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Carideorum Catalogus: the Recent Species of the Dendrobranchiate, Stenopodidean, Procarididean and Caridean Shrimps (Crustacea: Decapoda)

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Carideorum Catalogus: the Recent Species of the Dendrobranchiate, Stenopodidean, Procarididean and Caridean Shrimps (Crustacea: Decapoda) Carideorum Catalogus: The Recent Species of the Dendrobranchiate, Stenopodidean, Procarididean and Caridean Shrimps (Crustacea: Decapoda) S. De Grave & C.H.J.M. Fransen* De Grave, S. & C.H.J.M. Fransen. Carideorum catalogus: the recent species of the dendrobranchiate, stenopodidean, procarididean and caridean shrimps (Crustacea: Decapoda). Zool. Med. Leiden 85(9): 30.ix.2011: 195-589 figs 1-59.— ISSN 0024-0672, ISBN 978-90-6519-200-4. S. De Grave, Oxford University Museum of Natural History, Parks Road, Oxford, OX1 3PW, United Kingdom ([email protected]). C.H.J.M. Fransen, Department of Marine Zoology, Netherlands Centre for Biodiversity Naturalis, P.O. Box 9517, 2300 RA Leiden, The Netherlands ([email protected]). * The order of authorship is purely alphabetical, with both authors having equally contributed to this work. Key words: Crustacea; Decapoda; Dendrobranchiata; Stenopodidea; Procarididea; Caridea; checklist. A checklist of recent species of dendrobranchiate, stenopodidean, procarididean and caridean shrimps including synonyms and type localities. Also listed are unavailable names, larval names, nomina dubia and nomina nuda. A complete list of references to original descriptions of taxa listed is provided. Contents Introduction ............................................................................................................................................................ 196 The higher classification of shrimp .................................................................................................. 196 Fossil taxa ........................................................................................................................................................ 197 How many shrimp species are there? ............................................................................................ 197 Structure of the list ............................................................................................................................................. 197 Notes on type locality ....................................................................................................................................... 199 Notes on dating of some papers ................................................................................................................ 200 Notes on author names .................................................................................................................................... 201 Notes on authorship of taxa ......................................................................................................................... 202 Notes on some papers of nomenclatorial significance ................................................................. 203 Notes on the spelling of some names ..................................................................................................... 204 Notes on the gender of generic names and gender agreement in species names ........ 204 Acknowledgements ........................................................................................................................................... 205 Checklist ................................................................................................................................................................... 206 Suborder Dendrobranchiata ................................................................................................................ 206 Suborder Pleocyemata ............................................................................................................................. 248 Infraorder Procarididea .......................................................................................................................... 248 Infraorder Stenopodidea ........................................................................................................................ 249 Infraorder Caridea ..................................................................................................................................... 254 Unavailable names ..................................................................................................................................... 471 Penaeoidea Larvata .................................................................................................................................... 471 Penaeoidea Nomina dubia .................................................................................................................... 471 196 De Grave & Fransen. Carideorum catalogus (Crustacea: Decapoda). Zool. Med. Leiden 85 (2011) Penaeoidea Nomina nuda ..................................................................................................................... 472 Sergestoidea Larvata ................................................................................................................................. 472 Sergestoidea Nomina dubia ................................................................................................................. 472 Sergestoidea Nomina nuda .................................................................................................................. 473 Caridea Larvata ............................................................................................................................................ 473 Caridea Nomina dubia ............................................................................................................................ 475 Caridea Nomina nuda ............................................................................................................................. 476 References ................................................................................................................................................................ 478 Systematic index to currently valid families and genera ............................................................ 585 Introduction The higher classification of shrimp Over the last decade or so, much has been written on the classification of Decapoda, fuelled by a surge in molecular phylogenetic studies, as well as close scrutiny of inter- nal and external morphological characteristics. As discussed by Fransen & De Grave (2009), such studies on shrimps are still somewhat ”thin on the ground”, at least com- pared to the more extensive work done on the Brachyura and Anomura. At a higher level in decapod classification it has long been recognised that three distinct lineages of shrimps can be distinguished: Dendrobranchiata, Stenopodidea and Caridea, a system which has not been seriously challenged by recent studies. The internal classification of Dendrobranchiata and Stenopodidea alike has been stable for some time, with the only major addition being the family Macromaxilloca- rididae Alvarez, Iliffe & Villalobos (2006) to the Stenopodidea in recent years. A different picture has emerged for Caridea very recently with Bracken et al. (2009) and Chan et al. (2010), both drawing attention to the non-monophyletic status of certain superfamilies and families. Further, we are aware of work currently in progress (some by the authors of this compilation) corroborating the hypothesis that the current clas- sification of Caridea is unnatural, lines of study which will lead to the resurrection of certain family names as well as further refinement to other families. As one of our objec- tives for the current effort was to link this compilation of species level information with the earlier work by Chace (1992) for families and Holthuis (1993a) for genera, we have elected to largely follow the classification outlined by De Grave et al. (2009) which builds upon this earlier work. As such, it was deemed advisable to include the recently resurrected family Acanthephyridae Spence Bate, 1888 in the superfamily Oplopho- roidea, rather than in this catalogue to create a new superfamily, which would perhaps be more congruent with the results in Chan et al. (2010). Although we follow herein the classification scheme of De Grave et al. (2009), two recent changes have been implemented. The clarification of the status of Galatheacaris abyssalis Vereshchaka, 1997a, as the megalopal stage of Eugonatonotus chacei Chan & Yu, 1991a, by De Grave et al. (2010) resulted in the removal of the family Galatheacarididae and superfamily Galatheacaridoidea in the current listing. Bracken et al. (2010) clarified the status of the family Procarididae, resulting in the recognition of a fourth group of shrimp, Infraorder Procarididea. De Grave & Fransen. Carideorum catalogus (Crustacea: Decapoda). Zool. Med. Leiden 85 (2011) 197 Fossil taxa The current listing is restricted to extant genera, species and subspecies only. For readers interested in how the current classification of extant taxa compares with those of fossil taxa, we refer to De Grave et al. (2009) who employed a suprageneric classifica- tion of Decapoda across both fossil and extant taxa, as well as Schweitzer et al. (2010) who list all species (and their classification) of fossil species. Schweitzer et al. (2010) list 97 fossil species of Dendrobranchiata in 31 genera, two fossil species in Stenopodidae (each in its own genus) and 52 species of fossil Caridea in 33 genera. Their placement of the genus Udora Münster, 1839, within the superfamily Procaridoidea (herein consid- ered at infraordinal level) is interesting, as this group is currently only known from six species, restricted to anchialine habitats. If the five
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