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Naturalist

Volume 44 Number 2 Article 9

4-30-1984

Checklist of vascular of the Canyon and Church mountains

Sherel Goodrich USDA Forest Service, Intermountain Forest and Range Experiment Station, Ogden,

Follow this and additional works at: https://scholarsarchive.byu.edu/gbn

Recommended Citation Goodrich, Sherel (1984) "Checklist of vascular plants of the Canyon and Church mountains," Great Basin Naturalist: Vol. 44 : No. 2 , Article 9. Available at: https://scholarsarchive.byu.edu/gbn/vol44/iss2/9

This Article is brought to you for free and open access by the Western North American Naturalist Publications at BYU ScholarsArchive. It has been accepted for inclusion in Great Basin Naturalist by an authorized editor of BYU ScholarsArchive. For more information, please contact [email protected], [email protected]. CHECKLIST OF VASCULAR PLANTS OF THE CANYON AND CHURCH MOUNTAINS

Sherel Goodrich'

Abstract. A checklist of vascular plants of the Canyon and Church mountains, Juab and Millard counties, Utah,

is provided. Frequencv, distribution, and flowering time of taxa within the study area are indicated. Collection num- bers of voucher specimens are listed. Physiography, climate, and communities of the area are briefly described.

this study are deposited at New York Bot- People involved in rangeland and forest in- anical Garden, University of Utah, Utah State ventories studies are frequently faced with University, and Brigham Young University, at the need to identify plants. Plant checklists which institutions the above floras are being for rather small areas can be very helpful to written. these people or anyone who would like to the name of a plant. Local checklists know Study Area greatly reduce the number of choices possible in comparison to the numerous choices from The area included in this study is within manuals covering large areas. In the absence the Canyon and Church Mountains in Mil- of completed floristic treatments for the In- lard and Juab counties, Utah (Fig. 1). These termomitain Region, localized checklists are mountains are at the eastern edge of the often the main source of information about Great Basin. The Canyon Mountains are a the flora of an area. In addition, phytogeo- narrow, steep, fault-block range that is about graphy is of great interest to ecologists and 32 km (20 miles) long with a north-south axis. others. Checklists of specific mountain ranges Elevation varies from about 1463 m (4800 ft) provide a basis for many phytogeographic to 2962 m (9717 ft) at the top of Fool Peak. studies. The primary objective of this study is Much of the range is steep and rocky with to provide a plant list for the use of those massive rock outcrops forming cliffs and led- working in the Canyon Movmtains area. ges mainly of precambrian quartzite and Some other local checklists of the Inter- with some limestone. The Church Mountains moimtain area have been authored by Gra- are at the south end of Canyon Mountains ham 1937, Harris 1980, Lewis 1970, 1971, and consist of a few low hills with an east- 1973, Linsdale et al. 1952, MacMillian 1948, west axis. The boundary of the Fishlake Na Neese 1981, Preece 1950, and Welsh 1957. tional Forest encompasses the two ranges,

Work is currently imder way on three ma- and the National Forest Boundary was used jor floras (Intemiountain Flora, The Atlas of as a boundary for this study. Vascular Plants of Utah, and Utah Floral^) The climate is typically that of the Great that include the area of this study. These Basin, with cold moist winters and warm dry treatments are dependent upon plant speci- summers. Two weather stations (Oak City mens for descriptions and distribution data of and Scipio) with long-term records are adja- plant taxa. The secondary objective of this cent to the area. Average annual precipi- study was to provide specimens for these tation was 30.63 cm (12.06 inches) for 65 projects. Many specimens collected during

'USDA Forest Service, Intermountain Forest and Range Experiment Station, Ogden, Utah 84401; stationed at the Sciences Laboratory. Provo, Utah 84601. 1977). The Atlas 'These floras are in various stages of completion. Two of the six volumes of the Intermountain Flora are pubhshed (Cronquist et al. 1972, published separately or a few together (Welsh 1978, 1980, of Vascular Pliints of Utah is in preparation. Several plant families of the Utah Flora have been 1982. Welsh and Reveal 1977).

277 278 Great Basin Naturalist Vol. 44, No. 2

shorter frost-free period. The average frostfree period was 103 days for 1956 to 1979, or 41 days shorter than at Oak City. At Oak City, 12 May was the average day for the last freezing temperature in the spring for 1956 to 1979. This date at Scipio was 2 June, for the same period, or about 20 days later than at Oak City. Figure 3 shows the average monthly temperature for the two stations. Most of the lower elevations in the study area are out of the zone of cold-air drainage

in which Scipio is located, and the climate of

the area is probably more nearly like that at Oak City. The upper elevations likely have considerably more precipitation and colder temperatures.

Plant Communities

Pinyon-juniper

Pinyon pine (P/ni/5 edulis), singleleaf pin-

yon (P. monophyUa), and juniper (Juniperus osteospenna) communities cover the Church Mountains and lower hills of the Canyon Mountains. They extend up to near the high points of the Canyon Mountains on south ex- posures before being replaced by mountain Ijrush communities. On north exposures, they are replaced at much lower elevations. Juni- per dominates, and the pinyons are nearly lacking at lower elevations, but at upper ele- vations the pinyons are most abundant. The limits of pinyon and juniper seem controlled by cold temperatures of low-level inversions in the valleys and by increasing cold temper- atures with increasing elevation. The lower limits are likely also influenced by lower pre- cipitation. Reference to the pinyon-juniper

zone is frequently made in the text. This re- fers somewhat loosely to that elevational range at which either or both pinyon and juniper are common. This varies, roughly, at Fig. 1 Checklist collection area. the lower limit from 1585 to 1707 m

(5200-5600 ft) to an upper limit of about years at Oak City, and 31.34 cm (12.34 inch- 2438 m (8000 ft) on south exposures, and up es) for 76 years at Scipio. Figure 2 shows the to only about 1830 m (6000 ft) on north monthly distribution of precipitation at these exposures. two stations. The average frost-free period was 144 days Mountain Brush at Oak City for 1956 through 1979. Scipio is in a basin where cold air drainage is com- Mountain brush communities border the mon, and this station has a considerably upper edge of pinyon-juniper communities April 1984 Goodrich: Canyon and Church Mountain Plants 279 and follow canyons and drainages well down 4 r OAK CITY into the pinyon-juniper zone. They are domi- nated by one or more of the following plants: Oak {Querctis gambelii), curlleaf mountain mahogany {Cercocarpiis ledifolitis), true - 1.0 moimtain mahogany (C. rnontanus), greenleaf CM 2 - manzanita {Arctostaphylos pattila), and chokecherry {Prunus virginiana). Oak and 0.5 curlleaf moimtain mohagany are by far the most abimdant of these plants. Communities dominated by curlleaf mountain mahogany are typically found on shallow rocky soils and J FMAMJ J ASOND often on warm exposures, while those domi- nated by oak are on somewhat more favor- able soils and sometimes on cooler exposures. SCIPIO ni.5 Along canyon bottoms, oak often becomes tree like, and it is sometimes associated with bigtooth maple (Acer grandidentatwn). As re- 1.0 ferred to in the text, the mountain brush zone 2 is that area typically above the pinyon-juni- CM per zone where any of the above-listed plants are dominant.

Sagebrush J FMAMJJASOND

Wyoming big sagebrush {Artemisia triden- Fig. 2 Average monthly precipitation at Oak City and tata ssp. wtjomingensis) communities cover Scipio. extensive areas just beyond the boundary of this study area, such as in Little Valley on the slopes and in canyons over much of the east side of the Canyon Mountains, where middle and upper elevations of the area. most of this sagebrush was removed in an ex- Vasey sagebrush-grass communities are quite tensive burn in 1981. Soils of these commu- limited. They are mostly at the upper eleva- nities often have a hardpan that is high in tions in small areas of concave topography. carbonates. This pan typically occurs from 30 Black sagebrush forms communities on to 55 cm (12 to 22 inches) below the surface. convex topography with juniper, cliffrose Much of the area covered by big {Cowania mexicana), and bluebunch wheat- within the pin- sagebrush is below the pinyon-juniper zone. grass {Agropyron spicatum) An abundance of this sagebrush over a large yon-juniper zone. Occasionally it is common area probably indicates that average annual up in the mountain brush zone, but in this limestone. precipitation is between 20.32 and 25.40 cm zone it is usually restricted to (8-10 inches). Basin big sagebrush (A. tridentata ssp. tri- Vasey sagebrush (A. tridentata ssp. va- dentata) communities are restricted mostly to seyana) becomes abundant at about the deep alluvial soils along drainages at the lower edge of the pinyon-juniper zone and lower elevations of the area. extends to the highest elevations of the area. Low sagebrush forms communities of con- xeric grasses and With juniper, it covers extensive areas at the siderable size with various lower elevations of the pinyon-juniper zone forbs on convex, windswept slopes and ridges pinyon-jvmiper zone. where, it is most common on concave topog- in and above the raphy. On convex topography, where soils are more shallow, it is replaced by black Aspen sagebrush (A. nova). Vasey sagebrush forms communities with mountain brush Aspen [Populus tremuloides) is quite limit- and with low sagebrush (A. arbuscula) on ed in the Canyon mountains, and the stands a

280 Great Basin Naturalist Vol. 44, No. 2

Cystopteris fragilis, coynpacta, canum, nauseosus. Gera- nium parryi, Haplopappus watsonii, Hetero- theca villosa, Heuchera rubescens, Lepidium montanum var. heterophylltim, Macheran- thera kingii var. barnebyi, Pellaea breweri, Petrophytum caepitosurn, Polystichum sco- pulinum, debile, Selaginella watsonii, Sphaeromeria diversifolia, Solidago spar- siflora, and Trifolium andinum. Other plants that occur mostly in rocky places are: Allium brandegii, Echinocereus triglochidiatus, Epilo- J FMAMJ JASOND bium nevadense, brevicaule, Pet- Fig. 3 Average monthly temperature at Oak City radoria pumila, Potentilla glandulosa, Silene (lined) and Scipio (shaded). douglasii, and Xanthocephahwi petradoria. are usually rather small. These commimities are usually found along drainages and on Plants of Old Lake Deposits concave slopes, and often on cool exposures. above the Sevier River

Lacustrine deposits of old Lake Bonneville Fir above the Sevier River at the north edge of the area provide some of the most unusual White fir {Abies concolor), sometimes in as- habitat of the area. These deposits are saline, sociation with Douglas-fir (Pseudotsuga men- or alkaline and seleniferous in places. Several ziesii) forms closed communities on cool ex- plants of salt-desert shrub regions enter the posures at upper elevations of the Canyon area on these deposits, including: Atriplex Mountains. confertifolia, A. falcata, Kochia americana ,Sarcobatus vermiculatus, and Thelypodium Plants of Unique Habitats sagittatum. Other plants on these deposits, and apparently nowhere else in the area, are: Plants of Aeolian Sand Astragalus bisulcatus, A. calycosus, Hedysa- rum boreale, Hymenoxys cooperi, The following plants were found to be re- confusus, and Stanley pinnata. stricted, or nearly restricted, to wind-depos- ited sand: Abronia eUiptica, Ambrosia acanthicarpa, Amsinkia intermedia. Astra- New Tax galus ceramicus, Chrysothamnus nauseosus Macheranthera kingii var. barnebyi and ssp. turbinatus, Cryptantha circumscissa, C. Xanthocephalum petradoria have been de- confertiflora, Cyinopterus fendleri, Eriogo- scribed collections the Can- num keamey, Grayia spinosa, Lupinus brevi- from taken from this Collections caulis, densuni, Oenothera pallida, yon Mountains during study. Penstemon angustifolia, Prenanthella exigua, referred to here as Geranium parryi may be varietal those re- Psoralea lanceolata, Rhus trilobata, Streptan- worthy of distinction, and ferred to as Lesquerella occidentalis will thella longirosteris, Townsendia florifera, and Triterocalyx micranthus. likely be described as something new (pers. comm. from Reed C. Rollins, pro- fessor emeritus of systermatic botany, Har- Plants of Rocky Places vard University).

Much of the mid and upper elevations of the Canyon Mountains is composed of mas- Range Extension sive rock outcrops. The outcrops provide habitat for several specialized plants that tuberosa. Disjunct by about 150 grow mostly in rock crevices or on rock km (93 mi) northeast from near the Desert faces. Among these specialized plants are: Experimental Range in Beaver and Millard April 1984 Goodrich: Canyon and Church Mountain Plants 281 counties, and about 140 km (87 mi) south of a The Flora remarkable collection (B. Albee 2288) from Stansbury Island. A total of 65 families, 244 genera, 440 spe- CryptantJm cornpacta. Previously known cies, and 26 infraspecific taxa are listed. Of only from Western Millard County. the species, 20% are in . Poaceae, Cijmopterus acaulis. Collections from just , , , and Scro- south of Oak City apparently represent the phulariaceae each contain 10%, 8%, 6%, 5%, southernmost extent of the species in the and 4% of the taxa, respectively. Chenopo- Great Basin. These may represent a new diaceae, Polemoniaceae, , and variety. each contain about 3% of the Epilobium nevadense. Range extension of species. Apiaceae, Cyperaceae, and Ona- graceae each have about 2% and the remain- 230 km (143 mi) from Beaver Dam moun- ing 52 families each have less than 2%. tains of County. Penstemon leonardii. Range extention of 60 km (37 mi) from Mt. Nebo, Juab County. Literature Cited Polystichum scopulinwn. Third known lo- cation for Utah, previously reported for Salt Cronquist, A., A. H. Holmgren, N. H. Holmgren, and L. Reveal. 1972. Intermountain flora. Vol. 1. Lake and Washington counties (Cronquist et J. Hafner Publishing Co. Inc., New York. 270 pp. al. 1972). L. Cronquist, A., A. Holmgren, N. H. Holmgre.n, J. Trifolium andinitrn. Third known location Reveal, and P. K. Holmgren. 1977. Inter- for Utah, previously reported for north slope mountain flora. Vol. 6. Columbia Univ. Press, New York. 584 of the Uinta Mountains, Daggett County, and pp. Graham, E. H. 1937. Botanical studies in the Uinta Ba- (Welsh Navajo Mountain, Kane County sin of Utah and . Ann. Carnegie Mu- 1978), about 275 km (171 mi) from either of seum 26:1-4.32. Harris, G. 1980. A vascular flora of the San Rafael these locations. J. Swell, Utah. Unpublished thesis, Brigham Young Univ., Provo, Utah. 173 pp. The Checklist Lewis, M. E. 1970. Alpine rangelands of the Uinta Mountains. USDA Forest Service. Mimeo. 75 pp. plant communities of the The checklist is arranged alphabetically by 1971. Flora and major Rubv-East Humboldt Mountains. USDA Forest , , and species without regard for Service. Mimeo. 62 pp. mostly phylogenetic order. Nomenclature 1973. Wheeler Peak area species list. USDA For- follows that of Welsh et al. (1981). The list is est Service. Mimeo. 17 pp. Linsdale, M. a., T. Howell, and M. Linsdale. primarily based on collections and observa- J. J. 1952. Plants of the Toiyabe Mountains area, Ne- tions that I made during 1980, 1981, and Biol.' 10; 129-200. vada. Wasmann J. 1982. A very limited herbarium search was McMillan, C. 1948. A taxonomic and ecological study made to find a few expected taxa for which I of the Deep Creek Mountains of central western had no records. Unless otherwise designated, Utah. Unpublished thesis. Univ. of Utah, Salt Lake City, Utah. 99 pp. the collections numbers are mine. Taxa for 1981. va.scular flora of the Henry Moun- Neese, E. J. A are given were which no collection numbers tains, Utah. 370 pp. S. 1950. Floristic and ecological features observed but not collected. The most com- Ptrrvr, J., Jr. of the Raft River Mountains of Northwestern plete set of specimens are deposited at Brig- Utah. Unpublished thesis. Univ. of Utah, Salt ham Young University, Provo, Utah (BRY). Lake City, Utah. 86 pp. Many other specimens were also deposited Welsh, S. L. 1957. An ecological survey of the vegeta- at: Intermountain Forest and Range Experi- tion of the Dinosaur National Monument, Utah. thesis. Brigham Young Univ. Provo, ment Station, Shrub Sciences Laboratory, Unpublished Utah. 86 Provo, Utah (SSLP); New York Botanical pp. 1978. Utah flora: Fabaceae (Leguminosae). Great Garden, New York, New York (NY); Region Basin Nat. 38:225-367. Four, Forest Service, Ogden, Utah (OGDF); 1980. Utah Flora: Malvaceae. Great Basin Nat. Supervisors Office, Fishlake National Forest, 40:27-37. 1980. Utah flora: Miscellaneous families. Great Richfield, Utah; and at Garrett Herbarium, Basin Nat. 40:38-57. University of Utah, Salt Lake City, Utah 1982. Utah flora: Rosaceae. Great Basin Nat. (UT). 42:1-44. 282 Great Basin Naturalist Vol. 44, No. 2

L., 1977. Utah flora: Brassi- plants of the Moimtains are transitional Welsh, S. and J. L. Revel. Some Canyon caceaae (Cruciferae). Great Basin Nat. from var. grayi to var. depaupcratiim (Jones) Mathias. 37:279-.3&4. Lomatium macrocarpum (H. & A.) Coult. & Rose. In-

Welsh, S. L., N. D. .'Vtwood, S. Goodrich, E. Neese, frequent to occasional in sagebrush-juniper, pinyon-juni- K. H. Thorne, and B. Albee. 1981. Preliminary per, and mountain brush communities. April-June. index of Utah names. Great Basin 14082, 14208, 15286, 15.5.50, 15632. Nat. 41:1-108. Orogenia linearifolia Wats. Infrequent to locally common in various plant communities above the pin- yon-juniper zone. May-June (July at the edge of melting ' Species List snowbanks). 14.3,54A,^5394; Welsh 164.32. Osmorhiza chilensis H. & A. Occasional to locally Aceraceae common in aspen, fir, and oak-maple communities on canyon bottons. May-June. 14326. 15586. Acer glabrum Torr. Occasional on slopes, mostly Osmorhiza occidentalis (Nutt.) Torr. Locally common within moiuitain brush and aspen communities. in mesic mountain brush and aspen communities. May-June. 14359, 16843. June-July. 14314, 1.5800. Acer grandidentatum Nutt. in T. & G. Occasional to locally common along major drainages. May-June. .\p6cynaceae 10427, 15317, 17360; Welsh 16424.

Apocynum androsaemifolium L. Locally common in Amaranthaceae Oak Creek Canyon, apparently rather rare elsewhere, mountain brush communities. Jidy-.\ugust. 15791, Amaranthtis albus L. Seen at the mouth of Oak 17312; Foster 6552. Creek Canvon in a hurnedover cottonvvood--oak community. Jime-October. 17320. Asclepiadaceae Amaranthtis graecizans L. Introduced from the Old World, adventive with disturbance, an early succession Asclepias aspertda (Dene.) Woodson. Infrequent in plant on chared ground where slash piles from pinyon- sagebrush and pinyon-juniper communities. June-July. juniper chainings have been burned. June-July. 14111. 14282. Asclepias speciosa Torr. Seen at a single location in a chainedover pinvon-juniper woodland between Church .\nacardiaceae and Canvon mountains. fune-Julv. 14133. Rhus glabra L. Infrequent along ephemeral drainages of lower Eightmile Creek in the pinyon-juniper zone. .Asteraceae June-July. 14114, 14239. Rhus trilobata Nutt. in T. & G. Locally common to L. Occasional in various plant abundant on aeolian sand in and below the pinyon-juni- communities over a wide elevational range. per zone, infrequent elsewhere. May-June. 13779, June-September. 14297; Fo.ster 6.534. 14240. Agoseris aurantiaca (Hook.) Greene. Seen only at East Fork Eightmile Canyon, with oak and sagebrush. June. 14010. Apiaceae Agoseris glauca (Pursh) Raf. Common in several plant

Cymopterus acatdis (Pursh) Raf. Infrefjuent on aeo- communities over a wide elevational range. Mav-June. lian sand along the west edge of the area in pinyon-juni- 13792, 13803, 13928. per communities. April-May. 1.5279, 1531 1', 15313, Ambrosia acanthicarpa Hook. Locally common to 15404. abundant on sandy ground, mostly in and below the pin- Cymopterus purpurascens (Gray) Jones. Infrequent yon-juniper zone. August-September. 16391. T. to on sandy-clay deposits of old Lake Bonneville above the Antennaria dimorpha (Nutt.) & G. Occasional locally in sagebrush com- Sevier River at the extreme north limit of the area, rare common pinvon-jimiper and 15.380. in the pinyon-juniper zone elsewhere in the area. munities. Late April-June. 13768A, 13786, locally March-May. 138.38, 13852, 1,5272; Welsh 16420. Aritennaria microphylla Rydb. Occasional and Lomatium dissectum (Nutt.) Math. & Const, var. ea- common in various plant communities. Mav-July. 14005. tonii (Coult. & Rcse) Gronq. Infrequent in oak and oak- Antennaria parvifolia Nutt. Seen only at the head of maple-white fir commimities. Mav-June. 14199, 15324, 15605. Dry Creek in a snowbank area adjacent to a fir commu- Lomatium foeniculaceum (Nutt.) Coult. & Rose var. nity. June-July. 17237. macdougalii (Coult. & Rcse) Cronq. Occasional to com- cordifolia Hook. Occasional in white fir and mon in sagebrush-juniper, and low sagebrush commu- Douglas-fir communities. June. 15612, 15617. nities. Mav-June. 13781, 13847, 1.3894, 13902, 13909, Artemisia arhuscula Nutt. .\bundant to dominant on 1.39.32, 13933, 15378; Welsh 14605. exposed slopes and ridges at the upper edge of and Lomatium grayi Coult. & Rose. Occasional to com- above the pinvon-jiuiiper zone. .\ugust-September. mon in pinyon-juniper, sagebmsh, and mountain brush 15130, 15254, 16457, 16466. commimities, mostly on quartzite, but also on limestone. Artemisia dracunculus L. Collected at the mouth of May-June. 13976, 14020, 14365, 15287, 15325, 15391, Lyman Canyon in Oak Creek. Sagebrush community. 15553, 1.5559; Foster 6538; Welsh 16.397, 16410, 16598. August-October. 17961. April 1984 Goodrich: Canyon and Church Mountain Plants 283

Artemisia ludoviciana Nutt. var. incompta (Nutt.) along drainages in sagebrush and pinvon-juniper com- Cronq. Occasional at the upper edge and above the pin- munities. August-October. 16.381, 16.382, 16385. yon-juniper zone, usually in rocky places. Au- Chrysothamnus nauseosus (Pall.) Britt. ssp. turbi- 15137,' gust-October. 15136, 16461. natus (Jones) H. & C. Abundant on aeolian sand in and Artemisia ludoviciana Nutt. var. ludoviciana. Occa- below the pinvon-juniper zone along the west side of the sional along drainages and in rocky places mostly in the area, mostlv outside the boundary. August-September. pinvon-juniper zone. .'\ugnst-October. 1490, 15245. 16386, 17963. Artemisia nova A. Nels. Abundant to dominant on Chrysothamnus viscidiflorus (Hook.) Nutt. ssp. pub- shallow rocky .soils on slopes and ridges in and below the erulus (D.C. Eaton) H.& C. Common to abundant in pinvon-juniper zone, and occasionally extending to the pinvon-juniper and sagebrush communities, mostly on upper edge and perhaps above this zone, especially on slopes and ridges. Juiy-August. 14306, 14947, 15149, limestone. September-October. 15148, 15233, 15292, 15234. 16467. Chrysothamnus viscidiflorus (Hook.) Nutt. ssp. vis- Artemisia tridentata Nutt. ssp. tridcntata. .\bundant cidiflorus. Occasional at the south end of the area and elsewhere in juniper and vasey sagebrush to dominant on deep alluvial soils of drainage ways in perhaps and below the pinvon-juniper zone. Septem- communities. neomexicanum Gray. Occasional to locally ber-October. 16380. common in sagebrush and juniper communities. Artemisia tridetitata Nutt. ssp. vaseyana Beetle. June-August. 16876, 17133. Common to abundant over much of the area near the Cirsium vulgare (Savi) Ten. Introduced from the Old lower fringe of the juniper zone, to the crest of canyon World, adventive on disturbed ground, many plant com- range, in many plant commimities; most common and nnmities. Jime-Julv. Seen but not collected. sometimes the dominant plant on concave positions of Cirsium wheeleri (Gray) Petrak. Occasional in Gam- slopes. September-October. 16454, 16455, 16458. bel oak communities. July. 15792, 17311. Artemisiac tridentata Nutt. ssp. wyomingensis Beetle Conza canadensis (L.) Cronq. Seen at the mouth of & Young. Common to dominant on soils with a hardpan Oak Creek Canyon and upper Lyman Canyon in of from about 30 to 55 to 22 inches) some tvpe cm (12 burnedover cottonwood-willow-oak and mountain brush surface, mostly below the pinyon-juniper zone below the communities. May-September. 17325, 17952. most bevond the boiuidary of the area, and common Crepis acuminata Nutt. Occasional in Vasey sage- such as in Little Vallev. September. brush, pinyon-juniper, and mountain brush commu- Nels. Macbr. Occasional to very Aster perelegans & nities. June'. 14036, 14083, 15634. locally common in mountainbrush communities. Crepis modocensislo Greene. Infrequent in mountain July-September. 14924, 17354, 17953, 17957. brush and low sagebrush communities. Jinie-July. 14011, Aster wasatchensis (Jones) Blake. Seen at only a few 14365. stations among rocks near the crest of the range. .Au- Crepis occidentalis Nutt. Occasional in pinyon-juni- gust-September. 15135. per and mountain brush communities. June. 13925, Balsamorhiza saggittata (Pursh) Nutt. Abundant on 14037, 14132, 14172. open slopes, with big sagebrush and Gambel oak, domi- Erigeron aphanactis (Gray) Greene. Collected at a nant understory plant in curlleaf mountain mahogany single station at the north end of the Canyon Mountains. communities, and sometimes in other communities in Common just east of the area. June. 14045. and above the pinvon-juniper zone. Late May-June. Erigeron argentatus Greene. Seen at the divide be- 14034, 15381. tween Oak and Little Oak Creeks on rocky ridge with adjacent to the Brickellia californica (T. & G.) Gray. Seen on slopes curlleaf mountain mahogany; also seen above the Sevier above Wide Canyon, southeast exposure of outcrop of area on deposits of old Lake Bonneville Precambrian quartzite, moinitain brush community. River in Mills Valley. 14331. T. &. G. Occasional in oak and 16462. Erigeron divergens sagebrush communities, often along canyon bottoms, Brickellia grandiflora (Hook.) Nutt. Occasional to lo- and perhaps adventive on distiubed ground. June-July. cally connnon in rocky places in mountain brush com- 14017, 14432, 15239, 15820, 16874; Foster 6535. nuinities. Julv-September. 14430, 14926. Erigeron eatonii Gray. Occasional in various plant Brickellia microphylla (Nutt.) Gray. Seen only at a communities above the pinvon-juniper zone. 14029, single station near the southern limit of the area in 14356, 15373, 15556, 15608, 17238; Foster 6529. cracks of rocks within a black sagebrush-bluebunch Erigeron engelmanii A. Nels. Occasional in juniper- wheatgrass connnunitv. 15232. Julv-September. sagebrush communities. Mav-June. 13770, 13794, 13818, A. Occasional in Chaenactis douglasii (Hook.) H. & 1.3846, 14109, 16824. many plant communities throughout the area. Erigeron flagellaris Gray. Occasional along canyon May-June. 13901, 13916, 13938, 13951, 14084. bottoms with narrowleaf cottonwood, oak, and maple. Chrysothamnus greenei (Gray) Greene. Collected June-July. 14291, 14295; Welsh et al. 17439. from adjacent to the area in the Clay Springs drainage Erigeron nauseosus (Jones) Cronq. Occasional in from aeolian sand. August-September. 15249. rockv places, often on faces of rock outcrops, usually in Chrysothamnus nauseosus (Pallus) Britt. ssp. holo- mountain brush communities. June-August and to Octo- 15596, leucus (Gray) H. & C. Apparently uncommon in moun- ber, with sufficient fall moisture. 15242, 15589, Foster 6531. tain bnisii and Vasev sagebrush comnnmities. 16459, 15626, 15795, 16083, 16465; Occasional to locally 17960. Erigeron speciosus (Lindl.) DC. common in mountain brush and aspen communities. Chrysothamnus nauseosus (Pall.) Britt. ssp. graic- Julv-August. 14923, 17353, 17955. olens (Nutt.) Piper. Common to abundant, especially 284 Great Basin Naturalist Vol. 44, No. 2

Gnaphaliiim palustre Nutt. Seen only at East Fork Macheranthera kingii (D.C. Eaton) Cronq. & Keck Eiglitniile Creek alont; canyon bottom with oak, narrow- var. harnebyi Welsh & Goodrich. Infrequent to locally Cottonwood, and white fir. July. 14448. common in crevices of cliffs and ledges of precambrian Haplopappus acaulis (Nutt.) Gray. Occasional to lo- quartzite from mountain brush zone to the crest of the cally common rocky exposed slopes and ridges in pin- Canyon Mountains. August-September. 14929, 14933, yon-juniper and mountain brush and low sagebrush 15133, 15134. communities; perhaps most common in the northern Malacothrix torreyi Gray. Infrecjuent in juniper-sage- half of the Canvon Moimtains. May-June. 15366, 16841. brush coiumimities. June. 14149. Haplopappus armerioides (Nutt.) Gray. Seen only Microseris nutans (Geyer) Schultz-Bip. Occasional to near or along the divide between Weetches Canyon and locally common in oak communities. June. 14012. Dry Canyon among rocks and ledges in mountain brush Perityle stanshuryi (Gray) Macbr. Seen only near the and white fir-Douglas-fir communities. June-July. northern limit of the area on a limestone outcrop with 15796, 15810. scattered jvmiper and black sagebrush. June. 15638. Haplopappus watsonii Gray. Widespread and com- Petradoria pumila (Nutt.) Greene. Common to abun- mon throughout the Canyon Mountains in cracks of rock dant on rocky exposed slopes and ridges in low sage- outcrops at the upper edge of the pinyon-jimiper zone brush, pinyon-juniper, and mountain brush commu- to near the crest of the mountains. August-October. In- nities; infrequent in nonrocky areas. June-August.

tergrading into H. njdbergii Blake in Tidestr. 14930, 14100, 14182, 14.321, 14324; Welsh et al. 17451. 15132, 15140, 15236, 15241, 15244, 15251, 16081, 16460, Senecio douglasii DC. A single large plant found 16464, 17951. along a roadside at the south end of the Canyon Moun- Helenthella uniflora (Nutt.) T. & G. Occasional in tains. Julv-September. 17364. mountain bnish communities. June-August. 14431. Senecio integerrimus Nutt. Occasional to locally com- Helianthus annus L. Adventive on disturbed groimd mon in Vasev sagebrush and moimtain brush commu- usually in and below the pinvon-juniper zone. 15640, nities. May-June. 15384. 16383. Senecio multilobatus T.& G. ex Gray. Occasional in Heliomeris multiflora Nutt. Occasional in rocky many plant comnumities over a wide elevational range. places in mountain brush communities. .August- May- August. 13981, 14116, 15692, 16840. September. 14931, 15824. Senecio streptanthifolius Greene. Seen on Blue and in Hardscrabble Canyon, pinyon-juniper Hymenopappus filifoliits Hook. Occasional in pinyon- Moimtain communities. 15555, 168408. juniper communities, infrequent in dry mountain brush and mountain brush June. communities. Late Mav-June. 13919, 13950, 14098, Solidago sparsiflora Gray. Occasional to locally com- Au- 14180; Foster 6544; Welsh 17025. mon on rock outcrops in the moimtain brush zone. 14437, 14925, 14934, 15138, 15237, Htjmenoxtjs acaulis (Pursh) Parker. Occasional in gust-September. 17950. juniper conuuunities, locally common in low sagebrush 16085, 17359, the Old communities. May-June. 13947, 15397. Sonchus asper (L.) Hill. Introduced from at Eightmile Hymenoxys cooperi (Gray) Cockerell. Occasional on World, more or less weedy; seen only once 16875. old Lake Bonneville deposits of sandy clay above the Se- Ditch, disturbed ground. June-September. Eaton) Rydb. Occa- vier River at the extreme northern limit of the area. Sphaeromeria diversifolia (D.C. in crevices of ledges cliff, May-June. 13835. sional to locally common and usually the pinyon-juniper zone. Jidy-September. Iva axillaris Pursh. Infrequent in various plant com- above 14322', 14935, 15139, 16082. munities, usuallv in and below the pinvon-jimiper zone. 14349. Stephanortieria exigua Nutt. Infrequent or locally June-July. 14088. Layia ^landidosa (Hook.) H.& A. Seen only on sandy common on sandy ground (often stabilized dunes) in the zone. 14223, 14232; Foster 6549. ground in the Clay Springs Wash in juniper-sagebrush pinyon-jimiper June. communities. May-June. 14159, 15309, 15566. Taraxacum officinale Weber in Wiggers. Introduced usu- serriola L. Introduced from the Old World, froiu the Old World, adventive on disturbed ground zone. adventive on disturbed ground and locally common on allv in and below the pinyon-juniper charred ground where piles of pinyon and juniper slash May-October. 13905. sagebrush, have been b\inied after chaining. Jime-July. Tetradymia canescens DC. Occasional in communities. Leucelene ericoides (Torr.) Greene. Occasional to lo- pinvon-juniper, and dry mountain brush cally common in sagebrush-juniper conuuunities. June. June-August. 14948. 14039. Townsendia florifer (Hook.) Gray. Seen at two sta- tions in sand. Lygodestnia grandiflora (Nutt.) T. & G. Occasional in Clav Springs Wash on aeolian Mav-June. sagebrush-juniper communities, usually on .sandy soil; 14151, 14226. the .seen onlv at the southwest part of the area. May-June. Tragopogon dubius Scop. Introduced from Old 14127; Fo.ster 6550. World, adventive on disturbed ground and in various in- Macheranthera canescens (Pursh) Gray. Occasional in digenous plant communities. June. 13963. manv plant communities. August-October. 14938, Wyethia amplexicaulis (Nutt.) Nutt. Seen only in 15129, 15243, 16389, 17956. Fool Creek drainage where locally abundant in sage- Macheranthera grindelioides (Nutt.) Shinners. Seen at brush-mountain brush communities. May-June. 15623. the divide between Oak and Little Oak creeks, on rocky Xanthocephalum strumarium L. Collected along ridge with curlleaf moimtain mahogany; common adja- Highway 132 at the north edge of the area. Au- cent to the area on old river deposits above the Sevier gust-September. 17964. River on hills between Little Vallev and Mills Valley. Xanthocephalum petradoria Welsh & Goodrich. Lo- June-July. 14330. cally common on rocky slopes and ridges, mostly in the April 1984 Goodrich: Canyon and Church Mountain Plants 285

mountain bnish zone. August-October. 14928, 15127, Brassicaceae 151:31, 15145, 15240, 15256, 15257, 16080. Xanthocephalum sarothrae (Pursh) Shinners. Com- Alyssum alyssioides L. Introduced from , mon to abundant in pinyon-juniper comnnmities espe- somewhat weedy and perpetuated by disturbance, in cially on shallow soils; infrequent to occasional in many and below the pinyon-juniper zone. May-June. 14206. other plant comminiities; adventive on disturbed Alyssum desertorum Stapf. Introdviced from the Old ground. August-October. 14943, 15128. World, somewhat weedy and perpetuated by disturb- ance, mostly in and below the pinyon-juniper zone. Berberidaceae May-June. 1.3911. Arahis drummondii Gray. Infrequent in various plant Mahonia repens G. Don. Occasional in mountain communities, mostly in and above the oak zone.

iinish and aspen communities. April-May. 13725; Welsh May-June. 14362, 15.385; Welsh et al. 17041, 17442. 16401. Arabis glabra (L.) Bernh. Collected only from Oak Creek, cottonwood-oak-maple community. May-July. 17.363. Arabis holboellii Hornem. Occasional in various plant Seen only at Amsinckia intermedia Fisch. & Mey. communities over a wide elevational range. May-June. two stations on aeolian sand along the south and west 13903, 15365; Welsh 16593; Welsh et al. 17440. sides of the Canvon Mountains. May-June. 15310; Arabis microphylla Nutt. Seen at a single station, in Welsh 16590. cracks of rocks in Oak Creek Canyon. May-June. 15319. Cryptantha compacta Higgins. Occasional in rocky Arabis pendulina Greene. Occasional and locally places along the crest of the Canyon Mountains. common in cracks of rock outcrops and on windswept June-July. 14364, 15554, 15607. ridges in low sagebrush communities, mostly above 2285 Cryptantha circumscissa (H. & A.) Johnst. Infrequent m (7500 ft) elevation. May-June. 14.363, 15395, 15552, or occasional on aeolian sand; seen only at the south and 15591. west edge of the area in and below the pinyon-jimiper Arabis perennans Wats. Occasional in various plant zone. May-June. 14118, 14152. communities, often in rocky places. May-June. 13840, Cryptantha confertiflora (Greene) Payson. Seen at 14019, 15318, 15599; Welsh 16400. only one location at the extreme western limit of the Arabis pulehra Nutt. in T. & G. Infrequent or occa- area on aeolian sand, in a juniper community. June. sional in sagebrush and pinyon-juniper communities. 15314. Plants of the area are referable to var. miin- Cryptantha flavoculata (A. Nels.) Payson. Occasional May-June. ciensis Jones. 13789, 13801, 138.32. to common in upper pinyon-juniper and movmtain brush Andrz. Introduced from the communities, particularly with curlleaf mountain ma- Camelina microcarpa World, adventive on disturbed ground, mostly in hogany. May-June. 14166, 15362, 15560. Old and below the pinvon-juniper zone. 1.3913; Welsh 16602. Cryptantha gracilis Osterh. Occasional to frequent in (L.) Medic. Introduced from sagebnish and pinvon-juniper communities. May-June. Caspella bursa-pastoris the World, adventive on disturbed ground, mostly 13895, 14157, 15689. Old in below pinyon-juniper zone. May-June. 14440, Cryptantha humilis (Gray) Payson. Occasional to and the 15296; Welsh 16429.' common in sagebnish and pinyon-juniper conununities. Cardaria draba (L.) Desv. Introduced from the Old May-June. 13767, 13836, 1.394,5, 15347; Welsh 16417. adventive, seen only at a few locations in the Cryptantha kelseyana Greene. Occasional to com- World, area. 15354. mon, sandy ground in the vicinity of Clay Springs Wash. June. Chorispora tenella (Pallas) DC. Introduced from the May-June^ 14153, 14224. World, adventive on disturbed ground, usually in Cryptantha pterocarya (Torr.) Greene. Occasional to Old and below the pinyon-juniper zone. May-June. 15297; common in sagebnish and pinyon-juniper communities. Welsh 16428. May-June. 13842, 14119. californica (Gray) Schulz. Infrequent in Cryptantha watsonii (Gray) Greene. Occasional in fir communities. June-August. 17.352. pinyon-juniper, mountain brush, and sagebrush commu- aspen and Descurainia pinnata (Walt.) Britt. Adventive on dis- nities. May-June. 158.33. turbed ground, in various plant communities. 13793, Cynoglossum officinale L. Adventive on disturbed 13827, 13910. 13920; Welsh 16588. ground, in various plant communities. June-July. 14215. Descurainia sophia (L.) Webb in Engler & Prantl. In- Hackelia patens (Nutt.) Johnst. Seen only at a canyon from the Old World, adventive on disturbed on south side of Leamington Canyon, pinyon-juniper troduced mostly in and below the pinyon-juniper zone. commimity. May-June. 15361. ground, 13828. Lappula occidentalis (Wats.) Greene. Occasional in Nutt. ex T. & G. Infrequent to oc- various plant communities. May-June. 14124. Draba cuneifolia casional in pinyon-juniper communities. .\pril-June. incistim Lehm. Occasional to common 13914, 1.3974, 14131, 15278, 15298, 15678. in sagebnish and juniper communities, perhaps most nemorosa L. Occasional in various plant com- common in Church Mountains. May-June. 13766, Draba usually above the pinyon-juniper zone. 13787, 1.3897. Lithospermum ruderale Dougl. ex munities 14110, 1.5320, 15399; Welsh 16414. Lehm. Infrequent in various plant communities over a April-June. 13996, Fern. With the distribution of rather wide elevational range. June-July. 15625. Draba reptans (Lam.) the upper part of stems Mertensia oblongifolia (Nutt.) G. Don. Occasional in D. cuniefolia and, except that upper pinyon-juniper and mountain bnish communities. and the pedicels glabrous, not different from D. cunei- May-June. 14170, 15364, 15614. folia April-June. 13968A, 14207, 15298. 286 Great Basin Naturalist Vol. 44, No. 2

Erysismiim aspenim (Nutt.) DC. Occasional in many Capparidaceae plant coininunities over a wide elevational range. May-June. 1.3769, 13791, 13817; Welsh 16421. Cleome serrulata Pursh. Collected from along High- Ertjsismtim repandttm L. Introduced from the Old way 132 at the north edge of the area. Au- World, adventive on disturbed ground, usually in and gust-September. 17967. below the pinvon-juniper zone. 13810, 13829; Welsh lfv412. Caprifoliaceae Lepidium densiflorum Schrad. Seen at Eightmile Creek, juniper community, along an ephemeral drain- Sambucus caertdea Raf. Infrequent in pinyon-juniper age. May-June. 14138. communities. June-Julv. 14281; Foster 6551. Lepidium lasiocarpum Nutt. in T. & G. The one Symphoricarpos oreophilus Gray. Occasional to com- specimen seen (Welsh 16596) is from a pinyon-juniper mon in Vasey sagebrush, aspen, and fir communities. community at Leamington Pass. May-June. June-July. 14200, 1.5809. Lepidium montanum Nutt. var. heterophyUum (Wats.) C. L. Hitchc. Seen only at Lyman Canyon, in Caryophyllaceae cracks of rock outcrops, mountain brush community. June-July. 14316. Arenaria kingii (Wats.) Jones. Occasional to frequent Lepidium montanum Nutt. var. montanum Occasion- in pinyon-juniper, sagebrush, and mountain brush com- al in sagebrush and pinyon-jvmiper commimities. munities. May-June. 139.34, 15.346. May-June. 13827, 14095. Arenaria macradenia Wats. Locally common to Lepidium perfoliatum L. Introduced from the Old abundant in mountain brush communities, often in World, adventive, mostly in and below the pinyon-juni- rocky places. June- August. 14173, 14320, 15806, 15831; per zone. May-June. 13830. Welsh et al. 17450. Lesquerella occidentalis (Wats.) Wats. Locally com- Lychnis drummondii (Hook.) Wats. Seen only in a mon in upper Lyman Canyon on a massive quartzite Vasev sagebrush community at the top of the Canyon 17949.' outcrop. June. 16883, Mountains. July. 14.334. Malcolmia africana (L.) R. Br. in Ait. Introduced Silene antirrhina (L.) Desv. Seen in a juniper-black from ., adventive on disturbed ground, usually in sagebrush community among rocks. May-June. 1535.5; and below the pinyon-juniper zone. 13957A, June. Welsh et al. 17449a. 14136. Silene douglasii Hook. Infrequent in rocky places in Physaria chambersii Rollins. Occasional to common mountain brush and white-fir communities. Late in sagebnish, pinyon-juniper, and dry mountain brush June-July. 14424, 15794, 17351. communities, extending to near the crest of the Canyon SteUaria jamesiana Torr. Occasional to frequent in Mountains on limestone. May-June. 137982, 13819, mountain brush, aspen, and fir connnunities. May-July. 1.3936, 14097, 15396; Foster 66.58. 13985. Sisymbrium altissimum L. Introduced from the Old SteUaria media (L.) Cyrill. Introduced from the Old World, adventive on disturbed ground, usually in and World, seen only in Lyman Canyon, wet ground along a below the pinyon-juniper zone. June-July. 14125. small stream in fir-snowberry community. Stanleya pinnata (Pursh) Britton. Infrequent on de- May-September. 17942. posits of Lake Bonneville, above the Sevier River at the extreme northern limit of the area, on seleniferous, alka- Celastraceae line, sandy clay soil. Mav-June. 14046. Streptanthella longirostris (Wats.) Rydb. Occasional Forsellesia nevadensis (Gray) Greene. Infrequent in on aeolian sand in sagebnish and juniper communities. rocky places in the pinyon-juniper zone. 15359. June. 14229. Pachystima myrsinites (Pursh) Raf. Occasional in Streptanthus cordatiis Nutt. ex T. & G. Occasional to mountain brush, aspen, and fir communities. May-June. frequent in sagebnish and pinyon-juniper communities, 13724, 14032, 14169, 16845. infrequent in mountain bnish communities. May-Jiuie. 13788, 13849, 13939, 13984, 15600; Welsh 16603.' Thelypodium sagittatum (Nutt.) Endl. Occasional on Chenopodiaceae sandy clay deposits of old Lake Bonneville above the Se- Atriplex confertifolia (Torr. & Frem.) Wats. Occa- vier River at the north limit of the area. Mav-Jiuie. sional on Lake Bonneville deposits above the Sevier Riv- 15353. er at the north edge of the area. Mav-Jime. Thlaspi arvensis L. Introduced from the Old World, Atriplex falcata (Jones) Standi. Occasional on Lake adventive on disturbed areas. Mav-June. 15572. Bonneville deposits of .sandy clay above the Sevier River at the north limit of the area. Jime. 14048. Cactaceae Chenopodium album L. Introduced from the Old World, adventive disturbed ground, usually in and Echinocereus triglochidiatus Engelm. Occasional in on rocky places, usually within the pinyon-juniper zone. below the pinyon-juniper zone. July-.4ugust. May-June. 1.5.363; Foster 6542; Welsh 16600. Chenopodium botrys L. Introduced from the Old World, in area Opuntia polyacantha Haw. Occasional in various adventive on disturbed ground, seen the plant connnunities except mesic mountain brush, aspen, on a recent rangeland seeding where juniper had been chained. 14946. and fir. June 17134; Foster 6572. July-.-\ugust. April 1984 Goodrich: Canyon and Church Mountain Plants 287

Chenopodium capitatum (L.) Asch. Adventive dis- on Carex douglasii F. Boott in Hook. Occasional, mostly turbed ground in various plant communities over a wide along roads in the pinvon-juniper zone. May-June. elevational range. June-Julv. 13993, 14347, 15827, 138.33. 17347. Carex getjeri F. Boott. Occasional to dominant under- Gratjia spinosa (Hook.) Moq. in DC. Seen onlv at the story plant in oak, aspen, and fir communities. northern limit of the area on aeolian sand. Mav-June. May-June. 1.3723, 14371, 15374, 15616. 1.3851. Carex hoodii F. Boott in Hook. Rare, moist places, Halogeton glomeratus C. A. Meyer in Ledeb. In- mountain brush communities. 17232. 17358. troduced from the Old World, adventive on disturbed Carex microptera Mack. Occasional to common along ground, u.sually in and below the pinvon-juniper zone. drainages of permanent or near permanent stream flow. July-August. 14233. June-July. 14194, 14289. Kochia americana S. Wats. Occasional on Lake Bon- Carex nebrascensis Dewey. Verv locallv common to neville deposits of sandy clay above the Sevier River at abundant in wet places. June-July. 14000. the extreme northern limit of the area. Mav-June. Carex petasata Dewey. Infrequent in Vasev sage- 13.S.37, 14047, 15639. brush communities. June-Julv. 14.337. Kochia scoparia (L.) Schrad. bitroduced, weedv, col- Carex rossii F. Boott in Hook. Occasional to common lected from along Highway 132 at the north edge of the in many plant communities in and above the pinvon- area. August-September. 17965. jimiper zone. June. 14016. Monnlepis mittalliana (Schult.) Greene. Occasional in Carex vallicola Dewey. Occasional in Vasev sage- nianv plant communities over a wide elevational range, brush and mountain brush communities. June-July. usuall)' on disturbed soil, June-August. 1.3988, 14445. 14197, 14325, 144.34, 15624. Salsola ibcrica Sennen & Pau. bitroduced from the Eleocharis palustris (L.) R. & S. Infrequent or verv lo- Old World, adventive on disturbed ground, usuallv in cally common in wet places. June. 14001, 14191. and below the pinyon-juniper zone. June-Julv. 14230. Sarcobatus lermiculatus (Hook.) Torr. Occasional on Lake Bonneville deposits above the Sevier River at the Elaeagnaceae north edge of the area. June-Julv. Shepherdia argentea (Pursh) Nutt. In aeolian sand at the north edge of the area; more common adjacent to Convolvidaceae the area along the flood plain of the Sevier River. Jime. Convolvulus arveiisis L. Introduced from the Old 18047. \\'orld, adventive especially on disturbed ground, mostly in and below the pinvon-juniper zone. 14214. Ephedraceae

Ephedra nevadensis Wats. Occasional in rockv places Cornaceae in the pinyon-juniper zone. Cornus stohnifera Mich.x. Occasional along drainages with permanent stream flow. June 13987. Equisetaceae

laevigatum A. Br. Infrequent in moist Equisetum places along drainages. 14195. Seditm debile Wats. Occasional on rock outcrops in and above the pinvon-juniper zone. 14312, 14346, Julv. Ericaceae 15821, 17.356; Welsh et al. 17443. Sedum lanceolatiim Torr. The one specimen seen Arctostaphtjlos patala Greene. Locallv abundant and (Foster 6532) is from a sagebrush-jimiper community in sometimes dominant on slopes at the upper edge of the Oak Creek Canyon. June-July. pinyon-juniper zone, up through the mountain brush zone. April-June. 1401.5, 149.39, 15291, 16847. Cupressaceae Juniperus osteospenna (Torr.) Little. Dominant over- story plant over much of the area around the base of the gltjptospcrma Engelm. Infrequent or occa- mountains, growing on every substrate in the area from sional in sagebrush-juniper communities, perhaps most sand dunes to cracks of rock outcrops. common on sandy soil. June. 14141, 141.55. Juniperus scopidorum Sarg. Occasional in mountain Euphorbia ocellata Dur. & Hilg. Rare on aeolian sand bnish communities and with scattered fir. 15375, 15812. at the southwest part of the area. .\ugust-September. 152.50. Cyperaceae

Carex aurea Nutt. Seen onlv along a nioss\ springlet Fabaceae at Ledge Spring. June-.\ugust. 17231. Astragalus argophtjllus Nutt. var. martinii Jones Oc- Carex deweyana Schwein. var. bolanderi W. Boott in casional in sagebrush, pinvon-jimiper, and oak-maple Wats. Rare, only a few plants seen at the mouth of Davi- communities. May-June. 14201, 14.3.32, 15326, 15601; son Canyon and in Drv Canyon, along permanent Foster 6518, 6.5.30,' 6558. streamlets. Julv. 15826. 17317. 288 Great Basin Naturalist Vol. 44, No. 2

Astragalus histilcatus (Hook.) Gray. Locally common Medicagn sativa L. Introduced from the Old World, to ahuiidaiit on old Lake Bonneville deposits of seleni- used extensivelv in rangeland seedings. June-Jidy. ferons sandy clay or clay groimd above the Sevier River 14280. at the extreme north limit of the area. Jnne. 14044. Psoralea lanccolata Pursh. Connnon to abundant and Astragalus cahjcosus Torr. Locally common on old sometimes dominant on sand dunes in and below the pinvon-juniper zone. 14144; Foster 6.546. 1 ,ake Bonneville deposits of sandy clay above the Sevier June-Julv. River at the extreme north limit of the area; common Trifoliutn andinum Nutt. in T. & G. Locally common of rocks rock crevices in the just to the east of the area. May-June. 1.383.3. to abundant on faces and Astragalus ccramicus Sheld. Occasional, restricted to mountain brush zone and upward to the highest eleva- tions on the Canyon Mountains, mostlv north of Oak aeolian sand in and below the pinyon-juniper zone. Creek. Mav-June^ 14342. 14357, 14932, 1,5377, 15592, June. 14123. 1,5606, 1,5829. Astragalus ciharius Sheld. Occasional to connnon in Trifolium gymnocarpon Nutt. in T. & G. Seen once sagebnush, pinyon-juniper, and mountain brush comnui- along a trail to Lvman (>anvon, Vasey sagebrush-white nities; one of the most widespread and common loco fir community. June. 16881. weeds of the area. Mav-June. 13785, 13881, 13884, Trifolium pratense L. Introduced from the Old 13940, 15293. 16882; Foster 6528; Welsh 16409. World, seen around a spring in Eight Mile Canyon. July. Astragalus convallarius Greene. Occasional in sage- 17240. l)rush. pinvon-juniper, and mountain brush commu- Trifolium repens L. Occasional in moist soil along nities. 1.3944, 13962. June'. drainages. Mav-July. 14188. Astragalus eurekensis Jones. Occasional to connnon Vicia americana Muhl. ex Willd. Occasional to fre- in sagebnish and pinyon-juniper communities, occasion- quent in Vasev sagebrush-jiuiiper, mesic mountain brush communities. May-June. 13764, al in mountain brush, and aspen communities. Mav-fune. 13790, 13848, 13774, 13784, 1.3815, 1.3834, 13952, 15294; Welsh 16419, 13880, 14092; Weksh 16,5,39a. 16589, 16604. Astragalus geyeri Gray. Rare or infrecjuent on .sandy Fagaceae soils in and below the pinvon-jvmiper zone. June. 14227. Astragalus marianus (Rydb.) Barneby. Infrequent in Quercus gambelii Nutt. Connnon to dominant over various plant commimities, at the upper edge and above much of the mountain brush zone, occasional well down the pinvon-juniper zone. June-July. 16886; Welsh 16433. into the pinyon-juniper zone. May-June. 13812, 14942. Astragalus oophorus Wats. Occasional to common in .sagebnish and pinyon-juniper comnnuiities. May-June. Funiariaceae 1.3816, 13887, 1.5.352; Welsh 16444, 16,587. Astragalus scopulorum T. C. Porter. Infrequent in Corydalis aurea Willd. Infrequent in various plant mountain bnish, aspen, and fir communities. June-July. communities, usually above the pinyon-jimiper zone, 144,36. 1,5551. rather adventive on disturbed ground. June. 13986. Astragalus utahensis (Torr.) T. & G. Occasional to lo- ainindant in sagebrush and pinvon-juniper commu- callv Geraniaceae nities. Mav-June. 13765, 13806, 1.3953, 1.5351, 1,5397; Welsh 16403. Erodium cicutarium (L.) L'Her in Ait. Introduced Hedysarum boreale Nutt. Seen onlv at the north limit from the Old World, adventive with disturbance, occa- of the area on old Lake Bonneville deposits of sandy clay sional in sagebrush and pinvon-juniper communities. above the Sevier River, juniper connnunitv. Mav-June. May-Julv. 1,3798, 1.3807. 15,349. Geranium parryi (Engelm.) Heller. Rare, seen only at brachycalyx Rydb. Occasional to common two locations in cracks of massive quartzite outcrops in in sagebrush, pinvon-juniper, and mountain brush com- East Eightmile Creek. July-August. Plants vary from numities. Mav-June. 13808, 1.3949, 14025, 1,5333, 1,5360; typical G. parn/i in having whitish rather than purple Welsh 16402,' 16407, 16408. , and thev average somewhat smaller. Similar Lathyrus lauzwertii Kellogg. Occasional in mountain plants have been found in the Stansburv Mountains far- iirush and aspen communities. Jime-July. 14024, 140.31. ther to the north in Utah (A. Taye & B. Wall 1213 and These plants be worthy Lathyrus pauciflorus Fern. Infrequent in moiuitain \. Taye & J. Herrick 1431). may brush and tir-snowberry communities. June-Jidy. 1,5618, of varital status. 14423, 14916. 1.5799. Lotus utahensis Otiey. Occasional in sagebrush and Hvdrophvllaceae sagebrush-juniper communities, seen only in Church Movmtains and south end of Canyon Mountains. occidentale (Wats.) Gray. Common to June-July. 14085, 14278, 14444. dominant understory plant in oak and aspen commu- Lupinus hri'vicaulis Wats. Seen only on aeolian sand nities, occasional in other connnunities above the pin- at i'Ady Springs Wash in the pinvon-juniper zone. von-juniper zone. May-June. 13999, 15372. June-July. 141,50, 14225, 14228. Phacelia ivesiana Torr. Infrequent in juniper-sage- Lupinus prunophilus Jones. Occasional in pinyon- brush connnimities. Mav-June. 15,564, 15677. juniper connnunities, seen only on south end of Canyon Phacelia linearis (Pursh) Holz. Infrequent in pinvon- juniper brush comnnuiities, very locally Mountains. May-June. 14165, 1,5,3.32. and mountain aeolian in a pinyon-juniper stand in Lupinus sericeus Pursh. Seen only in Oak C^reek common on sand Whiskey Creek drainage. May-June. 15558, 15565, drainage, sagebrush-oak-white fir connnunit\'. June-Julv. 16838. Foster 65,36; Welsh et al. 17452. April 1984 Goodrich: Canyon and Church Mountain Plants 289

Jiincaceae Loasaceae

Junctts articus VVilld. Occasional to locallv conmion Mentzelia laevicaulis (Dougl.) T. &. G. Seen only at in wet places. June-July. 14002. one location between the Canyon and Church moun- Junctts htifonius L. Infre(juent along epiienieial tains in a juniper-sagebrush community. Jidy-August. drainages in pinvon-juniper connnunities. fune. 14086, 14311. 14140,17326. Mentzelia tncmtana (A. Davids.) A. Davids. Occasion- Junciis ensifoHus VVikstr. Occasional along permanent al in juniper and juniper-sagebrush communities. streams, around seeps and springs, over a wide eleva- June-July. 14221, 14112. Much like M. alhicaulis Dough, tional range. June-July. 14193, 14284. which might also be in the area. Juncus tenuis VVilld. Infrecjuent in moist and wet places. June-July. 14288. Malvaceae ,

Lamiaceae Iliamna rivularis (Nutt.) Rvdb. Seen at two stations: Buck Hollow north e.xposure, white fir-oak-niaple com- Agastache urticifolia (Benth.) Kiintze. Occasional in munity, and mouth of Oak Creek, burned over sage- connnunities, locally abun- mountain hnish and aspen bru.sh-moiuitain brush commimitv. June-July. 15602, 14361. dant in snow accunnilation areas. June-July. 17318. Dracocephalum parviflonim Nutt. Introduced from Sphaeralcea coccinea (Nutt.) Rydb. Occasional in the Old World, adventive on disturbed ground, usually sagebrush and juniper communities in and below the in and below the pinvon-juniper zone. Jiuie. 14216, pinvon-juniper zone. Mav-Jime. 1.3930, 14101. 17.321. Sphaeralcea grossulariifolia (H. & A.) Rydb. Occa- Leonarus cardiaca L. Introduced from the Old sional in sagebrush and jimiper communities in and be- World, adventive; seen only at Oak Creek, on canyon low the pinyon-juniper zone. May-June. 13930, 14101; bottom with oak; most abundant where cattle have Welsh 16595. made beds under oak. June-July. 14313, 17959. Oxyhaphus linearis (Pursh) Robins. A single speci- Marruhium vulgare L. Introduced from the Old men collected from month of Oak Creek-Dry Creek, World, adventive on disturbed groimd, usually in and burned over jiuiiper community. July-September. below the pin\'on-jiiniper zone. June-July. 14164. 17.331. Monardella odoratissima Benth. Occasional to locally Tripterocalyx micranthus (Torr.) Hook. Occasional in common in rockv places usually in the mountain brush dunes at the southwest edge of the area. June. zone. July-September. 14319, 15143, 17.357. sand 14146.

Liliaceae Allium acuminatum Hook. Occasional to common in various plant communities over a wide elevational Camissonia boothii (Dougl.) Raven ssp. alyssoides range. Mav-June. 13961, 14014, 14081. (H. & A.) Raven. Rare, seen only at one station at the Allium hrandegei Wats. Infrequent or occasional in south end of Canyon Mountains in the pinyon-juniper various plant communities, mostly above the pinvon- zone. May-June. 14128. juniper zone. June-July. 14353, 13989 15069. Epilobium angustifolium L. Seen at two locations Calochortus nuttallii T. & G. Occasional to common where very locally common, along streamlets in moun- in sagebrush and pinvon-jiuiiper communities. June. tain brush and fir communities. July-August. 17943. 13968. 13978, 14091, 14105. 14171, 14333, 15690. Epilobium brachycarpum Presl. Occasional in sage- Erythronium grandiflorum Pursh. Occasional to local- brush, pinyon-juniper, and mountain brush commu- lv common in various plant communities, usually above nities. June', July. 14090, 14142, 14298, 15802. the pinyon-juniper zone. April-Mav. 13991, 15316, Epilobium canum (Greene) Raven ssp. garrettii (A. 15386; Welsh 16422. Nels.) Raven. Occasional to common in rocky places, of- Fritillaria atropurpurea Nutt. Infrequent or rare in ten on rock outcrops in mountain brush communities. Vasev sagebrush and mountain brush commiuiities. July-September. 14315, 14457, 14917; Welsh et al. June-July. 13998, 14338. 17449. Very locally com- californicum Durand. Epilobium ciliatum Raf. Occasional in moist places in wet places in Drv (Janvon and Bowens Canyon. mon along drainages. June-July. 14190, 15835; Welsh et al. August. 17315. July- 17446. Zigadenus panicidatus (Nutt.) Wats. Occasional to Epilobium nevadense xMunz. Rare in rocky places in rather common in Vasey sagebrush-juniper, pinyon-jimi- mountain brush communities. .\ugust-September. per, and moiuitain brush communities. May-June. 14918, 15144. 1380.5, 13923. 14120, 14145. Epilobium saximontanum Hausskn. Occasional in Zigadenus venenosus Wats. Seen only at Ledge moist places along drainages. June-July. 15822. Spring along a springlet. July. 17226. Gayophytum racemosum T. & G. Occasional, some- what adventive on disturbed ground, various plant com- Linaceae munities, often on moist soil. June-July. 14303, 14.358. Gayophytum ramosissimum Nutt. Occasional, some- Linum percnne L. var. lewisii Hulten. Occasional to what adventive on disturbed ground, various plant com- common in sagebrush, pinyon-jimiper, and mountain usually on dry soil. 13964, 14121. bnish connnunities. munities, June-JuK'. 290 Great Basin Naturalist Vol. 44, No. 2

Oenothera caespitosa Nutt. Occasional in sagebrush Agropyron eristatum (L.) Gaertn. Introduced from and pinvon-juniper communities. May-June 13954, the Old \Vorld, used extensively in rangeland seedings in 14209. pinyon-juniper and sagebrush communities. May-June. Oenothera pallida Lindl. Common to abundant on 13912. aeiolian sand and locally dominant on active sand dunes. Agropyron dasifstachyum (Hook.) Scribn. Infrequent In and below the pinyon-juniper zone. June-September. in various plant comnumities in and above the pinyon- 14148, 16388. juniper zone. June. 13967. Agropyron intermedium (Host) Beauv. var. inter- Orchidaceae medium. Introduced from the Old World, used exten- sively in rangeland seedings in pinvon-juniper and sage- Hahenaria sparsiflnra Wats. Rather rare in white fir brush communities. Late May-July. 14279, 14.304. cximmunities. Julv. 14339, 14344. Agropyron intermedium (Host) Beauv. var. trichopho- Habenaria unalascensis (Sprengel) Wats. Seen only at rum (Link) Halacsy. Introduced from the Old World, the following location; mouth of Davison Creek in shade used extensively in rangeland seedings in pinyon-juniper of white fir. July. 15828. and sagebrush communities. Late May-July. 14304. Agropyron smithii Rydb. Locally common to abun- Orobanchaceae dant along swales with moderately deep and somewhat fine-textured soils in the Church Mountains and south along Orohanche corymbosa (Rydb.) Ferris. Seen only end of Canyon Mountains, pinyon-juniper and sagebrush iirush conimiinitv in Can- a trail in a mountain Lyman communities. June-July. 14089, 14279. yon. June-September. 17941. Agropyron spicatum (Pursh) Scribn. & Smith. Co- Orobanche fasciculata Nutt. Infre(iuent in sagebrush dominant with sagebrush and often the dominant under- Fos- and pinvon-juniper communities. June-Julv. 15568; story plant in pinyon-juniper communities, increases ter 6547. rapidly after fire, occurs in awned and awnless forms. May-june. 13844, 13922, 13921. Agropyron trachyeauhim (Link) Malte. Locally com- mon in Vasev sagebrush, aspen, and fir communities, lo- munita Dur. Hilg. Occasional to locally Argemone & cally abundant in areas of snow accumulation. in pinyon-juniper communities, common, sagebrush and June-August. 14367, 15804. roadsides, in burn areas and other places mostly along aequalis Sobol. Seen along the stream at have been disturbed. 13972, 14038, 14181, that June. the mouth of Oak Creek Canyon. June-August. 17328. 14204. Bouteloua gracilis (H. B. K.) Lag. ex Steud. In- frequent to verv locally common in rocky places in low Pinaceae sagebrush and mountain brush communities near the crest of the range. August-September. 15146. Abies concolor (Gord. & Glend.) Lindl. Common to Bromus brizaeformis Fisch. & Mey. Introduced from dominant at upper elevations mostly on cool exposures. Europe, seen only at one station on southwest exposure Pinus ediilis Genelm. in Wisliz. Dominant overstory in a sparsely vegetated mountain brush connnunity. plant in the middle and at the upper edge of the pinyon- May-June. 15573. juniper zone, growing on most, if not all, substrates in Bromus carinatus H. & A. Occasional to locally abun- the area. May-June. dant in Vasev sagebrush, aspen, and moimtain brush Pinus mono})hylla Torr. & Frem. in Freni. Locally communities at upper elevations and often in snow ac- common in the central canyons of the west side of the cumulation areas. June-July. 14018, 15803. Canyon Mountains and occasional on the east side of Bromus diandrus Roth. Introduced from the Old these moimtains, apparently crossing some with P. World, seen onlv on distiubed ground around Mud edtilis as some trees show single- and two-needle bun- Spring, pinvon-juniper-oak community. May-June. dles on some twigs. May-June. 15637, 16456. 142 lo" Laws. Planted at Plantation Flat in the Old Oak Creek Canvon, where the trees have done very Bromus hordeaceus L. Introduced from World, adventive disturbed ground in and below the well. .\lso seen in Fool Creek. Mav-June. on Pseudotsuga menziesii (Mirb.) Franco. Occasional to pinvon-jvmiper zone, apparently cjuite limited in the co-dominant with white fir at upper elevations, usually area. May-June. 14211, 14286, 14294. on cool exposures. May-June. 15562. Bromus inermis Leyss. Introduced from the Old World, used in rangeland seedings, best adapted for cooler, more moist environments than pinyon-juniper communities, but occasional seeding in such commu- Plantago lanceolata L. Introduced from the Old nities. June-July. 13942. World, seen at the mouth of Oak Creek Canyon, burned Bromus rubens L. Seen only at two stations where lo- over cottonwood-willow-oak community. July. 17319. cally common in pinyon-juniper-mountain brush com- munities, on south and southwest exposures. May-Jime. 15574. Poaceae 15357, Bromus teetorum L. Introduced from the Old World, Aegilops cijlindrica Host. Introduced from tlie Old adventive on disturbed ground in many plant connnu- World, seen along a roadside at the mouth of Oak nities, but most common in and below the pinvon-juni- Creek. June. 18733^ per zone. May-June. 13795; Welsh 16601. April 1984 Goodrich: Canyon and Church Mountain Plants 291

Dactylis glomerata L. Introduced from the Old Secale cereale L. Introduced from the Old World, ad- World, used in rangeland seedings, best adapted for sites ventiye on disturbed ground, sometimes cidtivated as a that are more mesic than pinvon-juniper communities, cereal crop, used in the past for seeding rangelands in but occasionally seeded in such areas. Jmie 13941. sagebrush and juniper communities. May-Jime. 13926, Danthonia unispicata (Thurb.) Munro ex Macoun. 16827.

Infretjnent in drv rock places in mountain brush com- Sitanion hystrix (Nutt.) J. G. Sm. Occasional to com- numities. fulv. 14351, 14428. mon in many plant communities over a wide elevational Elymus cinereiis Scribn. & Merr. Apparently rare, range. May-June. 13937. seen in Oak Creek Canyon near the mouth of Lyman Sporobolus cryptandrus (Torr.) Gray. Occasional on Canyon, Va.sey sagebrush-oak community. June-August. sandy ground, seen only in Church Moimtains and south Elymus glat4cus Buck!. Infrequent in shade of aspen end of Canyon Mountains in and below the pinyon-juni- per zone. and Cottonwood, usually along drainages. June-July. July-September. 14126. 14301, 17327. Stipa Columbiana Macoun (S. nelsoni Scribn.). Occa- sional in Vasey sagebrush and aspen Elymus junceus Fisch. Introduced from the Old coninumities, usual- ly above the pinyon-juniper zone. June-Jidv. 14022, World, u.sed in rangeland seedings in sagebrush and pin- 14198, 14335, 156.33, 15805. yon-juniper communities. June. 16826. Stipa comata Trin. & Rupr. Occasional in sagebrush Festuca ovina L. Infrequent in juniper and perhaps and pinyon-juniper communities, usuallv on sandy soil. mountain bnish communities. May-Jiuie. 16828. May-June. 13969. Festuca pratensis Huds. Introduced from the Old Stipa lettcrmanii Vasey. Occasional in Vasey sage- World, seen around a spring in Eightmile Canyon. brush and mountain brush commimities above the pin- June-August. 17241. yon-juniper zone. June-July. 14.336. Glyceria striata (Lam.) A. S. Hitch. Infrequent along Vulpia ocfoflora (Walt.) Britt. Infrequent in sage- permanent streams in canyons. 14287. June-July. brush and pinyon-juniper communities. May-June. Hilaria jamesii (Torr.) Benth. Infrequent to occasion- 13891, 1.3971. al in juniper-sagebnish communities, perhaps most com- mon in the Church Mountains and extreme south limit Polemoniaceae of the Canyon Mountains. June. 13970, 14129.

Hordeum murinum Huds. Introduced from the Old Collomia linearis Nutt. Occasional in Vasey sagebru.sh World, seen only at Oak Creek Campground. May-July. and mountain brush conununities. May-Jime. 14026. 17363. Eriastrum diffusum (Gray) Mason. Seen only at Clay Melica bulbosa Geyer ex Port. & Coult. Occasional in Springs Wash, chainedover juniper woodland, sandy Vasey sagebrush and mountain brush commiuiities. soil; specimen has anthers (0.4-0.6 mm long) and the June-July. 14023. habit of E. diffusum, but the lobes of the are like Muhlenhergia wrightii Vasey in Coult. Rather rare those in E. sparsiflorum (Eastw.) Mason. May-Jime. along or near the crest of the Canyon Mountains on rock 14154. outcrops. September. 15147, 17948. Gilia aggregata Pursh Spreng. Occasional to rather Oryzopsis hymenoides (R. & S.) Ricker in Piper. Oc- common in sagebrush and pinvon-juniper communities. casional to common in sagebru.sh, pinyon-juniper, and June-September. 13935. mountain brush communities. 13983. Jime-July. . Gilia inconspicua (J. E. Sm.) Sweet Occasional to Oryzopsis micrantha (Trin. & Rupr.) Thurb. Rare in rather common in sagebrush and pinvon-juniper com- rocky places in mountain bnish coninumities and with munities. May-June. 13892, 13943, 13979, 13982. scattered fir. July. 14.343, 15811. Gilia leptomeria Gray. Occasional to rather common Fhleum pratense L. Introduced from the Old World, in sagebrush and pinvon-juniper communities, apparent- used in rangeland seedings, best adapted for areas more ly increasisng after chaining of jiiniper stands. mesic than pinyon-juniper communities. June. 14179. May-June. 13939, 13946, 14096, 14162, 15688. Foa annua L. Introduced from the Old World, adyen- Gilia polycladon Torr. in Emory. Occasional just to tive on disturbed ground, often along ephemeral and the east of the area in Little Valley, and to be expected permanent streams, and around springs. June-July. on the area on sandy soils in sagebrush and pinyon-juni- 14004, 14446, 15588. per commiuiities at lower elevations. Foa bulbosa L. Introduced from the Old World, ad- Leptodactylon pungens (Torr.) Nutt. Occasional in ventiye on disturbed ground, mostly in and below the many plant communities over a rather wide elevational pinyon-juniper zone, used in the past for seeding range- range. May-June. 13915, 14094. lands. 13780, 13796, 13904, 13907, 13977, 14130. Microsteris gracilis (Hook.) Greene. Occasional to Foa fendleriana (Steud.) Vasey. Common in pinyon- common in juniper and juniper-sagebrush communities, juniper, Vasey sagebnish, and mountain brush commu- increasing with disturbance and locally abundant on nities. April-June. 13809, 14028, 15327, 15328; Welsh chained areas. May-June. 13776, 13965; Welsh 16413. 16398. Fhlox austromontana Gov. Infrequent and local in Foa pratensis L. Occasional to locally common along pinyon juniper-juniper and mountain brush commu- drainages. nities, usually in rockv places. April-June. 13843, 15289; Foa secunda Presl. Common to abundant in sage- Welsh 16418. brush and pinvon-juniper communities. April-June. Fhlox hoodii Richards. Occasional in sagebrush and 13811. 13773, 13896, 13908, 15801; Welsh 16399, 16586. pin von-jimiper communities. March-May. 13783, Nutt. Occasional in sagebrush, pin- Polypogon semiverticillatus (Forsk.) Hylander. Seen Fhlox longifolia yon-juniper, and mountain brush communities. only in moist groimd around Mud Spring and mouth of May-June. 13772, 13797; Foster 6541; Welsh 16406. Oak Creek Canyon. June-July. 14212, 17.329. 292 Great Basin Naturalist Vol. 44, No. 2

Polygonaceae Pellaea brcweri D.C. Eaton. Infrequent in crevices of ledges. 1.5813. hrevicaule Nutt. In many plant comnni- Eriogpnum Polystichum scopulinum (D.C. Eaton) Maxon. Seen nities in and above the pinyon-juniper zone, abundant in onlv at East Fork Eightmile Creek, cracks of massive rockv places in mountain brush communities and open rock outcrop, in mountain brush zone. 14422. windswept ridges and slopes. Julv-October. 14318, 14345, 14368, 14427, 14921, 14922, 15246, 17.349, 17350. Small specimens from Fool Peak approach £. gratji Re- Portnlaceae veal, but these are probably depauperate plants of £. Claytonia lanceolata Pursh. Occasional in mountain hrevicaule. brush connnunities, and at the edge of aspen and fir Eriogonum cernuum Nutt. Occasional to abundant in communities. April-.May. 1.5290, 1,5.387; Welsh 16431. sagcbnish and pinyon-juniper communities, adventive Lewisia rediviva Pursh. Occasional on rocky ridges in and increasing on disturbed ground. June-September. the momitain brush zone. June-Julv. 14369. 14122, 1414.3, 14222, 14949; Foster 6545. Montia perfoliata (Donn) Howell. Locally common Eringomtm heracleoides Nutt. Seen only at a few lo- to abdunant imder juniper trees, especially along drain- cations where locally common on rocky slopes in Vasey ages; occasional in moimtain brush and fir communities. sagebrush and mountain brush communities. July. 13777, 14113, 15.398, 16844; Welsh 14323, 14429, 17225; Foster 6.5.33. Late April-May. Eriogonum hookeri Wats. Seen only at one location, 16411. in bare soil where a juniper had been uprooted in a chaining, .\ugust. 17725. Pyrolaceae Eriogonum kearneyi Tidestr. Locally abundant on ac- tive and partlv stable sand dunes at the west edge of the Chimaphila umbellata (L.) Bart. Seen only in one fir area; widespread on dunes to the west of the area. .Au- community in Davison Canyon. July. 15610. gust-October. 16.387. Pyrola asarifolia Michx. Seen at a single location in a Eriogonum maculatum Heller. Locally common on fir comniunitv in Davison Canyon. July. 156II. sandv soil in Clay Springs Wash area, apparently in- Pyrola secunda L. Seen only along the crest of the creasing after chaining of pinvon and juniper. June-July. Canvon Mountains above Lvman Canyon. July-.\ugust. 141,56, 14220. 179,54. Eriogonum microthecum Nutt. Occasional in low sagebrush and mountain brush communities. Ranunculaceae August-September. Eriogonum ovalifolium Nutt. Occasional in sagebrush columbianum Nutt. In T. & G. Occasional and pin\on-juniper communities. May-June. 13900, along drainages and around springs; a fairly good in- 1.3918, 14099. The yellow flowered phase (var. multi- dicator of permanent springs and streams. 14438. scapwn Gand.) is common just east of the area. Anemone tuberosa Rydb. Seen only at one station at Eriogonum pahnerianum Reveal in Munz. Occasional the north end of Canvon Mountains with sagebrush, in sandv soil, most conmion in Cla\' Springs Wash area scattered cliffro.se, and juniper. April-May. 153.56. in places cleared of juniper trees bv chaining. Jnne-Jidv. caerulea James. Seen only in cracks of 14161, 14219. rocks at Weetches Canvon, and along a springlet at Eriogonum racemosum Nutt. Infrequent but wide- Ledge Spring. June-July.'l,5807, 17228; Foster 6537. spread within various plant comnnmities over a rather Aquilegia formosa Fisch. in DC. Locally common wide elevational range. ,\ngust-Octolier. 14919, 15141, along drainages where stream flow is permanent or 17.3.55. nearly so. June-August. 14,300, 1,5587, 17229. Eriogonum umbeUatum Torr. var. dichrocephalum Gray. Infrequent in various plant com- Grand. Collected from Weetches Canvon from rockv munities over a wide elevational range. May-June. ground in a Douglas-fir-white fir comniunitv. 15797. 138.50, 1,5402A, 1.5619; Welsh 16,594. Eriogonum umbellatum Torr. var. umheUatum. Occa- Delphinium nuttallianum Pritz. Occasional in sage- sional in pinvon-jimiper-sagebnish and moimtain brush brush, pinvon-jnniper, and mountain brush commu- communities.' August. 14920, 14944, 1.58.32. nities. May-June. 1,5383; Welsh 16,591. or less weedy along Polygonum aviculare L. More Delphinium occidentale Wats. Locally common in roads and other areas of disturbance. Julv-September. mesic Vasev sagebrush, mountain brush, and aspen com- 17310. munities; seen at verv few stations in the area. July. Polygonum douglasii Greene. Occasional to common, 14.327, 15798. adventive on disturbed groimd in many plant commu- Myosurus minimus L. Seen only at East Fork Eight- nities. June .'\ugust. 14.302. mile Creek, among rocks in oak-sagebrush community. Pohfgonum sawatchense Small. Occasional to locallv June. 14026A. common, adventive on disturbed ground in nianv plant Ranunculus cymhalaria Pursh. Infrequent in wet communities. June-,\ugust. 14139. places. May-August. 14187; Foster 6.5,39. Rumex salicifolius Weinm. Infrequent along drainage Ranunculus sceleratus L. Occasional in seeps, bottoms in canyons. 14192. springs, and along streams, in mud and water. May-June. 14189, 14213, 16879, 17,322. Polvpodiaceae Ranunculus testiculatus Crantz. Introduced from the Old World, adventive on disturbed ground, usually in Cystopterus fragilis (L.) Bernh. in Schrader. In- and below the pinyon-juniper zone, common to abun- frecjnent or occasional in rock outcrops, mostly above dant under juniper.May-June. 13778, 13889. the pinyon-juniper zone. 14013, 14421, 1.5604. April 1984 GodDRicH: Canyon and Church Mountain Plants 293

Thalictrum fendleri Engelm. in Gray. Occasional in Potentilla biennis Greene. Infrequent along drainage rockv snowbank areas and aspen communities. bottoms in the mountain brush zone. June-August. June-July. 14354, 14360, 15593. 14433, 15584. Potentilla concinna Richards, var. bicrenata (Rydb.) Rhamnaceae Welsh & Johnston Seen only at the divide between Ly- man Canvon and Dry Creek. June. 16887. Ceanothus martinii Jones. Occasional in mountain Potentilla glandulosa Lindl. Occasional in rockv bnish conununitics. June-July. 14350, 15636, 15834. places in mountain brush zone. June-August. 14003, Ceanothus vehitintis Dougl. in Hook. Collected at 15595. Eightmile Creek in a mountain brush community. June. Potentilla gracilis Dougl. Seen only at the windswept, 13995. gravelly pass between Lyman Canyon and Dry Creek. June. 16885. Rosaceae Prunus virginiana L. Occasional and locally abundant along drainages and on slopes in the moutain brush Occasional in Amelanchier alnifoUa (Nutt.) Nutt. zone. May-June. 13992. 16880. mountain brush communities. May-June. 13994, Purshia tridentata (Pursh) DC. Occasional to abun- Occasional in pinyon- Amelanchier utahensis Koehn. dant in Vasey sagebrush-juniper, pinyon-juniper, and juniper and mountain brush communities, mostly on mountain brush commimities; sometimes common on drier sites than A. alnufolia. May-June. 16605; Foster aeolian sand with juniper. May-June. 13898, 13917, 6613. 14106, 14107. Cercocarpus ledifoUus Nutt. Dominant overstory Rosa woodsii Lindl. Occasional to locally common over wide areas above the pinyon-juniper zone, on plant along drainages. June-July. 14296, 17946. dryer sites than Gambel oak, aspen, or fir; occasional or Rubus leucodermis Dougl. Infrequent along drain- other plant com- common in upper pinyon-juniper and ages, usually in mountain brush and aspen communities. munities. May-June. June-July. 14435. Cercocarpus montanus Raf. Locally common to abun- Sanguisorba minor Scop. Introduced from the Old in pinyon-jiuiiper dant and mountain brush commu- World, used in recent rangeland seedings, particularly overstory at a few stations nities; the dominant shrub in the pinyon-juniper zone. June. 15691, 16823. such as in Oak Creek Canyon and near Scipio Pass. May-June. Rarely crosses with C. ledifoUus An apparent Rubiaceae of such origin was found above Wide Canyon.

The plant was treelike and had persistent, thick leaves Galium aparine L. Occasional to common in pinyon- as in C. ledifoUus but the leaves had the shape and ser- juniper, oak, and narrowleaf cottonwood comnuinities, rations of C. montanus. 13820, 15635 (16463, the appar- often along ephemeral drainages. May-July. 13802, ent hybrid). 14009, 14137, 14196, 14447. (Torr.) Cmcania mexicana D. Don. var. stansburiana Galium bifolium Wats. Seen at only one location on a to abundant in pinyon-juniper and Jeps. Common rock outcrop in a moimtain brush community near Pauls mountain bnish communities, occasionally dominant on Meadow. June-July. 15630. slopes and ridges; often associated with black sagebrush on shallow, rocky soils. Late Mav-June. 14093, 14108, Salicaceae 14307, 15569; Welsh 16599. Crategus douglasii Lindl. Seen at Oak Creek Camp- Populus angustifolia James. Common to dominant ground, cottonwood-oak-maple communitv. Mav-June. tree along some drainages of permanent or nearly per- 17361. manent waterflow. May-June. Fragaria vesca L. Infrequent in various plant commu- Populus tremuloidcs Michx. Common to dominant nities, usually above the pinyon-juniper zone. June-July. tree on mesic slopes and along drainages. April-May. 14426, 17233. Salix exigua Nutt. spp. melanopsis (Nutt.) Cronq. Oc- Ceum triflorum Pursh. Seen only at an ephemeral casional or locally common along drainages; usually in meadow of forbs at an ecotone between white fir and and below the pinyon-juniper zone. April-May. low sagebrush commvmities, above divide between Salix lasiandra Benth. Rather rare, along streams. Little Oak and Oak Creeks; very locally common. 17313. May-June. 15388. May-J\me. Salix lasiolepis Benth. Infrequent along drainages Holodiscus dumosus (Nutt.) Heller. Occasional in the pinyon-juniper and mountain brush zones. rocky places in pinyon-juniper and mountain brush com- within 16391, 16829. munities. June-July. 14469, 17348. April-May. 13990, 15315, in Creek Canyon, wet Ivesia gordonii (Hook.) T. & G. in Newberry. Seen Salix lutea Nutt. Seen Oak only at head of Eightmile Creek, rock outcrops, moun- places. May-June. 16878. tain brush-fir zone. June-July. 14936, 17236. Salix scouleriana Barratt in Hook. Infrequent along (Nutt.) Rydb. Very locally drainages in mountain brush, aspen, and fir commu- common on faces of rock outcrops, probably most com- nities. April-May. 15252, 15622, 17314, 17944. mon on limestone. July-August. 15830. Physocarpus malvaceus (Greene) Kuntze. Seen only Santalaceae at Buck Hollow and Lyman Canyon, rocky canyon sides, north exposure, white fir-oak-maple community, locally Comandra umbellata (L.) Nutt. Occasional in many common at both places. June. 15603, 17947. plant communities. May-July. 13831, 13899, 15814. 294 Great Basin Naturalist Vol. 44, No. 2

Saxifragaceae Penstemon cyananthus Hook. Rare in pinyon-jimiper, Vasey sagebrush, and mountain brush communities. in pinyon-jnni- Heuchera parvifolia Null. Occasional June-July. 14352, 17239. Plants of the area might be per and mountain brush communities, usually in rocky referable to ssp. longiflorits Pennell. places. June-July. 14033, 15594, 15598. Penstemon eatonii Gray. Occasional in pinyon-juniper Heuchera rubescens Torr. Occasional to common in comnnmities. June. 13948, 15567. crevices of ledges and cliffs at the upper edge of the pin- Penstemon humilis Nutt. Occasional in pinyon-juni- yon-juniper zone and upward. June-July. 14168, 15597, per and mountain brush commiuiities, often in rocky 15823. places. May-June. 13882, 13893, 14167, 15631, 16825. Lithophragma glabra Nutt. Infrequent or occasional Penstemon leonardii Rydb. Very locally common in in sagehnish, pinvon-juniper, and mountain brush com- rocky places near the crest of the Canyon Mountains munities. .\pril-Mav. 14353A, 15295, 15321; Welsh with mountain brush and scattered fir. Jidy. 14340, 16592. 15793. Lithophragma pariiflora (Hook.) Nuft. The two Verbascum thapsus L. Occasional, usually in and be- specimens seen are from oak communities in Oak Creek low the pinyon-juniper zone, adventive on disturbed Canyon. .'Kpril-May. 18734; Cottam & McMillan 9616. ground. June-August. 14283. Mitella stauropetela Piper. Seen only in Eightmile Veronica americana Schwein. in DC. Infrequent,

Creek in a white fir community. June. 14425, 17227. aquatic or in mud. Jime-August. 14185, 17323. aureum Pursh. Infrequent to very locally com- Veronica anagallis-aquatica L. Infrequent, aquatic or mon along drainages. Mav-June. 17453; Welsh et al. in mud. June-August 14183, 17324; Welsh et al. 17445. 17453. Veronica biloba L. Common to abundant in many Ribes cereum Dougl. Collected once from Hard- plant communities, most common along drainages and scrabble Canyon from a pinyon-juniper-mahogany com- in canyons, adventive on disturbed ground. Mav-Jime. munity. June. 16842. 13775,'l3890, 13931, 15400; Welsh 16430. Ribes velutinum Greene. Infrequent, usually in rocky Veronica peregrina L. Infrequent along drainages, places in the mountain brush zone. June. 14328, 15323; usually in mountain brush, aspen, or fir communities. Welsh 16423. June-July. 14184. 14443. Ribes wolfii Rothr. Infrequent, Douglas-fir-white fir communities, usually in rocky places. July. 14372, 17230, Selaginellaceae 17945. Saxifraga rhomboidea Greene. Seen in a drying Selaginella watsonii Underw. Occasional in rocky meadow of forbs at an ecotone between a low sagebrush places in and above the pinyon-juniper zone. 14355. and white fir community at the divide between Oak Creek and Little Oak Creek, and in pockets of soil in a Solanaceae massive quartzite outcrop in upper Lyman Canyon. May. 15389, 16884. Nicotiana attenuata Torr. Occasional to locally com- mon in juniper-sagebrush communities, adventive on disturbed ground, and locally abundant on charred soil Scrophulariaceae where slash piles from chainings have been burned. Castilleja chromosa A. Nels. Occasional to common June-July. 14202. in sagebmsh, pinyon-juniper, and mountain brush com- Solanum triflorum Nutt. Occasional adventive on dis- munities. May-June. 13768, 13814, 14030; Welsh 16416. turbed groiuid; the few plants seen were in juniper and Castilleja linariifolia Benth. in DC. Occasional in sagebrush areas that had been chained. June-September. Vasey sagebrush and moimtain brush communities. 14163, 14945, 16384. June-July. 15142. Collinsia parviflora Dougl. ex Lindl. Occasional to Ulmaceae abundant in many plant communities over a wide eleva- tional range. April-June. 13771; Welsh 16415. Celtus reticulata Torr. Occasional along intermittent Limnosella aquatica L. Seen only at East Fork Eight- drainages of lower West Eightmile Creek. June. 14102. mile Canyon, wet soil along the stream and around springs. 14186. Urticaceae Mimetanthe pilosa (Benth.) Greene. Seen only at West Fork Eightmile Creek, ephemeral drainage bottom Urtica dioica L. spp. gracilis (Ait.) Seland. Occasional in juniper community. June. 14114. along drainages. June-July. Mumuliis guttatus Fisch. Occasional in wet places, usually in and above the pinvon-juniper zone. Valerianaceae June-August. 14290, 17233. T. Occasional and locally Nama densum Lemmon. Seen only at single location Plectritis macrosera & G. in pinyon-juniper communities and especially in Whiskey Creek drainage on aeolian sand in a juniper common under juniper trees and along drainages. May-June. community. June. 15563. 13799, 13927, 14115; Welsh 16404. Penstemon angustifolitis Nutt. Very locally common on aeolian sand in C'lav Springs Wash, sagebrush and jimiper communities. May-June. 15402, 15403. Verbenaceae Penstemon confusus A. Nels. Seen only on old Lake Verbena bracteata Lag. & Rodr. Common to abun- Bonneville deposits of sandy clay above the Sevier River dant usually in and below the pinyon-juniper zone, ad- at the extreme north limit of the area. May. 15360. April 1984 Goodrich: Canyon and Church Mountain Plants 295

ventive on disturbed ground, and mostly along roads. Viola nuttallii Pursh var. major Hook. Seen at two June-July. 13973, 14087, 14203. stations in Davison Canyon-Buck Hollow area of Fool Creek drainage, in white fir-aspen-snowberry and oak- Violaceae maple communities. June. 15613, 15620; Welsh 16426. Viola purpurea Kell. Infrequent along dry drainage Viola nephrophylla Greene. Infrequent along drain- bottoms in sagebrush-juniper communities, occasional in ages, usually in the mountain brush zone. May-June. rocky snowbank areas up to the crest of the Canyon 15376. Mountains. April-June. 13813, 14352a, 15288.