Taxon Representative Source Morphology Age/Record Elasmobranchii Reif (1973); Motta (2004) Non-Cone-Shaped Teeth Cretaceous Ichthyodectiformes Schwimmer Et Al

Total Page:16

File Type:pdf, Size:1020Kb

Taxon Representative Source Morphology Age/Record Elasmobranchii Reif (1973); Motta (2004) Non-Cone-Shaped Teeth Cretaceous Ichthyodectiformes Schwimmer Et Al Taxon Representative source Morphology Age/Record Elasmobranchii Reif (1973); Motta (2004) non-cone-shaped teeth Cretaceous Ichthyodectiformes Schwimmer et al. (1997b) not all species possess procumbent unknown from Euro- (e.g., Xiphactinus) anterior teeth pean Maastrichtian Euteleostei Friedman (2009) often numerous fanglike conical teeth Late Cretaceous alternated by small teeth Dyrosauridae Khosla et al. (2009) narrow arc of front teeth, different no contemporary tooth sizes records in Europe Pliosauridae Ketchum and Benson (2010); procumbent anterior teeth known disappeared more than Fischer et al. (2015) 20 Ma before Echinoco- rys record Aristonectes parvidens (and the Cabrera (1941); Gasparini et relatively large cranium (0.73 m Maastrichtian of the Wed- following D Elasmosauridae) al. (2003); O’Gorman (2016) length); jaw bones containing nu- dellian Sea merous tiny alveoles Elasmosaurus platyurus Sachs (2005) prognathous dentition early Campanian of Kansas Hydrotherosaurus alexandriae Welles (1943) skull length only 0.33 m; remarkable Campanian-Maastrichtian irregular dentition, not procumbent of California Kaiwhekea katiki Cruickshank and Fordyce small and homodontous needle- Maastrichtian of New (2002) shaped teeth; skull length 0.62 m Zealand Libonectes morgani Carpenter (1997, 1999) premaxillae and dentary with prog- early Turonian of Texas nathous anterior tooth pairs Styxosaurus snowi Welles (1952); Carpenter prognath, but skull too short (0.47 m) Coniacian–Santonian of (1999) and slender teeth Texas Terminonatator ponteixensis Sato (2003) skull length 0.26 m; anteriorly procum- late Campanian of bent teeth Saskatchawan Tuarangisaurus keyesi Wiffen and Moisley (1986); long and narrow teeth, skull length Maastrichtian of New Carpenter (1999) 0.37 m; premaxillary teeth inter- Zealand locked with dentary teeth which are meagerly prognath Dolichorhynchops herschelensis Sato (2005) extremely narrow arrangement of late Campanian-early (D Polycotylidae) teeth (in parallel) Maastrichtian of Saskatchewan.
Recommended publications
  • Estimating the Evolutionary Rates in Mosasauroids and Plesiosaurs: Discussion of Niche Occupation in Late Cretaceous Seas
    Estimating the evolutionary rates in mosasauroids and plesiosaurs: discussion of niche occupation in Late Cretaceous seas Daniel Madzia1 and Andrea Cau2 1 Department of Evolutionary Paleobiology, Institute of Paleobiology, Polish Academy of Sciences, Warsaw, Poland 2 Independent, Parma, Italy ABSTRACT Observations of temporal overlap of niche occupation among Late Cretaceous marine amniotes suggest that the rise and diversification of mosasauroid squamates might have been influenced by competition with or disappearance of some plesiosaur taxa. We discuss that hypothesis through comparisons of the rates of morphological evolution of mosasauroids throughout their evolutionary history with those inferred for contemporary plesiosaur clades. We used expanded versions of two species- level phylogenetic datasets of both these groups, updated them with stratigraphic information, and analyzed using the Bayesian inference to estimate the rates of divergence for each clade. The oscillations in evolutionary rates of the mosasauroid and plesiosaur lineages that overlapped in time and space were then used as a baseline for discussion and comparisons of traits that can affect the shape of the niche structures of aquatic amniotes, such as tooth morphologies, body size, swimming abilities, metabolism, and reproduction. Only two groups of plesiosaurs are considered to be possible niche competitors of mosasauroids: the brachauchenine pliosaurids and the polycotylid leptocleidians. However, direct evidence for interactions between mosasauroids and plesiosaurs is scarce and limited only to large mosasauroids as the Submitted 31 July 2019 predators/scavengers and polycotylids as their prey. The first mosasauroids differed Accepted 18 March 2020 from contemporary plesiosaurs in certain aspects of all discussed traits and no evidence Published 13 April 2020 suggests that early representatives of Mosasauroidea diversified after competitions with Corresponding author plesiosaurs.
    [Show full text]
  • On the Cranial Anatomy of the Polycotylid Plesiosaurs, Including New Material of Polycotylus Latipinnis, Cope, from Alabama F
    Marshall University Marshall Digital Scholar Biological Sciences Faculty Research Biological Sciences 2004 On the cranial anatomy of the polycotylid plesiosaurs, including new material of Polycotylus latipinnis, Cope, from Alabama F. Robin O’Keefe Marshall University, [email protected] Follow this and additional works at: http://mds.marshall.edu/bio_sciences_faculty Part of the Animal Sciences Commons, and the Ecology and Evolutionary Biology Commons Recommended Citation O’Keefe, F. R. 2004. On the cranial anatomy of the polycotylid plesiosaurs, including new material of Polycotylus latipinnis, Cope, from Alabama. Journal of Vertebrate Paleontology 24(2):326–340. This Article is brought to you for free and open access by the Biological Sciences at Marshall Digital Scholar. It has been accepted for inclusion in Biological Sciences Faculty Research by an authorized administrator of Marshall Digital Scholar. For more information, please contact [email protected], [email protected]. ON THE CRANIAL ANATOMY OF THE POLYCOTYLID PLESIOSAURS, INCLUDING NEW MATERIAL OF POLYCOTYLUS LATIPINNIS, COPE, FROM ALABAMA F. ROBIN O’KEEFE Department of Anatomy, New York College of Osteopathic Medicine, Old Westbury, New York 11568, U.S.A., [email protected] ABSTRACT—The cranial anatomy of plesiosaurs in the family Polycotylidae (Reptilia: Sauropterygia) has received renewed attention recently because various skull characters are thought to indicate plesiosauroid, rather than plio- sauroid, affinities for this family. New data on the cranial anatomy of polycotylid plesiosaurs is presented, and is shown to compare closely to the structure of cryptocleidoid plesiosaurs. The morphology of known polycotylid taxa is reported and discussed, and a preliminary phylogenetic analysis is used to establish ingroup relationships of the Cryptocleidoidea.
    [Show full text]
  • Morphologic and Ontogenetic Patterns in Elasmosaur Neck Length, with Comments on the Taxonomic Utility of Neck Length Variables F
    View metadata, citation and similar papers at core.ac.uk brought to you by CORE provided by Collection Of Biostatistics Research Archive Marshall University Marshall Digital Scholar Biological Sciences Faculty Research Biological Sciences 6-1-2006 Morphologic and Ontogenetic Patterns in Elasmosaur Neck Length, with Comments on the Taxonomic Utility of Neck Length Variables F. Robin O’Keefe Marshall University, [email protected] Norton Hiller Follow this and additional works at: http://mds.marshall.edu/bio_sciences_faculty Part of the Aquaculture and Fisheries Commons, and the Ecology and Evolutionary Biology Commons Recommended Citation O’Keefe FR, Hiller N (2006) Morphologic and ontogenetic patterns in elasmosaur neck length, with comments on the taxonomic utility of neck length variables. Paludicola 5:206–229. This Article is brought to you for free and open access by the Biological Sciences at Marshall Digital Scholar. It has been accepted for inclusion in Biological Sciences Faculty Research by an authorized administrator of Marshall Digital Scholar. For more information, please contact [email protected]. Paludicola 5(4):206-229 June 2006 by the Rochester Institute of Vertebrate Paleontology MORPHOLOGIC AND ONTOGENETIC PATTERNS IN ELASMOSAUR NECK LENGTH, WITH COMMENTS ON THE TAXONOMIC UTILITY OF NECK LENGTH VARIABLES F. Robin O'Keefe1 and Norton Hiller2 1Department of Anatomy, NYCOM 2 rm. 321, New York College of Osteopathic Medicine Old Westbury, New York 11568, [email protected] 2Canterbury Museum, Rolleston Avenue Christchurch, 8001 New Zealand, [email protected] ABSTRACT Elasmosaur cervical vertebrae are common fossils, but their taxonomic utility is limited due to a lack of understanding concerning their shape within and among taxa.
    [Show full text]
  • Cranial Anatomy and Taxonomy of Dolichorhynchops Bonneri New
    Marshall University Marshall Digital Scholar Biological Sciences Faculty Research Biological Sciences 2008 Cranial anatomy and taxonomy of Dolichorhynchops bonneri new combination, a polycotylid (Sauropterygia: Plesiosauria) from the Pierre Shale of Wyoming and South Dakota F. Robin O’Keefe Marshall University, [email protected] Follow this and additional works at: http://mds.marshall.edu/bio_sciences_faculty Part of the Animal Sciences Commons, and the Ecology and Evolutionary Biology Commons Recommended Citation O'Keefe, F. R. (2008). Cranial anatomy and taxonomy of Dolichorhynchops bonneri new combination, a polycotylid (Sauropterygia: Plesiosauria) from the Pierre Shale of Wyoming and South Dakota. Journal of Vertebrate Paleontology, 28(3), 664-676. This Article is brought to you for free and open access by the Biological Sciences at Marshall Digital Scholar. It has been accepted for inclusion in Biological Sciences Faculty Research by an authorized administrator of Marshall Digital Scholar. For more information, please contact [email protected], [email protected]. CRANIAL ANATOMY AND TAXONOMY OF DOLICHORHYNCHOPS BONNERI NEW COMBINATION, A POLYCOTYLID (SAUROPTERYGIA: PLESIOSAURIA) FROM THE PIERRE SHALE OF WYOMING AND SOUTH DAKOTA F. ROBIN O’KEEFE ABSTRACT The taxonomic identity of two well-preserved polycotylid plesiosaur skeletons from the Pierre Shale of far northern Wyoming and southern South Dakota has been controversial since their discovery. Originally referred to Dolichorhynchops osborni, the material was almost immediate-ly christened Trinacromerum bonneri Adams 1997; more recently the material has been referred to Polycotylus. Recent preparation of the well-preserved skull of one specimen permits detailed examination of the cranial morphology of this animal for the first time, and allows for its inclusion in a cladistic analysis of the Polycotylidae.
    [Show full text]
  • Remarks on the Pectoral Girdle of Hydrotherosaurus Alexandrae (Plesiosauria: Elasmosauridae)
    Sachs, Pectoral girdle Hydrotherosaurus alexandrae www.PalArch.nl, vertebrate palaeontology, 4, 1, (2005) Remarks on the pectoral girdle of Hydrotherosaurus alexandrae (Plesiosauria: Elasmosauridae) S. Sachs Institut für Paläontologie Freie Universität Berlin, Malteser Strasse 74–100 Haus D,12249 Berlin, Germany [email protected] ISSN 1567–2158 3 figures Abstract The pectoral girdle of Hydrotherosaurus alexandrae Welles 1943, an elasmosaurid plesiosaur from the Upper Cretaceous (Maastrichtian) of California, USA, is redescribed. Some differences to the reconstruction presented in the original description, as well as newly discovered features of the pectoral girdle are discussed and a new reconstruction is provided. Contents 1. Introduction 2. Description 2.1. General comments 2.2. Scapula 2.3. Coracoid 3. Discussion 4. Conclusions 5. Acknowledgements 6. References Abbreviations AMNH American Museum of Natural History, New York, USA RMF Richmond Marine Fossil Museum, Richmond, Australia SDSM South Dakota School of Mines, Rapid City, USA UCMP University of California Museum of Paleontology, Berkeley, USA PalArch Foundation Sachs, Pectoral girdle Hydrotherosaurus alexandrae www.PalArch.nl, vertebrate palaeontology, 4, 1, (2005) 1. Introduction In his description of Hydrotherosaurus alexandrae, an elasmosaurid plesiosaur from the Upper Cretaceous (Maastrichtian, Moreno Formation) of Fresno County, California, Welles (1943) presented an unusual restoration of the pectoral girdle with asymmetrical coracoids. At the time, the about 8 m long, almost complete skeleton of the type specimen (UCMP 33912) had already been mounted at the University California Museum of Paleontology at Berkeley (UCMP). The exhibit was taken down in the 1960s and apparently has not been studied since. The skeleton was examined by the author during a visit to the UCMP in 2001 and some differences in the description and interpretation by Welles (1943) became apparent.
    [Show full text]
  • Morphologic and Ontogenetic Patterns in Elasmosaur Neck Length, with Comments on the Taxonomic Utility of Neck Length Variables F
    Marshall University Marshall Digital Scholar Biological Sciences Faculty Research Biological Sciences 6-2006 Morphologic and Ontogenetic Patterns in Elasmosaur Neck Length, with Comments on the Taxonomic Utility of Neck Length Variables F. Robin O’Keefe Marshall University, [email protected] Norton Hiller Follow this and additional works at: http://mds.marshall.edu/bio_sciences_faculty Part of the Aquaculture and Fisheries Commons, and the Ecology and Evolutionary Biology Commons Recommended Citation O’Keefe FR, Hiller N (2006) Morphologic and ontogenetic patterns in elasmosaur neck length, with comments on the taxonomic utility of neck length variables. Paludicola 5:206–229. This Article is brought to you for free and open access by the Biological Sciences at Marshall Digital Scholar. It has been accepted for inclusion in Biological Sciences Faculty Research by an authorized administrator of Marshall Digital Scholar. For more information, please contact [email protected], [email protected]. Paludicola 5(4):206-229 June 2006 by the Rochester Institute of Vertebrate Paleontology MORPHOLOGIC AND ONTOGENETIC PATTERNS IN ELASMOSAUR NECK LENGTH, WITH COMMENTS ON THE TAXONOMIC UTILITY OF NECK LENGTH VARIABLES F. Robin O'Keefe1 and Norton Hiller2 1Department of Anatomy, NYCOM 2 rm. 321, New York College of Osteopathic Medicine Old Westbury, New York 11568, [email protected] 2Canterbury Museum, Rolleston Avenue Christchurch, 8001 New Zealand, [email protected] ABSTRACT Elasmosaur cervical vertebrae are common fossils, but their taxonomic utility is limited due to a lack of understanding concerning their shape within and among taxa. In this paper, we analyze data from complete elasmosaur necks in an attempt to quantify and understand the variation in centrum dimensions.
    [Show full text]
  • Sea Monsters: a Prehiistoriic Adventure Summatiive Evalluatiion Report
    Sea Monsters: A Prehiistoriic Adventure Summatiive Evalluatiion Report Prepared for Natiionall Geographiic Ciinema Ventures By Valleriie Kniight-Wiilllliiams, Ed.D. Diivan Wiilllliiams Jr., J.D. Chriistiina Meyers, M.A. Ora Sraboyants, B.A. Wiith assiistance from: Stanlley Chan Eveen Chan Eva Wiilllliiams Daviid Tower Mason Bonner-Santos Knight-Williams Research Communications November 2008 This material is based on work supported by the National Science Foundation under Cooperative Agreement No. 0514981. Any opinions, findings, and conclusions or recommendations expressed in this material are those of the authors and do not necessarily reflect the views of the National Science Foundation. Knight-Williams Table of Contents CREDITS............................................................................................................................................................................................ 1 INTRODUCTION................................................................................................................................................................................ 2 THEATER CONTEXT ........................................................................................................................................................................ 4 METHOD............................................................................................................................................................................................ 8 SAMPLE INFORMATION...............................................................................................................................................................
    [Show full text]
  • Late Cretaceous) of Morocco : Palaeobiological and Behavioral Implications Remi Allemand
    Endocranial microtomographic study of marine reptiles (Plesiosauria and Mosasauroidea) from the Turonian (Late Cretaceous) of Morocco : palaeobiological and behavioral implications Remi Allemand To cite this version: Remi Allemand. Endocranial microtomographic study of marine reptiles (Plesiosauria and Mosasauroidea) from the Turonian (Late Cretaceous) of Morocco : palaeobiological and behavioral implications. Paleontology. Museum national d’histoire naturelle - MNHN PARIS, 2017. English. NNT : 2017MNHN0015. tel-02375321 HAL Id: tel-02375321 https://tel.archives-ouvertes.fr/tel-02375321 Submitted on 22 Nov 2019 HAL is a multi-disciplinary open access L’archive ouverte pluridisciplinaire HAL, est archive for the deposit and dissemination of sci- destinée au dépôt et à la diffusion de documents entific research documents, whether they are pub- scientifiques de niveau recherche, publiés ou non, lished or not. The documents may come from émanant des établissements d’enseignement et de teaching and research institutions in France or recherche français ou étrangers, des laboratoires abroad, or from public or private research centers. publics ou privés. MUSEUM NATIONAL D’HISTOIRE NATURELLE Ecole Doctorale Sciences de la Nature et de l’Homme – ED 227 Année 2017 N° attribué par la bibliothèque |_|_|_|_|_|_|_|_|_|_|_|_| THESE Pour obtenir le grade de DOCTEUR DU MUSEUM NATIONAL D’HISTOIRE NATURELLE Spécialité : Paléontologie Présentée et soutenue publiquement par Rémi ALLEMAND Le 21 novembre 2017 Etude microtomographique de l’endocrâne de reptiles marins (Plesiosauria et Mosasauroidea) du Turonien (Crétacé supérieur) du Maroc : implications paléobiologiques et comportementales Sous la direction de : Mme BARDET Nathalie, Directrice de Recherche CNRS et les co-directions de : Mme VINCENT Peggy, Chargée de Recherche CNRS et Mme HOUSSAYE Alexandra, Chargée de Recherche CNRS Composition du jury : M.
    [Show full text]
  • Reptilia: Sauropterygia)
    Records of the Westem Australian Museum Supplement No. 57: 201-205 (1999). Recent developments in Australasian sauropterygian palaeontology (Reptilia: Sauropterygia) Arthur R.I. Cruickshankl, R. Ewan Fordyce2 and John A. Long3 1 Earth Sciences Section, Leicester City Museum, New Walk, Leicester LE1 7EA, UK, and Geology Department, Leicester University, Leicester LII 7RH, UK; email: [email protected] 2 Department of Geology, Otago University, PO Box 56, Dunedin, New Zealand }Western Australian Museum, Francis Street, Perth, WA 6000, Australia Abstract -A brief review of recent research into Australasian sauropterygians is given. The earliest record is an Anisian pachypleurosaur from South Island, New Zealand. An Early Jurassic record from Queensland, hitherto included in the Plesiosauria is shown to be the last known pachypleurosaur. A diverse record of Pliosauroidea is known from the time of the break-up of East Gondwana, in marginal rift sediments of Western Australia and South Australia, dated to the earliest Cretaceous. The record of the marine incursion into the Eromanga Basin, Queensland, contains Albian-age pliosaurs and elasmosaurs. The New Zealand plesiosaurian record contains mostly members of the 'long-necked' Plesiosauroidea, all from the Haumurian stage of the Campanian-Maastrichtian. One specimen represents a unique record of a cryptoclidid in Australasia, more closely related to the European Callovian Cryptoclidus than to the other late cryptoclidids known from South America and the Antarctic Peninsula. INTRODUCTION NEW ZEALAND MATERIAL The sauropterygians were an entirely aquatic Sauropterygians have been known for over a group of reptiles which trace their ancestry to the century from the Haumurian Stage of the Late Permian tangasaurids of Madagascar, and Campanian-Maastrichtian in South Island, New whose intermediate evolutionary stages, the Zealand (Welles and Gregg 1971; Hiller et al.
    [Show full text]
  • O'keefe, F. R. and M. T. Carrano. 2005
    Paleobiology, 31(4), 2005, pp. 656±675 Correlated trends in the evolution of the plesiosaur locomotor system F. Robin O'Keefe and Matthew T. Carrano Abstract.ÐThis paper investigates trends in the evolution of body size and shape in the Plesiosau- ria, a diverse clade of Mesozoic marine reptiles. Using measures from well-preserved plesiosaur specimens, we document and interpret evolutionary patterns in relative head size, body size, and locomotor variables. Size increase is a signi®cant trend in the clade as a whole, and in constituent clades. The trend in relative head size is of variance increase; observed head sizes are both smaller and larger than ancestral values. In the locomotor system, changes in propodial and girdle pro- portions appear concomitant with body size increase and are interpreted as allometric responses to the physical constraints of large body size. Other trends in the locomotor system are signi®cantly correlated with both body size and relative head size. These locomotor trends evolved convergently in several clades of plesiosaurs, and may have had an ecomorphological basis, although data are lacking to constrain speculation on this point. The evolution of the locomotor system in plesiosaurs sheds new light on the response of aquatic tetrapods to the physical constraints of foraging at large body size. F.Robin O'Keefe. Department of Anatomy, NYCOM II, Room 326, New York College of Osteopathic Med- icine, New York Institute of Technology, Old Westbury, New York 11568. E-mail: [email protected] Matthew Carrano. Department of Paleobiology, Post Of®ce Box 37012, MRC 121, Smithsonian Institution, Washington, D.C.
    [Show full text]
  • The Macroevolutionary Landscape of Short-Necked Plesiosaurians Collapsed to a Unimodal Distribution
    www.nature.com/scientificreports OPEN The macroevolutionary landscape of short‑necked plesiosaurians Valentin Fischer1*, Jamie A. MacLaren1, Laura C. Soul2, Rebecca F. Bennion1,3, Patrick S. Druckenmiller4 & Roger B. J. Benson5 Throughout their evolution, tetrapods have repeatedly colonised a series of ecological niches in marine ecosystems, producing textbook examples of convergent evolution. However, this evolutionary phenomenon has typically been assessed qualitatively and in broad‑brush frameworks that imply simplistic macroevolutionary landscapes. We establish a protocol to visualize the density of trait space occupancy and thoroughly test for the existence of macroevolutionary landscapes. We apply this protocol to a new phenotypic dataset describing the morphology of short‑necked plesiosaurians, a major component of the Mesozoic marine food webs (ca. 201 to 66 Mya). Plesiosaurians evolved this body plan multiple times during their 135-million-year history, making them an ideal test case for the existence of macroevolutionary landscapes. We fnd ample evidence for a bimodal craniodental macroevolutionary landscape separating latirostrines from longirostrine taxa, providing the frst phylogenetically-explicit quantitative assessment of trophic diversity in extinct marine reptiles. This bimodal pattern was established as early as the Middle Jurassic and was maintained in evolutionary patterns of short‑necked plesiosaurians until a Late Cretaceous (Turonian) collapse to a unimodal landscape comprising longirostrine forms with novel morphologies. This study highlights the potential of severe environmental perturbations to profoundly alter the macroevolutionary dynamics of animals occupying the top of food chains. Amniotes are ’land vertebrates’, but have nevertheless undergone at least 69 independent evolutionary transi- tions from land into aquatic environments 1. Sea-going (marine) amniotes are textbook examples of inter- and intraclade convergent evolution, with repeated acquisitions of short, hydrodynamic body plans 2–9.
    [Show full text]
  • Revision of the Genus Styxosaurus and Relationships of the Late Cretaceous Elasmosaurids (Sauropterygia: Plesiosauria) of the Western Interior Seaway
    Marshall University Marshall Digital Scholar Theses, Dissertations and Capstones 2020 Revision of the Genus Styxosaurus and Relationships of the Late Cretaceous Elasmosaurids (Sauropterygia: Plesiosauria) of the Western Interior Seaway Elliott Armour Smith Follow this and additional works at: https://mds.marshall.edu/etd Part of the Biology Commons, Paleobiology Commons, and the Paleontology Commons REVISION OF THE GENUS STYXOSAURUS AND RELATIONSHIPS OF THE LATE CRETACEOUS ELASMOSAURIDS (SAUROPTERYGIA: PLESIOSAURIA) OF THE WESTERN INTERIOR SEAWAY A thesis submitted to the Graduate College of Marshall University In partial fulfillment of the requirements for the degree of Master of Science In Biological Sciences by Elliott Armour Smith Approved by Dr. F. Robin O’Keefe, Committee Chairperson Dr. Habiba Chirchir, Committee Member Dr. Herman Mays, Committee Member Marshall University May 2020 ii © 2020 Elliott Armour Smith ALL RIGHTS RESERVED iii DEDICATION Dedicated to my loving parents for supporting me on my journey as a scientist. iv ACKNOWLEDGEMENTS I would like to thank Dr. Robin O’Keefe for serving as my advisor, and for his constant mentorship and invaluable contributions to this manuscript. I would like to thank Dr. Herman Mays and Dr. Habiba Chirchir for serving on my committee and providing immensely valuable feedback on this manuscript and the ideas within. Thanks to the Marshall University Department of Biological Sciences for travel support. I would like to thank curators Ross Secord (University of Nebraska), Chris Beard (University of Kansas), Tylor Lyson (Denver Museum of Nature and Science), and Darrin Paginac (South Dakota School of Mines and Technology) for granting access to fossil specimens. Thanks to Joel Nielsen (University of Nebraska State Museum), Megan Sims (University of Kansas), Kristen MacKenzie (Denver Museum of Nature and Science) for facilitating access to fossil specimens.
    [Show full text]