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ANNUAL REPORT FOR AWARD # 9977958
Michael E Irwin ; U of Ill Urbana-Champaign PEET: A World Monograph of the Therevidae (Insecta: Diptera)
Participant Individuals: CoPrincipal Investigator(s) : David K Yeates; Brian M Wiegmann Senior personnel(s) : Donald Webb; Gail E Kampmeier Graduate student(s) : Kevin C Holston; Martin Hauser Post-doc(s) : Mark A Metz Undergraduate student(s) : Amanda Buck; Melissa Calvillo Other -- specify(s) : Kristin Algmin Graduate student(s) : Hilary Hill Post-doc(s) : Shaun L Winterton Technician, programmer(s) : Brian Cassel Other -- specify(s) : Jeffrey Thorne Post-doc(s) : Christine Lambkin Other -- specify(s) : Ann C Rast Senior personnel(s) : Steve Gaimari Other -- specify(s) : Beryl Reid Technician, programmer(s) : Joanna Hamilton Undergraduate student(s) : Claire Montgomery; Heather Lanford High school student(s) : Kate Marlin Undergraduate student(s) : Dmitri Svistula Other -- specify(s) : Bradley Metz; Erica Leslie Technician, programmer(s) : Jacqueline Recsei; J. Marie Metz
Participants' Detail
Partner Organizations: Illinois Natural History Survey: Financial Support; Facilities; Collaborative Research
Schlinger Foundation: Financial Support; In-kind Support; Collaborative Research
The Schlinger Foundation has been a strong and continuing partner of the therevid PEET project, providing funds for personnel (students, scientific illustrator, data loggers, curatorial
assistant) and expeditions, including the purchase of supplies, to gather unknown and important taxa from targeted areas around the world.
CSIRO Division of Entomology: In-kind Support; Facilities; Collaborative Research
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University of Illinois at Urbana-Champaign: Financial Support; In-kind Support; Facilities
North Carolina State University: Financial Support; In-kind Support; Facilities
NC State College of Agriculture and Life Sciences (CALS) are providing matching funds to support graduate student Hilary Hill. The university also employs co-PI Brian Wiegmann and the NCSU-based staff.
USDA Systematic Entomology Laboratory: Collaborative Research
Smithsonian Institution Museum of Natural History: Collaborative Research
North Carolina Agric. Research Service: Financial Support
The North Carolina Agricultural Research Service is providing matching funds to support graduate student Hilary Hill.
Australian Biological Resources Study: Financial Support
PI’s Irwin and Yeates have been awarded a grant to contribute to the costs of field expeditions by ABRS for 2001 and 2002.
Other collaborators:
The Therevid PEET project has borrowed therevid specimens from over 100 collections from around the world. A number of collaborators, worldwide, are servicing Malaise traps and capturing therevid specimens for our studies. Among these, we highlight the following: Gerardo Barría, Roberto González, Carlos Quiroz, Patricia Larraín, Hernan Vasques [Chile]; Tim Maggs, Ray Miller [South Africa]; Erick Yabar [Peru]; Karl Spitzer [Czech Republic] Frank Parker [Utah]; Evert Schlinger, Sally Regan, Saul Frommer [California]; Joseph Maddox [Missouri].
With this grant, specific contacts have been made with the following researchers: Dr. Christian Schmid-Egger (Germany/Berlin); Dr. Peter Tschorsnig (Germany/ Stuttgart) Museum for Natural History, Stuttgart (SMNS); Fritz Geller-Grimm (Germany/Frankfurt) Museum for NAtural History Wiesbaden; Stephan Blank (Germany/Eberswalde) DEI Deutsches Entomologisches Institut; Dr. Joachim Ziegler (Germany/Eberswalde) DEI Deutsches Entomologisches Institut; Dr. Manfred Niehuis (Germany/Albersweiler); Dr. Leif Lyneborg, emeritus, Zoological Museum, Copenhagen; Dr. Karel Spitzer (Czech Republic); Dr. Jean Chazeau (New Caledonia); Dr. Frank Parker; Dr. Steve Goodman, World Wildlife Fund (Madagascar); Dr. Patricia Wright (Madagascar); Drs. Brian https://www.fastlane.nsf.gov/cgi-bin/NSF_PrjRpt?@@@___0212___ps1v4APbehh%3Aw%3AKNBQ85Q2Q%3At%3APAqbu%3AyMm9hem3uF%3A5N2KH558 ... Page 2 of 25 Preview before Submission[NSF Project Report - Version 1.2] 5/30/03 15:25
Fisher and Charles Griswald (California Academy of Sciences); Rasolondalao Harin'Hala Hasinjaka (Schlinger Foundation, Madagascar); Evert I Schlinger (Schlinger Foundation); F. Christian Thompson (USDA-SEL, Smithsonian Institution); and Scott J. Robinson [Beckman Institute, UIUC, Urbana, IL].
The following people have been added to list of collectors and collection curators with whom we have collaborated: Ben Brugge [Natural History Museum, Amsterdam, Netherlands]; Dr. Fritz Gusenleitner [Landesmuseum, Linz, Austria]; Menno Reemer [Nationaal Natuurhistorisch Museum, Leiden, Netherlands]; Tony van Harten [General Department of Plant Protection, Sana'a, Yemen]; Mohamed El Habib Ben Jamaa [INRGREF, Ariana, Tunisia]; Dr. Karl Adlbauer [Landesmuseum, Graz, Austria]; Bernhard Merz [Museum d'Histoire Naturelle, Geneva, Switzerland] ; Jeanne Charbonnel [Museum National d'Histoire Naturelle, Paris, France]; Dr. Jens-Hermann Stuke [Leer, Germany] ; Dr. Franco Mason [State Forestry Corps, Verona, Italy]; Prof. Dr. sc. nat. Bernhard Klausnitzer [Dresden, Germany]; Dr. Amnon Freidberg [Tel Aviv University, Israel]; Dipl.-Biol. Joerg Weipert [Institut fuer biologische Studien, Plaue, Germany] ; Claus Claussen [Flensburg, Germany]; Juergen Wiesner [Wolfsburg, Germany]; Dr. Wolfgang Schawaller [Naturkundemuseum, Stuttgart, Germany]; Dr Ante Vujic [Institute of Biology, Novi Sad, Yugoslavia]; Dr. Oleg G. Gorbunov & Prof. Dr. Sergei Golovatch [Institute for Problems of Ecology & Evolution, Russian Academy of Sciences, Moscow, Russia]; Prof. Dr. Horst Aspoeck [Vienna, Austria]; Jose A Rafael [INPA - Entomologia, Manaus, Amazonas Brazil]; Dr. Diego Medan [Buenos Aires, Argentina]; Dr. Matthias Buck, [University of Guelph, Ontario Canada ]; Weston Opitz [Kansas Wesleyan University Department of Biology, Salina, Kansas]; Luciana Musetti [Ohio State University, Columbus, Ohio]; Magdi S. A. El Hawagry [Entomology Dept. Cairo University, Giza, Egypt]; Wolfgang Schacht [Zoologische Staatssammlung Munic, Germany]; Uwe Kallweit [Staatliche Naturhistorische Sammlungen, Dresden, Germany]; Dr. Karla Schneider [Institut fuer Zoologie der Martin-Luther-Universitaet, Halle, Germany]; Claudio José Barros de Carvalho [Universidade Federal do Paraná, Brasil]; Helmut Heider [Orchidarium Villa Tunari, Bolivia]; Noel Starick [CSIRO, Canberra, Australia]; Roger Buttermore [Tasmanian Museum & Art Gallery, Hobart, Australia]; Andras Szito [Western Australian Depart. of Agriculture, Perth, Australia]; Keith Arakaki [Bishop Museum, Honolulu, USA]; Gavin Dally [Museum & Art Gallery of the Northern Territory, Darwin, Australia]; John Donaldson [Queensland Depart. of Primary Industries, Brisbane, Australia]; Terry Houston [Western Australian Museum, Perth, Australia]; Susan Wright [Queensland Museum, Brisbane, Australia]; Catriona McPhee [Victorian Museum, Melbourne, Australia]; Murray Fletcher [Agricultural Scientific Collection, Orange, Australia]; Jan Forrest [South Australian Museum, Adelaide, Australia]; Greg Baker [SARDI, Adelaide, Australia]; John Chainey [Natural History Museum, London, England]; Margaret Schneider [University of Queensland Insect Collection, Brisbane, Australia]; John Marris [Lincoln University, Canterbury, New Zealand]; Trevor Crosby [Landcare, Christchurch, New Zealand]; Daniel Bennett [The University of Washington Fish Collection, Seattle, Washington]; Yoshizawa Kazunori [Systematic Entomology, Hokkaido University, Japan]
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[Added in Yr 3] Dr Ken Green: Australian Institute of Alpine Studies,
Jindabyne, NSW, AUSTRALIA; Emily Sherbrooke (Southwest Research Station, Portal, Arizona); Rustem Hayat (The Ataturk University, Erzurum, Turkey), Alvaro Herrera (INBio, Costa Rica); Manuel Zumbado (INBio, Costa Rica); Christel and Hans Werner Hoffeins (Hamburg, Germany); Augusto Henriques (INPA, Manaus Brazil), J. Hutto (INHS, Champaign, IL), Jeff
Cumming (CNC, Ottawa, Canada); Drs Arkady Lelej and Sergey Storozhenko
at the Russian Academy of Sciences, Department of Entomology, Institute of Biology and Pedology, Vladivostok, Russia. Mikhail B. Mostovski, Paleontological Institute, Russian Academy of Sciences, Moscow. Thomas Pape (Swedish Museum of Natural History, Stockholm).
Activities and findings:
Research and Education Activities:
see attached pdf
Findings:
see attached pdf
Training and Development:
Michael E. Irwin, PI, University of Illinois at Urbana-Champaign, has been elected to two boards that involve biodiversity issues: the All Species Advisory Board, and the Discover Life in America Governing Board.
David K. Yeates, Co-Principal investigator on the therevid PEET team, has relocated to CSIRO Entomology in Canberra where he has taken a position as Principal Research Scientist and Head of the Australian National Insect Collection (ANIC), the world's largest collection of Australian insects (See http://www.ento.csiro.au/research/natres/anic.htm). David will also continue his systematics research in Diptera at ANIC as part of the Natural Resources and Biodiversity Program.
Longlong Yang was awarded a Ph.D. at NCSU at the end of the earlier PEET award (DEB 9521825). He had been supported by matching funds from NCSU College of Agriculture and Life Sciences. Longlong successfully defended his dissertation entitled, 'Molecular phylogenetics of the Therevidae and their position among the families of the Asiloidea (Insecta: Diptera)' under the direction of co-PI Wiegmann. Dr. Yang accepted a postdoc in Bar Harbor, Maine.
Hilary Hill, masters student at NCSU with Wiegmann, began
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coursework and laboratory research on Therevidae at NCSU. Her assistantship is supported as matching funds provided by the North Carolina Agriculture Research Service and NC State College of Agriculture and Life Sciences (CALS). Ms. Hill is currently taking classes toward the MS degree in entomology, including ENT502 (Insect Systematics in Fall 2000), ENT503 (Insect Morphology and Physiology in Spring 2001) and ENT 591O (Molecular Systematics in Spring 2001). Her MS research involves molecular characterization and sequencing of the opsin gene in Therevinae. Her laboratory training has included DNA extraction and purification, PCR, automated sequencing, molecular cloning, sequence data editing, alignment and phylogenetic analysis. A graduate committee at NCSU consisting of 3 Entomology faculty members, B. Wiegmann, L. Deitz, E. Vargo, has met and approved an official 'Plan of Work.'
Shaun L. Winterton, continues postdoctoral work begun in 2000 in the Wiegmann lab on the molecular phylogenetics of the orthorrhaphous brachyceran family Acroceridae. He is funded through a postdoctoral research fellowship to NCSU from the Schlinger Foundation. Dr. Winterton remains active in therevid systematics and is a collaborator/consultant to the current project.
Kevin Holston and Martin Hauser, UIUC graduate students, visited PI Wiegmann’s lab for 3 months, Feb – April , 2001. Both students attended the class in molecular systematics taught by Wiegmann. Each student designed projects and conducted a laboratory rotation involving amplification and sequencing of the mitochondrial and nuclear genes for therevines (Holston) and phycines (Hauser). Training included PCR techniques, agarose gel fractionation, PCR purification, automated DNA sequencing and DNA sequence analysis using STADEN package software and GDE 2.2. Nucleotide data was collected for approximately 70 species and several outgroups.
Jeffrey Thorne, Associate Professor, NCSU, Department of Statistical Genetics, is developing methods for inferring divergence times from molecular phylogenetic data without the assumption of a molecular clock. These methods are being used to develop a time scale for major events in brachyceran fly evolution based on data generated in the Wiegmann lab for therevids and other Brachycera.
Christine Lambkin joined the therevid PEET team as a Postdoctoral Fellow at CSIRO Entolomology Section, Canberra, Australia, in May 2001, to continue the classification of Australian stiletto flies begun by Shaun Winterton. Christine completed her doctorate at the University of Queensland in Brisbane, which comprised a systematic revision of a tribe of bee flies (Diptera: Bombyliidae: Asiloidea), a close relative of the Therevidae. Chris, who has provided some technical assistance to the therevid team in the past, continues the work on the phylogenetics the Therevidae, concentrating on the genus Ectinorhynchus and its allies, in arid and semiarid environments of Australia. Chris is also investigating the web-based dissemination of interactive keys, data, and monographic work. She examines the systematic methods that we use in phylogenetic analysis and is heavily involved in the training and development of scientific knowledge in this area in Australia, America, and Europe.
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Christine Lambkin successfully garnered Official Travel funding from CSIRO to visit and obtain type specimens from the Natural History Museums in London and Paris, attend the ‘Tree of Life’ conference in New York in May 2002 and PEET IV Conference in June 2002.
Ann Coddington Rast is an intern working on scientific illustration with Jill Marie Mullett. She is working on Diptera illustrations for the therevid PEET project primarily for the experience and knowledge she can gain by illustrating fly images. Ann is an assistant professor in the Department of Fine and Applied Arts at UIUC. She is learning to produce professional illustrations of Diptera using both traditional and digital means.
In year 1 of the therevid PEET grant, Mark A. Metz advanced to candidacy of his Ph.D. by passing his oral examinations in the department of entomology; thereafter, he transferred to the department of Natural Resources and Environmental Sciences where he expects to obtain his Ph.D. degree by summer 2002. In 2001, Mark was listed among the teachers rated outstanding by their students and received an award for his teaching of Entomology 301, General Entomology, for which he was a teaching associate and guest lecturer during the summer of 2000. He published his first therevid article (Ann. Ent. Soc. Am. 93: 977-1018) and has a second article in press in Environmental Entomology on the bioecology of microtrombidiid mites and brachyceran Diptera (See Journal Publications). His dissertation study is focusing on the generic analysis of higher therevines based on morphology.
Mark A. Metz obtained his Ph.D. from the Dept of Natural Resources & Environmental Sciences at UIUC in May 2002, after successfully defending and depositing his dissertation on Systematics within the Therevinae. He published his second therevid article (Env. Ent. 30: 903-908), published a revision within Syrphidae (Studia Dipterologica 8: 225-256) in collaboration with F. Christian Thompson, submitted a co-authored manuscript on the revision of the therevine genus Pandivirilia in collaboration with D. W. Webb, and has another publication reviewing the species of Psilocephala ready to submit (See Journal Publications). He plans to continue as a post-doctoral fellow with Mike Irwin studying the homology of the male genitalia in the therevoids and collaborating on several other generic revisions within therevids.
Martin Hauser gave an invited presentation about the use of the enviromental electron microscope for entomologists at a symposium on scientific illustration in Montreal, in December 2000 (see Invited Presentations). At NCSU in 2001, he analyzed the DNA data to create a phylogenetic tree with the emphasis on the basel members of this family, the Phycinae and Xestomyzinae using genes ef1 and 28S were used. In April Martin participated on a field trip to Naggs Head (NC) to collect a species of Tabuda. In May/June 2001 he was part of an international expedition to the Republik of Kazakhstan and in June/July he led a collecting expedition to Tunisia.
During year 2 of the therevid PEET, Martin visited the collections of the Zoological Museum in Moscow and St. Petersburg in December 2001-January 2002 where he met amongst others https://www.fastlane.nsf.gov/cgi-bin/NSF_PrjRpt?@@@___0212___ps1v4APbehh%3Aw%3AKNBQ85Q2Q%3At%3APAqbu%3AyMm9hem3uF%3A5N2KH558 ... Page 6 of 25 Preview before Submission[NSF Project Report - Version 1.2] 5/30/03 15:25
the most important Russian therevid worker, Prof. Vladimir Zaitzev. In April, Martin Hauser successfully defended and deposited his Masters thesis on the 'Revision of the genus Ammonaios' and will receive his diploma in May 2002. He will present his work at the Evolution meetings in Champaign, IL in July 2002 and the 5th International Congress of Dipterology in Brisbane in September 2002.
Kevin Holston travelled to Stockholm, Paris, Berlin, St. Petersburg, Vienna, and Copenhagen from August-December 2000 to examine type specimens of the family Therevidae, particularly those in the genus, Thereva. Although he was fortunate enough to see numerous Krober types in Berlin, Krober’s types stored in Hamburg were destroyed during WWII. Kevin’s Copenhagen trip was supported by the American Scandinavian Society and the Schlinger Foundation. While in Copenhagen, he worked with Leif Lyneborg and expanded the character matrix for the genus Thereva from 77 to more than 160 characters. From February-April 2001, he used molecular techniques and applied them to the analysis of the genus Thereva to create a phylogenetic tree of this speciose genus.
During year 2 of the therevid PEET grant, Kevin Holston, graduate student at UIUC, has focused on developing molecular and morphological datasets for phylogenetic analyses of the genus Thereva Latreille. Collecting trips (see Expeditions in Activities) between May and August, 2001, substantially increased the taxonomic scope of his phylogenetic work. His dissertation will include a higher-level molecular analysis of therevine genera using 28s rDNA, EF 1-alpha nDNA, and 16s mtDNA. Morphological characters and mitochondrial DNA sequences will be used in a combined analysis of the worldwide Thereva fauna and will allow Holston to address trans-continental biogeographic questions concerning this genus. A chapter of his dissertation will be devoted to a revision of the North American species of Thereva.
Gail Kampmeier took the FAST3 (at UIUC) beginning and advanced workshops on DreamWeaver 4.0, the webpage design software [July 2001]. She is Chair of the INHS WebTeam and helps supervise hourly web designer’s activities and interactions with INHS WebTeam. This training and work with a designer will help in improving the therevid PEET site.
Jill Marie Mullett received her Masters in Art Education in May 2001. As the scientific illustrator based at UIUC, she has progressed rapidly in the rendering of computerized images over the past year. Her rapid progress made it possible for her to take the place of George Venable (Smithsonian Instition) in an invited talk about computerized rendering of scientific illustrations in Montreal (see Invited Presentations under Activities), when he was forced to cancel. Six months earlier, this would have been inconceivable. She has now been invited to present this talk at the annual meeting of the Guild of Natural Science Illustrators in August 2001, in Bar Harbor, Maine.
Don Webb is working on a revision and phylogeny of the genera Pandivirillia and Dichoglena and a revision of the genus Spiriverpa. In a related project, he is also describing the genus Chrysopilus (Rhagionidae), which has 15 new species from New Caledonia. https://www.fastlane.nsf.gov/cgi-bin/NSF_PrjRpt?@@@___0212___ps1v4APbehh%3Aw%3AKNBQ85Q2Q%3At%3APAqbu%3AyMm9hem3uF%3A5N2KH558 ... Page 7 of 25 Preview before Submission[NSF Project Report - Version 1.2] 5/30/03 15:25
Kristin (Mrozinski) Algmin, a recent graduate in Biology at the University of Illinois (UIUC) and former student hourly worker on the database team, is now working full-time as an Academic hourly in the Irwin lab. She continues to work on the database team, but is also learning more in depth functioning of Mandala and FileMaker Pro, and spends more time troubleshooting problems, training new students, and working to assure data integrity. Kristin is also taking care of curatorial needs in the collection, pinning of specimens from alcohol samples and identifying therevid flies to genus level. She is also studying the feeding patterns of therevid larvae from Western California. She plans to submit a manuscript on her work later this summer [2002].
Joanna Hamilton, Technical Officer employed by CSIRO, has been working for David Yeates curating and labeling the Therevidae collected from expeditions and testing a key to Australian genera developed by Chris Lambkin.
Beryl Reid, Australian National Insect Collection Volunteer since November 2001, has spent 2-days per week assisting Chris Lambkin with the curation of the Therevidae.
Steve Gaimari led a 7-week expedition to Bolivia in April 2001 to the Andean foothills north of La Paz, to the towns of Mapiri, Guanay, Coroico, and Chulumani; and also to the Cochabamba area. The expedition was partially funded by another grant from NSF. The other participants were Wayne Mathis (PEETster from the Smithsonian Diptera group), Amnon Freidberg, Frank Parker, Allen Norrbom, Brian Brown, and Giar-Ann Kung. In July 2001, Steve Gaimari accepted the position of Associate Insect Biosystematist (Dipterist) for the California State Collection of Arthropods, California Department of Food and Agriculture, Sacramento.
F. Christian Thompson (USDA-ARS-SEL at the Smithsonian) visited Gail Kampmeier and the therevid PEET group at UIUC 17-21 October, collaborating on database and web issues, upcoming publications, and plans for therevid PEET II.
YEAR 3 Additions: Wiegmann lab continued data collection and phylogenetic data analyses in collaboration with student projects of Hauser, Holston, and Hill.
Martin Hauser received his MS in May 2002 and since then he is working on his PhD about the phylogeny of the basal therevid groups Phycinae and
Xestomyzinae. Martin Hauser's assistantship is supported by Schlinger
Foundation funds.
Kevin Holston completed his dissertational research on Thereva and received his degree from the University of Illinois at Urbana-Champaign (Department of Entomology) in May 2003. Holston was a collaborator on the Fauna Europaea project, contributing an annotated checklist of European Therevidae. He was also a student delegate to the 5th International Congress of Dipterology, University of Queensland, Brisbane, https://www.fastlane.nsf.gov/cgi-bin/NSF_PrjRpt?@@@___0212___ps1v4APbehh%3Aw%3AKNBQ85Q2Q%3At%3APAqbu%3AyMm9hem3uF%3A5N2KH558 ... Page 8 of 25 Preview before Submission[NSF Project Report - Version 1.2] 5/30/03 15:25
Australia. Dr. Holston was awarded a postdoctoral fellowship in the NSF’s International Research Fellowship Program. He will be revising the European species of Thereva and building species pages for the web. Holston will be based in Stockholm, Sweden and working under the supervision of Thomas Pape. The fellowship is for two years starting June 2003.
Mark Metz finished a 6-mo post doc in the Irwin lab working on the homologies in the male and female genitalia of the therevoid families of Asiloidea, the Therevidae of New Caledonia, and the functional morphology of the genitalia of Prorates ballmeri Nagatomi and Liu (see Metz et al. 2002 in publications list). In October 2002 he accepted a post doc at the Pennsylvania State University’s Frost Entomological Museum. He continues to work with the therevid PEET project, primarily collaborating with Don Webb on papers and research. He was a co-PI along with other therevid PEET
graduates in the ambitious proposal to the NSF Planetary Biodiversity
Inventory, “Crossing the Finish-Line: A World Monograph of the Therevidae (Insecta: Diptera).
Hilary Hill completed her masters project in the Dept. of Entomology at North Carolina State University in March 2003. Her assistantship was supported as matching funds provided by the North Carolina Agriculture Research Service and NC State College of Agriculture and Life Sciences (CALS). Her work
evaluated the phylogenetic utility and information content of nuclear protein coding genes, opsin rh1 and CAD in the Therevidae and among their closest relatives. Hilary is currently a Research Associate with USDA/APHIS/PPQ in Raleigh NC.
During PEET IV at Berkeley, CA (June 2002), J. Marie Metz collaborated with illustrators working with the PEET project of Dr. James (Steve) Ashe at Kansas State University. She was subsequently invited to Kansas in August to continue this collaboration, which resulted in improved methodologies for producing digital illustrations. Ms. Metz subsequently taught these improved methodologies in invited workshops that she was invited to present in St. Paul, MN and Sacramento, CA.
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The workshop in Minnesota also sprung out of interactions at PEET IV, where J. Marie Metz taught workshops in digital illustration. She was asked by fellow PEETster, Ralph Holzenthal of the University of Minnesota, to conduct a 3-day workshop in November 2002, teaching digital illustration techniques to entomology students.
Therevid PEET collaborator Dr. Stephen Gaimari (California Department of Food & Agriculture, Sacramento), funded a visit to California by J. Marie Metz in December 2002 to give a one-week workshop on rendering digital illustrations.
Jacqueline Recsei has been working for David Yeates sorting, specimen
curation, labeling, and databasing the Therevidae collected from expeditions, participating fully in field work, and testing a key to Australian genera developed by Chris Lambkin.
Erica Leslie, a new Australian National Insect Collection (ANIC) Volunteer, spent 2-days per week assisting Chris Lambkin with the return and assimilation of Anabarhynchus material into Australian collections following publication of the revision. Through the experience gained, Erica has now obtained a part-time position databasing Australian fruit flies in ANIC into the CSIRO Biolink.
Brad Metz obtained his B.S. in Biology at UIUC in 2002 and has been working half time with the Irwin lab in 2003, learning to curate flies, enter specimens in the Mandala database, retrospectively obtain geographic coordinates and verify locality information for specimens in the database so that they may be mapped, electronically archive illustrations created by traditional means, and do minor database troubleshooting. Brad enjoys being able to transfer the knowledge he gains in the Irwin lab working with Photoshop and engaging in discussions about science and other computer applications to other jobs in his life. Brad will be graduate school in entomology at Texas A&M starting in fall 2003 with a one-year fellowship.
Outreach Activities:
Great Smoky Mountains National Park ATBI. Co-PI Wiegmann is a co-organizer of the Diptera TWIG for the All Taxon Biotic Inventory of the Great Smoky Mountains National Park. https://www.fastlane.nsf.gov/cgi-bin/NSF_PrjRpt?@@@___0212___ps1v4APbehh%3Aw%3AKNBQ85Q2Q%3At%3APAqbu%3AyMm9hem3uF%3A5N2KH55 ... Page 10 of 25 Preview before Submission[NSF Project Report - Version 1.2] 5/30/03 15:25
Wiegmann has worked to build support for the project among dipterists and other entomologists through presentations at the Entomological Society of America National Meeting, Entomological Collections Network Annual Meeting, ESA- South Eastern Branch Meeting, and North American Dipterists Society Meeting. The Wiegmann lab at NCSU has been collecting and sorting Diptera collected in the park as part of the ATBI.
Gail Kampmeier participated in the ACES Open House at the booth promoting the upcoming Biodiversity Blitz at Allerton Park in June 2001 (http://www.conted.uiuc.edu/rapcc/bioblitz/). The display encouraged visitors to match the identifications in a list with a large photo collage display of Illinois biota.
David Yeates and the therevid PEET team in Australia are organizing the 5th International Congress of Dipterology, Brisbane Australia 30 Sept-5 Oct 2002, http://www.uq.edu.au/entomology/dipterol/diptconf.html
Jill Marie Mullett and Carie Nixon (Illinois Natural History) have been invited to area junior high schools to work with young women in the GEMS (Girls in Engineeering, Math, & Science) program and introduce them to scientific illustration through the medium of Beatrix Potter. The students then have a chance to try their skills at drawing from natural objects.
Mullett and Nixon have also taught two 6-week short courses,'Beginning to Picture Nature' (2000) and 'Drawing in Colored Pencil' (2001), as part of the Illinois Wilds Institute for Nature. These courses were open to the public and Kampmeier overcame her fear of putting pencil to paper in the first course.
Year 2: Although many of the outreach activities are conducted on a volunteer basis during non-working hours, those involved feel very committed to the importance of sharing knowledge, expertise, and an appreciation for science and art with members of the community, particularly the youth.
Gail Kampmeier served on the Allerton Biodiversity Blitz organizing committee; organized database component, modifying Mandala for use in the Biodiversity Blitz at Allerton Park, June 29-30; she trained and supervised 12 volunteers and participated in data entry, management, & reporting.
In April 2002 J. Marie Mullett traveled back to her hometown to give two presentations on Digital Illustration Techniques at John Glenn High School and Urey Jr. High. She was asked to present her work as a result of having received the Connie Verkler Scholarship in Art when a senior at John Glenn High School. She prepared a presentation for upper level art students and science students at the high school and 300 students at the jr. high teaching digital illustration in the sciences. Junior high students at Urey Middle School were especially interested in understanding how microscopes they have available in their classes can aid in biological illustration.
Gail Kampmeier is on listserves for TAXACOM, TDWG (Taxonomic Databases Working Group), and UIUC-WebMasters and distributes pertinent information to others at INHS, UIUC, and in our PEET project. https://www.fastlane.nsf.gov/cgi-bin/NSF_PrjRpt?@@@___0212___ps1v4APbehh%3Aw%3AKNBQ85Q2Q%3At%3APAqbu%3AyMm9hem3uF%3A5N2KH55 ... Page 11 of 25 Preview before Submission[NSF Project Report - Version 1.2] 5/30/03 15:25
Gail Kampmeier and J. Marie Mullett helped interpret entomological exhibits in the Illinois Natural History Survey’s Mobile Science Unit at the Agricultural, Consumer, and Environmental Sciences (ACES) Open House on the campus of the University of Illinois. (March 2002)
An article was published in the latest issue of the ABRS journal Biologue (No 26, April 2002, pp. 14-15) on the grant entitled 'ABRS, NSF, CSIRO Entomology and the Schlinger Foundation: A partnership for the discovery of Australia's Insect Fauna'
In conjunction with the outreach team at the Illinois Natural History Survey, J. Marie Mullett spoke to grade school students about entomological illustration and introduced students to live insects from the tropics.
J. Marie Mullett has served two years as President of the Illinois Prairie Chapter of the Guild of Natural Science Illustrators, a local chapter that she organized, of this internationally based group. The Prairie Chapter’s main goals are in promoting art and science education and preserving archived biological illustrations. The group holds monthly meetings and sponsors workshops in scientific illustration, including an opportunity to learn techniques of sketching in the field at a local prairie (June 2002).
YEAR 3 Additions: Brian M. Wiegmann volunteered in the “Ask the Experts” exhibit at BugFest 2002. North Carolina Museum of Natural Sciences, August 2002.
Martin Hauser presented insects together with May Berenbaum to children at the Orpheum children’s science center in Champaign (http://www.m- crossroads.org/orpheum/).
Martin Hauser began working as an entomological advisor with the UIUC
BugScope
with school children, answering questions and explaining functions of insect body structures that they see from photos taken with the ESEM (environmental scanning electron microscope).
Gail Kampmeier talked with visitors at ACES Open House (March 2003) to the Illinois Natural History Survey’s Mobile Science unit, which featured
entomological exhibits.
Gail Kampmeier was invited to participate in a dinner at the Florida Avenue Residence Hall with WIMSE (Women in Math, Science, & Engineering) students and share her experiences as a scientist. Female undergraduate students may choose this housing option, which features extra activities and support in the sciences (October 2002)
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Survey as a PBS phone volunteer at WILL-TV's Winterfest fundraiser on 6 December.
Gail Kampmeier was interviewed by phone with Patricia Murphy of Pattycake Productions for a childrens' book on entomologists and their careers. (March 2003)
Gail Kampmeier was invited to participate in the 1st annual Building a
Presence for Science Institute sponsored by the Illinois Science Teachers Association. She dined with K-12 educators from all over the state who had been selected as Key Leaders that were working toward building a 'community of learners' for the improvement of science education. (July 2002)
Gail Kampmeier was a judge for the student poster competition at national Entomological Society of America meeting in Ft. Lauderdale, FL (November 2002)
An exhibit of several of J. Marie Metz’s stiletto fly and acrocerid habitus drawings is on public display at the Verde Gallery in Champaign (May-June 2003).
As one of five founding members of the Prairie Chapter of the Guild of Natural Science Illustrators, J. Marie Metz keeps the Guild’s mission in mind in many facets of her work: furthering education in natural scientific illustration as well as enhancing understanding of the importance of illustration in the sciences. Due to her efforts and those of the group, the chapter recently received a grant of $1,000 for mounting an exhibition at Allerton House in Monticello, IL. The historic house and grounds of Allerton Park were a legacy to the University of Illinois and are widely visited. Plans are being made for a traveling show that can be used by schools to help educate youth not only in natural science illustration, but also to emphasize the importance of
conservation, ecology and species identification within the sciences.
Therevid PEET collaborator Dr. Stephen Gaimari (California Department of Food & Agriculture, Sacramento), funded a visit to California by J. Marie Metz in December 2002 to give a one-week workshop on rendering digital illustrations.
Following interactions at PEET IV in Berkeley, CA in June 2002, where
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J. Marie Metz taught workshops in digital illustration, she was asked by fellow PEETster, Ralph Holzenthal at the University of Minnesota, to conduct a 3-day workshop in November 2002 teaching digital illustration techniques to
entomology students.
As an adjunct Professor at Parkland College, J. Marie Metz is using parts of the Powerpoint presentation designed to teach “PEETsters” illustration, for her community college students. In addition, while working with the Parkland College for Kids Program Marie has passed on information garnered during her therevid PEET experiences about insects, ecology, conservation, and species identification.
Journal Publications: Irwin, M. E., and B. M. Wiegmann, "A review of the southern African genus Tongamya (Diptera: Asiloidea: Mydidae: Megascelinae), with a molecular assessment of the phylogenetic placement of Tongamya and the Megascelinae", African Invertebrates, vol. 42, (2001), p. 225. Published Winterton, S. L., L. Yang, B. M. Wiegmann, and D. K. Yeates, "Phylogenetic revision of the Agapophytinae subf.n. (Diptera: Therevidae) based on molecular and morphological evidence", Systematic Entomology, vol. 26, (2001), p. 173. Published Yang, L., B. M. Wiegmann, D. K. Yeates, and M. E. Irwin, "Higher-level phylogeny of the Therevidae (Diptera: Insecta) based on 28S ribosomal and elongation factor–1alpha gene sequences", Molecular Phylogenetics and Evolution, vol. 15, (2000), p. 440. Published von Tschirnhaus, M. Irwin, M. Hauser, N. Evenhuis, & T. Pape., "Provisional checklist of the Agromyzidae, Therevidae, Mythicomyiidae, Sarcophagidae and Stratiomyidae (Diptera) of the Brandberg Massif, Namibia", Cimbebasia Memoir, vol. 9, (2000), p. 383. Published Wiegmann, B. M., S. C. Tsaur, D. W. Webb, D. K. Yeates, and B. K. Cassel, "Monophyly and relationships of the Tabanomorpha (Diptera: Brachycera) based on 28S ribosomal gene sequences", Annals of the Entomological Society of America, vol. 93, (2000), p. 1031. Published Metz, M. A & M. E. Irwin, "Microtrombidiid mite parasitization frequencies and attachment site preferences on brachyceran Diptera with special reference to Therevidae (Asiloidea) and Tachinidae (Oestroidea)", Environmental Entomology, vol. 30, (2001), p. 903. Published Metz, M. A. & M. E. Irwin, "Revision of Lindneria Krober, with the description of two new genera Insulatitan and Ambradolon, a fossil from Dominican amber (Diptera: Therevidae: Therevinae)", Annals of the Entomological Society of America, vol. 93, (2000), p. 977. Published Hauser, M., "A new species of Adoxomyia Kertész, 1907 (Diptera: Stratiomyidae) from Socotra, Yemen", Fauna of Arabia, vol. 19, (2002), p. 463. Published Hauser, M. & M. Niehuis, "Waffenfliegen (Diptera: Stratiomyidae) und Xylomyiden (Diptera: Xylomyidae) eines xerothermen Standortes im Mittelrheintal (Rheinland-Pfalz)", Fauna Flora Rheinland-Pfalz, vol. 9, (2001), p. 963. Published Irwin, M.E., "Species composition and seasonal flight periodicity of stiletto flies (Diptera: Therevidae) occurring along the Kuiseb River, Gobabeb, Namibia", Cimbebasia, vol. 17, (2001), p. 169. Published Metz, M.A. and F. C. Thompson, "A revision of the larger species of Toxomerus (Diptera: Syrphidae) https://www.fastlane.nsf.gov/cgi-bin/NSF_PrjRpt?@@@___0212___ps1v4APbehh%3Aw%3AKNBQ85Q2Q%3At%3APAqbu%3AyMm9hem3uF%3A5N2KH55 ... Page 14 of 25 Preview before Submission[NSF Project Report - Version 1.2] 5/30/03 15:25
with description of a new species", Studia Dipterologica, vol. 8, (2001), p. 225. Published Yeates, D.K., "Relationships of extant lower brachycera (Diptera): a quantitative synthesis of morphological characters", Zoologica Scripta, vol. 31, (2002), p. 105. Published Winterton, S. L., and M.E. Irwin, "Phylogenetic revision of Agapophytus Guerin (Diptera: Therevidae: Agapophytinae)", Invertebrate Taxonomy, vol. 15, (2001), p. 467. Published Petersen, J.F.T. & M. Hauser, "Family Stratiomyidae. In Petersen, J.F.T. & R. Meier (eds.). A preliminary checklist of Danish Diptera", Steenstrupia, vol. 26, (2001), p. 176. Published Lambkin, C.L., Lee, M.S.Y., Winterton, S.L., and Yeates, D.K., "Partitioned Bremer Support and multiple trees", Cladistics, vol. 18, (2002), p. 436. Published Webb, D.W. and M.A. Metz, "The Nearctic species of Pandivirilia Irwin and Lyneborg (Diptera: Therevidae: Therevinae)", Annals of Ent. Soc. America, vol. , (2003), p. . Accepted Gaimari, S.D., and Raspi, A., "The species of Leucopis, subgenus Leucopella Malloch (Diptera: Chamaemyiidae) from northeastern Africa and Yemen.", African Entomology, vol. 10, (2002), p. 241. Published Holston, K.C. and M. Niehuis, "Stiletto flies (Diptera: Therevidae) from a xerothermic locality in the Middle Rhine Valley (Rhineland-Palatinate)", Fauna und Flora in Rheinland-Pfalz, vol. 9, (2002), p. 1193. Published Metz, M.A., Winterton, S.L. and Irwin, M.E., "Notes on the functional morphology of terminalia from Prorates ballmeri Nagatomi and Liu (Diptera: Scenopinidae: Proratinae) collected while in copula, with a description of the previously unknown female.", Zootaxa, vol. 76, (2002), p. 1. Published Thiele, K. and Yeates, D.K., "Tension arises from duality at the heart of taxonomy", Nature, vol. 419, (2002), p. 337. Published Yeates, D.K., "Relationships of the lower Brachycera: A quantitative synthesis of morphological characters", Zoologica Scripta, vol. 31, (2002), p. 105. Published Yeates, D.K., Bouchard, P. and Monteith, G.B., "Patterns and levels of endemism in the Australian Wet Tropics rainforest: Evidence from flightless insects", Invertebrate Systematics, vol. 16, (2002), p. 605. Published Yeates, D.K., Merritt, D.J. and Baker, C.H., "The adult ventral nerve cord as a phylogenetic character in Diptera", Organisms, Diversity and Evolution, vol. 2, (2002), p. 89. Published Anderson, M.S., and Gaimari, S.D., "Raman-Atomic Force Microscopy of the ommatidial surfaces of dipteran compound eyes", Journal of Structural Biology, vol. , (2003), p. . Accepted Holston, K.C., "The systematic database of Thereva Latrielle names: an answer to the nomenclatural challenge in Therevidae (Insecta: Diptera)", Memoirs of the American Entomological Society, vol. 44, (2003), p. . Accepted Holston, K.C., M.E. Irwin, and F.C. Thompson, "Case No. 3251: Thereva Latreille, [1797] and Phasia Latreille, 1804 (Insecta, Diptera): proposed conservation by designation of Musca plebeja Linnaeus, 1758, as the type species of Thereva", Bulletin of Zoological Nomenclature, vol. 60, (2003), p. . Accepted Lambkin, C.L. and Yeates, D.K., "Genes, morphology, and agreement: Congruence in Australian anthracine bee flies (Diptera: Bombyliidae: Anthracinae)", Invertebrate Systematics, vol. , (), p. . Accepted Metz, M.A., Webb, D.W. and Irwin, M.E., "A review of the genus Psilocephala Zetterstedt (Diptera: Therevidae) with the description of four new genera", Studia Dipterologica, vol. , (2003), p. . Accepted Papoucheva, E., Proviz, V, Lambkin, C.L., Goddeeris, B. and Blinov, A., "Phylogeny of the endemic baikalian Sergentia (Chironomidae, Diptera)", Molecular Phylogenetics and Evolution, vol. , (), p. . Accepted Webb, D.W., "The genera Cliorismia, Psilocephala, and Dichoglena (Diptera: Therevidae) in the Nearctic Region", Journal of Kansas Entomological Society, vol. , (2003), p. . Accepted Webb, D.W., and Metz, M.A, "The Nearctic species of Pandivirilia Irwin and Lyneborg (Diptera:
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Therevidae: Therevinae)", Annals of the Entomological Society of America, vol. , (2003), p. . Accepted Wiegmann, B. M., D. K. Yeates, J. L. Thorne and H. Kishino, "Time flies: A new molecular time scale for brachyceran fly evolution without a clock", Systematic Biology, vol. , (2003), p. . Accepted Yeates, D.K., Irwin, M.E., B.M. Wiegmann, "Ocoidae, a new family of asiloid flies (Diptera: Brachycera: Asiloidea), based on Ocoa chilensis gen. and sp. nov. from Chile, South America", Systematic Entomology, vol. , (2003), p. . Accepted Hauser, M. and M.E. Irwin, "Revision of the genus Ammonaios Irwin and Lyneborg 1981 (Diptera: Therevidae)", Annals of the Entomological Society of America, vol. , (), p. . Submitted Lambkin, C.L., and Yeates, D.K., "PBS reveals significant conflict", Systematic Biology, vol. , (), p. . Submitted Metz, M.A. and Webb, D.W., "Distostylus gen. nov., a monotypic therevine genus (Diptera: Asiloidae: Therevidae) from the Caribbean island of Dominica", Zootaxa, vol. , (), p. . Submitted Webb, D.W., and Metz, M.A., "The New World genus Protothereva Malloch (Diptera: Therevidae: Therevinae) with the description of two new species", Zootaxa, vol. , (), p. . Submitted
Book(s) of other one-time publications(s): Michael E. Irwin, Evert I. Schlinger, and F. Christian Thompson, "The True Flies of Madagascar (Insecta: Diptera)" , bibl. University of Chicago Press, (). Chapter in book Accepted of Collection: Steven M. Goodman and Jonathan P. Benstead, "The Natural History of Madagascar" Michael E. Irwin, "Stiletto Flies of Madagascar (Insecta: Diptera: Asiloidea: Therevidae)" , bibl. University of Chicago Press, (2001). Chapter in book Accepted of Collection: Edited by Steven M. Goodman and Jonathan P. Benstead, "The Natural History of Madagascar" Winterton, S.L., "Scales and setae" , bibl. Academic Press, (2003). chapter in book Published of Collection: R.T. Carde & V.H. Resh, "Encyclopedia of Insects" Lyneborg, L., "The Australian stiletto-flies of the Anabarhynchus genus-group (Diptera: Therevidae)" , bibl. Apollo Books, Stenstrup, Denmark. Illustrations by Jill Mullett., (2001). Book Published of Collection: , "Entomonograph Volume 13" Hauser, M., "Revision of the genus Ammonaios Irwin and Lyneborg 1981 (Diptera: Therevidae)" , bibl. M.S. Thesis, University of Illinois at Urbana-Champaign, (2002). Thesis Published Hill, H.N., "Investigation of the Phylogenetic Utility of Two Nuclear Genes, Opsin and CAD, within the Stiletto Flies (Diptera: Therevidae)" , bibl. M.S. Thesis, North Carolina State University, (2003). Thesis Published Holston, K.C., "A Phylogenetic Assessment of Thereva Latreille (Insecta: Diptera: Therevidae)" , bibl. Ph.D. Thesis, University of Illinois at Urbana-Champaign, (2003). Thesis Published Lambkin, C.L., Power, N., and Starick, N.T., "Mt Kosciuszko: Biodiversity of Flies (Diptera)" , bibl. pp. 76-87, AIAS Jindabyne, (2002). book chapter Published of Collection: K. Green, "Biodiversity in the Snowy Mountains" Fisher, E.M., and Gaimari, S.D., "Simplified digital photography for museum studies" , bibl. http:// www.cdfa.ca.gov/phpps/ ppd/Entomology/Diptera/ digphot.htm. (also, Page 68, Abstracts Volume, Fifth International Congress of Dipterology, Brisbane, Australia, 29, (2002). abstract Published Gaimari, S.D., "Aiming for a phylogenetic classification of the Lauxanioidea" , bibl. Pages 76-77, Abstracts Volume, Fifth International Congress of Dipterology, Brisbane, Australia, 29 September-4 October 2002., (2002). abstract Published Hamilton, J., Peakall, R., and Yeates, D., "Sex, flies and videotape: pollination and sexual deception in the Australian terrestrial orchid Pterostylis" , bibl. Page 96, Abstracts Volume, Fifth International Congress of Dipterology, Brisbane, Australia, 29 September-4 October 2002., (2002). abstract Published Hauser, M., "Comparison of pupal morphology of Asiloidea with special respect of pupal sensory organs" , bibl. Page 99, Abstracts Volume, Fifth International Congress of Dipterology, Brisbane, https://www.fastlane.nsf.gov/cgi-bin/NSF_PrjRpt?@@@___0212___ps1v4APbehh%3Aw%3AKNBQ85Q2Q%3At%3APAqbu%3AyMm9hem3uF%3A5N2KH55 ... Page 16 of 25 Preview before Submission[NSF Project Report - Version 1.2] 5/30/03 15:25
Australia, 29 September-4 October 2002., (2002). abstract Published Hauser, M., "New molecular and morphological evidence for the phylogenetic placement of Phycini and Xestomyzini (Diptera, Therevidae)" , bibl. Page 98, Abstracts Volume, Fifth International Congress of Dipterology, Brisbane, Australia, 29 September-4 October 2002, (2002). abstract Published Holston, K.C., and Irwin, M.E., "From polyphyly to biogeography of Thereva (Asiloidea: Therevidae)" , bibl. Page 104, Abstracts Volume, Fifth International Congress of Dipterology, Brisbane, Australia, 29 September-4 October 2002., (2002). abstract Published Irwin, M.E., "Embracing specimen-level databases: expectations and anxieties!" , bibl. Page 106, Abstracts Volume, Fifth International Congress of Dipterology, Brisbane, Australia, 29 September-4 October 2002, (2002). abstract Published Irwin, M.E., " Introduction to the Symposium: Modern innovations" , bibl. Page 105, Abstracts Volume, Fifth International Congress of Dipterology, Brisbane, Australia, 29 September-4 October 2002, (2002). abstract Published Kampmeier, G.E., and Holston, K.C., "Meeting the interrelated challenges of specimen, nomenclature, and literature tracking with Mandala" , bibl. Page 110, Abstracts Volume, Fifth International Congress of Dipterology, Brisbane, Australia, 29 September-4 October 2002, (2002). abstract Published Lambkin, C.L. and Yeates, D.K., "Modern innovations in systematic inquiry" , bibl. Page 131, Abstracts Volume, Fifth International Congress of Dipterology, Brisbane, Australia, 29 September-4 October 2002, (2002). abstract Published Lambkin, C.L. and Yeates, D.K., "Significant incongruence can be informative: Insights from partitioned support analyses" , bibl. Page 129, Abstracts Volume, Fifth International Congress of Dipterology, Brisbane, Australia, 29 September-4 October 2002, (2002). abstract Published Lambkin, C.L. and Yeates, D.K., "The Exoprosopini, Australia and the world: Revelations from systematic studies" , bibl. Page 130, Abstracts Volume, Fifth International Congress of Dipterology, Brisbane, Australia, 29 September-4 October 2002, (2002). abstract Published Lawson, A.E., Yeates, D.K., McGuire, D.J., Clarke, A.R., and Drew, R.A.I., "DORSALIS—an interactive key to fruitflies of the Bactrocera (Bactrocera) dorsalis complex" , bibl. Page 132-133, Abstracts Volume, Fifth International Congress of Dipterology, Brisbane, Australia, 29 September-4 October 2002., (2002). abstract Published Meier, R. and Wiegmann, B.M., "The struggle over reconstructing the phylogenetic relationships of the Sciomyzoidea (Diptera)" , bibl. Page 153, Abstracts Volume, Fifth International Congress of Dipterology, Brisbane, Australia, 29 September-4 October 2002, (2002). abstract Published Moulton, J.K. and Wiegmann, B.M., "Mining the dipteran genome: tools and resources from GenBank/ EMBL and NCBI" , bibl. Page 162, Abstracts Volume, Fifth International Congress of Dipterology, Brisbane, Australia, 29 September-4 October 2002, (2002). abstract Published Moulton, J.K. and Wiegmann, B.M., "Multiple data sets in addressing complex evolutionary questions within Diptera: case studies within Empidoidea and Simuliidae" , bibl. Page 163, Abstracts Volume, Fifth International Congress of Dipterology, Brisbane, Australia, 29 September-4 October 2002, (2002). abstract Published Savage, J., Wheeler, T.A., and Wiegmann, B.M., "Phylogeny of the genus Thricops Rondani (Diptera: Muscidae) based on molecular and morphological evidence" , bibl. Page 210, Abstracts Volume, Fifth International Congress of Dipterology, Brisbane, Australia, 29 September-4 October 2002, (2002). abstract Published Schlinger, E., "Interpreting spatio-temporal patterns among select clades of Diptera" , bibl. Page 211, Abstracts Volume, Fifth International Congress of Dipterology, Brisbane, Australia, 29 September-4 October 2002, (2002). abstract Published Skevington, J.H., Cumming, J.M., Sinclair, B.J., Wiegmann, B.M., and Moulton, J.K., "Molecular and morphological evidence supporting relationships within Eremoneura: focus on Cyclorrhapha" , bibl. Page 221, Abstracts Volume, Fifth International Congress of Dipterology, Brisbane, Australia, 29 September-4
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October 2002, (2002). abstract Published Wiegmann, B.M., "Molecular biology: a set of new characters for phylogenetic assessment" , bibl. Page 263, Abstracts Volume, Fifth International Congress of Dipterology, Brisbane, Australia, 29 September-4 October 2002, (2002). abstract Published Wiegmann, B.M., Yeates, D.K., Moulton, J.K., and Thorne, J.L., "Genes, fossils and morphology: combined data analysis and a revised time scale for higher- level phylogeny of the Brachycera" , bibl. Page 264, Abstracts Volume, Fifth International Congress of Dipterology, Brisbane, Australia, 29 September-4 October 2002, (2002). abstract Published Winterton, S.L. and Hill, H., "Phylogenetic position of Paracraspisa, a new genus of stiletto-fly from Australia (Diptera: Therevidae: Agapophytinae)" , bibl. Page 266, Abstracts Volume, Fifth International Congress of Dipterology, Brisbane, Australia, 29 September-4 October 2002, (2002). abstract Published Winterton, S.L., Wiegmann, B.M., and Schlinger, E.I., "Phylogeny of the small headed- flies (Diptera: Acroceridae) using multiple molecular markers" , bibl. Page 265, Abstracts Volume, Fifth International Congress of Dipterology, Brisbane, Australia, 29 September-4 October 2002, (2002). abstract Published Yeates, D.K., Bickel, D.K., and McAlpine, D.K., "An estimate of dipteran species- richness in Australia" , bibl. Page 271, Abstracts Volume, Fifth International Congress of Dipterology, Brisbane, Australia, 29 September-4 October 2002, (2002). abstract Published Yeates, D.K., Bickel, D.K., McAlpine, D.K., Schneider, M., Cranston, P., and Marshall, S., "An interactive key to fly families: from Australia to the world!" , bibl. Page 272-273, Abstracts Volume, Fifth International Congress of Dipterology, Brisbane, Australia, 29 September-4 October 2002, (2002). abstract Published Yeates, D. K., M. E. Irwin, and B. M. Wiegmann, "A new family-level taxon of asiloid flies (Diptera: Brachycera: Asiloidea), based on a new genus and species from Chile, South America" , bibl. Page 273- 274, Abstracts Volume, Fifth International Congress of Dipterology, Brisbane, Australia, 29 September-4 October 2002, (2002). abstract Published Yeates, D.K., Merritt, D.J., and Baker, C.H., "Neuromere fusion: a useful phylogenetic marker in brachyceran Diptera" , bibl. Page 275, Abstracts Volume, Fifth International Congress of Dipterology, Brisbane, Australia, 29 September-4 October 2002, (2002). abstract Published
Other Specific Products:
Data or databases
Information from >100,000 therevid specimens found in nearly 15,000 collecting localities has been entered into the therevid Mandala database. The taxonomic names datafile contains over 4700 entries and the histories of all of the therevid names are being catalogued. Nearly 8,500 records have been created that dissect and index the nearly 800 entries of therevid literature. Over 6500 people are listed as collectors, illustrators, authors of taxonomic names, lenders of material from collections or museums, authors of literature, and determiners of taxonomic names attached to specimens, and copyright holders of illustrations. Many apparently duplicate names exist due to misspellings and different name formats given under various circumstances and a system was established to designate junior synonyms and senior synonyms where appropriate.
Data can be entered and shared over the internet (TCP/IP connection) from the live Mandala database on a server to those with Filemaker Pro 5 and who are cleared by password. Otherwise, data may be searched and browsed by anyone from https://www.fastlane.nsf.gov/cgi-bin/NSF_PrjRpt?@@@___0212___ps1v4APbehh%3Aw%3AKNBQ85Q2Q%3At%3APAqbu%3AyMm9hem3uF%3A5N2KH55 ... Page 18 of 25 Preview before Submission[NSF Project Report - Version 1.2] 5/30/03 15:25
the WebMandala website http://pherocera.inhs.uiuc.edu/index.htm
Software (or netware)
Mandala is a database system for systematics and biodiversity research. It catalogs specimens, batches of specimens (in v5), nomenclatural history, tracks loans by specimen and by batch, and tracks and dissects taxonomic literature. It has context sensitive help at the field and file level, it tracks user questions and their answers, and tracks changes to the database structure over time. It was originally developed under the first NSF PEET project 9521925, and the first year of this current project has seen two substantial upgrade releases, Mandala 4.0 in November 2000, and Mandala 5.0 in April 2001. Mandala 4.0 was a major upgrade using FileMaker 5.0 and also represented a major facelift to the user interface. It also provided a time for major housekeeping of fields, layouts, scripts, that were no longer in use. Mandala 5 became a more viable tool for biodiversity studies that may need to deal with batches of specimens, and in so doing also became a more valuable tool to collections curators who must track loans. Increased flexibility in the format of unique specimen numbers, as well as the ability to record and track batches of specimens, has improved Mandala as a tool in managing loans of specimens and data from biodiversity studies. Loan management layouts aid in the tracking and details of shipping of loan requests and receipts. Literature records may be linked to illustration records (e.g., taxa,
habitats, and distribution maps), records of specimens examined, and taxonomic names records so the relevance of literature citations answering a database query may be evaluated. Detailed referencing of digital illustration archives has also been recently added to Mandala. Georeferenced data associated with specimens may be exported to generate distribution maps. Many specialized layouts and scripts within each datafile simplify data entry, searches, exports, and printing. Customized calculated fields facilitate export of concatenated data. Several auxiliary datafiles, such as the museum collections, journals, people (as collectors, authors, etc), and biogeographic regions datafiles, organize additional sets of data related to each of the three main datafiles, thereby improving the sophistication and accuracy of queries and reducing data entry errors.
Mandala is free for the asking and can be modified for use with many different types of organisms. Contact Gail Kampmeier at [email protected] for more information or see the WebMandala site at http://pherocera.inhs.uiuc.edu/index.htm
Journal cover
Winterton, S. L. 2001. Journal Cover Illustration- Systematic Entomology: 2001, vol. 26 (issues 1-4). Taenogerella platina Winterton & Irwin, (Therevidae) male habitus, black & white.
Illustration shared with all subscribers to the journal Systematic Entomology in 2001.
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Internet Dissemination:
http://www.inhs.uiuc.edu/cee/therevid/
This site chronicles the activities of the therevid PEET team, including reports, meetings, outreach, expeditions, interactive keys to newly revised genera, team member profiles, databasing efforts and WebMandala, and background information for users of all ages. It was begun in 1996 with DEB 95-21925 and continues with the current award.
Contributions:
Contributions within Discipline:
Our phylogenetic research provides an historical context on which hypotheses can be tested concerning the evolution and diversification of flies and other insects. Our characterization and analysis of gene sequences is critical to many research programs in molecular systematics, entomology, and evolutionary biology that will apply these genes in tests of phylogenetic hypotheses. Development of methods of estimating divergence times from nucleotide sequence data can be used to infer ages for many groups of organisms.
Mandala. Systematics research on stiletto flies (Diptera: Therevidae) has been facilitated by the cross-platform database, Mandala
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Contributions to Education and Human Resources:
A major contribution of our project is training the next generation of systematists. In the first year of our project we added to the systematics and dipterological communities the through the graduation of a student with his doctoral degree (LL Yang), employment of two postdocs trained in the PEET project (S. L. Winterton at NCSU and Christine Lambkin at CSIRO) and employment of a new masters student (Hilary Hill). In the summer of 2001, Steve Gaimari became an Associate Insect Biosystematist (Dipterist) for the California State Collection of Arthropods, California Department of Food and Agriculture, Sacramento. In year 3, Mark Metz accepted a postdoctoral position in the fall of 2002 at the Frost Entomological Museum at the Pennsylvania State University. After finishing her MSc. at North Carolina State University in March 2003, Hilary Hill is currently a Research Associate with USDA/APHIS/ PPQ in Raleigh NC. Kevin Holston was awarded a 2-year NSF postdoctoral fellowship to work on European therevids starting June 2003 at the Swedish Museum of Natural History under the mentorship of Dr. Thomas Pape. J.
Marie Metz has been teaching digital illustration and design both in workshops to other PEET groups and at a local college.
This project has also seen the rapid advancement in computerized rendering of illustrations by scientific illustrator, Jill Marie Mullett. Having only begun this technique of rendering in January 2000, Ms. Mullett's work is admired among her peers in the Guild of Natural Science Illustrators, and among those most highly respected in the field. She obtained her Masters in Art Education in May 2001, started the Prairie Chapter of the Guild of Natural Science Illustrators in East Central Illinois, and is mentoring an intern in scientific illustration, Ann C. Rast, a faculty
member in the Department of Fine and Applied Arts at the University of Illinois at Urbana-Champaign.
The therevid PEET project has also provided opportunities for undergraduate students to learn about databases while working with data entry in Mandala (http://www.inhs.uiuc.edu/cee/therevid/dbtop.html). We nominated our top student, Amanda Buck, for the campus-wide award of Student Employee of the Year, and she won one of six of these awards in 2001.
In year 2 of this grant:
Over 20 students and scientists from all over Australia attended a weekend illustration workshop in Melbourne run by Chris Lambkin in collaboration with Ken Walker the invertebrate curator from the Victorian Museum. The majority of attendees were able to achieve a complete scientific illustration of a specimen of interest in the time available.
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In year 3 of this grant:
Members of the therevid PEET team have volunteered at local open houses, bugfests, a childrens museum, and the UIUC BugScope, spreading knowledge and enthusiasm about science, nature, and entomology to school aged children and the general public. Further mentoring opportunities with the WIMSE (Women in Math, Science, and Engineering) group on the UIUC
campus, opportunities to interact with science teachers, all provide avenues for sharing experiences and enthusiasm for the sciences.
J. Marie Metz has been teaching digital illustration and design both in workshops to other PEET groups and at Parkland College. As an adjunct
Professor at Parkland College, she is using parts of the Powerpoint presentation designed to teach “PEETsters” illustration for her community college students. In addition, while working with the Parkland College for Kids Program Marie has passed on information garnered during her therevid PEET experiences about insects, ecology, conservation, and species identification. She has also had the opportunity with contacts made at PEET IV to collaborate with other scientific illustrators and to use techniques developed in that collaboration in the teaching methods presented at the illustration workshops in Minnesota and California.
Contributions to Resources for Science and Technology:
Through the efforts of Jill Marie Mullett and Carie Nixon (Illinois Natural History), young women from three local junior high schools in the GEMS (Girls in Engineeering, Math, & Science) are being introduced to scientific illustration. By teaching illustration through the life of Beatrix Potter, students are motivated to learn how to draw. Training children in aspects of scientific illustration teaches them to hone their observational skills and translate their observations to a rich visual medium.
The database system, Mandala, was used in the Biodiversity Blitz at Allerton Park at the end of June 2001 (http://www.conted.uiuc.edu/rapcc/bioblitz/). Volunteers were trained by Gail Kampmeier to input data from this 24-h exercise to identify as many species as possible within the confines of the park. The data were reported during the Blitz on a special website, and will later be searchable by subsequent visitors to the park.
Chris Lambkin will contribute training at the PEET IV meeting in 2002 by coordinating a workshop on ‘Character Incongruence between Data Partitions’ and contributing to the workshop on Morphological Character Analysis. J. Marie Mullett will be https://www.fastlane.nsf.gov/cgi-bin/NSF_PrjRpt?@@@___0212___ps1v4APbehh%3Aw%3AKNBQ85Q2Q%3At%3APAqbu%3AyMm9hem3uF%3A5N2KH55 ... Page 22 of 25 Preview before Submission[NSF Project Report - Version 1.2] 5/30/03 15:25
coordinating and presenting the workshop on Digital Rendering of Scientific Illustrations, which will be repeated 4 times during the conference and includes hands-on training for participants. Gail Kampmeier will be presenting information on ITIS (Integrated Taxonomic Information System) in the workshop on the Tree of Life and PEET that is being organized by David Yeates. This workshop will include presentations on the All Species Foundation
Chris Lambkin is organizing a all day symposium ‘Incongruence, Data Partitions, and Phylogenetic Signal’ for the Sixth International Congress for Systematic and Evolutionary Biology VI in Patras, Greece on the 9-16 of September, (http://www.icseb-vi.biology.upatras.gr)
J. Marie Mullett has mentored the three graduate students in the Irwin lab on digital illustration techniques that they can use to create their own drawings for publication. These drawings have started to appear in their publications. Ms. Mullett was also invited to return to her hometown to give presentations on Digital Illustration Techniques to John Glenn High School and Urey Jr. High.
David Yeates has developed and delivered a final year undergraduate course on Entomology (code BIOL3115) in collaboration with colleagues in CSIRO Entomology and the Australian National University (http://www.anu.edu.au/BoZo/undergrad/units2002.html).
David Yeates is Chair of the organizing committee of the 5th International Congress of Dipterology (Brisbane, Sept 29-Oct 4, 2002) http://www.uq.edu.au/entomology/dipterol/diptconf.html.
Building scientific infrastructure and collaborations in Madagascar. An offshoot of the therevid PEET project is the collaboration with MICET and the California Academy of Sciences in Madagascar, and supported by the Schlinger Foundation of a 3-year inventory of the higher Diptera (Brachychera) of Madagascar. Rasolondalao Harin'Hala Hasinjaka, 'Rin'ha', is checking 30 Malaise traps, sorting the contents, and sending them on to UIUC.
Developing an outreach program for Madagascar's National Park system has involved considerable effort on the part of Irwin and Schlinger in 2001-2. Four activities were initiated through expeditions to that biologically diverse island continent during the 2001 year. The first is to utilized our Therevid PEET informatics databasing activities to help develop the databasing prospects for Ranomafana National Park. To this end, we have worked with the Patricia Wright, Director of Institute for the Conservation for Tropical Environments, to have the park's research director, Razafindratsita Tiana, join our laboratory for about two months during the summer of 2002 to gain first hand knowledge of our Mandala database and how to use and modify it to suit the purposes of the park. The second is to help enhance visitor awareness of the natural history in selected Madagascar national parks through interpretive center displays and activities. To this end, and working with Parker and through Rin'ha, Irwin and Schlinger are helping to make known for the https://www.fastlane.nsf.gov/cgi-bin/NSF_PrjRpt?@@@___0212___ps1v4APbehh%3Aw%3AKNBQ85Q2Q%3At%3APAqbu%3AyMm9hem3uF%3A5N2KH55 ... Page 23 of 25 Preview before Submission[NSF Project Report - Version 1.2] 5/30/03 15:25
eco-tourists some of the fascinating but smaller wildlife (i.e., arthropods) in the parks. We have begun to put together our thoughts on a butterfly house and on some static displays. The third is to help develop an entomological research laboratory at the Centre ValBio, partly funded through NSF and currently being constructed adjacent to the Ranomafana National Park. Irwin has been the consultant to the development of the entomology lab and has been working with architect Peter Ozolins of Blacksburg, VA. A fourth endeavor is to help develop bioprospecting activities so that the local population living in the buffer areas around Ranomafana National Park can make a living utilizing the resources of the park in a sustainable way. Irwin has located a bioprospecting firm from France, Entomed, that is extremely interested in a collaborative arrangement. The head of research of that firm visited the University of Illinois during April 2002 to discuss the possibilities and arrangements. These will have to be negotiated with ICTE and ANGAP, but, if successful, should go a long ways towards stabilizing the economy of the local population in the buffer zone around the park.
Irwin was elected to the Board of Directors of Discover Life in America and has participated in the evaluation of over 40 proposals to discover the biodiversity of the Great Smoky Mountains National Park. He is also a member of both the Science and
Informatics Committees.
Irwin was a member of a US panel evaluating crop protection proposals submitted to the Binational Agriculture and Research Development (BARD) program, which links US and Israeli agricultural research initiatives.
Irwin and Yeates were appointed to the All Species Foundation advisory committee. This committee establishes the operational procedures for making all life on earth known within the next generation.
Irwin is a member of the Illinois Natural History Survey (INHS) Collections Committee; Kampmeier is a member of the INHS Database Team and chair of the INHS WebTeam.
Chris Lambkin was elected to the Council for the International Organization for Systematic and Evolutionary Biology (IOSEB) for the term 2002-2008, and the Council for Society for Australasian Systematic Biologists (SASB) for the term 2003-2005.
J. Marie Metz won first prize for her Henicomyia sp. habitus illustration at the 5th International Congress of Dipterology held in Brisbane Australia (Oct. 2002).
The Guild of Natural Science Illustrators selected both of J. Marie Metz’s entries for their 2003 GNSI Annual juried exhibition for their https://www.fastlane.nsf.gov/cgi-bin/NSF_PrjRpt?@@@___0212___ps1v4APbehh%3Aw%3AKNBQ85Q2Q%3At%3APAqbu%3AyMm9hem3uF%3A5N2KH55 ... Page 24 of 25 Preview before Submission[NSF Project Report - Version 1.2] 5/30/03 15:25
international meeting in July 2003 in Denver. Selected were H. Fly Habitus and the study, Tongamya Head. Both works were rendered digitally using Adobe Illustrator and Photoshop.
Michael E. Irwin was appointed to the Advisory Board (March 2003- ) of the Archbold Tropical Research and Education Consortium (ATREC), at Clemson University's Springfield Biological Research Station, Dominica.
Stephen Gaimari was elected a Fellow of the Linnean Society of London
(Feb. 2002)
David Yeates was elected chair of the Council of the International Congresses of Dipterology, October 2002.
Special Requirements for Annual Project Report:
Unobligated funds: less than 20 percent of current funds
Categories for which nothing is reported: Contributions to Other Disciplines Contributions Beyond Science and Engineering Special Reporting Requirements Animal, Human Subjects, Biohazards
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We welcome comments on this system
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Project Participants
Irwin E Michael : Principal Investigator Has worked for more than 160 hours : Yes Contribution to project : Michael E. Irwin is professor in the Dept. of Natural Resources & Environmental Sciences at the University of Illinois at Urbana-Champaign, with the title, Schlinger Research Scholar in Arthropod Biodiversity and Systematics; he is also affiliated with the Center for Economic Entomology at the Illinois Natural History Survey, and the Dept. of Entomology at UIUC.
Yeates K David : CoPrincipal Investigator Has worked for more than 160 hours : Yes Contribution to project : David Yeates has just relocated to the CSIRO Entomology Section in Canberra where he has taken a position as Principal Research Scientist and Head of the Australian National Insect Collection (ANIC), the world's largest collection of Australian insects (See ). David will also continue his systematics research in Diptera at ANIC as part of the Natural Resources and Biodiversity Program.
Wiegmann M Brian : CoPrincipal Investigator Has worked for more than 160 hours : Yes Contribution to project : Brian Wiegmann is an Associate Professor in the Dept. of Entomology at North Carolina State University. His lab coordinates the molecular research on the relationships of the family Therevidae and their closest relatives.
Webb Donald : Senior personnel Has worked for more than 160 hours : Yes Contribution to project : Participates in collecting trips; writes monographs; mentors graduate students, particularly those at UIUC. Funded by the Illinois Natural History Survey.
Kampmeier E Gail : Senior personnel Has worked for more than 160 hours : Yes Contribution to project : Designs and maintains database system, Mandala; designs and maintains the therevid website http://www.inhs.uiuc.edu/cee/therevid/, including WebMandala http:// pherocera.inhs.uiuc.edu/index.htm; mentors undergrad and graduate students; drafts reports and pertinent manuscripts. Funded through the Illinois Natural History Survey, this PEET grant, and the Ag Experiment Station at the UIUC.
Gaimari Steve : Senior personnel Has worked for more than 160 hours : No Contribution to project : Steve Gaimari obtained his Ph.D. under the first therevid PEET project in 1998; after teaching a graduate course in systematics in the Dept. of Entomology at the University of Illinois at Urbana-Champaign, he accepted a postdoc at the Smithsonian Institution from 1999-2001; starting in July, 2001, he will be an Associate Insect Biosystematist, California State Collection of Arthropods, CDFA, Sacramento. Gaimari retains a strong interest in the family Therevidae and continues to publish and collaborate with the therevid PEET team.
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Metz A Mark : Post-doc Has worked for more than 160 hours : Yes Contribution to project : Ph.D. candidate in the Dept. of Natural Resources & Environmental Sciences at the University of Illinois at Urbana-Champaign, working on revising the higher therevine genera. His assistantship is supported through Schlinger Foundation funds. Mark graduated with his Ph.D. in May 2002 and began a 6-month post doc in the Irwin lab working on the homologies in the male and female genitalia of the therevoid families of Asiloidea, the Therevidae of New Caledonia, and the functional morphology of the genitalia of Prorates ballmeri Nagatomi and Liu (see Metz et al. 2002 in publications list). In October 2002 he accepted a post doc at the Pennsylvania State UniversityÆs Frost Entomological Museum. He continues to work with the therevid PEET project, primarily collaborating with Don Webb on papers and research. He was a co-PI along with other therevid PEET graduates in the ambitious proposal to the NSF Planetary Biodiversity Inventory, ôCrossing the Finish-Line: A World Monograph of the Therevidae (Insecta: Diptera).
Winterton L Shaun : Post-doc Has worked for more than 160 hours : No Contribution to project : Shaun Winterton is a Postdoctoral Research Fellow, Department of Entomology, North Carolina State University. Dr. Winterton began postdoctoral work in the Wiegmann lab on the molecular phylogenetics of the orthorrhaphous brachyceran family Acroceridae. He is funded through a postdoctoral research fellowship to NCSU from the Schlinger Foundation. Earning his Ph.D. on the first therevid PEET grant, Dr. Winterton remains active in therevid systematics and is a collaborator/consultant to the current project.
Lambkin Christine : Post-doc Has worked for more than 160 hours : Yes Contribution to project : Christine Lambkin joined the therevid PEET team in May 2001 as a Postdoctoral Fellow at CSIRO Entolomology Section, Canberra, Australia, to continue the classification of Australian stiletto flies begun by Shaun Winterton. Christine has recently completed her doctorate at the University of Queensland in Brisbane, which comprised a systematic revision of a tribe of bee flies (Diptera: Bombyliidae: Asiloidea), a close relative of the Therevidae. Chris, who has provided some technical assistance to the therevid team in the past, will continue the work on the phylogenetics the Therevidae, concentrating on the genus Ectinorhynchus and its allies, in arid and semiarid environments of Australia. Part of Chris's time will be spent investigating the use of web- based dissemination of interractive keys, data, and monographic work.
Holston C Kevin : Graduate student Has worked for more than 160 hours : Yes Contribution to project : Ph.D. candidate in the Dept. of Entomology at the University of Illinois at Urbana-Champaign, working on the old and speciose genus, Thereva. Kevin Holston's assistantship is supported by Schlinger Foundation funds. He also obtained travel support from the American- Scandinavian Foundation for two months in Copenhagen, Denmark.
Hauser Martin : Graduate student Has worked for more than 160 hours : Yes Contribution to project : After completing his masters project in the Dept. of Entomology at the University of Illinois on the therevid genus Ammonaios, he will pursue his Ph.D. on the xestomyzine genera. Martin Hauser's assistantship is supported by Schlinger Foundation funds.
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Hill Hilary : Graduate student Has worked for more than 160 hours : Yes Contribution to project : Hilary Hill completed her masters project in the Dept. of Entomology at North Carolina State University in March 2003. Her assistantship was supported as matching funds provided by the North Carolina Agriculture Research Service and NC State College of Agriculture and Life Sciences (CALS). Her work evaluated the phylogenetic utility and information content of nuclear protein coding genes, opsin rh1 and CAD in the Therevidae and among their closest relatives. Hilary is currently a Research Associate with USDA/APHIS/PPQ in Raleigh NC.
Buck Amanda : Undergraduate student Has worked for more than 160 hours : Yes Contribution to project : A senior in actuarial science at the University of Illinois at Urbana- Champaign, she is on the database team and contributed to the generation of distribution maps, set up a way of calculating displacement of latitude/longitude from a point source, and won in 2001 a prestigious campus award as one of 6 Outstanding Student Employee of the Year.
Calvillo Melissa : Undergraduate student Has worked for more than 160 hours : Yes Contribution to project : A junior in psychology at the University of Illinois at Urbana-Champaign, she is on the database team working through categorizing data in the literature.
Montgomery Claire : Undergraduate student Has worked for more than 160 hours : Yes Contribution to project : Claire Montgomery joined the database team in June 2001, working throughout the summer and after school in the fall. She graduates in June 2002 and will be starting at the University of Illinois in the Physics Dept. in the fall. She plans to continue working with the database team in the summer of 2002, and if we're lucky, continue once she becomes an undergraduate. Claire also volunteered her time at the Allerton Biodiversity Blitz in June 2001, taking over the graveyard shift with other members of the database team and allowing her very tired supervisor a few hours of sleep!
Lanford Heather : Undergraduate student Has worked for more than 160 hours : Yes Contribution to project : Heather Lanford joined the database team in December 2001 and has been working with us after school. She graduates in June 2002 and will be starting at the University of Illinois in the Physics Dept. in the fall. She plans to continue working with the database team in the summer of 2002, and if we're lucky, she too will continue once she becomes an undergraduate.
Svistula Dmitri : Undergraduate student Has worked for more than 160 hours : Yes Contribution to project : Dmitri joined the database team in June 2001 and worked with us throughout the summer and during holidays from the University of Illinois at Chicago. He will continue to work with the database team in the summer of 2002. Because of his knowledge of Russian, we have had him translating and transliterating fly specimens with labels written in Cyrillic, in addition to the normal duties of a database team member. He also volunteered his time at the Allerton Biodiversity Blitz in June 2001, taking over the graveyard shift with other members of the database team and allowing his very tired supervisor a few hours of sleep!
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Cassel Brian : Technician, programmer Has worked for more than 160 hours : Yes Contribution to project : Agricultural Research Technician II, Department of Entomology, NCSU, serves as manager for the molecular systematics laboratory of CO-PI Wiegmann. Cassel conducts amplification and sequencing for the project and supervises trainees in the laboratory.
Hamilton Joanna : Technician, programmer Has worked for more than 160 hours : Yes Contribution to project : Technical Officer employed by CSIRO, has been working for David Yeates curating and labeling the Therevidae collected from expeditions and testing a key to Australian genera developed by Chris Lambkin. Joanna has now (2003) left the project, returning to full-time studies and working, part-time, on an interactive key to fly families of Australia
Recsei Jacqueline : Technician, programmer Has worked for more than 160 hours : Yes Contribution to project : Technical Officer employed by CSIRO Entomology since November 2002, has been working for David Yeates sorting, specimen curation, labeling, and databasing the Therevidae collected from expeditions, participating fully in field work, and testing a key to Australian genera developed by Chris Lambkin.
Metz J. Marie : Technician, programmer Has worked for more than 160 hours : Yes Contribution to project : Scientific illustrator (formerly listed as Jill Marie Mullett) on the therevid project funded by the Schlinger Foundation through the Dept. of Natural Resources & Environmental Sciences at the University of Illinois. Also mentors students teaching techniques of scientific illustration.
Marlin Kate : High school student Has worked for more than 160 hours : Yes Contribution to project : Kate Marlin joined the database team in June 2001 and worked throughout the summer with us. She volunteered her time at the Allerton Biodiversity Blitz in June 2001, taking over the graveyard shift with other members of the database team and allowing her very tired supervisor a few hours of sleep!
Algmin Kristin : Academic hourly Has worked for more than 160 hours : Yes Contribution to project : Kristin (Mrozinski) Algmin joined the Irwin lab as a senior in biology at the University of Illinois at Urbana-Champaign. She started out on the database team and moved to curation of insects trapped in malaise samples. She also helps rear therevid larvae. Now that she has graduated, she returned in the fall of 2001 to work as a full-time hourly assistant working on many of the projects she did as an undergrad, but now with the increased responsibility of troubleshooting and checking data quality of the therevid Mandala database, learning to identify therevids to genus and species, and documenting various operations. She also helps train and answer questions of other workers and interacts with grad students.
Thorne Jeffrey : Collaborator Has worked for more than 160 hours : Yes
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Contribution to project : Jeffrey Thorne is an Associate Professor, North Carolina State University, Department of Statistical Genetics. He is developing methods for inferring divergence times from molecular phylogenetic data without the assumption of a molecular clock. These methods are being used to develop a time scale for major events in brachyceran fly evolution based on data generated in the Wiegmann lab for therevids and other Brachycera.
Rast C Ann : Intern Has worked for more than 160 hours : Yes Contribution to project : Ann Coddington Rast is an intern working on scientific illustration with Jill Marie Mullett. She is working on Diptera illustrations for the therevid PEET project primarily for the experience and knowledge she can gain by illustrating fly images. Ann is a professor at the University of Illinois in the Department of Fine and Applied Arts. She is working with our team to produce professional illustrations using traditional and digital renderings of Diptera images.
Reid Beryl : Volunteer Has worked for more than 160 hours : Yes Contribution to project : Beryl Reid, an Australian National Insect Collection Volunteer since November 2001, has spent 2-days per week assisting Chris Lambkin with the curation of the Therevidae.
Metz Bradley : Academic hourly Has worked for more than 160 hours : Yes Contribution to project : Brad is working half time with the Irwin lab, learning to curate flies, enter specimens in the Mandala database, retrospectively obtain geographic coordinates and verify locality information for specimens in the database so that they may be mapped, electronically archive illustrations created by traditional means, and do minor database troubleshooting. He is supported through Schlinger Foundation matching funds. After obtaining his B.S. in Biology in 2002, he was awarded a one-year fellowship to attend graduate school in entomology at Texas A&M starting in fall 2003.
Leslie Erica : Volunteer Has worked for more than 160 hours : Yes Contribution to project : Erica, a new Australian National Insect Collection (ANIC) Volunteer, spent 2-days per week assisting Chris Lambkin with the return and assimilation of Anabarhynchus material into Australian collections following publication of the revision. Through the experience gained, Erica has now obtained a part-time position databasing Australian fruit flies in ANIC into the CSIRO Biolink.
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https://www.fastlane.nsf.gov/cgi-bin/NSF_PrjRpt Page 5 of 5 Year 3 Activities Towards a World Monograph of the Therevidae (Insecta: Diptera) Michael E. Irwin, Brian M. Wiegmann, and David K. Yeates, PI’s NSF PEET DEB99-77958 Sept. 2002-Aug. 2003
Relationships of the Therevidae and their closest relatives: In the Wiegmann lab, gene sequences were obtained for Therevidae from nuclear 28S ribosomal DNA, rh1 opsin, CAD, and mitochondrial 16S ribosomal DNA, and cytochrome oxidase genes. Particular emphasis was placed on sampling taxa from Australia, New Caledonia, Chile and South Africa. Taxa were added to existing datasets for Xestomyzinae and Therevinae. Laboratory and computational work was conducted to build sequence datasets, alignments, and perform phylogenetic analyses. These data were added to existing therevid datasets. Christine Lambkin continues work on the phylogenetics of the genus Ectinorhynchus and its allies in Australasia. Chris is also investigating supertree analyses and is involved in the training and development of scientific knowledge in systematics in Australia, America, and Europe. Martin Hauser is revising the Xestomyzinae from Madagascar. Several new species were found in the malaise trap material collected by Irwin and Schlinger in Madagascar. Xestomyzinae have not been previously known from this island. Hauser will be sorting Malaise trap material from Madagascar, identifying specimens to species, and starting the work on a publication about the Therevidae fauna of this island. He will also be finishing up a revision of the genus Henicomyia and other aspects related to therevid research Hauser received the first known amber (Baltic) fossil of a Xestomyzinae from the Hoffeins collection (Hamburg, Germany) and he is in progress of describing this new species. The discovery of a Xestomyzinae outside of the present distribution (South Africa and the New World) has great implications and the historical biogeography of Therevidae has to be reevaluated. Kevin Holston has worked to develop a framework for biological research on the genus Thereva (1) using molecular data to test a phylogenetic hypothesis that posits a monophyletic Thereva with respect to a diverse sample of genera in Therevinae, (2) using molecular and morphological data to propose a species-level phylogeny for Thereva to identify biogeographical patterns, and (3) using specimens and specimen label data to complete a taxonomic revision of Thereva for the Nearctic Region. Publications from Holston’s dissertational work are scheduled for 2003 and 2004. After the completion of his Ph.D. thesis in May 2003 on the phylogeny and taxonomy of Thereva, Holston begins a 2 year International Research Fellowship (National Science Foundation) to continue research on Thereva at the Swedish Museum of Natural History at Stockholm (SMNH) in collaboration with Thomas Pape. Building on results from worldwide phylogenetic work on Thereva and the revision of the Nearctic Thereva species, a revision of the approximately 70 European Thereva species is proposed for the tenure of the fellowship at the SMNH. The goal of the first year of the project is to compile and organize specimen-level data fundamental to taxonomic revision of Thereva, with results relevant to future interdisciplinary research programs involving the genus. Taxonomic data will be acquired as specimen label data from museum collections and field collections in southern Spain and Greece, with the organization of these data in Mandala. The goal of the second year is to provide the international scientific community with the results of this research in a published revision of European Thereva and species web pages for the North American and European Thereva. The results from this postdoctoral work will broaden our understanding of the European Therevidae and augment research programs with an emphasis on the North American and European fauna. In addition, the lack of species-level taxonomic data for Diptera, and many other insect groups, provided via the internet will make the web-based information source for Thereva species a test case project for developing and producing viable internet taxonomic resources for Diptera.
Expeditions • Kevin Holston received financial support from the Dipterology Fund and a Dissertation Travel Grant from the Graduate College, UIUC, to collect Therevidae for molecular phylogenetic research (25 Aug to 31 Aug 2002). Although Euphycus bocki Kröber, the target species for research on the genus Thereva, was not obtained due to unseaonably cold and wet conditions throughout the summer, adult stiletto flies of the genus Spiriverpa were collected from the 13 collecting sites visited in Primorskiiy Kray. Besides the adults specimens of Spiriverpa, 26 stiletto fly larvae were collected from Anisimovka at the Sukhodol River. From these larvae, adults of Spiriverpa and four specimens of Dialineura lyneborgi Zaitzev were reared in the laboratory. Molecular sequences for D. lyneborgi have recently been obtained at NCSU, and will be added to the molecular dataset compiled for Therevinae (dissertational research by Holston). Logistical support during the collecting trip in Russia was generously provided by Drs Arkady Lelej and Sergey Storozhenko at the Russian Academy of Sciences, Department of Entomology, Institute of Biology and Pedology, Vladivostok, Russia. Mikhail B. Mostovski, Paleontological Institute, Russian Academy of Sciences, Moscow, offered additional advice and logistical support during the initial planning of this project. • Wiegmann collected therevid specimens in Australia at Cooloola National Park. October 2002. • In early October following the International Congress of Dipterology Chris Lambkin, Noel Starick, Gail Kampmeier, and Narelle Power trapped therevids on Moreton Island, near Brisbane, Queensland. Many therevids were observed, photographed, and caught mating on the sand. • Chris Lambkin and Noel Starick successfully trapped the therevid genus Ectinorhynchus in Mullion Range National Park and Giralang Nature Reserve near Orange NSW in late October 2002. • Extensive fires in Carnarvon National Park, Queensland meant quickly revised plans for Dave Yeates, Jacquie Recsei, Noel Starick, and Chris Lambkin in November 2002. Instead the group attacked the semi-arid desert regions of northwestern Victoria over two weeks. Twenty-three Malaise traps were set and serviced at Wyperfeld, Murray Sunset, Little Desert National Parks, Lawloit State Recreation Reserve, and Big Desert State Forest. Much of the collecting activity centered on Little Desert National Park where the myrtaceous heath was at the peak of spring flowering. Because of the severe drought numbers were down with about 700 therevids collected, but biodiversity remained high, with at least one new genus collected. • After the massive bushfires in January- February 2003 Christine Lambkin, Jacqueline Recsei, and Noel Starick returned to Kosciuszko National Park in March to assess the effect of wildfires on Diptera diversity. The results will be compared to the base-line New South Wales National Parks and Wildlife Biodiversity Blitz study in January 2002. They have been also asked to return to replicate their survey in January 2004. • David Yeates, Michael Irwin, Jacqueline Recsei, and Christine Lambkin are part of a large group that spent five weeks in the field in April and May 2003 in the isolated Karajini and Millstream National Parks in the Pilbara region of Western Australia to observe the ecological, and behavioral characteristics of therevids in their environments, fill taxon and geographic gaps in extant therevid material, and provide well- preserved material for molecular studies. The ANIC group will erect 42 malaise traps as well as collect therevids using flight intercept traps, pan traps, hand nets, fogging and soil sieving. • In July 2002 Martin Hauser visited the South West Research Station in Portal, Arizona (http://research.amnh.org/swrs/) and successfully searched for specimens of Henicomyia hubbardii, which he needed for his DNA dataset. • Michael Irwin and Ev Schlinger mounted an expedition to the Fiji Islands expedition to establish relationships and set up network of collecting stations for long-term monitoring of Fiji arthropods (Sept. 19-27). Much to their excitement, the first therevid was caught from that island during this trip. • Michael Irwin went to the Western Cape of South Africa expedition in search of key therevid species for molecular studies. (Oct 6-15). • Michael Irwin was joined by former Madagascar intern, Rasolondalao Harin'Hala Hasinjaka (Rin’ha), on a Southern Madagascar expedition to continue survey of flies on the island (Nov. 2002) (see also Dipteran Biodiversity in Madagascar, below) • In September and October 2002 Martin Hauser joined Fritz Geller-Grimm on a collecting trip through Queensland, Australia, which yield several undescribed species of Therevidae, with some specimens collected in alcohol for DNA studies. • In January 2003 Martin Hauser visited the Museum in Paris to examine type specimens for his revisionary work. • Martin Hauser was invited to join an ALAS expedition to Costa Rica together with Stephen Gaimari in February 2003. During his stay in Costa Rica, he visited Manuel Zumbado and Alvaro Herrera at INBIO and borrowed all the unidentified Therevidae from the institute’s collection. (http://www.evergreen.edu/ants/alascollns/2003.expeditions/Feb03expedition/index.h tml)
Dipteran Biodiversity in Madagascar Over the past four years, Irwin and Schlinger have been conducting a survey of flies, particularly those belonging to the families Therevidae and Acroceridae, throughout Madagascar. Many hundreds of specimens have been obtained so far, representing a large diversity of these flies. This effort is in support of a detailed project on the evolution and systematics of those families of flies from a world perspective. The major method of collecting, deploying a Malaise trap network in selected regions and allowing them to collect material for about a year, has been successfully operated by Harin'Hala Hasinjaka (Rin’ha). This year’s survey being conducted in the South Eastern and South Western portions of the country (mid-October 2002-mid October 2003). An overlapping survey encompassing the West Central region of Madagascar begins in June 2003 through mid-October 2004. This region appears to be rich in species although neither of these target families have yet been collected in that part of Madagascar. Irwin will join Rin'ha for nearly a month starting 1 June 2003 to select strategic localities in which to place the Malaise traps. Thereafter, and for the remainder of the year, Rin'ha will return to the area about every month, work with local people he has employed to monitor the traps, and help service the traps. He will take the collected specimens to Antananarivo where he will remove adult Lepidoptera that contaminate the samples with wing scales, otherwise clean the samples and provide them with fresh 95% ethanol, and sort out the target insect groups using the laboratory facilities of the California Academy of Sciences (CAS) Guest House. The target material will be sent to the respective specialists. Once the survey has been completed, the material will be worked up and monographs of these groups of flies produced for the whole of Madagascar. Support for this research is through the Schlinger Foundation, but the results have a direct bearing on this NSF PEET project.
Specimen curation. Therevid specimens captured in Malaise traps from expeditions described above and subsequently stored in alcohol are being mounted and labelled by Kristin Algmin. The bulk of the specimens are from Madagascar and South Africa, although others have been curated from the Western U.S., Chile, Namibia, and Dominica as well. Kristin estimates between 6-7,000 therevids have been processed since August 2002, and she predicts that up to another 4,000 will be finished by August of 2003. She is training Brad Metz in the techniques.
Laboratory Exchanges and Visitors One of the strengths of our therevid PEET is the ability of members to participate in short term laboratory exchanges to learn new techniques, take advantage of expertise in various laboratories, and cement working relationships among students and scientists on the project. • During the third year of this therevid PEET grant, many PEET members Kevin Holston, Steve Gaimari, and Martin Hauser visited the CSIRO ANIC lab in Canberra Australia in September/October 2002. They examined specimens and obtained a preliminary understanding of the Australian therevids. Shaun Winterton also visited CSIRO at this time, and helped identify material that had accumulated since his studies and publications. Gail Kampmeier visited CSIRO as well in September and talked with Steve Shattuck about BioLink. She also met with other CSIRO staff about databasing issues. • Holston and Hauser continued periodic visits to the Wiegmann lab during year 3 of the project, collaborating on phylogenetic data analyses with lab personnel and graduate student, Hill. • Mikhail B. Mostovski, Palaeontological Institute, Russian Academy of Sciences, Moscow, visited the Irwin lab to collaborate on fossil flies. (July 2002) • Norman and Luciana Johnson, Ohio State University, visited the Irwin lab to discuss collaborative projects involving biodiversity of arthropods. (Sept. 2002) • Gail Kampmeier worked from 6-23 August 2002 with Vololontiana ("Tiana") Rivo Razafindratsita, the biodiversity coordinator at the Institute for Conservation of Tropical Environments (ICTE), working in Ranomafana National Park in Madagascar. Tiana worked to gain the knowledge to support a strong bioinformatics program in Madagascar's national park system.
Database Data entry in Mandala. Nearly 107,000 specimens have been entered into therevid Mandala from nearly 16,000 localities. Label data from each specimen is recorded along with a myriad of information about the specimen itself. Over 4,900 taxa have been entered, over 1,100 representing valid names from the family Therevidae. Nearly 800 literature citations have been entered, as well as about 3,250 illustrations documented, although not yet with images. We are still working on the backlog of specimens from various museum collections, and of course the entry of newly curated specimens from recent expeditions. Improvements to Mandala. Mandala 5.2 was released in September 2002. In response to many novice users and in an effort to facilitate the location of reporting features of Mandala, the features of the control file (control.fp5) were expanded to act as the gateway into Mandala, funnelling all users through it to sign in and initially navigate to their destination. It provides navigation to commonly used features as well as resources available for help and troubleshooting. The original intention of this file still remains in place, where developers can quickly make all files single or multiuser (for using with FMP Server) and mount files for WebMandala The new interface was introduced at the 5th International Congress of Dipterology, with demo files freely available upon request subsequent to that meeting. The meeting also sparked interest in collaborations for three NSF proposals submitted in 2003 that would use and extend the features of Mandala. Mandala 5.2 also now facilitates the importing of large numbers of taxa from other data sources. This is particularly useful for biodiversity studies and surveys where the status and detailed nomenclature of taxonomic names is not the primary focus of the use of Mandala. Through a series of scripts invoked by the user, the data can be massaged and taxonomic hierarchies created with relative ease. This system was developed first for the Allerton Park Biodiversity Blitz in June 2001, and was further enhanced in this release of Mandala 5.2, which will be used for biodiversity studies in the national parks of Madagascar. The ILLUS file now supports images and an image folder has been added to facilitate the referencing of images in Mandala. As always, any taxon, specimen, piece of literature, etc. that is referenced in ILLUS will support the viewing of these related images. Ensuring Data Integrity and Quality in Mandala. Kristin Algmin continues to work on a quality assurance project in the Mandala database, looking for obvious errors and inconsistencies. During year 3 approximately 15,100 specimen label records have been compared with the data interpreted from them in the locality and collection event datafiles, bringing the total proofed records to 30,000. Specimens examined list. Kristin Algmin has helped Martin Hauser and Kevin Holston consolidate and verify their specimens examined lists for upcoming publications. Verification consists of double checking the specimens with the database and specimens examined list, indicating a determination, and parsing them out into their respective loans. This allows for greater accuracy in the upcoming publications. Manual for Georeferencing Specimens. With the prospect of finishing data entry for therevid specimens borrowed from museums and collected in expeditions by our therevid PEET team and other collaborators, Kristin Algmin has finished a manual for georeferencing specimens. This manual will help ensure a certain uniformity of the process by which latitude and longitude are determined by workers when such coordinates are not specified on the label. It will also serve as a measure of confidence in the reliability of the data. Sections of the manual have already been used to aid new workers as they learn to georeference. Interactive Mapping & Keys. Kampmeier worked with Norm Johnson in Sept. 2002 to implement mapping capabilities with WebMandala. In 2003 she also worked with John Pickering, DiscoverLife.org on implementing mapping of therevid specimen data from Mandala with his system that uses TopoZone to map. Elementary searches functional that will bring user back to Mandala output for a single specimen from a map http://pherocera.inhs.uiuc.edu/. She also learned to create interactive keys on the web for general public consumption. These are under construction.
Illustrations Type photos. Primary type specimens (holotypes, neotypes & lectotypes) are being digitally photographed by Kristin Algmin. These type photos will be linked by specimen number and taxon to data recorded in the Illustrations file in Mandala and eventually be able to be viewed on the internet. About 75 specimens have been photographed. Digital archiving of illustrations. The illustrations produced over the course of 6 years by J. Marie Metz and others produced prior to 1996, are now in the process of being digitally archived. This type of archiving provides greater accessibility and economical storage, while providing a way to maintain “healthy” images for years to come. The archiving project is expected to be completed in the summer of 2003 by Brad Metz.
Honors and Awards • Chris Lambkin was elected to the Council for the International Organization for Systematic and Evolutionary Biology (IOSEB) for the term 2002-2008, and the Council for Society for Australasian Systematic Biologists (SASB) for the term 2003- 2005. • J. Marie Metz won first prize for her Henicomyia sp. habitus illustration at the 5th International Congress of Dipterology held in Brisbane Australia (Oct. 2002). • The Guild of Natural Science Illustrators selected both of J. Marie Metz’s entries for their 2003 GNSI Annual juried exhibition for their international meeting in July 2003 in Denver. Selected were H. Fly Habitus and the study, Tongamya Head. Both works were rendered digitally using Adobe Illustrator and Photoshop. • Kevin C. Holston was awarded a postdoctoral fellowship in the NSF’s International Research Fellowship Program. He will be revising the European species of Thereva and building species pages for the web. Holston will be based in Stockholm, Sweden and working under the supervision of Thomas Pape. The fellowship is for two years starting June 2003. • Michael E. Irwin was appointed to the Advisory Board (March 2003- ) of the Archbold Tropical Research and Education Consortium (ATREC), at Clemson University's Springfield Biological Research Station, Dominica. • Michael E. Irwin was appointed to the Board of Directors of Discover Life In America, (2001- ) and is also a member of both the Science and Informatics Committees. • Michael E. Irwin served on the panel of Discover Life In America for funding proposals to the All Taxa Biodiversity Inventory program in the Great Smoky Mountains National Park. • Stephen Gaimari was elected a Fellow of the Linnean Society of London (Feb. 2002) • Martin Hauser’s poster, “Pupal morphology of Asiloidea – an unused character set” received the second prize in the student’s competion at the 5th International Congress of Dipterology in Brisbane, Australia (Oct. 2002). • Martin Hauser’s oral presentation, “New molecular and morphological evidence for the phylogenetic placement of Phycini and Xestomyzini (Diptera, Therevidae)” received 3rd prize in the student competition at the 5th International Congress of Dipterology in Brisbane, Australia (Oct. 2002). • Kevin Holston, Martin Hauser, and Gail Kampmeier each received $2,500 from the Williston Fund to support travel to the 5th International Dipterology Congress in Brisbane, Australia. Each presented papers as well as taking advantage of opportunities to visit colleagues at CSIRO in Canberra and go out to collect and observe the Australian therevid fauna first hand. • Kevin C. Holston was awarded a Dissertation Travel Grant to collect Therevidae in Primorskiy Kray, Russia (17 Aug.-1 Sept. 2002) from the University of Illinois Graduate College with 12% funding match from the UIUC Dept. of Entomology. $1,670. • Kevin C. Holston was awarded $1,000 (Canadian) from the Dipterology Fund to support dissertational research to collect Therevidae in Primorskiy Kray, Russia (17 Aug.-1 Sept. 2002) • David Yeates was elected chair of the Council of the International Congresses of Dipterology, October 2002.
Conference Organization • Irwin, M.E. chaired the Entomological Collections Network meeting in 2002 and organized the two day meeting program for about 200 professional collections managers and museum curators, Ft. Lauderdale, FL, Nov. 2002. • Irwin, M.E. Documenting the Planet's Biodiversity: An Interactive Symposium. Organized symposium and presented opening remarks. Entomological Collections Network meeting, Ft. Lauderdale, FL, Nov. 2002. • Lambkin, C.L. chaired the section on “Phylogeny of Cyclorrhapha” at the 5th International Congress of Dipterology in Brisbane Australia October 2002. • Lambkin, C.L. organised a one and a half day Symposium ‘Incongruence, Data Partitions, and Phylogenetic Signal’ for the Sixth International Congress for Systematic and Evolutionary Biology VI in Patras, Greece, 9-16 September 2002. • Yeates, Irwin and Wiegmann co-organized the symposium, “The Influence of Modern Innovations on Systematic Inquiry with Emphasis on Diptera” at 5th International Congress of Dipterology, Brisbane, Australia. • Yeates, D.K. chaired the Organising Committee of the 5th International Congress of Dipterology in Brisbane Australia October 2002. Christine Lambkin was also a member of that committee. • Yeates co-chaired with Brent Mishler (UC Berkeley) the plenary session, “Connectivity: PEET and the other big initiatives in Biology” at the PEET IV meeting, Berkeley, CA. June 2002.
Invited presentations Fisher, E.M., and Gaimari, S.D.. 2002. Simplified digital photography for museum studies. 5th International Congress of Dipterology, Brisbane, Australia, 29 September-4 October 2002. Hauser, M. 2002. The Therevidae PEET project - a students perspective. In symposium, "Fueling the Renaissance in Descriptive Systematics: the NSF PEET program” held at the National meeting of the Entomological Society of America, Ft. Lauderdale, FL, December. Irwin, M.E. 2002. Casting a sticky web: How partnerships enhance a PEET project. In symposium, "Fueling the Renaissance in Descriptive Systematics: the NSF PEET program” held at the National meeting of the Entomological Society of America, Ft. Lauderdale, FL, December. Irwin, M.E. 2002. Discovering Madagascar's Diversity. Entomological Collections Network meeting, Ft. Lauderdale, FL, December Kampmeier, G.E. 2002. ITIS (Integrated Taxonomic Information System) and PEET. Plenary session at PEET IV organized by David Yeates (co-PI on therevid PEET project) on "Connectivity: PEET and other Big Initiatives in Biology." University of California, Berkeley, June. Kampmeier, G.E., and Holston, K.C. 2002. Meeting the interrelated challenges of specimen, nomenclature, and literature tracking with Mandala. In Program Symposium, “Cataloguing and Databasing,” 5th International Congress of Dipterology, Brisbane, Australia, 29 September-4 October 2002. Lambkin, C.L. and Metz, M.A. 2002. For a future in systematics and the marketability of the discipline: A student perspective on current attitudes and how to prepare for a supply and demand world. Sixth International Congress for Systematic and Evolutionary Biology VI in Patras, Greece, 9-16 of September. Lambkin, C.L. & Yeates, D.K. Modern Innovations in Systematic Inquiry. International Congress of Dipterology in Brisbane October 2002 Lambkin, C.L. & Yeates, D.K. Significant incongruence can be informative: Insights from partitioned support analyses. International Congress of Dipterology in Brisbane October 2002 Lambkin, C.L. & Yeates, D.K. Systematics, evolution, and incongruence: Insights from the World Exoprosopini (Diptera: Bombyliidae). Sixth International Congress for Systematic and Evolutionary Biology (ICSEB) in Patras, Greece on the 9-16 of September 2002 Lambkin, C.L. was filmed in August 2002 for the Australian-wide television program ‘Totally Wild” titled ‘Stiletto Flies’. Moulton, J. K. and B. M. Wiegmann. 2002. Mining the Dipteran Genome: Tools and Resources from GenBank/EMBL and NCBI. Program Symposium, “Cataloguing and Databasing,” 5th International Congress of Dipterology, Brisbane Australia, September 2002. Wiegmann, B. M. 2002. Molecular Biology: A Set of New Characters for Phylogenetic Assessment”, Program Symposium, “ The Influence of Modern Innovations on Systematic Inquiry with Emphasis on Diptera”, Fifth International Congress of Dipterology, Brisbane Australia, September 2002. Wiegmann, B. M., D. K. Yeates, J. K. Moulton, and J. L. Thorne. 2002. Genes, Fossils and Morphology: Combined Data Analysis and a Revised Time Scale for Higher- Level Phylogeny of the Brachycera. Program Symposium, “Multiple Data Set Analyses for Dipteran Phylogenetics,” 5th International Congress of Dipterology, Brisbane Australia, September 2002. Wiegmann, B.M. The Evolutionary History of Flies in the Fossil and Molecular Record. Department of Biological Sciences, Wayne State University, 10 February 2003. Wiegmann, B.M. The Evolutionary History of True Flies. Department of Entomology, The University of Maryland, College Park, MD, 25 April 2003. Winterton, S. L., B. M. Wiegmann, and E. I. Schlinger. 2002. Phylogeny of the small- headed flies (Diptera: Acroceridae) using multiple molecular markers. Program Symposium, “Multiple Data Set Analyses for Dipteran Phylogenetics,” Fifth International Congress of Dipterology, Brisbane Australia, September 2002.
Other Presentations Gaimari, S.D. 2002. Aiming for a phylogenetic classification of the Lauxanioidea. 5th International Congress of Dipterology, Brisbane, Australia, 29 September-4 October 2002. Hamilton, J., Peakall, R. & Yeates, D.K. 2002. Sex, Flies and Videotape: Pollination and Sexual Deception in the Australian Terrestrial Orchid Pterosylis. 5th International Congress of Dipterology, Brisbane, 29th Sept-4th Oct. Hauser, M. 2002. Pupal morphology of Asiloidea – an unused character set. Poster presented at Evolution Society of Systematic Biologists in July 2002 in Champaign, IL Hauser, M. 2002. Pupal morphology of Asiloidea – an unused character set. Poster presentation in Student Competition at 5th International Congress of Dipterology in Brisbane, Australia in Sept. 2002. Hauser, M. 2002. The phylogenetic placement of genera in Phycinae and Xestomyzinae based on DNA data. Oral presentation in Student Competition of 5th International Congress of Dipterology in Brisbane, Australia in Sept. 2002. Lambkin, C.L. & Yeates, D.K. 2002. Significant Incongruence can be Informative: Insights from Partitioned Support Analyses. 5th International Congress of Dipterology, Brisbane, 29th Sept-4th Oct. Lambkin, C.L. & Yeates, D.K. 2002. The Exoprosopini, Australia and the World: Revelations from Systematic Studies. 5th International Congress of Dipterology, Brisbane, 29th Sept-4th Oct. Lambkin, C.L. and Metz, M.A. 2002. For a future in systematics and the marketability of the discipline: A student perspective on current attitudes and how to prepare for a supply and demand world. Sixth International Congress for Systematic and Evolutionary Biology VI in Patras, Greece, 9-16 of September. Lambkin, C.L. and Yeates, D. 2002. Problem solving through integrated systematics. Specimen data bases; NRM, and MORE. CSIRO’s Eaglehawk I in August 2002 Lawson, A.E., Yeates, D.K., McGuire, D.J., Clarke, A.R. & Drew, R.A.I. 2002. DORSALIS – An Interactive Key to Fruit Flies of the Bactrocera (Bactrocera) Dorsalis Complex. 5th International Congress of Dipterology, Brisbane, 29th Sept- 4th Oct. Weigmann, B.M., Yeates, D.K., Moulton, J.K. & Thorne, J.L. 2002. Genes, Fossils and Morphology: Combined Data Analysis and a Revised Time Scale for Higher-Level Phylogeny of the Brachycera. 5th International Congress of Dipterology, Brisbane, 29th Sept-4th Oct. Wiegmann, B.M., D. K. Yeates, J. L. Thorne, and H. Kishino. 2002. Time Flies: A revised time scale for the Diptera. Evolution Society meeting, University of Illinois, Urbana-Champaign. June 2002. Yeates, D.K. 2002. Blowies, Mossies and the Great Aussie Salute. The Cultural History of Flies in Australia. 5th International Congress of Dipterology, Brisbane, 29th Sept- 4th Oct. Yeates, D.K. and Williams, S.E. 2002. Patterns and Levels of endemism in Australia’s wet tropics: concordant signals from invertebrates and vertebrates. Conference of the Australian Ecological Society, Cairns, December 2002. Yeates, D.K., Bickel, D., McAlpine, D., Schneider, M., Cranston, P. & Marshall, S. 2002. An Interactive Key to Fly Families: From Australia to the World! 5th International Congress of Dipterology, Brisbane, 29th Sept-4th Oct. Yeates, D.K., Bickel, D.K. & McAlpine, D.K. 2002. An Estimate of Dipteran Species- Richness in Australia. 5th International Congress of Dipterology, Brisbane, 29th Sept-4th Oct. Yeates, D.K., Irwin, M.E. & Wiegmann, B.M. 2002. A New Family- Level Taxon of Asiloid Flies (Diptera: Brachycera: Asiloidea) Based on a New Genus and Species from Chile, South America. 5th International Congress of Dipterology, Brisbane, 29th Sept-4th Oct. Yeates, D.K., Merritt, D.J. & Baker, C.H. 2002. Neuromere Fusion: A Useful Phylogenetic Marker in Brachyceran Diptera. 5th International Congress of Dipterology, Brisbane, 29th Sept-4th Oct.
Professional Meetings Attended Evolution Society Meetings, University of Illinois, Champaign, IL, June 2002. Attending: Hauser, Holston, Irwin, Metz, Wiegmann, Winterton. Sixth International Congress for Systematic and Evolutionary Biology VI in Patras, Greece, Sept. 2002. Attending: Lambkin. Fifth International Congress of Dipterology, Brisbane, Australia, Sept.-Oct 2002. Attending: Gaimari, Hamilton, Hauser, Holston, Irwin, Kampmeier, Lambkin, Thompson, Wiegmann, Winterton, Yeates. Entomological Society of America, National Meeting, Fort Lauderdale, FL Nov. 2002. Attending: Hauser, Irwin, Kampmeier, Thompson, Wiegmann Entomological Collections Network meeting, Fort Lauderdale, FL, Nov. 2002. Attending: Hauser, Irwin, Kampmeier, Thompson, Wiegmann North American Dipterists Society field meeting, Hocking Hills State Park, OH, May 16-20, 2003. Attending: Wiegmann Australian Ecological Society, Cairns, Dec. 2002. Attending: Yeates. All Taxa Biodiversity Inventory meeting, Great Smoky Mountains National Park, Gatlinburg, TN, Dec. 2002. Attending: Irwin.
Proposals submitted Gaimari, S.D., Metz, M.A., Skevington, J., Lambkin, C.L., and Yeates, D.K. 2003. Crossing the Finish-Line: A World Monograph of the Therevidae. Submitted to NSF program on Planetary Biodiversity Inventories for $4,392,595 for 9/1/03-8/31/08. Subcontract to INHS included M.E. Irwin, M.E. and G.E. Kampmeier.for $139,121.00. Unfunded (2003) (Steve Gaimari, PI, Calif. Division of Food and Agriculture, Sacramento) Irwin, M.E., G.E. Kampmeier. Fiji Arthropod Survey. NSF program on Biotic Surveys and Inventories. 9/1/03-8/31/06. $58,656.00. Unfunded (2003) (Neil Evenhuis, PI, Bishop Museum) Irwin, M.E.. Travel to Fiji. ACES Global Connect Grant Application. $2,500 over 1 year. Unfunded (2003) Pape, T. and Irwin, M.E.. Populating the Therevid Mandala Database. Pre-proposal to GBIF's DIGIT program (Global Biodiversity Information Facility's Digitisation of Natural History Collections). $49,228 over 1 year. Cooperators: Kevin C. Holston, Gail E. Kampmeier. Unfunded (2003) Wiegmann, B.M., Courtney, G.W., Friedrich, M., Meier, R., Yeates, D.K. 2002. Building the Dipteran Tree: Co-operative Research in Phylogenetics and Bioinformatics of True Flies (Insecta: Diptera). For $2,700,275 for 5 yr to National Science Foundation to DEB-Systematic Biology. Subcontract from North Carolina State University to INHS included for G.E. Kampmeier, co-PI with M.E. Irwin. unfunded Wiegmann, B.M., Courtney, G.W., Friedrich, M., Meier, R., Yeates, D.K., Beckenbach, A., Irwin, M.E. 2003. Building the Dipteran Tree: Co-operative Research in Phylogenetics and Bioinformatics of True Flies (Insecta: Diptera). For $2,978,756 for 5 yr (1/1/04-12/31/08) to National Science Foundation to DEB-Systematic Biology, Assembling the Tree of Life program. Includes subcontract for $193,012 to Michael Irwin, PI; Gail Kampmeier (Co-PI), pending. Year 3 Findings Towards a World Monograph of the Therevidae (Insecta: Diptera) Michael E. Irwin, Brian M. Wiegmann, and David K. Yeates, PI’s NSF PEET DEB99-77958 Sept. 2002-Aug. 2003
New Family of Flies from Chile Discovered One of the most significant finds this year was the discovery and publication (in press) of a completely new family of flies from Chile. These newly discovered flies are closely related to stiletto flies (Therevidae) but appear to represent a more ancient lineage that probably predates the early dinasaurs on earth. Adult flies of this new family inhabit dry, hilly terrain, but the immature stages remain unknown and uncollected. New families are discovered only about once in ten years, and the rate of discovery of this level taxon is diminishing exponentially. That this family is closely related to the family Therevidae is an added bonus to our program because the findings can further refine the phylogenetic inferences of the superfamily, better placing the family Therevidae into phylogenetic context.
Relationships of the Therevidae and their Closest Relatives Phylogenetic analysis of 4 genes, 28S rDNA, EF1alpha, CAD, opsin, resulted in trees partially congruent with previous results. Parsimony and Bayesian likelihood analyses of the combined data gave similar results and agreed with previous higher-level classifications. These data support a basal position for the New World xestomyzine genus Henicomyia. Phylogenetic analyses of separate gene partitions yielded trees with lower resolution, weak support and topological conflict. 42 sequences of rh1 opsin were obtained for Therevidae and related Diptera. A gene tree placing the new sequences relative to all previously characterized insect opsins identified two paralogous copies of opsin within the sample --a short wavelength rh1, and a long wavelength opsin. Analyses of just the short wavelength gene suggest that these are orthologs of other known dipteran rh1 genes. The partial fragments generated show some agreement with expected dipteran and therevid relationships, but are limited in their information content for resolving major phylogenetic issues at the subfamily level and above in flies.
Monophyly of Thereva The first phase of phylogenetic analyses in Kevin Holston’s dissertation addressed previous assertions that Thereva that is a polyphyletic assemblage (composed of several unrelated evolutionary therevine lineages), and involved study of 59 species representing 33 genera in the subfamily Therevinae. Using a molecular dataset of two nuclear genes (28S rDNA and elongation factor 1-alpha) and one mitochondrial gene (16S rDNA), a monophyletic circumscription of Thereva was identified that includes Afrotropical, Nearctic, and Palaearctic species and excludes Afrotropical and Neotropical species treated as Thereva by previous authors.
Biogeographical patterns in Thereva The second phase of phylogenetic analyses in Kevin Holston’s dissertation examined species relationships within the monophyletic nominal clade of Thereva using two mitochondrial genes (16S and cytochrome oxidase I) and 33 morphological characters; these analyses included six outgroups and 37 Thereva ingroup species. The position of two major clades of Nearctic species and the position of a Far East Palaearctic species provided evidence for at least three Nearctic evolutionary radiations of Thereva that are derived with respect to Palaearctic species. The phylogenetic relationships from combined analyses support a biogeographic connection between the Nearctic and the eastern Palaearctic, and ten species groups of Thereva were identified.
A revision of Nearctic Thereva Kevin Holston’s dissertation includes a revision of the Nearctic Thereva, providing dichotomous keys, illustrations, character spreadsheets, and descriptions for recognizing and distinguishing among the 35 Nearctic Thereva species. Eleven of the 35 species were previously undescribed. One species, Thereva nobilitata (Fabricius), is common in the western Palaearctic Region and has been collected in British Columbia near Vancouver; this is the first record of a transcontinental introduction of Therevidae. Characters of the male genitalia and the female frons are of primary importance in distinguishing species of Thereva and are illustrated for all species in the Nearctic Region including T. nobilitata. The nomenclature for Nearctic species of Thereva was updated, and lists of specimens examined and distribution maps were provided that were based on specimen data housed in Mandala for over 9500 specimens.
Revision of Henicomyia and Ataenogera Martin Hauser and Don Webb have begun revising the genera Henicomyia and Ataenogera and found 7 new species so far of Henicomyia and 2 new species of Ataenogera in the material currently housed at UIUC and other specimens requested by Hauser from several museums in Canada, Brazil, Costa Rica, and Denmark. To make names of new species available for the publication of the DNA results, Hauser is describing new species in the genera Pentheria, Stenogephyra and Neotabuda. This publication will use the new journal Zootaxa (http://www.mapress.com/zootaxa/index.html), which has a quick turn around time for publication.
Therevid Biodiversity in Madagascar During and after the October 2002 expedition in Madagascar, the number of therevid specimens collected by the network of Malaise traps expanded greatly. We now have over 500 specimens representing several genera and numerous described and new species. The southwestern part of the country is truly rich in therevid taxa. There are now three species of Xestomyzine therevids collected in Madagascar, six species of Phycines, and within the Therevines, three genera have been collected, Stenopomyia, Irwiniella, and a yet to be described genus.