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Walker&RPPLWWHH0HPEHU 'DWH Dr. Sal Sanders$VVRFLDWH'HDQ6FKRRORI*UDGXDWH6WXGLHV 'DWH '\ODQ57KRPDV ABSTRACT 7HUUHVWULDO RSRVVXPV XVH WKHLU VHPLSUHKHQVLOH WDLO IRU JUDVSLQJ QHVWLQJ PDWHULDOV DV RSSRVHG WR ORFRPRWRU PDQHXYHULQJ 7KH REMHFWLYH RI WKLV VWXG\ LV WR UHODWH WKH GHYHORSPHQWRIWKLVDGDSWLYHEHKDYLRUZLWKRQWRJHQHWLFFKDQJHVLQP\RVLQKHDY\FKDLQ 0+& JHQH UHJXODWLRQ DQG LVRIRUP H[SUHVVLRQLQ WKH WDLO IURP ZHHNV WR DGXOWKRRG Monodelphis domestica LVH[SHFWHGWRGHPRQVWUDWHDSURJUHVVLYHDELOLW\WRIOH[WKHWDLOXS WR DJH PRQWKV ZKHQ LW ZLOO KDYH IXOO XVH RI LWV WDLO DQG VKRXOG H[KLELW URXWLQH QHVW FRQVWUXFWLRQ:HK\SRWKHVL]HWKDWMXYHQLOHVWDJHV ±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µWXQLQJ¶ IRU WKH UHSHWLWLYH QHVWUHPRGHOLQJWDVNVUHTXLULQJVXVWDLQHGORZIRUFHFRQWUDFWLRQVRIWKHFDXGDOIOH[RUV LY ACKNOWLEDGEMENTS ,VLQFHUHO\WKDQNP\DGYLVRU'U0LFKDHO7%XWFKHUIRUDOOKLVJXLGDQFHDQGPHQWRULQJ WKURXJKRXW P\ 7KHVLV UHVHDUFK SURMHFW DQG 0DVWHUV 'HJUHH , WKDQN P\ JUDGXDWH FRPPLWWHHPHPEHUV'UV0DUN':RPEOHDQG*DU\5:DONHUIRUFULWLFDOUHYLHZVRI P\ 7KHVLV DQG WKHLU KHOS ZLWK WKH FKDOOHQJHV RI P\ H[SHULPHQWDO SURWRFROV , DP H[WUHPHO\ JUDWHIXO WR -RKQ DQG -DQH 9DQGH%HUJ 7H[DV %LRPHGLFDO 5HVHDUFK ,QVWLWXWH 7%5, IRUFRRUGLQDWLRQDQGDVVLVWDQFHZLWKRSRVVXPFROOHFWLRQDQGWKHHQWLUHVWDIIDWWKH 0RQRGHOSKLV )DFLOLW\ 7%5, IRU DVVLVWDQFH ZLWK DQLPDO FDUH DQGKDQGOLQJ$YHU\ VSHFLDO WKDQNV WR - %XGGH <68 %LRORJLFDO 6FLHQFHV ZKR SURYLGHG DVVLVWDQFH ZLWK T3&57KDQNVWR-<RXQJ 1(20(' DQG7'LJJLQV <68%LRORJLFDO6FLHQFHV IRU KHOS ZLWK VWDWLVWLFDO DQDO\VLV 7KDQNV WR - 0LQ <68 %LRORJLFDO 6FLHQFHV IRU KHOS SURWHLQVHTXHQFLQJDQGSULPHUGHVLJQ7KDQNVWR-5XSHUW ,8620 IRUJXLGDQFHZLWK H[SHULPHQWDO SURWRFROV 1&%, SURYLGHG VRIWZDUH %/$67 IRU SURWHLQ VHTXHQFH PDWFKLQJ 7KLV ZRUN ZDV SDUWLDOO\ VXSSRUWHG E\ 1DWLRQDO 6FLHQFH )RXQGDWLRQ 16) IXQGLQJWR07%XWFKHUDQG<6867(0&ROOHJHIXQGLQJIRUVWXGHQWWUDYHO7KH<68 'HSDUWPHQWRI%LRORJLFDO6FLHQFHVLVDOVRJUDWHIXOO\DFNQRZOHGJHG Y DEDICATION ,GHGLFDWHWKLV7KHVLVWRP\IDPLO\HVSHFLDOO\P\PRWKHU7RQLDQGP\IDWKHU'DQQ\IRU WKHLU XQIDOWHULQJ VXSSRUW ERWK HPRWLRQDOO\ DQG ILQDQFLDOO\ WKURXJKRXW P\ DFDGHPLF FDUHHUDQGGXULQJWKHFRPSOHWLRQRIWKLV7KHVLV YL TABLE OF CONTENTS $SSURYDO3DJHLL &RS\ULJKW3DJHLLL $EVWUDFWLY $FNQRZOHGJPHQWV Y 'HGLFDWLRQYL 7DEOHRI&RQWHQWVYLL /LVWRI7DEOHVYLLL /LVWRI)LJXUHVL[ ,1752'8&7,21 0$7(5,$/6DQG0(7+2'6 Animals Muscle harvesting and Protein preparation Protein analyses: SDS-PAGE and densitometry Genomic analyses: rtPCR and qPCR Behavioral observations Data analysis and Statistics 5(68/76 MHC expression: mRNA and isoform proteins Tail behavior: Correlations with MHC and body size ',6&866,21 Evolutionary implications and Conclusions 5()(5(1&(6 $33(1',; 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