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ZOBODAT - www.zobodat.at Zoologisch-Botanische Datenbank/Zoological-Botanical Database Digitale Literatur/Digital Literature Zeitschrift/Journal: Arachnologische Mitteilungen Jahr/Year: 2011 Band/Volume: 40 Autor(en)/Author(s): Deltschev [Deltshev] Christo Artikel/Article: The faunistic diversity of cave-dwelling spiders (Arachnida, Araneae) of Greece 23-32 1 © Biodiversity Heritage Library, http://www.biodiversitylibrary.org/; Arachnologische Mitteilungen 40:23-32 Nuremberg, January 201 The faunistic diversity of cave-dwelling spiders (Arachnida, Araneae) of Greece Christo Deltshev doi: 1 0.543 1/aramit4004 Abstract: Until today, from Greek caves a total of 1 09 species of spiders belonging to 25 families are known. One species, the linyphiid Porrhomma convexum (Westring, 1861) was recorded here for the first time in Greece. The 109 species are distributed in caves of different geographic territories as follows:Thrace - 8 species, Macedonia - 1 8, - 1 - Central - Attiki-Saronic Islands - Peloponnese - 1 Evoia-Vories Sporades Epirus , Thessaly 6, Greece 3, 24, 5, - 1, Eastern Aegean Islands - 5, Cyclades - 3, Dodecanese - 6, Ionian Islands - 23, Crete - 47. The largest fraction of troglobite species were encountered mainly in the territories of Crete - 15 species (5 of which are anophthalmic), the Ionian Islands - 4, Thrace - 2 (both anophthalmic), the Attiki-Saronic Islands - 2 (both anophthalmic), the Pe- loponnese - 2 (one anophthalmic), and Macedonia, Thessaly, and the Cyclades - each with 2 species. The richness of the troglobitic spiders in these regions strengthens the assumption that they were major centres of speciation and evolution for the species of this group. According to their current distribution, the established 1 09 species can be classified into 12 zoogeograpical categories, grouped into 4 complexes (widely distributed, European, Mediterranean, endemics). The largest number of species belong to the endemic complex (53.2 %) and are also the most characteristic and reflect the local character of the cave-dwelling spiders. Key words: cave-spiders fauna, endemics, troglobites, zoogeography The earliest data on Greek cave-dwelling spiders were on the southern end of the Balkan Peninsula. The presented by SIMON (1885), KULCZYNSKI (1903), country has borders with Albania, the Republic of ROEWER (1928, 1959), DRENSKY (1936); Kra- Macedonia and Bulgaria to the north, and Turkey to TOCHVIL (1937, 1938), HADJISSARANTOS (1940), the east. The Aegean Sea lies to the east and south and FAGE (1945). More recent publications derive of mainland Greece, while the Ionian Sea lies to the from the investigations of BRIGNOLI (1968, 1971a, west. Both parts of the Eastern Mediterranean basin 1971b, 1972, 1974a, 1974b, 1974c, 1976, 1977, 1978, feature a vast number of islands, islets and rock is- 1979, 1984), Deeleman-Reinhold (1971, 1977, lands (Fig. 1). Two -thirds of the territory of Greece is 1983, 1985, 1989, 1993), DEELEMAN-REINHOLD dominated by limestone, many ofwhich are karstified & DEELEMAN (1988), SENGLET (1971, 2001), (CLENDENON 2009). Deltshev (1979, 1985, 1999, 2000, 2008), Beron The territory of Greece can be divided into 13 (1985, 1986), Beron & Stoev (2004), Thaler & geographical regions (BOSMANS & CHATZAKI 2005; Knoflach (1995), Wunderlich (1995), Bosse- Fig. 1). There are 7 geographical regions on the main- LAERS (1998), BOSSELAERS &HENDERICKX (2002), land: Thrace, Macedonia, Epirus, Thessaly, Central GASPARO (2003, 2004a, 2004b, 2005a, 2005b, 2006, Greece, Attica and the Peloponnese. The Ionian 2007, 2008, 2009), CHATZAKI et al 2002, BOSMANS Islands are situated on the western border of Greece dt CHATZAKI (2005), CHATZAKI &ARNEDO (2006), in the Ionian Sea. There are several island groups in and PLATNICK (2009). The critical incorporation of the Aegean Sea on the eastern side of Greece: Evoia all available literature records and the accumulation of and the Sporades, the Saronic Islands (grouped with new data are now sufficient to allow a critical analysis Attica), the Cyclades, the Eastern Aegean Islands, of the distribution of spiders established in the caves the Dodecanese and Crete (Fig. 1). of Greece. Results Study area and material Species composition Greece is a country in south-eastern Europe, situated The spiders established in the caves of Greece (Main- land and Insular part) are represented by 109 species, Christo DELTSHEV, Institute of Biodiversity and Ecosystem included in 52 genera and 25 families: Ctenizidae Research, 1 Tsar Osvoboditel Blvd; 1 000 Sofia, Bulgaria - 1, Filistatidae - 1, Sicariidae — 1, Scytodidae - 1, E-Mail: [email protected] Leptonetidae - 9, Pholcidae - 10, Segestriidae - 3, - - - submitted: 30. 12.20 1 0, accepted: 9.4.20 1 0; online: 10.1.2011 Dysderidae 12, Oonopidae 1, Mimetidae 1, Ere- © Biodiversity Heritage Library, http://www.biodiversitylibrary.org/; 24 C. Deltshev (6), Dysderidae (3), Nesticidae (2), and Linyphiidae (1). The genera with the largest number of species are Tegenaria (8), Lepthyphantes (6), Harpactea (5), and Histopona (5). The species are distributed in caves belonging to the geographic territories of Greece as follows: Thrace - 8 species, Macedonia - 18, Epirus - 1, Thessaly - 6, Central Greece - 3, te Attiki- Saronic Islands - 24, the Peloponnese - 15, Evoia-Voroies Sporades - 1, the Eastern Aegean Islands - 5, the Cyclades - 7, the Dodecanese - 6, the Ionian Islands - 23, Crete - 47 (Table 1). We also have to emphasise that the degree of exploration in these territories is not equal: the territories of Evoia- Sporades, the Cyclades, the Dodecanese and Central Greece are less explored. Cave-dwelling spiders can be categorised into four ecological groups (SKET 2008): • troglobites: species limited to a life cycle in caves. Often they show a suite of characters, associated with their adaptation to subterranean life: loss of LIV1KÖ8 f>ELAQOS pigment, loss of eyes and elongation of appenda- ges. Figure 1 : Map of different geographical regions in Greece. • eutroglophiles: species which can live their entire life in caves, but also occur in other environments. sidae - 1, Uloboridae - 1, Nesticidae - 4, Theridiidae • subtroglophiles: species which utilise caves, but - 4, Anapidae - 1, Linyphiidae - 16,Tetragnathidae must leave the caves to complete their life cycle. - 4, Araneidae - 1, Lycosidae - 1, Agelenidae — 21, • trogloxenes: species which occur underground Amaurobiidae — 4, Gnaphosidae — 6, Philodromidae sporadically. - 1, Thomisidae - 2, Salticidae - 2 (Table 1). One The largest fraction of troglobite species was encoun- species is new for the Greek spider fauna: Porrhomma tered mainly in the caves of the territories of Crete convexum (Westring, 1861) (marked in the list with *), -15 (5 anophthalmic), the Ionian Islands - 4, Thrace a spider with a Holarctic distribution and widespread - 2 (2 anophthalmic), the Attiki- Saronic Islands 2 (2 in European caves. It is also well represented in the anophthalmic), the Peloponnese -2(1 anophthalmic), caves of the Balkan Penisula - and not only estab- and Macedonia, Thessaly and the Cyclades each by 2 lished in the caves of Croatia, Romania and Turkey species (Table 1). All troglobites are endemics for the (DELTSHEV 2008). The number of species is high and territory of Greece or the Balkan Peninsula. represents about 13 % of the Greek spiders. This is Very important is the presence of eutroglophiles also evident from a comparison with the number of (35 species), because together with troglobites (29 cave-dwelling spiders recorded from the other coun- species), they can be considered as dependent faunistic tries of the Balkan Peninsula: Bulgaria - 99, Croatia elements of caves. The largest number of species is - 63, Serbia - 59, Bosnia and Herzegovina - 52, established in the caves of Crete (14 species), the Io- Macedonia - 44, Montenegro - 44, Slovenia - 43, Al- nian Islands (11 species), the Attiki- Saronic Islands (9 bania - 10, Turkey - 8, and Romania - 4 (DELTSHEV species), Macedonia (8 species), and the Peloponnese 2008). The established number of species, however, (8 species). Here, the endemics are represented by depends not only on the size of the regions, but also 14 species (35 %). A present day example of active on the degree of exploration. The most characteristic subterranean colonisation and cave penetration are are the families: Leptonetidae (8.2 %), Pholcidae the species Lepthyphantes centromeroides and Pallidup- (9.2 %), Dysderidae (11 %), Linyphiidae (14.6 %), hantes spelaeorum, widespread in the Balkan Peninsula and Agelenidae (19.2 %). The families with largest (DEELEMAN-REINHOLD 1978). Here, the species number of anophthalmic species are Leptonetidae Palliduphantes istrianus should also be included. 1 ; © Biodiversity Heritage Library, http://www.biodiversitylibrary.org/; Cove-dwelling spiders of Greece 25 ifi "Ö i_ "O C O C ra CR X X X X X X X X x X X uo u c E Eastern 03 0) CD "O Q- c - O uj II O X X X X X 2? EAI s/ A3 ra ^ CD 111 DO X X X X 4—' cd" 15 U c O 03 c Evia-Sporades; o u Q. - 03 |- O £ .y Ö £ - E o ^ ESI HI CD 111 ~o EAI X in c 2 ^ c oc Peloponnese; S £ ESI v -o E c 13 U . ”3 .. m - E PE 1 1 CD "Ö PE X X X c. - c ca uj Islands; ASI X X X X to E CD 03 i/i -a "o c Attiki-Saronic c CGR to £ ca cjj CD — Q_ T3 u — -C O jk LU U Q. ^ Io C 13 - O LU -O O ^ THE X 1 03 E ra ASI < CD J2 ua U £ - -6 c < ^ Greece; EP X X £, to 2 ^.P o a CD HI to Z ™ Central E X X X t c > CD MA v/i fa 03 “Ö CD <D - >5 S Crete. 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  • 19 4 273 282 Tanasevitch2 for Inet.P65

    19 4 273 282 Tanasevitch2 for Inet.P65

    Arthropoda Selecta 19(4): 273282 © ARTHROPODA SELECTA, 2010 On synonymy of linyphiid spiders of the Russian fauna (Arachnida: Aranei: Linyphiidae). 1 Î ñèíîíèìèè ïàóêîâ-ëèíèôèèä ôàóíû Ðîññèè (Arachnida: Aranei: Linyphiidae). 1 Andrei V. Tanasevitch À.Â. Òàíàñåâè÷ Centre for Forest Ecology and Production, Russian Academy of Sciences, Profsoyuznaya Str. 84/32, Moscow 117997 Russia. E-mail: and- [email protected] Öåíòð ïî ïðîáëåìàì ýêîëîãèè è ïðîäóêòèâíîñòè ëåñîâ ÐÀÍ, Ïðîôñîþçíàÿ óë. 84/32, Ìîñêâà 117997 Ðîññèÿ. E-mail: and- [email protected] KEY WORDS: Spiders, Linyphiidae, new synonym, new combination, Russian fauna. ÊËÞ×ÅÂÛÅ ÑËÎÂÀ: Ïàóêè, Linyphiidae, íîâûé ñèíîíèì, íîâàÿ êîìáèíàöèÿ, ôàóíà Ðîññèè. ABSTRACT. Seven new synonyms are established Introduction for the Russian fauna: Agyneta yakutsaxatilis Marusik et Koponen, 2002, syn.n. = Agyneta amersaxatilis Hundreds of new taxa of the linyphiid spiders have Saaristo et Koponen, 1998; Bolyphantes palaeformis been described during the last three decades from the (Tanasevitch, 1989), syn.n. = Bolyphantes bipartitus territory of the former Soviet Union, mainly from the (Tanasevitch, 1989), both comb.n. (ex Lepthyphantes Caucasus, Siberia, the Far East and Central Asia. A Menge, 1866); Epigytholus tuvensis Tanasevitch, 1996, significant progress in the taxonomy of the family syn.n. = Epigytholus kaszabi (Wunderlich, 1995), Linyphiidae, as well as abundant material from various comb.n. (ex Lepthyphantes); Hybauchenidium holmi Palearctic regions which has been recently accumulated, Marusik, 1988, syn.n. = Hybauchenidium aquilonare allow one to not only critically consider some of the (L. Koch, 1879); Poeciloneta yanensis Marusik et Ko- earlier described species and genera but also, in some ponen, 2002, syn.n. = Poeciloneta variegata (Black- cases, to dispel any doubts in their invalidity.
  • Revision of the Histopona Italica Group (Araneae: Agelenidae), with the Description of Two New Species

    Revision of the Histopona Italica Group (Araneae: Agelenidae), with the Description of Two New Species

    TERMS OF USE This pdf is provided by Magnolia Press for private/research use. Commercial sale or deposition in a public library or website is prohibited. Zootaxa 3640 (1): 023–041 ISSN 1175-5326 (print edition) www.mapress.com/zootaxa/ Article ZOOTAXA Copyright © 2013 Magnolia Press ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.3640.1.2 http://zoobank.org/urn:lsid:zoobank.org:pub:2865EE2A-01A9-45D9-B299-AAF6E118D5C4 Revision of the Histopona italica group (Araneae: Agelenidae), with the description of two new species ANGELO BOLZERN1,2, PAOLO PANTINI3 & MARCO ISAIA4,5 1Naturhistorisches Museum Basel, Augustinergasse 2, CH-4001 Basel, Switzerland 2American Museum of Natural History, Invertebrate Zoology, Central Park West at 79th Street, New York, NY 10024-5192, United States. E-mail: [email protected] 3Museo Civico di Scienze Naturali “E. Caffi”, Piazza Cittadella, 10. I-24129 Bergamo, Italy. E-mail: [email protected] 4Dipartimento di Scienze della Vita e Biologia dei Sistemi, Università di Torino, Via Accademia Albertina, 13. I-10123 Torino, Italy. E-mail: [email protected] 5Corresponding author Abstract During a large survey of agelenid spiders from different private and museum collections, a closer examination of material from all over Italy (including type material and fresh material) previously identified as Histopona italica Brignoli 1977, disclosed two new species for science, both belonging to the italica group. Based on the results of the present work, we describe Histopona leonardoi sp. n. and H. fioni sp. n. and revise the distribution pattern of H. italica group in Italy and Switzerland.
  • Spiders from the Ionian Islands of Kerkyra (Corfu) and Lefkada, Greece (Arachnida: Aranei)

    Spiders from the Ionian Islands of Kerkyra (Corfu) and Lefkada, Greece (Arachnida: Aranei)

    Arthropoda Selecta 23(3): 285–300 © ARTHROPODA SELECTA, 2014 Spiders from the Ionian islands of Kerkyra (Corfu) and Lefkada, Greece (Arachnida: Aranei) Ïàóêè Èîíè÷åñêèõ îñòðîâîâ Êåðêèðà (Êîðôó) è Ëåâêàäà, Ãðåöèÿ (Arachnida: Aranei) Anthony Russell-Smith Ý. Ðàññåë-Ñìèò 1, Bailiffs Cottage, Doddington, Sittingbourne, Kent ME9 0JU, the UK. KEY WORDS: Aranei, Greece, Ionian islands, faunistic list. КЛЮЧЕВЫЕ СЛОВА: Aranei, Греция, Ионические острова, фаунистический список. ABSTRACT. A list of spiders collected from the remains limited compared to that for most of central Ionian islands of Kerkyra and Lefkada is provided and NW Europe, as is the case for all areas of the together with a list of all previously published records. eastern Mediterranean. An important recent advance Information is provided on collection localities, habi- was the publication of an annotated catalogue of the tats and geographic distribution of all species record- Greek spider fauna [Bosmans & Chatzaki, 2005]. This ed. A total of 94 species were collected in Kerkyra, of listed a total of 856 valid species for the country, which 37 had not been previously recorded. 98 species although that figure has been substantially increased by were collected in Lefkada, of which 71 were new records subsequent work. Since then, provisional checklists for the island. Currently, 243 spider species are record- have been published for the islands of Lesbos [Bos- ed from Kerkyra and 117 species from Lefkada. Five mans et al., 2009], Chios [Russell-Smith et al., 2011] species collected were new records for Greece: Agyne- and Crete [Bosmans et al., 2013]. These checklists ta mollis, Tenuiphantes herbicola (Lefkada), Trichon- apart, there has been little published on the spider cus sordidus (Kerkyra), Tmarus stellio (Kerkyra) and faunas of individual regions of Greece.