . ! , , ! ! ! . Q 1. F.., W F.., F ^.F. W ! Y- . – F: «G». – 2005. - W. 55-56. 2. M! K.H., V F.G., ^! I.H., ^ H.F., W .F., X H.I., R Z.F., K H.U., S.S., R K.Z. V S U!. – W: - W. -, 2007. – W. 36-37. 3. G! S.I., X.. M ! S.F. G //R . . . «S F G (1861-1937). Y ! V, W^F». W-U, 2009. – W 10- 11. 4. F.., U J.H. !! Y M- V // R . -. . «S ! ». F, 2010. –X. 2. – W. 383-385. *** G ''' L-P %' '. L '' ' ' '. 2009-2011 ' ' '' . 2009 '' 3+ . P'' ' 4+ % 5+ . G %'' ''. *** In the article shows data by recovery Ural-caspian populations of Alosa kessleri kessleri. Spawning migrations of species in the river Ural not registered several ten years. In 2009-2011 in this river become apparent spawning herd . In 2009 spawning fishes were at 3+. The next two years spawning fishes were at 4+ and 5+. That would indicate about recovery field populations.

YJ 597.554.3(282.25) T.. RQ RTZTTQTI]TO Q (AlBURNUS AlBURNUS) M 8 M] JQ TR TJTRT TIT O XTT « S !», F

, , . J , . V -N . H ! ! ! 36 , !! ! ! ! , ! , ! ! , !, , ( – Abramis brama, – Sander lucioperca, – Coregonus albula, – Cyprinus carpio carpio, ! – Coregonus peled, 82 Y . . 1 (33). 2012

– Pseudorasbora parva, Carassius auratus auratus, !! – Pungitius pungitius, !, – Hypophthalmichthys molitrix, – Cottus poecilopus. H ! ] , . U ( – Astacus leptodactylus, !), ( – Eriocheir sinensis), ! ! , , ! ! . H ! ! , " , ! . RQ RTJ QJTO W ! 2009-2011, , ! ! 037 «I ». T ! H- . : – 176 ., – 5 . R ! , ! ..U [1]. . H ! . H - ! ! [1,2]. H ! ! !! , ! , 11-12 . MQb QJTO TLJ Y, !, H ! , ! . Y! – U . J " K, V, , -, ! . H ! H, Y, c, W, . ] -W . H , ! , ! [3]. T ! ! " 90- X. U, ! 2000- . ! H- , , ^ . H H , ! ! ! ! , ! . G , - , , ! . U - 2006 , ! ! 10-11 % ! , 0,54 /, 2009 . " ! 23%, 1,0 /. H – 2010- 2011., ! ! ! ( 12-20) " ! 90-95 % , 1,86 /. Y! , ! . H! ! ! , ! . Y! - !, ! ! ! 15-16 , ! , ! ! . U , , 6 . V ! , ^ ; ! 1-2 , ! [4]. ! ! ! ! . W - 10,8 - 11,0 16,6-17,9 ( 1). Y ! !, !, ! – 60 %. Y ! " ! 1,3-1,5. X 1 J H W!! , W!! , Y . 2009. 2010. 2011. 2009. 2010. 2011. 2009. 2010. 2011. 2 11 - - 18 - - 1,4 - - 3 13,3 10,5 9 34,8 15,6 10 1,5 1,3 1,4 4 14 10,9 10,1 36,6 16,8 13,7 1,3 1,3 1,3 5 - 11,2 11,2 - 17 18,2 - 1,2 1,3 6 - 12 12,4 - 20 26,2 - 1,1 1,4 W 13,3 10,8 11,00 34,0 16,6 17,9 1,5 1,3 1,3 H SY. . 1 (33). 2012 83

U! ! ! , . Y! !!! . S ! ! ! !. H ! ! 3-4 , 7-9 . R! ! 11,69 . . H ! ! , !, ( 2). X 2 J , (. .) I H W!! FU 4 5 6 2010 1,88 2,64 2,39 2,10 2011 2,8 11,69 4,49 8,36 V , , , 0,9-6,3 ./3, !! , , . R! ! 3. X 3 R U U R ± m .. W min max W 11,52 – 0,14 0,61 5,33 10,9 12,3 J 9,8 – 0,10 0,45 4,56 9 10 F 19,6 – 0,12 0,55 2,79 19 20 P 12,0– 0,16 0,71 5,89 11 13 V 8,0 – 0,10 0,45 5,10 8 9 48,4 – 0,46 2,07 4,28 45 50 . . I . 20,0 – 0,16 0,71 3,53 19 21 6,90 – 0,05 0,22 3,24 6,5 7 40,20 – 0,24 1,10 2,72 39 42 % 20,50 – 0,33 1,47 7,19 17,9 21,5 15,98 – 0,21 0,92 5,78 15,1 17,5 ! 25,02 – 0,30 1,33 5,31 23,7 27 ! 8,04– 0,17 0,77 9,62 7 9,1 ! 56,68 – 0,29 1,31 6,30 54,6 58 ! 33,78 – 0,28 1,23 3,65 31,7 34,6 ! 17,92 – 0,41 1,83 10,19 15,4 20,4 ! J 11,14 – 0,47 2,09 18,77 9,7 14,8 ! J 16,64 – 0,39 1,74 10,43 14,4 18,6 ! F 18,70 – 0,12 0,55 2,92 18,1 19,4 ! F 12,60 – 0,27 1,19 9,44 11 14 V 18,82 – 0,25 1,10 5,85 17,1 19,8 V 13,70 – 0,24 1,06 7,70 12,8 14,9 ! V V 18,64 – 0,65 2,90 15,55 15,3 22,4 ! V F 16,68 – 0,32 1,45 8,71 14,9 17,9 % 25,50 – 0,67 3,00 11,76 21,7 30,1 27,78 – 0,58 2,61 9,38 24,9 31,9 39,26 – 0,83 3,72 9,48 34,3 43,9 30,04 – 0,61 2,71 9,02 27,2 33,7 42,14 – 0,55 2,44 5,79 39,1 45,1 84 Y . . 1 (33). 2012

T -!, -!, , ; -. ]! !, ! !!, . S , ! ! . F , ! . Y !! . V . X , ! ! , ! ! !. 45 50 , !! 48,4. H 2 8 , ! 9 10. H 3 19,6 , ! 19-20 . ] 12 ., 8 . ! 20,0, ! 19-21, -40,0 (39-42) ( 3). M ! , , ! (). M! , ! : 2.5 – 5.2, ! 2.5-4.2. X , ! ( % ) ! 25,02 % 3 (8,04 %). J ! ! –20,5 %, " -15,98 %. M ! " -39,26%, -25,5 %. Q - 30,04% . I , ! –27,78 %. F ! ! – 56,68 %, – 33,78 %, ! – 17,92 %. W , 1,8 ! – 24,79% – 13,87. I , – 20,58% – 17,74%. W - , ! ! – 18,7 %, ! -12,60%. V! , - – 18,64 % 16,68 % . . U! H - , , ! ! , ! ! ! , ! , ! . U! , ! . H ! , !, , ! , , , ! , , ! !. Q 1 U .. V . – R.: U! , 1966. – W.376. 2 Q I.. G!. – R.: H! , 1980. – 296. 3 R H.U., J I.R., . V . – F-F: S, 1989. – X. 3. – 312 . 4 ^ I.. G! . R.:S,1964, 176. *** R ' ' % '''' % ' ' ''. G ' ', %' % ' '' '. L K' '' ' % . *** In this article are mentioned some features of the ecology of the area, biology and morphological characteristic of the bleaks that belong to the alien species category for the settling reservoirs. There are given the analysis of the modern status of the “invader’s” population, its influence to the ichtyocenosis of the settling reservoirs and also the degree of the biological diversity. There are worked out its role in the ecosystems of the reservoirs of the Upper-Irtysh basin and the prospects of its usage in farm.

H SY. . 1 (33). 2012 85

YJ 597-19 1Z.. , 2.. R TRO T TZ . Q ITTO UJITTO MT 1 XTT« », F 2JIU «S » VIU «SY .F-», F

U .^ . T - Diptychus dybowskii, Triplophysa stoliczkai, T.labiata - Paraslmo mykiss (Salmo gairdnerii). U . H ! ! ! [1,2]. H ! , - , , ! !!! ! V . T ! ! G , ! ! .G, . [3,4]. V.^ !!! . V . W! . \ ! !! ! ! .^ ! . RQ RTJ QJTO W ! 2008-2010 . , ! [5]. J! ! ! , ! [6]: .1-4 ^ G , .5-8 – . H ( !! ) ! . T ! 5070 , . X . R , ! , , ! ! [7, 8]. H 2010 . , , ! 8 . MQb TLJ . ^ 4 (.1). I Dipthychus dybowskii Kessler, 1874, Triplophysa labiata (Kessler, 1874), T.stoliczkai (Steindachner, 1866) !! ! . S ! - Diptychus maculatus Steindachner, 1866 ! Schizothorax argentatus pseudaksaiensis Herzenstein, 1889. T !! , ! !, - , !!! , V [9] F [10] , !, .^. X 1 - ! (! !). W ! ! . T [11,12] ! Salmo (Parasalmo) gairdnerii Richardson, 1836, [13,14] Parasalmo mykiss (Walbaum, 1792). H .^ . V! , 1966 G !, [15]. H 1970- ! .^ Y , [16]. W ! . T ! G . ^ - . V ! - ! !, , . U G ^. V- ! 2. 86 Y . . 1 (33). 2012

X 1 . Q V ! Parasalmo mykiss (Salmo gairdneri) V! H, Diptychus dybowskii I c, Triplophysa labiata T c, Triplophysa stoliczkai X c, V! ! . ^. U ^ G . W 3-4 10-13 , – - 12-14 . S! - [15]. H 2010 . - 5 . (. 2). H 0+ - 2+ . V ! , .

X 2 - . U min max !! . . (! ), 5 . L, 13,4 29,5 26,2 7,4 9,8 l, 8,9 22,4 16,6 5,2 6,8 Q, 119 335 244 3,8 4,2 g, 94 286 194 2,6 3,4 Y 1,42 1,56 1,52 0,1 0,12 I (! ), 3 . L, 28,1 123 70,6 20,1 26,75 l, 22 101 56,8 16,69 22,14 Q, 0,18 19,45 4,543 3,13 4,89 g, 0,13 14,82 3,488 2,41 3,73 Y 1,41 1,92 1,71 0,11 0,14 Y 1,02 1,5 1,3 0,10 0,12 I ( G ), 4 . L, 62,7 87,3 74,15 10,4 12,25 l, 49,5 69,5 59 8,6 10,08 Q, 1,77 6,01 3,61 1,57 1,95 g, 1,52 5 3,01 1,36 1,66 Y 1,49 1,79 1,62 0,13 0,15 Y 1,25 1,48 1,35 0,17 0,10 I (! ), 18 . L, 104,9 108 106,45 1,55 2,19 l, 81,5 86,2 83,9 2,35 3,32 Q, 8,95 11,3 10,12 1,18 1,67 g, 7,33 9,41 8,37 1,04 1,47 Y 1,65 1,76 1,71 0,06 0,08 Y 1,35 1,47 1,14 0,06 0,08 T (! ), 4 . L, 64,6 121 88,07 17,23 21,17 l, 53,1 101,3 73,32 14,62 17,99 Q, 1,22 12,03 4,51 2,90 4,08 g, 1,12 8,98 3,48 2,09 2,95 Y 0,77 1,16 0,90 0,12 0,15 Y 0,66 0,86 0,73 0,06 0,08 X (! ), 5 . H SY. . 1 (33). 2012 87

L, 56,9 97,4 75,35 8,33 10,63 l, 46,1 77,2 62,05 6,90 8,62 Q, 1,11 4,98 3,07 0,79 1,06 g, 0,95 4,56 2,4 0,71 0,93 Y 1,04 1,52 1,24 0,13 0,16 Y 0,76 1,14 0,97 0,08 0,10 X (! , G ), 7 . L, 47,1 57,5 53,3 3,01 3,69 l, 38,2 47,5 43,3 2,28 3,03 Q, 0,60 1,3 1,00 0,22 0,27 g, 0,57 1,07 0,86 0,18 0,21 Y 1,06 1,45 1,21 0,10 0,14 Y 0,86 1,21 1,04 0,10 0,12 I ! . ^ T !! - - . Y ! - ! , ! . H 2010 . ! 0+ - 2+ . W 1:1. S ! , ! . U ! 0,066 /2, 0,012 /2. T : !, !; , ! Triplophysa strauchii (Kessler, 1874), !. Y ! !. H 2010 ^ 4 ! 0+ - 2+ . U ! 0,036 /2. X : , , -, -, ! ! !; - ! , , , ! !. H 2010 ^ 12 . 0+ - 2+ . X ! . W 1:1. U ! 0,054 /2. MQM] H ! .^ 4 : , , ! – ! . ! ! ! ! . H , . U G ! ! . Q 1. Jenkins M. Prospects for biodiversity// Science. - 2003. - Vol. 302. - P.1175-1177. 2. Revenga C., Campbell I., Abell R., de Villiers P., Bryer M. Prospects for monitoring freshwater ecosystems towards the 2010 targets// Philosophical Transactions of the Royal Society B-Biological Sciences. - 2005. - Vol. 360. - P. 397-413. 3. R H.U., J I.R. S // V – F-F: I - 1992. -X.5. - W.329-371. 4. X H.I., W F.W. F G // H . - 1995. -X.35. - H.1. - W.71-77. 5. U .. V . – R.: U! , 1966. – 376 . 6. Feunteun E., Ombredane D., Bagliniere J.L. Ecologie des poisons en hydrosystemes continentaux// Atlas des poisons d’eau douce de France. – Paris, 2001. - P. 36-55. 7. R H.U., J I.R. . V – F-F: S-I, 1986-1992. – X.1-5. 8. G F.F., X W.V. - . – F: SY, 1999. - 347. 9. ! V . . 4-, . X 1: L. ] 1: U.- F: DPS, 2010.- 324. 10. ! F (L) – F, 2006. - 520 . 11. ] .F. F F // H – 1996. – X.36. - 35. – W.597-608. 12. Scott W.F., Crossman E.J. Freshwater fishes of Canada// Fish.Res.Board Can.Bull. – Ottawa, 1998. – 970p. 13. Mecklenburg C.W., Mecklenburg T.A., Thorsteinson L.K. Fishes of Alaska – Betsheda, Maryland: American Fisheries Society, 2002. – 1037 p. 14. G! S.I.. S F.R. V ! – R.: X R , 2004. - 389 . 88 Y . . 1 (33). 2012

15. W F.. Salmo gairdneri Richardson – ! , ! !// V – F- F: I, 1992. – X.5. – W. 56-119. 16. G Z.F. Salmo mykiss Walbaum – // V – F-F: I, 1992. – X.5. – W.119-125. *** ^ ''' '' %' '. X- ''' ' ' ' ' '' ''' ' % '' : ' Diptychus dybowskii, X Triplopysa stoliczkai, '' T. labiata, () Paraslmo mykiss (Salmo gairdnerii). G %' '. *** The data on current composition of the Shlek River ichthyofauna as well as its vertical distribution was provided. Due to natural- geographical reasons and low anthropogenic load in this river, the favorable conditions for existence and reproduction of native fish species like scaleless osman Diptychus dybowskii, Thibet stone loach Triplophysa stoliczkai, plain stone loach T.labiata as well as alien Kamchatka steelhead Paraslmo mykiss (Salmo gairdnerii). The data on density of fish distribution and their brief biological characteristics were presented.

YJ 597.19 1 Z.. , 1 . R., 1 . J, 2 .. J TZ .LR 1XTT« », 2XTT «SV»,

U .L. - . U . RQ RTJ QJTO I ! .L 2011 ( – ! !, – !). U - ! 523 . - 1039 . . MQb QJTO TLJ H L (c) 2011 . [Q.W. G,1949; S G.X, 1970; U.U,1974; R! «V », 1986-1992, 5 ] 23 6 ! 7 : , , -, , !, , , , , , , !, , , , ( 1). J !!! - 14 . X 1 . L 3 1949-1992 .* 2011 .** Esocidae – _ 1 _ – Esox lucius (Linnaeus) + + . Cyprinidae – 2 U- Gobio gobio (Linne) + - 3 F – Abbottina rivularis (Basilewsky, 1855) (Dybowski)* - + 4 Q – Abramis brama (Linnaeus) + + 5 G – Abramis sapa (Pallas) + + 6 Y! – Alburnus alburnus (Linnaeus) + + 7 I – Rhodeus sericeus amarus (Bloch) + - 8 L – Aspius aspius (Linnaeus) + + 9 H – Chondrostoma nasus variabile Iakowlew + + 10 F- – Carassius auratus (Linnaeus, 1758) + + 11 - Carassius carassius (Linne) - - 12 W – Cyprinus carpio carpio (Linnaeus) ++ 13 – Leuciscus idus (Linnaeus) + + 14 U- – Rutilus rutilus fluviatilis (Jakovlev) ++ 15 – Scardinius eryhrophthalmus (Linnaeus) ++ . Cobitidae – 16 ^ - Cobitis taenia Linne +- 17 U! - Cobitis aurata (Filippi) ++ Siluridae – 18 W – Silurus glanis Linnaeus + + W. Gasterosteidae- H SY. . 1 (33). 2012 89

19 R! ! - Pungitius platygaster (Kessler) ++ W. Poeciliidae - 20 T! ! - Gambusia affinis (Bajard et Girard) +- . Percidae – T 21 W – Sander lucioperca (Linnaeus) - + 22 T – Perca fluviatilis (Linnaeus) + + Gobiidae – 23 G – Neogobius syrman (Nordmann) - + :23 18 17 U: * - , ** - ! XTT « » 2011 . H 2011 . , , , 2 ( ! ), !. H ! ! , , .L . H !, ! - . , [J ., 2010] !!! ! F, G, F!, ], X . W !, . L ! . H 2011 . . L. H . L 2011 . ! , !, !. U X- .L, , ! , ! ! . S ! ! (-) ! – , , !, ! ! . W, , , !! , ! 2 4 , , ! [V 5 ]. Y ! ! ! - 1,00 /3. H .L, . S ! !, ! () ! ! . H ! ! 7,27 /3, 2,14 /3 3,45 /3. W ! . L 2. X 2 (/3) ( ) 2011 . T 2011 . Q 0,15 2,140 U 0,30 3,450 - 0,124 Y! 0,40 0,962 0,10 0,003 W 0,04 T 0,105 G - 0,01 F 0,486 1,00 7,270 6 7 S .L !: - 2011 ., ! , !, !; 90 Y . . 1 (33). 2012

- , L , !! ! . W .L 20-50 /. ! ! ! !. H 2011 . . L ! - 64,1% - 31,9%, -1,8%, ! -1,5%, 0,4%. T (58%) (15%), ! 10%. ! ! .L 3. S , , . X 3 L, 2011 . H H, J, J H H Y- U- W- min- !!, min-max, !!, , max, min-max , % . Q 2-7 8 - 24 14,2 6,4 - 284 54 1,42-2,92 19,4-85,6 53 U - 4-6 14-16 15,4 54-89 71 1,97-2,19 1,2-12,4 63 6-8 18-22 19,5 127-218 167 1,84-2,06 49,7-93,2 75 7 21-22 21 229-231 229 2,47 17,8 50 L 3-4 18-26 23 165-255 204 1,42-4,03 19-176 50 U 5-6 19-22 20,8 121-190 159 154-1,91 1,4-8,3 42 Q 0,6-2,9 1,3 0,7-3,2 1,4 0,6-1,3 0,08-1,4 50 Y! 3,1-7,6 4,7 0,3-5,9 1,5 0,6-1,8 0,6-1,0 62 W! 5-9 13-27 18,8 65-646 268 2,58-3,74 69-345 83 K 3-5 20-35 28 410-902 656 2,1-5,1 126-637 50 _ 4-6 35-54 46 210-1041 540 0,84-0,87 12-38 66 W 4-6 54-86 72 1700-4500 3100 0,7-1,1 5,4-68,9 50 T 5-6 17-23 19,5 103-259 170 1,94-2,99 33-151 57 W 3-4 29-36 31 428-605 517 1,30- 1,75 50 U- 4-7 3,4-7,1 5,3 1,2-2,4 1,6 0,7-3,0 0,54-1,5 50 ! R! ! 4-6 3-7 4 1,2-2,8 1,4 0,8-4,4 0,3-0,5 50 G- 3 1,0 1,0 16 16 1,6 0,1-0,3 50 MQM] X , 2011 L 23 6 ! 7 : , , -, , !, , , , , , , !, , , , . G . T , ! ! !, , ! !!. Q 1. .G Q.W. V WWWV . – R.-Q., 1948 – 1949. 2. J I.R., R S.^., R .H. F V (W 1) //! SFS V. W! !, 2010. 33.- W. 36-49 3. ! F.. T . - R.: U! . - 1980. - 139 . 4. R H.U., J I.R. . V – F-F: S-I. - 1986-1992. – X.1-5. 5. S G.X. V . c: ! / IY. – F-, 1970. – 51 . 6. U.U., T X.V. T W, M H : W – Q.: I, 1974. – 137 . *** L '' '' ''. X- % ' %' ' '' % ' ' . G %' '.

H SY. . 1 (33). 2012 91

*** The data on current composition of ichthyofauna of the Zhem River are provided. By virtue of natural-geographic reasons and high anthropogenic load the breeding and development conditions of the fish remain difficult. The data on fish density distribution and their brief biological characteristics are provided.

YJ 597:625.21+639.214.(925.21.) . . U UTb _ ESOX LUCIUS L., 1758 TJTQ_ Q R. . U V -F \ "Q! J U"/ W - ! e-mail: [email protected]

U (Esox lucius L., 1758) . P. . T . T . U 2000- 2012 . T (Rutilus rutilus (L., 1758)) (Perca fluviatilis L., 1758), 3 11- (C. auratus (L., 1758)). J , . . . W (-) 1974 . ! ! ! \ . S 13 237 2, 1016 . 3. 11 214.3 2, 972.4 . 3 ^. T! ! !- 458 . U! - 2000 . 3/, ! - 1720 3/. Y ! ! 416 . H ! ! 25 [1]. H ! ! !! ! (Rutilus rutilus (L., 1758)), (Perca fluviatilis L., 1758) (Esox lucius L., 1758). \ ! ! . . W. RQ RTJ QJTO R 2011 . H 122 . 13 . . W. H ! [2, 3]. U [4]. W! [5, 6]. MQb QJTO TLJ _ . ! . ! - 0 55.6 % ( 1). S ! , ! ! !. S ! . (IY) 3 3, 8, 11 (HH) 3 29. U ! ! ! ! ! . ! ! .(r= -0.928). !! (r= 0.744). G ! . T, ! . _ !! ( ! W > 0.05) ( > 0.01). U - ! , , !, ! 2011 . , !. U , , 2000 . . IY 3 7-10 [7, 8] 2005 . . IY 33 7-11 HH 3 29 [9]. H 2000 ! : ! IY 3 7 – 11.1 %, . IY 3 8- 0 %, . IY 3 9 – 50 %, . IY 3 10- 25% [8]. X , ! , , . T - : - 33.3%, - 25.4 % - 6.1 %, - 64.8 % . J ! – 26.9 %. Y ! 8.1 %. G 92 Y . . 1 (33). 2012

, !, . ! ! 0.05 % ( 2). T ! ! ! . IY 3 11, ! 50 %. ! !! ! . H , ! ! ! . IY 3 11. X ! - . Q ! ! . S . IY 3 11 ! !. H ! . T ! . X 1 . . U: - Y Y !, % M±m M±m R- H . IY 3 1 11 27.3 0.97±0.03 0,09 0.91±0,02 0.07 . IY 3 2 3 0 0.89±0.05 0,09 0.84±0,04 0.07 . IY 3 3 13 46.2 0.92±0.01 0,05 0.87±0.01 0.04 . IY 3 4 2 0 0.89±0.002 0.003 0.85±0,004 0.01 . IY 3 5 3 0 0.89±0,04 0.06 0.85±0,03 0.05 . IY 3 6 3 0 1.00±0.04 0.07 0.91±0.02 0.03 . IY 3 7 6 33.3 0.92±0.03 0.07 0.873±0.02 0.05 . IY 3 8 8 50.0 0.99±0.03 0.08 0.93±0,03 0.08 . IY 3 9 1 0 1.06 - 1.01 - . IY 3 10 13 15.4 0.96±0.03 0.12 0.88±0,03 0.11 . IY 3 11 24 50.0 1.11±0.02 0.08 1.02±0.02 0.08 . HH 3 29 11 55.6 0.94±0.02 0.07 0.88±0.02 0.06 . c 1 0 1.06 - 0.97 - T : 99 34.3 0.99±0.01 0.11 0.92±0.01 0.09 I 2+ 5 40.0 0.88±0,03 0.06 0.76±0.02 0.05 3+ 26 50.0 0.95±0,02 0.12 0.90±0.02 0.10 4+ 46 34.8 1.02±0,02 0.11 0.95±0.01 0.08 5+ 16 12.5 0.96±0,02 0.07 0.90±0.02 0.07 6+ 3 33.3 1.03±0.07 0.11 0.96±0.05 0.08 7+ 2 0 1.00±0.08 0.11 0.96±0.10 0.14 8+ 1 0 1.06 - 0.97 - U 48 29.2 1.01±0.02 0.11 0.94±0.01 0.09 45 40.0 0.97±0.02 0.10 0.91±0.01 0.08 W!- ! . IY 3 1 1 0 0.91 - 0.85 - . IY 3 4 5 20.0 0.93±0.02 0.05 0.88±0,02 0.04 . IY 3 8 13 23.1 0.95±0.03 0.11 0.89±0.02 0.09 . IY 3 11 4 75.0 0.96±0.02 0.04 0.88±0.01 0.03 T : 23 30.4 0.94±0.02 0.09 0.88±0.01 0.07 R ! 45 1 . IY 3 8 37.8 550 ., - 4 ( 2 ! !) 39.6 713 . IY 3 11. ! 2000 . [7, 8] 1 , [10], 72 ! 20 ! 5-8 . X 2 U . . H T H J! , % T! , (%) - Gammarus lacustris 2 0.07 0.009 0.0014 Paramysis lacustris 45 0.315 0.039 0.14 H SY. . 1 (33). 2012 93

Ischnura elegans 1 0.03 0.004 0.0003 Carassius auratus 16 203.0 25.4 32.79 Rutilus rutilus 10 266.0 33.3 26.85 Abramis brama 1 49.0 6.1 0.49 Esox lucius 1 64.4 8.1 0.65 Perca fluviatilis 18 215.0 26.9 39.07 !-! Carassius auratus 2 94 22.6 19.85 Rutilus rutilus 2 56 13.6 11.83 Esox lucius 2 119 28.7 24.46 Perca fluviatilis 3 143 37.9 44.71 MQM] X , ! . . W. H ! - ! . H . IY 3 11 ! !, !!! . U, , ! . HH 3 29, . , , ! ! , ! . W, . . W 2000 . [7, 8] 2005 . [9], ! ! . U ! . H , (!, ) - !, - [8]. U , , !. W 2012 !! , ! . . W. T, ! !, , . ! ! , ! ! , !- !. ] - , ! !. T ! !! ! ! ! ( ). T! ! . Q 1. H. S. F (Actinopterygii) ! // H IY . K. F. G, !: G!, ! , -2011. -3 3. -W 47-56. 2. U H. Q. (.) ! . –Q.: ISTV, -1953. -27 . 3. S I. H. c! . –R.: H! , -1974. -376 . 4. V Z. W., W F Q., U I. Z., X H. Q. U . Y!.// c! . –R.: S, -1993. –W. 66-143. 5. L Q. F. U!! !. -R.: S, -1991.- 271 . 6. U S. F. G!. –R.: RIY, -1970, -367 . 7. Krainyuk V. N. Rules for sustainable fishing in the rivers and lakes of the Karaganda region (Final report).-Karaganda, -2001.- 78 p. 8. H. S., Z. H. ], ! Sander lucioperca (L.) (Osteichthyes; Percidae) - W // Tethys Aqua Zoological Research, – 2002. – X. 1. – W. 108-114. 9. T - ! !! . V: H : T SV. H.S., T Z.H., R Q.H./ VF\ "Q! J U" - 3IV 0105V00071 - ,- 2005. -198 . 10. H. S. S ! Esox lucius L. (Osteichthyes; Esocidae) ! \ . // . . \SX, -1995, 3 86-95, 2 . *** . % ''' (Esox lucius L., 1758) ' %' '. M ' ''' '' ''' ' . L '' . 2000-2012 ' '' . F '' ' (Rutilus rutilus (L., 1758)) % (Perca fluviatilis L., 1758), 3 11 ' (C. auratus (L., 1758)) . '' ' %'''' ''' . *** Actual materials on a feeding and state of nourishment of a pike (Esox lucius L., 1758) in reservoirs of the K. Satpaev's channel are resulted. The variability of feeding parameters into population of this species from investigated reservoirs is discussed. Age dependence of decrease in intensity of a feeding is marked. At comparison chronological groups from 2000 - 2012 years on distinctions of feeding intensity not revealed. As the basic objects of a pike feeding in channel reservoirs serve roach (Rutilus rutilus (L., 1758)) and perch (Perca fluviatilis L., 1758), in reservoir 3 11 - goldfish (C. auratus (L., 1758)). The conclusions about satisfactory character of a food of a pike in reservoirs, and as about necessity of strengthening of a catching press on this species and its feeding object's populations become.

94 Y . . 1 (33). 2012

YJ 597.5:59.31 Q.. TQTI] ZT UTQR T MTTM TM -Qb Q! G- S F V « , , , , » F.H.^ , N-P, , , . In the article, there is given an analysis of reasons and problems resulted after human activity in Issyk-Kul Lake Basin, in particular after introduction and acclimatization of some fishes into this drainless water body. Also there are considered positive and negative consequences of any works on acclimatization, and some measures are recommended which can promote to conservation and renewal of aboriginal fish populations. T -, ! , ! . K! , !. H !, WWWV, -!. R , , ! [1] ! () , ! - [2,3]. T . - ! , , ! , – , , . – – (! ! , ) ( , ) ! !, ! . RQ RTJ QJTO H , ! ! , ! , , , , . MQb QJTO TLJ ! . U ! ! , 70- !. T, F , G . T - - ! . S!! 30- ! ! . - ! [4], ! ! !. ! ! ! , !! ! ! -, , , , , . W! , «» - !, ! - , -!. ] . T , ! , . J , V. S ! «» - ! , «! ». c ! 1 . X 1 J - (J J R PV) I 1965 32 1257 20 26 1335 H SY. . 1 (33). 2012 95

1970 19 803 116 5 5,5 7 956 1975 77 686 112 47 2 3 927 1980 36 224 36 40 5 1,5 2 344 1985 14 86 22 13 23 15 1 174 1990 32 163 32 18 21 7 5 278 1995 123 2000 51 2005 13,5 2010 3,4 2006 3,5 2007 53 2010 156 , ! - , . W – , ! - – , , . G ! ! ! !, ! ! 15 ! ! , J ! ! (.). U: S ! , ! -, ! ! , !!: 1. , . 2.T !. 3.V ! U. 4.Y . 5.V . 6.! ! , . 7. G ? H , , . F.H.^ 1979 ! ! -: ) S !! , . ) H - . ) W , , . ) M . ! : 1) W «M a-», . 2) Y - , ! . 3) M -. 4) Y - 300. . 5) H . 6) W , . 7) V . 8) W ! . 9) V - - . TJ 1.K «» - ! , , : , , -, . 2. F! – . 3.U ! , ! -, ! ! . 96 Y . . 1 (33). 2012

4.S ! - !! ! ! , ! , ! ! . Q 1. X .F. U -. H F WWWV, 1961, 3 2, W. 55. 2. F.T., ! Q.F.. U ! ! . // G ! W F , 1978, - , , W.340-342. 3. F.T., X F.., ! Q.F., U R.H. // A! WWWV. 1972, , W.43-45. 4.U .F. V . , - , 1990,W. 95-96. *** G a' ' %' % ' , ' '', '' % , ' ' % . *** In the article, there is given an analysis of reasons and problems resulted after human activity in Issyk-Kul Lake Basin, in particular after introduction and acclimatization of some fishes into this drainless water body. Also there are considered positive and negative consequences of any works on acclimatization, and some measures are recommended which can promote to conservation and renewal of aboriginal fish populations.

YJ: 561.26.016 Q.U. Q, J .. TURM\ TT UT\T QTQQ UQ UTQbMT ] T ] TQTI] JTT TRR . -, F, T - . S , . X . U . . H ! ! !!! ! ! , . G! !, !, ! , , , , !. \ ! Chlorella vulgaris Spirulina sp . J! ! : 1. Y ! Chlorella vulgaris ( X-4) Spirulina sp 2. U Chlorella vulgaris ( X-4) Spirulina sp. 3. T Chlorella vulgaris X-4. 4. V . RQ RTJ QJTO J! ! , 100 26-28 \!. U ! 30 . J! ! M, X!, K ]-10. J! !! ! ( ), . T 5 30- !. H I!. J! ! ! , 105 1 , CaCl2 , !! ! ! ! . W ! 0,0001. J! ! ! (V), ! , V= 2/1, 2 – ( ! ! ), 1 – ( ! ! ). U ! , . J! !! !! , 540 590 , .. . H SY. . 1 (33). 2012 97

F ! . 45 °W ! !. W ! 70 . U ! . W ! . ! GYH. T ! , . U! ! 0,1 R CaCl2. U ! c 420 . U ! , U ! , . MQb QJTO TLJ V ! ! W . W . G! , 20 , ! 200 , !. S ! ! ! Spirulina sp !. J! ! ! , ! M 30 24 . ] ! ( 1) ! ( 2). U ! 15 . H ! . T! ! ! ! 0,01 /.

1 R " M 30 24 .

2 " M 30 24 . – ! ! , ! , , ! ! ! . Y ! , ! . T . M ! ! !. R ! ( ) !!! ! . S Chlorella Sp F . J! I, X!, ]Y-10, K 30 ( 23-25 °W 2000 ). X ! ( 590 ). ! ( 3). 98 Y . . 1 (33). 2012

3. J Chlorella sp. -4 Ue , ! ! , . S X! ! ! , ! 110 7 /, ! 25 . S ! ! ]Y-10 , 3,4 10 7 \. . X, 15- ! 5,9 10 7 /. U ! 20 ! !, «». T 25 ! 2,5 , 14,7 10 7 /. S 30- ! H !, ! - ! . MQM] F , ! . R! ! 20- !. 20- , ! ! 33,3107 /. T ! !!! , 20-21 . X , , ! ! I Chlorella sp. T , ! . S X-4 Chlorella sp. ^ . V ! 1. X 1 -4 Chlorella sp. DL-- " --! DL - DL -! –- L - DL -! L - DL - DL - DL - DL - ! DL - DL – L - ! DL -!- , , ! . c ! ! ! . Q 1. Beijerinck M.W. Culturversuche mit Zoochlorella, Lichenengonidien und anderen niederen Algen // Bot. Zeit., 48, 47, Idem in: Verzamelde Geschriten van M.W. Beijerinck, 1921, 3 2. 2. G S.. W ! «Chlorella vulgaris V 3 W-111»: . V. ! 3 2176667 / S.. G, R.H. ; G . 3 34. – 2001. 3. F F. U , ! ! ! // - ! Y. X: Y WWV, 1976. – W. 92-93. 4. Beijerinck M.W. Culturversuche mit Zoochlorella, Lichenengonidien und anderen niederen Algen // Bot. Zeit., 48, 47, Idem in: Verzamelde Geschriten van M.W. Beijerinck, 1921, 3 2. 5. G S.. W ! «Chlorella vulgaris V 3 W-111»: . V. ! 3 2176667 / S.. G, R.H. ; G . 3 34. – 2001. 6. F F. U , ! ! ! // - ! Y. X: Y WWV, 1976. – W. 92-93. F H.R. V CLORELLA. R!, , / H.R. F – Q.: - «S», Q. ., 1975. – 110 . H SY. . 1 (33). 2012 99

8. R F.R. / F.R. R, X.X. X. – X: YWWV, 1984. – 136 . 9. Q F.U. X! ! ! . Y ! / F.U. Q, H.S. R, S.I. G // Y . – R.: - SQ. 1997. – 184 . *** L '- % ''' ''' % . [ ' '' , % '' % - ' . J %'' %' ''' ''' %. ^ %' % . G ' ''' ' % . *** Described work on optimizing of breeding processes of blue-green and green water-plants on the example of spirulina and . An optimal environment for growing of present stamm of , and optimal concentration of cages and duration of growing is found for spirulina. The increase of culture is also well-proven the experienced way at the permanent bubbling by oxygen. On present methodologies amino acid composition of and amount of clean albumen was certain in spirulina. All data were treated and presented in a table form.

YJ 65.012:504.06 (262.81) . R T RTU T] IUU UT UT, UQ TTQ T_ T UOTIT RT U Ic-UVTTS FWUcT, [email protected]

, U UVTTS-IZ PFUPT, P . U U P . U PFUPT (YGU), . , !!! , !. W ! ! (! ! !, ! , ) (!, , ! , ) [U ., 2005; TDA, 2007; R, 2009]. U , ! !, 0,3 W/ ! ! 7,5 / [! M, 2007]. G , ! ( ., 2008) ! ! !, Mnemiopsis leidyi, . H ! . H ! ! , , !!! ! ! ! ! . U !, ! ! [Ward et al., 2002; Royal Society, 2003]. W , !! , ! ! . S! ! , . M 95 % ( 65 ) c 39 . 0,6 . 2010 [ , 2010]. S ! ! ! ! !!! ! F-] ! Mnemiopsis leidyi. H 430 . . 2000 . 135 . . 2004 . [Mamedov and Daskalov, 2007]. H ! ! 6 ! , ! !, IUCN/RWTU. G , RWTU (2010/IUCN), 85 % !! !, ! RWTU. Y! , ! ! , 100 . . , , !. J , , ! ! F V V . U , ! ! 2 7 . W^F [TDA, 2007], ! ! ! . 100 Y . . 1 (33). 2012

W !: U!, ! ! W , , ! ! !! H - (Mnemiopsis leidyi), , , , ! ! M! () ! ! S !! W ! H ! , ! , ! , . H , ! U c U (cU) !, ! ! 10 Ic, FWUcT (2009-2012). H !! ! ! ! ! , W (WUJ) (2006), . T !!, ! ! «FWUcT», !! !: ! ; , ; ! ; ; ! ! !, !! . H FWUcT ! : 1. Y! ! ! ! ( 1) 2. Y! ( 2) W ! ! «FWUcT» Y ! (YGcU), ( ), !, ! - ! U . U , !! ! ! , !. H YGcU , !. , YGcU , , , , , , - ! (1990-2010). U !! , ! !! . S , ! , ! !, ! ! ! Ecopath (c) Ecosim (c) (EwE) ! W !. W , c c !!! ! () , ! . , c c ! !, , ! ! ! ! ! !! ! . T ! ! !! . H ! ! - H SY. . 1 (33). 2012 101

! ! , ! ! ! !. G , ! ! ! ! - !. U ! ! ! (, 5, 10 20 , ! !) ! , !!; ( , ! , !, ); F ( F). H ! ( !) - . H ! !! !, , . T ! ! !! , ! ! !! ! , !, ! . , ! , !, ! . X , ! ! FWUcT, !, !! !. , ! ! c c ! c W !. W ! , ! , , W !, ! . U ! - ! ! ! (! ) ! !, ! ! ! ! . V ! , !!, ! ! , ! ! , ! ! ! . F , , ! ! !!. H , ! ! ! . U , ! ! , ! , (!) , , . H ! ! , , , ! ! . U , ! !, ! ! ! ! U !, , , ! ! . Q 1. ! R.I., M .W. // R. R - «U ! ! ! ! ». – F, 2007. – W.56-57 2. F. R. U ! H- / F.R. , T. F. ], .I. J, .S. H // R - « ! ! ». – F, 2008. – W.99-106 3. R c.H. W, ! . // R - «U ! ! ! ! ». – F: SV, 2009. – W.143-147 4. U, I. S. W ! ! /I.S. U, V.R. R, . H. R // U . R. I. !. – R.: S, 2005. – 356. 102 Y . . 1 (33). 2012

5. Daskalov G. M., Mamedov E.V. 2007. Integrated fisheries assessment and possible causes for the collapse of anchovy kilka in the Caspian Sea. – ICES Journal of Marine Science, 64: 503-511 6. Ward, T., Tarte, D., Hegerl, E. & Short, K. 2002. Ecosystem-based management of marine capture fisheries. World Wide Fund for Nature Australia, 80 pp. 7. Royal Society, 2003. Environmental effects of marine fisheries. Policy Document 18/03 (http://www.royalsoc.ac.uk/policy). 2010. IUCN, Gland, Switzerland. http://www.iucn.org/?4928/Sturgeon-more-criticallyendangered- than-any-other-group-of-species Accessed 20/03/20. 8 Transboundary Diagnostic Analysis for the Caspian Sea (TDA) Revisit (2007). Caspian Environment Programme// (http://www.caspianenvironment.org/newsite/Data-MajorDocuments.htm) *** R X'' % ' '' P ''' ' UVTTS-IZ PFUPT P '' '' '- . ' '' PFUPT '' %' ''. *** The activities of working groups comprised of national experts from all the Caspian littoral states of the UNDP-GEF CASPECO project aimed at restoring depleted fish stocks of the Caspian Sea on ecosystem-based management of aquatic bioresources and the strengthening of the regional environmental governance are discussed. The project supports the Caspian countries efforts to further halt the decline in bioresources and to restore depleted fisheries of the Caspian Sea. The main outputs of CASPECO’s working groups on ecosystem-based management of bioresources (EBBM), the inventory of fish spawning grounds and fish passage facilities are provided.

YJ 565.32 .. R, .U. RTITQ JR TL \ R (ARTEMIA sp. ) M TM TIT M S . -, e-mail: Nadezhda.Shakaeva@ mail.ru

P. U , . L ! (Artemia sp.) !!! ! , (I.. 1, F.H.W 2). F! ! , .. , . H ! , ! , . T ! !! ! , !, !, . V !, . H ! !! . RQ RTJ QJTO J ! 1991 R W W- , – – U . U !, ! ! , ! : 100 % !. – 10 .H ! ! ! (.H.G ., 2). H ! ! ! !. H ! . MQb QJTO TLJ ! ! - 6,1 34,8 %. U ! ! 100% ! , 58-76%. H ! ! ! 100 % !. H ! ! , ! (G..R, S.U. ^ 4). H ! !, ! !. , ! ! ! ! R - 75,9%. H ! ! ! 2-3 . H 2012 , .. ! ! W , 2008 . H ! 22,64%, R 5 %. , ! ! ! ! .

H SY. . 1 (33). 2012 103

X 1 (%), J R 07.05.2001 75,69 63,82 49,13 70,08 10.05.2002 62,96 57,33 52,28 63,30 17.05.2003 59,89 54,47 47,88 54,79 10.06.2008 35,43 19,27 22,39 32,25 08.02.2012 5,2 20,29 2,1 22,64 Q 1. I. . . –R:, V! . 1981. W. 7-13. 2. W F.H. T ! ! ! . – R:, HSVT, \SSXcV, 1984. 63 . 3. G .H., I K.K., ^ M.., V ( Frtemia salina L.).-R:.1980. W.1-30. 4. R G.., ^ S.U., H ! . Tethys agua zoological research, 1-Almaty, Tethys, 2002., W.183-186. *** R ' ' '' '''' ' %' '. F ''' '''' ''. *** In the article presented data’s by long experience of germination eggs of artemia from for lakes of north Kazakhstan. The data’s indicate that on long standing artemia embryo’s can viability.

YJ 639.3 R .. T\ TT RTQTJ TIT T UT TTJT-TQTI]R URR TR]R UTMQR T UTQ TRQ URQbR TRR XTT «S !»

T . S , - , . S (1,95) 32. S 33 2,1 3, 40% . U , , , , , , . N , , 32 , , , , - . . H ! ! ! !, ! . H ! !, ! , . T ! ! [1]. M !! ! ! ! ! ! ! - . I !! ! ! . Q , ! , , . . c ! . V ! ! ! [2]. H ! , XTT «S !» XTT «S » JIU « » ! !, . T !!! ! 104 Y . . 1 (33). 2012

! !. H !! !. RQ RTJ c ! , - !. U 30 . J! 10 . W 8-10 . 50 ./2. ! !. H ! : 18,40W; 7,65 /; S 7,17 8,04. Y ! 30 . V (7 – 8 /) 9,5 /, [3]. J! ! ! ! , !: 1 10 . c 3 1, 2 XTT « S ! », 3 3 – JIU « ». V 1. X 1 H 31 32 33 R ! 49,0 45,97 45,97 46,0 ^ 16,0 20,0 13,0 10,0 J 6,0 1,0 - 3,0 U! 5,0 5,0 8,0 6,0 T - 5,0 5,0 2,0 U 1,0 1,0 1,0 0,5 L 3,0 3,0 3,0 3,0 R 4,0 3,0 3,0 3,0 R !! 5,0 - - - R !! 5,0 - - - R ! 1,0 - - - W 5,0 5,0 5,0 5,0 \ - - 1,0 - G - - - 1,0 U- «G» - 0,03 0 ,03 - I ! - 10,0 15,0 - GFH - - - 20,5 : 100,0 100,0 100,0 100,0 W ! , : 3 1 – 10 %; 3 2 – 15 % ; 3 3 – GFH. W , ! 33, 2. X 2 - + S , U, % 49,8 H: /100 H- 2,5 W 110 J 400 H1 1,5 H6 2 H12 6

H SY. . 1 (33). 2012 105

MQb TLJ V- , , 3. X 3 2009 H U T V H , , , , /2 , /2 % 9,13±0,85 2,37 113,1 0,6 1,16 3 1 13,0±0,43 2,26 154,7 0,87 1,42 3 2 13,4±0,47 1,95 148,8 0,9 1,5 3 3 12,5±0,51 2,13 147,0 0,84 1,4 S (1,95) 32, !. S ! 33 2,1 3, 40% . W , 32, , . G 4. X 4 U 31 32 33 T ,/ 37,1±3,8* 38,1±0,3* 39,1±1,1* 40,1±5,5** I , / 3,7±0,3 4,9±0,2 3,27±0,2 6,14±0,3* , / 2,41±0,13 4,3±0,18* 2,56±0,15 4,63±0,14* X, / 199,3±18,6** 343,5±7,6*** 176,8±2,5* 329,6±6,2** _! , U/L 99,23±1,08* 132,6±6,4** 7,62±2,9*** 2,46±1,4*** U - : -<0,05, - <0,01, - <0,001 U 31,2 3 ! ! 2,7% 8%. W 31 33 32% 66% . U 32 ! 13%. S 78%, 6,2% 92% 31, 32, 33 . W 31 33 ! 72% 65% , 32 ! 12%. U ! 32 33 1202% 3933% , ! 24% 31. Y ! , , , ! , . Y 31 , ! ! ! , !, 32 33, ! ! . U ! ! ! , , , ! ! !, 33 31 ! ! . , ! , !! ! , . S! ! 33. Y 31 3, . MQM] , 32. X , ! : , . U !, ! . H ! , !!! . 106 Y . . 1 (33). 2012

! , , 32 , !!! , ! , !, - . H ! ! ! . ! . Q 1. I.I., J W.., Q! X.H. T !! !// R 1 R ! . -WU., 1996.- W.108. 2. Q K.X. J .//U , .102 (496), -2007. 3. H Q.R., F.F., U X.S. . G .-F: «F », 2010.-80 . *** R %' %' . G '' % ''' '' . F % , ' '''' '' ' %' %' . *** In this article described an experiment about the feeding the russian sturgeon by home man-made foods with various additions. Given estimation of the physiological condition of young russian sturgeons arcading to fish-breeding and biological parameters and biochemical parameters of blood. By virtue of got database found the food with addition, which more balanced and nutritious go the sturgeon fishes bred in basius.

YJ 639.3 R .., .. QJT Q TRT TQTI] R JTR T TT U OTTO TQTI _ XTT « - !», . F, [email protected]

. U , , , - , . H !! ! , , , , , !, ! , .. ! , , !, ! ! . T! !!! !. W ! ! . Q ! 9-12 , ! , , . S !!! , !, ! , , ! !. !!: , , , . S, ! (50 / ) H12 100% ; , ! , , , , . S ! ! H12 [1]. T ! !!! – !. , !! ! !, ! , W K, , . c , , .. . H ! , ! , , ! !. ! – .W! , . K" . T !! ! ! , ! (, !), V. \ ! ! , ! , ! , !! , , ! . X ! ! , ! , - , ! . c - , , , - , , .. H SY. . 1 (33). 2012 107

G ! ! ! , ! -!. T ! ! ! . F , (H, , ), !! ! (V!) [2]. RQ RTJ QJTO M ! 2009-2011 3 , , ! : 2009 - ; 2010 – ; 2011 - . c SH (- !). R . V ! 60 ./2. ! !. ! , TX-6. I ! : ! – 18,2-18,90W, S – 7,5-7,9, ( !) – 7,6-8,0 /, [3]. J! , ! ! , ! 1 10 . MQb QJTO TLJ H , 2009 8 , ! . M 30 3 3 !! 8 21,1 . U , 3,37 0,3 /2 ( 1 2).

1,4 14 1,2 12

10 0,8 8 0,6 6 , / 0,4 4 0,2 2 0 0 3 3 1 2 H , 2010 85 , ! . M 30 3 1 !! 86,32 143, 25 . U , 10 1,11 /2 ( 3 4).

7 60 6

50 5 40 4

30 3

20 , / 2

1 10 0 0 1 1 3 4 108 Y . . 1 (33). 2012

H , 2011 87 , ! . M 30 3 3 !! 85,84 140,9 . U 1,7 , 0,7 /2, ! ( 5 6).

57 4,8

56,5 4,6

56 4,4

55,5 4,2

55 , / 4

3,8 54,5 3,6 54 3 3 5 . 6.

MQM] X , ! , . V, , - , ! , , ! ! - . Q 1 ] R.W., I K.H., ] Z.S. H . -R.: ISY. V, 2004.- 136 . 2 T.H., I U.U., V S.S. c ! ! –//U : R. . . .- ../ R , 2005.-175 . 3 I .., I..\ .- Y. 1984.– 44 . *** R ' % '' ' '' % ' ' %'' ' %' ''. M ' % ' ' ' '' '''' '''. *** In article results of experiment on influence of forages with biologically active additives on growth and an exit russian and siberian sturgeons are reflected at basin conditions of cultivation. the received results have shown advantage of mixed fodders with additives, such as clorella and a vitamin additive, at feeding of sturgeon fishes that was reflected in improvement of rybovodno-biological indicators young sturgeon fishes.

YJ 639.371.2 .T. RQ UT _M QTT UTQQb R TTR T - ! «FH!U » e-mail: [email protected]

Acipenseriformes - Polodon spathula 3 . Polodon spathula Acipenser gueldenstaedtii. U . W ! , , ! , ! " . V !! ! . W , ! ! . c [1]. [2]. Y ! ! [3-5]. H ! ! ! ! H SY. . 1 (33). 2012 109

! [6-8]. H ! ! , , !, [9-14]. S !! (Polodon spathula W.), ! ", ! , !. H ! , ! , ! ! . U , !, , !!! ! ( ), Spongilla lacustris [15-17]. G ! . H ! 3 , ! [9, 18-21]. T, ! ! ! ! 10 . \ ! (10 ) . M : - ; . RQ RTJ QJTO V 2007 . IYU SU\ «GTW». H ! (Acipenseriformes). F ! F ! – " (Acipenser gueldenstaedtii .). T " – 3780 . 694 . . U! ! 62,0 . . . H! 51 !. H 2,02,00,7 . U! 5 . ./2. S ! – 40 ! ! [22- 25], 21,5 0W. . H! – 1100 , ! 20-25 . U 0,2 , ! ! 43 . . H ( 1,0 ). MQb QJTO TLJ c ! . S 34 , , ! - 17 . H !! 2,8 , 7,1 ( 77 ). U – 6,3 ./2. S ! , , , ! !. U . ! . H , " , ! ! [19, 26]. ! , 2 ! ! ! . W!! ! 21,4 0W, – 22,8 0W. W!! ! ! 8,4 . R , 1. F , . U ! 3 ! , ! 72,2%, !! ! 1,6 [23]. H - . T, . c [27-30]. T ! . X , !. ] . H , , ! ! .

110 Y . . 1 (33). 2012

X 1 - H! H W!! , T, , !, % % , % 1-8 0,03 36,73 63,27 0,00 18 0,24 4,30 44,00 26,17 25 0,55 2,18 43,09 0,00 32 2,35 5,01 23,03 54,31 39 5,03 16,24 20,16 7,41 51 10,00 5,24 17,89 23,02 MQM] X , ! !, ! ! [6]. U , .. ! ! . ! !, , ! . T ! ! ! ! "! – 17 " !! 0,4 . U ! - !. U ! !. J! ! , . S . S ! , ! . Q 1. ! V.U., V I.., U J.W. U, , H- . R.: X R. - 2007. – 242 . 2. S W.T., U H.. ! ! // U : ! ! . R.R. : . . . . . . . ! . (, 20-25 !! 2010 .). : - . . -. - 2010. – W. 436. 3. R I.I. ! ! V // F ! V: ! ! , ! , !. R R (27-30 !! 2011 ., V--J). V-/J: ZS\ VFS. - 2011. – W. 13-18. 4. L F.H. V // V : . R! -! !, 10-11 !! 2011 .: / ISY HSV V. R.: - VIFY-RWF . .F. X!. - 2011. – W. 11-14. 5. W I.K., S S.U. F ! ! // V : . R! -! !, 10-11 !! 2011 .: / ISY HSV V. R.: - VIFY-RWF . .F. X!. - 2011. – W. 24-27. 6. F c.F., S W.T. U ! ! // c ! : R ! ". F: «F . - 2010. – W. 90-93. 7. S W.T., Q F.R. R! ! ! ! ! // c ! : R ! ". F: «F - 2010. – W. 113 – 115. 8. X F.R., S W.T. ! // c ! : R ! ". F: «F . - 2010. – W. 69-71. 9. F H.H. H F. . . . ... F: SV. - 1997. - 28 . 10. H Q.R., S W.T., F H.H. W ! YMH // «F : ! !»: III R - . F. - 2004. W. 299. - W. 27-31. 11. S W.T. H 1,4 // U ! ! ! U! : R 2 R 31 ! 2004 ./ F! . c: IY. - 2004 . - W. 82-84. 12. S W.T. T ! ! Z V // S !: R . V /J.: V. -. - 2004 . - W. 131- 134. 13. S W.T. U ! Polyodon spatula V // U! : !, !. W , !" 60- W. ": «Eco-TIRAS». - 2005. - W. 47-50. H SY. . 1 (33). 2012 111

14. S W.T., F.U., J! F.S., W K.S. M // F . K , ] 11. !. S!. ! ! !. X 1- R (6- R ). W: W IXY. - 2005. – W. 124-126. 15. S W.T. H! c // V : . R! -! !, 10-11 !! 2011 .: / ISY HSV V. R.: VIFY- RWF . .F. X!. – 2011 . – W. 133-136. 16. S W.T. S – ! // F \ H K: ! // II NACEE (W \ \ H K) , , 17- 19 !! 2011 . : Pontos. - 2011 . - W. 197-200. 17. S W.T., K W.F. Y ! ! ! // V 3 105128 23.12.2010 ., 10.06.2011 . U S W.T. 18. H H.., K Q.H., R K.F. G ! ! (Polyodon spathula (Walbaum)). R.: ISY «V». - 2003. – 344 . 19. " F.Z., ^! F.H., ! F.Z., G Q.F., H... X! ! 1-2 !. R.: HSUV. - 1994. - 15. 20. R K.F., H H.., K Q.H., ] H.I., ! H.F., G R.H., W T.H., H.K., X.F., G c.H., T.G. T ! ! // V. -, , ., !: F. R.: HScV. – 1996. - . 1.- 67 . 21. Arkhangelskiy V.V., Sudacova N.V., Nekrasova S.O., Pismennaya O.A. Technology of Paddlefish Seeding and Commodity Rearing with Application of Industrial Methods, Including Recirculation. Proceeding the Sixth Imternational Conference on Recirculating Aquaculture July 21-23, 2006. The Hotel Roanoke & Conference Center Roanoke, Virginia, U.S. Virginia: Departament of Food Science and Technology. - 2006. - V. 448-458. 22. S W.T. U ! ! // H . – 2007. - X. 8. - 31 (29). - W. 130-137. 23. S W.T. U ! (Acipenser Stellatus Pallas) (Polyodon Spathula Walbaum) ! . F ..., F: SV. - 2006. – 24 . 24. S W.T., F.U. H ! // H . – 2006. - X. 7. - 34 (28). - W. 644-654. 25. S W.T., F.U., Q Q.. U - ! // H FIXY. – 2006. - 3 6 (35). – W. 245-253. 26. H Q.R., F.U., _ X.I., U X.S., X! H.H., F.F., F H.H., W S.H., F W.W., K.F. X VI / U . S.H. W. R.: HSVT. - 2006. – 100 . 27. U H.V. G ! . R.: cRcL FS WWWV. - 1972. – 230 . 28. U! .. I // S !. R.: cRcL FS WWWV. - 1982. – W. 140-152. 29. Wojtenek W., Pei X., Wilkens L. A. Paddlefish strike al artificial dipoles simulating the weak electric fields of planktonic prey // J. Exp. Biol. – 2001. – 204. - 3 8. – V. 1391-1399. 30. Wojtenek W., Pei X., Wilkens L.F. Detecting artificial sinusoidal electrical signals by the paddlefish, Polyodon spathula // 5th Int. Congr. Neuroethol., San Diego. Calif., Aug. 23-28, 1998: Program and Abstr. – San Diego (Calif.). - 1998. – P. 12-15. *** Acipenseriformes - ''- Polodon spathula 3 . F - Polodon spathula - %'' '' - Acipenser gueldenstaedtii . U - -. *** Improved breeding Acipenseriformes biotechnology now allows juveniles to increase Poluodon spathula in pools over 3 g. This paper presents the experience of growing a large sample of young Poluodon spathula in polyculture with Acipenser gueldenstaedtii. Preliminary data on the cultivation of polycultures in industrial environments.

YJ 591.8:576.895 .. , .. , .R. L, .., .. T ITUTQTI] MR TI QI TMTO U UMTO M . -, e-mail: Sabyr. [email protected] Rhabdias bufonis , Saplometra cylindracea (, , , , ) , S, - . N, 20 " P 2011 . N 12 (11 , 1 ). U . U, , Rhabdias bufonis, . I , . V . X, , , , . U . U ! , !! ! . ! ! [1]. G , ! ! !, , ! ! [2]. S !!! Rhabdias bufonis [3]. J !!! , 112 Y . . 1 (33). 2012

!. W , !. J ! ! , ! . H ! ! . U ! , ! !. , !, ! ! . Q ! !! . H ! [4]. S , , Saplometra cylindracea[7]. \ ! Rhabdias bufonis ! ! . RQ RTJ QJTO T ! ! !, ! S, - . , 20 " . U , ! . S! ! , ! 25 100 . 12 ! (11 , 1 ), 45,27±5,16 – 81,5±3,2 , ! ! , , , , . T V [5]. G ! . U R [6; 7]. MQb QJTO TLJ H !. U ! !, , ( 1). U Rhabdias bufonis, Rhabdiasidae. X , !! 0,8 7 . W ! (). U , . V! ( 1 F). H " ". H " " , . W ( 2). Y ! ! . U .

F G 1. Rhabdias sp. . – ; – W! , 2 !. W . Q! , , . R . U ! . W ! , !. U ! !, !! . H , , ! , . U . X ! ( 3 F). S ( 3 G). U ! !. X! H SY. . 1 (33). 2012 113

, ! , ! , ! ! !.

F G 2. Rhabdias sp. T () () H ! !, , " . U , , ! . X . S , ( 4 F). . , , !! ! ! . X ! , . H , ( 4 F).

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F G 4. U . - U , ; – , , . T: - ; – R. . 100 114 Y . . 1 (33). 2012

H !, , , ! , ! , ! , ! ! . X !, ! ( 4 G). S !! . J ! , . U , . X , , ! , , ( 5). U !! , , ( 5). Y ! , ! !. H ! , !, . H ! ! , ! . W ! , ( 6 F). U! ! ! ; ! , ! , ! , ! ( 6 G), !, . U . Y!, ! ! (21 ! 2011.) , , .

F G 5. . U . T, . T: - ; – R. . 100 X , !, S ! Rhabdias bufonis, ! ! ! ! [8]. U G (2008), ! ! , ! [9]. R.bufonis ! , Saplometra cylindracea.

F G 6. . – ; - . T: ) ; ) R. . 200 H SY. . 1 (33). 2012 115

I! : , , , ! ! . U , , , ! . H , , . W , ! ! , ! . Q 1. L T.S., W T.H., M Q.W. U ! M W // H , ! !. - X , 2002. - H. 3. - W.63-68. 2. X! S.K. M // H F . – 2009. - 3 10(60). – W. 76-79. 3. X! S.K., F F.R., ^ R.^. R Rhabdias bufonis // H! . – 2011. – W. 87-93. 4. Goater C, Ward P. Negative Effects of Rhabdias bufonis Nematoda on the Growth and Survival of the Toads Bufo-Bufo // Oecologia. - 1992. - 89(2). – V. 161-165. 5. V .R. I WWWV / .R. V, H.U. ^, S.S. ^. – R.: S, 1980. – 279 . 6. V G. R! . R.: - . - 1953. 718 . 7. .H., W.W. R T . . IY, 1999. - W.46. 8. I W.S. M X : !: ! /W.S. I, T.S. L, S.F. W, F.I. W , W.. ^, T.F. , F.Z. , F.H. G.- X : . X , 2006.-132. 9. G F.H. T ! ! // H TIY, 2008. - 381 – W. 111-116. *** 2011. '' = '' 20 S ' Rhabdias bufonis % , '' Haplometra cylindracea '' '' (, ', , , ') . M ' 12 (11 , 1 ). P' '' % '' %' ''. G ' '' Rhabdias bufonis . P' '' ' % %' ' 1 ' '. U' ''' ' % '' ' . R, ', , ' % ' '. U' , '' % '' %' ''. *** In work is studied influence of a parasitic invasion by nematode Rhabdias bufonis and trematod, presumably pulmonary Saplometra cylindracea the histologic structure of an internal (a stomach, an intestines, kidneys, gonads, muscles) the lake frog were collected in a channel Naryn, located in the northeast part of delta of the river Ili, in 20 km to the west of settlement Karaoj in the spring of 2011. In research have been subjected 12 individuals of a frog (11 males, 1 female). The data about distribution of frogs on sites channels and their number were cited. It were shown that at all frogs were studied in a body cavity parasites of a class of nematodes, namely Rhabdias bufonis localized, specific to amphibians. Histologic studying of an internal and fabrics has revealed presence of one more kind of the parasite who is belonging to the class trematod. Life cycles of parasites and it’s localization in an internal were considered. So, trematods localized in intestines, in muscles and gonads, nematodes were mainly in muscles, in kidneys and in spermary. Pathological influence of a parasitic invasion on the gisstructure and functional condition of bodies were shown.

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T ''' ' ' (, , , , ', ) 5 ' 21 ' . M' ' ', % '. F ' ' , '' '. L ' - Proteocephalus osculates– 100% , ' 2-128 ' ', '' '. T Ŀ Ľı , T- ı ı Ŀ . R , Ŀ, , , ! ı Ľ . T Ŀ ı ĳ Ŀ T- ı Ľ ı ! ĳ ĳ Ľ . T Ŀ ! ı Ŀ ĳ ı. Ľı Ŀ Ľĳ ı ĳ , Ľ Ŀ Ľ ı Ľ ı Ŀı ı Ľ Ŀ . G ı !! Ľ . G Ŀı Ľ ı !ı ı ĳ . ľ ı !- ! . Q ı Ľ ı ĳ Ľ 116 Y . . 1 (33). 2012

Ŀ ı Ľ ı ĳ. W ı !- ! ı Ľ ĳı ĳ ! Ŀ Ľ ĳ Ľ ı Ŀ Ľ . 2010 Ŀ T Ŀı ! ı ! ĳ. RQJ Lļ M ļJ== U! Ľ 2010 Ľ- T Ŀı G, K Ľ J ! . X ! Ľ ı ĳ (, , ĳ , Ŀ, ) 84 (1-). F ĳ ! -15, -15, ĳ -15, -15, Ŀ-15, 9 . W G ! 28, K-28, J-28, 84 . 1- T Ŀ'' Ľ'' ĭı ĭ M , G G K J W- Cyprinus carpio (Linne) 5 5 5 X -Abramis brama orientalis Berg 55 5 Wĳ - Rutilus rutilus 5 5 5 caspicus F - Aspius aspius (Linne) 5 5 5 Ŀ - Stirostedion luciperca (Linne) 5 5 5 L -S'lurus glan's 3 3 3 28 28 28 G 84 R Ľ Ŀı .K.G!-U! [1] ! Ľ ĳ. U ı ĳ Ľ WVT ı Ŀı 1,2,3 [2,3,4]. M ļLQ= 6 ĳ 84 5 ı 21 ĳ (2-). F : Monogenoidea, Cestoda, Trematoda, Nematoda, Acanthocephala. M ı , , Ŀ , ĳ, , . 2- T Ŀ'' Ľ'' ĭ ' 'ı ĳ' ĭĵ U ' ' Dactylogyrus extensus W D. minutus W /-/ D. wunderi /-/ D. crucifer /-/ D. sphyrna /-/ D. falcatus /-/ D. tuba /-/ Ancyrocephalus paradoxus Ŀ /-/ Siluridiscoides siluri /-/ Paradiplozoon pavlovskii /-/ Caryophylleus laticeps , C. fimbriceps C /-/ Proteocephalus osculatus /-/ Diplostomum spathaceum , , , , Ŀ Ŀ, Tylodelphys clavata , /-/ Phyllodistomum elongatum , Bucephalus polymorphus Ŀ /-/ Bunodera luciopercae Ŀ /-/ Trematoda sp. larva , ĳ, Anisakis sp. , Ŀ, , J , , Pomphorynchus laevis Ŀ, , =ı H SY. . 1 (33). 2012 117

G ı : Dactylogyrus, Ancyrocephalus, Silurodiscoides, Paradiplozoon Ľ . Tı Ŀ Ľ Ancyrocephalus paradoxus, c - 80%, – 7-42 , Silurodiscoides siluri, c – 88.8% Ľ – 7-1211 . Lı Proteocephalus osculates , c – 100% Ľ – 2-128 . G ı Ŀ ı ĳ . G Ŀ c 86,6%- , 1-20 , - c - 66,6%, - 3-12 , c – 53,3%, – 1-8, c - 40%, - 1-16, c – 46,6%, - 1-5, Ŀ c – 6,6%, – 1. W ı 5 ĳ, -7, -6, -7, Ŀ-6 Ľ 5 ĳ . TJ M : Monogenoidea, Cestoda, Trematoda, Nematoda, Acanthocephala. T Dactylogyrus, Ancyrocephalus, Silurodiscoides, Paradiplozoon , , Ŀ , .. ĳ . Kı ĳ ! Proteocephalus osculates, c – 100% Ľ – 2-128 , Ŀ !, c 86,6%, c - 66,6% . M ı Ŀ , ı Ŀ Ŀ ĳ Ŀ, ı , ĳ, ĳ . W Ľ . ļJ 1 G! – U! .K. R .. - Q.: S, 1985. 120 W. 2 T WWWV /T. . W.W. ^. Q.: S, 1984. - X. I. - 428 W. 3 T WWWV /T. . F.H. I. Q.: S, 1985. - X. II. - 424 W. 4 T WWWV /T. . T.S. G. X. Q.: S, 1987 - X. III. - 582 W. *** Y 2010 . : , , , , 21 , 5 : Monogenoidea, Cestoda, Trematoda, Nematoda, Acanthocephala. T , , . T . N Proteocephalus osculates 100 % 2-128 . . *** Our investigations in the spring of 2010 in six species of fish - carp, bream, roach, chub, perch and catfish were found 21 species of parasites belonging to 5 classes. Of the 21 species of parasites found in fish listed in the mouth of the Ural relatively high degree of invazirovannosti observed monogeneans, diplostomid metacercariae, cestodes and nematodes. However, the intensity of infestation is not high and they do not cause fish diseases. The exception is infection with pathogenic catfish cestodes Proteocephalus osculates to 100% with intensity of infection of up to 2-128 copies. which reduces the nutritional state and slows the growth rate of fish.

TL 597 R.L. U, .. ĭ LQ QĨİ (CHANNA ARGUS CANTOR) UĨO TOR TO Q « - » L^W

R 2011 %' '' ''' '. L '' % '' , '''' ' % '' ' ''. L '' % 2012 %'' ' ' , %'' ' ''. W! ^ ı ı Ŀ Ŀ . F ı Ŀ Ŀ Ŀ 1964 [1]. 1966 ^ [2]. 1967 ı ĳ F ıı ı 500 , ı 75% . RļQ=R R ļJ=RQ. R = Ŀı ! 2011 Ľ . L ( Ŀ 20-80 ) . G! ! Ľ ĳ. L RGW-2 Ŀ [3]. F ! .

118 Y . . 1 (33). 2012

ļLQ R QQQ L (Channa argus Cantor) – Ľ . Y, W Ŀı , F Ŀı Ľ Ŀ , -, Ŀ . 1960 V- Ŀ Ľ, F , X, ^ Ŀ Ľ W Ŀı Ŀ [4]. I.R. Jı Ľ F Ľ Ŀ ı Fı ı ı Ľ, F Ŀ, ! ı Ŀ . ı (! Ŀ) Ľ G = Ŀı Ŀ Ŀ ĳ [5, 6]. L ı Ŀ . Fı ĳ Ľ ĳ . Ĳ , ! 25-30 . Y , 2 ĳ. Y ! Ľ Ľ ! , . ı ı ĳ ĳ, , 2008 ı - . ı, K Ľ .. Ŀı , -, Ŀ ! . Ĩ- Ľ 2008 ! Ľ ı (l – 545 , Q – 1955 ) . 2010 = Ŀ ! !, - ı 29,5 21 . 2011 . - = Ŀı ! ı 39 . Ĵ ı 46,2% , 35,9% . RĽ - Ŀ 36,7 Ľ 644 (1-). 1- ĩ ĭ ĩ ĩı ĭ Ŀ''

G Ŀ ı F L n l, Q, . q, . Ŀ 28,5 247 220 3 0,6 0,6 51,5 1774 1625 6 1,6 1,5 39 36,7 644 599 - 1,1 1,1 X ı 2010 . M ı 4-6 . F ı 1:1,1– ı ı Ŀ. Y - 18°W- Ľ ĳ. L ı Ľ 2- . 2- 2010 . ĩ ĭ ĩ ĭı ' ĵĭĩ L Y ., FLX, ı n 4-6 0,8-1,5 1,2 59,2-70,0 64,6 5 ı . F ĳı ! Ľ ! , Ŀ Ŀ Ľ Ŀı ! Ŀ . L ı - Ľ Ľ ı . , ı Ŀ ! . W , ı Ŀ !! : Ŀ ı . Kı ıĳ Ľ ı Ŀ ĳ . G , = Ŀı ı ! Ŀ Ŀ ı , ı ı ĳ . ı ! Ŀ , , Ŀıı .. ı ı ı !!ı ı . W, H SY. . 1 (33). 2012 119

ı ĳ ĳ , , . G-= ı ĳı ĳ ı ĳ ĳ. T - (Noemacheilus strauchi, N. labiatus), ! (Phoxinus brachiurus) Ľ (Persa schrenki) . L Ŀ ĳ Ľ ĳ ı ı ı 2012 Ľ ı 1,6 Ŀ TFR (ı Ŀ) . ļJ 1. G S.K. T ! F !: X. . . . - . W. F . : . 1968. W. 34-35. 2. K H.. ! ! . W: X. . . . - . W. F . : . 1968. W. 62-63. 3. U :. V . R.: U! , 1966.-376 . 4. V . F.: I, 1992. X. 5. W.286-316. 5. J I.R. S Channa argus (Cantor, 1842) . // ! SFS V. W. . . – 2007. - 32 (260). – W. 15-22. 6. G-FĿ Ľ ı ı Ľ ı Ŀ , ı Ŀ (VKLGFR) ! Ľ Ľ Ŀ 2012 . GĿ: : ĨML /G^ĨM. –F, 2010.- 65 . *** 2011 . T, . .

*** In article the basic biological indicators of snakehead according to 2011 are resulted. It is noticed, that snakehead lives on a reservoir in the overgrown and shallow sites. Now snakehead increases the number and further it is recommended to catch as a trade kind of fishes YJ 597-14+597-19

.. , R.T. T TJ ITQb\ \T NEMACHILUS SEWERZOWII O TM.Q P -Z, SNN , F, P

Nemacheilus sewerzowii G.Nikolsky, 1938 ., .G . . U . , . . . I W Nemacheilus sewerzowii G.Nikolsky, 1938 ! !!! G . W! . J I.H.S [1] . I..G [2] - .^. S !, W, H.U.R .Y F [3]. U S.U.W [4], W ., .G, !!. U I.H.S [1] S.U.W [4], W !. H 2003-2008 . I..G [2] 5 ! W . F, ^. , ! ! ! G , ! [5]. H ! ! ! W [6]. \ ! !! W . W.

RQ RTJ QJTO V W F ! !! -W F-]. J 100 , ! 60 [7]. .W : Triplophysa labiata (Kessler, 1874), ! T.strauchii (Kessler, 1874), T.stoliczkai (Steindachner, 1866), T. dorsalis (Kessler, 1872). 120 Y . . 1 (33). 2012

T ! W 2011 . S 3 ! W. R ! [8,9]. J! ! ! : L - ! ; l - ; Q - ; q - ; Fulton - ; Clark - ; aD - !; pD - !; aA, aV, aP – ! ! , ; P-V - ! ! ; V-A - ; lca - !; lc - ; ao - ; o – ; hc - ; io - ; H - ! ; h - ! ; HTT - ! ; hTT - ! ; lD , lA – ! ; hD, hA – ; lP, lV – ; lCs – ; lCm – ; lCi – ; li- . C ! [10] c "Excel 98".

MQb QJTO TLJ I W W. X , ! S.U.W [4] .G . T, ! [2,3,6,11] , ! ! !. T !, ! !, . U : , !. Y !!, ! ( 1).

G 1. I . : – , – V : 50-56 , 41- 46 , 0,9-1,2 , 0,4-0,9 . Y 0,9-1,6, 0,4-0,9 ( 1). U [2,3], .W . Y !. ! 2 , III V !, 1 - III ! . Y V ! ! 512 . Y ! , , , [2,3]. S! . I . D II-III, 4-7 ( 2), F I 4-5, P 11-12, V I 5-7. 34-35. U : , U , ! . , ! ( 3). X 1 R . U min– max M ± m W () (CV) L, 5-5,6 5,27±0,65 1,12 21,3 l , 4,1-4,6 4,40± 0,65 1,12 25,5 Q , 0,9-1,2 1,07± 0,63 1,09 103 q , 0,4-0,9 0,58± 1,04 1,77 300 H SY. . 1 (33). 2012 121

Fulton 0,9-1,6 1,27±0,45 0,77 60,7 Clark 0,4-0,9 0,70±0,74 1,27 183 lD 0,4-0,5 0,48±0,58 0,88 214 hD 0,7-0,8 0,77±0,51 0,88 114 0,3-0,3 0,27 ± 0,58 0,97 333 l 0,9-1,1 0,95± 0,52 0,91 90,9 0,2-0,2 0,15±0,58 0,97 500 hc 0,4-0,5 0,45±0,46 0,80 171 i 0,4-0,5 0,45 ± 0,57 0,91 214 aD 2,3-2,5 2,4±0,63 1,09 45,9 pD 1,5-1,7 1,6±0,61 1,07 66,6 V 1,1-2,4 1,8±0,73 1,26 70,1 H 0,6-0,7 0,65±0,49 0,85 135 h 0,3-0,6 0,53±1,29 2,25 421 V-A 0,7-1,3 0,97±0,74 1,28 133 P-V 1,2-1,5 1,30± 0,72 1,25 0,72 ĻA 0,5-0,6 0,53±0,58 0,95 187 lA 0,4-0,5 0,43±0,58 0,95 230 lP 0,7-0,8 0,75±0,66 1,14 86,0 lV 0,5-0,5 0,45±0,58 0,95 200 Lcs 0,8-0,9 0,85±0,65 1,12 135 Lcm 0,6-0,9 0,73±0,74 1,28 175 Lci 0,7-0,8 0,75±0,66 1,14 155 li 20-30 23,3±0,38 0,66 2,85 HTT 0,5-0,7 0,56±0,41 0,71 126 hTT 0,2-0,2 0,15±0,57 1,01 500 Lca 2-2,1 1,90±0,61 1,05 55,2

2. : 3. U 1- , 2- W: 1-, 2-, 3- , 4- MQM] V ! W .G. W ! 122 Y . . 1 (33). 2012

! [2]. U , . H . U ! , ! ! ! ! . Q 1. S I.H. S (Nemacheilus sewerzowi) W F// G .RTU. S.. T.. – 1938. – X.11-12. H.5-6. 2. G G.. R W Nemacheilus sewerzow'// U. W! - 2010. - 32 (1) – W. 115-119. 3. R H.U. Noemacheilus sewerzowi G.Nikolsky - W// V – F-F: S, 1989. - X.4. - W. 27 -30. 4. W S.U. T ! G // X.. .- W.F – : FS WWV, 1961. – W.201-211. 5. R H.U., J I.R. S // V . – F-F: I. 1992. X.5. W. 329-371. 6. ! F (L) - F, 2006. 520 . 7. V W. http://ru.wikipedia.org/. 8. U .. V . R.: U! . - 1966. - 376 . 9. Holcik J. General introduction to fishes. 2. Determination criteria // The freshwater Fishes of Europe.- Aula-Verlag Wiesbaden. - 1989. - Vol.1. Vart 2. - V. 38-58. 10. Q I.. G! – R.: H! . - 1990. - 352 . 11. R S.^. ! ., XIX .// U. W! . - 2010.- 32 (1) –W. 91-95. *** K ''' Nemacheilus sewerzowii G.Nikolsky, 1938 '' ''' '' , ' G ' ' %'. '' ' ' '' . ''' % '', % %' ' '' . K '' '. X''' ' . *** The first time Severtzov’s stone loach (Nemacheilus sewerzowii G.Nikolsky, 1938) has been found in the Sarkand rivulet. This find showed to the large spread of Severtzov’s stone loach in the Balkhash Lake basin at the foremr time. Nowadays the species distribution area consists from a few separeted populations. Biological and morphological features of Severtzov’s stone loach from the Sarkand rivulet were presented in comparison with former data. Description of morphology of digestive system was given. Environmental condition in the Sarkand rivulet was evaluated as quite faborable for Severtzov’s stone loach.

YJ 597.5 .. ĩ, I.. , I.. QT QĨİ MICROPERCOPS CINCTUS RTZTQTIQ Lļ TQTIQ UR - , F,

% '' Micropercops cinctus % '' . ''' = '' % ^ : '' , , ''' ', , ''' , '' , , , % . '' '' . ı Micropercops cinctus XRJ - ı, Hı . XRJ – ı 1960 Xĳ! . ľ Ľ. W!, =, X, ^ Ľ G Ŀ, F ı ı . Y - , Ľı ı ĳı . L ĳ [1]. ľ ı , F ı . Ľ ı . RQJ R M ļJ== M ı Micropercops cinctus Ľ Ŀ 51 (. 1).

1 - ĩ H SY. . 1 (33). 2012 123

G ! ı Ŀ . M .. Uı Ŀ ! Ľ ! ĳ [2]. M ļLQ= Lļ QQQ Ľ Ŀ ĳ ! Ľ ! Ľ Ŀ: ı ı 29-56 , 39,6 ±0,9, 0,29-2,1 , 0,8 ± 0,06, ı 25 – , 9 – , 17 - (juveniles) ( 1, 2). ı Ŀ 1:3 ı. Ŀ = 1:2 , ^ 1:2 ı [3, 4]. 1- ĩı ĭ Ľ ĭı ĭ ĭ L W % F 9 17,6 25 49,0 Juv 17 33,3 G 51 100 c ı ! Ľı Ŀ 2 – Ŀ (2-). 2- ĩı ĭ '' (n=51) G min –max M ± m W H. () (CV) 1 2 3 4 5 L, 29 - 56 39,6 ±0,9 7 17,6 l , 24 – 46 33,3 ± 0,8 5,9 17,6 Q , 0,29 – 2,1 0,8 ± 0,06 0,5 58,7 q , 0,12 – 1,51 0,6± 0,05 0,4 56,4 I D 11 – 12 11,3 ± 0,2 1,7 23,7 II D 13 – 14 13,3 ± 0,2 1,8 18,9 P 12 -14 12,6 ± 0,3 2,2 23,1 A 10 -11 10,8 ± 0,2 1,7 21,3 sp.br 13-15 13,3 ± 0,3 2,4 23,0 Vert 28-33 29,6 ± 0,2 1,6 5,3 pdH 20-35 24,7± 0,6 4,6 18,6 W 6-14 9,3 ± 0,3 2,2 23,1 Io 2-5 2,5 ± 0,1 0,8 30,2 Hc 5-10 6,2 ± 0,2 1,3 21,2 St 3-7 3,6 ± 0,1 0,9 24,6 Ao 2-4 2,6± 0,1 0,7 25,2 O 2-3 2,3 ± 0,1 0,5 20,1 Po 1-8 4,5 ± 0,3 2,3 52,3 H 5-10 6,3 ± 0,2 1,4 22,1 h 2-4 2,6 ± 0,1 0,9 32,6 Pl 4-9 5,7 ± 0,2 1,4 23,6 aD 8-16 11,7 ± 0,3 2,2 19,2 pD 16-29 21,5 ± 0,5 3,6 16,6 aV 8-16 11,2 ± 0,3 2 17,9 aA 12-27 19,1 ± 0,6 4,1 21,6 l I D 2-7 4,2 ± 0,2 1,5 35,4 l II D 4-11 6,8 ± 0,3 1,9 28,3 I D – II D 1-4 2,3 ± 0,1 0,9 38,4 h I D 2-6 3,8 ± 0,1 1,1 27,7 h II D 4-7 5,5 ± 0,1 0,8 15,3 l V 4-8 5,3 ± 0,1 1,1 20,1 V – A 5-13 8,8 ± 0,3 2,1 23,5 l A 2-8 5,5 ± 0,2 1,6 28,9 h A 4-7 5,1 ± 0,1 0,9 17 124 Y . . 1 (33). 2012

* L – ı ; l – ı ; Q – ı ; q – ı ; I D – ı Ľ ; II D – ı Ľ ; P – ı Ľ ; A – ı Ľ ; sp.br.- ı Ŀ ; vert.- ı ; pdH – ı ; c – ı ; io – Ŀı ; Hc – ı ; st – Ŀı ı ; ao – ı ; o – Ŀı ; po – Ŀı ı ı ; H - ı ı ; h – ı ı ; pl – ı ; aD – ; pD – ; aV – ; aA – ; l I D – ı ; l II D – ı ; I D –II D – ı ; h I D – ı ; h II D - ı ; lV – ı ; V – A - ; lA – ı ; hA - ı . 2009 ! Ŀ 1988 K.J. H, F.U. Rı Ľ = ı Ľ ^ 2007 L L.Y. ĳ ı Ŀ Ŀ : ı , Ŀı , Ŀı , ı , ı ı , ı , ı , ı , ı , ı , ı Ľ ı /3,4/. G = Ŀ Ľ ^ 13 - Ŀ , Ŀ , ! ı 15 -, Ŀı 3 -, Ŀı 4 -, ı ı 6 -, 22 -, ı 5 -, ı 11 -, ı 7 -, ı 9 -, ı 7 -, ı 10 - ( 3). 3- ĩ ĩĩ Ľ'' Ľ Ŀ'' Ľ' = Ŀ ^ (K.J.H, G F.U.R,1988) (L.Y.L,2007) (Ŀ Ľ,2009) min – max min– M ± m min – max M ± m max L 23,7-40,0 23,5-33 26,4±1,49 29 – 56 39,6±1 Lc 28,0-34,2 20,9-34,8 24,69±2,38 6 – 14 9,3±0,3 Io 4,5-8,7 2,7-6,6 5,8±0,11 2 – 5 2,5±0,1 O 6,8-9,03 3,7-7,8 6,48±1,29 2 – 3 2,3±0,1 H 8,2-11,4 6,3-12,7 8,97±1,9 2 – 4 2,6±0,1 aD 36,1-42,2 24,8-44,1 34,13±2,96 8 – 16 11,7±0,3 lD1 9,3-16,4 9,3-14,2 9,98±1,04 2 – 7 4,2±0,2 lD2 15,9-25,9 15,8-25,8 18,59±1,86 4 – 11 6,8±0,3 hD1 9,3-16,4 6,8-17,4 10,87±2,78 2 – 6 3,8±0,1 hD2 10,5-21,6 13,9-27,9 14,98±2,15 4 – 7 5,5±0,1 La 7,5-15,9 9,9-18,7 12,95±1,79 2 – 8 5,5±0,2 lV 14,7-24,2 13,8-19,9 15,96±1,67 4 – 8 5,3±0,1 c ı ı Ľ . c ı ı Ľ Ŀ ( 4). 4- ĩı ĭıĭ ' = Ŀ (V.K.W, (Ŀ , ,1982) ,2009), n=51 min – max M ±m min – max M ±m 1,78-2,50 2,12±0,03 0,73-2,42 1,89±0,03 1,42-2,16 1,66±0,02 0,55-1,86 1,43±0,03 T ı ! Ŀ = Ŀ Ľ ^ Ŀ . Jı Ľ , ı ı ı 56 H SY. . 1 (33). 2012 125

. M ı , : ı , Ŀı , Ŀı , ı ı , , ı , ı , ı , ı , ı , ı . T = Ŀ 32-35, ^ 29-31, ı 28-33. Ŀ ı ! Ľ ! Ŀ Ŀ Ŀ Ŀ. ļJ 1. K H.S. W Eleotridae (Gobioidei, Perciformes), K WWWV, W F. // M. . 1981 .X. 60. 3 10. -W. 1517-1522. 2. U .. V . H.1. R., 1931 – W . 8 -23. 3. H K.J., R F.U. R ! (Pisces Eleotridae) WWWV // M.. 1988. X. 67. 38. - W.1194 - 11204. 4. L L.Y. ^ ı (Micropercops cinctus) ! . X II- R «R ». – F: S . F-. - 2008. –W. 34-35. *** N Micropercops cinctu) P - . N ] , , , , , , , , . *** Morphological and biological features of the beautiful sleeper (Micropercops cinctus) in ponds of the Kapshagai fish breeding farm had been investigated. Morphological and biological differences from other populations of the beautiful sleeper (Micropercops cinctus) from the rivers Ile and economy of Chilik's pond, on length of a body, on length of a head, on diameter of eyes, on undorsal distance, on length of the first back fin, on length of the second back fin, on length of an anal fin, on length of a belly fin, by quantity of vertebras are revealed, etc.

YJ 597 1.. , 2.. TQTI] QTIT T PERCA SHRENKII TM QTQbTO R 1XTT « - !, 2 . -, F,

Perca shrenkii F . U, F , P, " . F . P . J ! Perca shrenkii F . H ! ! G , ! !, !!! ! !, /1/. Q ! F /2/. J ! , ! !! ! F , . RQ RTJ QJTO R 2009 . F . J! ! 20 50 , 25 . G 490 ., 5558 ., 25 ! . T ! RGW – 10 /3/. J! «Excel». MQb QJTO TLJ H F : ( ) (). H ! ! . K 2008 . ! ! 7 %, 2009 . 21 % ( 1). H F !!! . M ! 10 40 %. H W !, ! ! . H ! ! !. 126 Y . . 1 (33). 2012

H F ! ! 3 - 4 , 16,5 80-125 /4/. W ! 9,1 20 . U! ! , ! (!), – , /4/. F ! ! F 2005 . !! 2:1 (. 1).

100 90 80 70 60 % 50 40 30 20 10 0 2008 2009

! () ! () 1. , (%) X 1 , 2005-2009 . % I W W Z N 2005 67,8 29,3 2,9 1089 2006 77,5 21,6 1,0 1683 2007 75,6 24,1 0,3 4607 2008 68,9 31,0 0,1 5105 2009 68,42 31,56 0,02 6048 G ! - . T ! , F ! . . , , , . T ! 5 ‰ /2/. S ! ! , ! !. U ! /5/. H 2009 !. T ! 2009 F ! ! ( 2). V , ! ! . X 2 U , 2009 . T! . FU, !! . . J J! H N ./. ./. , . , % 4+ 33,1 0,15 1,23 0,63-0,86 12 3 5+ 41,7 0,17 1,79 0,69-0,99 60 15 6+ 53,5 0,22 2,39 0,73-1,08 24 6 7+ 80,6 0,21 2,91 0,8-1,2 4 1 100 25 Y ! F , W , ! " ( 3). H 2009 . ! ! "! !. H ! 60 326 , 4 586 , 2 10 . T 4+ 6+ ( 3). H W ! ! 120 330 , 23-735 , 2-11 . H !! ! H SY. . 1 (33). 2012 127

! 147-287 , 56- 443 , 3-8 . R ! 5-6 ( 3). X 3 T , 2009 . F Y. Y. J! , H ! l, Q, q, ] % 2 65 4 3 1,52 1,13 0,03 2 3 107 16 15 1,33 1,22 2,60 157 4 130 30 28 1,32 1,24 37,52 2269 5 164 61 54 1,37 1,22 32,75 1981 6 198 111 96 1,42 1,23 17,69 1070 7 231 184 157 1,48 1,26 7,23 437 8 262 275 231 1,51 1,27 2,05 124 9 299 442 376 1,64 1,4 0,1 6 10 325 552 480 1,61 1,4 0,03 2 100 6048 W 2 120 23 22 1,31 1,28 1,96 1 4 153 63 60 1,76 1,68 1,96 1 5 194 133 123 1,84 1,7 27,45 14 6 212 174 156 1,82 1,64 45,1 23 7 238 250 228 1,84 1,69 11,76 6 8 257 304 283 1,79 1,66 5,88 3 9 282 411 371 1,84 1,66 3,92 2 11 330 735 672 2,05 1,87 1,96 1 100 51 3 147 56 53 1,76 1,67 1,67 1 4 161 77 74 1,91 1,78 10 6 5 196 130 121 1,75 1,59 40 24 6 221 191 174 1,8 1,61 25 15 7 244 247 222 1,67 1,51 16,67 10 8 276 378 333 1,8 1,59 6,67 4 100 60 V ! /5/. X ! ! F , . Q 2. F ! ! F , ! , ! ! . 35 30 25 20 15 10 5 0 234567891011 () . F . W .

2. ", 2009 .

128 Y . . 1 (33). 2012

H W ! ! , ! ! ". Q 1. R S.^., G I.., I.W. V G // V L G . - 3 2. – 2010. – W. 29-36. 2. F S.F., X W.V., F ^.F. ! F : F – 2006 . «G», - 368 . 3. U .. V . R:U! , 1966. -376 . 4. T !! , !! . V: F! : T SV / SV. - F, 2001. - 58 . 5. V : H 5 . - F-F: S, 1988. - X. 4. - 312 . *** R F' ' '' G ''' ''. F' G - ' ' ', F' ''' . F' '' ' '' . ' ' G '''' ' ', ''' . *** In research of the chain of the lakes Alakol, it is found out, that Fultin's condition factor a Perca Shrenki Kessler in lakes Sasykkol and P above, than the lake Alakol. Given the assumption is connected by that abundance and a biomass of a Perca Shrenki Kessler in the lake Alakol the high. As it is noticed, that an age abundance of a perch in an investigated reservoir was reduced.

YJ 597.745 (262.811) ..¹, R .² UQTO UT UT TMJM TTU MTIT UTLb UOTIT RT \QbM T UOTIT MQ ¹ , e-mail: [email protected] ²U FWUcT, e-mail: [email protected]

U . U . U P P, TTUX: P, P, P, G, P, 50 . Pusa( [Phoca] caspica (Gmelin, 1788)) !!! , !. K , ! 1 [1], ! ! 100 ! [2]. W !! 2008 W RWTU , !! ! Endangered (A2abd+3bd+4abd ver 3.1) [3]. ! , , !!! U . J! ! ! ! !. T , , TTUX !. W TTUX : 1. Y ! !; 2. T ; 3. Y . W !, R !, . S !- !! ! . U TTUX : 1. R ; 2. R ; 3. R - ; 4. R . T ! ! !! !: 1. I ; 2. F (!, ! ); 4. ! ; 5. K! ; 6. S !. I !!! , .. . S! 1993/1994 ! W , 1960-1970- [4]. K !!! !, ! , ! ! , ! ! , , . U . T !! !: ! !; ; ; ; ! . H SY. . 1 (33). 2012 129

Y! , ! ! ! . ! ! ! ! ! !! , , ! W , !! ! , ! ! [5]. V ! – !, ! . S! , ! ! W ! !!! . H ! ! , ! ! . H ! , , . ! , , !! ! ! , Clupeonella engrauliformis C. grimmi, !! ! !, . W, , ! ! !. ! !! !, ! ! . , ! !!! ! !, !!! 10% , !! !! . S! , ! , , !!, ! , . U ! TTUX !. H ! V M «T » 12 TTUX ! !. U ! TTUX !! ! : o : !, !! . T : , , ( ), " !. H TTUX , , ! « » !, ! ! . H ! TTUX . o : M ( «F-L»), ( – ); G ( - , ! ); o : ( ); ( ). o : ! ! W ! ! W ! 50 . T – , , !. H ! !, ! ! ! ! W- . W TTUX . T ! ! TTUX ! ! ! W- ! ! ! ! !. Q 1. Krylov V. I. Ecology of the Caspian seal // Finnish Game Reserve. – 1990. – 47. – pp.32-36. 2. Harkonen T., Jussi M., Baimukanov M., Bignert A., Dmitrieva L., Kasimbekov Y., Verevkin M., Wilson S., Goodman S.J. Pup Production and Breeding Distribution of the Caspian Seal (Phoca caspica) in Relation to Human Impacts // AMBIO: A Journal of the Human Environment. – 2008. – 37(5). – pp. 356-361. 3. Harkonen T. 2008. Pusa caspica. In: IUCN 2011. IUCN Red List of Threatened Species 4. Kouraev A.V., Papa P., Mognard N.M., Buharizin P.I., Cazenave A., Cretaux J.F., Dozortseva J., Remy F. Sea ice cover in the Caspian and Aral Seas from historical and satellite data // Journal of Marine Systems. – 2004. – 47. – pp.89-100 5. Kajiwara N., Watanabe M., Wilson S., Eybatov T., Mitrofanov I.V., Aubrey D.G., Khuraskin L.S., Miyazaki N., Tanabe S. Persistent organic pollutants (POPs) in Caspian seals of unusual mortality event during 2000 and 2001 // Environmental Pollution. – 2008. – 152. – pp. 431-442 *** P %' . N ''' '' %'''' '' . KXF ' P '' ' ' : ' P' , , P , G , P'' , 50 '' %''. *** The authors have analyzed data on numbers of Caspian seal and main risk factors of Caspian seal survival. Prospective sites within the Caspian Sea within its Kazakhstan sector for establishment special protected natural areas (SPNAs) have been proposed: ice floe in the Northern Caspian, Kulaly archipelago, Komsomolots gulf, Bautino bay, Kenderli gulf, migration corridor along the coastline at depths up to 50m. 130 Y . . 1 (33). 2012

YJ 639.3 1.. , 1,2Q.. UTJQ UR\ R\T TIT T M ] JTUTQQb ITRTQb `\O 1\! ! , IGY «W», . U. U, 24, W-U, 197022, V!, e-mail: [email protected]; 2IGYS « . .R. W V F S», . R. X, 44, W-U, 194223, V!, e- mail: [email protected]

, , . T , , . (IIU) – -- . V . U . , ( ) 4- . R 11- . U, , (12 . ), . U IIU , , . U . U p , , , . W ! . J! , , ! ! [1,2,3]. J « » - « » «» ( 1 ) ! H – 80 200 ./ . ! ! !, , , «» 200 ./, ! III-IV IV . c , ! , ! [4, 5]. Y ! - ! , . H ! ! 2000 [6, 7, 8]. H , ! [9, 10]. H ! ! ! ! . RQ RTJ QJTO W ! F ! H 2004 2011 . J! 15 (1,5-2 /). U ! 1- !, ! IIU ( 50 66 Q.K.) 20-25 . W ! ( ) ! 2004-2007 . ( ! ). J! G ( 0,50,51 ) . U . W 5-6 I. J! ! (ELISA). H! () 2004 2011 .

H SY. . 1 (33). 2012 131

MQb QJTO TLJ W! IV . 1. W !. H , ! 12 . 18-24 . , . ] 2-3 ! ! ! ( , !). M ! , (. 2), ! !.

. 3. 3- . . 4. , . , , , . . 400. . . 400. J! ! IIU ( 14-14,5 ºW) ! 6 . U 10 . . J IIU ( ! – 3-4 ! 13-13,5\W), IIU ( !). X! ! !! , 1-2- 2-3- !. W 3- ! IIU . W , 3- (. 3), 4- !. J ! (. 4), , IIU.

. 1. . 2. 1- . 1- . U « » . . 400. . . 400. J! IV 11- . U ! [11], 12 . (. 5 6). W , ! 7 . . 2-! 3-! ! ( IIU) 132 Y . . 1 (33). 2012

11-. 2- IIU ! – - (. 7 8). X , ! ( ) , ! .

. 5. . 6. 11- (n=9) (n=9) (+IIU+IIU). (+IIU+IIU). . T 2004 . . T 2004 . *** - V! *** - V! ! (P<0,001). ! (P<0,001).

. 7. . 8. 11- (n=6) (n=6) (+). (+). . T 2011 . . T 2011 . *** - V! *** - V! ! (P<0,001). ! (P<0,001). U ! IIU, . U ! 3 5 , ! , ! - , ! 8-12 . W ! , [12], 2-2,5 , ! ! . , ! , - ( ) ! ! , ! ! ! !, 2,5 . H SY. . 1 (33). 2012 133

Q 1. I S. Q. G (Acipenser gueldenstaedti persicus Borodin) ! // J. FS WWWV. – 1950. - X. 71. - 3 4. - C. 785-788. 2. G .F. G! ! ( ! H) // Y QIY. 1957.- 3 228. - H. 44.- C. 54-71. 3. G .F. . Q.: S. Q. T. - 1975. - 210. . 4. X .H. F ! - ! . H .: «S ! : ! !», R.: HSV. – 2010.- W. 398-404. 5. X .H., V Q.. U - !! ! // R. IV R. S.-. . «U ! ! ! ! ». F. – 2011. - W. 239-243. 6. I F.G., X .H.. H ! (Acipenser gueldenstaedti Br.) ! //R. 2- R. -. . «F : ! !». F. 21-22 !! 2001.- W. 22-23. 7. X .H. H ! ! // J R - «V : ». 10-11 !! 2011 ., R. HSV.- W. 186-194. 8. X .H., I F.G. H ! ! // R. J IV R. S.-. . «F : ! !». F, 2006.- W. 127-130. 9. I F.G., X .H. H! ! (Acipenser gueldenstaedti Br.) //R. R. . «W !». F. ! 2002. - W. 87-89. 10. X .H., I F.G., R T.I, G Q.H., C X.G. ! QI-VI // X VIII H “S !-2010”, W-U: “T-WU”, 2010.- W. 137-139. 11. Semenkova T.B., Barannikova I.A., Kime D.E., McAllister B.G., Bayunova L.V., Dyubin V.P. and Kolmakov N. Sex steroids profiles in female and male stellate sturgeon (Acipenser stellatus Pallas) during final maturation induced by hormonal treatment // J. Appl. Ichtiol. 2002. V. 18. - V. 375- 381. 12. R ! ! ! H (. .H. X)// W-U, IY W: «^». - 2010. - 44 . *** M ' %'' '' ' ' '''' , ' '' ''' '' '' ' '' ' % %' ''' %. G '' ' '' ''-' ' ' ' ''-' '' . L '-' (IIU) % – '' -- '' ''' . U '' % '''' . G' '' '' ''' ''. G '' , 4-' ' ( ) '. ' %'' (''''') 11- '' ''' . ^ % ' ( ' 12 .) ' '' ''' ', '''. ' '' IIU- ' '' '' ''' %'. G ''' % '' () . F %' ''', ', ' , ' % ''' ' ' ''. *** Reservation of natural populations by methods of artificial breeding needs in using all genetic diversity of breeders. However now aquaculturists have to reject the female sturgeons having oocytes with low degree of polarization and average weight below definitive value at the beginning of spring spawning season. The shortage of mature males is one of the factors limiting possibilities of spring breeding period prolongation for maturation of females with retarded vitellogenesis. The possibilities of Russian sturgeon male spermiation prolongation by the supplemental injections of sturgeon glycerol pituitary preparation (GPP) or synthetic superactive LH-RH analogue (surfagon) were investigated. The data showed that the optimal doses of both preparations and their comparative effectiveness depend on water temperature and intervals between injections. The histological changes of testis structure after multiple injections were demonstrated. The continued decline of spermatozoa content in seminal tubules took place during this process making impossible 4-th injection. The testosterone and 11-ketotestosterone blood serum levels had been measured by enzyme linked immuno-sorbent assay (ELISA). The slight increase of both androgens at the moment of induction of spermiation (12 hours after 1-st injection) was followed by their sharp decline. The supplemental injections of GPP or surfagon stimulated enforcement or even re-starting of spermiation but did not induce new androgen level’s elevations. Significant differences in influence of two preparations on androgen’s profiles were not observed. The data indicates the possibilities of supplemental hormonal treatment of Russian sturgeon males and likely the males of others sturgeon species for spermiation prolongation and their reproductive potential maximal using.

YJ 597-15 .. , .. R T JU\T TMRTLT TT TI ]LTJ JT QTIT O JIU «S- » VIU « -», . F, V . E-mail: [email protected]

Y, G . S - 134 Y . . 1 (33). 2012

Pseudorasbora parva. U - . W ! . G G , ! S-F [1,2]. H G . , [3]. ! ! ! ! [4]. M ! W F !! ! , [3, 5]. M !! ! !. RQ RTJ QJTO J! !! ! [6]. H ! : Diptychus dybowskii Kessler, 1874, ! Schizothorax argentatus argentatus Kessler, 1874, Triplophysa stoliczkai (Steindachner, 1866), ! Triplophysa strauchii (Kessler, 1874), Triplophysa dorsalis (Kessler, 1872), Pseudorasbora parva (Temminck et Schlegel, 1846), Micropercops (Hypseleotris) cintus (Dabry et Thiersant, 1872), Abbottina rivularis (Basilewsky, 1855), Carassius gibelio (Bloch, 1782). J! 2-5 : , Y F, F, G F, - ! ] !. V ! 5- !. U , ! ! , . T ! !. H 9 . U !! ! , ! !! , ( 5 ), , !. G : 1) - , ! , , ! , ; 2) - , , , ; 3) . H ! 3-5 ! 0,5 1,0 1 . J! ! Chironomus sp., Tubifex sp., 5 , ! , . H ! [7, 8] 1 2.5-3% . H ! 2- . U . MQb QJTO TLJ W! ! ! ! ! G (^-, ], , G-F, .): ! ! , ! ! ! ( 30-50 ). R , !, . T . X U H R! Y! H! () !1 50% 100% [ ] 10 4-12 14- 3- [9] + 4-18 9-74 +, 3-14 9-46 ! 9 14-35 71- 3- [10] + 14-31 92- 2- +, 6-18 36-65 6 4-36 21- 3- [11] + 8-24 28- 4- +, 6-19 25-54 ! 7 8-58 45- 5- H SY. . 1 (33). 2012 135

[11] + 8-34 38- 3- +, 8-24 20-63 6 12-26 72- 3- [11] + 8-26 68- 2- +, 6-19 23-72 5 16-94 72- 2- [12] +, 26-121 80 2 3 3-45 22-183 [13] +, 1-12 8- 80 2 5 6-72 42- 2- [14] +, 8-24 62- 80 2 7 14-94 2- [15] +, 16-28 72- 80 2 U: 1 – !: – ! , + – , +, – , 2 – U ! ! . T !! ! ! ! , : ! , . H, ! . ! – 2,5 !. H ! , ! , . U , . H, , . W , , !!! ! [16]. H ! . H ! ! – Chironomus sp. 3I. 3IV ! ! . F !! ! ! – ! , . F ( 5 20 ), 3 () 2 ( ). U! !! : , , . F (, ) ! ; . , . MQM] X , ! , ! , , !, . H!, , , !! ! ! . Q 1. G Q.W. V U F ! // – R., Q.: - FS WWWV, 1962. - X.5. - W.320-363. 2. R H.U. // V – F-F: S, 1986. - X.1.- W.6-40. 3. J I.R., R H.U. ! // V – F-F: I, 1992. - X.5. - W.6-44. 4. G // U . F F.., G S.I. – R.: X R, 2004. - 436 . 136 Y . . 1 (33). 2012

5. X H.I., W F.W. F G // H – 1995. - X.35. - H.1.- W.71-77. 6. W W. W – R.: Q! ! , 1983. - 192 . 7. W! H.., I K.F., V Q.U. – R.: Q! ! , 1984. - 120 . 8. U Z.F. – R.: F, 1991. - 368 . 9. W F.., X W.V. Diptychus dybowskii Kessler - // V . - F-F: S, 1988. - X.3. - W.93-105. 10. G F.F., R H.U., X W.V. Schizothorax argentatus Kessler – ! // X . 1988. X.3. W. 57-83. 11. R H.U. V Noemacheilus Van Hasselt, 1823 – I// V . - F-F: S, 1989. - X.4. - W.6-63. 12. G F.F. Pseudorasbora parva (Schlegel) – // V – F-F: I, 1992. - X.5. - W.159- 169. 13. I .H., J I.R., H.K. Hypseleotris cinctus (Dabry de Thiersant) – // V – F-F: I, 1992. - X.5. - W.254-269. 14. G F.F., R H.U. T // G – F-F: IY, 1975. - H.9. - W.121-127. 15. I F.. Carassius auratus gibelio (Bloch) – ! // V . - F-F: S. 1988. X.3. 212-231. 16. S I.H. V F – R.: - FS WWWV. 1956. 551 . *** F G Ľ Ľ Ŀ ı ĳ Ŀ. F ı Pseudorasbora parva ı Ŀ. G ĳı Ľ . *** Long term coexistence of many indigenous and non-commercial alien fishes was shown as result of experiments in aquariums. Stone moroco (topmouth gudgeon) Pseudorasbora parva has the largest adaptive possibilities. Unfavorable impact of stone moroco to the other fish species was observed.

YJ 597 J.. ] TRT TT MUT QT\ TQIT-UOTR O - ! V!, . F

-P . M . " . J . G (Stenodus leucichtys leucichtys,Guldenstadt, 1772) – ! ! , !. H W Q U ! [1, 2]. G ! ! 50 , - ! W . U! , , . M 5-7 . H. R ! ! , , H ! 2007 -2011 . «I!» (. G, ! F ). (485056) – ./. M ! 2007 - 2011 . 2490 . M ! F TVM 322 . , ! 218 . [3] (. 1). X. 1 , I 2007 2008 2009 2010 2011 - , 218 15 15 29 45 . T 61 58 41 30 28 , . R! ! . H !! , ! " ( 90%) – I . H . H 77% , : – 16%, ! – 19%, – 42%. H 23% , – 21%, – 2%. M ! . H ! " . V ! ! H 70- – 80- . H SY. . 1 (33). 2012 137

, 0,4 50- 50 90-. T, , ! !! , " 44 , 80- [4]. W, ! 2 - 3 , ! . J! ! F H ! " . T ! . H, . W 2001 . !! ! SV, , ! H . M 5 (2007-2011 .) ! 0.87 ./, 1.14 ./. J , : – 0.5 ./, 0.12 ./ (. 1). W !!! , ! , , .

1,2

0,8

0,6

0,4

0,2

,./ 0

. 1. J 2007-2011 . . «I», ./ H , !! , ! ! ! (. 2). X 2 U , . H, I U 5 6 7 8 9 10 J, 80.5 84.0 86.2 88.7 93.5 95.0 2007 R, 6.3 7.2 8.0 8.7 9.7 11.3 J, 78.0 82.7 86.6 90.8 91.0 96.0 2008 R, 5.9 6.4 7.8 8.5 8.7 12.0 2009 J, 78.0 83.6 89.5 89.0 92.0 97.0 R, 6.2 7.0 8.4 8.6 9.4 11.1 J, 78.6 86.8 87.8 91.0 - - 2010 R, 5.74 8.2 8.6 8.9 - - J, 78.4 83.6 87.2 92.0 99.0 - 2011 R, 5.92 6.4 8.2 9.1 10.2 - M 2007-2011 . 2009 ., !! ! ! 86.3 , – 7.9 . H (2010-2011 .) - !. J 2007-2011 . 70.0 99.0 , – 4.5 10.2 , , 85.6 7.52 (. 3).

138 Y . . 1 (33). 2012

X. 3 J 2007-2011 . . I J, R, min max min max 2007 76.0 97.0 86.1±0.9 5.5 10.4 7.6±0.2 2008 70.0 96.0 85.1±0.7 4.8 12.0 7.3±0.18 2009 75.0 98.0 86.3±0.85 5.7 11.3 7.9±0.22 2010 75.0 91.0 84.61.3 4.5 8.9 7.540.37 2011 78.4 99.0 85.81.04 5.92 10.2 7.520.26 H ! 1980 – 1989 . 13- . W 2001 . ! 10 . H! ! 2007-2011 . ! 5-10 (. 2). T ! ! 5-9 (97.7%). T! ! ! , . 35

P, % P, 10

0 5678910

. 2. , % H ! ! ! [5]. U ! , , 1970- [6], ! " ! . S ! ! !!. H " ! 20- 30% . F !! F , ! (. X) . !. H 5 8 835 040 . H 5 235 735 . (. 4). c ! ! ! . J! ! 10 . . X 4. , . I , . . 2007 1.79 2008 1.86 2009 0.000727 2010 0.837008 2011 0.748000 H ! ! ! (. 5). X. 5. ] , . H H I 10 4+ 5+ 6+ 7+ 8+ 9+ 2008 0.001285 0.026982 0.010667 0.000845 0.000738 0.000197 0.000075 0.040789 2009 0.001217 0.007525 0.022361 0.007985 0.000507 0.000286 0.000032 0.039914 2010 0.001226 0.007129 0.006236 0.016739 0.004797 0.000197 0.000043 0.036367 2011 0.000757 0.007179 0.005908 0.004668 0.010056 0.001862 0.000031 0.030460 W 0.0018184 0.0123374 0.0080128 0.0062932 0.003321 0.00061 0.0000412 0.0336814 2007-2011 H SY. . 1 (33). 2012 139

, ! ! 2007-2011 . 33 681 ., ! 322 . , 0,9 % ! . H ! ! F TVM 1000 . . , . U! ! . H , ! ! ! . H ! ! (. «!», . «\!», . «W!», .«9-! T"» H ). H " ! !! F . Q 1. V Z.W. c! . – R.: S. – 1980. 2. G Q.W. V WWWV // X. 1 . FS WWWV. – 1948. – 466. 3. U .. V ( ). - 4-, . // U . U.F. J! H.H. U. – R.: U! . – 1966. – 376 . 4. H T.S. G ! ! H. – F: SV. – 2002. – 114 . 5. J H.., W F.J., U H.., U Q.R. W ! ! ! ! H // G - V. – H. – 2000. – ].2. – W. 56- 57. 6. Q R.F. H . – R.: Q! . – 1983. – 112 . *** R K'-P '' '' . K' ''' ' '' . '' ''. *** In the article current state reserves of Stenodus leucichtys leucichtys,Guldenstadt, 1772 in Volgensis-Caspian basin are considered. The reserves of Stenodus leucichtys leucichtys,Guldenstadt, 1772after regulations of runoff Volga river formed at the expense of artificial reproduction.

YJ 597.19 ..], ..T JTTO T TZ UOTIT RT UJQ RITO TQ UQ M MS VFS, W-U, e-mail: [email protected] XTT « F «FUc» F, e-mail: [email protected]

P R . V (2007- 2010 .). J . 49 , 21 a 8 . , . H ! (2007- 2010 .) !! ! ! R : , , L, X -, J (W-H ), W, T, S, F (W ), . W W-H (WU-1, WU- 3, WU-5), – W (WU-7 WU-9). ! , 2007 ., W-H , F 2009 ., ! . RQ RTJ QJTO V . M 10 - , , L, X -, J (W-H ) S (W ). H ! , !! . X! 20 !, , , L, X -, J (W-H ), W, T, S, , F (W ) W-H W . Q ! ( !! ), [1, 2, 3, 4], ! ! R . ! [5,6].

140 Y . . 1 (33). 2012

MQb QJTO TLJ 100 126 . U , . S 76 47 17 , ! [1,7]. U , ! I R , . 1. M 49 , 21 a 8 7 ! (. 2). H ! , (47- 67%). H ! 47 74% , . X 1 , R , 2007- 2010 . Q : Q 1 T! T Asipenseriformes W T Acipenseridae Acipenser Linnaeus, 1758 Acipenser gueldenstaedtii A. gueldenstaedtii Brandt et V ii (Brandt) Ratzeburg, 1833 U ii A. persicus (Borodin) A. persicus Borodin, 1897 A. nudiventris Lovetsky, ^ Ľii i A. nudiventris (Lovetsky) 1828 W A. stellatus (Pallas) A. stellatus Pallas, 1771 Huso Brandt, 1869 G Huso huso (Linnaeus, 1758) H. huso (Linnaeus, 1758) T! W Clupeiformes W W Clupeidae Alosa Linck, 1790 ! ! Alosa brashnikovi brashnikovi A. brashnikovi brashnikovi (!) (Borodin) (Borodin, 1904) A. caspia caspia (Eichwald, W A. caspia caspia (Eichwald) 1838) H! ! A. kessleri volgensis (Berg) A. volgensis (Berg, 1913) (!) i A. saposchnikowii (Grimm, G A. saposchnikowii (Grimm) 1885) A. sphaerocephala (Berg, F A. sphaerocephala (Berg) 1913) Clupeonella Kessler, 1877 ]-! -ı – Clupeonella cultriventris C caspia Svetovidov, 1941 (Nordmann) T! Cypriniformes W Cyprinidae Abramis Cuvier, 1816 Q , Abramis brama (Linnaeus) A. brama (Linnaeus, 1758) Alburnus Rafinesque, 1820 Chalcalburnus chalcoides A. chalcoides ! ! , - (Guldenstadt) (Gueldenstaedt, 1772) Aspius Agassiz, 1832 T Ľii Aspius aspius (Linnaeus) A. aspius (Linnaeus, 1758) Ballerus Heckel, 1843 B. ballerus (Linnaeus, W Ŀ, Ŀ- Abramis ballerus (Linnaeus) 1758) G , - A. sapa (Pallas) B. sapa (Pallas, 1814) Rutilus Rafinesque, 1820 Rutilus rutilus caspicus R. caspicus (Jakovlev, W-! , -Ŀ (Jakowlew) 1870) R. kutum (Kamenetsky, i, i R. frisii (Nordmann) 1901) Pelecus Agassiz, 1835 H SY. . 1 (33). 2012 141

Q : Q 1 P. cultratus (Linnaeus, ] - Pelecus cultratus (Linnaeus) 1758) T! F Atheriniformes W F Atherinidae Atherina Linnaeus, 1758 , Atherina boyeri caspia F A. caspia Eichwald, 1831 Ľii (Eichwald) T! Syngnthiformes W Syngnathidae Syngnathus Linnaeus, 1758 -, ıi Syngnathus nigrolineatus ! - S. caspius Eichwald, 1831 i caspius (Eichwald) T! Mugiliformes W Mugilidae Liza Jordan & Swain 1884 T , ĳi Liza saliens (Risso) L. saliens (Risso, 1810) T! T Perciformes W T Percidae Sander Oken, 1817 S. lucioperca (Linnaeus, W Ŀ, ii- Sander lucioperca (Linnaeus) 1758) W ıi Ŀi S. marinus (Cuvier, 1828) S. marinus (Cuvier, 1828) W G Gobiidae Benthophilus Eichwald,

1831 F B. abdurachmanovi U F - Ragimov, 1978 U G G Benthophilus baeri (Kessler) B. baeri Kessler, 1877 B. casachicus Ragimov, ! B. casachicus (Rahimov) 1978 U I I B. grimmi (Kessler) B. grimmi Kessler, 1877 U B. kessleri (Berg) B. kessleri Berg, 1927 B. macrocephalus (Pallas, ! ĳ B. macrocephalus (Pallas) 1787) U ! B. magistri (Iljin) B. magistri Iljin, 19272 R B. mahmudbejovi Ragimov, U R B. mahmudbejovi (Rahimov) 1976 U ! ii B. spinosus (Kessler) B. spinosus Kessler, 1877 W B. svetovidovi (Pinchuk et B. svetovidovi Pinchuk et U W Rahimov) Ragimov,1979 U G G B. leobergius Berg, 1949 B. leobergius Berg, 1949 Caspiosoma Iljin, 1927 Caspiosoma caspium (Kessler) C. caspium (Kessler, 1877) Neogobius Iljin, 1927 - Neogobius caspius (Eichwald) N. caspius (Eichwald, 1831) - - N. fluviatilis pallasi (Berg) N. fluviatilis (Pallas, 1814) - N. melanostomus (Pallas G-! N. melanostomus (Pallas) 1814) Babka Iljin, 1927 - N.s gymnotrachelus B. gymnotrachelus (Kessler, - macrophthalmus (Kessler) 1857) Mesogobius Bleeker, 1874 ı - W Mesogobius nonultimus (Iljin) M. nonultimus (Iljin, 1936) Ponticola Iljin, 1927 142 Y . . 1 (33). 2012

Q : Q 1 Ľ Neogobius iljini (Vasiljeva et P. gorlap (Iljin in Berg - Vasiljev) 1949) P. syrman (Nordmann G- N. syrman (Nordman) 1840) G- N. ratan goebeli (Kessler) P. goebelii (Kessler, 1874) Proterorhinus Smitt, 1900 - Proterorhinus marmoratus P. nasalis (De Filippi, G- (Pallas) 1863)3 Hyrcanogobius Iljin, 1928 G G G - Hyrcanogobius bergi (Iljin) H. bergi Iljin, 1928 Knipowitschia Iljin, 1927 Knipowitschia caucasica G- - K. caucasica (Berg, 1916) (Berg) - G K. iljini (Berg) K. iljini Berg, 1931 J K. longecaudata (Kessler, K. longecaudata (Kessler) - 1877) U: 1 - : [1], ! [2,3]. 2 - ! ! [8]; 3 - Proterorhinus marmoratus ] ; ! - ! Proterorhinus nasalis (Filippi, 1863) [2,4] X 2 , R , 2007-2010 . 2007 2008 2009 2010 Q T T T 1. Acipenser gueldenstaedtii V + + + + + + + + 2. A. nudiventris ^ ----+ - -- 3. A. persicus U - - + + - + + + 4. A. stellatus W + + + + - + + + 5. Huso huso G ------+ + ! ! + 6. Alosa brashnikovi + + + + + + + W- + 7. A. caspia + + + + - - - 8. A. saposchnikowii G + + + + + + + + 9. A. sphaerocephala F + + + + + + + + H! ! , 10. A. kessleri volgensis - - + + - + - + ! (!) ]-! + 11. Clupeonella cultriventris + + + + + + + 12. Abramis brama Q + + + + + + + + 13. Ballerus ballerus W - - - - + - - - 14. B. sapa G + - + + + - - - 15. Aspius aspius T - + + + + + + + 16. Chalcalburnus - ! ! + + + - - - - chalcoides 17. Pelecus cultratus ] + + + + + + - + 18. Rutilus frisii - - + + - + + + 19. R. rutilus caspicus W-! + + + + + + + + 20. Liza saliens (Risso) T - + - + - - - - 21. Atherina boyeri caspia F + + + + + + + + 22. Syngnathus nigrolineatus + ! - + + + + + + + caspius H SY. . 1 (33). 2012 143

2007 2008 2009 2010 Q T T T 23. Sander lucioperca W - - + + - - - - 24. S. marinus c + - - - - + - + 25. Benthophilus U + - - - - + + + abdurahmanovi F 26. B. baeri U G - - - - + + + + 27. B. casachicus ! - - - - - + - + 28. B. kessleri U - + + + - + - + 29. B. macrocephalus ! - + + + + - - - 30. B. cf. mahmudbejovi U R + - + + - + + + 31. B. magistri U ! - - + + + - - - 32. B. spinosus U ! - - - - - + - + 33. B. svetovidovi U W - - - - - + + + 34. B. grimmi U I - + + + - - - - 35. B. leobergius U G - - - - + - - - / Benthophilus sp. U sp. - - - - + - - - 36. Caspiosoma caspium + + + + + - + 37. Hyrcanogobius bergi G G + + - + + + + + 38. Knipowitschia caucasica G- - + - + + - - - 39. K. iljini G - + + + + - - - J - 40. K. longecaudata + + + + + + + 41. Mesogobius nonultimus W + + - + - - - - 42. Neogobius - gymnotrachelus - + + + + + + + macrophthalmus 43. N. caspius + + + + + - + + - + 44. N. iljini - - - - - + + 45. N. melanostomus G-! + + + + + + + + - + 46. N. fluviatilis pallasi + + + + + + + 47. N. syrman eurystomus G- + - + + - + - + 48. N. ratan goebeli G- ------+ + 49. Proterorhinus - G- + + + + + + + marmoratus XTIT T! 7 6 7 6 7 6 6 6 6 W 8 78 7 8 7 7 77 V 21 1318 19 19 14 17 1417 H 49 2329 33 37 30 31 2735 H % 100 46.9 59.2 67.3 75.5 61.2 63.3 55.1 71.4 !!! ( !! – ! ), . S (9-18 ), (4-7 ) (4-6 ). T 2-4 . H !!! . H ! . H W-H , W . H W-H ! , J, , X -. H , , . H . W , , . H ! ! . S, R , . S , ! . , !! ! , !.

144 Y . . 1 (33). 2012

Q 1. Naseka A.M., Bogutskaya N.G. Fishes of the Caspian Sea: zoogeography and updated check-list //Zoosystematica Rossica. – 2009. – V. 18. - 3 2. - P. 295-317. 2. Freyhof J., Naseka A.M. Proterorhinus tataricus, a new tubenose goby from Crimea, Ukraine (Teleostei: Gobiidae) // Ichthyol. Explor. Freshwat. - 2007. V. 18. - 34. - P. 325-334. 3. Neilson M.E., Stepien C.A. Escape from the Ponto-Caspian: Evolution and biogeography of an endemic goby species flock (Benthophilinae: Gobiidae: Teleostei) // Molecular Phylogenetics and Evolution. - 2009a. - V. 52. - P.84-102. 4. Neilson M.E., Stepien C.A. Evolution and phylogeography of the tubenose goby genus Proterorhinus (Gobiidae: Teleostei): evidence for new cryptic species //Biological Journal of the Linnean Society. -2009b. - V. 96. - P.664–684. 5. J I.R., R S.^., R .H. F V . W 1. W R, T, W, Q, W, , _, Y, //«! SFS V», . . . - 2010. - 3 3 (279). - W. 36-49. 6. J I. R., R S. ^., R . H. F V . W 2. W ], G, H , W, F, U, F, S, , , , T, I, G, M, , // ! SFS V, . . . - 2010. - 3 4 (280). - W. 18-28. 7. K.S. V !. – R.: Q"! ! . - 1981. – 240 . 8. Boldyrev V.S., Bogutskaya N.G. Revision of the tadpole-gobies of the genus Benthophilus (Teleostei: Gobiidae) // Ichthyol. Explor. Freshwaters. - 2007. - V. 18. 3 1. V. 31-96. *** P ''' R . G ' (2007-2010 .) ' . G ' ' % '', ' % . G 49 ', 8 21 '. G ''' , % '' . *** The species composition of the Caspian fish fauna is presented, for waters of the Mangistau region . Fishes were collected during four-year State Ecological Assessment (2007- 2010). Gill nets and beam trawl were used, both in spring and autumn periods. Totally, 49 species of 21 genera and 8 families were caught. The list of the species is included, with Kazakh, Russian and Latin names.

YJ 597 ..], ..T T T TLJ RTTIT J SANDER MARINUS T-TT]TO ] UOTIT RT MS VFS, W-U, e-mail: [email protected] XTT « F «FUc», F, e-mail: [email protected]

S Sander marinus (Cuvier, 1828) (Perciformes), P (P ). T ! Sander marinus (Cuvier, 1828) !!! !! Z !, ! [1]. U R, – -G, W F [1]. H ! ! . H ! (2007- 2010 .) !! !, - ! – S F. RQ RTJ QJTO V . W F 2007 . FUc - .H. T F.W. J. H 2010 . .

1 R Sander marinus. TL 490 , VI-II. W S. J ! 9 ! 2010 ., 42°51'36.7'' .., 51°52'04.3'' ..; - 13.6 . S ! . _ . V! ! . J ! TL 423 , SL 382 , – 0.97 , 0.88 ; IV. H (22 ). Q 1D XII, 2D II 15, F II 10. J TL 490 , SL 450 (. 1), 1.3 , 1.2 ; , VI-II. L . Q 1D XIV, II D III 15, F II 10. H SY. . 1 (33). 2012 145

H !, ( 10), ! . U ! . c! !! M VFS (. 3 55034). Y! : 14.3°W, 11.2‰, 10.5 /; W 10.9, ! 14 /. W! , ! - , . Ceramium sp. (75%) Polysiphonia elongate (25%), ! Cladophora sp.; ! Dermatolithon sp. Mytilaster lineatus Balanus improvisus, Corophium chelicorne, C. mucronatum, Amathillina cristata, Dikerogammarus haemobaphes. T ! , Corophium. T Dreissena rostriformis M. lineatus. H Acipenser gueldenstaedtii Brandt et Ratzeburg, 1833, A. persicus Borodin, 1897 ! Alosa brashnikovi brashnikovi (Borodin, 1904). W 17% 36% . F. H 2007 . 3- !. T ! (UAKT-3, 09 !! 2007 .) ! , (43°36'23,81" .., 51°12'09,79" ..). J (12 !! 2007 .) F: ! Ah- 5 (43°36'05,26" .., 51°12'27,35" ..) - , ! Ah-1 (43°36'23,81" .., 51°12'09,79" ..) - , . S ! 7.5-14.5 , W 4.5-7.3 , 10.6-16.8°W, 9.34-10.04 /, S 8.07-8.68, 11.5-12.3‰. I – , ! (UAKT-3 Ah-5), , ! (Ah-5). S UAKT-3, 13 ! (TL) 260-450 ( 287.7 ), SL 220-390 ( 252.3 ), 150-350 ( 215 ); ! 145-330 ( 192 ). I II, ^. W , ! 5.5:1, 4+, 5+ 7+ . H 6+ . X 1 J S. marina ( UAKT-3) ] J R U TL, SL, , , 4+ W 1 II 260 220 170 155 260-290 230-250 160-200 150-190 5+ W 5 II 270.0 234 187.0 176 260-260 220-220 150-190 145-170 W 2 III 260 220 170 157.5 6+ 280-450 250-390 240-300 180-215 W 2 II-III 365 320 270 197.5 270-320 250-280 200-350 180-330 7+ W 3 II-III 293.3 270 270 250 U. U !, – . H 317 !/, - 94.4 /. W (52.3% 76.5% ), , A. caspia caspia (Eichwald, 1838) Rutilus caspicus (Jakovlev, 1870). , 4+ 7+ (. 1). V 5- . U ( III) ! 6+ 260-280 . H ! Ah-1 Ah-5, , ! Alosa brashnikovi brashnikovi (Borodin, 1904) - Neogobius iljini (Vasiljeva et Vasiljev, 1996), ! ( 90.3 % 98.9% ). ] ! Ah-1, Ah-5 . 2. X 2 ] S. marina ( Ah-1 Ah-5), / ], ./ / 20 30 40 50 Ah-1 44.3 27.7 125.9 19.4 217.3 Ah-5 21.4 49.0 0 7.1 77.6 146 Y . . 1 (33). 2012

H 98.3 , / Ah-1 9.30 9.97 96.59 14.09 129.95 Ah-5 7.57 25.47 0 10.36 43.40 H 57.8

X 3. J S. marina ( Ah-1 Ah-5) ] J R U TL, SL, , , 250-280 210-240 170-240 150-165 W 3 III 3+ 260 223.3 200.0 158.3 W ------260-340 230-300 175-470 155-420 W 11 III 4-4+ 280.0 247.3 245.5 219.5 W ------210- 270-440 240-390 190-990 W 12 III 1050 378.3 333.3 669.8 5-5+ 717.3 400-420 350-370 790-800 690-740 W 2 III 410 360 675.0 629.8 570- 350-450 310-400 525-980 W 6 III 1000 408.3 358.3 746.3 795.0 6-6+ 814- 390-480 340-420 710-1330 W 4 III 1480 435 380 1020.0 1147.0 U. U !, – . H ! Ah-1 Ah-5 38 ! TL 250- 480 ( 352.9 ), SL 210-420 ( 310 ), 170-1480 ( 588.1 ); ! 150-1330 . V 3+ 6+. I ^. W , ! 5.3:1, 3+ 4-4+ . H 5+ 6+ (. 3). X !, F ! , (. 2).

, 400

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) ! [3]. T! Sander volgensis (Gmelin, 1789) ! ( ) [3]. J 62 , 50 [3]; - 12 [1].

3 I . TL 423 . , , U !, ] ! ( J- G , G J ) [2]. H ] ! !. J 8- ! 55 , - 3 . U ! 3-4 . S! - 8-20°W, . , . H , ! 13 126 . . T ! . U ! (), (, , , ) [2]. R !, -, , : ! 49 [1]. W , ! !!, ! [2]. H 48 , – 7 . ] , , . W !. H 5-6- . X . ] - ! 5- ! ! ! [1]. T, ! , ! [1], ! !. U S ! TL 490 VI-II !. Y! ; ! ! [1, 3]. H Y [2]. H, . Q 1. M.R. 1981. R Stizostedion marinum (Cuvier) ! // H . - X. 21. - H. 5. - W. 816-822. 2. V Y // http: fish.kiev.ua/pages/ukrfish.htm 3. R U.W., Q ., W H. F : T K. – WU: XJ F. - 2009. – 287 . *** P ''' ' ( ) ' ' ''' Sander marinus (Cuvier, 1828) (Perciformes) . *** New records of the Estuarine Perch Sander marinus (Cuvier, 1828) (Perciformes) are given, from the north-eastern part of the Caspian Sea (Kazakhstan waters).

148 Y . . 1 (33). 2012

YJ 639.3 I.R. , .. , .. T\ I]TIT MTTM T _ TT RTQQT-I]R RTJR XTT «S !», XTT «SH», e-mail: [email protected]

- - (VR). H ! ! V !!! ! ! !. U ! ! !!! ! ! , ! , ! , ! . R!- . J! !! U\V- RAPD-U\V (U\V JS), JS , . \ – !- ! ! ! - (VRW). RQ RTJ QJTO J! ! 2- (33 ) 4- (80 ) (35 ), !. T 96% . , ! (! ), . H JS ! . J! ! ! ! , \ !- H S ! (HSVT) (S. R ., 2008, I 44 (7) . 913-919). U HSVT U\V- V 3 2332463 25 !! 2007 . F JSP. F ! U\V, VXW-225 MJ Research (W^F). V! (25 ) ! ! 2,5 10- ! Taq-, 1 50mM MgCl2, 2 , 10 R , 10 JS 1,5 Taq-. F ! : JS – 950C (10 .), U\V- (35 ): – 950W (20 .), – 580W (15 .), JS – 720W (15 .), ! ! – 720W (5 ). U ! [1, 2]. G ! [1, 2, 3]: U ! : Q U 5 -3 An201 F: AATAACAATCATTACATGAGGCT R: TGGTCAGTTGTTTTTTTATTGAT AfuG412 F: TGACGCACAGTAGTATTATTTATG R: TGATGTTTGCTGAGGCTTTTC AfuG512 F: ATAATAATGAGCGTGCTTTCTGTT R: ATTCCGCTTGCGACTTATTTA AoxD1653 F: TTTGACAGCTCCTAAGTGATACC R: AAAGCCCTACAACAAATGTCAC AoxD1613 F: GTTTGAAATGATTGAGAAAATGC R: TGAGACAGACACTCTAGTTAAACAGC F D- JSP (JSP). H JS ! ! ! - , R . (2008 ). F ! U\V MJ Research PTC-225 (W^F) (J, W, Fermentas .). U\V- ! 1 Taq- (W, R), 2,5 M MgCl2, 2,5 R dNTP (J, R), 2,5-5 M ( . 1), 2,8 - (3,5 M , 50% ), 2,5 . Taq- (W-R), 2 JS ! (milliQ) 15. Y! ! U\V H SY. . 1 (33). 2012 149

: 95 W – 2 ., U\V- (35 ): – 92 W (20 .), – 57 W (30 .), ! – 72 W (30 .), ! ! 72 W – 10 . U ! : U U S H AHR TATACACCATTATCTCTATGT U! ! H AGF GCACAGACTATGTGGTATCCFGGA T ABF CAGATGCCAGTAACAGGCTGA T V (baerii-like) ABRM TGTCTGTCTAGAACATAtG T W

JSP. U ! JS !! , . W ABI 3100 Genetic Analyzer (W^F) BigDye v.1.1. W [4, 5]. RAPD-U\V . U\V ! VXW-225 MJ Research (W^F) AmpliTaq Gold PCR Master Mix. V! , 25 ! 2,5 U\V , 2,5 2 R SX (), 0,25 (100 R/), 1 Xq-, 2 100 JS 16,5 . Y! U\V : ! ! – 94°C (5 .) , U\V- (36 ) – 94°C (30 ), 36°C (30 .), 72°C (1 .), ! ! – 72°C (10 .) . T JS 1,7 % . J! ! JS XHK () – 0,089R X, 0,089R ! 0,002R cJXF. H ! 0,5 / ! . V U\V- ! ! , 5-10% 70% 0,025% GW ( ). c 20 ! ! 10 H/. K-2 «» ! «c-4». I . MQb QJTO TLJ. F D- JSP (JSP). F ! JS. W , 2- – 24 !! «-» , 7 «» JS. W , 4-, 2 «-» , (78 ) – «» . JSP. H ! , , () JS. F , - , , 4- , , 2--, JS. X «-» , . U ! , – , – . F JSP. U ! . T ! , . S ! , ! . S ( 1 4) ! . Y . X , , ! . U ! - , ! ! . J! ! . H ( JS) ! , ! . X , , , , . X , , ! !! ! . RAPD-U\V . J! ! F01, F05, F09 H10. S JS 2 /. U\V ! . S . 1 RAPD- ! JS F05. 150 Y . . 1 (33). 2012

1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20

1 RAPD- J 05 S - . , ! !! . J! ! 20 . H . S . 2 JS F09. V !! ! , ! ! , ! , ! : 2, 4 - 7,9,11 – . V RAPD-U\V BioNumerix.

2 J 09

MQM] U !- ! ! ! - !, ! , , . H – , – . X , , ! !! ! . H ! , VRW. H RAPD-U\V, : CAG-GCC-CTT-C – A01; AGG-GGT-CTT-G – A05; GGG-TAA-CGC-C – A09; CTG-CTG-GGA-C – B10. S , !! . U ! RAPD-U\V (Acipenser gueldenshtaedtii) (A. baerii) . Q 1. Zane et al., Isolation and characterization of microsatellites in the Adriatic sturgeon (Acipenser naccarii); 2002, Mol. Ecol. Notes, 2, 586-588. 2. Welsh et al., Identification of microsatellite loci in lake sturgeon, Acipenser fulvescens, and their variability in green sturgeon, A. medirostris; Mol. Ecol. Notes, 3, 47-55. 3. Henderson-Arzapalo, King TL., Novel microsatellite markers for Atlantic sturgeon (Acipenser oxyrinchus) population delineation and broodstock management; 2002, Mol. Ecol. Notes, 2, 437-439. 4.X S.S., F.K.G, F.H.Y, S.W.R . H (Asipenser gueldenstaedtii) // I - 2009. - 39. - W.1250-1259. 5.X S.S. H . . .., 2009.118 .

H SY. . 1 (33). 2012 151

*** L '' (VFX) '' '''' ' ' - '' . *** In work carrying out of the molekuljarno-genetic analysis of polymorphism of grown up sturgeon fishes for the purpose of definition of suitability of the selected individuals for formation of repair-uterine herd (VR) is considered.

YJ 576.8 .. .ļ. JĽ', J.. Lĭ J TORJĨ Qİ LR R MRJ 2011 ^ (Cyprinus carpio), ' (Carassius auratus), (Abramis brama), (Aspius aspius) '', . T ' 6 ' ''' '''. Tıĳ ı ı – ^ . 1965 ĳ W! Ŀı ı . Wı 100 Ľ 20 , – 775 ², Ŀ – 5,2 ³, ı 6,5 [1]. RQ Lļ ļJ= 2011 ĳ ^ ! ĳ. U! Ľ Ŀı Ľ ĳ [2]. F ! ĳ . Gı Ľ Ŀ.. U! Ľ ı 40 . ļL Lļ T QQ ^ ı .H. Wı [3] ĳ ! ı Ŀ. M 10 ĳ ı 11 ĳ , ı ı ĳ (5 ĳ) Ŀ . ľ ı 70- ĳ ! Ľ [4] ı 12 ĳ 32 ı ĳ . Tı Ľ , !Ľı ı ! ĳ . G, ı Ŀ . 80- ı 14 ĳ ı 20 ĳ . Gı Ľ Ľ ı ĳı ı Ŀ. 90- ĳ ! ı Ľ ı ĳ 70- Ľ 80- ĳ 16-18 ĳ [6]. G : ı ! ĳı (, ), ıı ı Ľ ı ı Ŀ . 2000- ı 66 ĳ , ı 14 ĳ [6,7]. X ! (4 ĳ), ! (11), (21), (7) Ľ (11), (8) , ĳ (1), !Ľ (2) . M ı ĳ : - Thwaitia ovata (20 %), Philometra obturans (10 %), Raphidascaris acus (50 %), Ŀı - Thwaitia ovata (20 %), Philometra obturans (40 %), Rhabdochona hellichi (10 %), Raphidascaris acus (30 %), - Thwaitia ovata (30 %), Rhabdochona hellichi (20 %) Ľ Contracoecum sguali (20 %), Rhabdochona sulaki (20 %) ĳ (1-).

1 ĭ ĩ ĵĭ ĭĩ ĵ ĭĵı ĳ' ĳ''' ı Ŀ Ľ Ŀı . M Ŀı ı 10 ı 4 ĳ , ı Philometra obturans (40 %) . W ı 10 , ı 3 ĳ 152 Y . . 1 (33). 2012

. T ı ı Ŀ Raphidascaris acus (50 %) , 50 %. ļJ 1. F I.R., .H., K.G. ^ ı // L. – 2007. - 311. – W.58-60. 2. G!-U! .K. R . – Q.: S. – 1985. – 120. 3. W .G. R ] // G ! W F . – G. – 1967. – W.256-257. 4. ! ! // T SV / SV. – NIV.73044780. – F-F. – 1975. – W.64-75. 5. T !! ] : T SV / SV. – NIV. 02860064431. – G. – 1985. - W.67-75. 6. F .W., R K.H., T .H., .H. ^ // H SY, .. - 35(51). – F. – 2001. – W.18-23. 7. c , ! ! // V: ^ : T SV / SU\ V. – F. – 2004. – 121. *** U - (Cyprinus carpio), (Carassius auratus), (Abramis brama) (Aspius aspius) ^ . 6 . *** Results of investigation of nematodes distribution in the carp like fishes as carp (Cyprinus carpio), goldfish (Carassius auratus), bream (Abramis brama) and asp (Aspius aspius) in the Sharadara water reservoir were presented. 6 species of parasites had been discovered.

YJ 639.2.053.7 (262.81) .U. , .. , .. , M.. U , .. , .. , J.. I, .. M, J.. TT MUT RT TOTIT UTLb UOTIT RT - !, IYU «SV», . F, V!, e-mail: [email protected]

P . V , , , , , , , , (). S . Y, . ] , , . Y XXI P, , , . . . U , . \ . M , . U P . F . M , . – , , . ( ) . - . W (62,6 %) 100 (238,4 . 2), . T - , . H ! , !! (44 %) , . T !! ! ! !. W !! - ! !! 2011 . H, 36000 . , 6500 . , 18000 . 411 . !. U 2001 . ! (Clupeonella engrauliformis (Borodin, 1904)) (Clupeonella grimm (Kessler, 1877)), ! 99 %. W ! . H SY. . 1 (33). 2012 153

! . H 2001 . ! ! . H ! ! ! . R! ! ! , ! , [1]. G ! , , , , ! ! , , , [2]. U! ! , ! , , !!! ! , ! . F ! ! !!! !! ! . X, 2009 . 48 % 27 % ! , [3]. W 2002 . ! ! - , ] !. H ! ! 99.8 % [4]. !! ! ! ! , ! ! , . ] ! . S !! . U . H 2006 2010 . 2011 . ! 93.3 , – 4.3 , – 18.7 . J 6.0 14 , – 10.5 , - 1.1 20 , – 9.7 . W ! 7 0+ 6+ , ! . G! 9.6 , – 7.3 . T ! (96.4 %) ! , 2.2 , 4.4 % 2006-2010 . H! !! 2013 . ! 6,4 . , – 0,14 . . ! ! (Clupeonella deliatula caspia (Nordmann, 1840)) , ! . , , , . H !! ! , !!! , !, . G! ! ! , ! , ! !!. V , ! . H ! ! ! 85 %. U - W !. U!! ! !. R ! , ! ! ! ! ! W 2,5 214 ./ ! 546 ./ !. U!! ! !. X, «» W 2006 2010 . 298 326 ./ !. H 2011 . 316 ./, W ! 8.4 , – 5.4 . H! 2005-2010 . !. W ! ! ! , ! , . H , , ! . M 15- ! , !! 464 676 . , 540 . . J 2002 ., , ! (438.5 .). H , , , V . H ! !! " , ! !, . 154 Y . . 1 (33). 2012

F , ! !!! - W ! ( . ] – . J). H ! ! , 30-50 , . J! ! ! ! " . ! !! , " ! , ! !! 2013 . ! 56,6 . . U ! ! . J ( 60- .) ! ! ( 80- .) (Alosa saposchikowii (Grimm, 1887)). c ! - ! ! !. W Z . H W ! , W . H W !!! !, 5-7 Wº ! . . H ! ! ! ! . U - ! (). J 14 36 , – 37 630 . T ! ! 21-28 144-265 . U (1935 .) (1970-1971 .), ! ! . W , ! ! ! ! - !. V !! !!! !, ! , ! ! . U ! !, ! 1978 ., . H W 45-46 . 26 . 1969-1979 . 30- , 1,26 ./ ! 1970-1980 . 10-12 . U ! ! ! 15 %. H , . H .. , ! , ! . T ! 2,5 . . ! !! ! (Alosa braschnikowii (Borodin, 1904)) (Alosa caspia caspia (Eichwald, 1838)). J! !!! . : : 12 20,4 /, 17,8 /. Y 1,6 5,4 /, 3,5 /. c ! ! . H ! , . X, 2011 . ! , ! . S, ! !. H 2011 . . F! ! ! . U ! ! !! W !. T . T, , ! ! ! . H ! W !, ! ! !. U! !! ! , ! ! ! ! . X , , . V! ! H SY. . 1 (33). 2012 155

W ! ! !, !! ! . H !! 2013 . ! 2,5 . ! 6,8 . - ! . F (Atherina boyeri (Risso, 1826)) , !. F - !, ! . W !. U! . W ! W W ! 2006 . V !, !. T . F . 0 0 H! 6 25 W 0 16 /00, 0 0 3 12 /00. S! 4,2 /00. F ! . H ( ), (, ), (, ) . H ! ! ! . F !!! , !, ! . c! ! , , . U! ! !!! ! . X , ! ! ! ! ! . U 6 , ! – – . W ! ! 3.8 , 9-13 . U! , - 2-3 4.5-9.6 . H W W !, , !!! . G . R 4 (1044 ./ !). U ]! W!! L!. I! ! ! . J! ! J ! . U , . H . H! !! 2013 . ! 7,0 . . W (Liza aurata (Risso, 1810)) – , ! ! !-! !. W X- ! ( J) -!. U ! – , - ! . W !!! , ! , ! ! ! , , !. Y ! 2000-2010 . 1.5 849.4 , 158.6 . H (2009-2010 .) !! ! !, H W ! 15,8 ºW. S X 5.3 /, ]! 1.3 /. V 19 46 , – 36.9 . R ! 0.21 1.27 , 0.75 . U ! ! ( 70 %). T - ( -!) ! W W . H ! ! ! 36-40-45 – 50 : . Q !!! ! . U ! 2013 . ! 2,0 . . Q 1. J.S., I G.S., J.H. J W ! 2001 .// V! ! : V SV 2001 . - F: - - SV,- 2002. - W.41-55. 2. I U.U., R I.., ^ I.., U Z.S., F H.U. . // V! ! : V SV 2001 . - F: - SV, 2002.-W.510-517. 3. H K.F., J! F. H. !! (Clupeonella ngrauliforvis), «!» . // H!, 2009. - 3 9. - W.55-58. 4. F.R. T ! ! Mnemiopsis leidyi (A. Agassiz) (Ctenophora: Lobata) . – F. . … . . . - F., 2005. – 23 . 156 Y . . 1 (33). 2012

*** R P ''' '' %'' '' . R %''' ' – %'' %', '' , , P , % ' () , , . F %'' '' %'' '' '. *** Status of the main commercial marine fish stocks in the Caspian Sea is analyzed in the work. Problems of distribution, abundance, qualitative population structure of common sprat, anchovy kilka, big-eyed kilka, Dolginsk herring, Caspian shed, big-eyed shed, silverside and mullet (long-finned mullet) are considered. Based on received materials recommendations for presumable size of commercial catch of concerned biologigal water object are given. It was stated that modern sprat fishery catches minimum amount of anchovy and big-eyed kilka. Their number has extremely shortened, the effectiveness of natural spawning has reduced, commercial stocks are coming down rapidly. Environmental degradation in the Caspian Sea starting at the beginning of the 21st century, caused by the immigration of ctenophoran mnemiopsis, waterquakes and water eutrophication conduced diseases of anchovy kilka, which appeared in appearance of splanchnic tumors. The volumes of commercial catch are not going to increase in the short term. Effectiveness of reproduction of common spart is annually set at the long-time average annual level. Commercial stocks are increasing, it is recommended to increase the size of commercial catch of this species. It is reasonable to increase the number of commercial fishing gears at Dagestan seaside. The stocks of sea herring are at the stable level, dolginsk herring has the most abundance. The replenishment of populations is determined by existence of forage organisms and water temperature in the southern part of the Caspian sea. Silverside is caught by common sprat harvesting. The stocks are stable, it is recommended to increase species catching. There is only one mullet species is known in harvesting, it is long-finned mullet. Its stocks are stable, the volume of commercial catch is determined by catching management. Raw material resources of marine fish (except for anchovy and big-eyed kilka) let to increase their commercial catch.

1.2 IJTTQTI YJ 576.895 1.R. , 2.. M] MR QI TMJTTQ TTUT IT\TMT T IQbRT 1XTT « - » FT «F!» 2U

V . F ( ) : ( – ) . ! U 3 – Pleurogenes intermedius, Haplometra cylindracea, Opisthioglyphe ranae – Rhabdias bufonis Oswaldocruzia filiformis. . ! ! Sphaerirostris teres, (Strigea strigis, S.falconis), ! , , . Alaria alata, U . G ! ( ! !) : 1) ! ! ( ! ! !! !); 2) ! ! , !! ! ! ! -; 3) ! , !! ! , , ; 4) ! ! – ( , !, ! ). U !! ! ( H.F.J [1]), ! , ! ! ! . H Z.H. Q..G [2] ! – . U – ( ! ); – , . T , , ! , ! ( !), !. W , ! ! ! , [3], ! ! ! ! . H SY. . 1 (33). 2012 157

U !, ! ! ! !. X.R.G [4] , Fasciola hepatica ! ! ! Haplometra cylindracea, !. W , . M , , ! . c ! ! ! , . G , !, ! , !, , ! , . U , U Alaria alata, ! ! ! . T, , U , . , : 1985-1987 . 1993 ., 2004-2011 . ! ! 2005 . J , Rana arvalis U , ! . \ ! Sphaerirostris teres, ! ! 1993 , 100%- , . (Strigea strigis, S.falconis), ! ! 1984 2011 . , ! ( ) . c , , , ! ! ! . ! U 3 – Pleurogenes intermedius, Haplometra cylindracea, Opisthioglyphe ranae – Rhabdias bufonis Oswaldocruzia filiformis. P.intermedius ! , ! . J , , !! Plagiorchidae – H.cylindracea O.ranae – 80- . H 2005-2011 . . . Y O.ranae ! 2006 ( 2-3 ! !), H.cylindracea – 2007-2008 . ( 50 . ! 100%- ). H . H ! Lymnaeidae ! . H . - Lymnaea stagnalis, L.fragilis, L.auricularia, L.palustris, L.ovata, L.glutinosa, L.palustris, L.peregra, L.tumida, Radix sp. [5], a !! ! L.stagnalis L.auricularia. R (L.truncatula) , U . ! , , ! (Plagiorchis elegans, P.arvicolae, Notocotylus noyeri), ! . S Rhabdias bufonis, ! !, ! , [6]. X , , . Oswaldocruzia filiformis, , !!! ! : ! ! !, ! !, ! ! !, ! [6]. W ( ! ) ! . , [7], Q3 - ! , ! , ! ! . T, ! O.filiformis !, ! 158 Y . . 1 (33). 2012

. Q! !!! ! !, , , ! ! , ! ! ! . U ! ! ( !) ! !; . U !, ! ! , ! . J! ! ! ! , , !. X , ! ( , U , !!! ! !) : 1) ! ! ( ! ! !! !); 2) ! ! , !! ! ! ! -; 3) ! , !! ! , , ; 4) ! ! – ( , !, ! ). U ! , ! , : 1) ! !; 2) ; 3) ; 4) ! . U – , ! «! !» . H ! , . H ! ( ) !! ! , : 1) ! ; 2) – ! ; 3) , ( ) !! ! . M ! !!! ! , , . ! ! , , ! ! ! . W ! « » – ! ! , ! ! !. V ! . U , , ! , ! ! . Q 1. J H.F. . - Q.: Y, 1941. – 287 . 2. Z.H., G Q.. T , ! ! // ! ! : , U, 3-5 . 1990 . - R., 1990. – W. 75-77. 3. M H.. W . - II H. . X . - : S , 1983. - W. 118-119. 4. G X.R. Haplometra cylindracea (Zeder, 1800) . F. . . – R., 1986. 5. I X.H. U (W ) //III R! ! ! «G », 4-6 !! 2005 ., J. – J, JSY, 2005. – W. 30-31. 6. V .R., ^ H.U., ^ S.S. I WWWV. – R.: S, 1980. – 279 . 7. X! S.K., X W.H. V ! ! Oswaldocruzia filiformis //R R - «G, - », U, 22-23 !! 2009 . – U, 2009. – W. 103-106.

H SY. . 1 (33). 2012 159

*** L %' ' '' '' . % %''' ' ' '' (' ) '' ' ( '' ' ' % ') % % ' '' ' ' . U '''' ' 3' - Pleurogenes intermedius, Haplometra cylindracea, Opisthioglyphe ranae % 2 ' – Rhabdias bufonis % Oswaldocruzia filiformis ' ''' '. K' '' , ' '' %' Alaria alata , Strigea strigis % S.falconis , Sphaerirostris teres %'' . F '' Alaria alata , ' U ' . ( '' ' ) ' ' ' ' : 1) ( '''' ' ''' ) %' ' '; 2) % -' %' , ' '' ' ' '; 3) ' () , , ' ' '' % ' '; 4) ' ( ') ' ' %' ' . *** The role of different taxonomic and ecological groups of the wild animals as the eliminators of helminthes of domestic animals was considered. The participation of anural amphibian (on the example of moor frog) for the sanitation of the pastures and near-water landscapes from the helminthes of domestic and hunting animals: as the direct eliminators of non-specific parasites (including the consuming of insect and other invertebrates as intermediate hosts) and as the hosts of competitor worm species – was analyzed. Among the mature helminthes forms in the moor frog from the most landscapes of Pavlodar region 3 trematodes species Pleurogenes intermedius, Haplometra cylindracea, Opisthioglyphe ranae and 2 nematodes species – Rhabdias bufonis Oswaldocruzia filiformis parasitized. Among the larval helminthes stages in the flood-land of Irtysh river cystacanthes of Sphaerirostris teres, metacercaria Strigea strigis, S.falconis which are harmless for people, fishes and home animals were recorded. We can rank to the potentially dangerous parasites only larval stages of trematode A.alata which have increasing of the quantity in Pavlodar region so rarely. Brown land frogs (including moor frog) can play the noticeable role in the improvement of sanitary condition of pastures and near-water landscapes from the helminthes by several positions: 1) as the direct eliminators of larval stages of pasture parasites of home hoofed animals from suborder Strongylata and other nematodes with the simple life-cycle; 2) as the definitive hosts of harmless for the people nematodes which are the competitors of pasture parasites from suborder Strongylata; 3) as the definitive and intermediate hosts of several trematodes species which are the competitors of the trematodes of hoofed animals, aquatic birds, fishes on the stage of parthenogenetic larvae in fresh-water gastropod snails; 4) as the eliminators of larval stages of several practically important helminthes in the intermediate hosts – snails and arthropods (Eurytrema, Dicrocoelium, suborder Spirurata).

YJ 577. 472 (28) .. 1, .. I2, R.. 3 IJTZ TM 1 S F F, 2,3R F, e:mail: [email protected] 2011 . , .P. 34 , 34 (Rotatoria – 18, Cladocera – 10, Copepoda – 6 ) 84 , 11 . R . T W -F .. T! F !!, ! W, ! ! . U ! 6700 . T ! G-I ^-I. H 2010 . . ! , . H ! ! 2011 . . W . . W , W !!! ! . R ! [1,2]. W! W [3,4] . W 34 – ! , , , , , , , , , , , , ! !, -, , ! !, , !, !, , !, ! , , !, , , , , !, !, , ! !, , . H W 34 , !! 3 (Rotatoria – 18 , Cladocera – 10 , Copepoda – 6 ) ( 1).

160 Y . . 1 (33). 2012

X 1 M 2011 . M Q T Rotatoria 18 6 18 18 10 Cladocera 10 4 10 10 8 Copepoda 6 3 6 6 4 H 34 13 34 34 22 H W 84 , !! 11 ( 2). T ! W ! , ! 19 % , (15,2 %) (11,8 %), . S , . R (80-84) -, - (20-36) - . W , ! , ! 16 , – 14 , – 11 – 8 . X 2 2011 . 3 M Q T Oligochaeta 1 Stylaria lacustris (L.) + + - - 2 Nais communis Pig. + + + - 3 N.elinguis Müller - + + - 4 Pristina aequiseta Bour - + + - 5 Aulodrilus limnobius Bret. - + + - 6 A.pigueti Kowal + + + + 7 Limnodrilus udekemianus Clap. + + + + 8 L.hoffmeisteri Clap. + + + - 9 Potamotrix hammoniesis Mich. + - + - 10 Tubifex tubifex Müller - + + - 11 Eiseniella tetraedra Sav. - + + + Hirudinea 12 Protoclepsis tessulata Müller - + + + 13 Haementeria costata Müller - + + - 14 Helobdella stagnalis (L.) - + + - 15 Piscicola geometra (L.) - + + + Mollusca 16 Lymnea stagnalis (L.) - + + - 17 L.auricularia (L.) + + + + 18 L.ovata Germ. + - + + 19 L.palustris Müller - + + - 20 Costatella acuta Drap. - + + - 21 Planorbis planorbis L. - + + - Amphipoda 22 Dikerogammarus haemobaphes Eich. - + + - 23 Pontogammarus robustoides Grimm - + + - 24 P.sarsi Sow. - + + - 25 P.aralensis Schaf - + + - 26 G.balcanicus Der. - + + - Hydrocarina 27 Eylais hamata Koen - + + - 28 E.degenerata Koen - + + - Odonata 29 Lestes sponsa (Fabr.) - + + + 30 L.barbara Fabr. + + + + 31 L.nympha (Selys) - + + + 32 Enallagma cyathigerum (Charp) - + + + H SY. . 1 (33). 2012 161

33 Agrion virgo (L.) - + + + 34 Sympycna fusca V.d.L. - + + - 35 Platycnemis pennipes Pall. - + + - 36 Coenagrion puella (L.) - + + - 37 C.scitulum Ramb. -++ - 38 Anax imperator Leach. - + + - 39 Sympetrum flaveolum (L.) - + + - 40 S.danae Sultz - + + - 41 S.striolatum (L.) - + + - Ephemeroptera 42 Palingenia longicaudata Ol. - + + + 43 P.fuliginosa Boeb. - + + + 44 Ephemera vulgata L. - + + - 45 Heptagenia perflava Brod. - + + - 46 0sonychia ignota Walk -++ - 47 Cloen dipterum (L.) - - + - 48 Baetis rhodani Pict. - + - - 49 Ordella macrura Steph. - + + - Coleoptera 50 Peltodytes caesus Durf. - + + - 51 Noterus crassicornis Müller - + + - 52 N.clavicornis (Deg.) - + + - 53 Bidessus signatellus (Klug.) - + + - 54 Hypophorus musicus (Klug.) - + + - 55 Hydroporus planus (Fabr.) - + + - 56 Porphydrus lineatus (Fabr.) - + - - 57 Gyrinis minutus Fabr. - + + - 58 G.caspius Men. - + + + 59 G. suffrioni Scz. - + + + Hemiptera 60 Notonecta lutea (Müll) - - - - 61 N.glauca (L.) - + + - 62 Corixa punctata 0llig. - + + - 63 Hydrometra stagnorum (L.) - - + + 64 Sigara falleni Fieb. - - + + Trichoptera 65 Ecnomus tenellus Ramb. + - - - 66 Hydropsyche ornatula Mel. + + + - 67 Leptocerus tineiformis Curt - + + + 68 Oecetis furva Ramb. - - + + Chironomidae 69 Stempelina bausei Mg. + + + - 70 Chironomus plumosus (L.) + + + - 71 Ch.dorsalis Mg. + + + - 72 Chironomus thummi Kieffer + + + - 73 Cryptochironomus defectus Kieffer + + - - 74 C.conyugens Kieffer + + - + 75 Cryptopladelma virudula Kieffer + + + + 76 Leptochironomus tener (Kieffer) + + + + 77 Einfeldia pagana Mg. - + + + 78 Polypedilum nubeculosum Mg. - + + + 79 P.convictum Walker - + + + 80 Anatopynia plumipes Fries - + + + 81 Procladius choreus Mg. - + + + 82 P.ferrugineus Kieffer - + + + 83 Tanypus punctipennis Mg. - + + + 84 T.vilipennis Kieffer - + + + H 20 80 84 36 162 Y . . 1 (33). 2012

H !! . X ! . T ! , , ! W, , ! ! . U : Branchiura sowerbyi, Lymnaea auricularia, Costatella acuta, L.stagnalis, Pontogammarus robustoides, Coenagrion acitulum, Sympetrum striolatum, Chironomus plumosus, Ch.dorsalis, Ch.thummi, Cryptochironomus defectus, Polypedilum nubeculosum . W , W ! ! ! ! , , . . Q 1. L H.. R ! // L WWWV, 1956, . 4, . 1, . 226 – 288. 2. U .. V . – U ., R., 1966, 376 . 3. F.I. U! . – G: c. 1999. - 275 . 4. F F.S. Rotatoria // L F. G, 1951, .386-394. *** 2001 P ''' ''' '''. M %' 34 ', 34 ' (Rotatoria – 18, Cladocera – 10, Copepoda – 6 ') % 11 ' 84 ' . T' ' % '' . *** Studies on the hydrofauna of the Sarysu Lake on the right bank of the Kura River were conducted in 2011. As a result of studies there are registered 33 species and 2 hybrids of fishes, 34 species of zooplankton and 84 species of makrobentic organisms belonging to 11 taxonomic groups. Maximum development of organisms was recorded in spring and summer.

YJ 574.5 .. J T MTTM RTMTTT RTIT TJTQ_ 2005-2009 ITJ F SV, Y-, , e-mail: [email protected]

U , , G 2005-2009 . T 66 . X – Borystenia naticina, Agraylea multipunctata Tipula pierrei . J , . G !!! ! , ! !! . R 60-70- ! F ! S ! X W.. H-^ Q.U., 70-90- . ! ! F.Q. , !. W ! 1966-1973 . 10 , Paramysis lacustris Paramysis intermedia, ! Mysis relicta, Gmelinoides fasciatus Micruropus possolskii, U Pallasiola quadrispinosa. H 2009 . ! «W G », [1]. H – Unio pictorum, Viviparus viviparus Lithoglyphus naticoides, Micruropus kluki [2, 3]. V 2001- 2004 . SV [4]. S!! ! ! ! 2005-2009 . RQ RTJ QJTO ! ! [5]. H 259 , 124 135 , 18 . U. S () 2 ! 0.3 2. ! !, [6-11]. G ! ! . MQb TLJ H 66 , 15 , 3 – ! , 4 – , 5 – , 2 – , 1 – , , , 27 , , ! (. 1). H ! Borystenia naticina, H SY. . 1 (33). 2012 163

2009 ., Agraylea multipunctata, , Tipula pierrei, . X 1 2005-2009 . ] , % T- I- X T! I! ! ! Mollusca Unio pictorum (Linnus) 9 18 + + Anodonta piscinalis Nilsson 3 + + 10 Pisidium amnicum (Muller) - 12 - - Sphaerium nucleus (Studer) 1 - - - Euglesa sp. 3 - - - Borystenia naticina (Menke) 1 - - - Valvata depressa C.Pfeiffer 5 + 12 15 Valvata pulchella Studer - - 4 - Valvata piscinalis (Muller) - - - 5 Viviparus viviparus (Linnus) 6 12 + - Lithoglyphus naticoides C.Pfeiffer - 59 38 - Bithynia leachi (Sheppard) 1 - 4 - Lymnaea auricularia (Linnus) + + + + Lymnaea ovata (Draparnaud) + - + + Anisus acronicus (Ferussac) 5 - 12 5 Oligochaeta 79 88 83 100 Hirudinea Piscicola geometra (Linnus) 5 6 29 10 Hemiclepsis marginata (Muller) + - - - Erpobdella octoculata (Linnus) 1 - + 5 Mysidacea Mysis relicta Loven - - - 24 Paramysis lacustris (Czerniavsky) 58 50 44 43 Paramysis intermedia (Czerniavsky) 58 50 26 - Amphipoda Gmelinoides fasciatus (Stebbing) 14 12 33 35 Micruropus possolskii Sowinsky 9 - 33 5 Micruropus kluki (Dybowsky) - - 12 - Gammarus sp. - - - + Odonata Agrion hastulatum Charp. - - - + Erythromma najas Hans. - - + - Ephemeroptera Potamanthus luteus (Linnus) - + - - Ephemera orientalis McLachlan - 6 - 20 Caenis horaria (Linnus) - - 8 - Caenis miliaria (Tshernova) 3 - - - Brachycercus harrisella Curtis - 6 - - Heteroptera Nepa cinerea Linnus - + - + Trichoptera Ecnomus tenellus (Rambur) - 6 4 20 Agraylea multipunctata Curtis - - - 5 Tipulidae U 1 164 Y . . 1 (33). 2012

Tipula pierrei Tonnoir - - + - Psychodidae - - + - Ceratopogonidae - 6 8 5 Chironomidae Procladius sp. 4 6 21 35 Ablabesmyia . monilis - - - 5 Cricotopus . silvestris 4 24 12 5 Orthocladiinae - 12 - - Tanytarsus sp. 3 12 4 - Cladotanytarsus . mancus - 6 4 5 Cladotanytarsus sp. 1 6 8 30 Chironomus plumosus (Linnus) 38 35 29 35 Chironomus tentans Fabricius - 6 4 - Chironomus sp. 14 12 4 10 Cryptochironomus . defectus 6 6 8 20 Cryptochironomus sp. 3 6 4 5 Parachironiomus . pararostratus - 6 4 - Parachironiomus vitiosus Goetghebuer - - 4 5 Lipiniella arenicola Schilova - 6 - - Limnochironomus nervosus Staeger 5 - - - Limnochironomus sp. 9 6 8 20 Endochyronomus albipennis (Meigen) 3 - 4 5 Endochyronomus tendens Fabricius 2 - - 10 Glyptotendipes gripekoveni Kieffer 3 - 8 - Glyptotendipes glaucus (Meigen) 2 - - - Polypedilum . nubeculosum (Meigen) 3 6 21 30 Polypedilum . scalaenum (Schraenck) - - 4 - Polypedilum . convictum (Walker) - - 8 10 Polypedilum sp. - 6 12 5 Pentapedilum . exectum 1 - - - Chironomini - - 4 5 H 36 34 44 38 U – «+» , R - (34 ), – - (44 ). U Anodonta piscinalis, Valvata depressa, Lymnaea auricularia, , ! Piscicola geometra, Procladius sp., Cricotopus . silvestris, Cladotanytarsus sp., Chironomus plumosus, Cryptochironomus sp., Limnochironomus sp., Polypedilum . nubeculosum, ! P. lacustris G. fasciatus, U. pictorum. H . X, V. viviparus ! - !, -; L. naticoides - - !. U! P. intermedia , , ! M. relicta, , . G M. possolskii ! - ; M. kluki, 2006 ., - . ] ( 79-100%), P. lacustris (43-58%) P. intermedia (26-58%, ), C. plumosus (29-38%), G. fasciatus (12-35%), - - ! – L. naticoides (38-59%). Q 1 J! H.. R. // W G – T - 2009 - W. 95-119. 2 J! H.. G – H . // c - F, - 2004 - . 1 - W.83-86. 3 J! H.. G – H- // K ! ! - V--J – 2007 - W. 106-108. H SY. . 1 (33). 2012 165

4 J! H.., K F.F. W! G . // V! ! V : ! ! - F: 2005. - W. 417-427. 5 R ! ". M ! - Q. 1984.-51. 6 L H.. R WWWV. - R.-Q. 1952. -376. 7 T K WWWV.- Q. 1977 -512. 8 T V . V, - W.-U. 1995 -628. 9 U H.. Q Podonominae Tanypodinae WWWV. - Q. 1977 -154. 10 U H.. Q Chironominae WWWV -. Q. 1983 -296. 11 ] F.F. T Tendipedidae - R.-Q. 1949. - 186 . *** R 2005-2009 G ' , , % ''. 66- . ' – Borystenia naticina, Agraylea multipunctata % Tipula pierrei ''' *** Data on species composition, dominant groups, abundance and biomass of benthic macro invertebrates of Bukhtarma reservoir in 2005- 2009 are given. About 66 species are determined. Three species – Borystenia naticina, Agraylea multipunctata and Tipula pierrei are recorded in Bukhtarma reservoir for the first time. Oligochaetes, larvae of chironomids and mollusks dominated.

YJ 574. 582 .. MTTUQT b-RTITTIT TJTQ_ H- , : 76 73 91, e-mail: [email protected]

Y-P , . Y- 1952 . ! ! , !. V H- . M ! 71 , 37 2, " 0.65 3. ^ 400-750 , ! 1200 . H , !! ! 17 . I 6 46 . Y- ! : 27-41 ( , 23 ). V -. U , ! ! : – G IcW (GIcW) U ( ); – U R!; ( ) – R! Y IcW. H!! GIcW . W! ! !, . S . T ! T . Y- – , !! G , ! 8.0 0W. H ! , 22.0 0W . Y- !!! ! ! . T – , , [1]. RQ RTJ QJTO ! 2004 . F «SV» H \ . C 2005-2011 . . ! Y- - 2005-2006 . F «S- !», - 2009-2011 . H \ . H 7 (19 ), 105 . «R ! ! (, )» [2]. T !. J! !, K.H. G I.I. H [3]. Y ! « » W.U. [4]. H , , . U Y- . 166 Y . . 1 (33). 2012

U ! H.F. (1958., 1964-1966 .). V « Y- » [5]. H ! Y- 1956-1963 . . H (1956-1960 .) , ! , (!! 4.4 . /3 0.14 /3). c !! , ; ; , , ! ! , . H (1962-1963 .) ! (!! – 117.8 . /3 2.4 /3). U, , : G ! ; . U H.F. [5] Y- Kellicottia longispina, Keratella cochlearis, Synchaeta sp., Bosmina longirostris. S! Mylilina spinifera, Platyias militaris, Alona gutata . U ! G K. longispina. U , Sida crystallin. X , , ! 2-3 , Y- ! 10 . H 5 , . H ! , !! ! J J. H 1964-1966 . K. longispina, B. longirostris, Daphnia hyalin, Mesocyclops oithonoides. H , Y- ! ! : 17.7 . ./3, – 1076 /3; – 122.2 . ./3 3427 /3; – 61.5 . ./3 1197 /3. H , – , – , ! . U ! ! F.Q. [6] 1993 . ! , 25 30 . U ! 12 (Brachionus angularis, B. bakeri, Brachionus sp., Keratella cochlearis, K. quadrata, Notholca longispina, Asplanchna sp., Synchaeta sp., Polyarthra sp., Filinia sp., Lepadella sp., Mytilina sp.) 12 (S. crystallina, Diaphanosoma brachiurum, D. pulex, D. longispina, D. cucullata, Ceriodaphnia reticulata, B. longirostris, A. gutata, A. restraugula, A. affinis, Chydorus sphaericus, Leptodora kindti). , : Mesocyclops dubowekii, M. rylovi, Cyclops abyssorum, Macrocyclops albidus. H – 318.7 . /3 680 /3. T! – 49% , – 36% . U Y- . H 1990-1996 . ! Q.G., J! H.. [7,8] H ! , , ! . H 17 : – P. dolichoptera, A. priodonta, K. cochlearis, K. quadrata, K. longispina, Filinia aseta, - Neutrodiaptomus incongruens, C. vicinus, M. leuckarti, Thermocyclops crassus, - D. brachyurum, Simocephalus vetulus, D.cucullata, D. galeata, D. hyalina, B. longirostris, Leptodora kindti. H 1990-1996 . : P. dolichoptera, D. longispina, K. quadrata, Thermocyclops crassus, M. Leuckarti. S G Y- IcW. T (90%) (88%) ! , . H 1995 . ! 12.4 . /3, - 109 /3; 1996 . – 10.3 . /3 159 /3 [7,8]. H 2005-2011 . Y- 38 , : - 19 , - 6, - 13. J : P. dolichoptera, Asplanchna priodonta, K. longispina, K. cochlearis, K. quadrata; - Cyclops vicinus M. leukarti; – D. brachyurum, D. H SY. . 1 (33). 2012 167 cucullata, Ch. schaericus, B. longirostris. X Y- 1. X 1 (%) - 2005-2011 . X ] , (%) 2005 . 2006 . 2009 . 2010 . 2011 . Rotifera Rotifera gen sp. 33 83 3 4 - Notommata saccigera Ehrenberg -67-- - Eothinia sp. --311 - Trichocerca sp. -17-- - Synchaeta sp. 17--- - S. pectinata Ehrenberg 33--- - S. kitina Rousselet - 17 3 - - S. tremula (Muller) - 17 - - - Polyarthra dolichoptera Idelson 83 100 30 70 30 Bipalpus hudsoni (Imhof) - 17 9 - 11 Asplanchna priodonta Gosse 58 100 64 71 33 Lecane sp. ---4 - L. luna (Muller) 8 50 - - - Keratella cochlearis (Gosse) 58333930 67 K. quadrata (Muller) 58 33 76 48 41 Kellicottia longispina (Kellicott) 92 83 88 67 93 Notholca acuminata (Ehrenberg) 17 33 - - - Conochilus sp. 8 - 36 15 26 Filinia longiseta (Ehrenberg) 8--- 7 Copepoda Neutrodiaptomus incongruens (Poppe) 8 - 24 7 4 Macrocyclops albidus (Jurine) 17 - 6 4 - Cyclops vicinus (Uljanine) 58 83 94 89 67 Mesocyclops leuckarti (Claus) 506752100 93 Thermocyclops crassus (Fischer) 25 - - 4 - Harpacticoida gen sp. --94 - Cladocera Sida crystallina (Muller) - - 6 - 4 Diaphanosoma brachyurum (Lievin) 50 - 58 37 41 Ceriodaphnia quadrangula (Muller) 8 - 9 7 7 Daphnia sp. ---4 - D.cucullata Sars 33 - 30 37 52 D. longispina (Muller) 18 4 - - 7 Chydorus schaericus (Muller) 25 33 18 22 4 Alona quadrangularis (Muller) - - 3 - - Graptoleberis testudinaria (Fischer) - - 3 - - Acroperus harpae (Baird) 17 - 3 - - Macrotrix laticornis (Jurine) - - 18 4 - Bosmina longirostris (Muller) 33 33 39 63 67 Leptodora kindti (Focke) 8 - 12 22 11 H - 24 17 26 23 19 U ! Y- !! ! . U !!, !. X, 63% , – 50% – 44%, , , 36%, 47% 55%. H 2005-2011 . ! 205-973 /3, , . : - ; : - !- ( 2). X 2 ] (N, . ./3) (, /3) - 168 Y . . 1 (33). 2012

2005 . 2006 . 2009 . 2010 . 2011 . I ( - ) ( ) ( -) ( -) ( -) M N H N H N H N H N H Rotifera 19.9 158 24.9 275 7.5 24 7.1 9 25.9 70 Copepoda 53.6 704 33.8 223 17.6 271 15.5 140 15.4 201 Cladocera 6.1 111 24.5 103 6.0 184 2.4 56 14.5 366 H 79.6 973 83.1 601 31.2 480 25.0 205 55.7 637 , , , , , !-- !-- -- - !-- [4] R ! : 2009 . «F 8» - 3131 /3, . vicinus N. ingongruens.; 2011 . – «T 4» - 1600 /3, B. longirostris. H. M Y- . M 30 . M! ( 1 /3), ! (1962-1963 .), ! 5 /3 G . H ! . H , . M !!! , Y- . Q 1. T - «c , ! ! ! . V: H- ( ) 03.03.03.S3», 3 IV (V) 0101V00134 2. ^ Q.., F.U. R ! ! (, ). – F, 2006. – 27 . 3. G K.H., H I.I. M //T ! . – Q.: S, 1979. – W.169-172. 4. W.U. T « » //X. . V HIGT, . II. – , 1986. – W. 254-255. 5. H.F. Y- . F . F-F. 1967. – 23 . 6. T - «G ! !. V: H H ». - Y- , 1994. – W. 75-80. 7. K ! ! 1995 // H- , . Y-, 1995 . – W.17-23 8. K ! ! 1996 // H- , . Y-, 1996 . – W.20-23 *** R & '''' ', ''' ''. *** In article dynamics of quantity indicators of a zooplankton of Ust-Kamenogorskogo of a water basin is considered, the list structure is resulted

YJ 574.5 I.. , M.. QbT UJQ UQT Z\TTR IUIQTR TM TQbT TT, QOO O WT VFS, . G, [email protected] , . . G (F ) , . J H !! ! , ! ! ! . ! . J! , !, ! , ! . c 100 / , ! H SY. . 1 (33). 2012 169

( Artemia sp.) , ! ! ! !. H M W «!» , ! 51-56º 61-82º . H [1] ! !, ! ! , ! ! 80- , ! - ! ! . I M W, , , , , . H ! !! . G – (! 10 ) , . T , ! ! . H ! ! , , ! , . T! ! !!! ! . RQ RTJ QJTO W 19 ! (30 ) 31 ! (33 ) 2010 . T ! W. S 1 ! , ! , . W ! , [2]. F .

. 1. . ( ). J 0.2h, 0.6h, 0.8h , h – . MQb TLJ . . U F ! [3] G - 100W ! 2200–24000W IX = 0.8–0.6. U ! 155-160 , -500W. W!! 25 . G ! 115-120 , ! ! 253 , – 140-175 . W!! 5 /. T !!! \– . 25 , . U ! 4.0 – 4.4 ( 6.8–8.4) ! 63.0 66.7 2, – 251 300 . 3 [4]. J 11 , 7.9 . H , ! – –. Y !, ! ! . T !!! 1210 2 [5]. G ! , 10–25 , ; – , . Q !, ! ; 250–2000 , , . H !! . W Cl– Na+. S ! Mg2+ Ca2+. M 1957–1958 .. ! ! 188.50–203.13 /, 80 . 171.89 /, 1996 ! 154.31 / [4]. H 2007–2008 . 150–156 / [6]. . H ! (2 ) ! 11.2-13.90W. X -5.80W . H ! ! ! : 22.60W (. 1). X ! 5 6 , 10W – , ! . S ! -2.40W . X , . G 170 Y . . 1 (33). 2012

– . U [7] H . X ! (004 026, 006 027, 007 029) 1.5-20W, , . U W, ! , – 0.5- 0.6 . U ! 1.8 4.8 . M. H ! . G (.2). R ! ! . H . U!! , , - . .2. . , 19.05.2010: 25 4-7 , 30 , 26-29 ; – 0 , – 1.0-5.0 , – 5.1-7.0 20 .

15 H ! ! 10 114.5 ./ (. 3). H ! : , , - 5 . U –

, ./ , 0 4 6 8 7 30 29 28 27 26 , !! 30.7 ./ ( 2.6–114.5 ./) 0.95 ./ ( 0–13.7 ./), . 140 V.3. N 120 . G , 31.07.2010: 2-7 , 30 , 26-29 100 ; – 0-2.0, – 2.1-5.0 , – 5.1–7.0 , – 80 7.1-9.0 . , ./ ./ , U 60 0.05 ./ ( 0–0.3 ./) 0.38 40 ./ ( 0–1.4 ./), . 20 H 0 5.5 (. 4-). U 2 4 6 8 7 3029282726 : 5.5 , – 5.5 , – – 3.5 (. 4-). V. 4. . G (. 030) 31.07.2010 .: – , – W ! ! ! . G - , F ! : H SY. . 1 (33). 2012 171

(0-2 2.1-5 ), (5.1-7 7.1-9 ). U! . V ! (=0.001) ! (=0.018) ! . ! , . H ! ! , ! . W! , ! [8]. T ! ! . G ! [9]. Y, ! ! 7.5 , ! – !. W (Q.F. J, ). R !! ! , ! [10]. Y 5-6 ! , ! , -, ! ! ! ! . X « ! » (IFD – ideal free distribution with costs) . T . V! ! , , , , ! , . H !! , . H ! ! [11]. X , . G ! ! !, ! ! . V . Q 1. Q Q.., Q F.., G K.I. F! M W. – S: S. – 2009. – 304 . 2. V / U . H.F. F – WU: I. – 1992. – 318 . 3. F F !. – Q.: I. – 1971. – 155 . 4. W H.U., U R.F., ! X.Q. L !: ! ! . – G: TFT «F ». – 2001. – 144 . 5. V ! / U . H. F. Y. – Q.: I. – 1962. – H. VI. V F ! ! S . – 978 . 6. G Z.F., \ I.F., K I.. J . G // H FIFY. – 2009. – 3 12 . – W. 57–62. 7. W.U. T ! . – U: VFS. – 2007. – 395 . 8. I R.Z. H! ! U!// ! W VFS. – 2007. – X. 9. – 3 4. – W. 973–986. 9. F.H., X.H. U! G // R , , !. – 2011. – 3 6 (31). – W. 422–428. 10. I R.. W! ! W ! // ! WT FS WWWV. W. G. S. – 1990. –H. 3. – W.78–85. 11. Lampert W. Vertical distribution of zooplankton: density dependence and evidence for an ideal free distribution with costs// BMC Biology, 3:10. – 2005. – http://www.biomedcentral.com/1741-7007/3/10. *** R '' ''' '' ' ' . L '' (F '') , ' ' '. *** In article it is considered vertical plankton distributions at a temperature gradient, as an ode from the major characteristics of an ecosystem of shallow salty lakes. During the summer period in the lake Big Summer (Altay territory) is established thermal stratification which causes the authentic stratified distribution of population Artemia sp.

172 Y . . 1 (33). 2012

YJ 574.583+633 (282.25) .. TT MQb QJT MTTUQT TJTRT TIT M WT VFS, e-mail: [email protected]

P. N 2009 2010 . . N, 17 -P . \ . , . N, , , , . , . ] , , Ergasilus sieboldi Nordmann. U , . RQ RTJ H - 2009-1010 . 17 W . 82 . T !! ! 50 F ( 3 64). W 4% . U - G. MQb QJT TLJ T. U (N 55º10'12" E 69º14'49"). ] 285030 ./3 15897.6 /3 ! Mesocyclops crassus (Fisch.), Eudiaptomus graciloides Lill., Daphnia longispina Müller, Keratella quadrata (Müller). T Bythotrephes longimanus Leydig, ! ! . U G 1.60, .. !- . ! ^-Y 2.37 . H ! , !! ! . T. Q (N 55º08'30" E 69º12'16"). R! ! 493440 ./3 1674.4 /3. J Mesocyclops leuckarti Claus 43 %, Brachionus calyciflorus Pallas 16 %, Filinia major (Golditz) 22 %. U G 1.73 –1.76. T !-, !. W , ! !. ! ^-Y 1.84 . T (N 54º58'31" E 70º50'19"). ] 59.8 . ./3, – 2 /3. U M. leuckarti E. graciloides, D. longispina Müller K. quadrata - . U G ! 1.58-1.64. H !!, !-. ! ^-Y 2.37 . H , !! ! . T I ( . ) (N 54º43'39" E 66º58'20"). R! 140888 ./3 ! M. leuckarti K. quadrata, - 3961.6 /3 D. longispina Daphnia pulex (De Geer). U G ! 1.72-1.79. H !-, !. ! ^-Y 1.94 . S ! - !- . T U (N 54º42'58" E 66º57'17"). ] 577.5 . ./3, - 27.5 /3. T . U ! . W ! M W . U G 1.58. T ! !-, ! . ! ^-Y 2.09 . T J (N 54º48'15" E 68º50'52"). ] 31.3 . ./3, - 1.3 /3. J M. leuckarti, Cyclops kolensis Lill., E. graciloides Lill., D. longispina, Ceriodaphnia quadrangula (Müller) Bosmina longirostris (Müller), K. quadrata. U G 1.65 - 1.68. H ! !-, !. ! ^-Y 2.04 - 2.57 . H , !! ! . H SY. . 1 (33). 2012 173

T (N 54º36'14" E 69º41'30"). ] 89.2 . ./3, – 2 /3. Copepoda , . H , D. longispina Chydorus sphaericus (Müller). W ! Asplanchna priodonta Gosse ( 27 . ./3). U G 1.59 - 1.62. H ! !-, !. ! ^-Y ! 1.79 2.09 . T (N 54º36'14" E 69º41'30"). ] 156.2 . ./3, - 8.7 /3, ! D. longispina. U G 1.53. T, !!! . H ! !-, ! . ! ^-Y ! 1.59 2.13 . S ! - !- . T U (N 54º49'53" E 69º50'60"). ] 245000 ./3, – 19739.3 /3. U Mesocyclops oithonoides Sars. 71%; Sida crystallina (Müller) 6 %. U G ! 1.63-1.64. H !!, !-. ! ^-Y 1.77 . H ! , !! ! . T (N 54º41'30" E 68º57'45"). R! 354900 ./3, - 941.3 /3. J M. leuckarti (42 %), Polyarthra minor (Rousselet) 14 %, K. quadrata 28 %. U G ! 1.63-1.64. H !-, !. ! ^-Y 1.94 . H - !- . T U" (N 52º59'36" E 68º17'10"). ] 565000 ./3, 8179.7 /3. J M. leuckarti 7 %, K. quadrata 74 %. U G 1.64 - 1.65. !-, ! . ! ^-Y 1.59 . H ! 1-2 , !! ! . T (N 52º35'30" E 66º40'20"). ] 497940 ./3, - 4220.6 /3. J M. oithonoides 17 % - . U G 1.65. H ! !-, !. W , ! !. ! ^-Y ! 1.84 1.87 . S ! , !! ! . T (N 53º31'46" E 67º50'30"). ] 299500 ./3, - 1787.6 /3, ! M. leuckarti 17 %, Polyarthra remata Skorikov 52 % Br. angularis Gosse 17 %. T . U G 1.69 - 1.72. H !-, !. ! ^-Y ! 1.69 1.83 . H ! - !- . T (N 53º00'52" E 68º14'17"). R! 508560 ./3, - 2707.1 /3. H ! M W. J C. kolensis, Cyclops strenuus Fisch, M. leuckarti 22 %, D. longispina, Keratella cochlearis (Gosse) 4 %, K. quadrata 17 %. W Kellicottia longispina (Kellicott), ! . U G 1.57 - 1.61. H ! !-, ! . ! ^-Y 2.08 . H ! , !! ! . H Ergasilus sieboldi Nordmann. c – . T (N 50º54'50" E 67º00'10"). ] 601.5 . ./3, - 21.4 /3. T . J M. leuckarti Claus, Ch. sphaericus, K. quadrata. U G 1.63. H ! !-, !. W , ! !. ! ^-Y 2.48 . T (N 52º56'37" E 66º52'30"). ] 63.2 . ./3, - 3.9 /3. J C. kolensis, C. strenuus, D. longispina. U G 1.53. H ! !-, !. ! ^-Y ! 1.62 2.11 . M . T , . (N 54º11'51" E 67º48'35"). ] 1120820 ./3, - 15552.1 /3. T 174 Y . . 1 (33). 2012

!. J M. leuckarti 17 %, Br. angularis 51 %, K. quadrata 4 %. U G 1.69 - 1.70. H !-, !. ! ^-Y 1.52 . T , !! ! 1 - 2 - . S . . . (N 55º22'30" E 69º22'48"). ] - 3060 ./3, - 22.3 /3. J M. leuckarti 39 % Ch. sphaericus 15 %. M, , . U G 1.59, .. !- . ^-Y - 2.44 . X.. ! , !! ! . . R (N 55º03'04" E 69º08'39"). ] - 6840 ./3, - 38.5 /3. J Br. calyciflorus 17 %; M. leuckarti 24 %. U G 1.67. Y !-, !. H !!! M. oithonoides - !!. ^-Y 2.16 . S ! !- . . U, . (N 55º56'05" E 69º07'06"). ] 176740 ./3, – 1179.9 /3. J M. leuckarti 60 %, F. major 4 % Polyarthra remata Skorikov 16 %. U G 1.66. Y !-, !. ! ^-Y 1.99 . G . . U (N 54º52'35" E 69º30'49"). M ! . U . ] - 46840 . ./3 1345.5 /3. J Cyclops scutifer Sars, Acanthodiaptomus denticornis Wierz., B. longirostris, C. quadrangula, K. quadrata (Müller), .. . M, , . U G 1.57 –1.58. Y !-, !. W , ! !. ! ^-Y 2.44 . 35 . U (N 54º41'50" E 68º58'42"). ] 10540 ./3 42.0 /3. J M. leuckarti 30 %, P. remata 15 %, F. major 15 %. M . U G 1.65, .. ! !- . ^-Y 2.42 . S ! , !! ! . 5 . (N 52º21'21" E 66º40'21"). ] 2500 ./3, - 9.7 /3. J M. leuckarti 20 %. U G 1.60. Y ! !-, !. ^-Y - 1.72 . S ! - . . M (N 52º56'59" E 66º37'32"). ] 3120 ./3, - 22.3 /3. U ! . J M. oithonoides 13 % ( !!), K. quadrata 13 %, Euchlanis deflexa Gosse 12 %. U G 1.62. Y !-, !. ^-Y - 1.56 . S ! - !- . H Ergosilus sieboldi Nordmann. . (N 53º20'49" E 66º59'25"). ] 1300 ./3, – 9.8 /3. T M. leuckarti 2 % Colurella obtusa Gosse 20 %. - 1.60. Y !-, !. ^-Y 1.69 . S ! . (N 53º24'55" E 67º30'47"). ] 87.3 . ./3, - 4.6 /3. U ! , ! , , ! . J M. leuckarti, D. longispina, Diaphanosoma brachyurum (Lievin). U G 1.66. Y ! !-, !. H !!! M. oithonoides - !!. ^-Y 1.56 . S ! - !- . . ] 375160 ./3, - 4816.7 /3. J M. leuckarti 13 %, Br. angularis 28 %, D. longispina 8 %. S Leptodora kindtii (Focke) – 300 ./3. , Ergasilus sieboldi Nordmann 50 - 100 ./3, H SY. . 1 (33). 2012 175

. U G 1.55 - 1.68. H ! !-, ! ! ^-Y 2.04 – 2.25 . H ! , !! . (N 55º90'43" E 69º15'47"). ] 5.3 . ./3, – 156.6 /3. T M. leuckarti Claus C. strenuus ! K. quadrata. U G 1.53. Y ! !-, !. ! ^-Y 1.61 . S , ! ! , ! - !- . TJ S , ! , ! , ! . T . c , ! ! ! . G !! ! !. c (!, ! !, ), . T ! ! ! ! , . ( W ) Ergasilus sieboldi Nordmann. *** L ' ' ' %' ''. M 2009-2010 . K' '' ' % ' 17 ' '''. L'' ' : % %'' . M' %' ', '' ''' % K' ''' ' '' . T , ' ' . M '' '' . G ''' Ergasilus sieboldi Nordmann ' . P' '' ''-'' %' , ' ' ' ' . *** In the presented work are cited results of inspections an ecological condition of some lakes and rivers of Northern Kazakhstan. Zooplankton researches were spent to a summer season 2009 and 2010 on the river Ishim, the Sergeevsky reservoir and 17 lakes of the Northern-Kazakhstan area. The estimation of an ecological condition and the trophic status of reservoirs bioindication methods was the purpose of the spent work. As a result of research it is shown that the most part of the investigated lakes and considerable sites of the river Ishim, including the Sergeevsky reservoir, are in an unsuccessful ecological condition, a zoocenosis in them are transformed, the dominating complex of a zooplankton is very poor in the specific relation. As a part of a zooplankton predominate as a rule eurybiontic species, which have wide ecological lability. The part of the lakes, including having the fishing status, is infected with ectoparasite Ergasilus sieboldi Nordmann. The potential of self-cleaning of lakes and the river isn't definitive yet, however additional nature protection actions are necessary for restoration of the safe ecological status.

YJ 574.5 . M. M, R.. TRTIT TT TJ TR T- MTO TQ (UQ M) UT RTZR WT VFS, . G, V!, e-mail: [email protected]

16 , -P . T , . W-! V (WT) ! ( 3500) ! . T , , , , . H !! , !! – ! !- , WT ! [1, 2, 3]. H! ! (), !!! , !! . ! !!, , !! . S !! ! , ! , ! !! , , Potamogeton pectinatus, Lemna minor, L. trisulca, Spirodela polyrhiza, Ceratophyllum demersum, Najas marina, !! ! !! [4, 5]. 176 Y . . 1 (33). 2012

\ – ! , W- . J! ! HcU WT VFS 2009 - 2010 . 16 W- . H . V VIU «W- . R. RTS V» WT VFS. ! , ( ! - ) [6, 7, 8]. J! ! ! [9]. U : 1). W- , « », ! (G J); 2). X (G X, , Y); 3). I: ( . – F, , U, V!, W), . (I, Q!, U, U), W (Y, Y, U). U , !! , , ! [10]. U ! 0,1 48,9 2, (. 1). ! – ! !! . J ! : , ! - . V ! ! (, ), (, U). X 1 - (2009-2010 .) U R! R!, ] S! , 2 , /3 S F 0,10 2,9 0,5 7,35 20 G J 0,15 1,8 0,5 9,00 12 G X* 34,80 3,2 1,1 8,30 13 I* 0,70 3,0 3,1 8,70 3 * 48,90 10,0 1,1 7,55 11 0,22 5,3 1,5 7,45 8 Q!* 5,80 3,0 2,9 7,35 3 U* 1,30 3,0 2,6 8,85 12 U* 3,70 1,5 1,1 8,70 9 U* 1,50 1,5 1,8 8,7 20 U 0,30 7,5 0,6 7,60 19 V! 2,50 1,8 0,8 8,60 24 W 1,04 1,0 1,2 9,10 12 Y 1,12 0,7 1,2 9,30 11 Y L 7,00 1,8 2,9 8,70 12 Y* 20,00 4,5 0,7 8,40 10 *– : [9]. U !! ( [11]), , [12], - (F, G J, U, U, V!, Y, Y) - (G X, , I, Q!, U, U, W, Y L). F! ! (S) 7,35-9,3 ! [13]. MQb TLJ H 64 40 , 28 4 . S ! ! Cladophora glomerata, – (Thelypteris palustris) ! (61 ). R (11 ), (9 ) (5 ). X H SY. . 1 (33). 2012 177

! [14, 15], M- [16] T- ! [17], ! , , , ! . T , ! !- ! . T ( ) . . T – «» , ! – 80-85 %. H ! – : 20 . U! , (Alisma plantago-aquatica), (Sagittaria sagittifolia), (Butomus umbellatus) (Typha angustifolia), . H (Ceratophyllum demersum), , (Potamogeton perfoliatus, P. pectinatus), ! ! (Miriophyllum sibiricum) ! (Utricularia vulgarus). S ! (Nuphar lutea), ! (Nymphae candida), (Stratiotes aloides). ] - 8 ./2, 280 /2. H ! ! (Lemna trisulca). G , !! ! , ! . T J (R ) - .. F! 50 % . W ! . J 80-100 %, S !! - ! (Elodea canadensis), !, (Potamogeton lucens, P. praelongus), (Ceratophyllum submersum), ! !. U! , ! 10 , (Phragmites australis) . ] ! 20 ./2, – 256 /2, ( ) – 28 ./2 560 /2 . G , !! ! , ! , F, . T U ( ) . !!!, !, . X ! – -. F! . J , . S ! - , (Batrachium circinatum), , (Chara sp.). S ! ! : !, !, ! !. H 20 , (28 ./2) , – 1000 /2. W , ! . T ( R L) – . , V! – 24 15 . X ! – -. F! 40 % , (Scirpus tabernaemontani) (Carex) . ! . Y 80-90 % ! (Spirodela polyrhiza.) , (Hydrocharis morsus-ranae). T! ! – 70 %. ] ! 212 ./2 () – 524 ./2 (), 2120 600 /2 . H 100%, ! 60 ./2, 112 /2. U – . T (\ ) . W - (, , , , ) ! . W - ! . U . ] ! 36 ./2, 112 /2. H ! ! 30 80 %, 24 ./2 328 /2. G 2 ! ! , , (M. spicatum). X ! – T! ! 5 %, ! 2-3 %. T (F ) !. H ! 50-200 3 , 178 Y . . 1 (33). 2012

! 5 – 10-13 65-70 %. S , ! !, !, – . ] ! 48 ./2, 480 /2. X ! – . T! ! 5 %. T I (L ) ! . H , 3 . H 1,5 ( 10 ). H , , . G 0,4 0,6 . ] ! 92 220 ./2, 176-280 /2. T ( ) – , . U! , , . H , ! !, . T Q ( ) 2 . W ! . H (84%) [9]. T , ! ! 50-300 . U ! – 70 ./2 , – 7924 / 2. H ! . T! ! 15%. T U ( ) ! U !! . S , ! 65 %, 204 ./2 1120 /2. V ! (1096 /2). H ! ( 92 ./2, – 104 /2). U ! 15 – 17%. T U (R ) (L ) . H 9 . J , 5 , ! ! , – G (P. berchtoldii). G , ! – 76 ./2, – 736 /2 . H ! !. T! !, Z.R. [9] – 40 %. T U ( ) 2,5 . H. U ! [9]. X ! - . G , 10 30 , X. ] – 160 ./2 1584 /2 , - 44-56 ./2 848-2272 /2 . S ! . H ! ! 20-25 %. T ! . T ( ) ! . G . 40% - !. X ! - - . U ! ! Batrachium( circinatum). ] 104 ./2, – 448 /2. ^ ! ! . H ! !. V , , - 616 /2, - 800 /2. T (X! ) . . J ! !- !. W ! – 80%. X ! – -. X , ! (Zannichelia palustris), (P. pusillus), 952 /2. ^ !. T (F ). T 30 % . T! ! ! 50 %. ] 188 ./2, – 816 /2. H ! ( 128 ./2, – 136 /2), !, ! – 480 ./2 344 /2 . T ( ) . T – , 50 250 60% . U! , , , ! !. H ! H SY. . 1 (33). 2012 179

(Persicaria amphibia). U ! ! 20-25 %. ] - 52 ./2, 272 /2. TJ X , . H ! 3 24 . S , !, . S , !! ! !! - , , , ! !. R !! - ! !, !! !! . U ! ! 5 85%. S F, U, V! , , . ! Y, !!, !, . !, - !!, , ! . Q 1. J Q.. I! WWWV ( ). ]. 2. I! . – Q.: QIY, 1955. – 600 . 2. U F.I. T T- (M! ! ). – S: M.-W. . -, 1967. – 350 . 3. ! WT // T ! ! ! W- [ ] – http://dpr.sko.kz/rus/bulleten.htm 4. G .S. R c. – F. J…... – R, 1994. – 22 . 5. U W.W. c- G. – F. J…... – J, 1992. – 17 . 6. G! F. U. ! . // G. . 1979. X. 64, 3 1. W. 32-41. 7. ! H. R. H! ! WWWV. R !. – Q.: S, 1981. - 188 . 8. U H.I. U! ! // I: ! : R ^ . – V: TFT «V J », 2003. – W.137-145. 9. Z.R. T W – . - U, 2004. – 106 . 10. U F.I. M! ! T- // H M W. – S: M.-W. . -, 1965. – W. 13-42.. 11. T T.U., L H.S., G U.S. . ! ! ! // I. . - 1993. - X.29, 34. - W. 62-76. 12. F T. F. T . - Q.: I, 1953. - W. 109. 13. J Q.F. T ! . // R , , !, 2011. - 3 1. – W. 347–351. 14. ! H. R. V W // T WWWV. Q.: S, 1970. W. 922135 15. W, G.. H W- ( , !, ). – F. J…... – X, 1987. – 17 . 16. ^, S. H. M- : F. . . . . X. / S. H. ^. – , F, 2007. – 22 . 17. M K.Z., W R.., I J. Z. W T- ! (H F !) // R , ! – 2009, 37(19) – W. 15-21. *** T ' , %'' ' 16 '' '. &'' ' ''''' % ' ''' . *** In this research it is the description of the modern status of 16 water bodies, which are at the Nord-Kazakhstan’s region. The specific diversity and the productivity of the vetable communities is describes.

YJ 577.472 Q . " TT TZ QTQbTO R TM 2009-2011 II. XTT « S !» FT «F!»», . F, [email protected]

F 2009-2011 . . J . S ! !!! !! . M , ! , !! ! . 180 Y . . 1 (33). 2012

F! – . ! F , 40- , ! , !, [1]. , , . J! ! !! SV 1963 . . F W 1 . . H . F, ! , . U ! . W [2]. H !! ! F SV. U , , [3, 4]. U !! , , ! , , , . S ! ! , ! . J! ! !! [5]. \ !!! !! F 2009-2011 . !! (GW). T!! ! . RQ RTJ QJTO J! , 2009-2011 . . F, W ( 147 ). H ! ! – !. Q 2009-2010 . ! – !!, 2011 . – . R ! ! ! [6-9]. T !! !! [5]. U ! ! , (N), (B), (n), ^ (H), H (V) U (D). H H ! G (), ! . [10]. M! ! . Y! ! . MQb TLJ G F 2009-2011 . , !, , , , , , , ( 70 ). H , ! 60-80% !. ] , Caenis qr. macrura (Stephens), . Chironomus Procladius. H , !! - . S . F (50 ). T ! ! - , . W. H . W 38 36 , . ] , , , . H ! ! 2009 2010 . . H!, ! 2010 . ! , . W . W, F - (). X 1 T, 2009-2011 . H I R! ], ./2 G, /2 n S, / D, % GW T. F 2009 I 2294 0.92 25 1.9 5.9 8 2.2 III 1269 1.03 34 1.8 7.1 7 2.0 2010 I 633 1.37 23 1.1 6.7 12 1.6 III 517 1.09 18 0.9 6.5 2 1.6 2011 I 857 2.63 27 1.1 7.7 2 1.7 II 437 0.72 21 0.8 6.5 2 1.7 T. W 2009 I 598 0.83 14 0.9 7.0 70 1.4 III 513 0.18 11 1.3 7.2 65 1.4 2010 I 426 0.48 8 0.5 7.2 63 1.4 H SY. . 1 (33). 2012 181

III 428 0.62 10 0.6 7.5 52 1.7 2011 I 1098 1.26 20 1.1 6.8 14 1.8 II 601 2.37 12 0.6 7.7 46 1.6 T. 2009 I 3666 2.17 23 1.0 6.8 51 2.0 III 1320 1.07 5 1.4 6.5 47 1.9 2010 I 1144 0.96 7 0.1 6.7 98 1.3 III 1120 3.02 7 0.9 6.9 51 1.8 2011 I 1627 1.27 18 2.0 8.4 53 2.2 II 3123 3.94 6 0.9 6.9 18 1.9 U: I – , II , III– V . F , ! . H 2009 . ! , (24-49% , 43-45% ). H 2010 -2011 . . Chironomus . V - 16 6% . ! 2009 2011 . (). U 2009 2011 . 3 , ! - ! 1,5 . H! 2009 ., !, ! . W, !! , !, - . U 2011 . 3 ! ! !. H , . G . ! . T , - Chironomus plumosus (Linne). R! ! (! . ) 2010 . . ! !! ! !, ! . S 2009 . ( ) 2011 . ( ! Ch. plumosus). W - 2010 . ! Ch. plumosus. G . W ! !. T , , , Ch. plumosus. X ! 2009 2011 . 2011 ., . G 2009 2011 ., ! 2011 . ! Ch. plumosus ! !. S ! , , ! ^-Y. W, ! !. . F 2009 . . 2011 ., . T ! ! !! . F (« – !») ! ! . W . W . F «», . W ! «» 2009- 2010 . «! !» 2011 . S GW F (). T !! . ! . S 2009 2011 . W GW 2010 . . G . F , , !! . H GW . F ! 2009 2011 . !, 2009 . ! - , . R ! 2010 . ! ! Ch. behningi Ch. lumosus. R , ! ! . W, ! , GW. 182 Y . . 1 (33). 2012

S GW 2009 2011 . ! . W, !, !! . X , ! F ! GW. W ! ! . Q 1. R! F. W ! F . // W . F- F, 1959 . H. 2. W. 119-144 2. Q c.H. ! F . // F! . F-F: S,1965, H. 12. W. 223- 235 3. c Z. H. R F . // X F . F: R, 2004. W. 137-172 4. ^ Q. . , F. U., K. I., c Z. H., X X. X. F . // V! ! V : ! !. F: G. 2005 . W. 441-450 5. G! !! /U . T.H.G, I.W.V. – R., 2007.- 403. 6. R ! ! (, ) F, 2006. – 27 . 7. V . – Q., 1983. – 240 . 8. T V : S (J).- WU, 1999.-X.4.– ].1, ].2.- 998 . 9. T V : H .- WU, 2001.-X.5.- 836 . 10. G K.H. , !! . R . M FS WWWV. - Q., 1976, - W. 106-118 *** G 2009-2011 . F' ' '' '' %' ''. M , ''' ' ''' . F ''' . *** The article provides a comprehensive evaluation of the biological state of Alakol lakes’ benthic fauna in the period between 2009 and 2011. The mean results obtained are comparable in all the lakes due to the similarities in taxonomic and quantitative characteristics of the zoobenthos. The range of values obtained corresponds to the low well-being of benthocoenosis.

YJ 574. 582 .I. MTTM TQ] UTMQ MTTUQT RQ U TJTRT MUJT-MTO TQ VIU « », e-mail: [email protected]

U M-P , (45 ). K (76%) – Cladocera, Copepoda, Anostraca, Conchostraca. R -«», . N P . Calanoida – Arctodiaptomus ulomskyi Chechura A. spinosus (Daday). M . H ! !! . Y, ] G [1-3]. R , ! , ! !. H 30 ! 3 ! 2010 . 5 , 400 . Y L M- . U 0.001- 0.563 2. J . F (/) , ! ! ! («»). R, 0.2 . H . F . L ! . U !! ! . H, I , !. H « (12 ZM . ) !! . U . H !, !!! !, (0.21-0.32 /3), , !. H . L (2.90 /3), , !. W 0.064 /3. U ! ! ! ! , , . U . ! ! – 0.013-0.036 /3. H ! 45 ( 1). S – 17 . R (11) (12). H ! Anostraca (L) Conchostraca (Q). U ! H SY. . 1 (33). 2012 183

8 22. R ! -«», . T ! !! D. similis, D. magna, C. reticulata, M. brachiata, D. bicuspidatus, M. rubellus. H ! ! . Calanoida – Arctodiaptomus ulomskyi Chechura A. spinosus (Daday). T ! - . F. X 1 M- , 2010 . M L Bdelloida gen.sp. + Notommatidae gen.sp. + Asplanchna sieboldi Leydig + Platyias quadricornis Ehr. + Polyarthra sp. + + Brachionus rubens Ehr. + Keratella quadrata Mull. + + Euchlanis phryne Myers + Hexarthra sp. + Hexarthra intermedia (Wiszniewski) + Filinia longiseta Ehr. + Diaphanosoma brachyurum (Lievin) + Polyphemus pediculus (Linne) + Daphnia galeata G.O. Sars + D. magna Straus + + + D. longispina O.F.Muller + + + Daphnia pulex Leydig + D. (Ctenodaphnia) similis Claus + + + + Moina brachiata (Jurine) + + + M. micrura Kurz + Simocephalus expinosus (Koch) + + S. mixtus Sars + Ceriodaphnia reticulata (Jurine) + + + C. setosa Matile + Macrothrix odiosa Gurney + Alona rectangula Sars + Chydorus sphaericus (O.F. Muller) + + Bosmina longirostris (O.F. Muller) + Microcyclops rubellus (Lilljeborg) + + + Cyclops strenuus Fischer + + + Diacyclops bicuspidatus (Claus) + + + + Megacyclops viridis (Jurine) + + Arctodiaptomus salinus (Daday) + + A. ulomskyi Chechura + + A. spinosus (Daday) + Eudiaptomus graciloides (Lilljeborg) + + Diaptomidae 1 + Diaptomidae 2 + + Diaptomidae 3 + Hemidiaptomus sp. + Ostracoda 1 + Ostracoda 2 + Ostracoda 4 + Anostraca + + + 184 Y . . 1 (33). 2012

Conchostraca + + : 13 8 22 18 18 !, (, ) ! ! . W! 100.4 456.0 . /3, 7.1-26.9 /3 ( 2). X 2 M- , 2010 . I F WM F F L ] G ] G ] G ] G ] G Rotifera 1.1 <0.01 0.01 <0.01 67.8 0.1 25.9 0.1 1.9 0.0 Cladocera 3.6 1.6 5.3 1.6 133.5 9.6 104.1 8.7 80.8 23.7 Copepoda 95.7 1.9 20.6 1.4 254.7 2.2 38.6 0.6 69.2 0.4 U 0.06 3.6 0.2 10.3 0.0 0.0 0.0 0.0 0.08 2.8 100.5 7.1 26.1 13.3 456.0 11.9 168.6 9.5 152.0 26.9 U – ]. – , . /3, G. – , /3. H (95%) , ! A. salinus A. ulomskyi. U . U («U»), 51.0% . S 26.5%, – 22.5% !. J ! – 0.071 . V ^-Y (S) (S) ( 3). 3 M- , 2010 . U F WM F F L 11.0 11.0 17.0 14.5 18.0 S, / 1.67 1.29 2.40 2.25 2.39 S, / 2.31 1.37 2.14 1.71 1.99 !! . , 0.071 0.359 0.026 0.052 0.181 (M . ) , (. . 2). T ! (79 %) , A. spinosus. W – 20,3%, D. similis. U , – 77.2% !. S 12.1%, – 10.6 % . V ^-Y (. . 3). M . ! . J – 55,8%. W E. graciloides (33,8%), C. reticulata (20,0%), D. bicuspidatus (17,8%), Polyarthra sp. (14,5%). U 11.9 /3, – 80.7%. U ! - , ! ^-Y (. . 3). . L . F !! , 92.2% 61.7% . W D. magna C. reticulata. D. bicuspidatus, – K. quadrata. W ! ^-Y, (S) (S) . M «» ! ! . U D. longispina D. similis. W M. rubellus D. bicuspidatus. H 53.2 % 88.2 % . U – 45.5%. U , H SY. . 1 (33). 2012 185

. V ^-Y . X , M- , ! (45 ). K (76%) – Cladocera, Copepoda, Anostraca, Conchostraca. R ! -«», . ! . Calanoida – Arctodiaptomus ulomskyi Chechura A. spinosus (Daday). V ! , !. . Q 1. F .R., G M.G., V! S.F, I ! Y // V! ! .- F: SV, 1999. - W. 102-105. 2. X X.X., ! U.H., I T.U. M ] ( , 2005) // R., . 75- «G , ! !». - F, 2007. - W.81-83. 3. X X.X., ! U.H., I T.U. W- G . L ( 2006) // R., . 75- «G , ! !».- F, 2007. - W. 84-86. *** G '' ' ''' (45 ) '. T '' (76 %) 4 %'' : Cladocera, Copepoda, Anostraca, Conchostraca. '''' ' ' «» – ' . J% ' . Calanoida ' ' - Arctodiaptomus ulomskyi Chechura % A. spinosus (Daday) . *** At inspection of small steppe reservoirs of the West Kazakhstan area, the zooplankton was characterized by rather high variety (45 ). Its basis (76 %) was formed by crustaceans of four groups – Cladocera, Copepoda, Anostraca, Conchostraca. The minimum number of kinds is revealed in a zooplankton of reservoirs - of the "cubes" having very low transparency of water. In these reservoirs two Kazakhstans of a kind new to fauna island Calanoida – Arctodiaptomus ulomskyi Chechura and A are found out. spinosus (Daday).

YJ:574 1.Z. ., 2R.. Q RUTIT TQTIT LR TQ]T M TT IUU ZTUQT MTTUQT T-TT]TO ] TIT U 1F I (FIY), V!, . F 2XTT« », F,

U - P . U , - P, . J . S , ! . U , . [4;6]. \ ! !! ! ! - ! R ! ! - 2010. - ! SW «FQXFO». U ! . J! ! ! «HORIBA U-10». U (0,5 ) R ! 40% . U ! ! ! 10-14 , ! ! . T 30-80 . U ! S, 0,1 . V ! 13 . R , ! , /3. W ! J 370, ! . U 40% 10% . T . X – ! , ! !, . R 186 Y . . 1 (33). 2012

( ) ! . T , ! ! , ! ! . X , !! , ! ! ! [1;3;5]. H ! , ! !!, . H ! , 23.2°W ! 21.7°W, ! ! 20.0°W. , ! , . U ! ! !!! () ! , ! , ,– !, . . [2]. W ! ! . W , !, ! ! T! ! - W ! !! 0.07 0.11% [7]. R ! - W ! 0.82% ! 0.74%, 0.66% - , ! ! . H . Y. H ! ! (X.1). X 1 I - ! 2010 . X, W,% M 3 °W . H Q H Q H Q H Q

4644’ 52,107”N 1 23.2 27.3 0.66 0.73 556.1/181.5 385.1/2170.8 419.0/1016.4 116.7/455.7 5045a22,379bE 4644’ 40,989”N 2 21.8 27.3 0.76 0.86 125.1/283.0 210.8/618.2 199.8/1266.6 53.1/208.2 5115’ 7,235”E 3 4634’ 39,867”N 23.0 28.4 0.66 0.71 411.2/1001.7 311.2/2181.9 296.2/665.0 77.9/203.7 5025’ 22,882”E 4 4634’ 21,970”N 21.5 27.1 0.79 0.88 144.8/1596.9 214.1/1868.4 48.5/331.6 18.3/50.7 5115’20,392”E 5 4634’ 42,865”N 21.7 27.0 0.77 0.87 48.8/1006.8 88.0/448.7 75.0/1055.4 68.5/433.0 5155’ 8,989”E 6 4614’ 57,001”N 21.6 28.5 0.66 0.83 36.2/937.0 431.5/1967.0 115.2/241.3 281.0/954.2 5025’ 3,452”E 7 4615’ 2,246”N 20.0 28.4 0.81 0.88 29.7/786.4 87.1/2701.0 68.6/550.2 25.2/112.1 5055’19,913”E 8 4614’ 48,537”N 21.7 27.8 0.77 0.87 143.3/898.8 31.5/687.2 41.8/246.8 25.4/303.1 5125’4,455”E 9 4614’ 49,200”N 22.0 25.4 0.76 0.86 36.1/1228.5 57.4/1910.6 41.4/343.8 38.5/396.2 5154’48,994”E 10 4554’ 37,579”N 21.7 28.5 0.70 0.70 125.0/256.7 128.9/635.6 568.0/1430.2 99.9/179.5 5015’ 14,446”E 11 4554' 46,229”N 22.4 26.4 0.79 0.84 44.5/963.9 29.5/280.3 34.3/489.0 3.6/12.2 515’ 7,600”E 12 4554’ 44,283”N 22.0 24.1 0.70 0.82 29.6/214.6 114.4/2540.9 91.2/1606.1 9.7/136.1 5154’ 45,003”E 13 4535’ 3,338”N 21.0 24.1 0.77 0.90 18.2/902.1 35.0/986.2 59.7/383.5 4.7/30.6 5055’ 7,501”E 14 4534’ 35,967”N 21.0 25.2 0.82 0.83 68.7/182.8 35.4/1016.7 132.6/369.7 19.7/68.9 5135’ 0,419”E 15 4524’52,644”N 21.0 28.4 0.68 0.99 25.9/259.7 50.0/1198.4 70.5/357.8 21.5/132.5 5034’ 59,667”E H SY. . 1 (33). 2012 187

X, W,% M 3 °W . H Q H Q H Q H Q

W 21.7 26.9 0.74 0.84 122.8/713.4 147.3/1414.1 150.8/690.2 57.6/245.1 *U: – ./3, ., /3; – , /3 Q - ! 28.9°W, ! 26.9°W, ! 24.1°W. R ! - W ! ! 0.99%, ! ! 0.70% ! 0.84%. H ! - - W ! . H . Y, ! 556.1 ../3 1596.9 /3 ! 181.5 /3 18.2 ../3 (X. 1). U , Cyanophyta ( 202.0 ../3, 45.5 /3), Bacillariophyta ( 74.5 ../3, 1560.5 /3), Pyrrophyta ( 8.2 ../3, 30.1 /3) Chlorophyta ( 330.0 ../3, 20.6 /3). R W ! Cyanophyta ! 0.2 ../3, 6.0 /3, Bacillariophyta 7.1 ../3, 150.0 /3, Pyrrophyta 10.4 ../3, 0.8 /3, Chlorophyta 2.8 ../3, 0.1 /3 (X.1). Q ! 26.9°W, - . T ! - . R - W !, W ! ! 18.2 556.1 ../3, 181.5 1596.9 /3. U , - W !, , W !. X ! (-) ! !. ] ! ! , 23.0-28.0°W ! - , ! 21.0-24.1 °W. V - W ! ! , ! W !. R ! - 281682.0 . 954.2 . , , ! 3636.0 ., 12.2 /3. V! , Rotifera ( 290359.0 . 331.1 /3), Copepoda ( 150429.0 . 463.8 /3), Cladocera ( 7513.0 . 736.4 /3) - ! W !, ! 23.5-28.5°W, ! 0.6-0.7%. R ! - W !. X , ! 23.2 °W. Q 28.5°W, , ! . 28.5°W, 21.6°W. V , ! - !, !! . H . Y ! . H . Q 1. F K.I. X !// X. ITS, 1957. H. 35 W. 3-101 2. G Q.. W ! ! ! !// X. . . FS WWWV, 1959 X. 5 W. 95-101 188 Y . . 1 (33). 2012

3. S.U. S // I. L. 1984, 36, W. 50- 53 4. W R.F. c! ! !, G, 1999, 400. 5. X S.F. H! ! W ! // X. HSVT 1971 X.86 W. 56-89 6. Y .. F W ! // . FS FWWV. 1960 34 W. 79-92 7. ] T.H., Q F.H. I- ! ! // H. V. , 2004., 33, W. 299-317 *** ' P' '- '' % - . ' P' '- % , ' '' . Z '' %' . *** The ecological characteristics of the NE Caspian Sea and assessment of the planktonic community status were provided. Abiotic and biotic factors that determine formation of favorable conditions in the NE Caspian Sea as well as peculiarities of distribution and habitats of main planktonic organisms were analyzed. The impact of temperature and salt regime on development of phytoplankton and zooplankton was assessed. 1 YJ 574.5 +597-19 1Q T.., 1U J.., 2R .., 2 I.., 1R .. T MTTM IJTZ U UTT Q UJQ UQ M 1VIU «N » PS RTS VP, 2 JIU «SNN » VIU «PSY .-Z» .F, V P. E-mail: [email protected]

2007-2011 .. N V P. 21 78 – . U . U ! -G ! ! ! 20 [1]. M ! ! ! ! V SV ! . H [2-4]. ! ! !. H! ! !!! ! ! ! ! ! . T !! – , ! . !! , ! ! ! !! . T ! . \ – ! ! , !! !. RQ RTJ QJTO W - ! - 2007 – 2011 . , V . G .: .X-, .G, .Y, .G, .Y, ., .^. H ! ! !, ! .. U ! [5-10]. S ! ! ! ! ! !. U ! 1318 , 20 . U ! ! . J! , 15 ! 5 , . V 15 6% , , ! . T . ... K.J.H (M RIY, .R). H !, . H , , ! . F H SY. . 1 (33). 2012 189

, !!! : . J! ! . MQb TLJ T ! ! !. S , – . S ! ! . H ! : Hydridae; Turbellaria: Polycelis felina; Hirudinea: Erpobdella octoculata (Linnaeus, 1758); Helobdella stagnalis (Linnaeus, 1758); Oligochaeta: Limnodrilus sp., Tubifex tubifex, g.Pelascolex; Mollusca: Limnae ovata. Limnaea corvus (Gmelin 1791), Lymnaea stagnalis, Planorbis carinatus, Viviparus contectus (Millet,1813). Pisididae. Sphaerium corneum; Crustacea: Podocopid: Cyprididae, Amphipoda: Gammarus lacustris, Palaemon modestus(Heller, 1862); Leander sp., Acariformes: Hydracha sp., Insecta: Odonata: Gomphus flavipes, Onychogomphus forcipatus, Sympetrum flaveplum, Sympetrum sanguineum, Aeschna juncea ( Linnaeus , 1758 ), Ischnura elegans, Sympecma fusca, Lestes sponsa, Ephemeroptera: Cloen dipterum, Baetis rhodani, Heptagenia flava, Iron (Epiorus) sp., Epiorus (Iron) g.momtanus, Epiorus (Iron) g.rheophilus, Rhithrogena spp,Ameletus spp, Olygoneuriella sp., Plecoptera: Isoperla difformis, Diura difformis, Trichoptera: Hydropsyche pellucidula (Curtis, 1834), Hydropsyche ornatula, Phryganea sp., Sericostomatidae, Hydropsyche pellucidula, Heterptera: Sigara nigrolineata, Sigara lateralis (Leach, 1817), Sigara striata, Corixa punctata (?) (Illiger, 1807), Ilyocoris cimicoides, Nepa cinerea, Notonecta glauca, Diptera: Chironomidae: Camptochironomus tentans, Chironomus plumosus, Chironomus riparius, Cryptochironomus gr.conjungens, Glyptotendipes gr. gripekove, Sergentia g. longiventris, Stictochironomus gr. histrio, Procladius sp., Orthocladius g. saxicola, Psectrocladius gr.dilatatus, Syndiamesa gr. nivos; Diptera: Sphaeromias pictus, Odontomia sp., Iiponeura sp., Odagmia, Tetisimulium, Wilhelmia (Enderlein, 1921); Coleoptera: Coelostoma orbiculare, Cercyon sp., Cybister laterimarginalis (DeGeer, 1774), Hydaticus transversalis (Pontoppidan, 1763), Laccophilus sp., Haliplus sp. W ! 1. U Diptera, ! !! ! ! Nematocera. S , . U U. H , , ! 43% . S , ! , ! Ephemeroptera, , ! . W . H !, !! ! . S G. S ! . X, X- ! . X 1 T 2007-2011 .

X U I . H Vermes 5 0 2 7 Mollusca 4 2 1 7 Crustacea 3 1 2 6 Acariformes 1 0 0 1 Insecta 30 13 19 57 : Odonata 7 4 2 9 Ephemeroptera 1 3 5 9 Plecoptera 0 0 2 2 Trichoptera 1 2 3 6 Heterptera 6 2 0 8 Coleoptera 5 2 0 6 Diptera 10 0 7 17 190 Y . . 1 (33). 2012

: Chironomidae 8 0 3 11 43 16 24 78 H ! (Ephemeroptera, Odonata) , , !. U – , , , ! , ! ! . S Wilhelmia, . H G 2011 Gomphus havipes Ophiogomphus sp. ! !. U ! , ! ( 438 ). R , ! !! . H ! : Ophiogomphus forcipatus, Palaemonidae ( 339 / ) – , Limnae pereger ( 328 / ). H ! Anodonta sp. (24,7 / ). H ! 78 , - 7, – 6, -7, – 1, - 57, Diptera (17 ), 65% ! . Odonata, Ephemeroptera Heterptera ! ! . J! ! ! . T , . X ! ! ! ! ! - . H ! . I . U 2. X 2 , . S S V X SV [11] . F : W ! Phoxinus brachyurus Berg, 1912 + - - G! Schizothorax argentatus argentatus Kessler + + - ] Diptychus maculatus Steindachner, 1866 + - + I Diptychus dybowskii Kessler, 1874 + - + X Triplophysa stoliczkai (Steindachner, 1866) + - + C Triplophysa dorsalis (Kessler, 1872) + - + U! Triplophysa strauchii (Kessler, 1874) + + + T Triplophysa labiata (Kessler, 1874) + - + G Perca schrenkii Kessler, 1874 + + - ] : ^ Acipenser nudiventris Lovetsky, 1828 + - - R Salmo mikiss mykiss (Walbaum, 1792) + - ? U Rutilus rutilus (Linnaeus, 1758) - + + G Ctenopharyngodon idella (Valenciennes) + + + W () Leuciscus leuciscus baicalensis (Dybowski) + + - T Aspius aspius aspius (Linnaeus, 1758) + + - Q Tinca tinca (Linnaeus, 1758) - - - F Abbottina rivularis (Basilewsky, 1855) + + + F Pseudorasbora parva (Temminck et Schlegel) + + + Barbus brachycephalus brachycephalus F + - - Kessler, 1872 H SY. . 1 (33). 2012 191

Q Abramis brama (Linnaeus, 1758) + + + I Rhodeus ocellatus ocellatus (Kner, 1865) - ? ? T Rhodeus sericeus sericeus (Pallas, 1776) - + ? Rhodeus sinensis Gunther, 1860 + ? ? C! Carassius gibelio (Bloch, 1782) + + + C Cyprinus carpio Linnaeus, 1758 + + + T! Hemiculter leucisculus (Basilewsky, 1835) + + + G Hypophthalmichthys molitrix (Valenciennes) - + - U Hypophthalmichthys nobilis (Richardson, 1846) + - - H Misgurnus bipartitus (Sauvage et Dabry) ? + - + T Silurus glanis Linnaeus, 1758 + + + ! Orizias sinensis Chen, Uwa et Chu, 1989 - + + ! Oryzias latipes (Temminck et Schlegel, 1846) + ? ? T Sander lucioperca (Linnaeus, 1758) + + + G Sander volgensis (Gmelin, 1789) - + - Micropercops (Hypseleotris) cintus (Dabry et c ? + + Thiersant, 1872) Hypseleoris swinhonis + ? ? F Rhinogobius cheni Nichols, 1931 - ? ? G Ctenobius bruneus + ? ? G Ctenobius cliffordpopei + ? ? M Channa argus (Cantor, 1842) - + - H . ! . : ! Schizothorax argentatus pseudaksaiensis Herzenstein, 1889, ! Lagowskiella poljakowii (Kessler, 1879), W Nemacheilus sewerzowi G.Nikolsky, 1938. S . !! - Ictiobus cyprinellus, Mylopharyngodon piceus (Richardson, 1845), Gambusia affinis holbrookii (Girard, 1859). K.J.H . . H, . U [11], . !! Oryzias latipes (Temminck et Schlegel, 1846), . ! Orizias sinensis Chen, Uwa et Chu, 1989. ^ V , !! . SV. S, , , V . H , ! ., !. I.R.J W.. [12,13] ! Rhinogobius similis Gill, 1859. W! , Z.W.V ..R [14] ! Z , Rhinogobius brunneus (Temminck et Schlegel, 1845). I.-S.Kim [15] K.J.H [16] Gobius brunneus Rhinogobius similis. S.I.G! F.R.S [17] ! . U! Rhinogobius, ! [19] ! 10 , ! . X , . ! 21 78 . . F : – SV ... K.J.H (M RIY), .G ... J X.S., ... I F.c. F.. 192 Y . . 1 (33). 2012

Q 1 S- ! ! U!: R . 3 ! – F-F: S, - 1990. 2 H.K., W.W. R! ( Cyprinidae, Rhodeus sp.) .// V! ! V : ! !. – F: G. 2005. W.168-173. 3 J I.R. S G- (V ) // W!, , : R- - – H: H .ISY ISTV. 2007. W.95-96. 4 .., W H.F. W ! ! , ! // c! – F: G,, 2008. - W.273-280. 5 R ! ! (, ). - F: SU\ !, 2006. - 27 . 6 T V . X 1-6. U W. . \. W-U, 2000. – 1994-2004 – 528 . 7 .., ^ F.., .K. . ! Chironomini. F. - S, 1991. – 115 . 8 U H.. Q Chironominae WWWV (Diptera, Chironomidae). – Q.: 1983, S. – 309 . 9 Ecology and Biogeography of high Altitude insects. M.S Mani, Kluver, 1968. - 531 p. 10 Zasypkina I.A., Ryabinkhin A.S. Amphibiotic Insects of the Northeast of Asia. Pensoft, Backhuyes 2001. - 185 p. 11 Ren M., Guo Y., Zhang Q., Zhang R., Li H., Cai L., Yong W., Ren B., Gao H., Deng G. Fisheries resources and fishery of river Yili – Harbin: Heilongjiang Science and Technology Press, 1998. – 345 p. 12 W.., J I.R. R! ! Rhinogobius similes Gill, // H – 1981. X.21. H.4. W.600-607. 13 J I.R., W.. Rhinogobius similis Gill – // V – F-F: I. 1992. X.5. W. 269-286. 14 V Z.W., R .. Rhinogobius brunneus (Temminck et Schlegel, 1845) – // F V – R.: S. 2003. X.2. W.133-136. 15 Kim I.-S. Freshwater fishes// Illustrated encyclopedia of fauna and flora of Korea. Vol.37 – Seul: Ministry of Education. 1997. 629 p 16 H K.J. Gobiidae// F V – R.: S, 1998. - W.125-148. 17 G! S.I., S F.R. V !. – R.: X R, 2004. – 389 . 18 Nakabo T. (ed.). Fishes of Japan with pictorial keys to the species. English edition. – Tokyo: Tokai University Press. 2002. V.2. 1748 p. *** 2007-2011 . V = ''' % ''. G 21 ', ' % ' 78 ' . *** By the period 2007-2011 the macrozoobenthos and fishpopulation were ivestigated at the right tributaries of the River Ili in the Republic of Kazakhstan. 21 species of fish and 78 bottom and benthic invertebrates are identified. An analysis of the current species composition of floodplain waterbodies and watercourses lowlandand mountain areas, in comparison with previously published data are described in the artical.

YJ 577472 L.T.R TZ TJTQ_ - 28 QRTO TQ QTR 2011 I. XTT « - !» FT «F!», . F, [email protected]

Q 2011 . P – 28. G . , - - «». . J H ! ( F !, G . .) , ", , . F! . V . H ! ! , ! ! . W! ! . T ! ! - 28. H ! F . T ! J - X. H - 72 , !! . ^ - (]! ). G , . H ! ! ! . : ^ - 43° 17'3,08''W; 43° 17'7,14''W; 43° 16'24,88''W; 43° 16'29,02''W. J - 76°32'10,78''H; 76°31'58,76''H; 76°31'47,20''H; 76°31'32,66''H H SY. . 1 (33). 2012 193

\ – ! ! ! , , ! ! ! . RQ RTJ QJTO U ! U 0.025 2. I ! 3 23 ! . L ! , 4% . H !. T ! [1-5]. U 1 2. T ! W.U. [6]. MQb QJT TLJ H ! ( ) 1.5 . U ! . ^ 0.5 , ] – 0.3 , 1.0 , 12.5 , 1 . X – 20.50W. H ! ! ( 1, 2). H 2011 . – 28 12 , . c 3 9 – ! ( 1). U T. punctipennis, Ch. plumosus, P. ferruqineus ( 1). H – – C. Gr. viridulus 67 %. H – 1 – .^, 2 – . «]! » 3 – Y (). X 1 (%) Vermes – % Nematoda sp. 33 Oligochaeta sp. 67 Limnodrilus sp. 33 Insecta – S Diptera – J Tanypus punctipennis (Meigen) 100 Procladius ferruqineus (Kieffer) 100 Chironomus plumosus (Linne) 100 Cryptochironomus gr. viridulus Fabricius 67 Cryptochironomus gr. conjungens Kieffer 33 Cryptochironomus sp. 33 Microchironomus tener (Kieffer) 33 Polypedilum gr. onvictum Walker 33 Chironomus sp. Pupa 33 H: 12 H 1 ( . ^) 6 ( 2). c – Oligochaeta sp., – T. punctipennis, Ch. plumosus, P. ferruqineus, C. Conjungens Chironomus sp. upa. J! ( 1 ). M (97.9 % , 98.6 – ) ( 2). W 2- ( . «-») (9 ). J , -, . Q (94% 99 % ). U – P.ferruqineus P. onvictum (37 %). G 44 % – Ch. lumosus, 13% !. G (3 ) , 1- , 6 . T ( 2). S - . R! ! - 61% , 84 % – , 12 /2. X 2 -28, 2011 . I 1 2 3 W ], ./2 194 Y . . 1 (33). 2012

] 120 200 4720 1680 J 5640 3040 3000 3893 H: 5760 3240 7720 5573 G, /2 ] 0.08 0.02 10.0 3362.7 J 5.56 2.54 1.94 3344.0 H: 5.64 2.56 11.94 6706.7 ^ –Y 1.7 1.8 0.8 G 7.0 8.8 9.0 I Y!, % 2.1 6.1 61.1 H , - , ! « » . J! !! , !! G [7]. M! ! , «!». H «!». J! ! , (I! Y!) [7]. U – «» 1 2 « !» (3 ) ( 2). H!, I-Y! ! !! 1 2 . , , ! , ! . H , !! ! ( 2). H ! ^-Y . W ! , ! 0.8 / (3 !) 1.7 1.8 (2 3 , ), ^ ( 2). Q 1. R !. G !. – Q., ISTV, 1983. – 50 . 2. V . – Q., I, 1983. – 239 . 3. R G.R. T .- R.,1972.-399 . 4. T K WWWV ( ). – Q., 1977. – 511 . 5. T V : S (J).- WU, 1999.-X.4.– ].1, ].2.- 998 . 6. W.U. T « » " .- X. . V HIGT.- ].2- ,1986. – W. 254-255. 7. R . – Q.1976. – 168 . *** 2011 '' P-28 '. ' ''', % ''' '. '''' ' , '' «» ''. ''' ' . *** The summer of 2011 investigated a condition macrozoobentoc reservoir K – 28. The quantitative and qualitative characteristics of benthis fauna were given in comparative aspect. Pollution degree ground reservoir on zoobentoc to organisms which has been estimated - as - "polluted" is revealed. The size of a residual biomass defines a reservoir as average forage.

YJ574.5 .R. R TT TITR] TTT TM RTT TR MUJTO U QR] MR S X , e-mail: [email protected] V V _ 11-05-90726, \U «S - V» I 3 14.740.11.0935 M T- M W !! ! , ! ! [1; 2]. J ! !. S ! «!» , H SY. . 1 (33). 2012 195

! . T! ! : – « » , , , , ! ( ), , – ! . c ! , ! , ! ! M W ! ! [1]. W !! , ! ! ! ! , ! [3]. X ! ! M W, ! , [4; 5; 6; 7]. RQ RTJ QJTO ! ! , ! ! . X ! ! , " - . H ! ", , " ! 2–4 . S ", !, 1 [8]. H , . H ! , ! [9]. U (30–35 ) 250 , 4 (MILLEX Filter Unit) 0,45 . H ! Menyanthes trifoliata L. – H ! (V 1). X (-) - , ! ! W [10; 11]. H ! ! ! - . U !! 97 % [12]. H ! , Osmofilm ! ! - !! ! !. c ! ICP-MS (Agilent Technologies, 7500 ) In + Re. U ! 1 /-1 / . U 0,1 /.

1. M MQb TLJ U ! ! . U H , !, ! !: J! – K>Ca>Mg>Na>Fe>Mn>Al>Zn>Ba>Rb>Sr>B>Ni >Pb>Co>V>As>La>Cr>Ce>Mo>Zr>Cs>Y>Th>Dy>Yb>Hf>U 196 Y . . 1 (33). 2012

J! ! – K>Na>Fe>Mg>Ca>Al>Mn>Zn>Ba>Sr>Rb>Pb>B>Ni>Co>As>V> Zr>Cr>Ce>La>Y>Th>Mo>Cs>Hf>Dy>Yb>U J! ! – K>Na>Ca>Mg>Fe>Mn>Al>Zn>Sr>Rb>Ba>B>Ni>Co>Pb>As>V>Ce >La>Cr>Y>Zr>Cs>Mo>Th>Dy>Yb>U>Hf J! ! – K>Ca>Na>Mg>Mn>Fe>Al>Zn>Sr>Rb>Ba>B>Ni >Co>Pb>V>As>La>Ce>Zr>Cs>Cr>Mo>Y>Th>Sb>Dy>Yb>Hf>U T! ! H . S ! !, H , , ! : K, Sb, Pb Ba. H ! Mg, Ca, Mn Co, Rb, Sr, Cs. R ! ! ! , ! Tb, Dy, Ho, Er, Yb, U. H ! H () ! ! (V 2). S ! ! ! ! K, Mn, Na, Rb, Sr, Ba, Pb. ] ! !, ! .

2. M () 1. X M W ! , ! . ! ! ! ! ( ), , . 2. H ! !! ! ! ! (Fe, La, Ce, Pr, Sm, Eu, Gd, Tb, Dy, Ho, Er, Yb, Cr, Mn, Co, Ni) ! [13]. 3. T! ! H . S ! !, H , , ! : K, Sb, Pb Ba. H ! Mg, Ca, Mn Co, Rb, Sr, Cs. 4. S ! () ! , ! ! ! . 5. H H ! ! , , Pb, Zn, Sr , ( , .), !! ! M W (! «», « » .) [14]. H SY. . 1 (33). 2012 197

Q 1. W.S., U Z.R., G S.F. J M W ! ! // H X . – 2008. – 3 133. – W. 185– 189. 2. H.., G F.I. V 30 // . – 2009. – X. XIII, 3 2. – W. 16–26. 3. S.S., U Q.., W F.S., X .H., Q.F., .H., X T.F. R . G (V!) // F!. – 2008. – X. 18, 3 3. – W. 244– 255. 4. Q I.F. G! ! !! ! // H . – 2004. – X. 31, 3 2. – W. 215–222. 5. Q I.F., F I.S., G H.F. G // I!. – 2005. – 3 2. – W. 182–196. 6. Pokrovsky O.S., Shirokova L.S., Kirpotin S.N., Audry S., Viers J., and Dupre B. Effect of permafrost thawing on organic carbon and trace element colloidal speciation in the thermokarst lakes of western Siberia // Biogeosciences. – 2011. – 3 8. – V. 565–583. 7. Audry S., Pokrovsky O.S., Shirokova L.S., Kirpotin S.N., Dupré B. Organic matter mineralization and trace element post-depositional redistribution in Western Siberia thermokarst lake sediments / S. Audry, O.S. Pokrovsky, L.S. Shirokova, S.N.Kirpotin, B. Dupre // Biogeosciences Discuss. – 2011. – 3 8. – V. 8845–8894. 8. W.F. T ! F // S , – 2005. – 3 2, W. 15–24. 9. T U.X. F M W // M. – 2010. – X. XIV, 3 2. – W. 23–28. 10. W G.., R Z.W., W X.H. !! M-W . – T: F, 2011. – 231 . 11. W: W ! / W. Q.. R, I.F. U, .W. G . – S: S, 2005. – 362 . 12. U K.. J! M W. – F. . … . . . – X, 2009. – 24 . 13. W / U. . F.U. W. – R.: S, 1990. – 335 . 14. Q I.F., F I.S., F S.H., G L.T., H.S. c! !! -S // c! W, J H F: X . X. – 2001. – W. 153. *** L '' '' G '' '' ''' ' '' . *** In work it is considered some biogeochemical features of lake thermokarstic ecosystems of subarctic region of Western Siberia at climatic changes

YJ 574.5 .. MTTUQT O–ROTO R TM G XTT « - !», e-mail: [email protected]

2008- 2010 . N–R . N. ^ . –R! . ! ! ! 10 . S !! ! F, G ^ ( 1).

1. -R 198 Y . . 1 (33). 2012

T –R !! ! , , !. U !! – ! 452-492 /3. ! ! 68-102 % !, !!! ! ! . W – 4,5-7,8 T/3, !! !, ! ! . H . ! ! !, !!! . G, . M !! ! !. \ – ! –R . RQ RTJ QJTO R ! ! , – R ^, G F 2008-2010 . R ! ! ! [1, 2]. U F J. ! ! [3-6]. J! !! ! , , , . G ! ^ [7] W [8]. T 27. MQb QJT TLJ M ! ! 96 , 60 , 22 14 ( 1). X 1 (%) -R 2008-2010 . F G ^ X 2008 . 2009 . 2010 . 2008 . 2009 . 2010 . 2008 . 2009 . 2010 . 1 2 3 4 5 678 9 10 Rotatoria Notommata copeus Ehrnberg 100 Cephalodella gibba gibba Ehrnberg 33 66 33 Trichocerca (s. str.) longiseta Schrank 100 100 66 33 100 66 66 66 Trichocerca (s. str.) rattus rattus (Mull.) 33 33 T. (D.) bidens Lucks 66 33 Eosphora najas Ehrenberg 66 100 33 100 E. ehrenbergi Weber 33 Ascomorpha ecaudis Perty 33 33 33 Sunchaeta pectinata Ehrnberg 66 100 66 66 100 66 Polyarthra dolichoptera dolichoptera 66 66 66 33 100 66 Idelson P. minor Voigt 33 P. luminosa Kutikova 33 Bipaltus hudsoni Imhof 33 66 33 Dicranophorus forcipatus (Mull.) 100 66 100 66 33 Paradicranophorus hudsoni Glass. 33 33 66 Asplanchna priodonta priodonta Gosse 66 100 100 33 100 33 66 100 A. girodi Guerne 66 A. sieboldi Leydig 33 Lecane (s. str.) luna presumta Ahlstrom 66 33 66 L. (s. str.) luna luna Muller 66 100 33 66 66 66 L. (s. str.) ungulate Gosse 33 L. (M.) bula bula Gosse 100 33 66 66 L. (M.) quadridentata Her. 66 33 L. (M.) bula diabolica Hauer 33 L. (M.) cornuta cornuta Muller 33 L. (M.) crenata (Harring) 33 33 Habrotrocha bidens Gosse 100 100 100 33 66 Dissotrocha aculeata Ehrenberg 33 66 66 33 66 33 Adineta gracilis Janson 33 33 33 33 66 H SY. . 1 (33). 2012 199

Epiphanes macroura Barrois et Daday 33 66 Trichotria truncata truncata Whitelegge 66 66 66 33 33 66 33 66 T. pocillum pocillum Muller 66 100 100 33 66 T. tetractis Ehrnberg 33 Mytilina ventralis ventralis (Her.)0 33 66 66 33 33 Lophocharis rubens Wulfert 33 Colurella adriatica Ehrnberg 33 Lepadella (s. str.) ovalis Muller 33 66 66 E. incisa Carlin 100 33 100 100 66 E. dilatata dilatata Leydig 100 100 100 66 100 100 33 100 100 E. deflexa deflexa Gosse 33 66 E. pyriformis Gosse 33 66 33 Brachionus q. cluniorbicularis Skorikov 33 Br. q. hyphalmuros Tschugunoff 100 Br. q. ancylognathus Schmarda 33 Br. q. brevispinus Ehrnberg 33 100 100 33 Br. q. melheni Barrois et Daday 100 Br. angularis angularis Gosse 33 Br. plicatilis plicatilis Muller 33 Platyias guadricornis guadricornis 33 33 66 66 33 Ehrnberg P. patulus patulus Muller 33 Keratella cochlearis cochlearis (Gosse) 33 66 100 33 66 33 66 66 100 K. quadrata quadrata Muller 66 100 100 33 100 100 100 K. q. dispersa Carlin 66 K. v. brehmi Klausener 33 K. v. heteraspina Klausener 33 K. testudo testudo Ehrenberg 33 Notholca acuminata acuminata 66 33 66 Ehrnberg Testudinella patina patina Hermann 33 100 33 33 Filinia longiseta longiseta Ehrnberg 100 33 100 33 100 33 Hexarthra oxyuris (Zernov) 100 Cladocera Sida grystalline (Muller) 33 33 Diaphanosoma lacustris Korinek 33 Daphnia galeata Sars 100 100 66 100 66 100 Ceriodaphnia reticulata (Jurine) 66 33 C. laticaudata O. F. Muller 33 66 Simocephalus vetulus Muller 100 66 33 66 Moina brachiata Jurine 66 33 33 Scapholeberis sp. 33 Camptocercus rectirostris Schoelder 66 Macrothrix spinosa King 33 Ilyocryptus sordidus Lievin 33 33 Acroperus harpae (Baird) 66 33 33 66 Chydorus sphaericus Muller 100 100 100 66 100 100 66 66 100 Pleuroxus trigonellus O. F. Muller 66 33 66 33 P. striatus Schoedler 33 33 P. similes Sars 33 Alona costata Sars 100 33 33 66 Alona affinis Leydig 33 33 A. quadrangularis O.F. Muller 33 A. rectangula Sars 100 33 100 66 66 100 66 66 Bosmina longirostris O. F. Muller 100 100 100 100 100 66 100 100 Polyphemus pediculus Linne 33 100 66 33 66 100 Copepoda Macrocyclops albidus (Jurine) 33 33 33 M. fuscus Jurine 33 200 Y . . 1 (33). 2012

Eucyclops macrurus (Sars) 33 E. serrulatus Fischer 33 33 33 66 33 E. macruroides Lilljeborg 33 E. denticulatus Graeter 66 33 33 3333 Paracyclops fimbriatus Fischer 66 66 66 66 100 100 33 P. poppei Rehberg 33 33 Cyclops vicinus Uljanin 33 33 C. kolensis Lilljeborg 33 Acanthocyclops vernalis Fischer 33 Mesocyclops leuckarti Claus 33 Arctodiaptomus salinus Baird 100 66 33 33 Harpacticoida 100 100 66 66 100 66 100 H 96 46 23 44 21 27 53 30 24 37 H . F ! 68 , . G – 67, . ^ – 58. H ! – 53 78 %%, . T - ! 18 – Tr. longiseta, S. pectinata, P. d.dolichoptera, A. p. priodonta, L. l. luna, H. bidens, T. t. truncata. E. incisa, E. d. dilatata, K. c. cochlearis, K. q.quadrata, D. galeata, Ch. sphaericus, A. rectangula, B. longirostris, P. pediculus, P. fimbriatus, Harpacticoida. 44 89 %%. H ! 32 . . F – 12, . G – 13, . ^ – 7 ( 1). T W [7] -R 0,66 0,73, ! . ! ^ [7] (/ .) – . F (3,93-2,78-3,86) . ^ (2,60- 3,11-3,10), . G (1,98-2,70-3,92), ! . T ! – 56,1 %, – 51,4 %. H 75 % ! ( 2). X 2 -R 2008-2010 ., – (. ./3), – (/3) H H H H ] 2008 . F 5,907 0,016 1,20 0,018 2,027 0,027 9,134 0,061 G 1,280 0,002 0,146 0,001 0,962 0,004 2,388 0,007 ^ 1,927 0,003 0,532 0,003 1,310 0,008 3,760 0,014 2009 . F 3,070 0,011 3,583 0,029 1,317 0,004 7,970 0,044 G 2,040 0,027 4,373 0,085 1,220 0,015 7,633 0,127 ^ 2,797 0,012 0,827 0,009 0,753 0,004 4,377 0,025 2010 . F 5,467 0,023 1,443 0,035 1,670 0,026 8,580 0,084 G 9,280 0,064 1,60 0,078 3,62 0,022 14,50 0,164 ^ 6,617 0,019 1,713 0,053 1,70 0,008 10,030 0,080 1994-1996 . F 0,60 <0,001 3,0 0,052 0,90 0,017 4,50 0,069 G 1,7 0,006 5,2 0,106 2,4 0,061 9,30 0,173 ^ 0,6 0,003 2,4 0,099 3,0 0,089 6,0 0,191 K , , . X 2008 . – 59,6 %, F. longiseta – 51,9 %, – 47,6 %, 23,5 % Ar. salinus. H 2009 . – 44 62,8 %% , B. longirostris P. pediculus. H 2010 . ! – 64,5 %, 27,1 % ! K. q. quadrata, – 50,5 %, 23,8 % D. galeata. H SY. . 1 (33). 2012 201

T -R ! . - , ! !. W ! 1994-1996 ., (11,4 3 15,7 3) , , ! , ! ! ! , ( 2). S! , , ! [8]. M ! 2008-2010 . !!, ! . X . ! «!!» ! , !!! . V, ! , ! – , , [9]. H ! , . Q 1 R ! (, ). – F., 2006. – 27 . 2 R ! ". M !. – Q., 1984.– 33 . 3 Q. F. WWWV (Rotatoria). – Q., 1970. – 744 . 4 T V . V / W. . \. – W.-U.: S, 1995. – X. 2. – 628 . 5 T K WWWV / T. . Q. F. . . W. – Q.: I, 1977. – 512 . 6 G K. H., H I. I. M // T ! . – Q.: S, 1979. – W. 169-172. 7 F. W. T! !. - R.: H! , 1986. – W. 286 – 348. 8 W. U. c – R.: S, 1984. – W. 129-131. 9 I! F. R. J . R.: - S, 1987. – 189 . *** G' N'-R % = 2008-2010 . % '. P' ' ^ ' % ' . *** In the present work results of researches of zooplanktonic community during 2008-2010 with the account taksonomy structure and distribution in reservoirs Ijr-Majtansky systems of delta of the river are stated Or. In interannual aspect have been calculated an information index of a specific variety on Shennonu and an index of specific similarity on Serensenu.

YJ 576.895 .. RQ UT M]M IQbRTZ T RZO M U

P. U (Rana arvalis) 5 : Opisthioglyphe ranae, Haplometra cylindracea, Pleurogenes intermedius, Rhabdias bufonis Oswaldocruzia filiformis. N Alaria alata, Strigea strigis S.falconis, Sphaerirostris teres. Q, . Y : Haplometra cylindracea, Rhabdias bufonis Oswaldocruzia filiformis, Y- P – : Opisthioglyphe ranae, Rhabdias bufonis Oswaldocruzia filiformis. Y F 4 : Opisthioglyphe ranae, Haplometra cylindracea, Rhabdias bufonis Oswaldocruzia filiformis. Y ! (Rana ridibunda) Y- – Opisthioglyphe ranae Oswaldocruzia filiformis, . F – 3 : Skrjabinoeces sp., Strigea falconis, Alaria alata. Y ! W K H- . Y (Bufo viridis) . F 4 , !: Acanthocephalus falcatus, Rhabdias bufonis, Strongyloides sp., Cosmocerca commutata, Agamospirura magna. Y K U – Oswaldocruzia filiformis. Y (Bufo bufo) H- : Rhabdias bufonis Oswaldocruzia filiformis. U W U . 202 Y . . 1 (33). 2012

I , ! ! . H X.S.W [1] ! (Rana ridibunda) 7 3 . 6 : Gorgodera pagenstecheri, G.asiatica, Gorgoderina vitelliloba, Opisthioglyphe ranae, Haematoloechus (Pneumonoeces) variegates Skrjabinoeces similis – , – Codonocephalus urnigerus – . – Rhabdias bufonis Oswaldocruzia filiformis – , Filaria gen.sp. – . I ! ! (Rana temporaria) ! : O.ranae Pleurogenes intermedius. , ! – R.bufonis O.filiformis. H U , . , R ! ! ! 80- . H.I.H S.K.X [2, 3, 4, 5, 6]. H U U ! 5 , : Opisthioglyphe ranae, Haplometra cylindracea, Pleurogenes intermedius : Rhabdias bufonis Oswaldocruzia filiformis. Alaria alata, Strigea strigis S.falconis, 1993 . S.K.X ! Sphaerirostris teres [7]. RQ RTJ H 2005-2011 . U 1241 . !, 2005 . – 170, 2006 . – 250, 2007 . – 200, 2008 . – 152, 2009 . – 201, 2010 . – 116, 2011 . - 142 . G !, !, . Y – . , . H H- ! 3 . T! ! Y- 1987 (28 . Rana ridibunda). W ! Y- ! Q ( . Y) 1986 . H ! Q (H-! ) 1986 6 . Bufo bufo, Y-, ( 1987 ), ! . Q 2011 . G H- 50 . ! – . H F 2010 . ! K 35 . !. H ! 1987 . F ( «» - «») 36 . , . F – 46 . !. G 40 . Rana ridibunda . W SFS V. 9 . 17 ! 2010 . . F – W. H , , , !. 33 . . F.R K U 2009 . Q 1985 U , W 2 . . J ! [8]. U , .R.V . [9]. MQb QJT TLJ H . , 2005-1011 ., ! 5 , 80- .: Opisthioglyphe ranae, Haplometra cylindracea, Pleurogenes intermedius, Rhabdias bufonis Oswaldocruzia filiformis. X P.intermedius, ! , 2006 2011 . Strigea strigis S.falconis, Alaria alata, O.ranae H.cylindracea, ! ! !. I ! ( Y- ) , X.S.W [1]. H ! ! – Opisthioglyphe ranae Oswaldocruzia filiformis. c ! O.ranae (67.86±8.33% 4.75±1.17 . ), O.filiformis (17.86±7.24% 0.29±0.13 .). R ! ( ), 2011 . G , . H ! Q, . Y ! : Haplometra cylindracea, Rhabdias bufonis Oswaldocruzia filiformis. M ! R.bufonis O.filiformis , H SY. . 1 (33). 2012 203

! H.cylindracea . Y H.cylindracea ; ! O.filiformis R.bufonis . H, , . Y ! Y- : Opisthioglyphe ranae, Rhabdias bufonis Oswaldocruzia filiformis. ] , O.ranae . H !, ! , ! O.ranae (4.75±1.17 .) O.filiformis (0.29±0.13 .), R.bufonis Rana ridibunda ! . V ! Y- ! : Rana ridibunda , R.arvalis ! . H ! . Q, . Y, : Rhabdias bufonis Oswaldocruzia filiformis. U ! O.filiformis ; ! R.bufonis . Y , 1987 . Y- ( ! , ) – 60 – 2 . R.bufonis 20 . O.filiformis. H H : , . Y ! F 4 : : Opisthioglyphe ranae, Haplometra cylindracea : Rhabdias bufonis Oswaldocruzia filiformis. Q , ! , . W ! F U ( 2005-2011 .) ! – , U 2006 2011 . ! Pleurogenes intermedius. W ! F U . H ! – ! ! , , «!» ! ! – !. Y ! . F : 46 . Rana ridibunda 2 . Skrjabinoeces sp. Strigea falconis, ! , Alaria alata. Y ! W . T , , . Y . F 4 , !: Acanthocephalus falcatus, Rhabdias bufonis, Strongyloides, Cosmocerca commutata. Y Agamospirura magna. R.bufonis, C.commutata A.magna – . A.falcatus Strongyloides sp. .R.V . [9] ! ! . H ! A.falcatus !, Strongyloides sp. – ! !. H .R.V . [9] , ! ! , . U ( , .R.V, H.U.^, S.S.^), G.I- Strongyloides spiralis – . U-, , F , . M ! , 70 . A.falcatus. c ! R.bufonis Strongyloides sp. ; C.commutata. H 2010 . 3 – Acanthocephalus falcatus, Rhabdias bufonis, Cosmocerca commutata. U , W U 1985 ., . H K U – Oswaldocruzia filiformis , ! ( 33 1, 2 6 .). Y !, ! ! , 3 – R.bufonis O.filiformis O.ranae. Y ! ! ! R.bufonis, , , ! . K , , ! Q3 - . T! – , ! ! ! ! : ! !! . M, ! 204 Y . . 1 (33). 2012

! !. T O.filiformis ! ! !, . X W.H.X S.K.X! [10]. Q 1. W X.S. . – H .: c! . – F-F, 1975. – W. 186-192. 2. H H.I., X! S.K. G! Rhabdias bufonis W U. - J. HSX, 1988 ., 3 4146-H88. - 17 . 3. H H.I., X! S.K. G! Oswaldocruzia filiformis H W U. - J. HSX, 1988 ., 3 4147-H88. - 27 . 4. H H.I., X! S.K. G! Opisthioglyphe ranae W U. - J. HSX, 1988 ., 3 4148-H88. - 21 . 5. H H.I., X! S.K. M ! Rana arvalis R. - J. SX 12.08.93 ., 3 3971-93. 6. H H.I., X! S.K. I . - J. SX 12.08.93 ., 3 3969-93. 7. X! S.K. U!! ! . J. … . . : 03.00.19. – F: SH FT «F!», 2007. – 281 . 8. G W.S., W .G., U H.. I . - F-F: - FS WWV, 1962. X.1. – 377 . 9. V .R., ^ H.U., ^ S.S. I WWWV. – R.: S, 1980. – 279 . 10. X W.H., X! S.K. V ! ! Oswaldocruzia filiformis //R R - «G, - », U, 22-23 !! 2009 . – U, 2009. – W. 103-106. *** R %' '' ' . U ' (Rana arvalis) ' 5 ' – Opisthioglyphe ranae, Haplometra cylindracea, Pleurogenes intermedius, Rhabdias bufonis Oswaldocruzia filiformis '. J%' Alaria alata , Strigea strigis % S.falconis , Sphaerirostris teres %'' . Q ''', '' ' 3 ' – Haplometra cylindracea, Rhabdias bufonis % Oswaldocruzia filiformis, & 3 ': Opisthioglyphe ranae, Rhabdias bufonis % Oswaldocruzia filiformis . F ' ' 4 ': Opisthioglyphe ranae, Haplometra cylindracea, Rhabdias bufonis % Oswaldocruzia filiformis . & ' ' 2' – Opisthioglyphe ranae Oswaldocruzia filiformis, F ' 3 ': Skrjabinoeces sp., Strigea falconis , Alaria alata '. ^ '' % K' ' ' ' . F (Bufo viridis) ' ''' ' ' 4 ' - Acanthocephalus falcatus, Rhabdias bufonis, Strongyloides sp., Cosmocerca commutata % Agamospirura magna %'' '. U K' ' ' – Oswaldocruzia filiformis . ^ %'' (Bufo bufo) 2 ' - Rhabdias bufonis % Oswaldocruzia filiformis '. U '' '' ' %'' ' . *** In the article the original author’s materials on the helminthes fauna of anural amphibians in the different point of Kazakhstan were representative. In the moor frog (Rana arvalis) from Pavlodar region 5 mature helminthes species: Opisthioglyphe ranae, Haplometra cylindracea, Pleurogenes intermedius, Rhabdias bufonis Oswaldocruzia filiformis – were registered. Among the larval forms mesocercaria Alaria alata, metacercaria Strigea strigis and S.falconis, larvae of acanthocephalan Sphaerirostris teres were discovered. In the neighbourhood of Leninogorsk, in flood-land of Ulba river in this host three helminthes species - trematode Haplometra cylindracea, nematodes Rhabdias bufonis and Oswaldocruzia filiformis, and in the city agglomeration of Ust-Kamenogorsk – also 3 worm species - Opisthioglyphe ranae, Rhabdias bufonis Oswaldocruzia filiformis – were recorded. In the moor frog from Akmola region 4 mature helminthes species – Opisthioglyphe ranae, Haplometra cylindracea, Rhabdias bufonis Oswaldocruzia filiformis – were registered. In the lake frog (Rana ridibunda) in Ust-Kamenogorsk 2 worm species - Opisthioglyphe ranae and Oswaldocruzia filiformis, on the outskirt of Almaty city 3 species - Skrjabinoeces sp., larvae Strigea falconis, larvae Alaria alata – were recorded. In the green toad (Bufo viridis) in Almaty city we found 4 helminthes species parasitized in the mature form - Acanthocephalus falcatus, Rhabdias bufonis, Strongyloides sp., Cosmocerca commutata, and larval stage of nematode Agamospirura magna. In green toad from Ekibastuz district of Pavlodar region only one parasite species – nematode Oswaldocruzia filiformis – was registered. In the common toad (Bufo bufo) from Eastern Kazakhstan region two nematode species - Rhabdias bufonis Oswaldocruzia filiformis – were registered. By the autopsy of 2 exemplars common toads from Selety river in Pavlodar region helminthes weren’t find.

YJ 574.587 (262.811) 2.. , 1T.. , 1J.. , 1M.. TT_ RTMTTT MTIT T UOTIT RT ¹ , e-mail: [email protected] ² , e-mail: [email protected]

U P 2008 . S P 2008 . U . T , , , , . V ! , , , [1, 2, 3]. H !, !! . \ ! ! ! 2008 . H ! : W W ; W W ! ; ! , . H SY. . 1 (33). 2012 205

RQ RTJ QJTO R ! , , 2008 . !, !! R (V 1). T ! ! H ( 0,1 2). S 366 GG , 10% 4% «G! ». H [4] , , 0,5 , – . J 1 ². W! ( ) Primer v5. U G!- , , . U . MQb QJT TLJ W !! : W ! W !. I X, ! X ! .

) ) 1. (./²) , R T W ! ( 75% ). H Oligochaeta (45,6 %), Hediste diversicolor (14,6 %), Manayunkia caspica (8,2 %), Hypaniola kowalewskii (8,0 %). T !! Abra ovat, ! : Oligochaeta (40,2 %), H. diversicolor (16,3 %), A. ovata (13,7 %), H. kowalewskii (11,4 %), M. caspica (5,3 %). Y ! 85% . W ! : ! 3,3 22,1 ( – 8,1 ), " – 1,5 11,7 ‰ ( – 7,5, 206 Y . . 1 (33). 2012

– 8,9 ‰). W ( !) ( ) / , ! . H! , (TUU) <1%. J! W ! , . H ( 70% ) : Corophium chelicorne (17,2 %), Oligochaeta (14,5 %), Corophium volutator (12,3 %), Dikerogammarus haemobaphes (7,7 %), Corophium mucronatum (6,4 %), Jaera sarsi caspica (5,7 %), H. diversicolor (4,9 %). T 80% Chaetogammarus pauxillus (10,4 %), Corophium nobile (8,4 %), D.haemobaphes (8,4 %), Mytilaster lineatus (4,2 %), Amathillina cristata (3,2 %). J ! 5,4-82,3 ( 33,9, 32,5 ), " 8,2 11,9 ‰ ( 10,5, 11,5 ‰). S ! ! / , ! , – . V! W W ! ! ! ! ! , ! (/ ). F , ! 90% ! ! , , W D. haemobaphes, J. sarsi caspica, Chaetogammarus ischnus, Ch. sowinskyi, Amathillina cristata, Am. pusilla, Corophium robustum, Fabricia sabella caspica; W Pterocuma pectinata. S Oligochaeta (7,7 %), C. chelicorne (7,3 %), M. caspica (5,6 %), H. diversicolor (5,2 %), C. volutator (4,8 %), H. kowalewskii (4,7 %), C. mucronatum (3,5 %), S. macrurus (3,3 %). M.lineatus (3,1 %). U 64% W W !. H , ! . . U! (V 1, , «») : Oligochaeta (34,4 %), M. caspica (18,2 %), H. diversicolor (16,9 %), H. kowalewskii (11,4 %), 80% . J! !, 3,7 20,9- (!! 9,1 ), 6,5‰ 11,7‰ ( 8,6‰). I , , . U 1 +11,7ºW +21,2ºW, 4 – +15,0 +21,0ºW, +19,5ºW. V , TUU<1c. H! (V 1, , «!») ! 3 , ! : Oligochaeta (46,83 %), Stenogammarus (Stenogammarus) macrurus (15,7 %), P .pectinata (10,6 %), H .diversicolor (7,6 %), S.(S.) similis (6,5 %). J! 3,5 12,0 ( 6,4 ) " 1,5 ‰ 10,8 ‰ ( 5,9 ‰). I , – . X 1 +16,8º – +22,3ºW, 20,1ºW, 4 – +16,1 – +21,5ºW, – 19,8ºW. V ! TUU<1c. V! ! W ! ! , . U 60% 7 : M. caspica (12,4 %), S.(S.) macrurus (9,6 %), H. diversicolor (8,8 %), H. kowalewskii (8,2 %), Oligochaeta (7,6 %), S.(S.) similis (7,1 %), P. pectinata (6,9 %). H W ! , . K 3 , ! . U! X . J! (V 1, , «!») : H. diversicolor (48,9 %), A. ovata (29,1 %), Schizorhynchus bilamellatus (7,9 %), Oligochaeta (7,9 %). c 6,2 16,2 (!! 11,5 ) 10,7-11,7‰ ( 11,4 ‰). I - , . X 1 +17,6 - +22,8ºW, +20,2ºW, 4 – +16,4-+22,4ºW, +19,7ºW. V ! , TUU <1c. H! W W ! X (V 1, , «#»). J! , , 35% . V H SY. . 1 (33). 2012 207

: Oligochaeta (31,4 %), S(S.).similis (19,1 %), A. ovata (15,3 %), M.lineatus (10,8 %), Cerastoderma lamarcki (9,7 %). W ! !: 3,3 9,0 ( 6,9 ), – 9,2-10,9 ‰, 10,2 ‰. I , , – ! , . X 1 +21,8 - +23,4ºW, +22,8ºW, 4 – +21,8 - +23,7ºW, +22,9ºW. H! TUU<1c. X, " (V 1, , «»), A.vata, 88% . V : Oligochaeta (42,9 %), H. kowalewskii (16,9 %), H. diversicolor (12,9 %) A. ovata (9,1 %), M. caspica (6,3 %). W 3,5 22,1 (!! 7,3 ), 6,4 11,7 ‰ (!! 8,2 ‰). I - , – . X 1 +16,5º - +23,3ºW, 4 – +18 - +23,3ºW ( +22,2ºW). V ! , TUU<1c. W! W ! . J! W ! ! ! . W ! , ! ! . H ! . X , W ! . U! F X (V 1, , « »). W – 43,5%. ^ , , , 75% : C.volutator (21,2 %), M.lineatus (14,5 %), Ch.pauxillus (12,6 %), Am.pusilla (12,1 %), D.haemobaphes (8,3 %), H.diversicolor (6,3 %). W ! : 7,9-32,5 , 19,6 , ! 8,3 11,0 ‰ (!! – 10,1 ‰). X 1 +11,0 - +19,8ºW (!! +14,1ºW), 4 – +9,4 - +19,5ºW (!! +13,0ºW). W , , !. H ! , (36,0%) (V 1, , « »). X , ! , , ! – C.chelicorne (25,79 %) Oligochaeta (19,03 %). U 13,0-78,0 (!! 40,0 ) " 8,2-11,1 ‰ (!! 10,7 ‰). I ( ), . X 1 +8,4 - +19,7ºW (!! +12,2ºW), 4 – +7,4 - +18,7ºW (!! 12,0ºW). H . H ! 3 . H , F . S! ! , ! ! M.lineatus (18,4 % ). H , ! - , ( ). I ! 5,4 18,5 ( 10,9 ), " – 11,7 11,9 (!! 11,8 ‰). H! , , ! , : Ch.pauxillus (28,2 %), C.chelicorne (11,8 %), Hypania invalida (11,7 %), Corophium robustum (11,6 %), Oligochaeta (9,7 %), Dreissena rostriformis (7,0 %), Balanus improvisus (5,7 %). Y 14,0 82,3 ( 32,0 ), " 10,6 11,9 ‰ ( 11,5 ‰). W , , ! . X 1 ! +12,9 - +22,6ºW (!! +18,0ºW), 4 – +12,1 - +21,8ºW (!! 17,3ºW). X! , ! : C.chelicorne (28,2 %), C.volutator (11,7 %), C.nobile (10,3 %), D.haemobaphes (9,1 %), Ch.pauxillus (8,4 %), Oligochaeta (6,9 %). Y! : 44,6-62,0 ( 55,0 ), " 10,6-11,7 ‰ ( 10,9 ‰), 1 – +10,8 - +16,8ºW (!! +14,6ºW), 4 – +9,0 - +15,2ºW (+13,2ºW). H ! ! ! , ! -" .

208 Y . . 1 (33). 2012

MQM] R ! ! – W ! W !. W ! ! ! , , ! , ! !! A. ovata. H W , ! , !! M.lineatus. I X, ! X ! . H W ! ! . H, ! – , , . T , – . T , H !, W ! ! !! , W ! . H! ! , W : ! , – . T , ! ! W ! !, !!! . H W ! ! 2 , ! ! !. H, ! - . T . H ! ! M.lineatus, ! ! [5]. H , ! . Q 1. I! ! . , 1996. - X. VI, 322 . 2. . ! . R., S, 1985 . – 278 . 3. K.F. !. G! !. R., HSVT, 2007. – 142 . 4. F !. U . .F. G. R., U! , 1968. 5. H.U. , F.. W. S ! ! !!. F, 2000. – 180 . *** R P ''' 2008 ' % '. 2008 , ' % T P . R '' '''. R '' , '' % '' ' . *** Data on distribution of macrozoobenthos within Kazakhstan sector of the Caspian Sea in spring and autumn 2008 are presented. Based on the analysis of data on number of macrozoobenthos borderline between Northern and middle Caspian Sea in 2008 has been determined. Season dynamics of macrozoobenthos community is shown. Main factors affecting distribution of macrozoobenthos within water body under study are: water salinity, soil characteristics, and depth.

YJ 574. 582 1X.X X., 2H.F. , 2K.H. MT JR MTTUQT IQTTTJTO T JIT U (]T «») 2009 I. 1XTT «P », [email protected] 2 XTT «P» . F, P , - «S» P 2009 ., . W ! 35.39 %, 36.63 % ! 175.6 770 . U! W ! 12 13 ‰. R ! W ! ( «S») !, ! 2009 . 22- !. U (66 ) J 50– ! ( 18- !, ! 50 4- !, 50 , ). M ! ( RGW–10 MICROS MC300) ! [2,3,4,5]. H G ! ! H SY. . 1 (33). 2012 209

. J! . ! . J! !! !! ^ – Y [6] S «S» : - I: 10 161.0 310.0 (!! 224.2 ) – II: 12 18.4 71.0 (!! – 43.6 ). H 2009 . W ! . H ! 34 , , !! : Protozoa – 3, Coelenterata - 2, Ctenophora – 1, Nematod - 1, Rotifera – 2, Cladocera – 13, Copepoda - 5, Cirripedia – 1, Molluska -1, Polychaeta – 1, Amphipoda – 1, – Osracoda – 1. ! ( 1). X 1 Z «» 2009 . X ] , % ! Protozoa - Foraminifera sp. -18.2- Arcella vulgaris Ehrenberg 4.5 - - Tintinnidium sp. 4.5 Coelenterata - Cordylophora sp. 9-- Blackfordia virginica - - 13.6 Mayer tenophora - Mnemiopsis leidyi Mayer - - 45.4 Nematodes - Nematoda sp. 13.6 - - Rotifera - Synchaeta cecilia fusipes Buchholz - - 9.0 Synchaeta sp. 4.5 - 4.5 Cladocera - Podon intermedius Lilljeborg 36.4 86.4 68.2 Pleopis polyphemoides Mord.-Boltovsk/ 45.4 86.4 68.2 Evadne anonyx G.O.Sars 100 100 45.4 Evadne anonyx deflexa G.O.Sars 4.5 - - Evadne anonyx prolongata G.O.Sars 40.9 100 9.0 Evadne anonyx producta G.O.Sars - 22.7 45,4 Podonevadne camptonyx typica (Sars) 4.5 4.5 - Podonevadne c. macronyx (Sars) 9.0 4.5 - Podonevadne angusta (Sars) - 4.5 - Cercopagis pengoi (Ostr. ) - 4.5 - Cercopagis micronix G.O.Sars - 77.2 - Apagis cylindrata G.O.Sars - 31.8 - Apagis beklemischevi M-Boltovskoi - 9.0 - Copepoda – Halicyclops sarsi Akatova 31. 8 9.0 - Calanipeda aquaedulcis Kritsch - 59.1 - Acartia tonsa Dana 100 100 100 Nitocra typica Boeck - 4.5 4.5 Ectinosoma sp. 4,5 - - U Ostracoda sp. 100 100 100 Gammaridae fam. gen. sp 4.5 4.5 - Cirripedia fam. gen. sp. 77.2 54.5 77.2 Mollusca fam. gen. sp. 77.2 100 72.7 Hediste diversicolor (larvae Nereis) 4.5 4.5 40.9 22.7 59.0 50.0 : 21 24 18 210 Y . . 1 (33). 2012

H , ! 2009 . ! P.intermedius, P.polyphemoides, E.anonyx, E.a. prolongata, A. tonsa, Ostracoda sp., Cirripedia fam. gen. sp., , ! H. diversicolor, . T !! . H , 13.1-14.9W, 21 , . S ( 100%) ! W Z ! , E. anonyx, U - , A. tonsa - Ostracoda gen. sp.. V (77.2%) - Cirripedia fam. gen. sp . T 4.5 - 41.0%. J !!! A.tonsa, 40.5 – 81.2 % . U ! E. anonyx, 68.5 – 85.2 % . H 2009 ., 23.1 - 23.8W, - 24 . U , ! Apagis Cercopagis: A.cylindrataa, A.beklemischevii, C.pengoi, C.micronix. H , , . U ( 100%) , , E. anonyx, , A.tonsa. Ostracoda gen. sp.. . J W ! P.intermedius, P.polyphemoides , C.micronix ( 86.4 -77.2 %). J Apagis Cercopagis – 4.5 - 31.8 %. W 59.1% ! , ! W !, C. aquaedulcis. H ( 90.0%). T 4.5 - 41.0 %. F ! A.tonsa, 92.4 – 96.6 % . U , , E. anonyx (52.8 - 61.2% ). H ! - 18 . W Cercopagis, Apagis, Podonevadne. T C.aquaedulcis . S ! !! ! - B. virginica – M.leidyi. !, M. leidyi «S» ! ! . S ! , , !. U ! ( 100%) ! A. tonsa Ostracoda gen.sp. S ( 68.2 – 77.2%) P.intermedius, P. polyphemoides, Cirripedia gen.sp. W 40.9%, 10 , , (0.425 – 0.950 ), ! , H.diversicolor. ! ! (13.6 – 50.0 %). J ! A.tonsa, ! 95.6-96.2%% . T , ! . U ! A.tonsa, 80.3– 91.9%. Y ! W ! . T W . U , ! !! ! «S» !! – 7.21 7.53 . ./3, , ! ! 5.28 . ./3. U ! ! , ! 2.8 , – !. c ! - . «S» !!! ! ! . W - I (!! – 224.2 ) - II (!! 43.6 ) 2. X 2 (N, ./3) (B, /3) I II «H» C 2009 . R! Rotifera Cladocera Copepoda U H I (h .=224.2 ) R 0.004 0.0044 1.42 586.79 9.60 319.45 0.33 16.52 11.37 922.78 H SY. . 1 (33). 2012 211

0 0 0.21 132.438.8126 0.03137. 1.34 9.07 271.08 W! 0 0 0.02 1.76 4.67 58.48 0.023 0.11 4.70 61.46 II (h .=43.6 ) R 0 0 1.26 604.24 1.16 35.99 0.56 5.86 3.04 646.09 0 0 0.29 203.535.52.92 0.2174 5.33 5.99 283.79 W! 0.03 0.004 0.08 11.84 5.54 54.68 0.21 1.18 5.85 67.71 J , - ! I, ! ! – 36.61 20.94 . /3 ! 300 310 . U - A. tonsa 8 , . S II 2 – 3 , , , ! , ! ! -. X! ! W ! ! [7]. T II !! . H I ! ! 2 . T , , ! – M. leidyi B. virginica, ! , ! . T (81.2 – 98.8 %) , A. tonsa. H! ! A. tonsa ! . H – , – 51.4 – 65.0 % – 31.6 – 37.26 % . Q 2.8 – 9.3 % ! – . H !! 13.2 – 17.4 % ! , 61.5 – 65.3 % ! 15.8-24.6%. H ! !! ! - – 7.6 – 8.5 %, ! ! 49.4-59.6 % 32,8 -41.4 % !!! – . U !! !. T . W! ! . H ! ! – 757.16 /3 . 271.36 /3. S – 64.12 /3, , !. U (68.5 – 85.2 %) - E. nonyx, . F 80.4 – 91.9 % A. tonsa. V ! ^-Y , ! 2009 . W ! ! – 1.9 – 2.06 – 2.48 /, ! ! ! . Q 1. R !. M !.- Q., 1984.- 34 . 2. T V . - WU, 1995.- X.1.- 590 . 3. T V .- WU, 1995.- X.2.- 632 . 4. F !. - R.: U! .- 1968. – 414 . 5. U S.Z., I F.J., ^ S.H., K.., X J.. Acartia tonsa (Copepoda, Calanoida) – F // Vestnik zoologii.- 2002. - 36(5). - W.65-68. 6. R c, c . – R.: R. – 1992. - 154. 7. M Q.F. G! WWWV. – R. – 1963. – 734 . *** 2009 T P' «S» , % ''', ' , ' ''. *** The species composition and quantitative development iIdetified of area-season .dinamic of zooplankton deeppor tof the Middle Caspian sea during the growing season in 2009, assessed of status association.

212 Y . . 1 (33). 2012

YJ 591.524.12.(28) Q.. MQb RTTI MTTUQT TIT U QT ORTMJ XTT « - !» E-mail: kazniirh @ mail. ru N - - P ( 2 4 ). U J – 2011 . 5 " : , ". U 3.5 / ., 8 7 /. U Brachionus quadridentatus hyphalmyros, B.plicatilis rotundiformis, Podonevadne trigona P. camptonyx, Acartia tonsa, . S 5 «» «», " . S " 2 11 . X " - " , , . " (202 172 . ./3) (113 . ./3). (3.2 – 3.7%) (1,5%), (5 – 6 % ). N ! – " . . . c ! !, , , . ! !, ! ! , , ! . V . \ ! – ! - !. RQ RTJ QJTO S ! R! Y - 2011 . U 5 ! ( 2.2 4.3 ) " . U 5 !, , ! , 5 – ! 5 – ! . W ! [1]. M ! J, , 4% . T SV. J! " " ! ! - [2]. W ! !! !! ! ! U G W [3]. S !! [1]. MQb QJT TLJ W - 2011 . 29 (. 1). c ": – 10, – 4, – 5, – 3, ! . H , , , , – , , . U B. quadridentatus hyphalmyros, B.plicatilis rotundiformis, Podonevadne, A. tonsa, (100% ). ^ K. tropica, B.angularis, F. longiseta, , C. aquaedulcis, C. maeoticus hircus (60 – 93%). U Collothecidae.

H SY. . 1 (33). 2012 213

X 1 , " (%) - , 1 – 5 " (S) - 2011 . X % 1 2 3 4 5 S 1 2 3 4 5 6 7 8 Ciliata – V 80 + - - + + + + + - - + + + + + Foraminifera - 13 - - + - + - - + - Hydrozoa – I Hydrida gen.sp. - - + Moerizia maeotica 13 - + + Ostroumov M. pallasi (Derzhavin) * 7 - + - Blackfordia virginica Mayer 7 + - - Nematoda – S 13 - - + - - + Rotifera - Trichocerca (s. str.) caspica 7 - + - caspica (Tschugunoff) * Asplanchna brightwelli Gosse 13 - - + - + + Brachionus quadridentatus 13 + - - + - - ! quadridentatus Hermann B q.hyphalmyros Tschyg. 100 + + + + + + + + + + + + + + + ! B.plicatilis rotundiformis 100 + + + + + + + + + + + + + + + ! Tschyg. B.angularis Gosse 87 + + + + + + - + + -+ + + + + ! - Keratella tropica tropica 67 + + - + + + - - + - ! (Apstein) K. tropica heterospina Fad. 93 + + + + + + + + + + + + + + + ! Filinia longiseta limnetica 67 - + + - + + - + + - + - + + + 0-! (Zachar.) Collothecidae gen. sp. 53 - + + - + + - + + - + + Cladocera – H Podonevadne trigona trigona 100 + + + + ++ + + + + + + + + + (Sars) P. camptonyx (Sars.) 100 - + + + + + + + + + + + + + + Cornigerius maeoticus 60 + + + - + - - + + - - + - + + hircus (Sars) * Cercopagis pengoi (Ostroum.) 7 - + - Copepoda – Acartia tonsa Dana 100 + + + + + + + + + + + + + + + Calanipeda aquaedulcis 73 + + + + + + - + + - + + - + + (Kriczagin) Halicyclops sarsi Akatova 7 - + - Ergasilidae gen sp. 20 - + + - - + - + - Harpacticoida gen sp. 33 - + + - + + - + + Others - J Cirripedia nauplii – S 100 + + + + + + + + + + + + + + + Mollusca Bivalvia larvae – 53 + + + + + + - + + - - + Q Polychaeta larvae – Q 100 + + + + + + + + + + + + + + + Ostracoda 7 - - + - - + H : 29 - 14-16-19 13-18-18 10-19-16 10-16 16 11-15-15 214 Y . . 1 (33). 2012

T " . T. caspica caspica !!! !. H " ! 8 !! . R ! ! , 10 14 , , ", 15 19 (. 1). S ! ! W !. , , " - , (. 2). J! , , " «». U , !, A. tonsa – 80.9 . ./3 0.31/3 (71.6 75.6% ). W , B.plicatilis -18.5 . ./3 0.04 /3 (16.4 9.8%, ). J! . Q P. trigona, «» - ! - 4.7 2.0 % , . X 2 2011 . I % W % U % ], . ./3 26.74 23.7 90.04 44.6 66.56 38.7 H 0.76 0.6 1.29 0.7 1.38 0.8 H 80.98 71.7 103.91 51.4 95.35 55.5 J 4.48 4.0 6.74 3.3 8.67 5.0 H 112.96 100 201.98 100 171.96 100 U 64.4 – 212.0 - 49.7 – 318.6 - 136.1 – 247.4 - X- 0.3 – 1.5 - 0 – 3.2 - 0 – 3.7 - , % 1.79 – 1.98 - 1.93 – 2.03 - 1.91 – 2.0 - H, /. 2.7 – 3.5 - 3.5 – 8.0 - 5.0 – 7.0 - G, /3 0.05 12.2 0.15 22.4 0.13 21.2 H 0.03 7.3 0.10 14.9 0.08 12.1 H 0.31 75.6 0.40 59.7 0.40 63.1 J 0,02 4.9 0.02 3,0 0.02 3.6 H 0.41 100 0.67 100 0.63 100 U 0.11 – 0.34 - 0.17 – 1.23 - 0.45 – 0.96 - W! , , " : – 3 , - 1.7 , – 1.3 . W! 1.8 . J , : B.plicatilis – 62.3 . ./3 0.13 /3 (30.8% 19.4% ) A. tonsa – 101.0 . ./3 0.36 /3 (50.0 53.7%). ! ! ( 1.2 ). R ( 1.4 ), (37.1 . ./3 0.10 /3). U , «» ( 1.3 ), . H !. U ! ! " " – 0.26 8.66 /3 [4]. ! ! . W! 1 – , – . ] ! !. - ! . H ! ! (8 – 7 /.) «». W! 2 – " 3,5 - 4 , 5 – 6 . X , - . H ! (7 – 6,8 /.), . W! 3 – ! 9 -11 , !! ! . ] H SY. . 1 (33). 2012 215

. X !. U (5,5 –5 /.). W! 4 – «» «». U 1,5 . W , . U ! (3,5 3,8 /.), (5 /.). W! 5 – 1,3-1,4 !. X . W (3,3 3,5 /.), (6,2 /.). ! . c , ! A. tonsa, ! ! B.plicatilis. V ! Keratella, 1 A. tonsa, ! ! . Podonevadne. H Cornigerius ! . Brachionus. S , 3.7% (. 2), ! . , ! 5 – 6 % ! 13 % - [5]. U ", ! – , !" , III , [1]. X , !! . S ! ! , ! ! !. S 5 (3 4) «» «», " . S " ! (3 1, 3, 5) 2 11 ! . X (3 2) ! " - " , , . H ! " !!. H , ! . H!! ! . M ! ! !. Q 1. V .- Q.: I.- 1983.- 240 . 2. G K.H., H I.I. M // T ! .- Q.: S.- 1979.- W. 169-172. 3. Y ! // R . - R.: WcH, 1975. - ].3. - 176 . 4.^ Q.. ! 2008 – 2010 . // R IV . -. . «U ! ! ! ! ». F: SV. – 2011.- W. 269 – 274. 5. F.W. W " G ! – 1987 . // G !. Q.- 1990.-H. 3.- W.128 – 135. *** P ''' '- '''' (2-4 ') '- ' ''' '' %' '. U 2011 '- 5 , , '''' ' % ' %' ' ' . M ' ' ' 3,5 /, ' 7-8 /- ' '. P Brachionus quadridentatus hyphalmyros, B.plicatilis rotundiformis, Podonevadne trigona, P. camptonyx, Acartia tonsa, % ' %'' ' . 5 '' « ''''» ' % « » ' ' , '' % ' . 3 ''''' % ' ' %' % 2- 11 ' '. X ' ''' ', ''' 3 – 4 , ' ' ' '' ''. M '' %'' ' % ' (202 % 172 ./3 ) ' (113 ./ ). ' %'' (3,2 – 3,7%) (1,5%) ' , ' ( 5-6%) . ! % ''' . 216 Y . . 1 (33). 2012

U ' ' ' ' % %' . G %' '' ''' %' . *** Effect of seismic survey on zooplankton resulting caused by engineering and geological prospecting in the North-East part of the Caspian Sea were researched (from 2 to 4 meters deep). The zooplankton was collected with the Juday net in July – August 2011 at 5 stations in three replications: baseline, during the seismic survey and 24 hours thereafter. In baseline studies the wind force did not exceed 3.5 m per sec., subsequently it rose up to 7-8 m per sec. Widespread were rotifera Brachionus quadridentatus hyphalmyros, B.plicatilis rotundiformis, cladocera Podonevadne trigona and P. camptonyx, copepoda Acartia tonsa, cirripedia nauplii and polychaeta larvae. At one of the five stations the plankton counts in the baseline studies were equal to the plankton counts in the seismic survey, with almost complete absence of damaged species and low wind force. At the three stations the plankton counts and weight increased from 2 to 11 times in the phase of earthquake effect and strong wind. At one station only during the seismic survey a three-fourfold decrease in the number of community species was identified, which significantly increased in the subsequent period, when there was a strong wind as well. On an average for the study are, diversity and number of the animals increased during the seismic survey and thereafter (202 and 172 thousand specimens per m3) as compared to those in the baseline studies (113 thousand specimens per m3). In the last two periods sensitivity of invertebrate animals was a shade higher as well (3.2 - 3.7%) as compared to that in the baseline studies (1.5%), but the percent of such species did not exceed a known level of natural distortion (5-6% of the weight). Indicators of ! – saprobic zone and saprobity indices allowed to unambiguously characterize a water column as moderately contaminated at all stages. The plankton dynamics identified was caused by a more intensive wind activity during the last two periods and water mass transport from usual aggregated zooplankton. This type of observations did not identify any relation between the zooplankton dynamics indices and the effect of earth shocks.

YJ 574.587 Q.. TQTI] UTQJ JQbT ITTJTM_ UJUO JQ JT TT_ TJTTT Q WT VFS, e-mail: [email protected] F, . T , (, , ) , . J . H ! - ! , .. , . H ! ! [1]. R ! - , , . M! ! !! ! [2]. F! ! , . T . , ! V , , , . H ! ! ! ! - F! (V F), ! ! . . R W-H Z- F ( .T), , !! . !! Z- F! 1990- F - ! (FIRU), ! . W 1990- ! ! ! . T , ! ! [2]. T !! !! !: ; , ! ; , [3]. T !! ! !! ! !, .. ! [4]. H ! , ! [5]. T !! !! ! , .. , , , , !! [6]. \ – F!, !! !! !. H SY. . 1 (33). 2012 217

T !! , ! TTT "F - !" (FIRU): . , ], .]!, . ( .!) «H» (W - ): . W , . W, . a, . W ( . G!). J! ! . Y ( ! ! «», «X- », «Y ») [7]. T ! ! ! ! . U! , , ! !!. H ! ! , !! ! . H ! !! ! , !, : KVX (!, ), , [8, 9, 10, 11]. J! ! !! , ! : – ! ; – !. U ! ; , ! . R ( , , , !), ! !! ! . G , KW W^F, ! ! ! !! . T ! ! !!, . H ! ! (. 1). X 1 M . . S ^ TBI BMWP ASPT EPT EBI . 2,0 6,0 36,0 7,0 4,0 9,0 . G () 2,2 6,0 9,0 4,5 1,0 9,0 . 1,5 6,0 25,0 6,2 3,0 11,0 1,9 6,0 23,3 5,9 2,7 9,7 0,21 0,00 0,58 0,22 0,57 0,12 U: S – ! ( ^), TBI – . X, BMWP – Biological Monitoring Working Party Index, ASPT – Average Score Per Taxon Index, EPT – !, , EBI – H, s – . H ! BMWP EPT , ! (. 2) ! !! . ]! . M! 1-2 . H ! , , ! H (TBI) (6). X 2 M -]-] S , ^ TBI BMWP ASPT EPT EBI . () 1,9 6,0 23,3 5,9 2,7 9,7 . 1,4 1,0 7,0 3,5 0,0 5,0 V.] () 2,8 6,0 42,0 8,4 5,0 11,0 V.]! () 2,5 9,0 53,0 7,7 7,0 14,0 V. . ! 2,8 8,0 55,0 7,5 7,0 14,0 0,25 0,51 0,51 0,29 0,64 0,34 M! (TBI, BMWP, EPT, EBI) ! ! -]-]! !! ! !! – FIRU. F ! TBI, BMWP, EBI, ASPT ! . 5-6 , . ] – 1-4 , .]! – 1-3 . T, ! ^ ! , BMWP . ! 218 Y . . 1 (33). 2012

! ! . ! (5-6 ), . ] – – ! (1-3 ), .]! – – (1-2 ). R! ! – ! !! ! TBI, ! ! !! . H EBI ASPT. – ! – ! ! .]! , . ] – ! . – ! (3-4 ). R ! ! «H» . W , ! – . W. V H , !, ! . W !! «V «H» ! , ! . T, ! ! . W, . W , !. T ! [12], . W – . M! «» . W, «» – . W . a, «» – . W . X , ! . T, !! !! !, , ! ( !, ) ! ! !! ! . S , ! . S . X ( .T) ! ! ! , , ! (, !, ) ( ) [13]. !, ^, W^F, c, U, , ! ! (, , , , !), !, ! [11, 15, 16, 14, 17]. c ! ! ! . W ! ! , ! Heptageniidae, Ephemerellidae, Taeniopterygidae, Capniidae Leptoceridae ([15, 18]. X , , ! , !! !! !. c ! !!! , ! . W ! !. TJ T !! , !! ! ! ! ! .]!, . , . W, . W . a, , . ] – ! . . W – !. U ! . T ! , ! (!, , ). W ! ! TBI, BMWP, EPT, EBI ASPT, ! KW W^F ! !!, ! ! ! !! ! F!. U . V , ! , !! !! !. Q 1. R F.S., ^ R.S., R F.., S.., c ! ! !. – H XTIY. – 2010. – 3 3 (18). – W. 115-122. H SY. . 1 (33). 2012 219

2. U F.H., V Z.H., I .H., G W.H., Q V.H. M! ! ! F! // U . - 36. – 2008. – W. 28-32. 3. ! F.H. c- ! ! ( TFT «V «H», V F). – F. ... ... – X, 2006. – 24 . 4. G H. J ! : !, , . – R.: S, 2006. – 176. 5. W H.U. U . – R: T, 2004. – 125 . 6. G, F.. ! / F.. G // G! . – 2000. – 3 1. – W. 68-82. 7. K F.F., Q.H. R Y (H ) ! // R , , !. – 2010. - 36, . 1. – W. 258-262. 8. Kaye A. The effects of mine drainage water from Carrock Mine on the water quality and benthic macroinvertebrate communities of Grainsgill Beck // Earth & Environment 2005. – 1. – P. 120-154. 9. Maret T.R., Cain D.J., MacCoy D. E., Short T.M. Response of benthic invertebrate assemblages to metal exposure and bioaccumulation associated with hard-rock mining in northwestern streams, USA // Journal of the North American Benthological Society. – 2003. – V. 22, Iss. 4. P. 598–620. 10. Tarras-Wahlberg N.H., Flachie A., Lane S.N., Sangfors O. Environmental impacts and metal exposure of aquatic ecosystems in rivers contaminated by small scale gold mining: the Puyango River basin, southern Ecuador // The Science of the Total Environment. – 2001. – v. 278. – P. 239–261. 11. Iwasaki Y., Kagaya T., Miyamoto K.-i., Matsuda H., Sakakibara M. Effect of zinc on diversity of riverine benthic macroinvertebrates: Estimation of safe concentrations from field data. Environmental Toxicology and Chemistry, 2011 v.30: 2237–2243. doi: 10.1002/etc.612 12. W F.W. !! ! F. – F.…..-.. - X, 2008. – 24 . 13. -G K. F. T !! X ! !. – F. . … . . . – G, 2004. – 24 . 14. Beltman D..J, Clements W.H., Lipton J., Cacela D. Benthic invertebrate metals exposure, accumulation, and community level effects downstream from a hard-rock mine site. // Environ. Toxicol. Chem. – 1999. – v. 18, 32. – P. 299–307. 15. Courtney L.A., Clements W.H. Sensitivity to acidic pH in benthic invertebrate assemblages with different histories of exposure to metals. // J. Nam. Benthol. Soc. – 2000. – v. 19, 3 1. – P. 112–127. 16. Malmqvist B, Hoffsten P.O. Influence of drainage from old mine deposits on benthic macroinvertebrate communities in central Swedish streams. // Water Res. – 1999. – v. 33, 310. – P. 2415–2423. 17. De Pauw N., Gabriels W., Goethals P.L.M. River monitoring and assessment methods based on macroinvertebrates. // Biological monitoring of rivers. Applications and perspectives. – Chichester: John Wiley & Sons, 2006. – P. 113-134. 18. Beasley G., Kneale P. E. Investigating the influence of heavy metals on macroinvertebrate assemblages using Partial Canonical Correspondence Analysis // Hydrology and Earth System Sciences. – 2003. – v. 7. – P. 221–233. *** X- %' %' F '' . ' ' '''' , ' ' ' (, , ) ''. ''' '. *** In work features of structure of communities of ground macroinvertebrate mountain water currents of Altai which is in a zone of influence of the mining enterprises are analysed. Decrease in the general specific riches of ground communities, reduction of number of kinds sensitive to pollution groups of macroinvertebrates (, green drakes, caddis flies) on the sites subject to pollution isn'ted. The estimation of an ecological condition of water currents is given.

220 Y . . 1 (33). 2012

W 2. -

YJ 502.72 Q.. J1, .R. 1, .. 2, U.. 2, .I. 2, M. 3, M.L. 4, J.. R5 T TMTTM TJT-TQT ITJO JQb Jb 1 RTS V, e-mail: [email protected] 2 RTS V, e-mail: [email protected] 3 F SV, e-mail: [email protected] 4 IUM «G», e-mail: [email protected] 5 F U c, e-mail: [email protected]

, - . U , , , P P P. N - . U ! ! , - ! - ! , ! ! !! !. c , , ! ! , , ! . H- , , ! ( !, !, !, !, , ! , .), . T ! ! . - - , , ! ! . H 2005 . , F R G F. U ! W , ! R F - (HGY). c ! , . H ! ! HGY !! ! ! ! (, , , .). X . S ! ! ! ! - , ], X F. ! TGWK/RWF «H ! F !: . V: R - » (2011 .). T ! , , , , , , W!, R F ! . \ : ! ! W!, , , ! , !! - , ! , !. c ! ! : , - . H 3 : (), , . F !! ( - - , ). V - W (R 1: 50000). ! Landsat, ! 148 , GPS. Q 24 . T !!! . T - - (Phragmites australis) ! (Typha angustifolia, Scirpus lacustris, S.tabernemontanii), - H SY. . 1 (33). 2012 221

(Sparganium simplex, Alisma plantago-aquatica, Cyperus fuscus, Butomus umbellatus) - (Potamogeton perfoliatus, P. crispus, Zostera noltii, Najas marina, Ceratophyllum demersum, Myriophyllum spicatum) . S W (Phragmites australis, Typha angustifolia, Scirpus littoralis, S. lacustris, S. tabernemontanii)), o- (Salix songarica, S. wilhelmsiana, Elaeagnus oxycarpa, Tamarix ramosissima, T.laxa, T.hispida, T.elongata, Halimodendron halodendron, Calamagrostis epigeios, Phragmites australis, Sphaerophyza salsula, Glycyrrhiza glabra, Lythrum salicaria, Xanthium strumarium, Calystegia sepium) (Halimodendron halodendron, Tamarix ramosissima, T.laxa, T.hispida, T.elongata, Lycium ruthenicum, Climacoptera brachiata, Suaeda linifolia, Bassia sedoides, Atriplex saggitata, Artemisia scopiformis, Leymus multicaulis, Phragmites australis, Aeluropus littoralis) . S ! (Atriplex pratovii, Salsola foliosa, S.nitraria) - (Eremosparton aphyllum, Ammodendron bifolium, Argusia sibirica) . H , 112 82 35 , , ! , !! , , . H [1] !! (Nymphoides peltatum), (Scirpus kasachstanicus), U (Atriplex pratovii). , , (Haloxylon aphyllum, H.persicum). ! ! ^ (Nitraria schoberi). c (5) - (Calligonum crispatum), ! (Atriplex pungens), ! ! (Petrosimonia hirsutissima), ! (Artemisia scopiformis), (Astragalus brachypus). T W , ! F !. H !! - () (Nymphoides peltatum). (Scirpus kasachstanicus). S ! (Typha angustifolia) Q (T.laxmannii), (Bolboschoenus maritimus), (Eleocharis acicularis), (Scirpus lacustris), X (S.tabernaemontanii), (S.litoralis), (Crypsis schoenoides), (Butomus umbellatus). H ! - Salix( songorica, S.wilhelmsiana, Elaeagnus oxycarpa, Calamagrostis epigeios, Sphaerophya salsula, Lythrum salicaria) . S U (Atriplex pratovii), ! ^ (Nitraria schoberii), : (Calligonum crispatum), Atriplex( pungens), (Petrosimonia hirutissima), (Artemisia scopiformis), (Haloxylon aphyllum, H.persicum). H . W R F ! ! 85 ( 52, 7, 15, 9). : (Lopocharis naias, Lecane nana, Lecane (Monostyla) acruata), (Neodiaptomus schmakeri). c : (Podonevadne camptonyx), (Halicyclops rotundipes aralensis). Z : (Lopocharis naias, Lecane luna, Brachionus angularis, B. calyciflorus, B. nilsoni, Keratella valga monospina), (Ilyocryptus agilis, Moina mongolica, Podonevadne camptonyx), (Acanthocyclops trajani, Thermocyclops taihokuensis), ! (Neodiaptomus schmakeri), (Onychocamptus mohammed), (Mesocylops leuckarti, Calanipeda aquaedulcis), Bivalvia. 172 13 !: U!, , V, U, L, ], U, W, G, W, V, J, V. [2]: Anax imperator, Hierodula tenuidentata, Chilocorus bipustulatus. : Balclutha chloris, Holonabis sareptanus, Solenoxyphus lepidus, Vachiria deserta, Centrocoris volxemi, Aethus comaroffii, Trigonosoma oschanini, Ochyrotylus helvinus, Desertomenid albula, Berytinus geniculatus, Hydrous piceus, Scarites subterraneus, Scarites bucida, Amphimallon solstitiale, Cardiophorus ebeninus, Papilio machaon, Vanessa cardui, Oxybelus gracilissimus, Philanthus variegates. c : Onthophagus (Exotonthophagus) haroldi, Onthophagus (Palaeonthophagus) flagrans, Tetramorium schneideri, Stizus annulatus. Z : Coenagrion pulchellum, Sympetrum sanguineum, Orthetrum brunneum, Gryllotalpa gryllotalpa, Labidura riparia, Aspilaspis pallida, Tarisa pallescens, Sigara lateralis, Sigara assimilis, Saldula saltatoria, Ilyocoris cimicoides, Cybister lateralimarginalis, Dytiscus marginalis, Rhantus pulverosus, Calosoma imbricatum deserticola, Calomera littoralis, Tabanus filipjevi, Calliphora vicina, Sarcophaga carnaria, Hemilepistus cristatus, Formica subpilosa, Cataglyphis pallidus, Pseudohadena immunda, Blaps granulata, Microdera convexa, Adesmia karelini, Paederus littoralis. 222 Y . . 1 (33). 2012

R F : - Barbus brachycephalus brachycephalus, ! ! – Chalcalburnus chalcoides aralensis, ! – Abramis sapa aralensis. R F ! 18 : ! - Clupea harengus, ! - Esox lucius, - Abramis brama, - Aspius aspius, ! - Carasius auratus, - Ctenopharyngodon idella, - Cyprinus carpio, - Hypopthalmichthyx molitrix, - Pelecus cultratus, - Rutilus rutilus, - Scardinius erythropthalmus, - Silurus glanis, - Perca fluvatilis, - Sander lucioperca, - Channa argus, - ! - Platichthys flesus. R F , , ! W Z U F F. H ! ! R F W (IBA). H ! 20-50 . , ! (F1, F4i b A4ii), R F ! . , RWTU (IUCN), ! 13 : ! (Pelecanus crispus), ! (Rufibrenta ruficollis), ! (Aythya nyroca), (Oxyura leucocephala), (Circus macrourus), (Aquila clanga), "- (Aquila heliaca), - (Haliaeetus leucoryphus), " (Aegypius monachus), (Falco cherrug), ! (Falco naumanni), - (Chlamydotis undulata), ! (Glareola nordmanni). S ! 250 , 100 !. T ! ! - () . W , " V [3], 27 , !! (10) (15) . c - (Pelecanus onocrotalus), ! (Pelecanus crispus), ! ! ! (Egretta garzetta), (Platalea leucorodia), (Plegadis falcinellus), (Phoenicopterus roseus), ! (Rufibrenta ruficollis), - (Cygnus cygnus), ! (Aythya nyroca), (Oxyura leucocephala), (Pandion haliaetus), ! (Circaetus gallicus), " (Aquila nipalensis), (Aquila heliaca), - (Haliaeetus leucoryphus), - (Haliaeetus albicilla), (Falco cherrug), (Falco pelegrinoides), (Falco peregrinus), (Grus grus), - (Anthropoides virgo), - (Chlamydotis undulata), (Larus ichthyaetus), ! (Pterocles orientalis), ! (Pterocles alchata), (Syrrhaptes paradoxus), (Bubo bubo). V (Phasianus colchicus turcestanicus), W. S ! 14 , . 5 : ( Sus scrofa), (Canis lupus), (Vulpes vulpes), (Vulpes corsac), !- (Lepus tolai). ! ! ! V ! - W !! !. J! ! ! . H TTUX ! W ! R F !. T! W, ! . J! ! TTUX " ! W 5 " !. S ! c " !! ! ! ! ! «F W» I «G». U ! 5770 , W ( ) ! ! 4074 . Q 1. U !! ! . Y U V. 31034 31 !! 2006. F. 2. Y . U . – F: F. - 2005. - 115 . 3. ! V . - . 4-, . - X.1.L. - ].1. U. - F: DPS. - 2010. - 324 . *** R '' - ' %' ''. ' ''' '', ' ' , ' ''. G ''' '' - '' # ''' '' . *** The results of the study were received during initial investigation of wetlands in the Syrdarya River forming avandelta. High biodiversity of ecosystems, rare, endemic, vulnerable, vanishing species from Red Data Book have been presented. Complex assessment of biodiversity significance has revealed global importance of wetlands and necessity of measures on biodiversity conservation. H SY. . 1 (33). 2012 223

YJ 581.9 Z.. , M.. , .I. , .. RQ\ LQ RQQT (MN, CO, NI) LIT ] . Q -

(Mn, Co, Ni) . N. , Glycyrrhiza glabra Xantium strumarium , UJP. H ! !! , ! . ! !! ! ! , . Q- ! -G , , . ! !! , . , - , ! [1]. V . U !! , ! . L, , , ! !. ! ! !. S! 0,07 1 , [2]. W! ! , !!! . W Ni Co ! , ! , ! [3]. RQ RTJ QJTO H ! ! , !, . V! , , , 105 °W ! - !!, , 3 1 . T ! ! !. H ! - . T !" - «AAS IN». X - 1400°W. F - ! : ! . H ! , . U ! !. J ! , !! ! !, ! . c ! , ! !. U ! ! !. MQb TLJ H 563 , !! 274 66 . V! !! ! . U ! ! ! ! ! . c ! Glycyrrhiza glabra Xanthium strumarium (). X 1 . U Ni Co Mn Glycyrrhiza glabra 40 ± 0,3 2,5 ± 0,5 70 ± 0,2 1 Xanthium strumarium 30 ± 0,6 2,4 ± 0,55 165 ± 0,4 Glycyrrhiza glabra 28 ± 0,4 1,3 ± 0,85 45 ± 0,32 2 Xanthium strumarium 18 ± 0,26 1,9 ± 0,57 50 ± 0,40 Glycyrrhiza glabra 10 ± 0,9 1,2 ± 0,9 65 ± 0,26 3 Xanthium strumarium 8 ± 0,8 2,3 ± 0,83 90 ± 0,2

UJ RI/I 8 – 2,8 2 – 0,98 25 – 26,8 224 Y . . 1 (33). 2012

1 , !" Ni, Co, Mn. H 1 () Ni Glycyrrhiza glabra UJ [4] 5 , Co 1,25 , Mn 2,8 . H Xantium strumarium Ni 10,7 , Co 2,2 , Mn 6,6 . H 2 (G) Glycyrrhiza glabra UJ Ni UJ 3,5 , Co 0,65, Mn 1,6 . W !" Xanthium strumarium Ni UJ 6,4 , Co 1,3 , Mn 1,8 . H 3 ( ) , Glycyrrhiza glabra UJ Ni 1,25 , Co 1,15 Mn 3,6 , Xantiums trumarium UJ Ni 2,8 , Co 2,1 , Mn 3,3 . U ! ! ! Glycyrrhiza glabra Xantium strumarium, , . H ! !! , ! . I! ! ! , , ! [5].

Glycyrrhiza glabra Xanthium strumarium X 1

Glycyrrhiza glabra Xanthium strumarium

X 2

Glycyrrhiza glabra Xanthium strumarium X 3 1. . H ! ! ! !! ! , , , , , , , . W "! " ! , ! 40 , , , ! , , ! ! , ! ! ! , ! . X! !! ! , [6]. H ! ! !! ! !! . H !, H SY. . 1 (33). 2012 225

! Glycyrrhiza glabra Xantium strumarium ! ! , UJ. Q 1. W.., X! X.G., G .G. R ! ! // Y. . ., 1995. X. 115, . 3. W. 261-275. 2. G V . - F, 1997, - 142 . 3. H . S ! V-VIII : W. ./ U . H.F. . – Q.: "!",1989. - 40. 4. G.H., W F.. R ! !.- S.:WT VFS, 2001.-216 .5; 5. G.H., W F.. R ! !.- S.:WT VFS, 2001.-216 . 6. G! ! . – R.: R, 1998. – 348 . *** G = ''' '' '''' (Mn, Co, Ni) %' ''. G'' %' Glycyrrhiza glabra % Xantium strumarium ''' UJP '' ' . *** The article presents the results of the study the accumulation of heavy metals (Mn, Co, Ni) in the plants of the lower reaches of the river. Ile. As a result of our research was found that the dominant plants in selected Glycyrrhiza glabra and Xantium strumarium actively accumulate heavy metals, nickel and manganese that exceed the MCL.

YJ 581.9 .. ., M.. , I.. , Z.. QJT TJL LQ RQQT UT] LIT ] . Q -

(Mn, Co, Ni) . N, , , Mn, Co, Ni, « – ». , UJP . U! ! , ! M . S !. K XX ! !, ! , [1]. H ! !! , ! . R ! !! !! ! , ! , , . H , ! !! , ! ! ! 40 J.. R 50 : V, W, R, Fe, W, Ni, Wu, Zn, Mo, Cd, Sn, Hg, Pb, Bi . [2]. W!, !!! ! [3]. U !!! ! . U ! ! !!! ! !, , , [1, 4, 5]. RQ RTJ QJT H ! ! , !, . - T ! : ! , ! 5 , ! ( ). U 0-20 20-40 5 !. H - 1 . T !. M - !! . T " , ! 0,5 . - J! ! - 1 5 ! . S , ! 15 5 , !, ! ! . - H ! ! : , . - H ! - . T !" - «AAS IN». X - 1400°W. F - ! : ! . H ! , . U 226 Y . . 1 (33). 2012

! !. J ! , !! ! !, ! . c ! , ! !. U ! ! !. MQb TLJ U !!! !! . W , [6]. H . - , , -, - – ! , -, - [7]. U UJ ! ! (), ! , ! UJ. X 1 . U Ni Co Mn 1 1 2,4 ± 0,2 4,7 ± 0,15 260 ± 0,5 2 2 2,8 ± 0,15 5 ± 0,12 225 ± 0,9 3 3 4,4 ± 0,2 5,2 ± 0,1 255 ± 0,7 UJ / 4 5 1500 H () ! UJ 0,94 , (G) UJ 1 ( ) 1,04 . ! UJ. S ! !. (, G, .) 0,15 0,18

0 0,7

4 Ni Co Mn Ni Co Mn

X 2 X 1

0,17

1,04

1,1 Ni Co Mn

X 3 1. . 1 , ! ! , , ! , 1,04. W ! ! 0,15. U !, ! , , , ! « – H SY. . 1 (33). 2012 227

!». H , ! UJ .

Q 1. G.H., W F.. R ! !.- S.: WT VFS, 2001. - 216 . 2. S F.R. I!: . - Q.: I,1989. – 450 . 3. G G.F. I. F-F, «S», 1978. - 287. 4. I! ! . - 2000. – 398 . 5. G! ! . – R.: R, 1998. – 348 . 6. H.G. T ! ! ! // ! .-.. – 1987. - 31. – W. 41-48. 7. U R.W. c – ! H . – F: . «c», - 2000. – W. 336 - 338. *** G = ''' '' (Mn,Co,Ni) %' ''. F %' , ' Mn, Co, Ni , % «-'''» ' ''' '''. M ' '' UJP- . *** In this article is researching influence of heavy metals (Mn, Co, Ni) on soils at Ili river downstream. According research data we identified that soil in this region accumulates heavy metals, especially Mn, Co, Ni, what causes entry them in plants via “soil-plant” system. In research results we identified that cobolt level in soil exceed Maximum Possible Concentration in all 3 points of current location.

TL 61124. 591. 424 . ļ. O= TRQQ ľUQ==İ Q ľQ=R==İ ļĲQ= TTU O=RJQ==İ ļM=QRJ I=MJ= - , - mail: saparovkyandyk @ mail.ru

G % ''' ' ' ''' '''' ' ''' %' '. ' '', % '', % ''' , ' ''' ', - '' % ' '' . c! Ŀ ı !ı: ! !, ! ! Ľ ! ! Ŀ . G- ! [1, 2] Ŀı Ľĳ ! Ŀ Ŀ. F, ! Ŀ ı Ŀı , Ľ . X ĳı ! ı Ľ ı ĳ ı . G ı, ı Ľ ĳı Ľ ĳ Ŀ , ĳ Ŀı . F Ľ ı Ŀı Ŀı, ! ĳ ı Ľ ı. RQJ R M ļ=QJ= W ! ı ĳ Ľĳ ı, ı, ĳı Ŀ . L F ı Ľ (G, V , ı 2800 X-Xĳ ) . – Amphibia - Caudata 1.L ! – Ranodon sibiricus 2. – Triturus vilgaris ! – Anura 1. J – Bufo danatensis / / 2. J - Bufo danatensis / / G – Reptilia ! – Sqvuamata T! – - Sauria 1. F ıĿ – Ablepharus alaicus Elpatiewsky 2. ^ı – Eremias grammica Lichtenstein L ! - Serpentes 228 Y . . 1 (33). 2012

1. - Ancistrodon halys pall / / 2. J – Vipera ursine Bunap Wĳ – Mammalia ! - Rodentia 1. Ľ – Clethrionomys rutilus pall 2. T !- - Clethrionomys flater Thomas ! Ľ – Lagomorha 1. ! – Lepus tolai pall 2. Ĳ ! – Oructolagus cuniculus L Ŀ ! Ľ ! . ľ ! Ľ ĳ Ŀ ĳ . G 10c . ı 3-5 - Ľ H-I Ľ !. ľı Ľ ĳ 2,5 c (pH 7,4-7,6) 2,5 Ľ 1c ı 2 . T Ľ Ŀı (812) . R! , Ŀı W Ľ ĳ [3]. SĽ Ľ (V Ľ) Ŀı. L cHR-100Q Ľ ĳ. W Ľ ĳ Ŀ (533) Ľ Ŀı , . ı ı Super – probe 733 ı Ĳ 800-4000x ĳ ĳ. M ļLQ= R TQJ QJ ı Ŀ ı ! ı , ı Ľ ı II ĳ «» ı !ı , ı . ı Ŀ ĳ . ^ Ľ «» ı ĳ Ľ. X ı, ı ! , ı Ľ ĳı ı , ! ı ! ı - ı ı ı (747,3 ± 14,2 Ľ 1310,5 ± 24,9 Ľ P<0,001). W Ŀı ı ı . J ı, Ŀ ı ı II ĳı , !ı Ŀ. W ı «ı» Ŀ , Ŀı Ŀ ı ĳ . X ı ı II ĳı . T , ı ı Ŀı . F- ĳ . Tı ı , ı (628,4 ± 11,5 Ľ 1120,2 ± 20,5 P<0,001). W ı , ĳı , Ŀı ı . J Ľ ı Ŀı Ŀı Ŀ, - ĳı ı « » Ŀ . G Ŀ ı ı ! ĳ . J ı ı Ŀı Ŀ . Y ı II ĳ Ľ «» ı ĳ Ŀ. L Ŀ, ı ĳ Ŀı Ŀ ı . c - - ı ı ı Ľ ı . ı Ŀ ı Ŀı . W ! ı Ľ ı Ŀ Ľ - ı ı Ľ. X ıĿı ı II ĳı ı . F- ĳı ı, ı ı Ľ ı ı ı H SY. . 1 (33). 2012 229

, ı ı . F ıĿı - ĳı ı (284,1 ± 5,2 Ľ 873,0 ± 5,5 Ľ P<0,001). W ı Ľ ı ı «» . F , ı : ı Ŀı Ŀı , ı ! ı - ĳı ı. Gı Ľ ı Ŀı Ŀı ! Ŀ ı ı Ŀ Ľ ı Ľĳ . J ı Ŀı ı , Ŀı ı ı , ı . G , Ľ C.X. Sı (1997) [1] Ľ. W , «» ı Ŀı ! Ľ ı ! ı , ı ı , ı ĳ . X ı Ŀı , - ĳı ı Ľ Ŀı ı . R, ı - ĳ ı (475,4 ± 8,4 Ľ 757,8 ± 14,4 Ľ P<0,001) . F Ľ , ı Ŀı , ı ! ı Ľ Ŀ ! . 1- ' ĭı -ĭ () ĩĭı ĭıĩ () Lı ĳ, M ± m L ! 1310,5±24,9 747,3±14,2 P<0,001 J ( ) J ( ) 1120,2±20,5 628,4±11,5 P<0,001 ^ı F ıĿ 873,0±5,5 284,1±5,2 P<0,001 L ( ) L ( ) 757,7±14,4 475,4±8,7 P<0,001 Ľ X!- X X 777,7±14,0 224,0±4,1 P<0,001 X ı ĳ [4]. G ĳ- ! , Ľ ı ĳ (, ! , ĳ ! ! ı ). K - , ı () Ľ ı () !. G ĳ (Ŀ), ĳ–ı , ı - . J Ľ ı Ŀı Ŀ ! ı ! Ľ ı . G Ŀı Ŀ ı . X ! ı - ĳ, Ľ ı - ĳ ı (224,0 ±4,1 Ľ 777,7 ± 14,0 Ľ P<0,001). 1 ı - ( ) ı ı () Ŀ. X Ŀı . X ı Ŀ Ŀ ı I ĳı ı ! . W Ŀ. G ı ĳı ı , Ľĳ Ľ ĳ ı ĳ [5]. 230 Y . . 1 (33). 2012

F.J.Wı [5] ĳı ! ı , Ŀ ı -Ľı Ŀ ĳ [5]. M Ľ ĳ !ı Ŀı Ŀ ! . X Ŀ !ı - ĳı Ľ ı . G !, ı Ľ ı ! . W, ĳ , ı ĳı Ľĳ ! , ĳ Ľ ĳ Ŀ. G Ŀ ı ı ( ı Ŀ Ľ ) ı Ľĳ ĳ . ļJ 1. S W. X. «G TETRAPODA » - F: - . -1997. - 384 . 2. W . ., G G. T., F G. F. «L W ı !ı Ŀ ı Ľ » // R R. S- . . «W » F: SY- 2010.- W. 268-271. 3. Q I. . «G! » - R.: H! . – 1990. – 352. 4. T S. J. «F! » // R . – R.: S, - 1992. – 33-47. 5. W F. J. «c ! ! » - Q.: - S. 1986. – 74 . *** . (, , ) , . *** This paper presents the results of the study of adaptation of respiratory part of lung of some terrestrial vertebrates at the cellular level. However, to identify the ultrastructural basis of evolutionary adaptation of lung examined the data of electron microscopic studies of representatives of some species of terrestrial vertebrates (amphibians, reptiles, mammals) from the steppe and mountain habitats. The main aim of the paper is studying of lung adaptation as main external gas exchange organ to a variety of habitats in the evolution in animals of different species.

YJ 502.72 .R. , .. , .. , .. , M.I. , .. I , J.. R, .. , .. TQTI]T MTTM UTRTIT Q-QTIT UTJTIT M N RTS VP [email protected]

U GEF/UNDP KAZ/00/G37-2010 N G J !!! ! \ F - !, ! W!, F!, ] - ! . T !! G, ! . H ( 2500 ): • U !! ; • U R1:200000; • H! ; • U «-G» ! T G !!! ! ( ), ! ! - . U- ( ) ! , , . V ! 1 2 «F» V . S ! 427 , 2 , 19 !, 284 , 39 . H SY. . 1 (33). 2012 231

T , - ", () . W . T , , - -, , , -, -, , , ! : , . V ! . M ! 427 241 68 . U : ! 33% ! U . \ ! : – 7; – 7; ! - 18; , ! – 6. V! – 80% , ! 31 . !! , : , , , , . T 284 53 17 !: I (1 ., 1 ); U (1 ., 5 ); H (2 ., 3 ); F (3 ., 9 ); I (1 ., 27 ); W (3 ., 29 ); (1 ., 3 ); L (3 ., 12 ); V (7 ., 46 ); I (2 ., 9 ); (1 ., 1 ); W (1 ., 7 ); (1 ., 1 ); W (1 ., 1 ); V (4 ., 5 ); J! (1 ., 2 ); H (20 ., 117 ). T ! ! 150 !! , 23 . ! 125, " – 6 . M , ! , !! ! 88 . , ! !, ! 5 : , , - , , ! . H ! ! 6 : ! ! , ! , !! , , ! . H- ! !! ! \ F ! , . U!! ! \ F, ! - . Y !! ! ! ! , , (43 ). U! ! W-K-T – ! . H V , 56 , 33 , 19 !! ! !!, 11 – 3 – . , " RWTU, ! 20 , 8 - !!. 39 , , 3 !! (! – Vormela peregusna, – Gazella subgutturosa, - Salpingotus pallidus). I ! . ! !! ! ! ! Rana( ridibunda) ! (Bufo pewzowi), ! ! (Rana asiatica), ! - G !, ! . J! ! ! ! R 1:200000 ( 1). W Q ! . H 3 : , - , . QIJ TR MR UTJ TR - U I c ! G - - - (1-8) U , ! - (9-15) U ! " (12-15) c ! - (16-17)

232 Y . . 1 (33). 2012

I c - ! (18-23) c - (22-23) c ! - (24-27) c (28-30) c ! (31) c

1. I «- » , - (32-34) c (35-40) c (41-42) H SY. . 1 (33). 2012 233

c ! (43-44) c -! , - - (45-48) F c -! - (49-51) c - (52-54) F c !- (55-57) c -- (58-60) c -! (, ) (61-65) Qb TR (66-67) MR TUTIT- TR (68-69) W – () ! ! . R 3 : -! , , J . H 3 : (), , . T , . I . 19 . T , , . W 12 , 3 , 4 . H 68 (- ) ! . S ! c " !! ! ! ! «-G». X! «-G», V TTUX [1], ! !, : - ! , , «J - F-c». U 2010 1:200 000. M X , L , , -F-Q ! . X! - X " !. T! ! 353,6 .. L! (! 1441,3.) L ". ! (! 3663,5 .) ! G . -F-Q! (! 2873,4 .) F-Q, , , . «-G» U , .. H ! ! ! G J!! , . . U ! F- c . H , ! ! - " , ! . Q 1. M V «T ! !» ! ! 10.12.2008 . W UFVFIVF, 24.02.2009. *** = ''' G ''' ' ''' '' ' GEF/UNDP KAZ/00/G37-2010 ' ' ''' . *** Assessment of biological diversity of ecosystems in the Ile river delta and the southern coast of Balkhash lake was carry out in the framework of GEF/UNDP KAZ/00/G37-2010 project for the goal of establishment of nature protected area 234 Y . . 1 (33). 2012

W 3. G

YJ 581.522.4.578.088.5 .. , .M. TTT TR]TO MQT O U TRUQTR TMJO TTT . -

J - TU "G". U RJ - . S Stipa capillata L. K ! ! , ! 165 , . T ! 45 ! ! [1]. H ! !, ! W ! . U ! ! ! . U - ! ! " - - - ". T ! ! !, ! , ! [2, 3]. X , !, ! , ! , , . , ! , ! W , ! , ! ! . T ! ! , , ! . X , ! ! ! ! , , !! ! - , , . \ !!! ! Stipa capillata L. ! W . RQ RTJ QJTO T ! - Stipa capillata L., !! ! WU - “G”. R ! ! - “G” ! - "T !! - ". G 2 ! 1080, 1050, 1328 !! (. 1). M! – - F ( W^ 49056/ 24//, HJ 79001/ 00//), ( W^ 490 53/ 49//, HJ 780 50/ 48//). ! ! ! . U (.1) RcJ ! - F (h0) - 2,58 M/, 1 (h1) - 1,69 M/ !! - 300 / ×2. X 1 U Stipa capillata L. R I F G I h=0 I h=1 / × 2 M/ 0 / // 0 / // - 49 56 13 79 00 19 < 1 775 18,5 26,4 F H SY. . 1 (33). 2012 235

- 49 56 24 79 01 00 < 1 300 2,58 1,69 F 1080 50 00 07 78 51 58 < 1 15 0,72 1,05 1050 49 56 13 78 56 06 < 1 < 10 0,12 0,12 1301 (!.) 50 01 48 78 55 30 < 1 200 1,05 0,76 1301 () 50 01 50 78 55 38 < 1 12 0,18 0,21 1328 49 56 11 78 49 37 < 1 15 0,13 0,12 49 53 49 78 50 48 < 1 < 10 0,13 0,11 J! ! - ! !, . I- ! S\ V (.). J! Stipa capillata L. - . 70% , W- (, , , 1:1:1). F ! U R.Q. (1960), U! F. (1988). W! Q I.. (1990). R RG-6 ( 63). V ! W !, Stipa capillata L. ! TcU "G". Stipa capillata L. – ! Poaceae L., , !, !, . Stipa capillata L. H ! Stipa capillata L. , ! ! . W . U – ! ! ! , . R !! . U! , , !. U! , . W , ! . c ! ! (8,76 , 5,83 ). X ! (38,84 , 33,06 ). T !!! , ! ! ! (.2). X 2 T Stipa capillata L. . 3 Y X X J U , ! ! , !, , 10-3 2 1 - 8,11 ± 0,125 38,01 ± 0,85 27,6 ± 1,18 16,01 ± 1,23 0,68 ± 0,01 F 2 - 8,76 ± 0,87 38,84 ± 2,17 26 ± 1,36 14,6 ± 1,01 0,63 ± 0,05 F 3 1080 7,25 ± 1,45 36,65 ± 2,15 26,5 ± 0,85 15,56 ± 2,05 0,69 ± 0,022 4 1050 4,67 ± 0,199 37,13 ± 0,62 30,5± 0,62 11,40 ± 0,67 0,57 ± 0,08 5 1301 (!.) 7,16 ± 0,33 38,88 ± 2,29 28,9 ± 1,78 14,7 ± 0,38 0,69 ± 0,001 6 1301 () 5,05 ± 0,215 34,56 ± 1,371 37,3 ± 1,73 11,25 ± 0,45 0,54 ± 0,034 7 1328 5,61 ± 0,23 32,62 ± 0,773 36 ± 0,68 12,83 ± 0,91 0,58 ± 0,04 8 5,83 ± 0,48 33, 06 ± 3,05 32,4 ± 0,86 23,5 ± 1,08 0,52 ± 0,02 Stipa capillata L. Q! Stipa capillata L. , - . c . . S! . R , , , . U! ! . T , 1- 2 ! , , !! . S! ! , ! . 236 Y . . 1 (33). 2012

X 3 T Stipa capillata L. 3 Y X , X U ! , , !! !! 10-3 2 1 - 7,052 ± 0,214 6,55 ± 0,187 56,95 ± 1,284 21,64 ± 1,001 F 2 - 6,9 ± 1,4 7,4 ± 1,58 56,71 ± 1,52 22,34 ± 0,028 F 3 1080 5,98 ± 1,87 6,79 ± 2,54 47,58 ± 0,98 24,54 ± 1,01 4 1050 4,61± 0,19 4,24 ± 0,35 41,75 ± 0,62 17,03 ± 0,721 5 1301 (!.) 8,85 ± 0,265 7,29 ± 0,265 47,664 ± 0,212 23,82 ± 2,025 6 1301 () 3,43 ± 0,212 5,234 ± 0,31 53,94 ± 2,673 18,06 ± 3,152 7 1328 4,05 ± 0,15 4,61 ± 0,233 48,63 ± 2,204 17,61 ± 1,654 8 4,75 ± 3,5 4,87 ± 2,07 41,04 ± 1,23 18,20 ± 0,022 U! . W , ! ! ! . X ! ! 6,9±1,4 7,4±1,58 , – 2 - 4,75±3,5 4,87±2,07 (.3). Y! (56,71±1,52 – !, 41,04±1,23 - ) ! ! (21,64x10-3±1,001 18,20x10-3±0,022 ). Stipa capillata L. Stipa capillata ! . S ! : , . c . c ! . T ! 3-4 ! . I ! ! ! , ! ! , . U , ! . , !. \ ! . U!! ! - . ! ! . T ! !!! , , (.4). X 4 T Stipa capillata L. 3 Y X X ! X J , , , , 1 - 242,69 ± 13,87 39,22 ± 0,386 12,09 ± 0,82 27,98 ± 0,79 F 2 - 239,4 ± 2,72 40,26 ± 1,27 12, 57 ± 1,08 25, 79 ± 1,19 F 3 1080 232,15 ± 0,65 39,54 ± 4,57 11,87 ± 2,54 27,68 ± 0,54 4 1050 232,59 ± 5,95 30,53 ± 0,39 12,46 ± 0,46 25,79 ± 0,9 5 1301 (!.) 244,55 ± 3,32 29,19 ± 0,84 11,4±0,54 22,056 ± 1,124 6 1301 () 228,22 ± 11,87 25,11 ± 1,39 10,01 ± 1,09 22,59 ± 0,831 7 1328 229,53 ± 7,48 22,96 ± 0,834 9, 097 ± 0,48 24,61 ± 0,42 2 221,7 ± 3,07 22,4 ± 1,3 10,73 ± 1,2 22, 36 ± 0,99 S ! ! , .. . H 5, 6, 7 , ! ! . W ! .

H SY. . 1 (33). 2012 237

X 5 Stipa capillata L. TU «». Y W - , G/ 40 232Th 238U 241Am 137Cs 60Co 152Eu 154Eu - 436±35 34±6 28±4 1702±85 14359±574 5745±230 9684±387 7500±300 F - 210±41 20±4 30±5 210±25 3890±272 979±39 1570±94 640±64 F 1080 340±42 16±4 22±4 1426±57 1050 499±35 22±3 24±3 14±2 1328 522±47 24±4 36±5 19±2 548±50 21±3 19±2 24±4 F , ! !! (- - F ) ! 238U, 137Cs, 232Th . U 241Am, 60Co, 152Eu, 154Eu , ! ! . Y! 40 Stipa capillata L. ! !, ! ! ! . H - F ! ! 241Am (1702±85 G/), - ! 210±25 G/. F !. T ! - 69±21G/. S! , ! . H ! ! - 137Cs. X 6 Stipa capillata L. TU «» Y W - , G/ 40 232Th 238U 241Am 137Cs 60Co 152Eu 154Eu - F 192±45 3±1 4 ±2 74 ± 9 c- F 148±27 < 2 3 ± 1 64 ± 7 ±±±± 1080 149±61 < 2 3 ± 1 100 ± 8 1050 173±52 < 2 3 ± 1 4 ± 1 1328 184±44 < 2 3 ± 1 < 3 141±33 < 2 3 ± 1 < 3 U! 137Cs (1860 G/ 18 G/ - ), !! 40, 232Th, 238U, 137Cs, 241Am, 60Co, 152Eu, 154Eu. X 7 Stipa capillata L. TU «» Y W - , G/ 40 232Th 238U 241Am 137Cs 60Co 152Eu 154Eu - 310 ± 78 7 ± 3 10 ±3 69 ± 21 1860 ± 130 311 ± 28 408 ± 33 170±24 F c- 120 ± 55 4 ± 2 8 ± 4 37 ± 13 1350 ± 108 154 ± 14 253 ± 50 63 ± 12 F 1080 100 ± 32 8 ± 2 10 ± 2 1620 ± 113 1050 150 ± 58 3 ± 1 4 ± 2 < 3 1328 250 ± 58 < 2 < 2 < 3 110 ± 25 6 ± 2 5 ± 2 18 ± 4 X , . T ! , ! . U RcJ -! , !, ! ; ! ! , , ! . H ! , . 238 Y . . 1 (33). 2012

W, Stipa capillata L. RcJ ! ! . Q 1. X W.X. U . F, I, 1997, 745 . 2. V F.H. V! !: Y..-R.:- RIY, 1990. - 88 . 3. I S.., U! I.Z., J F.R., I J.. // V. !. V !. 2002.X.42.H.2.W.155-157. 4. Q Q.., X F.. W! ! : Y.-..-R.:- R.-, 1989. - 80 . 5. c . F! . R.:R,1980.X.1.2. - 558. 6. Q I.. G!.-R.: H! , 1990. - 352. *** P ' ''. G “G” %'' %' % Stipa capillata L. ''''' '' % ' ' ''. *** The anatomical characteristic of Stipa capillata L. the dominant kind of plants of territory of semipalatinsk test site is given. Af increase of PED there is an increase of quantitative parameters in structure of overground and underground organs. The greatest accumulation of radioactive elements is revealed in near root soil and underground organs.

YJ 634.0.30 R.. , .R. , .. T T JTO QT T T Q-QTIT IUU (-F )

, , P P (Malus sieversii (Ledeb.) M. Roem.) o (Armeniaca vulgaris Lam.) N-F . U , -, U . U . -F U U V 22 ! 1996 3228. U . F ! – 199703 . , ! F 542400 . H 2001-2003 . -F F ! , ! . H M ! ! FISUU : - – 57786,0 ; - – 23280,0 ; - ! – 14991,0 ; - ! ! – 103646,0 . X! 4 , 12 132 , : -F – 56992 – 29 1965 ; -R – 41922 – 35 1 – 1998 ; -X – 27938 – 30 1 – 931 ; -X – 72852 – 31 , – 2512 . T ! 178 ( !). I ! -F - !. T 75207 , : ! 62262 , – 10969 . X , ! ! 37,7%. RQ RTJ QJTO ! «R ! Q ! !», ! RW V 18 ! 2007 ., !! ! ! ! [1]. MQb TLJ H S ! V , U V H SY. . 1 (33). 2012 239

1999 ., ! , , K..V, Q.., S.U.T ..V [2] , ! ! : 1) Picea schrenkiana ( ] , !! ! ! ); 2) – - (Celtis caucasica Wild.), . R! F; 3) ! , !! W (Malus sieversii (Ledeb.) M.Roem.), , [3]; 4) (Armeniaca vulgaris Lam.), . J , – !! – !! ! ! , , !!! , . Y . J !, . S ! , , ! . c -F ! . H -! (! (Crataegus) – , , ; , , , – R ), !. T !!! R (Atraphaxis muschketovii rasn.) - I, ( X) -F F . , ! . ! W ! – , R, W, X, , G R F, F, , X 800 1800 ! [4]. S ! G, F.J.J 60- - « », 200 , X-G R X . S !! , , , , ! !. U! ! (30-120 ), – !, -!, -!, -!, – - ! , – - -, !. H ! ! , , - , . S F.J.J, V.R.X, X.S.W 1990 20 !- !, ! V [5]. W ! ! , , , !. W ! ! - , , ! (0,066%), ! . X ! ! . G , , ! , . 1%, – 0,46%. H ! , ! ! ! . H , , !! ! ! ! . W 740 % , V, 7,4 , . H V- W- - !. , , ! , , ! !. U – - , ! , . H 2010 . -F . T « » ! RW V 16 !! 2010 . 240 Y . . 1 (33). 2012

H ! 1350 – 1550 . ! ! ( , W F). S ! ! 158 . U ! ! !! ! (Crataegus songorica, Armeniaca vulgaris), (Euonymus semenovii, Lonicera tatarica, Berberis sphaerocarpa, Rhamnus cathartica, Rubus caesius, R. idaeus) 20 – ! (Aconitum leucostomum, Agrimonia asiatica, Geum urbanum, Lavatera thuringiaca, Hypericum hirsutum, Bupleurum aureum, Phlomis pratensis, Prunella vulgaris, Origanum vulgare, Inula helenium, Brachypodium sylvaticum, Dactylis glomerata, Agrostis gigantea, Polygonum coriarium, Humulus lupulus, Melandrium viscosum, Thalictrum flavum, Viola suavis). ] (!, , , , , , !, , Ligularia macrophylla, Trifolium pratense, Geranium prtense, G. robertianum) ! , !!! [6]. ! ( ) (Galium aparine, Arctium leiospermum, Artemisia absinthium, Urtica dioica), ! . F ! ! G R F, X, F!, , , X! 800-1600 . ! !, . T! ! 1100-1300 . U!! ! [7]. S, . X ( ) 2009 . 2000 . K ! 1650 . !. K 80 M F . , ! X . S F.J.J, X.S.W 20 , 6 -. T «» ! RW V 16.09.2010 . R! . ( 45 ) - - (1500 . !). V – 50 50 . T -. H 3 – , !! W, ! (Crataegus songorica C. Koch) 4 – ! (Spiraea lasiocarpa Kar. et Kir.), (Rosa platyacantha Schrenk), ! (Lonicera tatarica L.), (Rhamnus cathartica L.). W - ! 30%. ! 55 22 , !! ! Asteraceae, Poaceae, Rosaceae, Fabaceae, Brassicaceae, Apiaceae. J (Artemisia santolinifolia (Turcz. ex Pamp.) Krasch., A. dracunculus L.), (Glycyrrhiza uralensis Fisch.), (Galium verum L.), – Melica transsilvanica Schur Bromus oxyodon Schrenk, Phleum paniculatum Huds. W 2008 . ! , , ! V.R.X F.F.. U!! ! ! !, , ! ! - ! . Y ! !! ! , ! !. W ! , ! ! ! «R ! FISUU», – RW V, S.L.F. G ! ! , . M ! 7 , : X – 3, F – 2, R X – 1. T! ! 6,5 . J! ! 25 2265,0 , : ! W – 1409,0 , – 826,0 , ! – 30,0 . S! 2008 -F ISUU : !! – 81,1 , – 115,2 . W! ! ( 50% ). H ! ! , F ( W.W.G). Q 1. R ! Q ! ! . Y ! R ! V 18 ! 2007 3 156. – 46 . 2. S! ! ! V .- .-1999.-W.91-93. H SY. . 1 (33). 2012 241

3. ! WWV. ].2. V!. F-F.-1981.-260 . 4. X V.R., F.F. -F // G . R. R. -. F, 2009.-W.109-111. 5. U! . T . F, 2006. 3 7. W.123-137. 6. F.F., U.H., W H.Z. T -F // R R. S-. . «F ! ! \ F». F.-2008.-W.229-231. 7. X V.R., F.F. F -F – // R H . ( ) « ». H. 6.-.-2010.- W.92-94. *** G = F ' ' '' (Malus sieversii (Ledeb.) M. Roem.) %'' '' (Armeniaca vulgaris Lam.) ' % ''' ' . R '' % -, U '' ' %' '. R' '' %' ' % ''' '' . *** Results of research by distribution, ecology and condition of apple tree and apricot woods with dominated rare species, which listed in Red Data Book of Kazakstan - Malus sieversii (Ledeb.) M. Roem. and Armeniaca vulgaris Lam. on the territory of the Ile-Alatau national natural. Park have been generalized in this article. The authors give information about the rich distribution of forms of mentioned species, about selected and patented variety-clones, about the genetic reservations of these breeds, which have been sorted out at the territory of Park. The authors emphasize the need for special protection of unique plant communities and bring a summary of the researches on the monitoring grounds.

YJ 633.81:575.1 .. , .. , .. , R. JTTO T UT JT R. IT\ T ZQT \QbTIT M K.F.G e-mail: [email protected]

I \ P. S 50 , 20 , 22 % P 42,6 % . U 2 : Thymus L. Scutellaria L. V Thymus L. \ P 12 : T. asiaticus Serg., T. crebrifolius Klok., T. eremita Klok., T. guberlinensis Iljin, T. kasakstanicus Klok., T. kirgisorum Dub., T. lavrenkoanus Klok., T. marschallianus Willd., T. minussinensis Serg., T. rasitatus Klok., T. roseus Schipz., T. stepposus Klok.et Schost.] . I \ P . 2 10 \ P: , G, P, M, R, I, I, G, M, S. F . Lamiaceae \ P 7 . , . T ! !!! . H ! I (Lamiaceae L.), !! ! . X, ! 233 , 45 [1-3]. S! , ! . . S , ! , ! . H , , ! , ! ! \ . U ! ( , , , W , . . , . .) !! , ! , . ! 2010-2011 . J! ! ! Lamiaceae ! . ! \U ! [4]. U ! ! ! [5, 6]. 242 Y . . 1 (33). 2012

W Lamiaceae \ 50 , !! 20 ( 1), ! 22,0 % 42,6 % . X 1 . I \ ( ) 3 V T T % \, . , . 1 Thymus L. 12 22 54,6 2 Scutellaria L. 5 35 14,3 3 Dragocephalum L. 4 20 20,0 4 Nepeta L. 5 14 35,7 5 Mentha L. 3 8 37,5 6 Phlomis L. 3 4 75,0 7 Galeopsis L. 2 3 66,7 8 Hyssopus L. 2 4 50,0 9 Lycopus L. 2 2 100,0 10 Salvia L. 2 8 25,0 11 Ziziphora L. 2 7 28,6 12 Glechoma L. 1 1 100,0 13 Lagochillus Bunge 1 16 6,3 14 Lallemantia Fisch. et Mey 1 1 100,0 15 Lamium L. 1 3 33,3 16 Leonurus L. 1 4 25,0 17 Lophanthus Adans. 1 3 33,3 18 Prunella L. 1 1 100,0 19 Sideritis L. 1 1 100,0 20 Stachys L. 1 5 20,0 U 2 : Thymus L. Scutellaria L. X, Thymus L. \ 12 : T. asiaticus Serg., T. crebrifolius Klok., T. eremita Klok., T. guberlinensis Iljin, T. kasakstanicus Klok., T. kirgisorum Dub., T. lavrenkoanus Klok., T. marschallianus Willd., T. minussinensis Serg., T. rasitatus Klok., T. roseus Schipz., T. stepposus Klok.et Schost. (. 1).

, 6 . 4

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V . I: 1 – !, 2 – , 3 – , 4 - , 5 – !, 6 – , 7 – , 8 – , 9 - , 10 – , 11 - , 12 – , 13 – , 14 – !, 15 – !, 16 - , 17 – , 18 – , 19 – , 20 - 1. ] . I \ V Scutellaria L. 5 : S. dubia Tab. et Schir., S. galericata L., S. grandiflora Sims., S. scordiifolia Fisch.ex Schrank., S. supina L. G! ! ! : - Dragocephalum L. – 4 (D. nutans L., D. peregrinum L., D. ruyschiana L., D. thymiflorum L.); - Nepeta L. – 4 (N. cataria L., N. micrantha Bunge, N. pannonica L., N. ucranica L.); H SY. . 1 (33). 2012 243

- Mentha L. - 3 (M. arvensis L., M. longifolia L., M. micrantha L.); - Phlomis L. – 3 (Ph. agraria Bunge, Ph. puberula Kryl. et Serg., Ph. tuberosa L.); - Galeopsis L. - 2 (G. bifida Boenn., G. ladanum L.); - Hyssopus L. – 2 (H. ambiguus (Trautv.) Iljin, H. macranthus Boriss.); - Lycopus L. – 2 (L. europaeus L., L. exaltatus L.); - Salvia L. – 2 (S. deserta Schang., S. stepposa Schost.); - Ziziphora L. - 2 (Z. bungeana Juz., Z. clinopodioides Lam.). T 1 : -Glechoma L. (G. hederacea L.); -Lagochillus Bunge (L. acutilobus (Ledeb.) Fisch. et Mey); -Lallemantia Fisch. et Mey (L. royleana Benth.); -Lamium L. (L amplexicaule L.); -Leonurus L. (L. glaucescens Bunge); -Lophanthus Adans. (L. schrenkii Levin.); -Prunella L. (P. vulgaris L.); -Sideritis L. (S. montana L.); -Stachys L. (S. palustris L.). U I ! 10 \ [87], ! 7 ( 2). U 4,0 % 4,1 % . X 2 10 \ 3 W ] , % ] , % . . 1 W 50 10,3 187 14,9 2 G 22 4,6 113 9,0 3 44 9,1 82 6,5 4 M 30 6,2 81 6,5 5 R 24 4,9 74 5,9 6 I 15 3,1 57 4,5 7 I 20 4,1 50 4,0 8 G 20 4,1 49 3,9 9 M 30 6,2 47 3,7 10 S 10 2,1 47 3,7 W 50 7 (Hyssopus ambiguus, Lagochillus acutilobus, Thymus lavrenkoanus, Th. crebrifolius, Th. rasitatus, Th. kasakstanicus, Th. eremita), ! 14 % . U ! (. 2-8).

1 - Salvia stepposa, 2 - Scutellaria grandiflora, 3 - S.scordiifolia, 4 - Thymus marschallianus, 5 - T.lavrenkoanus, 6 - T.minussinesis, 7 - T.crebrifolius, 8 - T.rasitatus, 9 – T.stepposus 2. . Lamiaceae \

244 Y . . 1 (33). 2012

1 - Thymus kasakstanicus, 2 – T.eremita, 3 - Ziziphora bungeana, 4 - Z. clinopodioides, 5 – Nepeta ucranica, 6 – Galeopsis bifida 3. . Lamiaceae \

1 - Hyssopus ambiguus, 2 - H.macranthus, 3 - Lagochillus acutilobus, 4 - Lamium amplexicaule, 5 - Leonurus glaucescens, 6 - Lophanthus schrenkii, 7 – Mentha micrantha, 7 - Nepeta cataria, 9 - Prunella vulgaris 4. . Lamiaceae \

1 – Phlomis argaris, 2 - Sideritis montana, 3 – Nepeta micrantha, 4 – Salvia deserta, 5 – Stachys palustris, 6 - Dragocephalum nutans, 7 - D.peregrinum, 8 - D.ruyschiana, 9 - Glechoma hederacea 5. . Lamiaceae \ H SY. . 1 (33). 2012 245

1 – Thymus roseus, 2 – Lallemantia royleana, 3 – Dragocephalum thymiflorum, 4 - Eryngium planum, 5 – Nepeta pannonica, 6 – Thymus asiaticus 6. . Lamiaceae \

1 – Scutellaria dubia, 2 – Thymus kirgisorum, 3 – T.guberlinicus, 4 – Scutellaria galericata, 5 – Mentha arvensis, 6 – Scutellaria supina 7. . Lamiaceae \

1 – Phlomis puberula, 2 – Ph.tuberosa, 3 – Galeopsis ladanum, 4 – Mentha longifolia, 5 – Lycopus exaltatus, 6 – L.europaeus 8. . Lamiaceae \ 246 Y . . 1 (33). 2012

X, ! , - (, T G-L ), – -. c !, !! , , . H , (F, L, Y) ! ; , , , , !. Q 1. . X. 7. – F-F: - FS WWV, 1964. – 495 . 2. U S.H. \ . ].2.-R.-Q.: - FS WWWV, 1935.-546 . 3. F W.F. W . – F, 1999. – 215 . 4. G G.F. H . - F-F: S, 1970. - 226 . 5. W! .I. c! ! . – R.: H! , 1962.-378. 6. G G.F. c . – F-F: S, 1983. – 216 . *** R T ' '' ' '. '' 20 50 ' 22 % % 42,6 %- . X' ''' 2 : Thymus L. % Scutellaria L. Thymus L. T 12 ' ': T. asiaticus Serg., T. crebrifolius Klok., T. eremita Klok., T. guberlinensis Iljin, T. kasakstanicus Klok., T. kirgisorum Dub., T. lavrenkoanus Klok., T. marschallianus Willd., T. minussinensis Serg., T. rasitatus Klok., T. roseus Schipz., T. stepposus Klok.et Schost. T ' '' ' ' '. 2 T ' 10 ' : P'', G', P', F , F, ', K'', '', ^', ''. F T Lamiaceae ' '''' ' ' % 7 . R % ' % ''' . *** This article contains the tables which offer a structure of the family of taxonomic Labiatae of Central Kazakhstan. At the present stage the family is represented by 50 species belonging to 20 types, which compose 22 % of the total number of species of Kazakhstan’s flora and 42.6 % of total births. The two types take a central place in species diversity: Thymus L. Scutellaria L. The sort of Thymus L. Represented by 12 species in Central Kazakhstan: T. asiaticus Serg., T. crebrifolius Klok., T. eremita Klok., T. guberlinensis Iljin, T. kasakstanicus Klok., T. kirgisorum Dub., T. lavrenkoanus Klok., T. marschallianus Willd., T. minussinensis Serg., T. rasitatus Klok., T. roseus Schipz., T. stepposus Klok.et Schost. A distribution of Labiatae flora species is offered as a chart.10 leading families of the flora of Central Kazakhstan is described in the table 2: Asteraceae, Fabaceae, Cruciferae, Cereals, Chenopodiaceae, Caryophyllaceae, Labiatae, Boraginaceae, Umbrella, Fig-wort. The authors of this article carry out a research some of the spreading plants of Labiatae family in Central Kazakhstan and 7 maps of the regions are also worked out by them. At the end of the article there is a conclusion and references which were used in writing this article.

YJ 581.8 .. , .R. R, .. , .. , . J TTT TR]TO I TIT JTIT JR]TIT J LIMONIUM MICHELSONII LINCZ. ]Qb J TTIM -, F,

Limonium michelsonii . J Limonium michelsonii M F ( G, X G). Limonium michelsonii, . , Limonium michelsonii , . H , , ! , ! ! , ! ! . K 303 , ! 450 ! ! [1]. T !!! Limonium michelsonii Lincz. ( R), , ! , ! ! ! - M F (, W , X) [2,3,4]. – !! ! !. U ! ! , , !! , .. ! ! , , ! . U !! ! !, . U ! ! Limonium michelsonii !, ! , ! . RQ RTJ QJTO S ! Limonium michelsonii, ! M F. ! Limnium michelsonii - H SY. . 1 (33). 2012 247

GPS-. I ! [5]. U ! !, ! X.F.V [6], .I.W! [7], F.F.Y [8]. H - ! ! Limonium michelsonii ! . H ! ! !! . ! W-. F ! U R.S. (1960) [9], U! F. (1988) [10], G V.U. (2004) [11]. J! !- RTH-1-15 ( 9, 10,7). R MC 300 CAM V400/1.3M. W! Q I.. (1990) [12] Y S.Q. (1976) [13], Microsoft Office Excel 2003. MQb TLJ U! !! Limonium michelsonii ! F R G. Y ! 1283 – 1294 .!, GPS : N 43027.651' E 078047.688'. V ! . U – - !, !, . Y . H! !! Limnium michelsonii G , H X!-^!: M F, G X. Y ! 1108-1141 .!, GPS : N 43020.723' E 078031.593'. V ! - , . W . U - !, (30 80 ) (-). U . X! !! Limnium michelsonii ! X F- S. Y ! 1301-1320 .!, GPS : N 43020.630' E 078047.864'. H ! ! ! (!) () Limonium michelsonii, ! , !!. Limonium michelsonii ( ) Q! Limonium michelsonii – , ! !, . c ! : . S . X . R – . R () , . U - , . X – , . I . U! , , ! ! . H ! , !! ! . H! ! . R! ! ( 1).

F G H . – , . – , . – , . – , – , . - ! ; F – !! 1; G – !! 2; H – !! 3 1 - Limonium michelsonii ( ) U Limonium michelsonii ! !, ! 3 1 248 Y . . 1 (33). 2012

!. V! ! 2 3 . Z ! 1 ! ! 2 3, , ! . Y ! 2 . S! ! 3 ! , ! ! ! ( 1).

X 1 R Limonium michelsonii ( ) X X , X , J U!! ! , , 1 166,65 15,13 12,19 78,82±3,75 54,84±3,67 41,86±2,32 ±6,03 ±1,83 ±1,73 2 123,86 9,81 12,27 51,35±4,92 52,80±2,23 43,92±4,75 ±4,97 ±0,67 ±0,66 3 120,56 9,61 10,60 52,27±2,34 42,59±2,94 35,63±0,53 ±5,57 ±0,12 ±1,06 U ! !! ! , ! , - , !, !. Limonium michelsonii ( ) Limonium michelsonii – , ! . U , ! , ! . !. H ! , ! ! ! – ( 2).

– , . - ! , - F – !! 1; G – !! 2; H – !! 3 2. Limonium michelsonii ( ) ! 1 ! ! ! ! 2 3. J ! 3. V! ! 2 , ! . W , ! ! . S! , . S ! !! ! , !! ! ! ( 2). X 2 R Limonium michelsonii ( ) U!! X , X , J , 1 31,44±2,27 44,47±5,48 105,99±8,25 2 23,42±0,93 20,05±0,87 73,45±4,89 3 22,28±3,82 35,29±1,36 56,09±1,27 H SY. . 1 (33). 2012 249

Limonium michelsonii ( ) c Limonium michelsonii ! !, , . S , ! . H! ! , ! !! ( ) , . U , . ! ! , ! . U – , .. R ! , ! ! . U! , ! ( 3).

F G H . – , . – , . – , . – , – , . - ! F – !! 1; G – !! 2; H – !! 3 3. Limonium michelsonii ( )

V!, ! 2, ! !. Y , , ! . V! ! !, ( 3). X 3 R Limonium michelsonii ( ) X X , X , J U!! ! , , 1 181,08 12,29 12,73 78,12±7,69 74,93±3,41 63,79±2,69 ±4,83 ±0,99 ±1,43 2 229,00 15,12 14,5 102,91±4,77 96,57±6,60 83,07±3,31 ±1,12 ±0,29 ±0,87 3 172,83 12,28 15,38 77,94±2,35 84,89±4,45 74,14±1,83 ±3,82 ±0,56 ±2,33

Limonium michelsonii ( ) Limonium michelsonii – , ! . U , ! . W ! , , . R ! - ! . H ! ( 4). 250 Y . . 1 (33). 2012

F G H – , . - ! , - F – !! 1; G – !! 2; H – !! 3 4. Limonium michelsonii ( ): H , ! ! 2 !, ! . V , ! ! ! , , !!! ! ! . V! ! 3 , , ! ! , ! ! ( 4). X 4 R Limonium michelsonii ( ) X , X J J U!! , , , 1 28,85±2,81 93,86±3,49 203,59±1,46 10,68±1,06 2 29,46±1,69 104,87±4,15 232,10±4,13 10,86±0,48 3 34,37±3,36 96,85±4,45 180,67±1,82 8,75±0,14 U Limonium michelsonii !, !! . U ! , ! . W, ! ! , !! ! !. H ! . U ! . V ! . X! ! ! !. S : 1. Limonium michelsonii !! . 2. H ! , ! , ! . X – . 3. F ! ! , ! ! . Y . 4. J! , ! , ! ! , ! , !! ! . 5. Y ! 1 ( , , ! ) ! ! ! 2 3. 6. V! ! 2 !! !, ! , . Y ! H SY. . 1 (33). 2012 251

! !! ! , ! ! . 7. ! 3 ! ! , . X , ! Limonium michelsonii ! ! , ! , ! . Q 1. http://fsbk.kz/node/24 2. ! WWV. ] 2. V!. - F-F.- 1981. -284 . 3. G R.W. H . - F-F. - 1985. – 176 . 4. F.F. \ ! - : . -F: F! ! ! . - 2008.-184 . 5. U! . -R. - FS WWWV. -1964. - .3.- 530 . 6. V X.F. L ! // X.GS FS WWWV. R.-Q.: FS WWWV. I.- 1950.-.3. -. 6. - W.7-204 7. W! .I. R! . - R.-1952. - 240 . 8. Y F.F. T ! // T ! .- R. -1967. - W. 1-12. 9. U R.S. G! . - R. - 1960. – 208 . 10. U! F.. R. – R.: . RIY. - 1988. – 58 . 11. G V.U. . W . T . - R.: - RIY. - 2004. – 312 . 12. Q I.. G!. - R.: H! . - 1990. - 352 . 13. Y! S.Q. H . - F-F: - «S» WWV. -1976. - 83 . *** R ' % ' Limonium michelsonii-' '' '' ' '' ' %' '. T ' Limonium michelsonii-' = F ''' (G'', X % G ) '. $ Limonium michelsonii- ' ' , '' , ' ' . Limonium michelsonii-' %' ' % ''' '' '' ' , ' ' '' . *** Results of a study of the anatomical structure of vegetative organs of rare and endemic species Limonium michelsonii in the early stages of ontogeny are presented in this article. Three populations of Limonium michelsonii in eastern part of Trans-Ili Alatau (mountain Bogutty and Toraigyr, Bartogay hole) were investigated for this. Age states of plants of Limonium michelsonii have been installed, the patterns of them were collected and fixated in each of the studied population for further study its anatomical structure. The internal structure of leaves and roots of individuals of Limonium michelsonii from different populations in the early stages of ontogeny has a great similarity was found and the specific differences in the anatomical structure of vegetative organs has not been established.

TL 631.46.467 .ļ. JĽ'1, Z.. 2, I.. 2 «ľLT» ZTZT Tİ TIJ=-ĲQ=I QJZJ TUĲM=Q ĲJ==JI= TU TRMJİ ľQ= - 1 ľ.T. T Ľ ! - [email protected], [email protected]

M ' ' '' % ' ''. '' '' ' Nothrus, Scheloribates ''' ' '. G ' . '' '' ''' Anurida, Folsomia ' ' '' ''. &'' '' ' '' '. L % '''' '' , ''' '' '', - , ' '' '' %', ' ' '. Xı Ľ Ŀı ĳ ĳ , ı L ĳ ı ĳ. Kı Ŀ ! , ı ı ĳ . G!ı ı ı , ı ĳ ĳı 70%- . K Ľ ı Ľ ı ı . G 252 Y . . 1 (33). 2012

ı Ľ ı , ! ĳı -ı [1]. Xı ĳ ı Ŀ ı . Xı ı Ľ XIX ı 70-ı ! 80- ı . F H. [2] Ŀ ı (ı) ı Ŀ Ŀ. ^ıı Ľ ı ı ı ĳ Ŀ . T ĳ ı Ľ H. I (Hansen, 1877) Ľ ]. J (1882) !. F ]. J [3] ı Lı ı ı ı ı . T H.H. J [4] Ŀı «V » ı, ı Ŀ Ŀ Ŀ Ŀ , ı , ı ı Ŀ . Jı U.F. [5], ı Ľ ĳ , ı ı Ŀ . X ĳ ĳ ! ı Ŀ Ľ Ľ ! ı . W- Ľ ı Ŀı . F, ĳ ĳ ı Ŀ Ľ Ľ , Ľ Ŀ ı . X Ľ ! ı, Ľ Ľ ĳ ! Ŀ [6]. Xı ı [1]. X ı 80%- . ľ Ŀ Ľ Ŀ , ı ĳ Ľ . T ı ı ĳ . W-, ı ĳ Ŀ . W Ŀ ı . T Ŀ Ŀı , - ı . X Ŀ - ! . Xı Ľ ı Ŀ Ŀ , ı Ľ Ŀ ĳ . Wı Ŀı ĳ [7]. T ı Ŀ ĳ Ľ ı Ľı ı Ŀ . X ı Ľ . Ŀ ı ĳ Ŀ ı Ŀ . ^ıı, ı Ľ ı , - - , ı Ŀı ĳ, Ľ Ŀı ı ĳ ı ı , Ŀ ĳ ! ı Ŀ ĳ [8]. ľ ı Ľ ı ı Ŀ ı , ĳ . Rı ı ! ĳı ĳ ĳ ı . F ı Ŀ ı !, ı ! ĳ. L – ı . L Ľ ı ı Ŀ ı ı Ľ ı ı ı . Ĵ ! ĳ, Ŀı ! ĳ [9, 10]. Ľ ! ı - ! Ľ ı Ŀ. ı Ŀ ĳ ! , ! Ľ Ŀ ı , ı . ľĽ ! ĳ Ŀ ı 90- , ı ı Ŀ Ľ . F, ĿĽ ı Ľ ĳı !!ı !, ! ĳ . c!ı Ľ ĳ Ľ ĳı ĳ Ŀ ĳı ! H SY. . 1 (33). 2012 253

ı . c! Ľ ı ĳ ĳı ı ĳ Ľ [11]. U Ŀ Ľ « » ĳ. R ı Ľ ! , ! . ľ , Ľ ı . X ı Ľ : 1 – ! ! ĳ ĳ Ľ Ľ; 2 – ĿĽ ı Ľ - . Lı : -ĳ ĳ ĳ ı ı , ĳ ĳ ĳ Ŀ, ı Ľ . M Ľ''. M Ŀ ı ĿĽ ĳ ĳ. Ĳı Ŀ - 50 . G ĳ Ŀ ĳ. Ĳ ĳ . X ĳ ĳ ĳ Ŀ ! ĳ. Ĳ Ľ Ŀ. T ı . R , ĳ «T K WWWV» [12]. F G – Mı «U » [13]. ı 0-5 ,5-10 ĳ , . X ĳ Ŀı Ľ ĳ. F ĳ ĳ G – X Ľ . R ĳ 0,25 ĳ Ŀ Ľ . R Ŀı Ľ 70°W , , Ľ . ĩ Ľ Ľ . «Ŀ» – Ŀ , Ŀ . «Ŀ» 2 . M Lı ıĳ- ^ Ľ Ĳ F ı ı 500-700 . Ŀ Ľ ı Ŀ (3-4 ), ı, ı Ľ ı . Ĳı , ı Ľ ĳ (2-7 ) ı . Ĳı Ŀ ı, Ŀ 3-4 ı Ŀ Ŀ 7 . ľ Ŀ Ŀ ĳı Ŀ ı Ŀ. X - ıı ı ! Ŀ . «Ŀ» ı Ŀ Ŀ Ŀ ĳ ĳ . ľ - Ŀ ĳ ĳ ! ĳ. Ŀ ĳ - Ŀ (, ) . ľ Ľ ĳ ı Ŀ ı . Iı Ŀ Ľ ĳ ĳ ĳı ı Ľ. L ı ĳ Ŀ Ŀ , ı Ŀ ı Ľ, ı ı . ı ! Ľ ĳı ! ĳ Ŀ. Ŀ Ľ 38 ĳ . R ı Isotoma, Entomobrya, Folsomia Ľ ı (Oribatea) ĳ : Oribatula, Oppia, Nothrus . R Ŀ Ŀ . S ı Carabidae – ı Ŀ Elateridae – ı ı Ŀ . X ĳ Ľ Ŀ : 1, 1 ı ( ) Ľ 13, 2 ĳ 8 ıı . F ı ĳ Isotoma, Entomobrya ı Ŀ Oribatula Ľ Nothrus ı Ŀ . 2-ĳ 8 ı ı Oribatula, Oppia , ollembola ı Folsomia Ŀ . K Ŀ Ŀ . 10 ı ı Ŀ , 36, 7, 9, 11 ı - Ŀ . L , ı Ŀı 50,7% 40,2%- . G () Carabidae – ı Elateridae – Ŀ . Aphididae – ( Ľ ĳ) ĳ ĳ ĳ, Ŀ ! . ĳ ĳ . 254 Y . . 1 (33). 2012

Ĳ , Ŀ ĳı , Ŀ Ľ Lı ı . Tı Ŀ L Ŀ ĳ Ŀ Ŀı ! Ŀ . ı Ľ ı . Ŀ Ŀ Lı ı ı Ľ . , Ŀ Ľ Ľ Ŀı ĳ ĳı ĳ ı Ľ. Q Ŀ ĳ . X -, ĳ ĳ . Xı Ľ ĳ ı ı ı ı . J ĳ ĳ , ĳ , ı Ľ . c! ! , , Ŀ ı Ŀ Ŀ. Rı Ŀ . ľ Ŀ Ŀı Ŀ . L ı Ľ Ŀı Ŀ , Ŀ ı , - ı ı , ĳ ĳ ĳ Ľ, ! ĳ . ļJ 1. I! F.R. ! ! !.//L . . - 1969. - X. 30. - 3 6. - W. 652 - 657. 2 W. Kirby W. !. H c . R.1826. -X 4. – 324 . 3. ] J T ! ! ! . R.-Q.: 1936. - X 2. – 389 . 4. J H.H. V . R.: . H . -, 1883. – 376 . 5. U.F. U " V. , . - R.:1886. - 322 . 6. H.F. T ! . - R.: S. 1973. - . 1.- 447 . 7. W G.V. U . - R.: S, 1980. - 243 . 8. I! F.R. !. - R.: S, 1982. - W.38 - 46. 9. G F. G., S M U: ! ! ! // M. . - 2002. - X. 81. - 3 7. - W. 779 – 796. 10. G F.G., G H.., 1997. S (Collembola) ! K // M. . - 1996. - X. 76. - 3 4. - W. 409 – 417. 11. R H.U. G (Collembola) ! !- //M! - . - V, 1985. - W. 149 - 209. 12. T K WWWV, . «S», R-Q.X 1. 1964. W. 42-110. 13. G –M «U ». R. S, 1988, 214 . *** J , . S Nothrus, Scheloribates. . N , Anurida, Folsomia. . U L , - , - , . *** Data show that on the investigated objects of microzoofauna very monotonous and poor. On the investigated areas of dump breeds found out a few of representatives of testacean claws of Nothrus, Scheloribates. It related to the hydrothermal mode of soils. Researches of quantity of Collembola on dumps showed, that on all dump breeds meet in very a few of Anurida, Folsomia. In cuts stopped up on poorly and not copsy dumps there are single copies of microzoofauna. Presence of representatives micro and mesofaunas in soils of deposit of Kokjon are given by founding to consider that from scarcity of vegetable food, hard natural and climatic terms the initial process of soil fotmation on dumps goes nonactive, slowly.

YJ 595.754 U.. RQ Z UTQLTQ (HETEROPTERA) UTJTIT U «ľQO ľQJ=» S RTS V

«P' ''». 10 47 . (25), (9), ( 4) J «Ŀ Ŀ» ! . R ! ! 2011 . U . , W . , ! . RQ RTJ QJTO [1, 2]. S , !. J! ! . H SY. . 1 (33). 2012 255

W Nabidae – Himacerus maracandicus (Reuter, 1890). T ! 400 3000 [3]. J! ! !, , , . M . . U! (, !, ). - . Nabis (Nabis) ferus (Linnaeus, 1758). T , , , . . ^ ! , ! , !, , . X . W Fnthocoridae – R Anthocoris confusus Reuter, 1884. S , !. H ! , , , . U! !, , . I . Anthocoris nemorum (Linnaeus, 1761). S !, ! (1000-3000 ..). M ( , ! !, , , , ! , ! Miridae). X . Orius niger Wolff, (1804). H ! . L ! !: , , . , ! , , , ! . I . Orius (H.) horvathi (Reuter, 1884). H ! . L ! !. U! !, , , ! , [4]. X . Orius minutus (Linnaeus, 1758). R! . S ! !, !. ! , , , ! . X . W Miridae– W! Deraeocoris (C.) punctulatus (Fallen, 1807. H ! !. M. U! (!, ). X . Lygus rugulipennis Popp. 1911. R!. H : , , . !. H [5]. I . Lygus pratensis (Linnaeus), 1758. c. R!. W , , . U . X . Lygus gemellatus (Herrich-Schaffer, 1835). R!, , ! !. U , , . X . Lygus punctatus (Zett. 1839). c. T. R!, , ! !. U ! . X . Adelphocoris lineolatus (Goeze, 1778). S , !. U . X . Stenodema calcarata (Fallen, 1807). V!, . U . X . Stenodema trispinosum Reuter, 1904. I. V!, . H. I . Trigonotylus ruficornis (Geoffr. 1785). S ! . U – , ! , , , ., – !, !, . X . Trigonotylus caelestialium (Kirkaldy, 1902). S . W , . I . Chlamydatus pullus (Reuter, 1870). H! . H !. R! . . . X . I . W Tingidae – Acalypta marginata (Wolff, 1804). H , !, . X . Agramma minutum Horvath, 1874. U , ! . X . W Reduviidae – Rhinocoris annulatus (Linnaeus, 1758). S !, ! , . , (, , , .). M . Rhynocoris iracundus Poda, 1761. H! !, ! . S , . R , ! !. 256 Y . . 1 (33). 2012

, ! (, , , .) [6]. M- . W Lygaeidae – Q Lygaeus equestris Linnaeus, 1759. H !. U! ! . X . Nysius ericae groenlandicus (Zetterstedt, 1838). S , , . U. I . Platyplax salviae Schilling, 1829. ] (Salvia stepposa), , . M . Rhyparochromus pini Linnaeus, 1758. T . S !. U. U Echium vulgare, Potentilla auserina, Thymus marschalianus. U! [7]. X . W Pyrrhocoridae – Pyrrhocoris apterus (Linnaeus, 1758). U. S , , !. I . W Coreidae – V, Syromastus rhombeus (Linnaeus, 1767). . S (Arenaria, Cerastium .) [8]. M . Coreus marginatus (Linnaeus, 1758). . S !. X . W Rhopalidae - G Corizus hyoscyami (Linnaeus, 1758). !. W! [9]. X . Liorhyssus hyalinus Fabricius, 1794. R!. !! ! . . Brachycarenus tigrinus (Schilling, 1829). S . . R!, ! !. X . Rhopalus maculatus (Fieber, 1837). H . S ! !. X . Rhopalus (Rhopalus) subrufus Gmelin, 1790. R. S . . Rhopalus (Rhopalus) distinctus Signoret, 1859. H !, . L Thymus [9]. X . Stictipleurus punctatonervosus (Goeze, 1778). H . . S !. X . Stictopleurus crassicornis (Linnaeus, 1758). H . ] ! . S . X . Myrmus calcaratus Reuter, 1891. . S (Stipa ssp.) ! ! Agropyron cristatum [10]. X . W Pentatomidae – S! Graphosoma lineatum Linnaeus, 1758. U . H! ! [11]. X . Sciocoris microphtalmus Flor, 1860. R. X ! . X! ! – !. I . Holcostethus strictus vernalis (Wolff, 1804). H , , 850-2300 , !. S , , , , ! . X . Palomena viridissima (Poda, 1761). ^ - !. X . Carpocoris melanocerus (Mulsant & Rey, 1852). R ( , !, ); ( ! !, ! Verbascum). M . Dolycoris baccarum Linnaeus, 1758. H . U. H . X . Eurydema ornat (Linnaeus, 1758). H ! ( 900 ) !. X . Eurydema dominulus (Scopoli, 1763). R ( , ). ^ ( ). X . Eurydema oleracea (Linnaeus, 1758). R (, , , , 800-2400 ). S . X . MQb TLJ H 2011 «Ŀ Ŀ» ! 47 10 . H (25), (9), ( 4) . H SY. . 1 (33). 2012 257

Q 1 F.S. R ! ! // - FS WWWV. - R.-Q., 1957. - 124 . 2 .R., X.Q. T! Heteroptera (Hemiptera) // T WWWV. - «S». - R.-Q. 1964. - X. 1. - W. 655-843. 3 .R. U Nabidae. WWWV. S // Q. 1981. - S. - X. 13. - H. 2. - 327 . 4 c c.W. U . Anthocoridae (Heteroptera) W F // c. . - «S». - Q., 1976. - X. 55. - H. 2. - W. 369-380. 5 Wagi O. 1954. Bladtaeger (Miridae) of forekomst of frouden kim hos skaermolomstrade (Umbelliferae) // Tidsskr. Planteave. - T. 58. - 31. - P. 58-90. 6 U H.I. U. . Y // . 1987. - S . - X. 21. - H. 5.- 248 . 7 U H.I. Qiï. Yï // , 1969. - X. 21.- H. 3. - 388 . 8 U H.I. . Y // . 1962. - H. FS YVWV. - X. 21. - H. 2. - 163 . 9 U H.I. U Rhopalidae (Heteroptera) WWWV // Q. 1986. - S. - 132 . 10 .R. S (Heteroptera) WWWV // S . X.M. - FS WWWV. - 1964. - X. 34. - W. 113-130. 11 U H.I. _ W F (Hemiptera, Pentatomidea) // . - . - 1965. - 329 . *** R «P' ''» '. M %' 10 47 ' . M (25), (9), % (4) . *** The article presents the materials of the fauna of Heteroptera of national park "Polsay kolderi". As a result of the studies identified 47 species of 10 families. In the zoogeographically predominantly transpalaearctics (25), holarctics (9), westpalaearctics and transeurasians (4) species. YJ 595.754 U. . TZ] M UTQLTQ (HETEROPTERA) U] Ub M S RTS V

P 155 , , 61 , - 41 , - 21 , - 3 , - 25 , - 3 , Periphima batesoni. J U . U !! ! ! , , ! . U - : M – , ! ! ; , , . L! - . M – , ! . M! ! !. – ! , ! ! !, . U ! . V! 4 : ) U – ! !, !! . ) ^ – ! ! , !! (, ). ) Y ! ! . ) R – ! ! . H ! ! – , , .. W ! ! . V! ! . RQ RTJ QJTO ! [1, 2, 3]. 155 ! , ! . S ! ! . W Lygaeidae U: Nysius thymi thymi (Wolff, 1804), Plinthisus ptilioides Puton, 1874, Horvathiolus heydeni (Puton, 1892), Horvathiolus syriacus (Reuter, 1885), Melanocoryphus albomaculatus (Goeze, 1778), Tropidophlebia costalis (Herrich-Schaffer, 1850), Diomphalus hispidulus Fieber, 1864, Camptocera glaberrima (Walker, 1872), Bleteogonus beckeri Frey-Gessner, 1863, Bleteogonus circumcinctus (Reuter, 1885), Emblethis verbasci (Fabricius, 1803), Hyalocoris pilicornis Jakovlev, 1874, Xanthochilus turanicus (Wagner, 1961), Lygaeus murinus (Kiritshenko, 258 Y . . 1 (33). 2012

1914) – ! ! , ! , ! . ^ : Oxycarenus pallens (Herrich-Schaeffer, 1850), Emblethis denticollis Horvath, 1878 – : Centaurea, Spiraea, Lepidium, Alyssum ., ! . Y : Blissus putoni Jakovlev, 1875, Jakowleffia setulosa (Jakovlev, 1874), Bianchiella sarmatica Kiritshenko, 1926, Bianchiella sarmatica Kiritshenko, 1926, Haploprocta bicolor Kiritshenko, 1911, Haploprocta pustulifera (Stal, 1860) - , Thymus marschalianus, ! . R: Paranysius fraterculus fraterculus Horvath, 1895 - Lepidium, Alyssum. M: Geocoris desertorum (Jakovlev, 1871), Geocoris erythrocephalus (Lepeletier & Serville, 1825), Geocoris pubescens (Jakovlev, 1871), Geocoris megacephalus (Rossi, 1790), Geocoris aspasia Linnavuori, 1972, Geocoris hirticornis Jakovlev, 1882, Geocoris pattakumenis Kiritshenko, 1914 - , !! , , W Rhopalidae U: Brachycarenus tigrinus (Schilling, 1829) – , , ! , ! !. ^ : Stictopleurus abutilon (Rossi, 1790), Stictopleurus angustus Reuter, 1900, Leptoceraea femoralis (Horvath, 1897), Chorosoma schillingii (Schilling, 1829) - , !. Y : Agrophopus suturalis Reuter, 1900, Chorosoma gracile Josifov, 1968, Chorosoma longicolle Reuter, 1900, Myrmus glabellus Horvath, 1901 - ! Aristida, : Aristida pennata, A. karelinii, A. dscensionis, : Agropyron, Stipa. R: Agrophopus lethierryi Stal, 1872 – c (Cynodon dactylon). W Coreidae ^ : Centrocoris spiniger (Fabricius, 1781), Enoplops disciger Kolenati, 1845, Bothrostethus annulipes (Herrich-Schaeffer, 1835) - , ; ! !. Y : Cercinthinus annulipes Kiritshenko, 1916, Haploprocta pustulifera (Stal, 1860), Phyllomorpha lacerat Herrich-Schaeffer, 1835 - Ceratula, G , , ! . W Cydnidae U: Aethus hispidulus (Klug, 1845), Aethus pilosus (Herrich-Schaeffer, 1834), Byrsinus flavicornis (Fabricius, 1794), Byrsinus fossor (Mulsant & Rey, 1866), Byrsinus laeviceps (Kerzhner , 1972), Byrsinus penicillatus Wagner, 1964, Byrsinus rugosus (Jakovlev, 1874), Byrsinus pilosulus (Klug, 1845), Geotomus elongatus (Herrich- Schaeffer, 1840), Geotomus ciliatitylus Signoretti, 1883, Microporus nigrita (Fabricius, 1794), Linospa candida (Horvath, 1889), Linospa orbicularis (Jakovlev, 1885), Ochetostethus nanus (Herrich-Schaffer, 1834) – , ! . ^ : Stibaropus henkei (Jakovlev, 1874), Stibaropus hohlbecki Kiritshenko, 1912, Byrsinus comaroffii (Jakovlev, 1879), Byrsinus discus Jakovlev, 1906 - , Elymus giganteus, E.arenaria. Y : Exosehirus validus (Jakovlev, 1877) - Eurotia ceratoides, Atriplex cana. W Scutelleridae U: Ellipsocoris tamerlani Kiritshenko, 1914, Melanodema carbonaria Jakovlev, 1880, Odontotarsus angustatus Jakovlev, 1880, Odontotarsus impictus Jakovlev, 1886, Odontotarsus obsoletus obsoletus Horvath, 1906, Odontotarsus rufescens Fieber, 1861, Phimodera fumosa Fieber, 1863, Phimodera bergi Jakovlev, 1905, Odontoscelis zarudnyi V.G.Putshkov, 1965, Odontoscelis byrrhus Seidenstucker, 1972, Odontoscelis dorsalis (Fabricius, 1798), Odontoscelis fuliginosa (Linnaeus, 1761), Polyphima koenigi Jakovlev, 1889, Odontotarsus rufescens Fieber, 1861 – . !; . ^ : Irochrotus lanatus (Pallas, 1773), Irochrotus turanicus Kerzhner, 1976 - ! , Secale, Agropyrum, Elytrigia, ! !. U ! ! Periphima batesoni Jakovlev, 1889. W Pentatomidae U: Antheminia lunulata (Goeze, 1778), Antheminia pusio pusio (Kolenati, 1846), Carpocoris coreanus Distant, 1899, Desertomenida albula Kiritshenko, 1914, Desertomenida jakowleffi (Horvath, 1907), Desertomenida quadrimaculata (Horvath, 1892), Sciocoris cursitans cursitans (Fabricius, 1794), Sciocoris deltacephalus (Fieber, 1861), Menaccarus arenicola (Scholtz, 1847), Brachynema germari (Kolenati, 1846) - , , , , , , , , , !. ^ : Aelia acuminata (Linnaeus, 1758), Sciocoris macrocephalus Fieber, 1851, Sciocoris sulcatus Fieber, 1851, Sciocoris maculatus Fieber, 1851, Menaccarus deserticola Jakovlev, 1900, Menaccarus H SY. . 1 (33). 2012 259 dohrnianus (Mulsant & Rey, 1866), Eurydema wilkinsi Distant, 1879, Eurydema maracandica Oshanin, 1871, Crypsinus angustatus (Baerensprung, 1859), Capnoda batesoni Jakovlev, 1889, Thologmus flavolineatus (Fabricius, 1798) - , , , , !. Y : Leprosoma tuberculatum Jakovlev, 1874, Ochyrotylus helvinus Jakovlev, 1885 - , ! Alyssum, Peganum harmala, Atrophaxis. W Berytidae U: Gampsocoris punctipes punctipes (Germar, 1822) – ! ! : , , , , , . ^ : Berytinus signoreti Fieber, 1859 - , . W Tingidae ^ : Dictyonota atraphaxius Golub, 1975, Dictyonota horvathi (Kiritshenko, 1914), Dictyonota rectipilis (Asanova, 1970, Dictyonota sareptana Jakovlev, 1874, Tingis biseriata (Horvath, 1902), Tingis pauperata (Puton, 1879), Tingis grisea Germar, 1835, Kalama tricornis (Schrank, 1801) - , , , . Y : Catoplatus cathusianus (Goeze, 1778), Tingis pusilla (Jakovlev, 1873), Oncochila scapularis (Fieber, 1844) - , ! Seriphidium, c Eryngium . . R: Dictyonota ephedrae (Kerzhner, 1964) - Ephedra distachya. W Reduviidae M: Ploiaria turkestanica P.V.Putshkov, 1984, Empicoris culiciformis (De Geer, 1773), Holotrichius bergrothi Reuter, 1891, Holotrichius ilius Dispons, 1964, Holotrichius kizilkumi Dispons, 1964, Holotrichius mesoleucus Kiritshenko, 1914, Holotrichius tristis Jakovlev, 1874, Pasira basiptera Stal, 1859, Reduvius disciger Horvath, 1896, Reduvius elegans (Jakovlev, 1885), Reduvius tenuicornis (Jakovlev, 1889), Reduvius testaceus (Herrich- Schaeffer, 1845), Oncocephalus impictipes Jakovlev, 1885, Vachiria prolixa Kiriyshenko, 1925 - ! : – , , , !, , . W Piesmatidae Y : Parapiesma variabile (Fieber, 1844) - : Caryophyllacaea, Herniaria spp., Atriplex cana. W Nabidae M: Nabis palifer Seidenstucker, 1954, Nabis sinoferus sinoferus Hsiao, 1964 - ! !, -!, !. W Anthocoridae M: Orius horvathi (Reuter, 1884), Orius vicinus (Ribaut, 1923) – ! !, , , , !, ! , . W Miridae U: Chlamydatus eurotiae Kerzhner, 1962, Atomophora pantherina Reuter, 1879, Compsidolon pumilum (Jakovlev, 1876), Orthotylus turanicus Reuter, 1883, Camptotylidea suturalis (Reuter, 1903), Camptotylus bipunctatus (Reuter, 1879), Glaucopterum vilgus V.G.Putshkov, 1977 - , . !. ^ : Maurodactylus albidus (Kolenati, 1845), Atomoscelis onusta (Fieber, 1861), Orthops pilosulus Jakovlev, 1877, Polymerus brevicornis (Reuter, 1879), Stenodema turanica Reuter, 1904, Solenoxyphus lepidus (Puton, 1874) - , , . Y : Hyoidea notaticeps Reuter, 1876 - . M: Phytocoris kazachstanicus Muminov, 1989, Phytocoris kyzylkumi Muminov, 1989, Phytocoris undulatus Reuter, 1877. MQb TLJ H ! ! 155 13 . W ! . Lygaeidae – 30 (19%), Pentatomidae – 23 (14,5%), Cydnidae – 19 (12%), Miridae – 17 (10,7%), Scutelleridae – 17 (10,1), Reduviidae – 14 (8,9), Tingidae – 12 (7,6%), Rhopalidae – 10 (7%), 5 1-2 . 12 (Nysius thymi, Oxycarenus pallens, Geocoris ater, Leptodemus minutus, Emblethis verbasci, Hyalocoris pilicornis, Liorhyssus hyalinus, Brachycarenus tigrinus, Irochrotus lanatus, Sciocoris deltacephalus, Thologmus flavolineatus, Brachynema germari) !! ! , 49 - , 94 - . U , 61 , - 41 , - 21 , - 3 , - 25 , - 3 , ! ! Periphima batesoni. U !, , . , – . Cydnidae: Cydnus aterrimus, Stibaropus hohlbecki, 260 Y . . 1 (33). 2012

S. henkei, S. hohlbecki, Aethus hispidulus, A. pilosus, Byrsinus discus, B. flavicornis, B. fossor, B.laeviceps, B. penicillatus, Geotomus elongatus, G. ciliatitylus, Microporus nigrita, Linospa candida, L. orbicularis, Exosehirus validus . – !, . , , ! ! . U – ! . M ! ! . Q 1 U H.. R ! . - H. - 1970. - 192 c. 2 .. U . – R: H^, 1971. - 424 . 3 .R., X.Q. T! Heteroptera (Hemiptera) . T WWWV. X. 1. - R.-Q.: - «S», 1964. - W. 655-843. *** M %' '' 155 ' , '' , 61 , – 41 , – 21 , – 3 , – 25 , – 3 , Periphima batesoni ' . *** As a result of researches of the fauna of sandy deserts of Kazakhstan 155 species of Heteroptera are revealed. Among them by fodder specialization prevail polyphytophages (61 species), wide oligophytophages - 41 species, narrow oligophytophages - 21 species, monophytophages - 3 species, zoophages - 25 species, zoophytophages - 3 species, food connections are not found out at Periphima batesoni.

YJ 581.9 M.. , .I ., .L. ], I.. TMTTM RO POACEAE BARNHART. LTIT Q S Y . -

Poaceae Barnhart L F. F , L F 214 Poaceae Barnhart., 32 16 . L F !!! ! . T X!-^! M L-F ! ^- , ! . W - L-F G, SV ! - L-F . W-! L-F L-F , ! - 500 . ! L-F F! , ! L-F L- F . H L-F ! . F, - – . c-S. L F W F W , ! ! 400 , , ! ! ! . J ! , , 4622 . , ! ! , ! . Z L-F 4442 ! , . U L-F 3800 1100 , 500 3000 . [1]. L-F ! - !, , - ! . I ! Y-X! !. W! J-G , ! . M . c ! ! !. S ! ! X!-^! ! ! V . L-F ! ! , - !! ! !. W! L-F ! 9 . ! ! L-F : -, - - . ! ! ! [1]. S ! , !! , ! . T !! - . H SY. . 1 (33). 2012 261

, !. H ! ! . Y ! ! , . H !. c !: ! ! , !! ! . H L-F ! . H - ! ! - , ! L-F. T ! . H ! , ! L-F . , ! L-F ! !! . J! ! ! ( ), . J ! , , , . ! ! () - – ! L-F. -F, L-F !! , , ! !. H L-F ! , ! 50-200 . ! - , -, . ! ! . I! -! - , - . U , ! -F L-F . T !! ! , ! . W ! ! - . I ! , , . U - !, -. S L-F – L-F ( 15 ), L-F . [1, W.85]. S . Voaceae Barnhart. 2007 . , 2010- 2011 . T ! -. T, - . U « WWWV», « », «T W F», « », «M WWWV», « » R.C. G [2-7]. V F.Q. X! [8]. S , ! W.. ] [9]. H 214 , !! 32 16 (). X Poaceae L T: Magnoliophyta : Liliopsida U: Poales : Poaceae Barnhart U : Pooideae A. Br 1: Brachypodieae (Hack.) Hayek 12 2: Triticeae Dum. U: Triticinae Trin. ex Griseb. 5 38 262 Y . . 1 (33). 2012

U: Hordeinae Dum. 521 3: Bromeae Dum. 310 4: Aveneae Dum. U: Aveninae C. Presl. 2 6 U: Koeleriinae Aschers. et 2 10 Graebn. U: Airinae Benth. 1 2 U: Miliinae Dum. 1 2 U: Agrostidinae Griseb 4 16 5: Phalarideae Benth. U: Anthoxanthinae Miq. 2 3 U: Phalaridinae Griseb. 1 1 6: Phleeae Dum. U: Beckmanniinae Nevski 1 2 U: Phleinae Benth. 1 4 U: Alopecurinae Dum. 1 3 7: Poeae R. Br. U: Festucinae C. Presl 2 16 U: Poinae Stapf 2 34 U: Dactylidinae Stapf 1 1 8: Meliceae Endl. U: Glyceriinae Dum 1 1 U: Melicinae Fries 1 3 9: Stipeae Dum. 5 25 10: Danthonieae Zotov 1 1 11: Aristideae C. E. Hubb. 1 1 12: Aeluropodeae Nevski ex Bor 1 2 13: Pappophoreae Woods 1 1 14: Cynodonteae Dum. U: Eragrostidinae C. Presl 2 6 U: Chloridinae C. Presl 2 2 U: Sporobolinae Benth. 1 2 15: Paniceae R. Br. U: Brachiariinae Butzin 2 2 U: Setariinae Dum. 1 3 16: Andropogoneae Dum. U: Andropogoninae C. Presl 1 1 : 32 214 H !, Poaceae Triticeae Dum., !! 59 10 , Poeae R. Br. (51 , 5 ) Stipeae Dum., ! 25 , !! 5 . Q 1 ! ! V : Y . - F: K.S. . . Q. S. I, «F», 2010. - W. 81-85. 2 WWWV. - R-Q., 1934-1964. - X. 1-30. 3 . – F – F: S, 1956-1967. - X.X. 1-9. 4 T W F. – X: FS, 1968-1996. - X.X. 1-10. 5 . – F-F: S, 1969-1972. - X.1-2. 6 \ S.S. M WWWV. - Q.: S, 1976. - 788 . 7 G R.W. . – F: Ĩ, 2001. - X. 1-2. 8 X! F.Q. W . - Q.: S,1987. – 439 . 9 ] W.. W ! WWWV. - Q.: S, 1981. - 509 . *** R L' F Poaceae Barnhart ''' . X ''' Poaceae Barnhart 214 ' % 32 , 16 . *** This paper is devoted to analysis of Poaceae Barnhart family of Zhetysu Alatau, Analysis has shown that there are 214 species of Poaceae Barnhart family, which are referred to 32 genuses and 16 tribs.

H SY. . 1 (33). 2012 263

YJ 594.1+594.3 (574) .Q. UJQb T\ TTR]TIT T Z RTQQMT M RTS V, e-mail: [email protected]

P, , . U . Y, P 496 , 2 , 42 121 , 385 92 33 , – 111 29 9 . J T «S-! ! ! V », RTS V 2010 ., . R ! . H ! «» ! , , , . RQ RTJ QJTO M ! V , Z.. F.H. W [1]. T ! .. Y [2]. G 100 .

MQb TLJ F , 2 : , ! ! , ! ! , , ! ( , F !). S ! ! ! ! . GASTROPODA NERITIMORPHA [= Neritopsina] S NERITOIDEA Rafinesque, 1815 W NERITIDAE Rafinesque, 1815 V Theodoxus (2 ). LITTORINIMORPHA S RISSOOIDEA Gray, 1847 W RISSOIDAE Gray, 1847 U RISSOINAE Gray, 1847 V Contectiana (1 ) LITTORINIMORPHA S RISSOOIDEA Gray, 1847 W BITHYNIIDAE Gray, 1857 U BITHYNIINAE Gray, 1857 V Bithynia (2 ); Opisthorchophorus (4 ), Paraelona (1 ) W HYDROBIIDAE Stimpson, 1865 U BELGRANDIINAE de Stefani, 1877 V Andrusovia (4 ); Martensamnicola (1 ); Caspia (2 ); Caspiohydrobia (25 ). U PYRGULINAE Brusina, 1882 (1869) V Pyrgula (22 ); Turricaspia (19 ); Pseudamnicola (2 ); Sibirobythinella (1 ). HETEROBRANCHIA S VALVATOIDEA Gray, 1840 W VALVATIDAE Gray, 1840 U VALVATINAE Gray, 1840 V Cincinna (4 ); Valvata (3 ) I OPISTHOBRANCHIA NUDIPLEURA 264 Y . . 1 (33). 2012

U NUDIPLEURA “I” NUDIBRANCHIA CLADOBRANCHIA W TERGIPEDIDAE Bergh, 1889 V Tenellia (1 ) PULMONATA S! BASOMMATOPHORA SIPHONARIOIDEA Gray, 1827 S ACROLOXOIDEA Thiele, 1931 W ACROLOXIDAE Thiele, 1931 V Patella (1 ); Acroloxus (2 ) S LYMNAEOIDEA Rafinesque, 1815 W LYMNAEIDAE Rafinesque, 1815 V Aenigmomphisla (2 ); Lymnaea (37 ); S PLANORBOIDEA Rafinesque, 1815 W PLANORBIDAE Rafinesque, 1815 U BULININAE V. Fischer et Crosse, 1880 V Planorbarius (5 ) U PLANORBINAE Rafinesque, 1815 V Ancylus (1 ); Anisus (15 ); Armiger (2 ); Choanomphalus (1 ); Hippeutis (3 ); Planorbis (5 ); Polypylis (1 ); Segmentina (4 ); Stenogyra (1 ) W PHYSIDAE Fitzinger, 1833 U APLEXINAE Starobogatov, 1967 V Aplexa (3 ); Euglessa (1 ) U PHYSINAE Fitzinger, 1833 V Physa (1 ) S ELLOBIOIDEA L.Pfeiffer, 1854 (1822) W CARYCHIIDAE Jeffreys, 1830 V Carychium (3 ) S! STYLOMMATOPHORA A.Schmidt, 1855 W SUCCINEIDAE Beck (1837) U SUCCINEINAE Beck (1837) V Novisuccinea (5 ); Succinea (2 ) U OXYLOMINAE Schileyko et Likharev, 1986 V Oxyloma (2 ) W COCHLICOPIDAE Hesse, 1922 V Cochlicopa (7 ) W ORCULIDAE Steenberg, 1925 U ARGNINAE Hudec, 1965 V Acanthinula (1 ) U VALLONIINAE Morse, 1864 V Vallonia (6 ) W PUPILLIDAE Turton, 1831 V Gibbulinopsis (1 ), Pupilla (9 ), Gastrocopta (1 ) W VERTIGINIDAE Fitzinger, 1833 V Vertigo (3 ) W TRUNCATELLINIDAE Steenberg, 1925 V Columella (5 ), Truncatellina (2 ), Vertigo (10 ) W PYRAMIDULIDAE Kennard et Woodward, 1914 V Pyramidula (1 ) U PSEUDONAPAEINAE Schileyko, 1978 V Laevozebrinus (2 ), Mastoides (1 ), Pseudonapaeus (27 ), Subzebrinus (4 ), Turanena (8 ) U CHONDRULOPSININAE Schileyko, 1978 V Chondrulopsina (1 ) U MERDIGERININAE Schi1eyko, 1984 V Merdigera (1 ) U ENINAE Woodward, 1903 V Chondrula (1 ), Ena (1 ) W PUNCTIDAE Morse, 1864 V Punctum (1 ) W DISCIDAE Thiele, 1931 V Discus (1 ) H SY. . 1 (33). 2012 265

W ORCULIDAE Pilsbry, 1918 V Sphyradium (1 ) W ZONITIDAE Morch, 1864 V Perpolita (2 ) U OXYCHILINAE Hesse, 1927 V Oxychilus (1 ) W VITRINIDAE Fitzinger, 1833 U VITRININAE Fitzinger, 1833 V Vitrina (2 ) U PHENACOLIMACINAE Schileyko, 1986 V Phenacolimax (1 ) W GASTRODONTIDAE Tryon, 1868 V Zonitoides (1 ) W EUCONULIDAE S. Baker, 1928 V Euconulus (1 ) W ARIOPHANTIDAE Godwin-Austen, 1888 V Macrochlamys (5 ) W AGRIOLIMACIDAE H. Wagner, 1935 V Deroceras (6 ), Arion (1 ) W LIMACIDAE Rafinesque, 1815 U LIMACINAE Rafinesque, 1815 V Turcomilax (6 ) W PARMACELLIDAE Gray, 1860 V Candaharia (3 ), Parmacella (1 ) W ARIONIDAE Gray U ARIONINAE Gray V Arion (2 ) W HELICIDAE Rafinesque, 1815 U ARIANTINAE Morch, 1864 V Cepaea (1 ) W BRADYBAENIDAE Pilsbry, 1939 V Fruticicola (22 ), Karaftohelix (1 ); Ponsadenia (5 ) W HYGROMIIDAE Tryon, 1866 U TRICHIAINAE Lozek, 1956 V Leucozonella (10 ), Nanaja (2 ), Odontotrema (1 ), Xeropicta (1 ). U HYGROMIINAE Tryon, 1866 V Chilanodon (1 ), Kalitinaia (1 ), Lindholmomneme (4 ), Monachoides (1 ), Pseudochondrula (1 ), Pseudotrichia (1 ), Xeropicta (2 ) U ARCHAICINAE Schileyko, 1978 V Archaica (2 ) U PAEDHOPLITINAE Schileyko, 1978 V Angiomphalia (5 ), Euomphalia (1 ), Paedhoplita (5 ). BIVALVIA TXVJ GASTROTRITEIA W MYTILIDAE Rafinesque, 1815 V Mytilaster (1 ) TXVJ GASTROPEMPTA W DREISSENIDAE Gray, 1840 V Dreissena (11 ) W UNIONIDAE Rafinesque, 1820 U UNIONINAE Rafinesque, 1820 V Tumidiana (2 ), Unio (3 ) U PSILUNIONINAE Starobogatov, 1970 V Crassiana (5 ) U ANODONTINAE Rafinesque, 1820 V Anodonta (3 ), Colletopterum (5 ) U PSEUDANODONTINAE Jaeckel, 1962 V Pseudanodonta (1 ) W CARDIIDAE Lamarck, 1809 V Adacna (5 ), Cerastoderma (2 ), Didacna (7 ), Hypanis (9 ) W SEMELIDAE Stoliczka, 1870 266 Y . . 1 (33). 2012

V Abra (1 ) W CORBICULIDAE Gray, 1847 V Corbicula (4 ) W SPHAERIIDAE Jeffreys, 1862 U SPHAERIINAE Jeffreys, 1862 V Amesoda (3 ), Sphaerium (2 ) U MUSCULIINAE Starobogatov in Stadnichenko, 1984 V Musculium (2 ), Rivicoliana (2 ) W PISIDIIDAE Gray, 1857 V Europisidium (1 ), Neopisidium (2 ), Odhneripisidium (2 ), Pisidium (1 ) W EUGLESIDAE Pirogov et Starobogatov, 1974 U EUGLESINAE Pirogov et Starobogatov, 1974 V Cingulipisidium (5 ), Conventus (1 ), Cyclocalyx (1 ), Euglesa (14 ), Hensloviana (6 ), Pseudeupera (9 ) U LACUSTRININAE Korniushin, 1989 V Tetragonocyclas (1 ) X , 496 , !! 2 , 42 121 , !! 385 92 33 , – 111 29 9 . G, ! , ! . W , , . Y! , , ! , - !, , , ! . Q 1. Z.., W F.H. V . - R.: X R. - 2005. - 627 . 2. Y .. S . – F_F: S. - 1990. - 224 . *** R %' ' '''' % %''' %' '' . P ', % ''', ''' '' . 2 , 42 , 121 496 , 33 92 385 ' , 9 29 111 ' . *** In the present work results of revision of the faunistic data on the molluscs of Kazakhstan taken from the literature are resulted, in view of advanced achievements of recent systematics. The list of all taxa from classes up to genera with the indication of quantity of species in each genus is resulted. It is established, that the fauna of Kazakhstan includes 496 species of the molluscs concerning to 2 classes, 42 families and 121 genera, and to Gastropoda – 385 species from 92 genera of 33 families, and to Bivalvia – 111 species from 29 genera of 9 families concern.

YJ 595.771 I.. TTTC_ TR (CULICIDAE) UTJIT UO TIT b- S RTS V

X-^ 21 . F , , : Anopheles maculipennis, Aedes caspius caspius, Ae. dorsalis, Ae. flavescens, Ae. cyprius, Ae. vexans, Culex modestus, Cx. pipiens pipiens, Cx. p. molestus. J ! !! ! , . : ! ( ) - ! ( ), ! . , , - - . ! ! ( Anopheles) ! ( Aedes, Culex, Mansonia, Culiseta .). Anopheles !! ! !. W ! Aedes - , Culex - . RQ RTJ QJTO W ! -- (-!) W X!-^! ! ! [1, 2, 3, 4, 5, 6]. H SY. . 1 (33). 2012 267

! 2009-2011 . W X!-^! ! , F, ^, Q, ^, ^, ^, F, X. H 186 , !! ! ! c . U 3-5 2 10-12 ( 0,2 3 ) . c : , , . X ! 16 26 W0, S ! 5,0-8,0. H ! 21 : Anopheles maculipennis, Anopheles pulcherrimus, Anopheles claviger, Anopheles messeae, Culiseta alaskaensis, Aedes caspius caspius, Ae. c. dorsalis, Ae. straminaeus, Ae. cantans, Ae. flavescens, Ae. cyprius, Ae. pionips, Ae. pullatus,, Ae. catarphylla, Ae. leucomelas, Ae. vexans, Ae. cinereus, Culex theileri, Culex modestus, Cx. pipiens pipiens, Cx. p. molestus. Anopheles , , 1 . 20 300 !. U Culex 100 450 . 1 . . Aedes , , ! !! , 50144 . 1 .. U Aedes ! ! !!! - ! , Culex Anopheles !! ! : – ! !; – !. H ! ! . W! !! ( 6 10 ) ( 19 23 .). MQb TLJ Y, 186 , !! ! ! c 21 , 4 . F ! , , : Anopheles maculipennis, Aedes caspius caspius, Ae. dorsalis, Ae. flavescens, Ae. cyprius, Ae. vexans, Culex modestus, Cx. pipiens pipiens, Cx. p. molestus. R !! ! : , , , , , , ^, ^, ^, , F X, ! . U! - ! , Culex Anopheles !! ! : – ! !; – !, !!. U! Aedes !, Anopheles - !. T (.. ) Anopheles ! !, Culicidae !. H . S! ! (6.00-10.00), (14.00-19.00) (20.00-23.00) . S . Q 1 I P.M. F (Diptera: Culicidae), WWWV. // U!. - 1997. - X.31, . 6. - W. 473-485. 2 I F.H., R F.W., ^ F.F. Culicidae // WWWV: S . S. Q . - X. M. H. 4. - Q. 1970. - 384 W. 3 J F.S. (Diptera, Culicidae) // F-F, - 1970. - 220 . 4 J F.R. G WWWV // F-F - 1978. 5 R F.W. Q (. Culicidae) WWWV // T WWWV. . FS WWWV. - 1936. - 383 . 6 R F.W. Q WWWV (. Culicinae) // T . 337. - R.-Q.: S. - 1951. - 290 . *** M %' ' X-^ ' 21 ' . ' : Anopheles maculipennis, Aedes caspius caspius, Ae. dorsalis, Ae. flavescens, Ae. cyprius, Ae. vexans, Culex modestus, Cx. pipiens pipiens, Cx. p. molestus '' '' . *** As a result, research in the piedmont steppes waters of the Northern Tien Shan were found 21 species of mosquitoes. Analysis of the percentage of mosquitoes, showed that the dominant species were: Anopheles maculipennis, Aedes caspius caspius, Ae. dorsalis, Ae. flavescens, Ae. cyprius, Ae. vexans, Culex modestus, Cx. pipiens pipiens, Cx. p. molestus. 268 Y . . 1 (33). 2012

T 581.19. .. R, .. , .M. , .. INULA L Jļ=Q= ĲQJ=İ RTZTQTIQ, TRQ Q== - G Inula helenium, Inula macrophylla, Inula britanica ''''' % '' . Inula helenium ''''' % ' . Inula macrophylla ''''' ' ' , ', '' . Inula britanica ''''' ''' , % '' . Ŀ Ľ Ŀ . ľ Ľ ı Ľ Ľ ı ı . Ľĳ ĳ Ľ Ŀ Ŀı Ŀ . Gı ĳ. X Ľ Ľ Ŀ Ľ Ŀ . JĽ Ŀ ĳı ! Ľ ! Ŀ ı ĳ . JĽ Ŀ ĳı Ŀ ı ĳ . G ı ı Ľ , Ŀ Ŀı ı , Ŀ ĳ Ľ Ŀı ı Ŀ ĳ . Ľ Ŀı ! ! ! ! ĳ [1]. RQJ Lļ ļJ= Xĳ ! ĳ ı , ĳ, ĳ , ! Ľ . 70% , W- Ľ (, , , 1:1:1) !. M ĳı ı ! Ľ ! ĳ , Ŀı ı . G Ŀı Ľ ĳ ı . X 1 ı ı Ŀ . F! Ľ ı (XTW-2) !. ı 10-15 . RG-6 ĳ ( 63; 280 ). F! Ŀ ! RTH 1-15 [ ( -15,4 , 8) . ľ Ŀı, ı ! Ľ ! c [2], V.F. G [3] ı . F! ĳ ı Ŀ ı ı Ŀ Ľ ı ı Ŀ ĳ. L Ŀı ĳ 1500-2000 !, ĳ. ľ Ŀı ! [4] Ľ ! ı ı .c Ľ Ŀı [5;] ı . W Ŀı «STATISTICA» . ľı ĳ ı , !, , !, ! Ľ ĳ ı ı ı [6]. ļLQ Lļ TQJ QJ Inula macrophylla - Ŀ 2 , Ŀ Ŀ Ŀ, ı 4-5 , , ı ĳ, 10-12 , . W , , Ŀ, . , - . XĿ ı Ľ 20-25 , 10-12 . F ı Ľ, 30-90 , 12-25 . Wı 2-3 . Wı ĳ , 20-25 , 10-12 . Wı 6 . G Ŀ 11-12 . Wı 2-3 . Iĳ ĳ, , Ľ, ı, 4-5 , , ı ĳ, 10-12 , . ľ ı Ŀ Ŀ Ľ ĳ Ľ , Ŀı Ľ , ı -ı Ľ . W , Ŀ, . X , - . L ı ı ı ĳ ĳ . Wı ĳ , . W Ŀ Ľ 20-25 10-12 . Wı 6 . G Ŀ 11-12 . Wı 2-3 . Iĳ , 5 , Ľ, ı 4-5 , , ı ĳ, 10-12 , (Xı 97,07±3,23 . L Ĳ ı H SY. . 1 (33). 2012 269

Ŀ . L ı 1- 40-70, 2- 25-37 , 3- 17-33, 4- 1-7 . Xı 40 ı ı. T ı . G ĳ ĳ , Ŀ Ŀ Ŀ. G 10-40 , Ľ, Ŀ, Ŀ Ŀ. W , ĳ, ı, . L ĳ, . W , Ľ, ı - . Ŀ ı - . Iĳ Ŀı, ı . ^ ĳ , , ĳ ĳ. T ĳ , ĳ. X Ľ, ı , Ŀ ĳ . R ĳ ĳ. Xı 38,75±2,11 . G 30-37 ı . G ı ĳ . L ı 1- 15-20 , 2- 18-22 , 3- 16- 18 . L ı . Inula macrophylla Ľ Inula britanica Ŀ''' ĩı ĭ ĭĵ ''. M Ŀ ĳı ! ı Ŀ Ŀ ıı ĳ . W ! ĳ Ŀ ĳ (, , ) ĳ ! . Inula macrophylla - Ŀ Ŀ. Wı Ŀı ı – . I ıı ĳ ı ! ĳ ! : , Ľ . c . c Ŀ , ı, . c ĳ 4-6 Ľĳ . c 3-4 . F Ľ . F ı 421,2±1,58 . F ı Ŀ Ŀı . F Ŀ ı ı Ŀ , . Wı ı Ŀ Ŀı . T ı . T , Ŀ , Ŀ . ľ ı Ŀ , Ŀ . ľ ı 1750,6±2,18 . ľ ı Ŀ Ľ ı . G Ŀ Ŀ Ŀ. Wı Ŀı Ŀ, , , 276,6 . Wı , ĳ Ľ . Wı Ŀı , ı, . c 1-2 . Ŀ 5-7 Ŀı, , Ŀ , , ı ı 232,6 . Wı ı ĳ Ŀ , ı 13-15, . W Ŀ Ŀ ĳ-, ı Ľı ĳ ĳ «» , ı Ŀ Ŀ ı Ŀ Ľ ĳ-. W ı - (-). Inula macrophylla, Inula britanica Ŀ''' ĩı ĭ ĭĵ '' Lı ı Ľ ı ! . W- ı Ŀ ĳ ı ı . ľ ı ! ĳ Ľ Ŀ Ŀ. ı ı Ľ Ľ . ľ Ľ , ı ı , !! . ľ . L ı ı ı Ŀ !! . L Ľ Ŀ ı . T Ŀı . L Ŀ . ľ ı ! , Ŀ . c ı . ľ . Inula macrophylla Ŀ ı ! , . Ĳı ĳ, ı , Ŀ ĳ ĳ ı. Ĳı , Ŀ Ľ ĳ ı Ŀı . G Ľ , 270 Y . . 1 (33). 2012

. G ı ı . XĿ Ŀ . G ı () ĳ . G ı Ľ , ı . G ı Ľ . U Ľ “ ” Ŀ . XĿ , , , ı. L , ı Ľ Ŀı, ĳı ı. Tı ĳ Ŀ 150±1,6 , 146±1,6 , 66,6±0,4 . Inula macrophylla 2- Ŀı ı ı , Ľĳ Ĳ ı Ľ ĳ ı- . R ı (ı )- ĳ ı , ı Ŀ ĳ . X 2-3 . X ı Ľ, Ŀ , ĳ . Inula macrophylla 3- Ŀı Ŀ ı Ľ ı, ı , . Yı Ŀ ı Ľ ĳ ı . X Ŀ, Ľĳ . XĿ ı ĳ , Ŀ, Ŀ ĳ. Inula macrophylla Ŀı ı - Ŀ , 5-6 . . U Ŀ Ľĳ. U ĳ - . L ĳ. X 6-7 . G ı Ŀ, 3-4 , , Ŀ Ľ . L ı ı 62,10±0,81 , Ŀ ı ı 22,54±1,23 . Inula britanica Ŀ''' ĩı ĭ ''. L , Ŀı , ı L ı. ľ Ľ ı ıı , !! . Lı ı ı Ŀ !! ( «»- , «» ) ! ı Ľ Ŀ ı . Rı ı Ŀ Ľ Ľ. ı Ľ ı Ŀ Ľ Ŀ. G Ŀı ı ! Ŀ . G , Ľ, - , ! . G Ŀ , . Ŀ. G Ŀ . G Ľ Ľĳ , ı, . G- ĳ Ŀ ĳ . G Ŀ 4-5 , , . XĿ , Ŀı . Y Ľ ı ı . Lı Ľ ĳ Ŀ, Ŀı ı Ľ , ı ( 14). L ı ı 36,71±0,21, Ŀ 18,58±0,22 .. c , Ŀı . L ı ĳı ı , , , ĳ . G . G ı ı 53,64±0,28 , 47,34±0,88 , 12,31±0,21. G Ľ ĳ Ŀ, , , 4-5 . G ı ı 56,37±0,21 .XĿ , , , . ľ , Ľ Ŀ ( 16). ľ ! Ŀ . ľ ĳ ĳı .L Ŀ ĳı , ! ı ĳ ! Ľ Ŀ ı . W Inula macrophylla Ľ Inula helenium Ŀı ı ! ! Ľ . 1. Inula helenium ı Ŀ ı Ŀ ĳ . Inula helenium Ŀı Ľ Ŀ . H SY. . 1 (33). 2012 271

2. Inula macrophylla Ŀı , ı (ı ) , ĳ 2-3 (2- Ŀ), 3 Ŀ 8 , Ŀ ı ĳ , Ŀ, Ŀ ĳ. 3. G Ŀı Ŀ ı Ŀ . ľ , Ľ Ŀ . ļJ 1 F W.R. G // ! . - 2004. - 30 . 2. c. F! . - R.: R, 1980. - X. 1, 2. – W. 2-558. 3. G V.U. W . T . – R.: R IY, 2004. - W. 312. 4. . F! ! . - R.: U, 1966. – X.1. - W. 423. 5. M I.S. R . – R.: S, 1973. - W. 250. 6. R Ĩ.Ĩ T- Ŀ. - F: !, 1978. – X. 2. – 589 . *** Inula , Inula macrophylla Inula helen'um. *** The paper presents the results of a comparative morphological and anatomical analysis of the three species of the genus Inula in vivo growth, defined diagnostic features of the plants and Inula Inula macrophylla helen'um.

YJ 598 .. Rĵ ĲOJİ T – M - , G!

R '' ' ' %' '. T ' = F '' K, G'' % . X'' ' '' - % ' c, % '' ' . ı Ŀ Ľ ĳ Yı ıĳ, Tıĳ F!, Wĳ , Rı!, G Ŀı, Ľ T F! [1]. H.W. Gı [2] ĳı ĳ : 1) W (Struthio camellus) Ľ, ! 5 . !ı ĳ . U ı ı ; 2) W (Struthio mongolicus), Ŀ Ľ ĳ ĳ- ĳ , 2 ıĳ Ŀı = Ŀı [3]. 1958 F ı Kı Ľ V ı ! = Ŀı = ĳ ı ı WWV ĨF ! - ı ! Ŀı G.W. H.H. . Lı = Fı - Xĳ (Gı), GĿ, ^ Ŀı W Ľ X Ŀı K [4]. 1959 X!-^ı T ı ıĳ (a Ŀ ! Ŀı ) Ŀı Ľı ĳı ı . = Ŀı ĳı ı ı 3,5 , ĳ - 2 - . Ĩ ĳı ı ĳ ı ı , ı , ĳ . , = Ŀı ĳı ĳ Ľ ĳ, - ĳ , , 4 [3;4]. T T F! X!-^ı Ľ . H.W. G [2] ı - ĳı ĳ ĳ , « » (U ) . W ı ı - X! ı ĳı ĳ S. mongolicus (T ) . ĳı X!-^ ı Ŀ ı ĳ ĳ ĳ , ĳ , ı ı K, GĿ Ľ ı . F, GĿ ı ĳ (S. mongolicus) . C, ! ĳ ı – ĳ (= F) ĳ . T Fı F! – S. syriacus Rothsch. Ľ Wĳ F – S. camellus L. ĳ . F ^ Ľ Tıĳ F ĳı ĳ – S. massaicus Naum. Ľ S. australis Gurney ĳı ı ı Ŀ , c!ı ıĳ ĳı [3]. 272 Y . . 1 (33). 2012

Wı ı 25 ĳı ı , K Ŀı (« », «») ı ı ı, X (K, F, L, F) ı ; = (^, Ĳ GĿ) ı [5]. F ĳı 4 ĳ Ŀ. K Ľ, T F! ĳ . R, F. Gı [6] T F! F. H 1863 F!ı Ŀ ıı ı ĳ Ľ “ĳ” “ ” . ı “ĳ” , ĳ ı ĳ. W, Q.U. ^ [7] Ľ ĳ Xĳ . ļJ 1. L . - R.: U, 1970. – X. 5. – W.52-53. 2. G H.W. H X!-^!. // X FS WWV. F-F, 1961. – X.15. – W.5-11. 3. G.W. H! W" G – . F, 2007.- 26 . 4. G H.W. W W X!-^!. // X. . H . . ., R.:RIY, 1959. – .1. – W. 21-22. 5. X U.F. X . // G : R Ŀ . -, 12-13 ! 2009 . – F: , 2009. – W. 174-175. 6. G F. L . – R.: - c, 2003. – 960 . 7. ^ Q.R. T! (, ! ). – Q., 1940. *** , P . T N F K, G . , , . *** This article provides evidence that Kazakhstan is the ancestral home of the modern ostrich. They lived in the spur of Ile Alatau Esekartkan, Boguty and elsewhere. The extinction of ostriches in this area was under the influence of a general increase in humidity of the climate cooling and the effort that created the snow cover in winter, in the second half of the bottom of Anthropogenesis.

YJ 581.9 .R. R, .. , .. , . J T\ TT \TUTUQ\O JTIT JR]TIT J LIMONIUM MICHELSONII LINCZ. P . -Z, . F, V P M F ( G, G, X 3 (9 ) , Limonium michelsonii. , . 25 40 , . S Limonium michelsonii , . X Limonium michelsonii. U , . Limonium michelsonii , . S Limonium michelsonii 2 3. 2 1 Limonium michelsonii, 1 3. Limonium michelsonii - . G Limonium michelsonii , .. . U Limonium michelsonii, P V P .. H ! ! , , . T ! I F R W G (1992) ZSKWT R! «DIVERSITAS» R! ! ! (V--L, 1992). U!! (1995) «G ». T ! !! ! !! ! !. U ! !! ! , ! ! . T !! !! ! ! . H SY. . 1 (33). 2012 273

! ! ! [1-4]. T ! ! , , ! ! , ! ! !, !! ! [5-7]. W ! , ! . c ! XX . ] !, !! , ! ! !, !. U !! ! ! ! -G , V . RQ RTJ QJTO T ! Limonium michelsonii – , , . R ! 10-25 . H! M F ( X, G, .]) (], .) [8]. Limnium michelsonii Lincz. U V 21.06.2007 . 3521 [9] , . M 2011 . ! !! ! Limnium michelsonii Lincz.. H 9 ! 3 !. I ! ! . ! . H! ! X.F.V [10] F.F.Y [11]. ! GPS GARMIN GPSMAP 60CSx. I Limnium michelsonii ! 1. H F.F.Y [11]: - ; j – ; imm- ; v- ; g1- ; g2- – ; g3- ; ss-; s- sc- . S ! Limnium michelsonii ! . J! ! . S 10 1 . ( 95 ) S Limnium michelsonii !!. U ! 1 ..

1. I Limnium michelsonii (L) ().

MQb TLJ U ! F R G 13 . K F . Y ! 1283 – 1294 .!, GPS : N 43027.651' E 078047.688'. V ! 274 Y . . 1 (33). 2012

. U – -!, !, . Y . H ! Limnium michelsonii. U - . U ! 55-60%. H 30-32 . T . - Parmelia vagans. F! ! 4 !: I ! ! Artemisia sublessingiana Krasch. ex Poljak, Stipa caucasica Schmalh., Kochia prostrata (L.) Schrad 20-25 ; II ! ! Limnium michelsonii, Tulipa albertii Regel 15-20 ; III ! ! Nanophyton erinaceum (Pall.) Bunge, Anabasis cretacea Pall., Goniolimon cuspidatum Gamajun. 5-10 ; IV ! ! Orostachys spinosa (L.) C.A.Mey., Tortula tortuosa Parmelia vagans, 2-5 . K . Poa bulbosa L., Eremopyrum triticeum (Gaertn.) Nevski, Papaver pavonium Schrenk, Trigonella arcuata C.A.Mey, Tulipa albertii Ziziphora tenuior L.. , , .. . T , . H 10 1 .. S . T , . M ! Nanophyton erinaceum, 656 , Stipa caucasica 187 , ! ! : Orostachys spinosa - 89 , Artemisia sublessingiana - 31 , Ceratocarpus utriculosus Bluk.– 9, Goniolimon cuspidatum – 8, Kochia prostrata – 4, Salsola australis R.Br. – 4 . R !. Limnium michelsonii !! 10 . V , Limnium michelsonii 93 ., – 63 ., – 21 . – 9 . Z, , , Limnium michelsonii . H ! Limnium michelsonii . - - ! F . Y ! 1288 .!, GPS : N 43027.651' E 078047.688'. U -, , . U !. U ! 55-60%. J !! Limnium michelsonii Peganum garmala L. 35-40% 20-25% . ! 20 . R Tortula tortuosa Parmelia vagans . H , ! , ! . T , !! Limnium michelsonii Peganum garmala. . T . c . H 4- ! . I ! ! Krascheninnikovia ceratoides (L.) Gueldenst., Karagana balchaschensis (Kom.)Pojark., Spiraea hypericifolia L. 42-56 ; II ! - Peganum garmala, Suaeda physophora Pall., Artemisia sublessingiana, Nitraria shoberi L. 23-28 , III ! – Anabasis cretacaea, Kochia prostrata, Salsola orientalis S.G.Gmel. 15-20 ; IV ! – Salsola nitraria Pall., Salsola foliosa (L.) Schrad., Nanophyton erinaceum 5-10 . H 10 1 . . U , Peganum garmala – 168 Limnium michelsonii- 152 . S Stipa caucasica- 83 Artemisia sublessingiana - 82 , Orostachys spinosa- 47 , Salsola australis – 39 . W !: Flyssum turkestanicum var. desertorum (Stapf)Botsch. (= A.desertorum Stapf) - 14 ., Eragrostis minor Host – 10 ., Eremopyrum triticeum - 8 ., Ceratocarpus utriculosus – 6 ., Bromus oxydon Schrenk – 2 ., Nanophyton erinaceum - 2 , Goniolimon cuspidatum – 2 ., Draba nemerosa L.– 2 . U Limnium michelsonii !! !. 152 Limnium michelsonii, 10 96 ! , 33 – , 23 – . Z, , , Limnium michelsonii . H SY. . 1 (33). 2012 275

- - - F, . ! 40-450 . Y ! 1294 .., GPS : N 43027.664' E 078047.618'. U – . H ! . U ! 60-65%, ! 70%. Y – . T, ! !. M ! , , 14-15 !. U , !. H ! . U !. J !! Artemisia sublessingiana (25%), Limnium michelsonii (20%) !: Suaeda altissima (R.)Pall., Suaeda physophora (15%), Salsola australis (10%). K 50. H 5 ! : I ! !: Achnaterum splendens (Trin.) Nevski, Tamarix hohenackeri Bunge 140-170 ; II ! – Agropyron sp., Allium galanthum Kar. et Kir., Krascheninnikovia ceratoides, Karagana balchaschensis, Nitraria shoberi 60-70 ; III ! - Stipa capillata L., Atraphaxis spinosa L. 40-55 ; IV ! - Limnium michelsonii, Suaeda physophora, Atriplex sp., Stipa caucasica 20-35 ; V ! - Salsola australis, Salsola foliosa, Petrosimonia sibirica (Pall.)Bunge - 5-10 . , ! . U , Suaeda altissima – 423 , Artemisia sublessingiana - 106 , Salsola foliosa – 61 . W : Stipa caucasica – 20 ., Krascheninnikovia ceratoides -5 ., Agropyron cristatum (L.)Beauv. – 5 ., Allium galanthum – 5 ., Petrosimonia sibirica – 5 ., Kochia prostrata – 2 . R , ! Limnium michelsonii, Nanophyton erinaceum, Anabasis cretacaea, Ziziphora tenuior . S 10 Limnium michelsonii ! 146 ., – 102 ., – 32 ., - 12 . Z, , , Limnium michelsonii . X , 30 , ! ! 391 Limnium michelsonii, – 261, – 86, – 44, , , , . G K F , H X!-^!: M F, G X. Y ! 1108-1141 .!, GPS : N 43020.723' E 078031.593'. V ! - , . W . U - !, (30 80 ) (-). U . H ! ! . U Artemisia lessingiana Bess. Y - . U ! 35-40%. H Artemisia sublessingian, Salsola foliosa, Salsola orientalis, Nanophyton erinaceum. U ! Nitraria shoberi, Atraphaxis virgata (Regel)Krasn., Pentaphylloides phyllocalyx (Juz.)Sojak (= Dasiphora phyllocalyx Juz.). H 3 ! : I ! ! Limnium michelsonii Artemisia sublessingiana 17-20 ; II ! - Salsola orientalis, Aristida heymannii Regel (=A.adsensionis L.) 10-15 ; III ! - Salsola foliosa, Nanophyton erinaceum 3-5 . V , ! Limnium michelsonii – 145 , 4 , – 12 , – 43 , – 50, – 32, - 3 , - 1 . U ! !. c , !! ! ! Limnium michelsonii. - 1141 .!, GPS : N 43021.287' E 078032.150'. M 10 1 . . U ! 60-65%, 276 Y . . 1 (33). 2012

! 70%. H Artemisia sublessingiana, Stipa caucasica, Ceratocarpus utriculosus, Flyssum turkestanicum var. desertorum (= A.desertorum), Salsola foliosa, Orostachys spinosa . H! ! 2 !: I ! ! Artemisia sublessingiana, Limnium michelsonii, Aristida heymannii (=A.adsensions) 15-20 ; II ! - Salsola foliosa, Flyssum turkestanicum var. desertorum, Ceratocarpus utriculosus, Orostachys spinosa 5-10 . H Limnium michelsonii ! , Limnium michelsonii ! . S . U , ! Nanophyton erinaceum – 476 , Stipa caucasica – 197 , - Aristida heymannii – 96 . T : Artemisia sublessingiana – 15 , Orostachys spinosa – 27 , Puccinelia s.- 28 , Draba sp. - 1 . ] ! Limnium michelsonii, 10 ! 111 ., – 2, – 36, – 66, – 7, , , . -- - . U -, . Y ! 1108 .!, GPS : N 43021.277' E 078032.178'. U ! 65-70%. H : Artemisia sublessingiana, Aristida adsensions, Orostachys spinosa, Eragrostis minor . , ! ! ! 2 !: I ! ! Limnium michelsonii, Aristida heymannii Artemisia sublessingiana 15-20 ; II ! - Nanophyton erinaceum, Flyssum turkestanicum var. desertorum, Orostachys spinosa 3-5 . M Limnium michelsonii ! 40-45% . S 10 578 Limnium michelsonii, – 35 , – 100 , – 73 , – 229, – 85, -12 , – 35 – 9 . X , !! Limnium michelsonii, ! ! . c , ! - ! ! ! () ! ! Limnium michelsonii. Orostachys spinosa – 442 , Nanophyton erinaceum – 147 , Puccinelia s. – 125 . W : Salsola foliosa – 71 ., Aristida adsensions – 13 ., Stipa caucasica – 9 ., Flyssum turkestanicum var. desertorum – 5 ., Artemisia juncacea Kar. et Kir.– 3 . X , 25 ! Limnium michelsonii ! 834 , – 345, – 152, – 124, – 114, 39, - 38, – 12 - 10. F! Limnium michelsonii , ! G . H ! !!, , . Limnium michelsonii ! X F-S. Y ! 1301-1320 .!, GPS : N 43020.630' E 078047.864'. M . U - . T . U -!, !, !. Y ! 1301 .!, GPS : N 43020.630' E 078047.864'. U ! 65-70%. J !! Nanophyton erinaceum Limnium michelsonii. H Artemisia sublessingiana, Stipa caucasica, Stipa kirghisorum P.Smirn., Poa bulbosa L., Trigonella arcuata C.A.Mey., Orostachys spinosa, Kochia prostrata . U , 35-49% ! Nanophyton erinaceum, Limnium michelsonii 25-30%. . H 3 !: I ! ! Artemisia sublessingiana 25 ; II ! - Kochia prostrata, Climacoptera lanata (Pall.)Botch., Limnium michelsonii, Poa bulbosa 10-15 ; III ! - Stipa kirghisorum, Trigonella arcuata, Goniolimon cuspidatum, Orostachys spinosa 5-7 . M 5 1 .. Limnium michelsonii ! 10 , – 9, – 1, H SY. . 1 (33). 2012 277

, , , , . - --. Y 1320 .!, GPS : N 43020.783' E 078055.484'. V ! X. U - !, !. U ! 70-75%. H : Stipa caucasica, Stipa kirghisorum, , Agropyron pectinatum (Bieb.)Beauv. (=A.pectiniforme Roem. et Shult.), Goniolimon cuspidatum, Orostachys spinosa . H ! : I ! ! Camphorosma lessingii Litv., Agropyron pectinatum, Artemisia sublessingiana 20-25 ; II ! ! Stipa caucasica, Stipa kirghisorum, Anisantha tectorum (L.)Nevski (=Bromus tectorum L.) 10-15 ; III ! - Nanophyton erinaceum, Goniolimon cuspidatum, Trigonella arcuata, Orostachys spinosa 5-7 . M 10 1 .. S Artemisia sublessingiana - 828 , Nanophyton erinaceum - 152 , Camphorosma lessingii - 1791 , Stipa caucasica - 226 , Agropyron pectinatum - 168 , Carex sp. - 26 , Orostachys spinosa - 10 , Kochia prostrata - 2, Flyssum turkestanicum var. desertorum - 1 , Anisantha tectorum - 10 . Limnium michelsonii ! 55 , -11, – 19, – 10, - 3, – 2, – 1, . J ! Limnium michelsonii. , , . -. Y ! 1301 .!, GPS : N 43020.630' E 078047.864'. U -, . U ! 60-65%., ! 70%. Y ! X. H : Krascheninnikovia ceratoides, Stipa kirghisorum, Lagochilus leiacanthus Fisch. et C.A.Mey., Kochia prostrata . H ! : I ! ! Artemisia sublessingiana, Camphorosma lessingii, Stipa caucasica 20-30 ; II ! ! Dianthus hoeltzeri C.Winkl., Agropyron pectinatum 15-20 ; III ! - Kochia prostrata, Lagochilus leiacanthus 10-15 ; IV ! - Nanophyton erinaceum, Orostachys spinosa, Ziziphora tenuior 3-5 . H 25 1 .. S Nanophyton erinaceum - 2044 , Artemisia sublessingiana - 1546 , Orostachys spinosa - 763 , Stipa caucasica - 212 , Eremopyrum triticeum - 109 , Petrosimonia sibirica - 438 , Trigonella arcuata - 17 , Kochia prostrata - 19 , Koeleria cristata (L.)Pers.(= K. gracilis Pers.) - 7 , Poa bulbosa - 8 , Meniocus linifolius (Steph.)DC - 6 , Salsola foliosa - 5 , Flyssum turkestanicum var. desertorum - 3 , Puccinella diffusa V.Krecz - 4 , Sedum sp. - 2 , Goniolimon cuspidatum - 2 , Ceratocarpus arenarius L. - 1 . Limnium michelsonii ! 270 , -1, – 9, -34, – 122, – 89, - 11, – 4, . S 1 , !! Limnium michelsonii !! 2 3. U Limnium michelsonii 1 ! 2, ! 3. W 1 ! 2 !. S Limnium michelsonii 1 . S () Limnium michelsonii , ! ! ! 1 !! 2, 3 , !! 2 1, 3 , , . R !! 1 3 ! Limnium michelsonii (, ). J ! !. S ! Limnium michelsonii. X , ! (! !) , !! ! Limonium michelsonii , ! ! . U ! . Limonium michelsonii , !, - ! . U , . T . V ! !! Limonium michelsonii ! , ! 278 Y . . 1 (33). 2012

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1. Limnium michelsonii 1 ( %). X 1 Limnium michelsonii 3 H, J. J. - - - ! - , .-., , ., . . . . 1 7.8±0.18 12.63±0.35 12.83±0.36 6.28±0.21 3.41±0.14 3.56±0.15 2 5,75 ±0,13 6,98 ± 0,17 6,93±0,17 4,36±0,11 2,26±0,07 2,88±0,08 3 5,77±0,15 11,6±0,37 11,42±0,37 6,19±0,22 3,57±0,15 3,38±0,14 W ! ! [11] ! ! 2 3 !! ( ! ). U ! Limnium michelsonii , ! V , ! , , . Q 1. R.H. Iris setosa Pall. ex Link. V J H // F..... H. - 2000, 28 . 2. Prentise H.S. Enzyme polymorphism morphometric variation and population structure in a restricted endemic Silene diclinis (Caryophillacece) H.S. Prentise // Biol. J.Linm.Soc. – 1989. - 32. - Vol.22.- P.125-143 3. Dalan R. Patterns of isozyme variation in relation to population size, solation and phytogeographic history in royal catchgly (Silene regia Caryophyllaceae). R.Dalan // Amer.J.Bot.- 1994. - Vol.81. - P.965-972 4. Ouborg N. The Significance of genetic erosion in the process of extinction. IV inbreeding Load and heterosis in relation to population Size in the mint Salvia pratensis. N. Ouborg, // R.Van Treuren Evolution. -1994. – Vol.48. - 34. - P.996-1008 5. L! I.I. \! ! ]. - A. . ... – J. -1981. - 26 . 6. I S.H. T !. // c!. -1983.- 31.- W. 310 H SY. . 1 (33). 2012 279

7. F.W. W! ! Luzula sylvatica (Huols) Caudin . - F. . . .. – J. – 1983. - 24 . 8. ! WWV. ] 2. V!. - F-F.- 1981. -284 . 9. U U V 21.06.2007 . 3521 10. V X.F. L ! // X.GS FS WWWV. R.-Q.: FS WWWV. I.- 1950.-.3. -. 6. - W.7-204 11. Y F.F. H ! ! // G.. – 1975. - 32. - W. 7-33 *** = F ''' (G , X , G ) ', ' ' % Limoniun michelsonii ''''' 3 (9 ) ', %' '''' 9 , '', ' . $ 25- 40- ' '' 1 2 '' , % % ' . ' Limoniun michelsonii ''''' ' '' , ' . ' Limoniun michelsonii ' ''' '''. M %' L. michelsonii ''''' '' '' . L. michelsonii '' % ''. L. michelsonii ' 2 % 3-' '. 2-' 1-' L. michelsonii ''''' , ' , . L. michelsonii ''''' ' ' ''', ' ''' ''' '. L'' %' %' L. michelsonii '''' ' % ' ' %'' '' '. *** Three populations (9 of cenopopulations) a rare and relict species close endemic species Limonium michelsonii were investigated in the eastern part of the Trans-Ili Alatau (mountain Bogutty, hole Bartogay, mountains Toraigyr). Geobotanical descriptions of nine cenopopulations, the characteristics of soil and moisture were made as a result of the study. From 25 to 40 accounting areas were put in each population, the number of individuals of dominant and subdominant species were counted on them. Individuals of Limonium michelsonii by their age composition were pick out on each accounting areas, a full calculation of them with distribution by age categories was taken. Also, some measurements of biometric parameters of individuals of Limonium michelsonii were held on the accounting areas. Areal and area of distribution of this species have been found negligible in the study, also Limonium michelsonii prefers the loosened soil, areas with wind and water erosion. Age states of Limonium michelsonii most fully were represented in populations 2 and 3. There were the largest number of individuals of Limonium michelsonii on average on one accounting area in population 2 and the smallest number of individuals - in the population 3. Reproduction of Limonium michelsonii in three populations studied is the natural way - the seeds. Overgrazing is a great danger for the populations of Limonium michelsonii, because it does not tolerate trampling and grazing. The results, that were obtained in research, can be used to work out preservation measures of Limonium michelsonii, creating a database for the new release of the Red Book of Kazakhstan, etc.

YJ 581.9 R. ., . I. , M. . , . . ROO U ZQT MI – TITIT R. .-, e-mail: [email protected] -G . 85 , 10 61,31 % . . Asteraceae, 129 , 52 . !!! ! . T ! ! , " ! - !, ! . U ! . !! ! ! ! !! . U F (W -G ) . c ! (2,5 .²) ! ! , ! . , - . ^, - . ^ X - . [2, 3 ]. T, W - G ! - F (M F). J! W -G . X! ! - F, . W ! , ! ! ! , -! . W -G , ! , . X! ! , ! ! [4]. 280 Y . . 1 (33). 2012

! ! ! , . H ! ! ! ! W , . X. I! , ! -! - 1850 ( W), - -! - 500 ( , ). M! ( ), ( ), ! , ! !, . M , ! ! -! , !! ! W, W , G-G, Y- G. U , ! G- Y-G, ! , ! , - (! ); ! . R W X ! ! W ! . T ! ! ( , G), ! .]. ! . S , ! . H W -G : , -, -, , - [5]. U ! W -G 1983 1984 . 2011 . T -. M ! 1500 !. U « WWWV» [6], « » [7], «T W F» [7], « » [9], «M WWWV» [10], « » (2011) [11]. [12, 13]. U G . H.Q. FS V (W-U), RTS V (F) ! -. : «W F-F I » [14], « » [9], « » [7], «T W F» [8]. V F.Q. X! [15]. S , ! W.. ] [16]. H W -G ! 85 c: Pottiaceae Schimp., Grimmiaceae Arnott., Bryaceae Schwaegr., Leskeaceae Schimp., Brachytheciaceae Roth., Athrie F1st., Equisetaceae Rich. Ex DC.,Cupressaceae Bortl., Ephedraceae Dum., Ranunculaceae Juss.; Berberidaceae Juss.; Papaveraceae Juss.; Hypecoaceae Nakai; Fumariaceae DC; Caryophyllaceae Juss.; Amaranthaceae Juss.; Chenopodiaceae Vent.; Polygonaceae Juss.; Limoniaceae Lincz. (Plumbaginaceae Juss.); Hypericaceae Juss.; Primulaceae Vent.; Tamaricaceae Link; Frankeniaceae S.F. Gray.; Aceraceae Juss., Salicaceae Mirb.; Capparidaceae Lindl.; Brassicaceae Burnett.; Malvaceae Juss.; Ulmaceae Mirb.; Cannabaceae Endl., Urticaceae Juss., Euphorbiaceae, Crassulaceae DC., Grossulariaceae DC., Rosaceae Juss.,., Onagraceae Juss., , Fabaceae Lindl., Rutaceae Juss., Peganaceae Tiegh., Geraniaceae Juss., Cynomoriaceae Lindl., Elaeagnaceae Juss., Apiaceae Lindl., Caprifoliaceae Juss., Valerianaceae Batsch., Dipsacaceae Juss., Rubiaceae Juss., Gentianaceae Juss., Apocynaceae Juss., Asclepiadaceae R. Br., Solanaceae Juss. , Convolvulaceae Juss. . S !! ! 16 . W (129 , 52 ) !!! Asteraceae, Fabaceae (66 17 ), - Poaceae, (63 34 ), - Brassicaceae (62 37 ). J Rosaceae, 41 12 . H Boraginaceae ! 36 15 , Lamiaceae - 32 18 , Chenopodiaceae - 29 19 , Caryophyllaceae - 28 18 Scrophulariaceae - 26 7 . H Apiaceae 25 , Ranunculaceae - 18, Liliaceae - 16, Alliaceae - 14, r Rubiace - 11 . X ! 10 ! ! , ! . H 10 W -G . 512 H SY. . 1 (33). 2012 281

. 10 61,31 % . 323 75 , 30 . 9 10. X ] 10 - W % Asteraceae Dumort. 129 15,45 Fabaceae Lindl. 66 7,90 Poaceae Barnhart. 63 7,54 Brassicaceae Burnett. 62 7,43 Rosaceae Juss. 41 4,91 Boraginaceae Juss. 36 4,31 Lamiaceae Lindl. 32 3,83 Chenopodiaceae Vent. 29 3,47 Caryophyllaceae Juss. 28 3,35 Scrophulariaceae Juss. 26 3,12 512 61,31 X , W - G 85 , 10 ! 61,31% !. Q 1. X R.W. G – ! W -G ( ). – . . ... . . . - X., 1987. – 20 . 2. U R.I. H ! M F // H .: V . - ].2.- 1941. - .216. 3. G G.F. T ! ! . // H FS WWV. -1954. - 3 2. - W.15-20. 4. X K. T!! K. - R.: - 1966. - 76 . 5. W W.., F .F., F.G. U WWV(F-F! ). - F-F, - 1962. - 3 4. - W.424. 6. WWWV. - R-Q.: - 1934-1964. - X. 1 – W. 30. 7. . – F – F: S, - 1956-1967. - X.X. 1-9. 8. T W F. – X: FS. - 1968-1996. - X.X. 1-10. 9. . – F-F: S. -1969-1972. - X.1-2. 10. \ S.S. M WWWV. - Q.: S. - 1976. - 788 . 11. G R.W. . – F: Ĩ. - 2001. - X. 1-2. 12. R.W., K.F. V. R.: - 2003. - X.1. - W.1-608. 13. R Y.. X WWV. // W- I. – J: - 1990. - 236 . 14. U R.I. F-F . – F-F: 1940. – 50 . 15. X! F.Q. W . - Q.: S. - 1987. – 439 . 16. ] W.. W ! WWWV. - Q.: S. - 1981. - 509 *** R ''-G'' '' ' . X ' %' 85 ', '' 10 ' , ' 61,31%- . 52 129 Asteraceae ' . *** In article is resulted analyzes the spectrum of familial flora Syugaty-Bogutinskogo low-mountain massif. A study of familial spectrum recorded 85 families, of which 10 leading family make 61,31 % from the flora of region. The leading place is occupied by the family Asteraceae, which includes 129 species belonging to 52 general.

YJ 595.76 .. RQ UT Z TJ LTQ (INSECTA, COLEOPTERA) ITJTIT \TQbTIT UTJTIT U «Q-RQb» VIU « » S RTS V, e-mail: [email protected]

ISUU «F-». 69 , 11 . 1 - Deronectes latus Stephens P . Coleoptera U ! ! [1-12], - ! . . R !! ! !. S !, !! ! . H 2010-2011 . ! ! U I -S U U «F-c» , ! . , W.H. V.. 282 Y . . 1 (33). 2012

. W ! . S !! . W Dytiscidae Leach 1815 – U 1. Agabus (Gaurodytes) basalis (Gebler, 1829) – I . R: ], X, 3.06.2010, , 10 W, 9 V; ], -F, 4-5.06.2010, , 6 W, 10 V, .. X. 2. A. (G.) dichrous (Sharp, 1878) - I. R: ], X, 3.06.2010, , 1 W; ], -F, 4-5.06.2010, , 3 W, 2 V, .. X. 3. Ilybius ater (De Geer, 1774) – X . R: U , Y , . , 7.06.2010, , 1 W, .. X. 4. I. cinctus Sharp, 1878 – X . R: U , Y , . , 7.06.2010, , 3 W, 1 V, .. X. 5. Rhantus exoletus Forster, 1771 - . R: U , Y , . , 7.06.2010, , 1W, 1V; ], -F, 6.06.2010, , 1 W, .. X. 6. Rh. frontalis (Marsham, 1802) – . R: U , Y , . , 7.06.2010, , 1 W, .. X. 7. Rh. suturalis (MacLeay, 1825) (=Rh. pulverosus Stephens, 1828) - !. R: U , Y , . , 8.06.2010, , 1 W; ], -F, 5.06.2010, , 2 W, .. X; . . , 29.05.2011., , 2 W, V.. . 8. Colymbetes dolabratus (Paykull, 1798) – U (). R: . . , 29.05.2011., , 1 W, V.. . 9. C. semenowi Jakovlev, 1896 – U W. R: R: , 17.06.2011., , 1 W, W.H. . 10. Deronectes abnormicollis Semenov, 1900 – J ! !. R: ], -F, 3.06.2010, , 2 W, .. X. 11. Deronectes latus Stephens, 1928. - J . R: ], X, 3.06.2010, , 2 W, 4 V, .. X. 12. Hydroporus angustatus Sturm, 1835 – S! !. R: U , Y , . , 8.06.2010, , 1 W, .. X. 13. H. goldschmidti Gschwentdner, 1923 – S! I. R: ], -F, 3.06.2010, , 1 W, .. X. 14. Hydroporus nigrita (Fabricius, 1792) – S! !. R: U , Y , . , 8.06.2010, , 1 W, .. X. 15. Hydroporus pubescens Gyllenhall, 1808 – S! !. R: ], -F, 4.06.2010, , 1 W, 2 V, .. X. 16. Nebrioporus airumlus (Kolenati, 1845) (=Potamonectes airumlus Kolenati, 1845) – V . R: ], X, 10.07.2011, , 1 W, 1 V; , 11.07.2011, , 2 W, .. X. 17. Hygrotus caspius (Wencke, 1873) – U !. R: ], -F, 3.06.2010, , 1 W; ], X, 11.07.2011, , 3 W, .. X. 18. H. enneagrammus (Ahrens, 1833) – U !!. R: U , Y , . , 8.06.2010, , 3 W, 2 V, .. X. 19. H. impressopunctatus (Schaller, 1783) - U !. R: U , Y , . , 8.06.2010, , 2 W, .. X. 20. H. lernaeus Schaum, 1857 – U !. R: U , Y , . , 8.06.2010, , 1 W, .. X. 21. Hydroglyphus geminus (Fabricius, 1792) – S . R: ], -F, 5.06.2010, , 1 W, .. X; U , Y , . , 7.06.2010, , 1 W, 2 V, .. X; 8.06.2010, , 2 W, 2 V, .. X. 22. Hyphydrus ovatus Linnaeus, 1761 - U. R: ], X, 11.07.2011, , 1 W, .. X. 23. Acilius canaliculatus Nicolai, 1822 – U . R: U , Y , . , 3.06.2010, , 2 W, 1V, .. X; , 8.06.2010, , 1 W, 1V, .. X. 24. Graphoderus cinereus (Linnaeus, 1758) - U . R: . . , 29.05.2011., , 1 W, 7 V, V.. . 25. Cybister lateralimarginalis (De Geer, 1774) – W . R: U , Y , . , 3.06.2010, , 1 W, 1V, .. X; 26. Hydaticus grammicus Germar, 1830 – G !. R: U , Y , . , 3.06.2010, , 2 W, 3 V, .. X. W Gyrinidae Latreille, 1810 – H! 1. Aulonogyrus concinnus Klug, 1834 – H! !!. R: ], X, 10.07.2011, , 1 W; , 11.07.2011, , 1 W, 1 V, .. X. H SY. . 1 (33). 2012 283

2. Gyrinus paykulli Ochs, 1927 – H! U!. R: ], -F, 4.06.2010, , 1 W, .. X; U , Y , . , 8.06.2010, , 2 W, 2 V; . , 27.05.2011, , 2 W, 2 V, V.. ; ], -F, 8.07.2011, , 1 W; , 9.07.2011, , 1 V, .. X; ], X, 10.07.2011, 10 W, 13 V, , .. X, W.H. ; , 11.07.2011, 5 W, 4 V, , .. X. 3. Gyrinus distinctus Aube, 1838 – H! !. R: , 17.06.2011, , 1 W, W.H. ; , , 18.06.2011, 2 W, 2 V, .. X; ], -F, 8.07.2011, , 1 W, .. X; , 9.07.2011, , 1 W; ], X, 10.07.2011, 1 W, 1 V, , .. X; , 11.07.2011, 2 W, 4 V, , .. X. W Spercheidae Erichson, 1837 1. Spercheus emarginatus Schaller, 1783 – W. R: U , Y , . , 8.06.2010, , 2 W, 1 V, .. X. W Hydrophilidae Latreille, 1802 – H 1. Berosus spinosus (Steven, 1808) - U . R: U , Y , . , 8.06.2010, , 2 W, 4 V, X ..; , 8.06.2010, , 2 W, 2 V, .. X. 2. Laccobius minutus Linneus, 1758 – G . R: ], X, 10.07.2011, 3 W, 5 V, ; , 11.07.2011, 5 W, 4 V, , X .. 3. Laccobius colon Stephens, 1829 – G !. R: ], X, 10.07.2011, 2 W, 3 V, ; , 11.07.2011, 1W, 2 V, , .. X. 4. Laccobius gracilis Motschulsky, 1855 – G . R: ], X, 10.07.2011, 1 W, , .. X. 5. Laccobius striatulus Fabricius, 1801 – G . R: ], X, 9.07.2011, 2 W, , 4 V, .. X. 6. Hydrobius fuscipes Linnaeus, 1758 – H . R: ], X, 10.07.2011, 1 W , .. X; W, . L, 29.05.2011., , 1 W, 1 V, V.. ; . . , 29.05.2011., , 1 W, 3 V, V.. . 7. Enochrus bicolor Fabricius, 1792 – X . R: U , Y , . , 8.06.2010, , 3 W, 4 V, .. X. 8. Enochrus quadripunctatus Herbst, 1797. – X !. R: U , Y , . , 7.06.2010, , 4 W, 7 V; , 8.06.2010, , 2 W, 5 V, .. X; . . , 29.05.2011., , 2 W, V.. . 9. Helochares obscurus Mueller, 1776 – T . R: . . , 29.05.2011., , 1 W, 2 V, V.. . 10. Hydrophilus piceus Linnaeus, 1758 – H , . R: U , Y , . , , 1 W, 7.06.2010., .. X. 11. H. aterrimus Eschscholtz, 1822 – H , !-. R: U , Y , . , , 2 V, 6.06.2010., .. X. 12. Hydrochara dichroma Fairmair, 1892 – H . R: U , Y , . , , 1 V, 6.06.2010., .. X. 13. Chaetarthria seminulum Herbst, 1797 – ! !!. R: ], X, 11.07.2011, 4 W, 2 V, , .. X. 14. Cryptopleurum subtile Sharp, 1884. R: U G, Y , 7.06.2010, , 1 W, 2 V, .. X; ], X, 10.07.2011, , 5 W, 7 V, .. X. 15. C. minutum Fabricius, 1775 - . R: U G, Y , 7.06.2010, , 2 W, 2 V, .. X; ], X, 10.07.2011, , 5 W, 5 V, .. X. 16. Sphaeridium substriatum Faldermann, 1838 – ^ !. R: ], X, 10.07.2011, , 1 V, .. X. 17. Coelostoma orbiculare Fabricius, 1775 - ]. R: U G, Y , 7.06.2010, , 2 V, .. X. W Hydraenidae Thomson, 1859 - H 1. Ochthebius minimus Fabricius, 1852 (=O. impressus Marsham, 1802) - U . R: U , Y , . , 8.06.2010, , 2 W, 2 V, .. X. 2. Ochthebius pusillus Stephens, 1835 – U . R: ], X, 11.07.2011, 1 W, , .. X.

W Helophoridae Leach, 1815 - R 1. Helophorus kayae Angus, 1985 – R !. R: ], X, 10.07.2011, 1 W, 1 V , .. X. 2. Helophorus similis Kuwert, 1887 – R . R: ], X, 11.07.2011, 1 W, , .. X. W Elmidae Curtis, 1830 284 Y . . 1 (33). 2012

1. Potamophilus acuminatus Fabricius, 1792. – U R: U , Y , . , , 1 V, 6.06.2010., .. X W Dryopidae Billberg, 1820 - U 1. Dryops caspius Menetries, 1832 – U . R: U , Y , . , , 1 V, 6.06.2010., .. X; . . , 29.05.2011., , 1 W, V.. W Heteroceridae MacLeay, 1825 – U 1. Augyles intermedius Kiesenwetter, 1843 – U . R: . . , 29.05.2011., , 1 W, V.. ; ], X, 10.07.2011, , 3 W, 4 V, .. X. 2. Augyles hispidulus Kiesenwetter, 1843 - . R: ], X, 10.07.2011, , 1 V, W.H. . 3. Heterocerus flexuosus Stephens, 1828 – U . R: ], X, 10.07.2011, , 2 W, 2 V, .. X. 4. Heterocerus marginatus Fabricius, 1787 – U . R: . . , 22.05.2011., , 2 W, 2 V, V.. ; ], X, 10.07.2011, , 4 W, 6 V, .. X. 5. Heterocerus obsoletus Curtis, 1828 – U . R: . . , 22.05.2011., , 1 W, V.. . 6. Micilus minutissimus Sahlberg, 1900 – U . R: U , Y , . , , 1 V, 2 W, 6.06.2010., .. X; W, . L, 22.05.2011., , 1 W, V.. . W Erirhinidae Schönherr, 1825 - c 1. Tanysphyrus lemnae (Paykull, 1792) – J !, !. R: ], X, 10.07.2011, , 1 W, 1 V, .. X. 2. Grypus equiseti (Fabricius, 1775) – J . R: ], X, 11.07.2011, , 1 W, .. X. W Curculionidae Latreille, 1802 - J 1. Bagous argillaceus Gyllenhal, 1836 – G . R: ], X, 10.07.2011, , 1 W, .. X. H ISUU «F-c» H 69 , !! 2 ! 11 . H ! , ! , . J !!! !! . H ! V . Q 1. F V.G. Helophorus F. (Coleoptera, Hydrophilidae). II. // c . – X. LXIV. – H. 4. – W. 716-747. 2. M .F. U !. WWWV. L. X. IV. - R.-Q., - FS WWWV. - 1953. - 378 . 3. T V . – X. 5. H (, , , , , ) // U . . W..\. - WU.: S, 2001. - 836 . 4. Hansen M. Helophoridae // Catalogue of Palearctic Coleoptera / Eds. I. Lobl and A. Smetana. - V. 2. - Stenstrup, Denmark: Apollo Books, 2004a. - P. 36-41. 5. Hansen M. Spercheidae // Catalogue of Palearctic Coleoptera / Eds. I. Lobl and A. Smetana. - V. 2. - Stenstrup, Denmark: Apollo Books, 2004b. - P. 43-44. 6. Hansen M. Hydrophilidae // Catalogue of Palearctic Coleoptera / Eds. I. Lobl and A. Smetana. - V. 2. - Stenstrup, Denmark: Apollo Books, 2004c. - P. 44-68. 7. Jach M.A. Hydraenidae // Catalogue of Palearctic Coleoptera / Eds. I. Lobl and A. Smetana. - V. 2. - Stenstrup, Denmark: Apollo Books, 2004. – P. 102-122. 8. Kodada J., Jach M.A. Dryopidae // Catalogue of Palearctic Coleoptera / Eds. I. Lobl and A. Smetana. - V. 3. - Stenstrup, Denmark: Apollo Books, 2006. - P. 440-443. 9. Mascagni A. Heteroceridae // Catalogue of Palearctic Coleoptera / Eds. I. Lobl and A. Smetana. - V. 3. - Stenstrup, Denmark: Apollo Books, 2006. - P. 446-449. 10. Mazzoldi P. Gyrinidae // Catalogue of Palearctic Coleoptera / Eds. I. Lobl and A. Smetana. - V. 1. - Stenstrup, Denmark: Apollo Books, 2003a. - P. 26-30. 11. Nilsson A.N. Dytiscidae // Catalogue of Palearctic Coleoptera / Eds. I. Lobl and A. Smetana. - V. 1. - Stenstrup, Denmark: Apollo Books, 2003b. - P. 35-78. 12. Nilsson A.N. Catalogue of Palearctic Dytiscidae (Coleoptera). Internet version 2010-01-01. *** L «F-K» Ri X i. L 9 -ii 69 i 'i. 1 – Deronectes latus Stephens i i. *** In article given materials on faunistic structure of water beetles of the State National Natural Park "Altyn-Emel". All in the studied area recorded 69 species of water beetles belonging to 9 families. 1 species - Deronectes latus Stephens are found for the first time in Kazakhstan.

H SY. . 1 (33). 2012 285

W 4. c

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K ' ''' %' ' ''' '''' ' '' . X Ľ ! Ľ ı . ĳ ı ! ı [1-5]. – , ! (Ľ !), Ľ Ľĳ ĿĽ ĳı Ŀ [6]. X - ı Ŀ Ľ [7-10]. CCl4-ı Ľ ı ı, ı , Ľı Ŀ . [11-13]. Q ĳ ı ı Ŀ , Ľ ! ! Ľ ! . cı , !ı Ŀ , ĳı ĳ Ŀ ı [14]. T ı Ľĳ ĳı ! Ľ Ľ ı ! ! . T Ŀ Ŀ ı ! Ŀ ĳ. $' . M Wistar ĳ Ŀ ĳ (Ľ 12-15 ). Giii - . Kii – ĳ ĳ Ŀ (CCl4) Ľ . CCl4 ĳ 4 ( 0,3 /). K ı . c ! ĳ ĳ Ľ . G ı K ! ii Ľiı . iı, ı, ı Ŀ «Bio-Lachema- Test» . ı Ľ : ı (FQX), ı (FWX), , ı Ŀ . XĽı Ľ cHR – W ı t- ! ! Ľ . K Ľ <0,01, <0,05 . S% % . Mı Ľ, CCl4 Ľ ı Ŀ ı Ŀ Ŀ . ı Ŀ 27,5%-, - 36%- Ŀ. T ı ! 8,3±0,4 5,2±0,3 / – , - 20%, - 26% Ŀ. Q Ŀ Ŀ: Ľ 8,5±0,4 5,4±0,6 / – , – 23% Ľ – 20% (1-). 1- ĭĵ ĭ ĭ ĩ ĭ Ŀ'''ı ĭĩ F 1- 2- 3- R, / 8,3±0,4 5,2 ±0,3** 7,5 ±0,5* , / 64,2±2,1 52,0±1,3* 57,0±1,4 F , % 26,6±2,0 20,8±1,2* 23,5±1,2 Q: R, / 8,5±0,4 5,4±0,6* 6,8±0,6 , / 90,0±1,7 70,2±2,3** 82,7±2,2* F , % 27,0±3,1 21,6 ±1,8* 24,5 ±1,5 K: * - , <0,05,* -<0,01** 286 Y . . 1 (33). 2012

G Ľ ı ! Ŀ , ı , ı ı Ľ FQX FWX . K ı Ŀ , Ľ FQX(310%-) Ľ FWX-ı (257%) ı ĳ , ı ı Ŀ. ı ıi 23% F Ľ Ŀ, CCl4 ı ! ĳ Ŀ : ı ĳ , Ŀı Ľ FWX FQX Ľı . ı ı ı ıı ı . F Ľ ı ! Ŀ Ŀ. ļJ 1. G Z.. U // X cQ. - S, - 2000, - X.8. - W. 5-9. 2. G Q.c., J I.F., F W.S., Q.Y., K I.W. X! ! // H Z- . -2005, -34. –W.55-57. 3. H F.., R.Z., ] H.W., W c.H. . R // W. . . – 2000, - 32, - W. 12-14. 4. K Z.F., U X.H. R ! . - R., R. – 1989, – 14 . 5. M U.. T! ! / . . G. F. !, H.F.. - R., - 2002, – W. 352-384. 6. H.H., F F.R. IFR- // G . . . ., - 1982, - 37, - W. 40-43. 7. .Z., R W.H. H «c» // H .: U . – S, - 2004, - X.1, – W. 180-181. 8. H.W, W - ! . – R., «R - », - 2004, - 920 . 9. R.F., U F.F., G.., F H.Q., W! W.U. S // X 1- R. . « ». – R., 2001, - W. 130-131. 10. Q K.F., Q.I. T !. -R.: R, 1989, - 43 . 11. T I.. . - R.: S, - 1991, - 319 . 12. W K.R., W Q.F., V .J., G I.R. X – .–Y-, -2001,170. 13. Jwai M., Morikowa T., Muramatsu A., Tanaka G. et al. Biological significance of AFP expression in liver injury induced by CCl4 // Acta. Histochem. et Cytochem.2000, - 33, - 31. – P. 17-22. 14. Melin A., Perromat A., Deleris G. The in vivo toxity carbon tetrachloride and carrageenan on heart mikrosomes. Analyses by Fourier transform infrared spectroscopy // Can. J. Physiol. and Pharmacol. – 2001. – 79, - 3 9, – P. 799-804. *** U , l4, . G , . *** The oppression infringement in biochemical content of a lymph have been observed at toxic hepatite, caused by l4 introduction. There was shown increase in cytolytic enzyme activity and tymol tests and decrease in a level of total protein.

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. W ĳ ĳ Ľı Ŀ . T ı ı Ľı Ľ . M Ľ Ŀ [8,9]. F ĳ « ı» ı - , ı Ŀ Ľ . «F ı» ı – . M '' Ľ''. Lı ĳ Ľı « ı» . «F ı» 1934 C.S.Hall . T ı Ŀı ĳ . «F ı» ı ı, ı 100. ^ı 40 ĳ () . ^ıı ĳ ı - 20 16 ! (!ı 3, ı 1). ^ıı ĳ ı ĳ Ŀ: , , !. Ľ ! ı . K ı ı Ŀ ı . K Ŀ ! Ŀ, Ľ . T – , , ı Ŀ ! , ! , , ! ( Ľ ), ı (ı ) XĽı Ľ cHR– W ı t- ! ! Ľ . K Ľ <0,01, <0,05 . M Ľ'. M ! . T Ŀ. Kı ĳ Ľı Ľ , ı «F ı» ı ĳ (1-). G Ľ 1934 W.S Hall ! ĳ . 1- ĩ ĩ Ľ ' ' '' ĭĵ ĭı ĭ L - c () - Ŀ 0.5 1 2 3 4 5 0,5 1 2 3 4 5 X 60 70 30 80 80 70 100 100 50 100 100 100 Ŀ- ı F ! 70 50 45 80 80 80 100 80 80 100 100 100 (%) F - 80 80 70 80 75 80 100 100 100 100 100 100 G 85 80 60 30 40 8 100 100 80 60 60 20 ^ - 35 40 20 9 4 9 60 60 40 40 10 20 - L 80 85 85 90 80 75 100 100 100 100 100 100 R – 85 75 90 85 93 7 100 100 100 100 100 16 ! W 40 40 10 9 0 0 60 60 20 20 0 0 Ŀ X Ŀ 75 80 70 60 75 70 100 100 100 100 100 100 Ŀ W Ŀ 60 40 45 30 10 9 80 70 70 70 20 20 Ŀ R± m 25,05±0, 7 24,8±0,9 24,8±0,9 5,8±0,78 23,6±0,52 23,6±0,52 21,9±0,32 15,4±0,83 10,9±0,42 K: Ľı ı Ŀ -*00,02; **00,001. 288 Y . . 1 (33). 2012

«F ı» J.. H!ı Ľ Ŀı – ĳ Ŀ, ! ı, , ! ! , , Ŀ . XĽ ı – Ŀ ı . M ĳ Ľ ı Ľ ĳ. W ĳ Ŀ ı !- ĳ Ŀ . ı Ľ ı ı Ŀ 25,05±0,7 . F ı Ľ ĳ, - Ŀ . H.U. Uı Ľ « ı» ı –ı , ! !, ! ĳ, ! ĳ . R–ı Ŀ 24,8±0,9; ĳ Ŀ 23,6±0,52; ĳ ĳ Ŀ 21,9±0,32; ĳ Ŀ 5,8±0,78 . , Ľı - ĳ - ı . ļJ 1. Z.W., F.H. "c ! ", R U!, 2000, 34, . 99-107 2. Z.W., F.H. "U ", X , 2001, 32, . 84-89 3. I W. R-! . R.: U, 2000. 460 . 4. R ! ! !. : V!- Y(, 2003. 158 . 5. F! R. I. . V ! ! // L. !. 1980. X. 30. 3 5. 904 . 6. G S.I. J ! !! ! : . ... . . . !, 1984. 180. 7. J! R.F. W // W : VI R . R., 2001. 8. R F. Q. ! " !" // L. !. 1981. X.31. N2. W.301-307. 9. O.Karen B. Friend, Paul F. Malloy and Holly A. Sindelar The effects of chronic nicotine and alcohol use on neurocognitive function. Addictive Behaviors Vol. 30. Iss. 1 . January 2005. P. 193-202. *** F ! !! , !, ! !. *** Analysis of this material shows the experimental animals enterosorbents consumption is not only a quantitative change in the state centers that regulate movement, but also qualitative changes in the structure of behavioral patterns.

TL 612;591.1.57.034 .. , .. , L. L, . Q Q IU M=JI= LO QRZ ĲO=J==İ LQ QJ=Q=I= M -

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G ı . X ! ĳ Ľ . Lı Ľ Ľ ı ĳı Ŀ Ŀ- , Ľ ĳı . Kı ĳı - . S-7,4, +37W : 95% T2 Ľ 5% WT2. Q ĳı ! : -, - Ľ -ı 10-8R-10-3R !. Q ĳı ĳ S339 Ľ S 3012 . XĽı Ľ cHR– W ı t-! ! Ľ . M %'. M , ı 15- , 15 WWl4- 340 ! Ľ 90 ĳı Ŀ ĳ. F Ľ 2 ı 40% Ŀ . WWl4- , . Kı Ľ ĳ Ŀ ı ı 10±0,3, 2- 12±0,4 . M Ľ WWl4 ı 44% ( 0,32±0,04 /). Y ! ĳı Ŀ Ŀ. G ĳı Ŀ . G ĳı 5,0±0,1 Ŀ 4,4±0,1 – , 2,0±0,3 Ŀ 1,8±0,1 – Ľ . R ĳı Ŀ Ŀ . XĿ Ŀ ĳı ı Ľ ĳ ! . Kı Ľ Ľ ĳı . G ĳ ı 3,8±0,4 / Ľ , 6,8±0,3, ĳ 5,0±0,2 / Ľ – 7,2±0,7 (1-).

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G: I – , II – , : /., . F : 1 – , 2 – . 1-. ĭ Ľ '' ĭĵ ĭı ĭ ĳ '''ı ''' Ľ 290 Y . . 1 (33). 2012

Y ĳ Ľı 70%- . XĽı 20% - ! . X 10% . ^ ĳı 1,3±0,2 / Ľ – 1,2±0,3 ı. Xĳ ı Ľ !. F ı (10-8- 10-3R) ı ı ĳ Ľ - 47±1,4%, - 29±1,0%- . F! ! (10-8-10-3R) . T ĳı ı 47±1,4%, ı 29±1,0% - . Iı ĳ Ľ ı - 32±1,2% , 27±0,9%- . XĿ Ŀı Ľ Ľ ! !ı Ľ Ŀ . F Ľ (10-9-10-3R) Ľı 33%- ĳ ! , Ľ (10-9-10-3R) - 28%, ı Ľ (10-9-10-3R) – 30%- . G Ľ ! ĳ (2-).

G: 1 – , 2 – Ŀ Ŀ . F – (1×10-6R), F - (1×10-6R), I - (1×10-6R). ĳ Ŀ. Tı . 2–. ĭ Ľ '' ĭ ĳ '''ı H ĳı ! Ŀ. G Ŀ 60-65% Ŀ. H ı ! ĳı ! Ľı Ŀ . , Ľı 2-5% . ! ı ĳ Ľ ı . X ı - Ľ !ı ĳ Ľ ĳı Ŀ ! ĳ: -ı , !-ı , R-ı Ľ S1-ı (10-6-10-5R) . Fı Ľ (10-7-10-4R) ĳ (10-6-10-5R) ĳı . T (10-6-10-5R) Ľ ı 10-7-10-4R ! Ľ ĳı , Ŀ. Xĳı ıı ı 10-6-10-5R ! Ľ . F Ľ ı ĳı , Ŀ (3-). Kı ĳ ı ı Ŀ (10-6-10-4R) (10-7-10-3R) Ľ Ľı 70% - . S1 – ĳı ĳ ı ! (3-). Y ı ĳı , ı ı Ľ ĳı H SY. . 1 (33). 2012 291

ĳ ! (10-5-10-4R) . G ı Ľ ĳı ı Ŀ . W , , Ľ -ı , R- S1-ı -, ı ! ı ĳ ĳı Ľ . G Ľ Ľ [7], ı CCl4 ĳı . F Ľ Ľ, Ŀ, ĳı , -, R- Ľ S1- ĳı ı ! ĳı .

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G: 1 – ı Ľ, 2 – . T Ŀ : ĳı !ı %-. F Ŀ : – Ŀ, 100% , - (10-6R), - (10-6R), - (10-6R), 2 – (10-5R), 2 G- (10-5R), 2 - (10-6R), 2 - (10-6R). 3 – . ĭ ĳ '''ı T , ı ĳı ı Ŀ ĳı ı Ŀ ı . ļJ 1. M U.. T! ! /.. G.F.!, H.F. . - R.,- 2002,– W. 352-384. 2. Q K.F., Q.I. T . –R.: R, 3. -1989, - 43. 4. T I.. . - R.: S, - 1991, - 319 . 5. G V., ., J . c . – R. R, - 1983, - 206 . 6. Melin A., Perromat A., Deleris G. The in vivo toxity carbon tetrachloride and 7. carrageenan on heart mirosomes. Analyses by Fourier transform infrared spectroscopy // Can. J. Physiol. and Pharmacol. – 2001. – 79, - 3 9, – P. 799-804. 8. H F.., R.Z., ] H.W., W c.H. 9. . R // W.. . - 2000, - 32, - W. 12-14. 10. .Z., R W.H. H «c» //H .: U . – S, - 2004, - X. 1, – W. 180-181. *** U , l4, . *** At toxic hepatite, caused by WWl4 introduction the oppression of spontaneous and caused contractile activity of mesenterium lymph nodes and infringement in biochemical content of a lymph have been observed.

292 Y . . 1 (33). 2012

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! ĳ. F ĳ «F» Ľ . - WĿ, !, ^ ĳ 10.04.04 . - W Ŀ ı Ľ X ; T ! !ı Ľ Ľ ! - ĳ , ı ,Ŀ . G - ı ı ı Ľ Ľ . Ĳ ı Ľ , «c! Ľ » ĳ ĳ Ľ : 1. F ı Ľ . 2. FĿ ı Ŀ ı - !, ı , ! Ŀ . 3. X !ı Ŀı ı . 4. F ı ı ı ı . T- ! , ĳ ĳ ı ĳ : 1. ĳ, Ľ . 2. ı ĳ ĳ - ! . 3. Ŀ ı Ŀ Ľ ı ı - ĳ . 4. WcW . 5. Ŀı , - . 6. - ! ĳ. 7. . 8. Ŀ ! . 9. ı Ŀ Ŀ , - . ļJ 1. G. F. S « » . W. ľ. F, I. F. F. - F : Ĩ, 2004. - 432 . 2. Ĵ.G. F «c! Ľ » [c ] : : / - c. . ( 27,5R). - F : S-, 2006. - . 3. U Z.U., H.H., M Q.W. I . - V /J: «».2002.- 512 4. J.F., R Q.F., V S.S. . c! !: ! / . . Q.F. R!. – 2000. – W. 67– 69. 5. G V.R. R– ! ! // H .: R– U! ! !. – S, 1991. – W. 65– 68. 6. I Z.U. M! ! : F. T / IUSXG WT VFS–S. (W. c!. H. 68), 2003. – W.138-142. 7. c! ! «I XXI » / . X. . F, - «S», 1999. – W. 125– 131. 8. V G.F. M! !. H : . / R.: - RScUY, 2001. – W. 264-266. *** F . *** In article introduced results from the edition of illness in population of Aktubinsk area connection with pollution is presented by the reference on improvement a water supply condition.

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Ľ ı ĳ : ^-W, W!, Wĳ (Ŀ), Rı ( ı), -^-G (G ı) Ľ =. Gı ĳ ĳ Ŀ ĳ . G ı , ĳ [3, 4]. W , Ŀ ı . W “I” !ı Ŀ 7 ı [5]. Wı Ŀı . G ^ Ŀ , . Rı , Ľ ı Ŀ [6, 7]. Y ı ı . W ı Ŀ . R ^ Ŀı ! Ľ . M ''. 1. ^ Ŀı ! Ľ . 2. Y Ŀı ! . M Ľ' : «VY-6» L^W-ı ı ı , ! Ŀı Ŀ . Y Ŀ Ľ Ŀ ĳ. Vı ı ^ ( ıĳ- 130 ). ı ĳ, ĳ- Ľ , ıĳ- Xĳ , ıĳ Ľ ıĳ - W! ! . L - Ŀ-^ Ľ ^ . F ĿĽ , . ľĽ ı ı . W! ! Ľ Ŀ «VY- 6» L^W ĳ. «VY-6» L^W «» , ı ! ı (IXFY), Ŀı (XKľY), Ŀ- ı (LLY) . T Ľ Ŀ . Ŀ ı ı ı ^ ı ĳ-, ^ 20 . L ĿĽ Ŀ – Wĳ ı V UH-1, 28 . «VY-6» L^W-ı Ŀ , ı ! ı (IXFY), Ŀı (XKľY), Ŀ- ı (LLY) . W-, Ľ Ŀ (=== ı), ^-Xı ı 82 . «VY-6» L^W Ŀ – ı ! Ŀ Ŀ Ľ Ŀı. Y ĳ Ľ ı Ľ Ŀı ĳ . L (! ı IXFY) ı Ŀı . X ! Ŀ . XKľY- Ŀı ı : FR Ľ FRU - ! Ŀ ; S - !; ĳ ; F ı ı ! (« ») ĳ; ! ĳ , ! !, ĳ Ľ ! ı , ! ĳ ( - ĳ ĳ ) - ı ı ĳ. G ! ĳ Ľ Ŀ . X ı ! (« ») Ŀ ĳ ı (^V^) ı ı Ŀ Ŀı Ŀ ĳ – 1000/, ĳ – 151 336 /. Wĳ Ľ Tıĳ Ŀ ! Ľ ! 42 Ŀ , ı 30- . 296 Y . . 1 (33). 2012

Ĵ Ŀ: ı ĳ , ĳ , , , Ŀ- ĳ , ı ı , , . Ĵ Ŀ – Ľ , , , Ľ , Ŀı . ľı Ŀı (FQ) (QM) . ı FQ- ı ı V- YUVMF cVF 1.7. -Ľ ! Ľ ĳ. ľı ı WM ĳ ! – 300 , ! ĳ Ľ ı ^V- Ŀ. F ı G ! ı Ŀ ^V ı - Ľ ı . Y Ŀ Ľ Ŀı ı Ľ ĳ. Q Ŀ ! Ľ IXFY-1 Ŀı ı ı 6 Ŀ, IXFY-2 Ŀı ı ı 2 Ŀ . QM ı Ŀ XKľY !! Ľ ! . ľ ı !! ĳ Ŀ ı , IHW . V! ! V WcW- ! Ľ Ŀ . ^ ! ı ĳ : U-238, Th-230, Ra-226, U-235, Ac-227, Pb-210. «c» L^W ı ^V^ ı . X ĳ ı Ŀ ı (VG Ľ VG) Ľ SVG-99 Ľ . ľĽ ıı !ı ! Ŀ Ŀ , ı Ľ ı . K , Wĳ Ľ Tıĳ ı WM VG . W , 0,091 ı ı Ľ ĳ Ŀ 30 Ŀ, Ŀ 300 1%- Ŀ. WM Ŀ (301-04/2671 12.09.2008 . IcW Ľ 31317 25.03.2009 WcW ) (1-). 1- ĳ' Ľ Tıĳ' ĵ ı M ĩ ĭ L V VG VG U-238 4,47x10 0,04 1,1 Th-230 77000 0,0088 0,2 Ra-226 21,8 0,00022 0,015 U-235 1,410¹\ 0,037 1,0 Ac-227 1600 0,03 2,5 Pb-210 22,3 0,11 9 ľĽ ıı !ı ! Ŀ Ŀ , ı Ľ ı . K , Wĳ Ľ Tıĳ ı WM VG . W , 0,091 ı ı Ľ ĳ Ŀ 30 Ŀ, Ŀ 300 1%- Ŀ. WM Ŀ (301-04/2671 12.09.2008 . IcW Ľ 31317 25.03.2009 WcW ) (2-). 2- V 2008 . Vı ^V^ V V 2009-2013 /3 /3 /3 /3 H SY. . 1 (33). 2012 297

V- 5001 0,004336 0,137 0,004336 0,137 V- 5002 0,0085 0,269 0,0085 0,269 ľĽ 0,0128 0,406 0,0128 0,406 F! Ľ Ŀ . (LG^) ! , ! ı QM-ı ! Ŀ Ŀ ĳ Ŀ. 2008-2012 . ««VY-6» L^W Ŀ ı ĿĽ Ŀıı Ľ » 750 / « » ĳ Ŀ Ŀ . L ! ı Ŀ ! Ľ Ŀ ı 9 Ŀ , ı 7- . Ŀ Ŀ : ĳ ı , ı ı, ! ı Ľ ! Ŀı ı Ŀ (ĳ ı Ľ , ı ); R Ľ ı , , Ľ , , Ŀ , , ı, Ŀı Ľ Ŀ ; FMW Ľ Ŀ ı ; F Ŀı, !ı JHW ı Ŀ – (), Ŀ , ĳ , , , Ŀ, ĳ. V TR -Ľ Ľ Ŀ ! ı . L ! Ľ «VY-6» L^W Ŀ Ŀ Ŀ ! ı ! , ^V.. ! ĳ. 11.1155 / Ľ 1.3151 / Ŀ ««VY-6» L^W Ŀ ı Ŀ ĳ . JcW Ŀ, Ŀ ı 0.4135 / Ľ 0.4135 / . Wĳ ĿĽ 30 , ! ĳ . W- ! Ŀ , Ŀ Ŀ . X ı Ŀ 39420 ³ , Ŀ Ľ , RcJ Ľ ı ı Ŀ ( +30 V/ , 48 V/), ĳ - ı ı Ŀ . «VY–6» L^W–ı ı ı ĳ Ŀ ! Ľ . . Ľı ĿĽ ı Ŀ ı WM–ı ı Ŀ ! (Ŀ , ĳ , Ľ ı) ĳ ^ ! Ľ ı . ( , JIVF–160 ! Ľ ) Ŀ Ŀ , ĳ Ľ Ŀ ı ! ^ . W , «VY–6» L^W–ı ı 2011 ĳ Ŀ ! ı Ľ , Ľ ı WM ! ı ! Ľ Ľ Ŀ . R Ľı ı Ľ . ļJ 1. F! S. F., S G. F., U S. S. c! ! !. – R. – F, 1996. – 224 . 2. F! S. F., X H. . c! : . – R.: c, 1994. – 225 . 3. R J. L. W! !. – R.: U, 1991. – 176 . 4. Ĵ.G. F «c! Ľ » [c ] : : / - c. . ( 27,5R). - F : S-, 2006. - . 5. T Z. T . – R.: R, 1975. – 740 . 6. U G. G. c! : W- / T. . F. I. H. – R.: S, 1991. – 122 . 7. W. H IR // U . – 2002. 3 12. W.12-14. 298 Y . . 1 (33). 2012

*** U , MP . S P . *** In results of ecological monitoring at the board of North Kazakhstan is big concentration of bad matters in the air and exhausting by the plants, is not overdraws norms. Karamuryn is not influence to the monitoring of pollution of the.

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, Ŀı, , ĳ Ľ Ŀ . F ı ı Ľ- ĳ Ľ . G . F ı ı , Ŀ Ŀ . R «LĿ» «XĿ» Ŀ ı F ı 1991 347 «» . Ŀ Ŀ Ľ ĳ. W ı – ı Ŀı , , . F ! Ŀ Ŀ «» Ľ «R», «FXT», «», «F» . , Ľ FSFL ı Ŀ ĳ ı, ı ı, ĳ ı . ı ! ĳ. T Ŀı (50 ) Ŀı , ı ı Ľ. ĳ Ľ Ŀ ĳ . ľ, Ŀ . ľ ı ı – ĳ Ľ . ĳ Ľ ĳ ĳ Ŀ Ŀ Ŀ Ŀ Ŀ . F, Ŀı . ĳ Ľ . L ! Ľ Ŀı , Ľ Ŀ Ľ . F ı Ŀ Ľı – ı. « ı» ĳ 120 . G Ŀ 1872 G V W . Lı ı ı , . F Ŀ – ĳ Ľ , Ľ, ĳ Ľ ĳ. SĽ ı , Ľ S Ľ 5,6- Ŀ – ı . F ! ı Ŀ ı Ŀ- ĳ , Ŀ, , . ĳı ı Ŀ . F W . ıı Ľ Ŀı, Ŀ-Ŀ ı Ľ ı Ľ Ŀ . , ! ı ı Ľ – . G ı ĳ ı Ŀı Ŀ, Ŀ . ı Ŀ ĳ , , ĳ . W ! Ľ Ľ Ŀ. ļJ 1. G .T. c //H RTSV SFSV,-2002, 31,-W.12-17. 2. «F» 2006, 326, 31.01. 3. W !// Tıĳ «R ĳ» 2000 06.07 4. ı ! //- «L »,- 2005, 09.09. 5. G I.G. c!, Ŀ //- F,- 2004. 6. T I.W., G I.X. F . «c!». – F.: c 2002. *** N , , , , , . *** Intensive pollution of air, water and soil, degradation of flora and fauna, depletion of natural resources led to the destruction of ecosystems, desertification, and a significant loss of biological and landscape diversity, increased morbidity and mortality.

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3 , , ! ! !: - H, ! ! , ! , ( 3) . ! !, ! ( ) . , ! ; - : , !! !. H ! !, (!) . c ! ! , ! ! . H" ! !! !. ! ! !, , ! !. ! (!) ! !!, ! ! !. U, 4 , ! ! . c ! ! . J . X ! . Y!, , KNO3, Ca(NO3)2, , NH4NO3, KCl 5(Ca(NO3)2·2H2O)·NH4NO3), , , ! ! () ;

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W Sne [5], ! !, !, ! !: - : ! ! ! ! . U ; - : ! ! ! ! !, ! . G , , , ! ! - !. J! ! !, . V !. U , ! , !! . U ! ! 4 [6]: - ! ! ; - !; - ; - ! . Y ! , , , pH. Hagin Lowengart-Aycicegi [7] , ! : - : !, ! ! , , ; - : ! ! – ! , . V !; - ! : ( 1). T , : - !, ! . H , ! , , , !; - ! . U ! ! 50 1 ! ! ! 1-2 . K ! , ! ! ! ! ! [8]; - ! !. S ! , ! ! (, KNO3, Ca(NO3)2, , NH4NO3, KCl 5(Ca(NO3)2·2H2O)·NH4NO3). T, ! !, !, , ; - ! . ! . ! , , , , . S 1-3 [9]. X 1 !, . H 2 ! !. X 3 ! . H ! 4 [10]. U. S , ! KCl ! ! . ^ 1. S 4 ! : 3,7 K2O/ . ^ 2. H!, ! : Y! ^ 1 , : 3.7 K2O/ × 30 / = 111 K2O/ ^ 3. H! ! KCl ( K2O KCl ! 60 %):

^ 4. J , !! 20% ! : 185 KCl/ + (0.2 × 185) = 222 KCl/

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U ! [11]. X , ! ! NPK . U ! , , ! , ! !, ! !. X 1 W Fe, Fe, CO(NH2)2 NH4NO3 (NH4)2SO4 Ca(NO3)2 KNO3 KCl K2SO4 (NH4)3PO4 Zn, Zn, MgSO4 H3PO4 H2SO4 HNO3 Cu, Cu, Mn Mn R + S + + ! W + + + ! S + + - + ! S + + + + + ! + + + + + + ! W + + ± - + ± + ! + + + - + + + + ! W Fe, Zn, Cu, + + + - + + ± - + Mn Fe, + + + ± + + + ± + + Zn, Cu, Mn W + + + - + + ± - + + + ! ! + + + - + + + + + ± + + W! + + + - + + ± + + + + + + F! + + + + + + + + + - + + + + «+» - , «-» - , «±» - X 2 , R (), - U 100 (%) (.) 20°

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!. J! !!! ! ! . Q 1. Bar-Yosef B. Fertilization under drip irrigation // In: Fluid Fertilizer, Science and Technology. Ed. by D.A. Palgrave. Marcel Dekker, New York. - 1992.- pp. 285-329. 2. Bar-Yosef B. Advances in Fertigation // Adv. Agron. - 1999. – V. 65. – pp. 1–77. 3. Hagin J., M. Sneh and A. Lowengart-Aycicegi. Fertigation – Fertilization through irrigation // IPI Research Topics No. 23. Ed. by A.E. Johnston. International Potash Institute, Basel, Switzerland. - 2002. 4. Elfuving D.C. Crop response to trickle irrigation // In: Horticultural Reviews. J. Janick ed. AVI publishing Co. Inc. Westport Conn. USA. - 1982. - pp.1- 48. 5. Sne M. Micro irrigation in arid and semi-arid regions. Guidelines for planning and design // Ed. by S.A. Kulkarni. ICID-CIID. International Commission on Irrigation and Drainage. New Delhi, India. - 2006. 6. Kafkafi U. Global aspects of fertigation usage // Fertigation Proceedings, International Symposium on Fertigation Beijing, China. - 2005. - pp. 8-22. 7. Hagin J. and A. Lowengart. Fertigation for minimizing environmental pollution by fertilizers. // Fert. Res. - 1996. – V. 43. – pp. 5-7. 8. www.omafra.gov.on.ca/english/crops/hort/news/vegnews/2006/vg0406a2.htm 9. Kafkafi U. and J. Tarchitzky. Fertigation: A Tool for Efficient Fertilizer and Water Management //First edition, IFA, IPI, Paris, France, May. - 2011. - P. 138. 10. Wichmann W. World Fertilizer Use Manual // BASF AG, Germany, 25 April -1992. – P. 600. 11. www.iclfertilizers.com/Fertilizers/Pages/KnowledgeCenter.aspx *** V (N) 2011 13 22 %' '' ''' ' '' % '' RF^F %' . '' '%'' %' N' 16 ' '' '. N '' '' . R ''', ' - % ' %' . Z ''': - , '' . &''' ' ' ' '' % %' , '' '' '. Z ' % . R %'' '' . *** From 13 November to December 22, 2011 in Rehovot (Israel) by MASHAV (Israel Ministry of Foreign Affairs) organized a training course on the management of crop production under saline stress as a result of climatic changes, fertigation for the efficient management of fertilizers. Through the use of industrial fertilizers and new technologies of the agrarian sector of Israel currently provides agricultural products more than 16 countries around the world. One of the modern farming methods, which successfully use Israel, is fertigation. This article provides the notion of definition of fertigation, fertigation equipment, dosing of fertilizers. Benefits of fertigation allow are saving labour and cost of equipment, effective (almost 100%) the use of expensive chemical fertilizers. Fertigation is the most efficient and cost-effective way to deliver nutrients to root systems and allows receiving high harvests. Fertigation allows systems management is easily achieved optimal concentrations of fertilizers, their relationship, and these settings can be controlled automatically. The lectures and demonstration materials, given by the organizers of course, were used for writing of the article.

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\ ! ! \U . H ! (UIIX) : (\U) (RJF). RQ RTJ QJT H ! 30-40 (n=14). U 15 . J ! ! \U [6] (UTQ) ( 2- ). V ! - H [7]. MQb TLJ T !!! , - , . T ! . T, , , 1,85-3,17 /. U \U. H – (1 ), – 2 ( 1). J ! ! . X 1 Y! W. W / 1 2 J M 2,29 2,77 ± m 1,30 1,96 U M 2,63 2,51 ± m 1,27 1,14 n 94 W \U 14,8%, 2,19 – 2,98 /, ! . H , ! \U 64% (1 ) 14,9%. Y 2 (36 %) 9,3 %. W , 2 , 1 ( 1).

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V ! , RJF 14,8%. ! ! RJF , RJF . S! , ! , !, ! . c ! ! . I , !!! ! , , . . 8. U UIIX ! , , . Y . X, ! \U ! . Y \U , . W ! !!!, , ! WVT . J, RJF . R , ! ! ! - . TJ 1. Y : - !, !; 2. I ! ; 3. T! ! - !!! !. Q 1. ! F.M, \ X.S, T T.F. S! ! ! . - R.: R, 2003. - 403 . 2. I H.H., W S.., F.T. I!. – TTT «cQG – WU», 2003. – 535 . 3. H S.., F.H., W H.. . ! ! . – R.: H! ! VHWS U H, 2000. – 91 . 4. R H.., U I.J., T K.H. . I // G . - F, 2009. - ].2. – W. 261-264. 5. M! .F. F! , . / .F. M!, R.H. G// H. V FRS. 1995. 36. W.53 – 60 6. Ravin H. A. An improved colometric enzymatic assay of ceruloplasmin // . Lab. Clin. Med. Biochem. 1961. Vol. 58, 3 1. P. 161168. 7. ! ! . U! I F HRF, 1964 ., 1975 ., 1983 ., 1989 ., 1996 ., 2000 . //www.soramn.ru. 8. Y . G., ]! .S.., ^ F. F., T S. . I! ! ! . S 2003. - 3 2. *** X'' '' '' '' % ' %' ''. T'' ' '' . *** It had been study the effect of ceruloplasmin and malonic aldehyd on the adaptation to interrupt hypobaric hypoxia. Individual character of the organism adaptive reaction to hypoxia had been ascertain.

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U!, ! ! , . J! ! ! , ! ! ! ! . c " , ! . J! ! ! ! ! . S , ! ! " ! . J! ! , ! , " ! PWC 170, " ( ", , ! ) ]WW, ! ! . R ! ! ( ) - (IV) ! - ! . V! ! ! - ! -- (F.I.R, H.R.] .), " ! . R ! - , . H 1- 16-18 U 7,6 12,8, U 15,3 21,4. - !! . U ! “” (! ) ! . ! IV. U ! , ! ! . U . I I U 1 7 11 64 3 . 80 95 55 2 7,3 9,5 6,0 4 . 70 67 50 , , ! , IV, , !. W " . J! 3 ! 15 . J " ]WW ! ! !. I U ] ]WW H! ! 1 63 210 U 125 2 65 237 U 138 X , " , ! ]WW !, ! ! , . U , ! ! ! , !! . c ! ! IV. Q 1. J H.W. X Z.U., U H.X. J . X! . , 1991, 310 2. J H.W., S F.F. W! R., 1975 3. X! I.W. W! . Y. . R., W , 1998. 4. M .. T! . J. . R.: 2007. 5. ] H.. M . X! . , 1960, 311. *** R '''' '', ''''' % ' ' ''' -' ''' '. *** This paper analyzes the relationship between the feature build, stamina, and an indicator of galvanic skin response, which characterizes the level of the hypothalamic-sympathetic-adrenal activity. H SY. . 1 (33). 2012 311

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F I, XTT «SV» [email protected] F S F ..., , 87054022000, [email protected] F W R, , F XTT « - !» 8(7232) 259860 , [email protected] F F, SY . - F R, SY . - G R X ..., Y « » e-mail: [email protected] G I , ..., , JIU «S » VIU «SY .-» (272)-377-34-37 () [email protected] G X. , SV G F.I., SY . - G S! G, , SV H X! W, SY . -, X.:87023227047 E-mail: [email protected] I F ..., , SV J K XTT «SV» 245-37-76; .: 87057135903 , K-m1: [email protected] J ! F, ..., SY . -, 8727-377-33-34 (.12-12) J Q! F, 4 SY . -, 87759773646 J I R, JIU «S » VIU «SY . - » [email protected] 375-71-53 K G K, ..., SY . -, 8727-377-33-34 (.12-12), [email protected] L R ..., , SY . F-, F[email protected] L J , (!), - !, 8(727)-245-37-76, [email protected] M Q F, ..., , XTT « », .: 8(727)250-58-71, [email protected] W! F, 4 SY . -, 8727-377-33-34 (.12-12), [email protected] G, ..., - !, 8-727-383-3-17-15 [email protected] Q X, XTT «SV»87770990702,[email protected] T R, SY . -, 8727-377-33-34 (.12-12), [email protected] Kazuyuki INUBUSHI, professor, Graduate School of Horticulture, Chiba University, Matsudo, Chiba 271-8510, Japan, T +81-47-308-8816, F +81-47-308-8720, [email protected] I G, SY . -, 8727-377-33-34 (.12-12), [email protected] F , ! , M.. XTT «SV», .8(7112)28-11-26 [email protected] T , ! , XTT__-_- _ F , 25-98-60, fishedu @ mil. ru " H ..., , XTT « », X.: 8(727)250-58-71, e- mail: [email protected] H S, / , V -F \ "Q! J U"/ W - !, X.: 2(7212)516144, E-mail: [email protected] W W, ..., , SY . -, Saidina. [email protected] W!, F, XTT «SV» H SY. . 1 (33). 2012 319

Q! F, ..., ! W +996 (312) 642639.J + 996 (312) 310621, E-mail [email protected] Q T K ..., S RTS V R S, ..., . S RTS V R c H .. c ! FWUcT, U UVTTS-Ic FWUcT X.:+7 7172 (79-83-18) (. 109), e-mail: Elchin_mamedov @caspeco.org; [email protected] R S ^ ..., , , JIU «S » VIU «SY .-» (272)-377-34-37 R H, ..., PhD, , Y RI (University MacGill) R K H, , XTT « », X.: 8(727)250-58-71. R F F, , SV S W T, ..., TTT SUU «FH!U » X. 8 927 551 61 10, e-mail: [email protected] S W X, ..., SY . - T L W, ..., SY . -, Zhanar.Omarova @kaznu.kz T S, SY . - T H I XTT « F » (FUc) X.: (727) 2582496/2582489 , -mail: [email protected] U R . , SV E-mail: [email protected] U J K, ... S RTS V V I SY . - W K X, , XTT «SV», 245-37-76; .: [email protected] W S W, ..., SY . -, 8727-377-33-34 (.12-12), [email protected] W ^ X, W F W, , , SV W L, .-., ., X ! - , X.: 8(7262)-46-42-41, 87776498389, E-mail: [email protected] X H, .., . . , \! ! , IGY «W», X.:…8-812-234-4609…+79214290334 , E-mail: …[email protected] X S S, , JIU «S- » VIU « -» mybiolife- [email protected] ^ I R, XTT «SV». ^ F, SY . - ] S! H, ..., PhD W Y V F- S M VFS, MS VFS, .: (812)328-06-12 , -mail: [email protected] c - ! K-: [email protected] X.8 927 570 10 63 ! V U, ..., .. - ! F W G, ..., ..., - ! F F! F, /, ... - ! U Z F, ..., ... - ! W W , /, ... - ! W H, /, - ! I J! F, .. ..., .. - ! M X! W, F J V, /, .. - ! 320 Y . . 1 (33). 2012

F F.R., XTT « - » FT «F!» X! S! K, ..., , U , . 8-7182-50-95-33 (.), 8-7182-65-16-06 () F W , ..., ..., S F F, . 050 574 67 80, . – 994 012 538 83 17, e-mail: [email protected] V.H.I, R.F.F, R F, e:mail: [email protected] J! H , , F SV, e-mail: [email protected] K F F, - I , H- , .: 70 14 48, : 76 73 91, e-mail: [email protected] K I.., G Z.F., WT VFS, . G [email protected] K S.., .., .., WT VFS, . 8-383-330-84-84, :…8-383-330-20-05, E-mail: …[email protected] M K! Z , ..., ... WT VFS, . G, V!, . 89132136812, : 8(3852)240214, e-mail: [email protected] " Q F, , XTT « S !» FT «F!»», .383-17-168 (.), 290-61-23 (.), 7016009817, 2831516, E-mail: [email protected], [email protected] K I, ..., VIU « », X. 375-45-82, e-mail: [email protected] X! , ..., , 1F I ,V!, . F, e-mail: [email protected] R L T, , XTT « - !», FT «F!», . F, 8-7016288853, [email protected] R V, , S X , e-mail: [email protected] R G! , ..., , . - ^ S U, S . - , e-mail: [email protected] W S! S, . . G XTT « - !», . 871036 41774, 817036 41474, e-mail: [email protected] X! S! K, ..., , U , . 8-7182-50-95-33 (.), 8-7182-65-16-06 () X W V, , XTT « , .: 226-27-91, e-mail:[email protected]; [email protected] W! T S, , , . 2682496, .108, 117, :23091049, e-mail: [email protected] W J F, ! , , .: 8(727)2582496 .115, : 8 (727)2391049, e-mail: [email protected] c Z! H, , X X! X, , XTT « !», ..383-17-16, .224-19-74, : 8(727)383-17-15, e-mail: [email protected], [email protected] R H F, R K H., XTT « » . F, ^ Q , ..., .., ! , XTT « - !», / 8 (727) 383-17-15, e-mail: [email protected] Q H, ..., , ..., WT VFS, X.: 8(3852)36-46-81, : 8(3852)240396, E-mail:[email protected] F W S, ..., ..., F Ľ ! ĨM F S X, ..., , Ľ- ĴY L L, F Q, 4- , Ľ- ĴY XĿ W X, ..., , Ľ- ĴY, . 8-72-737-76-06, . 12- 08, e-mail: [email protected] c R , R S F, , 1-, Ľ- ĴY Iĳ F, ..., Ľ- ĴY H SY. . 1 (33). 2012 321

Y L, R , V , K , «c!» ı 2- F IĳĽ F, K Iĳ K, F U, R Iĳ V, «c!» ı 2- F , F K, , 2- , Ľ- ĴY, F S X, ..., , Ľ- ĴY L W, F Ĵ , F , F T, M J, 2- , Ľ- ĴY H Z, .., . , XTT « - !», _, 8-7163641153 S G, .., . , XTT « - !», _, 8-7163641153 X J F, SY . -, .: 87051334979, e-mail: danara_ [email protected] K F G, ..., SY . -, .: 8-72-737-76-06, . 12-08 R F F, ..., SY . -, . 8-72-737-76-06, . 12-08, e- mail: [email protected] R W W, ..., SY . -, . 8-72-737-76-06, . 12-08, 8 705 239 45 25 I, SY . - T K H, ..., ..., ! » S RTS V F F K, 4 , SY . -, [email protected] ^ W F, ..., , SY . -, [email protected] S R G, 4 , SY . - F W S, . K.F. G, ..., . H . I \ . [email protected] J ! F, .K.2, G I.G.2. SY . -, .., . [email protected], [email protected] K U F, S RTS V, , G. [email protected] H Q, VIU , . , , . [email protected] R S R, ., .X.F, W.J. SY . -, R F.X., F W.W., F S.M., S F.W., Ľ- , F , . X , VIU , , [email protected] , 2-37-48-52, 8-777-352-64-96 F W.W., F S.M. SY . - C G R, [email protected] , 380-93-17

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Lĳ' L. L : Ľ ı : . – 2012. – 222 . ISBN 978-601-247-336-0 T ! Ľ ! ĳ Ľ Ŀ Ľ ı ı- , ı .

. Ľ .. c! Ľ : / W., V.G, R.^Ľ. – 2012. – 130 . ISBN 978–601–247–309–4 T !ı ı - ı ı, ĳı Ľ , ı , Ľ ı , ı -Ľı Ľ, Ľ- ! Ľ ! Ľ !.

. ľ ! . – 2011. – 112 . ISBN 978-601-247-265-3 G Ŀ ı !, Ŀı !, , , Ŀı , !, Ŀı Ŀ Ľ .

.I. Q : -. . – 2011. – 82 . ISBN 9965–29–693–6 H - , ! ! ! ! , ! !.

., . G! !: . - 2-, . – 2011. - 294 . ISBN 9965-29-763-0 H ! ! , ! !, ! , ! !! , , ! , !.

U ! ! « » I.M. .: 8(727)377-33-66, 377-33-37, . 11-68. R.. 8-777-316-03-97. E-mail: [email protected]. www.read.kz