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Redalyc.Redescription of Tijubina Pontei, an Early Anais da Academia Brasileira de Ciências ISSN: 0001-3765 [email protected] Academia Brasileira de Ciências Brasil Simões, Tiago R. Redescription of Tijubina pontei, an Early Cretaceous lizard (Reptilia; Squamata) from the Crato Formation of Brazil Anais da Academia Brasileira de Ciências, vol. 84, núm. 1, 2012, pp. 79-93 Academia Brasileira de Ciências Rio de Janeiro, Brasil Available in: http://www.redalyc.org/articulo.oa?id=32721622008 How to cite Complete issue Scientific Information System More information about this article Network of Scientific Journals from Latin America, the Caribbean, Spain and Portugal Journal's homepage in redalyc.org Non-profit academic project, developed under the open access initiative Anais da Academia Brasileira de Ciências (2012) 84(1): 79-93 (Annals of the Brazilian Academy of Sciences) Printed version ISSN 0001-3765 / Online version ISSN 1678-2690 www.scielo.br/aabc Redescription of Tijubina pontei, an Early Cretaceous lizard (Reptilia; Squamata) from the Crato Formation of Brazil TIAGO R. SIMÕES Setor de Paleovertebrados, Departamento de Geologia e Paleontolgia, Museu Nacional, Universidade Federal do Rio de Janeiro, Quinta da Boa Vista, s/n, 20940-040 Rio de Janeiro, RJ, Brasil. Manuscript received on August 19, 2011; accepted for publication on November 9, 2011 ABSTRACT The record of Gondwanan Mesozoic lizards is very poor. Among the few species described for this region there is Tijubina pontei, an Early Cretaceous lizard from the Crato Formation (late Aptian) of northeast Brazil. Its description is very brief and lacks most of its diagnostic characters and clear delimitation from other lizard species. Here, a full redescription of the holotype is provided. T. pontei is demonstrated to be a valid species and a new diagnosis is provided with reference to Olindalacerta brasiliensis, a contemporary species of the Crato Formation. It lacks the posteroventral and posterodorsal processes of the dentary; the tibial/fibular length equals the femoral length and its posterior dentary teeth are robust, cylindrically based, unsculptured and bear no cuspids. The systematic position of T. pontei still needs further clarification, but preliminary analyses indicate that it lies in a rather basal position among the Squamata, similarly to O. brasiliensis. Key words: Brazil, Crato Formation, fossil lizard, Lower Cretaceous, Tijubina pontei. INTRODUCTION marocensis Richter 1994 and Tarratosaurus Lizard remains from the Mesozoic of Gondwanan anoualensis Broschinski & Sigogneau-Russell 1996 derived continents are extremely rare. There is a from the Early Cretaceous (Berriasian) of Morocco; proportionally higher number of snakes relative Konkasaurus mahalana Krause, Evans & Gao to lizards taxa in these regions and with a pattern 2003 from the Late Cretaceous (Maastrichtian) of opposite to that seen in Laurasia, where fossil lizards Madagascar; Pristiguana brasiliensis Estes & Price are much more common than fossil snakes (Krause 1973 from the Late Cretaceous of southeast Brazil; . 2003). Among the few taxa described, there are the Brasiliguana prudentis Nava & Martinelli 2011 species Tikiguana estesi Datta & Ray 2006, from from the Late Cretaceous (Turonian-Santonian) the Late Triassic (Carnian) of India; Bharatagama of southeast Brazil; Olindalacerta brasiliensis rebbanensis Evans, Prasad & Manhas 2002, from Evans & Yabumoto 1998 and Tijubina pontei the Lower-Middle Jurassic of India; Paramacellodus Bonfim-Júnior & Marques 1997 from the Late Cretaceous (Aptian-Albian) of the Crato Formation Correspondence to: Tiago Simões E-mail: [email protected] of northeast Brazil. Whether this distribution An Acad Bras Cienc (2012) 84 (1) 80 TIAGO R. SIMÕES difference is real or represents differential research out, from top to base (Neumann and Cabrera 1999, efforts between areas in the northern and the southern Valença et al. 2003). The Romualdo and Crato hemisphere is still unknown (Evans 2003). formations are among the most fossiliferous and To this day, the only valid species for South important fossil deposits in Brazil and are considered America are the four above ones from Brazil. world quality lagerstätten (Maisey 1993, Kellner The two Early Cretaceous taxa are significantly and Campos 1999). The holotype of Tijubina pontei better preserved, both with cranial and numerous comes from the Crato Formation, which is known postcranial elements, whereas the Late Cretaceous to have produced several thousand specimens of ones are represented almost exclusively by cranial fishes, plants and arthropods, including insects fragments. Phylogenetically, Olindalacerta was (Martins-Neto 2002, Santos et al. 2011), crustaceans interpreted as a sister group to Scleroglossa, (Martins-Neto 1987) and chelicerates (Campos clustering close to Eichstaettisaurus (Evans & 1986). Among tetrapods, there were found turtles Yabumoto 1998), a basal squamate from the Late (Oliveira and Kellner 2007), crocodilians (Kellner Jurassic and Early Cretaceous of Europe. Tijubina 1998), pterosaurs (Frey and Martill 1994, Campos pontei was originally described as a teiid (Bonfim- and Kellner 1997), anurans (Leal and Brito 2006, Júnior and Marques 1997), then as a basal Squamate Báez et al. 2009), fossil feathers (Kellner et al. 1994, (Bonfim-Júnior and Avilla 2002) and even as a Sayão et al. 2011) and the two published species Rhyncocephalia (Martill and Frey 1998). It is a well- of lizards mentioned above. The Crato Formation preserved specimen, but T. pontei material remains consists mostly of thin laminated limestones with largely undescribed (Evans 2003); therefore, a variable clay content and dark-coloured calciferous redescription is necessary. This may allow Tijubina shales (Valença et al. 2003). This represents a to have a better supported systematic positioning carbonatic lacustrine environment and is late Aptian among the squamates and provide more data to to Aptian/Albian in age (Maisey 1991, Neumann and future phylogenetic analyses, thus contributing to Cabrera 2002, Valença et al. 2003), despite diverging an advance in the understanding of the Cretaceous opinions on its precise dating (see Maisey 1991 for lizard fauna in Brazil. further details). This ancient lake was formed due to Here a full redescription of Tijubina pontei tension stresses on the South American plate during is presented, along with a reinterpretation of its the rift phase at the opening of the Atlantic Ocean. It systematic position, based on new data from its was located in an arid region and was one of the last holotype housed at the Museu de Paleontologia points of contact between South America and Africa de Santana do Cariri (MPSC-V 010), which is before their full separation by the Aptian/Albian the only currently available material attributed transition (Maisey 1993). to this species. MATERIALS AND METHODS GEOLOGICAL BACKGROUND Considering the small size of the specimen, The Araripe Basin in northeast Brazil can be divided measurements were done digitally utilizing high into two main sub-basins: the West or Feitoria sub- resolution pictures, which provide greater precision. basin and the East or Feira Nova sub-basin (Viana The proportion between fore and hind limbs has and Neumann 2002, Martill et al. 2007). It contains been calculated based on the length of the propodials the Santana Group where the Arajara, Romualdo, plus the epipodials, excluding the autopodium. Ipubi, Crato and Rio da Batateira formations crop This is necessary so the digital formula, which is an An Acad Bras Cienc (2012) 84 (1) REDESCRIPTION OF Tijubina Pontei 81 independent character, does not affect the fore/hind Type species: Tijubina pontei Bonfim-Júnior & limbs length ratio. Furthermore, the propodial and Marques 1997, by monotypy. epipodial lengths make the biggest contribution to Diagnosis: as for species the limbs total length. The published phylogenetic analyses of the Tijubina pontei Bonfim-Júnior & Marques 1997 Squamata considered for the systematic positioning Revised and extended diagnosis: Tijubina pontei of T. pontei within this group were chosen based differs from other Cretaceous squamates by the on two main criteria. The first one is to include following combination of characters: surangular an extensive taxonomic sampling throughout the and coronoid processes of dentary absent; tall Squamata, and the second, to have an osteologically facial process of maxilla; jugal slender with based diagnoses for the major monophyletic groups pointed ends, curved and with partial overlap that were found. These are the works of Estes et al. with the maxilla anteromedially; ectopterygoid (1988), Lee (1998) and Conrad (2008). limited to posterolateral margin of suborbital Abbreviations used: acf, anterior coracoid fenestra, not extending further anteriorly; fenestra; ans, annulated scales; ante, anterior meckelian canal open medially; splenial absent dentary tooth/teeth; bd, basal portion of dentary; or much reduced; deeply implanted cylindrically c, coronoid; cbr I, ceratobranchials I; cl, clavicle; based pleurodont teeth; anteriormost teeth of the cr, carpal rows; d, dentary; dd, dorsal portion mandibles slender, unicuspidate and unsculptured; of dentary; dss, dentary-suragular suture; ec, very robust posterior dentary and maxillary teeth, ectopterygoid; epco, epicoracoid; I – IV, first to with a sharply pointed apical region that lacks any fourth digits; fe, femur; fi, fibula; hu, humerus; processes or ornamentation; cylindrical base of icl, interclavicle; il, ilium; ims, intramandibular
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