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THE NEW CLASSIFICATION OF GASTROPODS ACCORDING TO BOUCHET & ROCROI, 2005 Guido T. Poppe & Sheila P. Tagaro February 23, 2006 VISAYAV - FEB., 2006 THE NEW CLASSIFICATION OF GASTROPODS G.T. POPPE & S.P. TAGARO The New Classification of Gastropods according to Bouchet & Rocroi, 2005

A practical adaptation for conchologists, malacologists and shell collectors.

Guido T. Poppe & Sheila P. Tagaro

The Volume 47 of Malacologia (2005) counts 397 SPF Nacelloidea pages, entirely dedicated to the new “Classification Nacellidae – this is new, and Nomenclator of Gastropod Families” by Philippe before either in Bouchet & Jean-Pierre Rocroi. or or The work drastically changes existing systematics . and is an important step in gastropod nomenclature. SPF Lottioidea As each change needs a lot of adaptations of the Lottiidae mind, research and courage, we here attempt to Acmaeidae resume Bouchet & Rocroi’s majestic achievement so that workers in the field get an easy access and SPF Neolepetopsoidea memory guide to the new classification.

Volume 47 of Malacologia is indispensable in any Clade library of significance and can be acquired from Klaus & Christina Groh, Germany: SPF not assigned: [email protected] Ataphridae Pendromidae An overview with notes on the changes and topics SPF Fissurelloidea that will be followed by Conchology, Inc. is given Fissurellidae below. SPF Haliotoidea Haliotidae As our company and most registered members SPF Leptelloidea of Conchology, Inc. are mainly concerned with Lepetellidae recent , we filtered out the numerous Addisoniidae families. Bathyphytophilidae Caymanabyssidae In BOLD: families of importance to collectors Cocculinellidae In BLUE: remarks Osteopeltidae In RED: families we maintain – not as in Bouchet Pseudococculinidae & Rocroi. Pyropeltidae SPF Leptodrilloidea Clade Patellogasropoda Lepteodrillidae Clypeosectidae SPF (=Superfamily) Patelloidea Sutilizonidae Patellidae SPF Neomphaloidea Neomphalidae

2 WWW.CONCHOLOGY.BE G.T. POPPE & S.P. TAGARO THE NEW CLASSIFICATION OF GASTROPODS VISAYAV - FEB., 2006 Melanodrymiidae – now Peltospiridae contains also the SPF Pleurotomarioidea Tricolia (formely Pleurotomariidae Tricoliidae) SPF Scissurelloidea Clade Cocculiniformia Anatomidae – these were in Scissurellidae before. SPF Cocculinoidea SPF Seguenzioidea Cocculinidae Seguenziidae – now well defined, before Clade Cycloneritimorpha found in etc. SPF Chilodontidae–before in Trochidae, now a homogenous family Proserpinellidae of in general heavily sculptured dull SPF Hydrocenoidea colored shells. Hydrocenidae SPF SPF Trochidae Stomatellidae – Conchology, Inc. SPF Neritopsoidea maintains this as Neritopsidae a family – in B & R a Titiscaniidae subfamily of Trochidae. Zygopleuroid Group : Abyssochrysidae Calliostomatidae – before in Trochidae. Informal group Solariellidae – before in Trochidae. SPF Ampullarioidea SPF Ampullariidae SPF Angariidae – Conchology, Cyclophoridae Inc. maintains this as a family – in B & R Craspedopomatidae as a subfamily Diplommatinidae Turbinidae. Maizanidae – Conchology, Megalomastomatidae Inc. maintains this as Neocyclotidae a family – in B & R Pupinidae as a subfamily of SPF Viviparoidea Turbinidae Viviparidae – before often called Cyclostre- Clade Sorbeconcha matidae - now well defined. SPF

WWW.CONCHOLOGY.BE 3 VISAYAV - FEB., 2006 THE NEW CLASSIFICATION OF GASTROPODS G.T. POPPE & S.P. TAGARO SPF – formerly in – now contains Cerithiidae. Lacuninae. Dialidae – formerly in Cerithiidae. Diastomatidae Litiopidae Zerotulidae Melanopsidae SPF Naticoidea Modulidae Pachychilidae – formerly in SPF or Thiaridae (e.g. Faunus ater). Paludomidae SPF Rissoidea – now also contains Fossarinae. – now contains Pleuroceridae Baicaliinae. Potamididae Scaliolidae Siliquariidae Thiaridae Turritellidae – now also containsVermicu- Calopiidae lariinae. SPF Campaniloidea Campanilidae Emblandidae Ampullinidae Epigridae Plesiotrochidae Falsicingulidae Helicostoidae Clade Hydrococcidae SPF Calyptraeoidea SPF Capuloidea SPF Cingulopsoidea – now contains Vitrinellinae. Rastodentidae SPF SPF – now contains Jenneriinae and Seraphidae – Formerly in . Strombidae. SPF Ficoidea For .

4 WWW.CONCHOLOGY.BE G.T. POPPE & S.P. TAGARO THE NEW CLASSIFICATION OF GASTROPODS VISAYAV - FEB., 2006 SPF Buccinidae. – Conchology, Columbellidae Inc. maintains this Fasciolariidae as a family – in Nassariidae B & R a subfamily of Melongenidae . SPF Muricoidea Tonnidae Muricidae Babyloniidae – formerly in Buccinidae. Costellariidae Pisanianuridae – formerly Cysticidae – formerly in in . Marginellidae. Ranellidae Harpidae SPF Marginellidae Mitridae Pleioptygmatidae Strepsiduridae SPF Turbinellidae – now – now contains contains Lamellariinae. Columbarinae. – now contains Volutidae Eratoinae. Volutomitridae SPF SPF Olivoidea Olividae SPF Olivellidae – formerly in Olividae. SPF Pseudolivoidea “Group” Pseudolividae – formerly in Olividae. SPF Ptychatractidae Epitoniidae SPF Conoidea Janthinidae Conidae – now contains the following SPF Eulimoidea subfamilies that were formerly in Aclididae Turridae: Clathurel- SPF linae, Conorbinae, Mangeliinae, Oeno- potinae & Raphi- tominae. Conchology, Inc. will continue Clade to use “Conidae” as before and group the SPF Buccinoidea above subfamilies in Buccinidae – now contains Conidae cf. Busyconinae. Clavatulidae – formerly in Colubrariidae – formerly in Turridae.

WWW.CONCHOLOGY.BE 5 VISAYAV - FEB., 2006 THE NEW CLASSIFICATION OF GASTROPODS G.T. POPPE & S.P. TAGARO Drilliidae – formerly in Informal group Turridae. Pseudomelatomidae – Clade formerly in Turridae. Strictispiridae – formerly in SPF Bulloidea Turridae. Bullidae Terebridae SPF Diaphanoidea Turridae Diaphanidae SPF Cancellarioidea Notodiaphanidae Cancellariidae SPF Haminoeoidea Haminoeidae Clade Bullactidae Smaragdinellidae Unassigned to SPF SPF Philinoidea Cimidae Orbistellidae Tjaernoeiidae Xylodisculidae SPF Philinoglossidae Acteonidae Plusculidae Aplustridae – formerly placed in SPF Runcinoidea Hydatinidae. Runcinidae Bullinidae Ilbiidae SPF Architectonicoidea Architectonicidae Clade Thecosomata SPF Glacidorboidea SPF Cavolinioidea SPF Mathildoidea Cavoliniidae Mathildidae Limacinidae SPF Omalogyroidea SPF Cymbulioidea Omalogyridae Cymbuliidae SPF Desmopteridae Peraclidae Clade Gymnosomata SPF Ringiculoidea Ringiculidae SPF Clionoidea SPF Rissoelloidea Clionidae Rissoellidae SPF Valvatoidea Notobranchidae Valvatidae Cornirostridae Hyalogyrinidae SPF Hydromyloidea Hydromylidae Laginiopsidae

6 WWW.CONCHOLOGY.BE G.T. POPPE & S.P. TAGARO THE NEW CLASSIFICATION OF GASTROPODS VISAYAV - FEB., 2006 Clade Aplysiomorpha Clade Umbraculida

SPF Aplysioidea SPF Aplysiidae Umbraculidae SPF Akeroidea Tylodinidae Akeridae Clade “Group” ” Subclade Pleurobranchomorpha SPF Acochlidioidea Acochlidiidae SPF Pleurobranchoidea SPF Hedylopsoidea Hedylopsidae Ganitidae Subclade Nudibranchia Livorniellidae Minicheviellidae Unassigned SPF Parhedylidae Rhodopidae Tantulida SPF Palliohedyloidea Clade Euctenidiacea Palliohedylidae SPF Strubellioidea Subclade Gnathodoridacea Strubelliidae SPF Bathydoridoidea Pseudunelidae Bathydorididae Subclade Clade SPF Subclade Oxynoacea SPF Oxynooidea Oxynoidae SPF Phyllidioidea Juliidae Volvatellidae Mandeliidae Subclade Placobranchacea SPF Onchidoridoidea SPF Placobranchoidea Placobranchidae Boselliidae SPF Platyhedylidae SPF Aegiretidae Gymnodorididae Hexabranchidae Okadaiidae

“Group” Cylindrobullida Clade Nudibranchia

SPF Cylindrobulloidea Clade Pseudoeuctenidiacea Cylindrobullidae

WWW.CONCHOLOGY.BE 7 VISAYAV - FEB., 2006 THE NEW CLASSIFICATION OF GASTROPODS G.T. POPPE & S.P. TAGARO SPF Doridoxoidea SPF Doridoxidae Clade Glaucidae Piseinotecidae Unassigned SPF Charcotiidae Informal Group Informal Group Basommatophora Goniaeolididae SPF Heroidae Amphibulidae SPF Pinufiidae Siphonariidae Clade Subclade Euarminida SPF Chilinoidea SPF Chilinidae Latiidae Doridomorphidae SPF Acroloxoidea Acroloxidae Subclade Dendronotida SPF Lymnaeoidea Lymnaeidae SPF Tritonioidea SPF Planorboidea Planorbidae Aranucidae Physidae Clade SPF Trimusculoidea Trimusculidae SPF Otinoidea Otinidae Smeagolidae Subclade SPF Ellobioidea SPF Flabellinoidea Clade Notaeolidiidae SPF SPF Onchidioidea Onchidiidae SPF Veronicellidae Pseudovermidae Rathouisiidae

8 WWW.CONCHOLOGY.BE G.T. POPPE & S.P. TAGARO THE NEW CLASSIFICATION OF GASTROPODS VISAYAV - FEB., 2006 Clade SPF Achatinoidea Achatinidae Subclade Elasmognatha Ferussaciidae Micractaeonidae SPF Succineoidea Subulinidae SPF Aillyoidea SPF Athoracophoroidea Aillyiidae Athoracophoridae SPF Testacelloidea Testacellidae Subclade SPF Partuloidea SPF Papillodermatoidea Partulidae Papillodermatidae Draparnaudiidae SPF Streptaxoidea SPF Achatinelloidea SPF Rhytidoidea SPF Cochlicopoidea Rhytididae Cochlicopidae Chlamydephoridae Amastridae Haplotrematidae SPF Scolodontidae Pupillidae SPF Acavoidea Argnidae Chondrinidae Dorcasiidae Macrocyclidae Pleurodiscidae Megomphicidae Pyramidulidae Strophocheilidae Spelaeoconchidae SPF Plectopyloidea Plectopylidae Corillidae Valloniidae Sculptariidae Vertiginidae SPF SPF Enoidea Enidae Informal GroupSigmurethra SPF Clausilioidea Clausiliidae Thyrophorellidae SPF SPF Sagdoidea Orthalicidae Cerionidae Coelociontidae “Limacoid” clade Megaspiridae Placostylidae SPF Staffordioidea Urocoptidae Staffordiidae

WWW.CONCHOLOGY.BE 9 VISAYAV - FEB., 2006 THE NEW CLASSIFICATION OF GASTROPODS G.T. POPPE & S.P. TAGARO SPF Dyakioidea SPF Chronidae SPF SPF Zonitoidea SPF Helicarionoidea SPF Boettgerillidae SPF Arionoidea Anadenidae Ariolimacidae Binneyidae Oopeltidae SPF Bradybaenidae Epiphragmophoridae Halolimnohelicidae Helminthoglyptidae Humboldtianidae Monadeniidae

10 WWW.CONCHOLOGY.BE V V COUNCIL OF ADMINISTRATION PRESIDENT Philippe Poppe DIRECTOR Guido T. Poppe

ADDRESS Conchology, Inc. Cebu Light Industrial Park Basak, Lapu-Lapu City Cebu 6015 Philippines

DESIGN & GRAPHICS Dulce Rose P. Lada

TELEPHONE NO. +63 32 495 9990 FAX NO. +63 32 495 9991

SUBMISSION OF ARTICLES All Authors Are Invited

All conchological/malacological authors are invited to place their articles here. The net is a very visual medium, and numerous digital (scanned) photo- graphs are welcome. We have the space and tech- nology to serve thousands of viewers. When our article is online, we will instantly inform our 3,000 registered members. No paper journal today can match this number of readers. We propose our help for photography of specimens if needed. For details you may email [email protected]

Conchology, Inc. Cebu Light Industrial Park, Basak, Lapu-Lapu City Cebu 6015, Philippines website: www.conchology.be email: [email protected] ISSN 1656-4650