California, USA
Total Page:16
File Type:pdf, Size:1020Kb
Load more
Recommended publications
-
Comparative Dissection of Three Giant Genomes: Allium Cepa, Allium Sativum, and Allium Ursinum
International Journal of Molecular Sciences Article Comparative Dissection of Three Giant Genomes: Allium cepa, Allium sativum, and Allium ursinum Vratislav Peška 1,* , Terezie Mandáková 2 , Veronika Ihradská 1,2,† and Jiˇrí Fajkus 1,2,* 1 Institute of Biophysics, The Czech Academy of Sciences, Královopolská 135, 612 65 Brno, Czech Republic; [email protected] 2 Mendel Centre for Plant Genomics and Proteomics, CEITEC, Masaryk University, Kamenice 5, CZ-62500 Brno, Czech Republic; [email protected] * Correspondence: [email protected] (V.P.); [email protected] (J.F.); Tel.: +420-541-517-199 (V.P.); +420-549-494-003 (J.F.) † Current address: Institute of Plant Genetics and Biotechnology, Plant Science and Biodiversity Center, Slovak Academy of Sciences, Akademická 2, P.O. Box 39A, 950 07 Nitra, Slovakia. Received: 4 January 2019; Accepted: 2 February 2019; Published: 9 February 2019 Abstract: Knowledge of the fascinating world of DNA repeats is continuously being enriched by newly identified elements and their hypothetical or well-established biological relevance. Genomic approaches can be used for comparative studies of major repeats in any group of genomes, regardless of their size and complexity. Such studies are particularly fruitful in large genomes, and useful mainly in crop plants where they provide a rich source of molecular markers or information on indispensable genomic components (e.g., telomeres, centromeres, or ribosomal RNA genes). Surprisingly, in Allium species, a comprehensive comparative study of repeats is lacking. Here we provide such a study of two economically important species, Allium cepa (onion), and A. sativum (garlic), and their distantly related A. -
Allium Cepa) Populations Possessing the Cytoplasm of Allium Galanthum
J. AMER. SOC. HORT. SCI. 124(6):626–629. 1999. Seed Yield, Floral Morphology, and Lack of Male-fertility Restoration of Male-sterile Onion (Allium cepa) Populations Possessing the Cytoplasm of Allium galanthum M.J. Havey1 Vegetable Crops Unit, Agricultural Research Service, U.S. Department of Agriculture, Department of Horticulture, University of Wisconsin, Madison, WI 53706 ADDITIONAL INDEX WORDS. cytoplasmic male sterility, alien cytoplasm, introgression ABSTRACT. The primary source (S cytoplasm) of cytoplasmic-genic male sterility (CMS) used to produce hybrid-onion (Allium cepa L.) seed traces back to a single plant identified in 1925 in Davis, California. Many open-pollinated populations also possess this cytoplasm, creating an undesirable state of cytoplasmic uniformity. Transfer of cytoplasms from related species into cultivated populations may produce new sources of CMS. In an attempt to diversify the cytoplasms conditioning male sterility, the cytoplasm of Allium galanthum Kar. et Kir. was backcrossed for seven generations to bulb-onion populations. The flowers of galanthum-cytoplasmic populations possess upwardly curved perianth and filaments with no anthers, making identification of male-sterile plants easier than for either S- or T- cytoplasmic male-sterile onion plants. Mean seed yield per bulb of the galanthum-cytoplasmic populations was measured in cages using blue-bottle flies (Calliphora erythrocephala Meig.) as pollinators and was not significantly different from one of two S-cytoplasmic male-sterile F1 lines, a T-cytoplasmic male-sterile inbred line, or N-cytoplasmic male-fertile lines. Male-sterile lines possessing either the S or galanthum cytoplasm were each crossed with populations known to be homozygous dominant and recessive at the nuclear locus conditioning male-fertility restoration of S cytoplasm and progenies were scored for male-fertility restoration. -
Michael Havey Cv
Michael John Havey USDA-ARS and Department of Horticulture University of Wisconsin, Madison, WI 53706 USA Telephone: 608-262-1830, Fax: 608-262-4743 Email: [email protected]; Lab URL: http://haveylab.hort.wisc.edu Education Degree Major Year Institution B.S. Plant Pathology 1977 Iowa State University M.S. Plant Pathology 1983 University of Wisconsin Ph.D. Plant Breeding-Plant Genetics and 1984 University of Wisconsin Plant Pathology Refereed Publications 1. Havey, M.J., and K.J. Frey. 1978. Optimal sample sizes and number per plot and replicate number for seed weights in oats. Cereal Res. Comm. 6:113-122. 2. Havey, M.J., and C.R. Grau. 1985. Decline of established alfalfa in soils naturally infested with Phytophthora megasperma f. sp. medicaginis and level of correlation with a seedling assay. Plant Dis. 69:221-224. 3. Havey, M.J., and D.P. Maxwell. 1987. Inheritance of Phytophthora root rot resistance in two diploid alfalfa species. Crop Sci. 27:225-228. 4. Havey, M.J., D.P. Maxwell, and J.A.G. Irwin. 1987. Independent inheritance of genes conditioning resistance to Phytophthora root rot from diploid and tetraploid alfalfa. Crop Sci. 27:873-879. 5. Havey, M.J., and D.P. Maxwell. 1988. Transfer of disease resistance from diploid to tetraploid alfalfa using unreduced female gametes. Plant Disease 72:603-604. 6. Havey, M.J., J.C. Faria, D.P. Maxwell, and D.J. Hagedorn. 1988. Partial resistance to anthracnose in Brazilian land races of dry beans shows race specificity. Euphytica 39:167-174. 7. Havey, M.J., and F.J. -
MOLEKÜLER MARKIR KULLANARAK BAZI YEREL SOĞAN (Allium Cepa L.) ÇEŞİTLERİNDE Ve ISLAH EDİLEN SAF HATLARDA SİTOPLAZMALARIN BELİRLENMESİ
MOLEKÜLER MARKIR KULLANARAK BAZI YEREL SOĞAN (Allium cepa L.) ÇEŞİTLERİNDE ve ISLAH EDİLEN SAF HATLARDA SİTOPLAZMALARIN BELİRLENMESİ Elif ÖZOKUTANOĞLU T.C ULUDAĞ ÜNİVERSİTESİ FEN BİLİMLERİ ENSTİTÜSÜ MOLEKÜLER MARKIR KULLANARAK BAZI YEREL SOĞAN (Allium cepa L.) ÇEŞİTLERİNDE ve ISLAH EDİLEN SAF HATLARDA SİTOPLAZMALARIN BELİRLENMESİ Elif ÖZOKUTANOĞLU Prof. Dr. Erdoğan BARUT Yrd. Doç. Dr. Ali Fuat GÖKÇE (Danışman) (İkinci Danışman) YÜKSEK LİSANS TEZİ BAHÇE BİTKİLERİ ANABİLİM DALI BURSA 2013 i TEZ ONAYI Elif Özokutanoğlu tarafından hazırlanan “Moleküler Markır Kullanarak Bazı Yerel Soğan (Allium Cepa L.) Çeşitlerinde ve Islah Edilen Saf Hatlarda Sitoplazmaların Belirlenmesi” adlı tez çalışması aşağıdaki jüri tarafından oy birliği ile Uludağ Üniversitesi, Fen Bilimleri Enstitüsü, Bahçe Bitkileri Anabilim Dalı’nda YÜKSEK LİSANS TEZİ olarak kabul edilmiştir. Danışman: Prof. Dr. Erdoğan Barut İkinci Danışman: Yrd. Doç. Dr. Ali Fuat Gökçe, Niğde Üniversitesi Başkan : Prof. Dr. Erdoğan BARUT Uludağ Üniversitesi Ziraat Fakültesi, Bahçe Bitkileri Anabilim Dalı Üye : Prof. Dr. H. Özkan SİVRİTEPE Uludağ Üniversitesi Ziraat Fakültesi, Bahçe Bitkileri Anabilim Dalı Üye : Doç. Dr. Himmet TEZCAN Uludağ Üniversitesi Ziraat Fakültesi, Fitopatoloji Anabilim Dalı Üye : Doç. Dr. Nuray AKBUDAK Uludağ Üniversitesi Ziraat Fakültesi, Bahçe Bitkileri Anabilim Dalı Üye : Yrd. Doç. Dr. Ali Fuat GÖKÇE Niğde Üniversitesi Tarım Bilimleri ve Teknolojileri Fakültesi, Genetik ve Uygulamalı Islah Anabilim Dalı Yukarıdaki sonucu onaylarım Prof. Dr. Ali Osman DEMİR Enstitü Müdürü -
1 Evolution, Domestication and Taxonomy
1 Evolution, Domestication and Taxonomy R.M. Fritsch1 and N. Friesen2 1Institut für Pflanzengenetik und Kulturpflanzenforschung, D-06466 Gatersleben, Germany; 2Botanischer Garten der Universität, D-49076 Osnabrück, Germany 1. The Genus Allium L. 5 1.1 General characteristics 5 1.2 Distribution, ecology and domestication 6 1.3 Phylogeny and classification 10 2. The Section Cepa (Mill.) Prokh. 14 2.1 Morphology, distribution and ecology 14 2.2 Cytological limitations 15 2.3 Grouping of the species 15 2.4 Enumeration of the species 16 3. Allium cepa L. 19 3.1 Description and variability 19 3.2 Infraspecific classification 20 3.3 Evolutionary lineages 21 3.4 History of domestication and cultivation 22 4. Other Economic Species 23 4.1 Garlic and garlic-like forms 23 4.2 Taxa of Asiatic origin 24 4.3 Chives and locally important onions from other areas 25 5. Conclusions 26 Acknowledgements 27 References 27 1. The Genus Allium L. controversy. In early classifications of the angiosperms (Melchior, 1964), they were 1.1 General characteristics placed in the Liliaceae. Later, they were more often included in the Amaryllidaceae, The taxonomic position of Allium and on the basis of inflorescence structure. related genera has long been a matter of Recently, molecular data have favoured a © CAB International 2002. Allium Crop Science: Recent Advances (eds H.D. Rabinowitch and L. Currah) 5 6 R.M. Fritsch and N. Friesen division into a larger number of small mono- • Ovary: trilocular, three septal nectaries of phyletic families. In the most recent and various shape, two or more curved competent taxonomic treatment of the (campylotropous) ovules per locule, monocotyledons, Allium and its close rela- sometimes diverse apical appendages tives were recognized as a distinct family, the (crests and horns); developing into a Alliaceae, close to the Amaryllidaceae. -
Allium Cepa L. Familie Der Liliengewächse (Liliaceae)
Quelle: http://www.mpiz- koeln.mpg.de/oeffentlichkeitsarbeit/kulturpflanzen/Nutzpflanzen/Kuechenzwiebel/index.html Allium cepa L. Familie der Liliengewächse (Liliaceae) links: Entwicklung der Zwiebel: A ein Jahr alt, B älter, rechts: verschiedene Zwiebelsorten Quelle: Rauh,W. Morph. der Nutzpfl., Quelle-Meyer Verlag, Heidelberg 1950; Wolf Garte Verbreitung: Die Küchenzwiebel wird in allen Erdteilen angebaut. Hauptproduktionsländer sind China, Indien, ex-UDSSR, USA, Türkei und Japan. Verlangt Licht, Wärme und nährstoffreiche, nicht zu trockene Böden im neutralen bis alkalischen Herkunft (rot) Anbau (grün) Bereich. Verwendung: Nahrungsmittel (Gewürz, Gemüse, Suppen, Kuchen) Reife Küchenzwiebeln enthalten 85-90 % Wasser, 7-10 % Kohlenhydrate, 1-2 % Eiweiß, 0,25 % Fett; hoher Vitamingehalt. Erträge und Produktion: Produktion (1000 t) Erträge (dt/ha) Land 1979-81 2005 1979-81 2005 China 2650 19050 125 211 India 2550 5500 102 104 USA 1625 36700 344 544 ex-USSR 1940 3870 109 131 Turkey 1020 2000 142 256 UK 223 365 321 324 Welt 21330 57910 132 182 Kultivierung und Züchtung: Die Küchenzwiebel ist eine alte Kulturpflanze der Steppengebiete des west- bis mittelasiatischen Raumes. Die ältesten Berichte über die wirtschaftliche Nutzung von Zwiebeln kommt aus Ägypten. Zwiebeln und Porree bildeten in der Zeit um 2800-2300 v. Chr. einen wichtigen Bestandteil der Nahrung der Pyramidenarbeiter. Mit der römischen Kultur fand die Küchenzwiebel in Europa Verbreitung. Aus dem 15. Jahrhundert liegen Hinweise über den Zwiebelanbau in Holland vor. Es wurden vielfältige Sorten gezüchtet, die sich in Form, Farbe und Geschmack unterscheiden. Als Gewürz werden hellgelbe und bräunlichgelbe, runde Sorten bevorzugt. Für Gemüse finden große, mildschmeckende Sorten Verwendung. Produktionstechnisch unterscheidet man Aussaat von Sommer- und Winterzwiebeln und Steckzwiebelanbau. -
Lowcountry Linguine Clams Oconee with Vidalia Onions & Bacon
ROMANCING THE CLAM, 23 March 2009, Savannah GA GEORGIA lams conee with idalia nions acon C O 4 tbsp of butter V HOeat pan over medium & heat. B Add This recipe is prepared by 1 medium Vidalia onion diced butter, swirling until melting. Add Chef Cal Berry of Berry’s (with green tops) chopped onions and minced garlic Catering in Augusta, 4 tsp minced garlic until onions are translucent. Add Georgia. 1 cup beer 1 cup white wine bacon, wine, beer, Boursin cheese and ½ cup crumbled crisp bacon sugar and bring to a boil. Add clams, ¼ cup parmesan cheese cover and cook until clams open, 5-7 2 tbsp minced basil leaf minutes. Remove clams and with a 2 tsp sugar slotted spoon scoop bacon and onion 36 littleneck clams mixture, spoon over open clams and 3 tbsp Boursin cheese sprinkle with parmesan cheese. Serves 4-5 Clams Provided by: Dish Prepared by: Charlie Phillips, Sapelo Seafarms, Rt. 1 Box 1672, Townsend, GA 31331; Chef Cal Berry, Berry’s Catering & Floral, 520 Reynolds Street, 912-832-4423; [email protected] Augusta, GA 30901; 706-724-2171; [email protected]; John Pelli, Savannah Clam Company, 9515 Whiteld Ave., Savannah, GA 31406; www.berryscatering.com 912-352-7102; [email protected] ROMANCING THE CLAM, 23 March 2009, Savannah GA SOUTHWASHINGTON CAROLINA Lowcountry Linguine This recipe is prepared by 8 oz country ham, julienned Add rst 6 ingredients to large sauté 2 cloves garlic, minced pan. Bring to boil. Reduce heat and Chef Peter Kornack of the 8 oz white wine T. W. -
INDUKCIJA HAPLOIDNIH GINOGENETSKIH REGENERANTOV ČEBULE (Allium Cepa L.) IZ KRIŽANJA LINIJ OH-1 X 5225B
UNIVERZA V LJUBLJANI BIOTEHNIŠKA FAKULTETA ODDELEK ZA AGRONOMIJO Barbara OVTAR INDUKCIJA HAPLOIDNIH GINOGENETSKIH REGENERANTOV ČEBULE (Allium cepa L.) IZ KRIŽANJA LINIJ OH-1 x 5225B DIPLOMSKO DELO Univerzitetni študij Ljubljana, 2008 UNIVERZA V LJUBLJANI BIOTEHNIŠKA FAKULTETA ODDELEK ZA AGRONOMIJO Barbara OVTAR INDUKCIJA HAPLOIDNIH GINOGENETSKIH REGENERANTOV ČEBULE (Allium cepa L.) IZ KRIŽANJA LINIJ OH-1 x 5225B DIPLOMSKO DELO Univerzitetni študij INDUCTION OF HAPLOID GYNOGENETIC REGENERANTS OF ONION (Allium cepa L.) FROM CROSSING OF LINES OH-1 x 5225B GRADUATION THESIS University Studies Ljubljana, 2008 Srce že ve zakaj solze sreče, ko gledam te. Foto in besedilo: A. Kralj & B. Ovtar Ovtar B. Indukcija haploidnih ginogenetskih regenerantov čebule (Allium cepa L.) iz križanja linij OH-1 x 5225B. II Dipl. delo. Ljubljana, Univ. v Ljubljani, Biotehniška fakulteta, Odd. za agronomijo, 2008 Diplomsko delo je zaključek univerzitetnega študija agronomije. Poskus je bil opravljen na Univerzi v Ljubljani, Biotehniški fakulteti, Oddelku za agronomijo, Katedri za genetiko, biotehnologijo in žlahtnjenje rastlin. Študijska komisija Oddelka za agronomijo je za mentorja diplomskega dela imenovala prof. dr. BOHANEC Boruta. Komisija za oceno in zagovor: Predsednik: akad. prof. dr. Ivan KREFT Univerza v Ljubljani, Biotehniška fakulteta, Oddelek za agronomijo Član: prof. dr. Borut BOHANEC Univerza v Ljubljani, Biotehniška fakulteta, Oddelek za agronomijo Članica: prof. dr. Marijana JAKŠE Univerza v Ljubljani, Biotehniška fakulteta, Oddelek za agronomijo Datum zagovora: Naloga je rezultat lastnega raziskovalnega dela. Podpisana se strinjam z objavo svoje naloge v polnem tekstu na spletni strani Digitalne knjižnice Biotehniške fakultete. Izjavljam, da je naloga, ki sem jo oddala v elektronski obliki, identična tiskani verziji Barbara OVTAR Ovtar B. -
Triparental Origin of Triploid Onion, Allium
Fredotović et al. BMC Plant Biology 2014, 14:24 http://www.biomedcentral.com/1471-2229/14/24 RESEARCH ARTICLE Open Access Triparental origin of triploid onion, Allium × cornutum (Clementi ex Visiani, 1842), as evidenced by molecular, phylogenetic and cytogenetic analyses Željana Fredotović1, Ivica Šamanić1, Hanna Weiss-Schneeweiss2, Juraj Kamenjarin1, Tae-Soo Jang2 and Jasna Puizina1* Abstract Background: Reconstruction of the parental origins of cultivated plants from wild relatives, especially after long periods of domestication, is not a trivial task. However, recent advances in molecular phylogenetics, among other approaches, have proved to be very informative in analyses of the origin and evolution of polyploid genomes. An established minor garden crop, triploid onion Allium × cornutum (Clementi ex Visiani, 1842) (2n =3x = 24), is widespread in southeastern Asia and Europe. Our previous cytogenetic analyses confirmed its highly heterozygous karyotype and indicated its possible complex triparental genome origin. Allium cepa L. and Allium roylei Stearn were suggested as two putative parental species of A. × cornutum, whereas the third parental species remained hitherto unknown. Results: Here we report the phylogenetic analyses of the internal transcribed spacers ITS1-5.8S-ITS2 of 35S rDNA and the non-transcribed spacer (NTS) region of 5S rDNA of A.×cornutum and its relatives of the section Cepa. Both ITS and NTS sequence data revealed intra-individual variation in triploid onion, and these data clustered into the three main clades, each with high sequence homology to one of three other species of section Cepa: A. cepa, A. roylei, and unexpectedly, the wild Asian species Allium pskemense B. Fedtsh. Allium pskemense is therefore inferred to be the third, so far unknown, putative parental species of triploid onion Allium × cornutum. -
Antimicrobial Activity Test of Genus Allium: a Review
Fadhilah Arifa et al. Int. Res. J. Pharm. 2020, 11 (12) INTERNATIONAL RESEARCH JOURNAL OF PHARMACY www.irjponline.com ISSN 2230 – 8407 Review Article ANTIMICROBIAL ACTIVITY TEST OF GENUS ALLIUM: A REVIEW Fadhilah Arifa, Anzharni Fajrina*, Aried Eriadi, Ridho Asra School of Pharmaceutical Science (STIFARM) Padang, Indonesia *Corresponding Author Email: [email protected] Article Received on: 4/12/20 Approved for publication: 31/12/20 DOI: 10.7897/2230-8407.1112102 ABSTRACT The genus Allium plants are often used by people as a food flavoring. These plants are also used for medicinal purposes. The genus Allium plants are known to inhibit the growth of microorganisms such as bacteria, fungi, viruses, and parasites by having the Allicin, Ajoene main compounds, and secondary metabolites. This review article describes plants from the genus Allium that have antimicrobial potential. In the writing process, this review article used literature study techniques by finding literature in the form of official books, national journals, and international journals in the last 10 years (2010-2020). The literature search in writing this review article was conducted through online media search with keywords as follows: antimicrobial, Allium, and inhibition zone diameter. The search for the main references in this review article was done through Google Scholar, ScienceDirect, Reserachgate, and other published journals. Some plants of the genus Allium such as Allium ascalonicum, Allium cepa, Allium chinense, Allium porrum, Allium roseum, Allium sativum, Allium staticiforme, Allium subhirsutum, Allium tuberosum, Allium tuncelianum, and Allium wallichii showed antimicrobial activity. The potential antimicrobial activity of each part of the plant depends on the solvent used, the concentration and levels of secondary metabolites contained therein. -
Taufiq Hidayat
Ilmu Pertanian (Agricultural Science) Vol. 3 No. 2 August, 2018: 66–71 Available online at http://journal.ugm.ac.id/jip DOI: doi.org/10.22146/ipas.26749 The Growth and Yield of Shallot ( Allium cepa L. Aggregatum group) in Application of Beneficial Microorganisms T aufiq Hidayat 1, 2 , Prapto Yudono 1, Endang Sulistyaningsih 1*, Arif Wibowo 3 1Department of Agronomy, Faculty of Agriculture, Universitas Gadjah Mada Jln. Flora no. 1, Bulaksumur, Sleman, Yogyakarta 5528, Indonesia 2Department of Agrotechnology, Faculty of Agriculture, Universitas Muhammadiyah Yogyakarta JL. Brawijaya, Kasihan, Bantul, Yogyakarta 55183 3Department of Phytopatholoy, Faculty of Agriculture, Universitas Gadjah Mada Jln. Flora no. 1, Bulaksumur, Sleman, Yogyakarta 5528, Indonesia *Corresponding email: [email protected] Received: 19 th July 2017; Revised: 21 st March 2018; Accepted: 05 th May 2018 ABSTRACT Shallot ( Allium cepa L. Aggregatum group) is one of the most widely utilized vegetables in Indonesia. Some technologies have been adapted to improve its productivity, e.g. the application of beneficial microorganisms. Mycorrhizal fungi, Trichoderma sp., and Bacillus thuringiensis as well as the combination of these microorganisms were applied on shallot cultivations. The experiment aims to investigate the effect of those microorganisms on the growth and yield of shallot. The experiment was arranged in a Randomized Complete Block Design with two treatment factors: (1) two shallot cultivars, Biru Lancor and Crok Kuning and (2) six beneficial microorganism treatments, i.e., control, mycorrhizae, Trichoderma sp., Bacillus thuringiensis , mycorrhizae and Trichoderma sp., combination, and those three microorganism combination. The objective was to study the effect of those microorganism application and their combination on the improvement of growth and yield of Crok Kuning and Biru Lancor cultivars. -
Morphological, Cytological and Embryological Characterization of F1 A
ACTA BIOLOGICA CRACOVIENSIA Series Botanica 57/2: 98–105, 2015 DOI: 10.1515/abcsb-2015-0025 MORPHOLOGICAL, CYTOLOGICAL AND EMBRYOLOGICAL CHARACTERIZATION OF F1 A. CEPA × A. ROYLEI HYBRIDS ALICJA CHUDA*, KAROLINA KŁOSOWSKA AND ADELA ADAMUS Unit of Genetics, Plant Breeding and Seed Science, Institute of Plant Biology and Biotechnology University of Agriculture in Krakow, Al. 29 Listopada 54, 31-425 Kraków, Poland Received July 28, 2015; revision accepted November 17, 2015 In the previous study we obtained a population of interspecific F1 A. cepa × A. roylei hybrids. In this study, in comparison to the parental species: A. cepa and A. roylei, the F1 hybrids were evaluated in terms of plant mor- phology, pollen viability, microsporogenesis and female gametophyte. Most of the morphological characters of the F1 hybrids were intermediate as compared to those of both parental accessions. In pollen mother cells (PMCs) of the F1 hybrids abnormalities were observed in meiosis as well as at the tetrad stage. Pollen viability of F1 A. cepa × A. roylei hybrids was reduced to 30.1%. In the F1 hybrids, 45.8% of the analyzed ovules showed developmental disturbances, whereas in 26.7% of the ovules necrotic processes were observed. Key words: Allium cepa, A. roylei, embryo sac, interspecific hybrids, microsporogenesis, morpho- logical characters, pollen viability INTRODUCTION al., 2007). The first sexual hybridization between Allium cepa and A. fistulosum was reported by Interspecific hybridization has always been consid- Emsweller and Jones (1935). However, backcross- ered an important tool for broadening the genetic ing and successful introgression of desirable traits variation of cultivated Allium species (Kik, 2002; such as resistance to pink root and onion leaf blight, Chuda and Adamus, 2009).