Research Article

Ecologica Montenegrina 22: 50-89 (2019) This journal is available online at: www.biotaxa.org/em

Faunistic diversity of the spiders in Montenegro (Arachnida: Araneae)

MARIA NAUMOVA1*, STOYAN LAZAROV2, CHRISTO DELTSHEV2

1Institute of Biodiversity and Ecosystem Research, Bulgarian Academy of Sciences, Sofia, Bulgaria; E-mail: [email protected] 2 National Museum of Natural History, Bulgarian Academy of Sciences, Sofia, Bulgaria; E-mail: [email protected]; [email protected]

Received 30 June 2019 │ Accepted by V. Pešić: 2 August 2019 │ Published online 21 August 2019.

Abstract Although the European fauna of spiders is relatively well studied, in the Balkans still have places for which little or almost nothing is known, especially Kosovo, Bosnia & Herzegovina and Montenegro. The spider fauna of Montenegro is still poorly known, and the number of relevant publications is very low, even with catalogs and quoted information is about 90. Only for the cave fauna has more in-depth observations. The aim of this study is to summarize and presents all available data of the spiders of Montenegro, both from the literature and from original records. A total of 144 spider species were collected and after incorporating literature reports, the number of identified species is 289, from 38 families. From them 84 species and two families are new for the country. The presented study provides the first annotated checklist and catalog of the spiders in Montenegro (as a result of a database with 1150 rows/records) with additional taxonomic and faunistic data.

Key words: annotated catalog, Balkan Peninsula, checklist, taxonomy, photos.

Introduction

The Republic of Montenegro is located in southeastern Europe and is a territory with high biodiversity owing to the variety of climate, topography, and geology found here, and the long-term ecological and evolutionary history of this region as a biological crossroads and Ice Age refuge. This suggests a rich araneofauna, but the available data at the moment are very scarce. The first report about the spiders in Montenegro (according to Damin 1900) probably came before the half of the 19th century, from Carrara (1846), but the first sure records came from Doleschall (1852) and Doblika (1853), followed by L. Koch (1868), Canestrini & Pavesi (1868), Pavesi (1876), Simon (1878, 1882), Castelli (1891), Gasperini (1891) and Chyzer & Kulczyński (1891, 1897). The first list of the spiders in Montenegro can be found in Damin (1896, 1900) who presented a total of 66 species, followed by Drensky (1936) with 77 species and Nikolić & Polenec (1981) with 86 species. Later, within about of 50 publications, can be found mostly taxonomic information about Montenegro’s spiders, but till now not any review of the araneofauna of this part of the Balkans. These data are comprised also in the paper of Deltshev (1999), who mentioned 102 species and later (Deltshev 2008) listed 43 species in caves.

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This review is a critical incorporation of the available literature data, and unpublished records due to sporadic research from 2002 to 2015 year. We excluded all the records from Dalmatia and Yugoslavia, for which cannot certainly be argued that relate to Montenegro. All reports, contained any sure data on the Montenegrin spider fauna were included in a database, which consist 1150 rows/records. The aim of this study is to present annotated checklist and catalog of the spiders of Montenegro with additional taxonomic and faunistic data.

Materials and methods

The material treated herein comes from two major sources. The first part comprises critical incorporation of all available literature records concerning the distribution of spiders in Montenegro (953 database records). The second part comprises original collections made in the period 2002–2015 during from different colleagues (196 database records). The studied original material was collected by hand picking, under stones, by sweeping and sieving and with pitfall traps (with formaldehyde). The specimens were preserved in 70– 80% ethanol and deposited in the Institute of Biodiversity and Ecosystem Researches (IBER) and the National Museum of Natural History (NMNH) at the Bulgarian Academy of Sciences. The digital images were taken with a Lumix digital camera attached to Wild M5A stereomicroscope and prepared with Photoshop CS6 software. The localities (Fig. 1) were mapped on the basis of exact geographic coordinates (decimal), rounded to 4 decimal places (when they are taken in the field with a GPS-receiver), or with approximate coordinates, rounded to 3 decimal places (on the basis of the location of the settlements/geographic objects, which names are given in the older literature). The list of localities (alphabetically by municipalities) includes the sites with numbers 1 to 77 (with exact or approximate coordinates; mapped), followed by sites in Krivosije plateau (lested as 78 to 83, but visualisated with number 78), and followed by sites 84 to 94 (without precise locality; not mapped). Mapping and visualisation of the map were done by ArcGIS 10.1 (ESRI, Redlands, California, USA). The nomenclature follows the World Spider Catalog (2019) and all taxa are listed alphabetically. All new taxa are marked with asterisk (*). The literature sources are listed chronologically (except in cases where the authors have more than one cited publications – then the years are listed after the first mention). The general distribution of the species is given mainly according to WSC 2019 and Nentwig et al. 2019. The additional sources were citing. Abbreviations used in the text include: ♂–male; ♀–female; j/jj–juvenile/s; L–locality (with number in the List of localities); UM–University of Montenegro, Podgorica.

List of localities (L)

Andrijevica 1. Maĉak village, Komovi Mts., N42.6947˚, E19.6795˚, 1898 m a.s.l., 11.08.2015, coll. G. Hristov & A. Lambevska. 2. Trešnjevik village, Komovi Mts., N42.7339˚, E19.6851˚, 1574 m a.s.l., 09.08.2015, coll. S. Lukanov. Bar 3. Bar (Antivari), N42.107˚, E19.112˚, 16 m a.s.l. 4. Bar area, Sustaś village, N42.110˚, E19.124˚, 200 m a.s.l. 5a. Bes village, Rumija Mts., near a stream limestone rocks/almond trees; 5 soil traps, N42.1013˚, E19.1446˚, 250 m a.s.l., 07-14.05.2004, coll. M. Langourov. 5b. Bes village, Rumija Mts., near a stream limestone rocks/almond trees; 5 tree traps, N42.1013˚, E19.1446˚, 250 m a.s.l., 07-14.05.2004, coll. M. Langourov. 6a. Donji Murići village, Skadarsko Jezero lake, N42.1622˚, E19.2206˚, 15 m a.s.l., 11.05.2004, coll. M. Langourov. 6b. Đuravci village, above the church, Rumija Mts., N42.1753, E19.1862, 370 m a.s.l., 10-14.05.2004, coll. M. Langourov. 7. Lakiĉević pećina (Krivošije), N42.263˚, E19.033˚, 428 m a.s.l. 8. Peĉurice village (1 km W of; Olea europea L.), Rumija Mts., 10 soil traps, N42.0238˚, E19.1660˚, 230 m a.s.l., 06-14.05.2004, coll. M. Langourov.

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9a, Virpazar, Babatuša cave (Bobotuša, Baba Tuša, Babatusha), Trnovo village, N42.2292˚, E19.0955˚, 350 m a.s.l. 9b. Virpazar, Boljevići pećina, Central MN, N42.219˚, E19.085˚, 13 m a.s.l. 10a. Virpazar, Golubova pećina cave, Gornja Seoca village, Dragaljasko polje, Gornje Krivošije, N42.209˚, E19.131˚, 440 m a.s.l. 10b. Virpazar, Špilja Goluspa, Donja Seoca village, N42.209˚, E19.131˚, 435 m a.s.l. Berene 11. Đato pećina, Petnik near Berane, N42.822˚, E19.898˚, 870 m a.s.l. Budva 12. Budva, N42.296˚, E18.855˚, 163 m a.s.l. 13a. Paštrovska gora (Petrovaĉka gora) Mts., Petrovac-Virpazar, N42.214˚, E18.975˚, 600 m a.s.l. 13b. Petrovac, Budva, N42.208˚, E18.955˚, 124 m a.s.l. 13c. Rumija Mts., route Petrovac-Virpazar, N42.214˚, E18.976˚, 627 m a.s.l. Cetinje 14a. Cetinje, N42.401˚, E18.934˚, 665 m a.s.l. 14b. Cetinjska pećina, N42.401˚, E18.935˚, 798 m a.s.l. 15a. Crni nugli village, Dragaljsko (=Dvrnsko) polje, Ĉora pećina cave (Pećina u Selakovom dolu), Selakov dol, Risan distr., Gorno Krivošije, N42.5861˚, E18.6969˚, 590 m a.s.l. 15b. Crni nugli village, Dragaljsko (=Dvrnsko) polje, unnamed cave, Risan distr., Gornje Krivošije, N42.5861˚, E18.6969˚, 590 m a.s.l. 15c. Crni nugli village, Dragaljsko (=Dvrnsko) polje plain, N42.5951˚, E18.6999˚, 650 m a.s.l., 27. 03 2006, coll. S. Lazarov & B. Petrov. 16. Dragaljsko polje, Golubova Pecina, Selakov Dol, Gornje Krivošije, N42.6197˚, E18.6889˚, 625 m a.s.l. 17. Golubnjaĉka pećina, Lovćen planina Mts., N42.397˚, E18.841˚, 1463 m a.s.l. 18. Lipska pećina cave, Lipa village, N42.375˚, E18.952˚, 528 m a.s.l. 19. Rumija Mts., Obzovica village, Budva-Cetinje, 42.324˚, E18.925˚, 700 m a.s.l. 20a. Virpazar distr., Trnovo village, Špilia cave, N42.293˚, E19.036˚, 360 m a.s.l. 20b. Virpazar distr., Trnovo village, Trnovo-Komarno, Grboĉica pećina, N42.290˚, E19.042˚, 340 m a.s.l. 20c. Virpazar distr., Trnovo village, under stones, N42.2941˚, E19.0375˚, 340 m a.s.l., 25. 03. 2006, coll. Lazarov & B. Petrov. Danilovgrad 21. Prekornica Mts., Gornji Martinici village, N42.5604˚, E19.2112˚, 692 m a.s.l., 06-22. 08. 2002, coll. V. Pešić & D. Pešić. 22. Prekornica Mts, Kamenik Mt, env., Konjic hill, N42.5799˚, E19.2131˚, 994 m a.s.l., 01.08.2007, coll. R. Mlejnek. Gusinje 23a. Prokletije Mts., N42.499˚, E19.818˚, 1215 m a.s.l. 23b. Prokletije Mts., Koruns cave, Katun Bjelic, N42.538˚, E19.854˚, 1612 m a.s.l. 23c. Prokletije Mts., Ropojana valley, N42.499˚, E19.818˚, 1200 m a.s.l. Herceg Novi 24a. Herceg Novi (Castelnovo, Castelnuovo), N42.457˚, E18.532˚, 145 m a.s.l. 24b. Herceg Novi, Monastir Savina (Kotor distr.), N42.452˚, E18.554˚, 50 m a.s.l. 25a. Orjen Mts., N42.591˚, E18.556˚, 1515 m a.s.l. 25b. Pećina na Pržini, near Sveĉava vill, Orjen Mts., N42.591˚, E18.556˚, 1515 m a.s.l. 26. Radoštak, N42.492˚, E18.571˚, 1230 m a.s.l. 27. Vucja pećina cave near Vrbanje village, Orjen Mts., N42.551˚, E18.511˚, 1047 m a.s.l. 28. Zelenika, N42.456˚, E18.589˚, 120 m a.s.l. Kolašin 29. Kolašin, N42.804˚, E19.518˚, 1000-1100 m a.s.l. Kotor 30a. Crkvice, artificial gallery on the road to Han village, Risan distr., Dolno Krivošije, N42.564˚, E18.633˚, 1090 m a.s.l. 30b. Crkvice, Blagojevo village, Risan districtit, (near house of Shuto Blagojevic), N42.5538˚, E18.6177˚, 1040 m a.s.l., 17.08.2006, S. Lazarov & B. Petrov.

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30c. Crkvice, Pećina kod Blagojevića, 1 km W of Blagojević village, Dolnje Krivošije, N42.563˚, E18.620˚, 1042 m a.s.l. 30d. Crkvice, potholes, near Shuto Blagojević monastery, Crkvice village, Risan distr., Dolno Krivošije, N42.568˚, E18.623˚, 1045 m a.s.l. 30e. Crkvice, Vilna pećina, (ViIina pecina u Napode), Krivošije, N42.560˚, E18.629˚, 1075 m a.s.l. 31, Dobrota (Kotor), 42.455, E18.769, 100 m a.s.l. 32. Izeta pećina cave, Knezlać village, Risan distr., Krivosije (in the foothills of Mt. Orjen), N42.524˚, E18.646˚, 934 m a.s.l. 33a. Kotor (valley), N42.488˚, E18.767˚, 52 m a.s.l. 33b. Kotor, Skalni Prasklina split, N42.488˚, E18.767˚, 55 m a.s.l. 34a. Krivošije, Bor pećina, Han village, Dragalj, N42.569˚, E18.662˚, 910 m a.s.l. 34b. Krivošije, Lopata pećina, N42.569˚, E18.662˚, 908 m a.s.l. 34c. Krivošije, Matjaševica pećina, N42.569˚, E18.662˚, m a.s.l. 34d. Krivošije, Nilova pećina near Han village, Krivošije Mts., N42.569˚, E18.662˚, 908 m a.s.l. 34e. Krivošije, Pećina u Ivici, N42.569˚, E18.662˚, 908 m a.s.l. 34f. Krivošije, Strmena pećina near Han village, Krivošije Mts., N42.564˚, E18.633˚, 1100 m a.s.l. 35. Muo, N42.423902, E18.753247, 315 m a.s.l. 36. Pećina (pećinis) kod Napode, (cave near Napode, Careve Doline, Terbulov Do), Krivošije, N42.5192˚, E18.7160˚, 890 m a.s.l. 37a. Risan distr., cave, N42.560˚, E18.695˚, 90 m a.s.l. 37b. Risan distr., Golodražnica cave, N42.535˚, E18.689˚, 570 m a.s.l. 37c. Risan distr., Jamutina pećina, Kotor valley, N42.527˚, E18.692˚, 430 m a.s.l. 37d. Risan distr., Lednice village, under stones, N42.528˚, E18.706˚, 570 m a.s.l., 26.03.2006, coll. S. Lazarov & B. Petrov. 37e. Risan distr., Pokljuka Dolnja cave, Dolno Krivošije, N42.542˚, E18.693˚, 585 m a.s.l. 37f. Risan distr., Pokljuka Gornja cave, Dolno Krivošije, N42.542˚, E18.695˚, 580 m a.s.l. Mojkovac 38. Biogradska gora, SW Bijelo Polje, N42.929˚, E19.584˚, 1040-1200 m a.s.l. 39. Biogradsko jezero, Bjelasica Mts., N42.902˚, E19.591˚, 1000 m a.s.l. Nikšić 40. Bijela pećina, Stubica, N42.695˚, E18.982˚, 565 m a.s.l. 41. Boraĉka pećina, Borak, Ubla, N42.945˚, E18.523˚, 1272 m a.s.l. 42a. Carev Most, Budoška pećina, N42.726˚, E18.961˚, 613 m a.s.l. 42b. Carev Most, Velja pećina, N42.726˚, E18.961˚, 780 m a.s.l. 43. Cistavlada pećina cave, Velimlje village, N42.822˚, E18.642˚, 820 m a.s.l. 44a. Đakovića pećina cave (Pećina Vojvode Đakovića), Grahovo village, Gorno Krivošije, N42.6558˚, E18.6758˚, 707 m a.s.l. 44b. Đakovica pećina (Pećina Vojvode Đakovića) cave, outside, Grahovo village, Gorno Krivošije, N42.6562˚, E18.6774˚, 700 m a.s.l., 28.03.2006, coll. S. Lazarov & B. Petrov. 45. Jasenovo Polje, N42.9069˚, E18.9634˚, 1032 m a.s.l., 07.08.2015, coll. S. Lukanov. 46a. Krstac (Crni Vrh), Kerovaćka pećina cave near Nešište, N43.003˚, E18.710˚, 966 m a.s.l. 46b. Krstac (Crni Vrh), Stoška pećina, Javljen, N43.008˚, E18.709˚, 950 m a.s.l. 47. Ledena pećina na Kiti (Kita) cave, Trubjela, N42.738˚, E18.784˚, 1000 m a.s.l. 48a. Osjeĉenice, Bobjerska pećina, Grahovo village, N42.674˚, E18.662˚, 997 m a.s.l. 48b. Osjeĉenice, Elazova (Elesova) pećina cave, Krivošije, Zagora (Osjeĉenice), N42.706˚, E18.673˚, 1007 m a.s.l. 48c. Osjeĉenice, Kaloperska pećina, Kaloper Mts., Krivošije, N42.688˚, E18.656˚, 1100 m a.s.l. 48d. Osjeĉenice, Zagorska pećina, N42.688˚, E18.656˚, m a.s.l. 49. Pećina Kućarada, Vir village, N42.842˚, E18.912˚, 770 m a.s.l. 50a. Pećina na Troglav I, Rudine-Podbožur, N42.743˚, E18.691˚, 872 m a.s.l. 50b. Pećina na Troglav II, N42.743˚, E18.691˚, 872 m a.s.l. 51. Sirbaba cave, Podkita village, Gorno Krivošije, N42.7863˚, E18.8249˚, 940 m a.s.l., 28.03.2006, coll. S. Lazarov & B. Petrov. 52. Studenićka pećina cave, Nikšić distr., N42.826˚, E18.786˚, 900 m a.s.l. 53a. Vraćenovići (Bileca), Gnjatova pećina, Vuĉi Do, N42.836˚, 1E8.505˚, 840 m a.s.l.

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53b. Vraćenovići (Bileca), Stanina pećina, Vuĉi Do, N42.840˚, E18.502˚, 850 m a.s.l. 53c. Vraćenovići (Bileca), Vranova jama, Troskarica Planina Mts., N42.838˚, E18.551˚, 1295 m a.s.l. Pljevlja 54. Franjova Ledena pećina cave, N43.302˚, E19.353˚, 915 m a.s.l. 55a. Tara river gorge, NP Durmitor Mts., N43.233˚, E19.134˚, 800 m a.s.l. (in Thaler 1978, Kropf 1998). 55b. Tara river gorge, NP Durmitor Mts., N43.2326˚, E19.1342˚, 800 m a.s.l., 13-18.07.2003, coll. S. Lazarov & M. Langourov. 56a. Tepća village, Devoyacki Most bridge,Tara river gorge, NP Durmitor Mts., N43.2184˚, E19.0803˚, 588 m a.s.l., 13-21. 07 2003, coll. S. Lazarov & M. Langourov. 56b. Tepća village, NP Durmitor Mts., N43.2083˚, E19.0748˚, 730 m a.s.l., 13-21.07.2003, coll. S. Lazarov & M. Langourov. 56c. Tepća village, Yavorite place, Komarnica river gorge, NP Durmitor Mts., N43.2042˚, E19.0952˚, 1310 m a.s.l., 29. 07.2003, coll. S. Lazarov. 56d. Tmuša cave, Tara river gorge, NP Durmitor Mts., N43.2184˚, E19.0803˚, 588 m a.s.l., 17.07.2003, coll. S. Lazarov & M. Langourov. Pluţine 57. Bajovo polje, Bundos, Golija Mts., N43.017˚, E18.899˚, 1050-1450 m a.s.l. 58. Crnogorski Maglić peak, Maglić Mts., N43.272˚, E18.734˚, 2000 m a.s.l. 59a. Komarnica river gorge, Kešina place, NP Durmitor Mts., N43.1464˚, E18.9972˚, 1470 m a.s.l., 20- 23.07.2003, coll. S. Lazarov. 59b. Komarnica river gorge, Sopot spring, NP Durmitor Mts., N43.1385˚, E18.9936˚, 1580 m a.s.l., 20.07.2003, coll. S. Lazarov. 60. Trnovaĉko Jezero Lake, N43.251˚, E18.731˚, 1700 m a.s.l. Podgorica 61. Daljam village near Podgorica, Moraĉa gorge, N42.4880˚, E19.2141˚, 156 m a.s.l., malaise trap, coll. V. Pešić. 62. Pećina kod Vraninjo Brdo (Cave near Vraninjo Hill, Vraninjo village, Skadar lake), N42.272˚, E19.135˚, 165 m a.s.l. 63a. Podgorica (Titograd), Moraĉa gorge, N42.477˚, E19.317˚, 430-760 m a.s.l. 63b. Podgorica (Titograd), Pećina od Zavora, N42.472˚, E19.259˚, 54 m a.s.l. 64a. Podgorica (Titograd), Dućica pećina cave near Peuta, N42.467˚, E19.364˚, 550 m a.s.l. 64b. Podgorica (Titograd), Fundina, Pećina u pjesatici, (Ćafa Pješatica Cave, Fundine, near Medun), N42.467˚, E19.364˚, 550 m a.s.l. 64c. Podgorica (Titograd), Hani Grabom-Podgorica, N42.489˚, E19.401˚, 478 m a.s.l. Roţaje 65. Bisevo (Rožaje), N42.913˚, E20.216˚, 1233 m a.s.l. 66. Bukovica, N42.940˚, E20.218˚, 1090 m a.s.l. 67. Mokra Gora Mts., 11 km S of Rožaje, N42.778˚, E20.196˚, 1100 m a.s.l. 68. Rožaje-Peć, small cave near the road Rožaje-Peć, N42.782˚, E20.258˚, 1520 m a.s.l. Šavnik 69a. Skorupna Jama cave near Slatina, Ljubishnja Mts., env., Krsa Mts., N42.9862˚, E19.1472˚, 1640 m a.s.l., 22.09.2009, coll. R. Mlejnek. 69b. Snježnica cave, Vojnik Mts., N42.998˚, E19.146˚, 1800 m a.s.l. Tivat 70. Stradioti (San Marko) isl., N42.411˚, E18.689˚, 3 m a.s.l. Ulcinj 71. Drin, N41.992˚, E19.379˚, 3 m a.s.l. 72a. Klezna place, Rumija Mts., N41.9800˚, E19.2819˚, 80 m a.s.l., 06-14.05.2004, coll. M. Langourov. 72b. Klezna place, N41.965˚, E19.247˚, 200 m a.s.l. 73. Pećina kod Jame, Gorana, Rumija Mts., N41.999˚, E19.230˚, 207 m a.s.l. Ţabljak 74a. Bobotov Kuk peak, NP Durmitor Mts., N43.1285˚, E19.0345˚, 2480-2523 m a.s.l., 21.07.2003, coll. S. Lazarov. 74b. Bobotov Kuk, Crno Jezero-Bobotov Kuk, under stones, N43.1384˚, E19.0536˚, 1900-2100 m a.s.l., 18.08.2006, coll. B. Petrov.

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Figure 1. Map with localities of the spiders in Montenegro. For details, see List of localities.

75a. Crno Jezero lake, NP Durmitor Mts., N43.1463˚, E19.0875˚, 1400-1500 m a.s.l., 18.07.2007, coll. S. Lazarov. 75b. Crno Jezero-Ledena pećina, NP Durmitor Mts., N43.1439˚, E19.0791˚, 1660 m a.s.l., 18.08.2006, coll. B. Petrov.

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75c. Ivan Do village, NP Durmitor Mts., N43.1573˚, E19.0919˚, 1496 m a.s.l., 04.08.2015, coll. S. Lukanov. 76. Modro Jezero lake, NP Durmitor Mts., N43.0832˚, E19.0719˚, 1800 m a.s.l., 02.09.2010, coll. P. Beron. 77a. Virak village, Dobri Dol, NP Durmitor Mts., N43. 1102˚, E19. 0899˚, 1770-1900 m a.s.l., 22.07.2003, coll. S. Lazarov. 77b. Virak village, Vodena pećina cave, NP Durmitor Mts., N43.1126˚, E19.1042˚, 1550 m a.s.l. Additional localities Krivošije (The massive is a part of different municipalities such as Bar, Cetinje, Kotor and Nikšić; mapped with aprox. coordinates N42.624˚, E18.664˚ and number 78) 78a. Krivošije. 78b. Krivošije, caves. 79. Krivošije, Leskova pećina. 80. Krivošije, Meradova pećina. 81. Krivošije, Mujkovica (Mijukovica) pećina. 82a. Krivošije, pećina kod Dvorecka zdrijela. 82b. Krivošije, pećina na Jankovim (Jankovom) vrhu. 83. Krivošije, pećina u Manitoj Rupi, Ledenice. Montenegro (without precise locality, not mapped) 84. cave, S Monténégro 85. Deljina pećina, E Montenegro 86. Deveriĉka pećina 87. Jankova pećina 88. Ledenice (probably in Bosnia?) 89. Montenegro, without precise locality 90. Pećina kod Šunjevca 91. Pećina na Velu gredu 92. Studena pećina 93. Tomova pećina 94. Two caves, E MN

Results and discussion

Species composition. The study comprises 289 species belonging to 38 families. Of them 84 species and two families are new for Montenegrin spider fauna (marked in the list with *). The number of the established species is not high and does not represent a real picture of the faunistic diversity of the spiders occurring in the region. This is also evident from a comparison with the number of spiders recorded from other countries in Balkan Peninsula, such as Bulgaria (1043 species, Blagoev et al. 2018), Greece (1167 species, Chatzaki et al. Version 1.0), Republic of North Macedonia (820 species, Nentwig et al. 2019) and Serbia (725 species, Deltshev et al. 2003, Nentwig et al. 2019) Two species are excluded from the check list as nomina dubia: Stalita schioedtei Thorell, 1870 (Dysderidae) and Lepthyphantes tenoides Kolosvary, 1938 (Linyphiidae), and four species are excluded as erroneous identification: Cybaeus brignolii Maurer, 1992 (Cybaeidae), Domitius speluncarum (Pavesi, 1873) (Nesticidae), Kryptonesticus fagei (Kratochvíl, 1933) (Nesticidae) and Dolomedes fimbriatus (Clerck, 1757) (Pisauridae).

Annotated checklist & catalog of Montenegrin spiders

Agelenidae 1. Agelena labyrinthica (Clerck, 1757) Literary data: L31 (Castelli 1891), L31, sub Agalena l. (Damin 1896, 1900, Drensky 1936). New data: L56b (1 ♂), L56c (1♀). Global distribution: Europe to Central Asia, China, Korea, Japan.

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2. *Eratigena agrestis (Walckenaer, 1802) New data: L56b (1♂). Global distribution: Europe to Central Asia. Introduced to USA, Canada.

3. Eratigena fuesslini (Pavesi, 1873) Literary data: L28, sub Tegenaria ericarum Simon, 1875 (Kolosvary 1938), sub nomen nudum Tegenaria corsica Bremi-Wolff, 1849 (Nikolic & Polenec 1981). Remark: unsure after Bolzern et al. (2013). Global distribution: Europe, Turkey.

4. Histopona conveniens (Kulczyński, 1914) Literary data: L43, L46a, L64a (Deeleman-Reinhold 1983), L89 (Deltshev 2008, Mammola et al. 2018). New data: 37d (1 ♂), 57b (1♀). Global distribution: Bosnia-Hercegovina; Croatia, Serbia.

5. Histopona dubia (Absolon & Kratochvíl, 1933) Literary data: L27 (Deeleman-Reinhold 1983), L89 (Deltshev 2008, Mammola et al. 2018). Global distribution: Croatia, Bosnia-Hercegovina, Serbia.

6. Histopona krivosijana (Kratochvíl, 1935) Literary data: L15a, L30c, L34b, L34c, L34e, L48b, L77b, L 82b (Kratochvíl 1935b, 1938a), L16 [locus typicus] sub Hadites bidens krivosijanus (Kratochvíl 1935b), L16, sub Hadites krivosijanus (Kratochvíl 1938a), L16, sub Roeweriana k. (Kratochvíl 1978), L34f (Deeleman-Reinhold 1983), L89, sub Hadites krivosijanus (Nikolić & Polenec 19810, L89 (Deltshev 2008, Mammola et al. 2018), L92, sub Hadites bidens krivosijanus (Kratochvíl 1935b). Global distribution: Croatia, Montenegro.

7. Histopona luxurians (Kulczynski, 1897) Literary data: L89 (Deeleman-Reinhold 1983). Global distribution: Austria to Ukraine and South-eastern Europe.

8. Histopona palaeolithica (Brignoli, 1971) Literary data: L10a (Naumova et al. 2016). Global distribution: Italy.

9. *Inermocoelotes falciger (Kulczyński, 1897) New data: L56c (1♀). Global distribution: Eastern Europe.

10. *Inermocoelotes gasperinii (Simon, 1891) New data: L44b (2♀♀). Remark: The species was described from Dalmatia (Croatia). Morphologically, our material corresponds well to the description of Inermocoelotes gasperini and the presented photos of the female contribute to the taxonomic characteristic of the species (Figs. 2–4). Global distribution: Croatia.

11. Inermocoelotes inermis (L. Koch, 1855) Literary data: L8, sub Caelotes [sic] i. (L. Koch 1868, Pavesi 1876), sub, Coelotes i. (Nosek 1904, Bresjanceva 1907, Drensky 1936). Global distribution: Europe.

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Figures 2–4. Inermocoelotes gasperinii. 2 – epigyne, ventral view, 3 – vulva, ventral view, 4 – vulva, dorsal view. Scale: 0.2 mm.

12. Inermocoelotes karlinskii (Kulczyński, 1906) Literary data: L75a, sub Eurocoelotes k. (Wang et al. 2010). New data: L1 (1♀, 1sa♀), L75a (1♂, 1♀, 4jj). Global distribution: South-eastern Europe.

13. Lycosoides coarctata (Dufour, 1831) Literary data: L70, sub Textrix c. (Brignoli 1966). Global distribution: Mediterranean.

14. Tegenaria animata Kratochvíl & Miller, 1940 Literary data: L7, L9a [locus typicus] (Kratochvíl & Miller 1940), L9a (Nikolić & Polenec 1981), L15b, L44a, L51 (Naumova et al. 2016), L89 (Roewer 1955, Bolzern et al. 2013, Mammola et al. 2018), L89, sub Pseudotegenaria a. (Deltshev 2008). Global distribution: Albania (Deltshev et al. 2011), Montenegro, Republic of North Macedonia, Serbia.

15. Tegenaria annulata Kulczynski, 1912 Literary data: L89 (Kratochvíl, 1935b, Nikolić & Polenec 1981, Deltshev 2008, Bolzern et al. 2013, Mammola et al. 2018) Global distribution: Bosnia-Hercegovina, Croatia, Montenegro, Serbia.

16. Tegenaria bayeri Kratochvíl, 1934 Literary data: L9a (Naumova et al. 2016), L30c, L30e, L34e, L93 (Kratochvíl 1935b, Drensky 1936), L78a (Nikolić & Polenec 1981), L89 (Deltshev 2008, Bolzern et al. 2013, Mammola et al. 2018). Global distribution: Bosnia-Hercegovina, Montenegro, Serbia.

17. Tegenaria bosnica Kratochvíl & Miller, 1940 Literary data: L89 (Bolzern et al. 2013). Global distribution: Albania (Deltshev 2008), Bosnia-Hercegovina, Croatia, Montenegro, Serbia.

18. Tegenaria dalmatica Kulczynski, 1906 Literary data: L28 (Kulczyński 1906, Drensky 1936, Nikolić & Polenec 1981), L89 (Deltshev 2008). Global distribution: Mediterranean to Ukraine.

19. Tegenaria domestica (Clerck, 1757) Literary data: L89 (Deltshev 2008). Global distribution: Europe to China, Japan. Introduced to Australia, New Zealand, the Americas.

20. *Tegenaria ferruginea (Panzer, 1804) New data: L56a (1♀, 1 j).

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Global distribution: Europe, Azores. Introduced to Venezuela.

21. *Tetrix chyzeri de Blauwe, 1980 New data: L37d (1♀). The species was desctribed from Trebinje (Bosnia & Herzegovina). The new photos of the female correspond well with the pictures presented by De Blauwe (1980) and will contribute to better taxonomic knowledge of the species (Figs. 5–7). Global distribution: Bosnia and Herzegovina, Bulgaria, Hungary.

Figures. 5–7. Tetrix chyzeri, female. 5 – epigyne, ventral view, 6 – vulva, ventral view, 7 – vulva, dorsal view. Scale: 0.18 mm.

22. *Textrix denticulata (Olivier, 1789) New data: L56b (1♀). Global distribution: Europe, Turkey.

Amaurobiidae 23. Amaurobius annulatus (Kulczyński, 1906) Literary data: L28 [locus typicus] sub Ciniflo a. (Kulczyński 1906), L28 (Drensky 1936, Kolosvary 1938, Nikolić & Polenec 1981). New data: L30b (1♀), L37d (5♀♀). Remark: the species was described from Dalmatia (Croatia) only by female and the male is still unknown. The presented photos of the female correspond well with the single picture in the description (Kulczyński, 1906) and will contribute to better taxonomic knowledge of the species (Figs. 8–10). Global distribution: Known only from the type locality in Montenegro.

Figures 8–10. Amaurobius annulatus, female. 8 – epigyne, ventral view, 9 – vulva, dorsal view, 10 – vulva, aboral view. Scale: 0.18 mm.

24. *Amaurobius hercegovinensis Kulczyński, 1915 New data: L75b (1♀). Remark: the species was described from Bilek (Bosnia & Herzegovina). Morphologically, our material corresponds well to the description of Amaurobius hercegovinensis and the presented photos of the female contribute to the taxonomic characteristic of the species (Figs. 11–13).

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Global distribution: Bosnia-Hercegovina.

Figures 11–13. Amaurobius hercegovinensis, female. 11 – epigyne, ventral view, 12 – vulva, dorsal view, 13 – vulva, aboral view. Scale: 0.2 mm.

25. Amaurobius scopolii Thorell, 1871 Literary data: L28 (Kolosvary 1938), L89 (Nikolić & Polenec 1981). Global distribution: France, Italy, Slovenia.

Anapidae 26. Comaroma simoni Bertkau, 1889 Literary data: L55a (Thaler 1978, Kropf 1998). Global distribution: Europe.

27. Zangherella relicta (Kratohvil, 1935) Literary data: L24b (Deltshev et al. 2011), L37b [locus typicus] sub Pseudanapis r. (Kratochvíl 1935b, Nikolić & Polenec 1981), L89 (Deltshev 2008, Mammola et al. 2018). Global distribution: Bulgaria, Montenegro, Serbia.

Anyphaenidae 28. Anyphaena sabina L. Koch, 1866 Literary data: L28 (Kolosvary 1938). Global distribution: Europe, Turkey, Caucasus.

Araneidae 29. *Aculepeira ceropegia (Walckenaer, 1802) New data: L56c (1♂, 1♀). Global distribution: Europe, Turkey, Caucasus, Russia (Europe to West Siberia), Kazakhstan, Iran?

30. Agalenatea redii (Scopoli, 1763) Literary data: L 26, L28, sub Aranea r. (Kolosvary 1938). Global distribution: Europe, Turkey, Caucasus, Russia (Europe to South Siberia), Iran, Central Asia, China.

31. *Araneus angulatus Clerck, 1757 New data: L56c (1♀). Global distribution: Europe, Turkey, Russia (Europe to Far East), Iran, Central Asia, Korea.

32. Araneus circe (Audouin, 182) Literary data: L33a, sub Epeira c. (Chyzer & Kulczyński 1891, Damin 1896, 1900), L33a (Drensky 1936), L35, sub Epeira c. (Damin 1896, 1900), L35 (Drensky 1936), L70 (Brignoli 1966), L89 (Nikolić & Polenec 1981). Global distribution: Southern Europe, Egypt, Turkey, Caucasus, Iran.

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33. Araneus diadematus Clerck, 1757 Literary data: L28, sub Aranea diadema (Kolosvary 1938), L70 (Brignoli 1966). New data: L45 (1sa♂, 1sa♀), L56b (1♂), L56c (1♂, 1♀), L59b (1♀). Global distribution: Europe, Middle East, Turkey, Caucasus, Russia (Europe to Far East), Iran, Central Asia, China, Japan. Introduced to North America.

34. Araniella alpica (L. Koch, 1869) Literary data: L64c (Sacher 1984). New data: L2 (1j), L56a (1♀). Global distribution: Europe, Turkey, Caucasus, Russia (Europe to Western Siberia), Iran?.

35. Araniella cucurbitina (Clerck, 1757) Literary data: L28, sub Aranea c. (Kolosvary 1938). New data: L56b (1sa♀). Global distribution: Europe, Turkey, Russia (Europe) to Central Asia, China, Korea.

36. *Araniella opisthographa (Kulczynski, 1905) New data: L56c (2♀♀). Global distribution: Europe, Caucasus, European Russia to Central Asia, Iran.

37. Araniella proxima (Kulczyński, 1885) Literary data: L78a (Sacher 1984). Global distribution: North America, Europe, Russia (Europe to Far East), Turkey, Iran, Kazakhstan.

38. Argiope bruennichi (Scopoli, 1772) Literary data: L12, L33a (Damin 1896, 1900, Drensky 1936), L70 (Brignoli 1966). New data: L56b (1♂). Global distribution: Europe, Turkey, Israel, Russia (Europe to Far East), Iran, Central Asia to China, Korea, Japan.

39. Argiope lobata (Pallas, 1772) Literary data: L12 (Damin 1896, 1900, Drensky 1936). Global distribution: Southern Europe to Central Asia and China, northern Africa, South Africa, Israel, India, from Myanmar to New Caledonia and northern Australia.

40. Cercidia prominens (Westring, 1851) Literary data: L28 (Kolosvary 1938). Global distribution: America, Europe, Caucasus, Russia (Europe to Far East), Kazakhstan.

41. Cyclosa conica (Pallas, 1772) Literary data: L33a (Damin 1896, 1900, Drensky 1936), L89 (Nikolić & Polenec 1981). New data: L56b (1♀). Global distribution: North America, Europe, Turkey, Caucasus, Russia (Europe to Far East), Iran, Central Asia, China.

42. Cyrtarachne ixoides (Simon, 1870) Literary data: L28 (Kolosvary 1938). Global distribution: Mediterranean, Caucasus, Madagascar.

43. Cyrtophora citricola (Forsskål, 1775) Literary data: L89 (Mikhailov & Temereva 2014). Global distribution: Southern Europe, Africa, Near East, Pakistan, India, China, Japan. Introduced to Dominican Repubic, Costa Rica, Colombia, Brazil.

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44. Gibbaranea bituberculata (Walckenaer, 1802) Literary data: L33a, sub Epeira dromedaria (Damin 1896, 1900), sub Araneus dromedarius (Drensky 1936), L70, sub Araneus bituberculatus (Brignoli 1966), L89, sub Araneus bituberculatus (Nikolić & Polenec 1981). Global distribution: Europe, Turkey, Israel, Russia (Europe to Far East), Central Asia to China, Japan, India.

45. Glyptogona sextuberculata (Keyserling, 1863) Literary data: L12, sub G. 6-tuberculata (Damin 1896, 1900, Drensky 1936), L89 (Nikolić & Polenec 1981). Global distribution: Italy to Israel.

46. Mangora acalypha (Walckenaer, 1802) Literary data: L12, L33a, L35, sub Epeira a. (Damin 1896, 1900), L12, L33a, L35 (Drensky 1936), L26, L28, L33a (Kolosvary 1938), L70 (Brignoli 1966), L89 (Nikolić & Polenec 1981). New data: L21 (1♀). Global distribution: Europe, Madeira, North Africa, Turkey, Middle East, Caucasus, Russia (Europe to South Siberia), Central Asia, China.

47. Neoscona adianta (Walckenaer, 1802) Literary data: L33a, L35, sub Epeira a. (Damin 1896, 1900), L33a, sub Araneus adiantus (Giltay 1932, Drensky 1936), L35, sub Araneus adiantus (Drensky 1936), L70, sub Araneus adiantus (Brignoli 1966), L89 (Nikolić & Polenec 1981). Global distribution: Europe, North Africa to Central Asia, Russia (Europe to Far East), China, Korea, Japan.

48. Neoscona subfusca (C.L. Koch, 1837) Literary data: L3, sub Araneus dalmaticus (Giltay 1932, Drensky 1936), L33a, sub Epeira dalmatica (Damin 1896, 1900), sub Araneus dalmaticus (Drensky 1936), L70, sub Araneus subfuscus (Brignoli 1966), L89 (Nikolić & Polenec 1981). New data: L21 (1♀). Global distribution: Southern Europe, Africa, Turkey, Middle East, Ukraine, Caucasus, Russia (Europe) to Central Asia.

49. Zilla diodia (Walckenaer, 1802) Literary data: L12, sub Epeira d. (Damin 1896, 1900), L12, sub Araneus diodius (Drensky 1936), L28, sub Aranea d. (Kolosvary 1938), L33a, L35, sub Epeira d. (Damin 1896), L89 (Nikolić & Polenec 1981). Global distribution: Europe to Azerbaijan.

50. Zygiella keyserlingi (Ausserer, 1871) Literary data: L12, sub Zilla k. (Damin 1896, Drensky 1936), sub, Epeira k. (Damin1900), L89 (Nikolić & Polenec 1981). Global distribution: Southern Europe, Ukraine, Turkey.

51. Zygiella x-notata (Clerck, 1757) Literary data: L12, sub Zilla x-n. (Damin 1896, 1900), L33a, L35, sub Zilla x-n. (Damin 1900), L89, sub Z. xnotata (Nikolić & Polenec 1981). Global distribution: Europe, Turkey, Caucasus, Iran? Introduced to North America, Chile, Uruguay, Argentina, China, Japan, Réunion isl.

Atypidae 52. Atypus piceus (Sulzer, 1776) Literary data: L71 (Kraus & Baur 1974).

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Remark: very doubtful country record. In original as ―Drin-Ursprung‖= springs of Drin river, positioned deep in Albania. Global distribution: Europe (France to Russia), Iran.

Cheiracanthiidae 53. *Cheiracanthium elegans Thorell, 1875 New data: L56b (1♀). Global distribution: Europe, Turkey, Caucasus, Russia (Europe to South Siberia), Kazakhstan, Central Asia.

54. Cheiracanthium mildei L. Koch, 1864 Literary data: L28 (Kolosvary 1938). New data: L21 (1♀). Global distribution: Europe, North Africa, Turkey, Middle East, Caucasus, Russia (Europe) to Central Asia. Introduced to North America, Argentina.

*Clubionidae 55. *Clubiona caerulescens L. Koch, 1867 New data: L59a (2♀♀). Global distribution: Europe, Russia (Europe to Far East), Kazakhstan, China, Japan.

56. *Clubiona similis L. Koch, 1867 New data: L56a (1♀). Global distribution: Europe, Turkey, Caucasus.

57. *Porrhoclubiona genevensis (L. Koch, 1866) New data: L59a (1♀). Global distribution: Europe, Turkey, Caucasus, Russia (Europe to South Siberia), Iran, Central Asia.

Cybaeidae 58. Cryphoecina deelemanae Deltshev, 1997 Literary data: L13a (Deltshev 1997a). Global distribution: Montenegro.

59. *Cybaeus balkanus Deltshev, 1997 New data: L75a (2♀♀, 1 j), L75c (1♂). Global distribution: Bulgaria, Greece (Van Keer et al. 2010), Republic of North Macedonia, Serbia.

Cybaeus brignolii Maurer, 1992 Remark: doubtful and removed after Deltshev (1997b).

Dictynidae 60. Archaeodictyna consecuta (O. P.-Cambridge, 1872) Literary data: L28, sub Dictyna annulata (Drensky 1936, after Kulczynsky 1906?). Remark: In Kulczynsky (1906) no any data about this species. The record probably corresponded to Amaurobius annulatus. Global distribution: Europe, Caucasus, Russia (Europe to South Siberia).

61. Brigittea latens (Fabricius, 1775) Literary data: L28, sub Dictyna l. (Kolosvary 1938). Global distribution: Europe to Central Asia.

62. Nigma flavescens (Walckenaer, 1830) Literary data: L33a, sub Dictyna f. (Damin 1896, 1900, Drensky 1936). Global distribution: Europe, Caucasus, Iran.

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63. Nigma walckenaeri (Roewer, 1951) Literary data: L28, sub Dictyna viridissima (Kolosvary 1938). Global distribution: Europe, Turkey, Caucasus.

Dysderidae 64. Dasumia canestrinii (L. Koch, 1876) Literary data: L24a, sub D. istriaca (Simon, 1882). Global distribution: Southern Europe.

65. Dasumia chyzeri (Kulczyński, 1906) Literary data: L28, sub Harpactes c. (Kulczyński 1906, Drensky 1936, Nikolić & Polenec 1981), L28, sub Harpactocrates c. (Kolosvary 1938). New data: L72a (1♂). Global distribution: Eastern Europe.

66. Dysdera adriatica Kulczynski, 1897 Literary data: L13c (Deeleman-Reinhold & Deeleman 1988). New data: L56b (2♀♀). Global distribution: Austria, Balkans.

67. Dysdera bellimundi Deeleman-Reinhold, 1988 Literary data: L39, L67 (Deeleman-Reinhold & Deeleman 1988). Global distribution: Albania, Montenegro.

68. Dysdera dubrovninnii Deeleman-Reinhold, 1988 Literary data: L4, L13a, L19, L72b (Deeleman-Reinhold & Deeleman 1988). Global distribution: SE Europe (Balkans), Romania, Slovakia.

69. Dysdera granulata Kulczyński, 1897 Literary data: L12, L14a, L24a, L72b (Deeleman-Reinhold & Deeleman 1988). Global distribution: Albania, Balkans, Italy.

70. Dysdera kollari Doblika, 1853 Literary data: L33a [locus typicus] (Doblika 1853, Damin 1900, Nikolić & Polenec 1981, Deeleman- Reinhold & Deeleman 1988). Global distribution: Balkans, Greece, Italy, Turkey.

71. Dysdera pretneri Deeleman-Reinhold, 1988 Literary data: L13a, L19 (Deeleman-Reinhold & Deeleman 1988). Global distribution: Croatia, Greece, Montenegro.

72. Dysderocrates silvestris Deeleman-Reinhold, 1988 Literary data: L56c (Lazarov 2010), L89 (Deltshev 2008). Global distribution: Croatia, Greece, Montenegro, Serbia.

73. Dysderocrates storkani (Kratohvil, 1935) Literary data: L23a, L39, L54 (Deeleman-Reinhold & Deeleman 1988), L89 (Deltshev 2008). Global distribution: SE Europe (Balkans).

74. Folkia mrazeki (Nosek, 1904) Literary data: L9a, L14b, L18, L20b, L53b, L63b, L73 (Deeleman-Reinhold 1993), L18 [locus typicus] sub Stalita m. (Nosek 1904, Fage 1931, Nikolić & Polenec 1981), L18, sub Stalagtia m. (Kratochvíl 1970, 1978), L89, sub Stalita m. (Bresjanceva 1907, Reimoser 1929, Absolon &

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Kratochvíl 1933, Drensky 1936, Roewer 1942), L89 (Reimoser 1919, Deltshev 2008, Mammola et al. 2018). Global distribution: Montenegro.

75. Harpactea lepida (C. L. Koch, 1838) Literary data: L28, sub Harpactes lepidus (Kolosvary 1938). Global distribution: Europe.

76. *Harpactea saeva (Herman, 1879) New data: L15c (1♂). Global distribution: Hungary, Slovakia, south-eastern Europe to Ukraine.

77. Kaemis vernalis Deeleman-Reinhold, 1993 Literary data: L4 (Deeleman-Reinhold 1993). Global distribution: Bosnia-Herzegovine, Croatia, Montenegro.

78. Rhode magnifica Deeleman-Reinhold, 1978 Literary data: L13a, L19 (Deeleman-Reinhold 1993), L47 [locus typicus] (Deeleman-Reinhold 1978a, Nikolić & Polenec 1981, Pesarini 1984), L89 (Deltshev 2008, Mammola et al. 2018). New data: L55b (1♂). Global distribution: Montenegro.

79. Stalagtia hercegovinensis (Nosek, 1905) Literary data: L48a, sub S. folki (Kratochvíl 1970, 1978, Nikolić & Polenec 1981), L89 (Deeleman- Reinhold 1993, Mammola et al. 2018). Global distribution: SE Europe (Balkans), Turkey.

80. Stalagtia monospina (Absolon & Kratochvíl, 1933) Literary data: L32, sub Stalita m. & S. krivosijana (Kratochvíl 1935b, 1970, 1978, Nikolić & Polenec 1981), L37a, sub Stalita m. (Kratochvíl 1934), L37b, sub Stalita m. & S. krivosijana (Kratochvíl 1935b, 1970, 1978), L89 (Mammola et al. 2018). Global distribution: Montenegro.

81. Stalagtia skadarensis Kratochvíl, 1970 Literary data: В62 [locus typicus] (Kratochvíl 1970, 1978, Nikolić & Polenec 1981). L89 (Deeleman-Reinhold 1993, Deltshev 2008, Mammola et al. 2018). Global distribution: Montenegro.

Stalita schioedtei Thorell, 1870b Remark: nomen dubium (Kratochvíl 1970).

82. Stalitella noseki Absolon & Kratochvíl, 1933 Literary data: L25a (Absolon & Kratochvíl 1933, Nikolić & Polenec 1981), L25b (Deeleman- Reinhold 1971), L30e (Kratochvíl 1935b), L89 (Drensky 1936, Roewer 1942 Mammola et al. 2018). Global distribution: Bosnia-Hercegovina, Montenegro.

Eresidae 83. Eresus kollari Rossi, 1846 Literary data: L89 (Rezac et al. 2008). Global distribution: Europe, Caucasus, Russia (Europe to Far East), Turkey, Iran, Central Asia.

Filistatidae 84. Filistata insidiatrix (Forsskål, 1775) Literary data: L89 (Drensky 1936). Remark: only on a map (p. 89: map 1) for Kotor valley, without explanation.

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Global distribution: Mediterranean to Turkmenistan, Cape Verde isl.

85. Pritha nana (Simon, 1868) Literary data: L28, sub Filistata n. (Kolosvary 1938). Global distribution: Mediterranean, India.

Gnaphosidae 86. Aphantaulax trifasciata (O. P.-Cambridge, 1872 Literary data: 33a, sub A.semi-niger (Damin 1896), 33a, sub A.seminigra (Damin 1900, Drensky 1936). Global distribution: Southern Europe, North Africa, Turkey, Israel, Caucasus, Russia (Europe) to Central Asia, China, Japan.

87. *Drassodes cupreus (Blackwall, 1834) New data: L5a (1♂), L6b (1♂), L56b (1♂, 1♀), L56c (2♀♀, 1j). Global distribution: Europe, Caucasus, Kazakhstan, Russia (Europe to Far East).

88. Drassodes lutescens (C.L. Koch, 1839) Literary data: L33a, sub Drassus l. (Damin 1896, 1900), L89 (Nikolić & Polenec 1981). Global distribution: Mediterranean, Ukraine, Caucasus, Russia (Europe) to Central Asia, Pakistan.

89. Drassodes montenegrinus (Kulczyński, 1897) Literary data: L89 [locus typicus] sub Drassus m. (Kulczyński in Chyzer & Kulczyński 1897, Nosek 1904, Bresjanceva 1907, Reimoser 1919), L89 (Drensky 1936). Global distribution: Croatia, Serbia.

90. *Drassodes pubescens (Thorell, 1856) New data: L59a (1♀). Global distribution: Europe, Turkey, Israel, Caucasus, Russia (Europe to Far East), Iran, Central Asia, China, Japan.

91. *Echemus angustifrons (Westring, 1861) New data: L56b (1♂). Global distribution: Europe to Central Asia.

92. Gnaphosa lucifuga (Walckenaer, 1802) Literary data: L 33a, sub Drassus lucifugus (Carrara 1846?), L 33a (Damin 1896, 1900, Drensky 1936), L89 (Nikolić & Polenec 1981). Remark: The record from Carrara 1846 is sensu Damin 1900. We cannot correctly interpret this record in the publication. Global distribution: Europe, Turkey, Caucasus, Iran, Russia (Europe to South Siberia), Kazakhstan, China.

93. Haplodrassus minor (O. P.-Cambridge, 1879) Literary data: L28, sub Drassodes m. (Kolosvary 1938). Global distribution: Europe, Turkey.

94. *Haplodrassus signifer (C. L. Koch, 1839) New data: L6b (1♂), L8 (2♂♂). Global distribution: North America, Europe, North Africa, Turkey, Israel, Caucasus, Russia (Europe to Far East), Central Asia, China, Korea.

95. *Micaria coarctata (Lucas, 1846) New data: L56b (1♂).

NAUMOVA ET AL.

Global distribution: Mediterranean, Eastern Europe, Caucasus, Russia (Europe to Far East), Kazakhstan, Central Asia.

96. *Micaria fulgens (Walckenaer, 1802) New data: L56c (1♀). Global distribution: Europe, Caucasus, Russia (Europe to South Siberia), Central Asia, China.

97. Micaria rossica Thorell, 1875 Literary data: L56c (1♂). Global distribution: Europe, Caucasus, Russia (Europe to Far East), Turkey, Central Asia, Mongolia, China, North America.

98. Micaria sociabilis Kulxzynski, 1897 Literary data: L89 (Nikolić & Polenec 1981). Global distribution: Europe, Azerbaijan.

99. Nomisia montenegrina Giltay, 1932 Literary data: L14a [locus typicus] (Giltay 1932, Drensky 1936, Bosmans 2013), L89 (Nikolić & Polenec 1981). Global distribution: Montenegro.

100. Pterotricha lentiginosa (C.L. Koch, 1837) Literary data: L33a (Roewer 1928, Drensky 1936), L89 (Nikolić & Polenec 1981). Global distribution: Balkans, Greece, Turkey, Ukraine.

101. *Zelotes balcanicus Deltshev, 2006 New data: L6b (1♂, 1♀), L56b (1♀). Global distribution: Bulgaria, Greece, Italy, Republic of North Macedonia, Romania, Israel.

102. Zelotes oblongus (C. L. Koch, 1833) Literary data: L28 (Kolosvary 1938). New data: L6a (1♀), L6b (3♂♂). L56c (1♂). Global distribution: Europe, Turkey.

103. *Zelotes similis (Kulczyński, 1887) New data: L1 (1♀), L56c (1♀). Global distribution: Italy, Central Europe to Turkey.

104. *Zelotes talpinus (L. Koch, 1872) New data: L56c (2♂♂), L59a (1♂). Global distribution: Western to Central Europe, Italy.

Leptonetidae 105. Barusia hofferi (Kratochvíl, 1935) Literary data: L30c [locus typicus] sub Paraleptoneta h. (Kratochvíl 1935b, Kratochvíl & Miller, 1939, ), L30c, sub Barušia (Barušia) h. (Kratochvíl 1978), L30b (Nikolić & Polenec 1981), L89 (Deltshev 2008, Mammola et al. 2018). Global distribution: Montenegro.

106. Sulcia armata Kratochvíl, 1978 Literary data: L15a [locus typicus] (Kratochvíl 1978), L89 (Deltshev 2008, Mammola et al. 2018). Global distribution: Montenegro.

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107. Sulcia mirabilis Kratochvíl, 1938 Literary data: L15a [locus typicus], L16, L37f, L77b, L82b, sub Paraleptoneta orientalis (Kratochvíl 1935b), L15a, sub Paraleptoneta orientalis (Kratochvíl 1938b), L15a (Kratochvíl 1978, Nikolić & Polenec 1981), L16, L37f, L77b, L82b (Kratochvíl 1938b), L37f (Naumova et al. 2016), L89 (Deltshev 2008, Mammola et al. 2018). New data: L56d (1♂, 4♀♀, 4j). Global distribution: Montenegro.

108. Sulcia montenegrina (Kratochvíl & Miller, 1939) Literary data: L9b [locus typicus] sub Paraleptoneta m. (Kratochvíl & Miller 1939), L9a (Kratochvíl 1978, Nikolić & Polenec 1981), L89, sub Paraleptoneta m. (Roewer 1942, Brignoli 1967), L89 (Deltshev 2008, Mammola et al. 2018). Global distribution: Montenegro.

109. *Sulcia occulta Kratochvíl, 1938 New data: L56d (1♂). Global distribution: Bosnia-Hercegovina, Serbia.

Linyphiidae 110. *Agyneta rurestris (C. L. Koch, 1836) New data: L21 (1♂, 1♀). Global distribution: Europe, Turkey, Caucasus, Russia (Europe to South Siberia), Central Asia, China, Korea.

111. Asthenargus bracianus Miller, 1938 Literary data: L15a (Naumova et al. 2016). New data: L6b (1♂). Global distribution: Central, Eastern Europe.

112. Bolyphantes kolosvaryi (Caporiacco, 1936) Literary data: L38 (Helsdingen van et al. 2001). Global distribution: France, Switzerland, Italy, Balkans.

113. Centromerus acutidentatus Deltshev, 2002 Literary data: L89 (Mammola et al. 2018). Global distribution: Balkans.

114. Centromerus cavernarum (L. Koch, 1872) Literary data: L89 (Deltshev 2008). New data: L59a (1♀). Global distribution: Europe.

115. Centromerus europaeus (Simon, 1911) Literary data: L10a (Naumova et al. 2016). Global distribution: Portugal, Spain, France, Algeria, Balkans.

116. Centromerus obenbergeri Kratochvíl & Miller, 1938 Literary data: L18 (Deeleman-Reinhold 1974, Deltshev & Ćurĉić 2002), L89 (Kratochvíl & Miller 1938, Deltshev 2008, Mammola et al. 2018). New data: L37d (1♂). Global distribution: Montenegro, Republic of North Macedonia (Ćurĉić et al. 2000a), Serbia (Ćurĉić et al. 2007).

117. Centromerus silvicola (Kulczynski, 1887) Literary data: L89 (Kulczyński 1914, Nikolić & Polenec 1981).

NAUMOVA ET AL.

Global distribution: Central Europe to Balkans and Ukraine.

118. Centromerus subcaecus Kulczyński, 1914 Literary data: L36, L37f, L53c (Kratochvíl 1935b, Kratochvíl & Miller 1938), L37c (Kratochvíl 1934, Kratochvíl & Miller 1938), L89 (Kratochvíl, 1936, Nikolić & Polenec 1981, Deltshev & Ćurĉić 1997, Deltshev 2008, Mammola et al. 2018). Global distribution: Europe.

119. *Ceratinella scabrosa (O. P. -Cambridge, 1871) New data: L56b (1♀). Global distribution: Europe, Caucasus, Russia (Europe to South Siberia).

120. Cresmatoneta mutinensis (Canestrini, 1868) Literary data: L28, sub Formicina m. (Kolosvary 1938), L33a, sub Formicina m. (Damin 1896, 1900), L89 (Nikolić & Polenec 1981). Global distribution: Southern Europe, Russia (Europe), Turkey, Caucasus.

121. Diplocephalus cristatus (Blackwall, 1833) Literary data: L89 (Bosmans & Oger 2018). New data: L56b (3♀♀). Global distribution: Europe, Russia (Europe to Far East), Kazakhstan. Introduced to North America, Falkland Is., New Zealand.

122. Erigone atra Blackwall, 1833 Literary data: L28 (Kolosvary 1938). Global distribution: Europe, Caucasus, Russia (Europe to Far East), Kazakhstan, Central Asia, China, Mongolia, Korea, Japan, North America.

123. Erigone dentipalpis (Wider, 1834) New data: L59b (1♂). Global distribution: Europe, Turkey, Caucasus, Russia (Europe to Far East), Kazakhstan, Central Asia, China.

124. Fageiella ensigera Deeleman-Reinhold, 1974 Literary data: L68 (Deeleman-Reinhold 1974, Deltshev 1988), L89 (Deeleman-Reinhold 1974, Deltshev 2008), L94 (Mammola et al. 2018). Global distribution: Kosovo, Montenegro, Serbia.

125. Frontinellina frutetorum (C. L. Koch, 1834) Literary data: 33a, sub Linyphia f. (Damin 1896, 1900), L89 (Nikolić & Polenec 1981). New data: L56b (1♀). Global distribution: Europe, Caucasus, Russia (Europe to South Siberia), North Africa, Turkey, Kazakhstan, Central Asia.

126. *Gonatium orientate Fage, 1931 New data: L56c (1♂, 1♀), L59a (2♀♀). Global distribution: Bulgaria, Republic of North Macedonia (Komnenov 2017), Romania, Serbia (Ćurĉić et al. 2000b), Ukraine (Hirna et. al. 2016).

127. Hypomma bituberculatum (Wider, 1834) Literary data: L28 (Kolosvary 1938, Nikolić & Polenec 1981). Global distribution: Europe, Turkey, Russia (Europe to Far East), Kazakhstan, Kyrgyzstan, China.

128. Lepthyphantes centromeroides Kulczyn'ski, 1914 Literary data: L29, L57 (Deeleman-Reinhold 1974).

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New data: L69a (1♀). Global distribution: Balkans, Romania.

129. *Lepthyphantes leprosus (Ohiert, 1865) New data: L37d (1♀), L74b (1♂), L74a (1♀). Global distribution: North America, Europe, Turkey, Caucasus, Russia (Europe to Far East), Kazakhstan. Introduced to Chile.

130. *Lepthyphantes notabilis Kulczyński, 1887 New data: L59a (1♀). Global distribution: Europe.

Lepthyphantes tenoides Kolosvary, 1938 Remark: nomen dubium (van Helsdingen et al. 1977: 42, WSC 2019)

131. Linyphia triangularis (Clerck, 1757) Literary data: L35 (Damin 1896, 1900), L89 (Nikolić & Polenec 1981). New data: L45 (1♀). Global distribution: Europe, Turkey, Caucasus, Russia (Europe to Far East), Iran, Kazakhstan, China. Introduced to USA.

132. *Maso sundevalli (Westring, 1851) New data: L59a (1♂, 4♀♀). Global distribution: North America, Europe, Turkey, Caucasus, Russia (Europe to Far East), China, Mongolia, Japan.

133. *Microneta viaria (Blackwall, 1841) New data: L59a (4♀♀). Global distribution: North America, Europe, Turkey, North Africa, Caucasus, Russia (European to Far East), China, Mongolia, Korea, Japan.

134. *Mughiphantes pulcher (Kulczyński, 1881) New data: L59b (1♂, 2♀♀). Global distribution: Central Europe.

135. *Neriene peltata (Wider, 1834) New data: L56b (1♀), L56c (1♀). Global distribution: Europe, Greenland, Caucasus, Russia (Europe to South Siberia).

136. *Neriene radiata (Walckenaer, 1841) New data: L45 (1♀). Global distribution: Europe, Turkey, Caucasus, Russia (Europe to Far East), Kazakhstan, China, Korea, Japan, North America.

137. *Obscuriphantes obscurus (Blackwall, 1841) New data: L59a (1♀). Global distribution: Europe, Turkey, Caucasus, Russia (Europe to West Siberia), China.

138. *Oedothorax agrestis (Blackwall, 1853) New data: L56a (3♂♂, 2♀♀). Global distribution: Europe.

NAUMOVA ET AL.

139. Palliduphantes spelaeorum (Kulczyński, 1914) Literary data: L48a, sub Lepthyphantes s. (Deeleman-Reinhold 1974), L48b, sub Lepthyphantes s. (Kratochvíl 1935b, Miller 1978), L89, sub Lepthyphantes s. (Nikolić & Polenec 1981), L89 (Deltshev 2008). Global distribution: SE Europe (Balkans).

140. Palliduphantes trnovensis (Drensky, 1931) Literary data: L89 (Mammola et al. 2018). Global distribution: Bulgaria, Montenegro, Republic of North Macedonia, Serbia.

141. *Pityohyphantes phrygianus (C. L. Koch, 1836) New data: L56c (1♀). Global distribution: Europe, Russia to Kazakhstan, Japan.

142. Porrhomma convexum (Westring, 1851) Literary data: L11, L48d, L85, sub P. pygmaeum c. (Deeleman-Reinhold 1974). New data: L74b (1♀). Global distribution: Europe, Caucasus, Russia (Europe, West Siberia), Kazakhstan, Canada, Greenland.

143. *Scotargus pilosus Simon, 1913 New data: L75b (1♂). Global distribution: Europe, North Africa, Caucasus, Russia to Central Asia.

144. *Sintula retroversus (O. P.-Cambridge, 1875) New data: L61 (1♂). Global distribution: Europe to Azerbaijan.

145. Sintula roeweri Kratochvíl, 1935 Literary data: L32 [locus typicus], L34c [locus typicus], L53c [locus typicus], L86 [locus typicus], L90 [locus typicus], L91 [locus typicus] (Kratochvíl 1935b), L78a (Nikolić & Polenec 1981), L89 (Mammola et al. 2018). Global distribution: Montenegro.

146. *Tenuiphantes flavipes (Blackwall, 1854) New data: L15c (1♂), L56c (1♀). Global distribution: Europe.

147. *Tenuiphantes jacksonoides (van Helsdingen, 1977) New data: L6b (1♂, 2jj). Global distribution: Austria, Germany, Switzerland; Bulgaria (Blagoev et al. 2018), Italy (Pantini & Isaia 2018).

148. *Tenuiphantes tenebricola (Wider, 1834) New data: L56c (1♀). Global distribution: Europe, Turkey, Russia, China.

149. *Tenuiphantes tenuis (Blackwall, 1852) New data: L20c (2♀♀), L37d (1♀). Global distribution: Europe, Macaronesia, Northern Africa, Turkey, Caucasus, Central Asia. Introduced to USA, Chile, Argentina, New Zealand.

150. *Tiso vagans (Blackwall, 1834) New data: L56b (1♀). Global distribution: Europe, Madeira.

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151. *Trichoncus saxicola (O. P. -Cambridge, 1861) New data: L56b (1♀). Global distribution: Europe.

152. Troglohyphantes boudewijni Deeleman-Reinhold, 1974 Literary data: L9a (Deeleman-Reinhold 1978b), L10b, L62 [locus typicus], L63b, L64b (Deeleman- Reinhold 1974, 1978b), L89 (Nikolić & Polenec 1981, Deltshev 2008, Mammola et al. 2018). Global distribution: Montenegro.

153. Troglohyphantes lesserti Kratochvíl, 1935 Literary data: L40, L41, L42b, L43, L46b, L47, L48a, L50a, L50b, L53a, L53b, L64a (Deeleman- Reinhold 1974, 1978b), L48a (Kratochvíl 1935b, Nikolić & Polenec 1981), L77b (Kratochvíl 1935b), L78a (Deltshev 2008, Mammola et al. 2018). Global distribution: SE Europe (Balkans).

154. Troglohyphantes pretneri Deeleman-Reinhold, 1978 Literary data: L23b (Deeleman-Reinhold 1978b, Nikolić & Polenec 1981), L89 (Deltshev 2008, Mammola et al. 2018). New data: L22 (2♂♂, 4♀♀). Global distribution: Albania, Montenegro.

155. Troglohyphantes troglodytes (Kulczyński, 1914) Literary data: L7, L15a, L16, L17, L30c, L30e, L32, L34c, L34e, L36, L37b, L37f, L48c, L53c, L77b, L79, L81, L82a, L82b, L86, L87, L90, L93 (Kratochvíl 1935b), L14b, L40, L41, L42a, L42b, L44a, L46b, L47, L48a, L48b, L50a, L69b (Deeleman-Reinhold 1974), L15a, L44a, L51 (Naumova et al. 2016), L25b, L30e, L83 (Deeleman-Reinhold 1978), L34a, L34d, L34f (Deeleman-Reinhold 1974, 1978b), L89 (Nikolić & Polenec 1981, Deltshev 2008, Mammola et al. 2018). New data: L10a (1♀). Global distribution: Bosnia and Herzegovina, Croatia, Montenegro (Mammola et al. 2018).

156. *Walckenaeria corniculans (O. P.-Cambridge, 1875) New data: L37d (1♀). Global distribution: Europe, North Africa.

157. *Walckenaeria nudipalpis (Westring, 1851) New data: L74b (1♀). Global distribution: Europe, Caucasus, Japan.

Liocranidae 158. Sagana rutilans Thorell, 1875 Literary data: L17, sub Liocranum r. (Kratochvíl 1935b), L89, sub Liocranum r. (Nikolić & Polenec 1981, Deltshev 2008). New data: L37d (1♀). Global distribution: Europe, Caucasus (Russia, Georgia).

Lycosidae 159. Alopecosa albofasciata (Brulle, 1832) Literary data: L33a, sub Lycosa ocellaris (Doleschall 1852), L33a, sub Tarentula ocellaris (Canestrini & Pavesi 1868), L33a, sub Lycosa a. (Gasperini 1891), L33a, sub Tarentula a. (Damin 1896, 1900, Drensky 1936), L89, sub Xerolycosa a. (Nikolić & Polenec 1981). New data: L5a (1♀), L6b (20♂♂), L8 (1♀), L44b (3jj). Global distribution: Mediterranean to Central Asia.

NAUMOVA ET AL.

160. *Alopecosa farinosa (Herman, 1879) New data: L56a (1♀). Global distribution: Europe, Turkey, Caucasus, Russia (Europe to Far East), Kazakhstan.

161. *Alopecosa pulverulenta (Clerck, 1758) New data: L56b (2♀♀). Global distribution: Europe, Turkey, Caucasus, Russia (Europe to Far East), Kazakhstan, China, Korea, Japan.

162. Alopecosa striatipes (C. L. Koch, 1839) Literary data: L28, sub Tarentula s. (Kolosvary 1938). Global distribution: Europe, Turkey, Caucasus.

163. Arctosa variana C. L. Koch, 1847 Literary data: L28, sub Tarentula v. (Kolosvary 1938). Global distribution: Mediterranean to Central Asia.

164. Aulonia albimana (Walckenaer, 1805) Literary data: L28 (Kolosvary 1938). New data: L44b (1♀), L56b (1sa♀). Global distribution: Europe, Turkey, Caucasus.

165. *Pardosa agrestis (Westring, 1861) New data: L56c (1♀), L59b (1♀). Global distribution: Europe, Caucasus, Russia (Europe to Far East), Central Asia, China.

166. *Pardosa albatula (Roewer, 1951) New data: L56c (4♂♂, 18♀♀), L59a (1♂), L74b (2♂♂, 1♀), L75b (6♀♀), L76 (1♀). Global distribution: Italy to Ukraine and Greece, Turkey, Caucasus (Russia, Georgia).

167. Pardosa amentata (Clerck, 1757) Literary data: L28, sub Lycosa saccata (Kolosvary 1938) Global distribution: Europe, Turkey, Caucasus, Russia (Europe to South Siberia).

168. *Pardosa cavannae Simon, 1881 New data: L74a (1♂). Global distribution: Italy, Albania.

169. Pardosa hortensis (Thorell, 1872) Literary data: L28, sub Lycosa annulata (Kolosvary 1938). New data: L56a (2♀♀), L56b (1♀). Global distribution: Europe, Turkey, Caucasus, Russia (Europe to South Siberia), Japan.

170. *Pardosa lugubris (Walckenaer, 1802) New data: L45 (1♀), L56a (2♀♀), L56b (2♀♀), L56c (2♀♀). Global distribution: Europe, Turkey, Caucasus, Russia (Europe to Far East), Kazakhstan.

171. Pardosa mixta (Kulczyński, 1887) Literary data: L89, sub Lycosa m. (Kratochvíl 1935a). New data: L58 (3♂♂), L60 (1♀). Global distribution: Europe, Turkey.

172. *Pardosa morosa (L. Koch, 1870) New data: L56a (1♀). Global distribution: Europe to Central Asia, Iraq, Iran.

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173. Pardosa proxima (C. L. Koch, 1847) Literary data: L28, sub Lycosa p. (Kolosvary 1938). Global distribution: Southern Europe, Macaronesia, Northern Africa, Russia (Europe to Far East), Central Asia, China.

174. Pardosa tatarica (Thorell, 1875) Literary data: L13b, sub P. strigillata (Tongiorgi 1966). Global distribution: Southern Europe, Turkey, Caucasus.

175. *Pirata tenuitarsis Simon, 1876 New data: L75c (1♀, 1sa♂). Global distribution: Europe to Mongolia.

176. *Piratula latitans (Blackwall, 1841) New data: L56b (1♂). Global distribution: Europe to Azerbaijan.

177. *Trochosa ruricola (De Geer, 1778) New data: L44b (1♀). Global distribution: Europe, Caucasus, Russia (Europe to Far East), Turkey, China, Japan, Korea. Introduced to North America, Cuba, Puerto Rico, Bermuda.

178. *Trochosa spinipalpis (F. O. P.-Cambridge, 1895) New data: L56c (1♀). Global distribution: Europe, Caucasus, Russia (Europe to South Siberia), China, Japan.

179. Trochosa terricola Thorell, 1856 Literary data: L28, sub Tarentula t. (Kolosvary 1938), L66, sub Lycosa t. pallida (Nosek 1904), L89, sub Lycosa (T.) t. var. pallida (Bresjanceva 1907). Global distribution: Europe, Turkey, Caucasus, Russia (Europe to Far East), Central Asia, China, Japan, North America.

180. *Xerolycosa nemoralis (Westring, 1861) New data: L60 (1♀). Global distribution: Europe, Caucasus, Russia (Europe to Far East), Turkey, Kazakhstan, Central Asia, China, Korea, Japan.

Mimetidae 181. Ero tuberculata (De Geer, 1778) Literary data: L28 (Kolosvary 1938). Global distribution: Europe, Russia (Europe to Central Asia), Central Asia, China.

182. Mimetus laevigatus (Keyserling, 1863) Literary data: L33a (Damin 1896, Drensky 1936, Nikolić & Polenec 1981), L33a, sub M. laevigata (Damin 1900). Global distribution: Mediterranean to Central Asia.

Miturgidae 183. Zora nemoralis (Blackwall, 1861) Literary data: L28 (Kolosvary 1938). Global distribution: Europe, Caucasus, Russia (Europe to South Siberia, Kamchatka), Turkmenistan, China, Korea, Japan.

NAUMOVA ET AL.

184. *Zora spinimana (Sundevall, 1833) New data: L56b (1♀). Global distribution: Europe, Caucasus, Russia (Europe to Far East), Turkey, Central Asia, China, Japan.

Nemesiidae 185. Nemesia pannonica Herman, 1879 Literary data: L26, L28, L33a, sub N. radiata (Kolosvary 1938), L28, sub N. p. pannonica (Nikolić & Polenec 1981). Global distribution: Eastern Europe, Italy (Pantini & Isaia 2018).

Nesticidae Domitius speluncarum (Pavesi, 1873) Remark: reported from Deltshev (2008), rejected from Pavlek & Ribera (2017) as probable Kryptonesticus eremita (Simon, 1880) for which there is no data from Montenegro.

186. Kryptonesticus arenstorffi (Kulczyński, 1914) Literary data: L9a, L15a, L15b, L37f (Naumova et al. 2016), L14b, L37bl L44a (Pavlek & Ribera 2017), L15a, L16, L17, L30c, L30e, L32, L34c, L37b, L37f, L77b, L79, L80, L81, L82a, L86, L90, L92, sub Nesticus a. (Kratochvíl 1935b), L37c, sub Nesticus a. (Kratochvíl 1933, 1934), L50a, sub Nesticus a. (Deeleman-Reinhold 1974), L89, sub Nesticus a. (Deltshev 2008), L89 (Mammola et al. 2018). Global distribution: Croatia, Montenegro.

Kryptonesticus fagei (Kratochvíl, 1933) Remark: reported from Nikolić & Polenec (1981) and Deltshev (2008), rejected from Pavlek & Ribera (2017).

187. Nesticus cellulanus (Clerck, 1757) Literary data: L33a (Kratochvíl 1933, 1934), L33b (Drensky 1936), L89 (Deltshev 2008). Global distribution: Europe, Turkey. Introduced to North America.

188. Typhlonesticus absoloni (Kratochvíl, 1933) Literary data: L9a (Naumova et al. 2016), L14b, L20b, sub Nesticus a. (Deeleman-Reinhold 1974), L14b (Deltshev et al. 2014, Pavlek & Ribera 2017), L78a, sub Nesticus a. (Drensky 1936m Nikolić & Polenec 1981), L78a, sub Nesticus vejdovskyi (Nikolić & Polenec 1981), L78b, sub T. speluncarum (Kulczyński 1914), L78b, sub Nesticus (T.) speluncarum (Fage 1931), L84, sub Nesticus vejdovskyi (Kratochvíl, 1939), L89, sub Nesticus a. (Kratochvíl 1933, 1934), L89, sub T. speluncarum (Drensky 1936), L89, sub Nesticus vejdovskyi (Kratochvíl 1938b), L89 (Deltshev 2008, Mammola et al. 2018). Global distribution: Montenegro.

Oecobiidae 189. Uroctea durandi (Latreille, 1809) Literary data: L26, L28 (Kolosvary 1938). New data: L44b (1♀). Global distribution: Mediterranean.

Oonopidae 190. Tapinesthis inermis (Simon, 1882) Literary data: L89 (Seppälä et al. 2018). Remark: The record seems mistakable. Global distribution: Europe.

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Oxyopidae 191. Oxyopes lineatus Latreille, 1806 Literary data: L35 (Damin 1896, 1900, Drensky 1936). New data: L56b (1♂). Global distribution: Europe, Caucasus, Russia (Europe to Central Asia), Turkey, Middle East, Central Asia.

Philodromidae 192. Philodromus dispar Walckenaer, 1826 Literary data: L35 (Damin 1896, 1900, Drensky 1936). Global distribution: Europe, Caucasus, Russia (Europe to South Siberia). Introduced to USA, Canada.

193. Philodromus poecilus (Thorell, 1872) Literary data: L28 (Kolosvary 1938), L89 (Nikolić & Polenec 1981). Global distribution: Europe, Caucasus, Russia (Europe to Far East), Turkey, Central Asia, Korea, Japan.

194. Tibellus oblongus (Walckenaer, 1802) Literary data: L28 (Kolosvary 1938), L89 (Nikolić & Polenec 1981). Global distribution: Europe, North Africa, Turkey, Israel, Caucasus, Russia (Europe to Far East), Central Asia, China, Korea, Japan, North America.

Pholcidae 195. Holocnemus pluchei (Scopoli, 1763) Literary data: L33a, sub H. rivulatus (Damin 1896, 1900), L33a, sub H. pluchii (Drensky 1936), L89 (Nikolić & Polenec 1981, Deltshev 2008). New data: L56d (1♂). Global distribution: Europe, Northern Africa.

196. Pholcus opilionoides (Schrank, 1781) Literary data: L89 (Deltshev 2008). Global distribution: Europe, Caucasus, Egypt, Iran (?).

197. Pholcus phalangioides (Fuesslin, 1775) Literary data: L89 (Deltshev 2008). Global distribution: Western Asia. Introduced to both Americas, Europe, Africa, Asia, Australia, New Zealand and numerous islands.

198. Spermophora senoculata Duges, 1836 Literary data: L33a (Chyzer & Kulczyński 1891, Damin 1900, Drensky 1936), L89 (Nikolić & Polenec 1981, Deltshev 2008). Global distribution: Middle East. Introduced to USA, southern Europe, China, Korea, Japan.

199. Stygopholcus absoloni (Kulczyński, 1914) Literary data: L7, L15a, L17, L30c, L30e, L32, l34b, L34e, L36, L37b, L37e, L48a, L53c, L79, L81, L82b, L90, L92, L93, sub Holocnemus (S.) a. (Kratochvíl 1935b), L37c, sub Holocnemus (S.) a. (Kratochvíl, 1934). Global distribution: Bosnia and Herzegovina, Croatia.

200. Stygopholcus skotophilus (Kratochvíl, 1940) Literary data: L89 (Mammola et al. 2018). Global distribution: Bosnia and Herzegovina, Montenegro.

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201. Stygopholcus skotophilus montenegrinus (Kratochvíl, 1940) Literary data: L7, L9a, L15a, L17, L30c, L30d, L30e, L32, L34b, L34c, L34e, l36, L37b, L37e, L48a, L48b, L53c, L77b, L79, L81, L82a, L82b, L90, L92, L93, sub S. s. (Kratochvíl 1940), L9a, L10a, L14b, L15a, L15b, L30d, l37f, L44a, L51 (Naumova et al. 2016), L34e, sub S. m. & sub S.s. (Kratochvíl 1978), L52, sub S. m. (Kratochvíl 1940, 1978, Nikolić & Polenec 1981), L52, sub S. s. (Kratochvíl 1978, Nikolić & Polenec 1981), L89, sub S. m. (Roewer, 1942), L89 (Deltshev 2008). New data: L30b (1sa♂). Global distribution: Montenegro.

Phrurulitidae 202. Phrurolithus festivus (C. L. Koch, 1835) Literary data: L28 (Kolosvary 1938). Global distribution: Europe, Turkey, Caucasus, Russia (Europe to Far East), Kazakhstan, China, Korea, Japan.

Pisauridae Dolomedes fimbriatus (Clerck, 1757) Remark: removed from the country list after Naumova (2018).

203. Pisaura mirabilis (Clerck, 1757) Literary data: L 28, sub P. listeri (Kolosvary 1938), L33a (Damin 1896, 1900, Drensky 1936), L35 (Damin 1896, 1900), L89 (Nikolić & Polenec 1981). New data: L8 (1♀), L59a (1♂). Global distribution: Europe, Turkey, Middle East, Caucasus, Russia (Europe to Middle Siberia), Central Asia, China.

Salticidae Attulus penicillatus adriaticus (Kolosvary 1938) Remark: sub Sitticus p. adriaticus (Kolosvary 1938), nomen dubium after Breitling et al. (2016).

204. Ballus chalybeius (Walckenaer, 1802) Literary data: L33a, sub B. depressus (Damin 1896, 1900, Drensky 1936), L89, sub B. depressus (Simon 1878, Nosek 1904, Bresjanceva 1907). Global distribution: Europe, North Africa to Central Asia.

205. *Ballus rufipes (Simon, 1868) New data: L56b (1♀). Global distribution: Europe, Turkey, Cyprus, North Africa.

206. * Calositticus zimmermanni (Simon, 1877) New data: L56c (1♂). Global distribution: Europe to Central Asia.

207. Carrhotus xanthogramma (Latreille, 1819) Literary data: L12, sub Philaeus bicolor (Damin 1896, 1900), L12, sub C. bicolor (Drensky 1936), L89, sub Philaeus bicolor (Simon 1878, Nosek 1904), L89 (Nikolić & Polenec 1981). Global distribution: Europe, Caucasus, Russia, Turkey, China, Korea, Japan.

208. *Euophrys frontalis (Walckenaer, 1802) New data: L59a (1♀). Global distribution: Europe, Caucasus, Russia (Europe to Far East), Turkey, Central Asia, India, China, Korea, Japan.

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209. Euophrys rufibarbis (Simon, 1868) Literary data: L12 (Damin 1896, 1900), L89 (Nikolić & Polenec 1981). New data: L56b (1♀). Global distribution: Southern Europe, North Africa, Turkey, China.

210. *Evarcha arcuata (Clerck, 1757) New data: L56b (1♂). Global distribution: Europe, Turkey, Caucasus, Russia to Central Asia, China, Japan.

211. Evarcha falcata (Clerck, 1757) Literary data: L89, sub Hasarius falcatus (Simon 1878), L89, sub Ergane f. (Nosek 1904, Drensky 1936), L89, sub Ergane faleata [sic] (Bresjanceva 1907), L89, sub E. flammata (Nikolić & Polenec 1981). Global distribution: Europe, Caucasus, Turkey, Afghanistan, Russia to Kazakhstan, China.

212. Evarcha jucunda (Lucas, 1846) Literary data: L12, L33a, L35, sub Ergane j. (Damin 1896, 1900), L12, L33a, L35 (Drensky 1936), L20a (Naumova et al. 2016), L28 (Kolosvary 1938), L89, sub Hasarius jucundus (Simon 1878), L89, sub Ergane j. (Nosek 1904). Global distribution: Mediterranean, Canary Is. Introduced to Belgium, Germany.

213. Evarcha laetabunda (C.L. Koch, 1846) Literary data: L33a, sub Ergane l. (Damin 1896, 1900), L33a (Drensky 1936), L89 (Nikolić & Polenec 1981). Global distribution: Europe, Caucasus, Russia to Central Asia, Turkey, China.

214. Heliophanus auratus C. L. Koch, 1835 Literary data: L89 (Wesołowska 1986). New data: L56b (2♀♀). Global distribution: Europe, Caucasus, Russia to Central Asia, Turkey, China.

215. *Heliophanus cupreus (Walckenaer, 1802) New data: L56b (1♀). Global distribution: Europe, Caucasus, Russia, North Africa, Turkey, China.

216. Heliophanus equester L. Koch, 1867 Literary data: L35 (Damin 1896, Drensky 1936). Global distribution: Italy to Azerbaijan, Iran.

217. Heliophanus flavipes (Hahn, 1832) Literary data: L28 (Kolosvary 1938), L35 (Damin 1896, 1900, Drensky 1936). New data: L56b (1♀). Global distribution: Europe, Caucasus, Russia to Central Asia, Turkey, China.

218. Heliophanus kochii Simon, 1868 Literary data: L12, L33a (Damin 1896, 1900, Drensky 1936), L89 (Nikolić & Polenec 1981). Global distribution: Europe, Macaronesia, North Africa, Caucasus, Turkey, near East, Kazakhstan. Introduced to Canada, USA.

219. Heliophanus lineiventris Simon, 1868 Literary data: L33a , L35 (Damin 1896, 1900, Drensky 1936), L89 (Nikolić & Polenec 1981). New data: L56c (2♀♀), L59a (1♂). Global distribution: Europe, Caucasus, Russia (Europe to Far East), Turkey, China, Mongolia, Korea, Japan.

NAUMOVA ET AL.

220. Heliophanus melinus L. Koch, 1867 Literary data: L33a (Damin 1896, 1900, Drensky 1936). Global distribution: Europe, Turkey.

221. Heliophanus simplex Simon, 1868 Literary data: L28 (Kolosvary 1938). Global distribution: Europe.

222. Heliophanus tribulosus Simon, 1868 Literary data: L35 (Damin 1900), L89 (Nikolić & Polenec 1981). Global distribution: Europe to Kazakhstan.

223. Macaroeris nidicolens (Walckenaer, 1802) Literary data: L89, sub Dendryphantes n. (Simon 1878, Nosek 1904, Bresjanceva 1907). Global distribution: Europe, North Africa to Turkey, Macaronesia, Caucasus, Turkmenistan, Iran. Introduced to Sri Lanka.

224. *Marpissa muscosa (Clerck, 1758) New data: 56b (1♀). Global distribution: Europe, Caucasus, Russia (Europe to Central Asia), Turkey, Japan.

225. Neon pictus Kulczyński, 1891 Literary data: L28 (Kolosvary 1938). Global distribution: South-eastern Europe to Central Asia.

226. Pellenes tripunctatus (Walckenaer, 1802) Literary data: L89 (Nikolić & Polenec 1981). New data: L59a (1♀). Global distribution: Europe, Caucasus, Russia to Central Asia, China.

227. Philaeus chrysops (Poda, 1761) Literary data: L26 (Kolosvary 1938), L89 (Simon 1878, Nosek 1904, Bresjanceva 1907). New data: 5b (1♀), 56b (1♀). Global distribution: Europe, North Africa to Middle East, Turkey, Caucasus, Russia (Europe to Far East), Iran, Central Asia, Afghanistan, China, Mongolia, Korea.

228. Phlegra fasciata (Hahn, 1826) Literary data: L28 (Kolosvary 1938). New data: 56a (1♀). Global distribution: Europe, Turkey, Caucasus, Russia to Central Asia, Iran, Afghanistan, India, China, Mongolia, Korea, Japan.

229. Pseudicius encarpatus (Walckenaer, 1802) Literary data: L12 (Damin 1896, 1900, Drensky 1936), L89 (Nikolić & Polenec 1981). New data: 56b (1♂). Global distribution: Europe to Central Asia.

230. Saitis imitatus (Simon, 1868) Literary data: L33a, sub Euophrys i. (Damin 1896, Drensky 1936), L33a, sub Euophrys imitata (Damin 1900), L89 (Nikolić & Polenec 1981). Global distribution: Croatia, Montenegro.

231. Salticus cingulatus (Pancer, 1797) Literary data: L28 (Kolosvary 1938), L89 (Nikolić & Polenec 1981). Global distribution: Europe, Turkey, Iran, Russia to Kazakhstan, Mongolia.

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232. Salticus mutabilis Lucas, 1846 Literary data: L33a, sub Epiblemum mutabile (Damin 1896, 1900, Drensky 1936). Global distribution: Europe, Savage Isl., Azores, Morocco, Egypt, Georgia, Argentina.

233. Salticus zebraneus (C.L. Koch, 1837) Literary data: L33a, sub Epiblemum zebraneum (Damin 1896, 1900), L33a, sub S. zebrinus (Drensky 1936), L89 (Nikolić & Polenec 1981). Global distribution: Europe, Turkey, Iran.

Scytodidae 234. Scytodes thoracica Latreille, 1804 Literary data: L28 (Kolosvary 1938), L89 (Deltshev 2008). Global distribution: Europe, North Africa, Turkey, Iran, temperate Asia to China, Korea, Japan. Introduced to North America, Argentina, India, Australia, New Zealand.

Segestriidae 235. Segestria florentina (Rossi, 1790) Literary data: L33a (Damin 1896, 1900, Drensky 1936). Global distribution: Europe to Georgia. Introduced to Brazil, Uruguay, Argentina.

236. *Segestria senoculata (Linnaeus, 1758) New data: L75a (1sa♀). Global distribution: Europe, Caucasus, Turkey, Iran, Japan.

Sparassidae 237. Micrommata virescens (Clerck, 1757) Literary data: L28, sub M. viridissima (Kolosvary 1938), L33a (Damin 1896, 1900, Drensky 1936). New data: L56a (1sa♀), L56b (1♂, 1sa♀). Global distribution: Europe, Turkey, Caucasus, Russia (Europe to Far East), Central Asia, China, Korea, Japan.

238. Olios argelasius (Walckenaer, 1806) Literary data: L33a, sub O. spongitarsis (Damin 1896, 1900, Drensky 1936), L89 (Nikolić & Polenec 1981). New data: L6b (1♀), L61 (1j). Global distribution: Mediterranean.

Tetragnathidae 239. Meta bourneti Simon, 1922 Literary data: L49 (Deltshev 2008), L89 (Deeleman-Reinhold 1974, 1977). Global distribution: Europe, Georgia, North Africa.

240. Meta menardi (Latreille, 1804) Literary data: L89 (Deltshev 2008). Global distribution: Europe, Turkey, Iran.

241. *Metellina mengei (Blackwall, 1869) New data: L56c (1♂), 75c (1♂). Global distribution: Europe to Caucasus, Iran, Russia (Europe to Altai).

242. Metellina merianae (Scopoli, 1763) Literary data: L7, L15a, L16, L17, L30c, L32, L37b, L37e, L44a, L48c, L53c, L79, L81, L86, L90, sub Meta m. (Kratochvíl 1935b), L10a, L15a, L15b, L20a, L30a, L30d, L37f, L44a, L51 (Naumova et al. 2016), L42a, L62, sub Meta m. (Deeleman-Reinhold 1974), L89 (Deltshev 2008).

NAUMOVA ET AL.

New data: L56a (1♂). Global distribution: Europe, Caucasus, Russia (Europe to Central Asia), Turkey, Iran.

243. Metellina segmentata (Clerck, 1757) Literary data: L28, sub Meta s. (Kolosvary 1938), L89, sub Meta s. (Nosek 1904, Bresjanceva 1907). Global distribution: Europe, Caucasus, Russia (Europe) to Central Asia, Turkey, Israel, China, Japan. Introduced to Canada.

244. Pachygnatha degeeri Sundevall, 1830 Literary data: L28 (Kolosvary 1938). New data: L56b (1♂). Global distribution: Europe, Caucasus, Russia (Europe to Far East), Turkey, Central Asia, China.

245. Tetragnatha extensa (Linne, 1758) Literary data: L28 (Kolosvary 1938), L89 (Nikolić & Polenec 1981). Global distribution: Europe, North America, Greenland, Caucasus, Iraq, Russia (Europe to Far East), Turkey, temperate Asia including China, Korea, Japan.

246. Tetragnatha montana Simon, 1874 Literary data: L33a, sub T. solandrii (Damin 1896, 1900), L89 (Nikolić & Polenec 1981). New data: L56b (1♂). Global distribution: Europe, Caucasus, Russia (Europe to Far East), Turkey, Central Asia.

247. Tetragnatha obtusa C. L. Koch, 1837 Literary data: L28 (Kolosvary 1938). New data: L6a (1♂). Global distribution: Europe, Caucasus, Russia (Europe to Far East), Turkey, Central Asia.

Theridiidae 248. *Anelosimus vittatus (C. L. Koch, 1836) New data: L56b (1♀). Global distribution: Europe, Caucasus, Turkey.

249. Argyrodes argyrodes (Walckenaer, 1841) Literary data: L33a, sub A. gibbosus (Kolosvary 1938), L33a, sub Argyrodina a (Nikolić & Polenec 1981), L70, sub Conopistha argyrodes (Brignoli 1966). Global distribution: Mediterranean to West Africa, Seychelles.

250. Asagena phalerata (Panzer, 1801) Literary data: L28 (Kolosvary 1938), L89, sub Steatoda p. (Knoflach 1996). New data: L8 (1♂), L59b (1♀). Global distribution: Europe, Caucasus, Russia (Europe to Far East), Turkey, Central Asia, China, Korea.

251. *Crustulina guttata (Wider, 1834) New data: L37d (1♀), L59b (1♂). Global distribution: Canary Is., Europe, Caucasus, Russia (Europe to South Siberia), Central Asia, China, Korea, Japan.

252. *Enoplognatha latimana Hippa & Oksala, 1982 New data: L56b (1♀). Global distribution: Europe, North Africa, Turkey, Caucasus, Russia (Europe) to Central Asia, Canada.

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253. Episinus truncatus Latreille, 1809 Literary data: L28, sub E. lugubris (Kolosvary 1938). Global distribution: Europe, Caucasus, Turkey.

254. Euryopis episinoides (Walckenaer, 1847) Literary data: L12, sub E. acuminata (Damin 1896, 1900, Drensky 1936). Global distribution: Mediterranean to Turkey, Israel. Introduced to Reunion, India, China.

255. Lasaeola convexa (Blackwall, 1870) Literary data: L33a (Damin 1896, 1900), L33a, sub Dipoena c. (Drensky 1936), L89 (Nikolić & Polenec 1981). Global distribution: Mediterranean.

256. Latrodectus tredecimguttatus (Rossi, 1790) Literary data: L89, sub L. mactans (Nikolić & Polenec 1981). Remark: Doubtful. Global distribution: Mediterranean, Ukraine, Caucasus, Russia (Europe to South Siberia), Kazakhstan, Central Asia, China.

257. Parasteatoda lunata (Clerck, 1757) Literary data: L35, sub Theridium formosom (Damin 1896, 1900), L89, sub Theridium formosom (Kolosvary 1938), L89 (Nikolić & Polenec 1981). New data: L56b (1♂, 1♀). Global distribution: Europe, Caucasus, Russia (Europe to Far East), Turkey, Israel.

258. *Phylloneta impressa (L. Koch, 1881) New data: L59b (1♀). Global distribution: Europe, Caucasus, Russia (Europe to Far East), Turkey, Kazakhstan, Central Asia, China, India, North America.

259. Robertus frivaldszkyi (Chyzer, 1894) Literary data: L63a (Knoflach & Thaler 2000). Global distribution: Central and South-eastern Europe.

260. Robertus mediterraneus Eskov, 1987 Literary data: L38 (Knoflach & Thaler 2000). Global distribution: Mediterranean, Switzerland, Austria, Eastern Europe, Caucasus.

261. Simitidion simile C.L. Koch, 1836 Literary data: L31, L33a, L35, sub Theridion s. (Damin 1900), L33a, L35, sub Theridium s. (Drensky 1936), L89, sub Theridion s. (Nikolić & Polenec 1981). Global distribution: Europe, North Africa, Turkey, Israel, Caucasus, Central Asia, Canada.

262. *Steatoda bipunctata (Linnaeus, 1758) New data: L75b (2♂♂). Global distribution: Europe, Caucasus, Russia (Europe to Far East), Turkey, Iran, Central Asia, China. Introduced to South America.

263. Steatoda paykulliana (Walckenaer, 1806) Literary data: L12, L33a, sub Lithyphantes paykullianus (Damin 1896, 1900, Drensky 1936). Global distribution: Europe, Mediterranean to Central Asia.

264. Steatoda triangulosa (Walckenaer, 1802) Literary data: L89 (Deltshev 2008).

NAUMOVA ET AL.

Global distribution: Europe, Caucasus, Russia (Europe to Far East), Turkey, Central Asia. Introduced to Canada, USA, Canary Isl.

265. *Theridion betteni Wiehle, 1960 New data: L59b (2♀♀). Global distribution: Europe, Turkey.

Thomisidae 266. Bassaniodes graecus (C. L. Koch, 1837) Literary data: L33a, sub Xysticus g. (Chyzer & Kulczyński 1897, Damin 1900, Drensky 1936, Nikolić & Polenec 1981). Global distribution: Balkans, European Russia, Israel, Iraq, Turkey, Ukraine.

267. *Bassaniodes robustus (Hahn, 1832) New data: L23c (1♂), L55b (1♂). Global distribution: Europe to Central Asia.

268. Diaea dorsata (Fabricius, 1777) Literary data: L33a (Damin 1896, 1900, Drensky 1936), L89 (Nikolić & Polenec 1981). New data: L58 (1♂). Global distribution: Europe, Caucasus, Russia (Europe to Middle Siberia), Turkey, Japan.

269. Heriaeus hirtus (Latreille, 1819) Literary data: L28, sub H. savignyi (Kolosvary 1938), L33a, sub H. savignyi (Damin 1896, 1900, Drensky 1936), L89 (Nikolić & Polenec 1981). Global distribution: Europe, Turkey, Caucasus.

270. Heriaeus oblongus Simon, 1918 Literary data: L33a, L35, sub H. hirsutus (Damin 1896, 1900, Drensky 1936). New data: L56b (1♂). Global distribution: Europe, Caucasus, Russia (Europe) to Central Asia.

271. *Misumena vatia (Clerck, 1758) New data: L56b (1sa♀). Global distribution: Europe, Caucasus, Russia (Europe to Far East), Turkey, Central Asia, China, Korea, Japan, North America.

272. Ozyptila confluens (C. L. Koch, 1845) Literary data: L28, sub O. strandi (Kolosvary 1938, Nikolić & Polenec 1981). Global distribution: Southern Europe, Syria.

273. Pistius truncatus (Pallas, 1772) Literary data: L33a (Damin 1896, 1900, Drensky 1936). Global distribution: Europe, Caucasus, Russia (Europe to Far East), Turkey, Iran, China, Japan.

274. *Psammitis ninnii (Thorell, 1872) New data: L56b (1♂). Global distribution: Europe, Caucasus, Russia (Europe to South Siberia), Turkey, Central Asia

275. Runcinia grammica (C.L. Koch, 1837) Literary data: L33a, L35, sub R. lateralis (Damin 1896, 1900, Drensky 1936), L89 (Nikolić & Polenec 1981). Global distribution: Europe, Middle East to Iran, Russia (Europe to West Siberia), Central Asia, China, Japan. Introduced to St. Helena, South Africa.

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276. Spiracme striatipes (L. Koch, 1870) Literary data: L26, sub Xysticus s. (Kolosvary 1938). Global distribution: Europe, Caucasus, Russia (Europe) to Central Asia, Turkey, Iran, China.

277. Synema globosum (Fabricius, 1775) Literary data: L12 (Damin 1896, 1900, Drensky 1936), L28 (Kolosvary 1938). New data: L56a (1♂, 3♀♀). Global distribution: Europe, Caucasus, Russia (Europe to Far East), Turkey, Israel, Iran, Central Asia, China, Korea, Japan.

278. Thomisus onustus Walckenaer, 1806 Literary data: L12, L33a, sub T. albus (Damin 1896, 1900, Drensky 1936), L28, sub T. albus (Kolosvary 1938). Global distribution: Europe, Selvagens Is., North Africa, Caucasus, Russia (Europe to South Siberia), Turkey, Israel, Central Asia, Iran, China, Korea, Japan.

279. Tmarus piger (Walckenaer, 1802) Literary data: L28 (Kolosvary 1938), L33a (Damin 1896, Drensky 1936), L35 (Damin 1896, 1900, Drensky 1936), L70 (Brignoli 1966). Global distribution: Europe, Caucasus, Russia (Europe to Far East), Turkey, Kazakhstan, China, Korea, Japan.

280. Xysticus acerbus Thorell, 1872 Literary data: L12, L35 (Damin 1896, Drensky 1936). Global distribution: Europe to Central Asia, Russia (Europe to Far East).

281. Xysticus brevidentatus Wunderlich 1995 Literary data: L24a (Jantscher 2003). Global distribution: Balkans, Italy.

282. *Xysticus cristatus (Clerck, 1757) New data: L56a (4♀♀). Global distribution: Europe, Caucasus, Russia (Europe to Middle and South Siberia), Turkey, Iran, Central Asia, China, Korea, Japan.

283. *Xysticus gallicus Simon, 1875 New data: L60 (1♀). Global distribution: Europe, Caucasus, Turkey, Iran.

284. Xysticus kochi Thorell, 1872 Literary data: L3 (Giltay 1932, Drensky 1936), L89 (Nikolić & Polenec 1981). New data: L56b (1♀). Global distribution: Europe, Mediterranean to Central Asia.

285. Xysticus lanio C.L. Koch, 1835 Literary data: L33a (Damin 1896, 1900, Drensky 1936), L89 (Nikolić & Polenec 1981). New data: L59a (1♀). Global distribution: Europe, Caucasus, Russia (Europe) to Central Asia, Turkey.

Uloboridae 286. Hyptiotes paradoxus (C. L. Koch, 1834) Literary data: L33a, L35 (Damin 1896, 1900, Drensky 1936). Global distribution: Europe, Turkey, Caucasus.

NAUMOVA ET AL.

287. Uloborus plumipes Lucas, 1846 Literary data: L12 (Damin 1896, 1900, Drensky 1936). Global distribution: Europe, Africa, Yemen, Pakistan, Philippines. Introduced to Argentina, Japan.

288. Uloborus walckenaerius Latreille, 1806 Literary data: L33a, L35 (Damin 1896, 1900, Drensky 1936). Global distribution: Europe, Madeira, Caucasus, Russia (Europe to Far East), Turkey, Iraq, Iran, Central Asia, China, Korea, Japan.

*Zodariidae 289. *Zodarion aculeatum Chyzer, 1897 New data: L56a (10♂♂). Global distribution: Bulgaria, Republic of North Macedonia, Romania, Serbia.

Acknowledgements We would like to thanks to the colleagues who provided us spiders material, the late B. Petrov (NMNH), M. Langourov (NMNH), P. Beron (NMNH), S. Lukanov (IBER), G. Hristov (IBER), V. Pešić (UM), D. Pešić (UM) and A. Schönhofer (Mainz, Germany). Thanks to Fulvio Gasparo for the helpful review of the manusctipt. The research was partially supported by the projects ―Cybertaxonomic approach to phylogenetic studies of model invertebrate genera (Invertebrata, Arachnida, Insecta), clarifying the problems of origin, formation and conservation of the Invertebrate Fauna of the Balkan Peninsula‖ (National Science Fund, Ministry of Education, Youth and Science of the Republic of Bulgaria, Grant KP-06-Н21/1-17.12.2018).

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