The Cirripede Cretiscalpellum Paucistriatum (Crustacea, Thoracica) in the Lower Maastrichtian of NW Europe - Is It Stratigraphically Useful?* by John W.M
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BULLETIN DE L'INSTITUT ROYAL DES SCIENCES NATURELLES DE BELGIQUE SCIENCES DE LA TERRE , 77: 107-116,2007 BULLETIN VAN HET KONINKLIJK. BELGISCH INSTITUUT VOOR NATUURWETENSCHAPPEN AARDWETENSCHAPPEN, 77: 107-116,2007 The cirripede Cretiscalpellum paucistriatum (Crustacea, Thoracica) in the lower Maastrichtian of NW Europe - is it stratigraphically useful?* by John W.M. JAGT & Barry W.M. VAN BAKEL Jagt, J.W.M. & Van Bakel, B.W.M., 2007 - The cirripede paucistriatum (Woodward, 1906) ont été découverts dans le Cretiscalpellum paucistriatum (Crustacea, Thoracica) in the Maastrichtien inférieur (Zone à obtusa) en affleurement dans la carrière Satum à Kronsmoor lower Maastrichtian of NW Europe - is it stratigraphically useful? (Allemagne septentrionale) ; la carène Bulletin de l'Institut royal des Sciences naturelles de Belgique, de ce taxon est décrite pour la première fois. Ces spécimens, ainsi Sciences de la Terre, 77: 107-116, 1 fig., 2 pis, Brussels, October que ceux décrits précédemment dans la littérature, suggèrent que 15, 2007-ISBN 0374-6291. C. paucistriatum est un rameau à courte extension stratigraphique du groupe C. striatum, apparemment confiné dans un intervalle s'étendant de la * Publication No. 16 of the Working Group 'Vijlen and partie inférieure de la Zone à obtusa à la partie inférieure de la Zone à sumensis du Maastrichtien inférieur, Beutenaken members' avec, à ce jour, des découvertes dans le Norfolk (Angleterre orientale), au Limbourg méridional (Pays-Bas) et en Allemagne septentrionale. Cette espèce pourrait donc être utile d'un point de Abstract vue stratigraphique. Pour tester cette hypothèse, il faudrait tamiser des faunules à cirripèdes d'âge équivalent en Belgique (Bassin de New material of the calanticid cirripede Cretiscalpellum Mons, Formation des Craies phosphatées de Ciply-Malognes), au paucistriatum (Woodward, 1906) is recorded from the lower Danemark oriental (Mon, faciès à calcaire blanc), en Allemagne du Maastrichtian (obtusa Zone) as exposed at the Saturn quarry, nord-est (Rügen, faciès à calcaire blanc) et même dans des gisements Kronsmoor (northern Germany); the carina is described for the first plus éloignés (notamment en Pologne centrale et orientale ; faciès time. These specimens, together with material described previously à opoka et à calcaire blanc). Bien qu'on considère généralement in the literature, suggest C. paucistriatum to have been a short- que la valeur stratigraphique des cirripèdes est plutôt limitée, il lived offshoot of the C. striatum group, apparently confined to the existe une tendance chez les scalpellomorphes Cretiscalpellum et interval ffom the lower obtusa to the lower sumensis zones of the Arcoscalpellum à développer des espèces à durée de vie courte au lower Maastrichtian, with records to date from Norfolk (eastern cours du Crétacé supérieur (Campanien et Maastrichtien). De tels England), southem Limburg (The Netherlands) and northern taxons, considérés conjointement avec d'autres membres des lignées Germany. The species may thus be stratigraphically useful. To principales présents dans les mêmes niveaux, pourraient s'avérer test this assumption, cirripede faunules from coeval strata in utiles dans le cadre de corrélations (supra-) régionales; de plus, southern Belgium (Mons Basin; Ciply-Malogne Phosphatic Chalk des 'registres d'extension de taxons' et des 'zones d'assemblages' Formation), eastem Denmark (Mon; white chalk facies), NE pourraient être établies sur base de cirripèdes. Germany (Rügen; white chalk facies) and even further afield (e.g., central and eastem Poland; opoka and white chalk facies) need to Mots-clefs: Cirripedia, Cretiscalpellum, Maastrichtien, Europe be screened. Although the général view is that the corrélative value du nord-ouest, stratigraphie. of cirripede taxa is rather limited, a tendency may be noted in the scalpellomorph genera Cretiscalpellum and Arcoscalpellum to develop short-lived species during the Late Cretaceous (Campanian Introduction and Maastrichtian). Such taxa, considered in conjunction with co-occurring members of the main lineages, may prove useful in (supra-)regional corrélation, and cirripede 'taxon range' and ln November-December 2003, within the framework 'assemblage' zones could thus be established. of the 'Access to Belgian Collections (ABC)' grant scheme, one of us (JWMJ) started a revision of Keywords: Cirripedia, Cretiscalpellum, Maastrichtian, NW cirripede collections held at the Institut royal des Europe, stratigraphy. Sciences naturelles de Belgique (IRScNB, Brussels). The aim was twofold: to redescribe type material Résumé from the extended type area of the Maastrichtian Stage, mainly that of BOSQUET (1854, 1857) and De nouveaux fossiles du cirripède calanticidé Cretiscalpellum WlTHERS (1935), and to détermine in more detail its 108 John W.M. JAGT & Barry W.M. VAN BAKEL stratigraphie provenance as based on preservational limited stratigraphie value, it may be that certain type, adhering matrix and personal fieldwork short-lived species (offshoots of the main lineage) can experience. Old collections generally suffer from a in fact be used for (supra-)regional corrélations. In the lack of stratigraphie data, but the fact that almost literature, there are a few examples of such cirripede ail localities from which Bosquet (1854, 1857) 'taxon range' and 'assemblage' zones (COLLINS described cirripede material are still accessible means & Mellen, 1973; ZULLO, 1984; Canis & Zullo, that stratigraphie ranges of the various species may 1986). yet be refined by renewed collecting. In recent years, Of the scalpellomorphs Cretiscalpellum and fieldwork has resulted in the discovery of new species Arcoscalpellum, most species are long-ranging and (collins & Jagt, 1999; Jagt & collins, 1999) and widely distributed (e.g., C. glabrum (roemer, 1841), copious material of species described by bosquet and A. maximum (J. de c. sowerby, 1829) and A. fossula authors is now available. The by other newly collected (Darwin, 1851)), but a handful of others (see e.g., material will also enable a robust taxonomy to be Withers, 1935; Alekseev, 1974, 1979) have a more worked out. Results of this restricted study will be published in stratigraphie range. Potentially, these may a series of brief notes (Jagt, 2004, in prep.) and, later, be used for corrélations, but the type material of the in a various taxa monograph in two parts, on scalpellomorphs and needs to be restudied and its ranges refined verrucomorphs/brachylepadomorphs, respectively. and calibrated to modem biozonations, preferably A comparison of material collected (September based on belemnitellid coleoids and ammonites and at Saturn 1988) the quarry near Kronsmoor (northern inoceramid bivalves. In that way, also trans-Atlantic Germany) with wlthers's (1935) records of corrélations become possible. On some occasions Cretiscalpellum paucistriatum from Trimingham (Collins & Mellen, 1973; Jagt & Collins, 1989, (Norfolk, England) and Slenaken (southern Limburg, 1999) it has been demonstrated, or hinted at, that The Netherlands), has led us to reflect upon the possible species from the Gulf Coast of the United States and of that NW stratigraphie value species. Although précisé Europe are obviously closely related and occur at stratigraphie data are lacking for the Trimingham and comparable stratigraphie intervais. Slenaken records, recent work on other macrofaunal To illustrate what might be possible in this way, groups (e.g., belemnitellid coleoids, brachiopods) at C. paucistriatum is briefly discussed here. Only these localities allows us to state that c. paucistriatum screening of existing collections from, or renewed appears to be confined to a fairly narrow stratigraphie collecting at, Rügen, Mon, the Mons Basin and interval (lower obtusa to lower sumensis zones; sensu perhaps further afield (e.g., central and eastern Schulz, 1979) within the lower Maastrichtian. In Poland) can demonstrate whether or not our claim spite of the that C. général view that cirripedes are of rather paucistriatum is stratigraphically usefi.il, can kazimiroviensis Zone sensu auctt. baltica / danica Zone danica / argentea Zone argentea /junior Zone regulatus /junior Zone fastigata Zone cimbrica Zone sumensis Zone obtusa Zone pseudobtusa Zone lanceolata Zone Fig. 1 — Map of northern Europe showing localities (dots) from which Cretiscalpellum paucistriatum has so far been recorded, and biozonation of the Maastrichtian Stage, based on the white chalk standard section of northern (schulz & Schmid, et Germany 1983; SCHULZ al., 1984; Schönfeld et al., 1996; Christensen et al., 2004). The cirripede Cretiscalpellum paucistriatum in the lower Maastrichtian 109 be upheld. by Bosquet himself, or by someone else after his To denote the repositories of specimens referred death (June 1880) when his collection was transferred to and/or illustrated here, the following abbreviations to Brussels. For comparison, two left scuta (IRScNB MI are used: IRScNB MI - Institut royal des Sciences 11027, 11028) in the same lot from Slenaken, here naturelles de Belgique, Brussels [Mesozoic referred to Cretiscalpellum striatum (darwin, 1851), are illustrated Invertebrates]; NHM - The Natural History Museum, (PI. 2, Figs 4, 5). Department of Palaeontology, London; NHMM - Natuurhistorisch Museum Maastricht (JJ = J.W.M. Description Jagt Colln). Carina (Pl. 1, Figs 3-5): apical third broken, estimated length c. 25 mm, i.e. near-equalling 2.5 times width; slightly bowed inwards; tectum transversely strongly Systematic palaeontology