And Sequence Stratigraphy of the Middle Cambrian Spence Shale of Northern Utah and Southern Idaho
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Early Sponge Evolution: a Review and Phylogenetic Framework
Available online at www.sciencedirect.com ScienceDirect Palaeoworld 27 (2018) 1–29 Review Early sponge evolution: A review and phylogenetic framework a,b,∗ a Joseph P. Botting , Lucy A. Muir a Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210008, China b Department of Natural Sciences, Amgueddfa Cymru — National Museum Wales, Cathays Park, Cardiff CF10 3LP, UK Received 27 January 2017; received in revised form 12 May 2017; accepted 5 July 2017 Available online 13 July 2017 Abstract Sponges are one of the critical groups in understanding the early evolution of animals. Traditional views of these relationships are currently being challenged by molecular data, but the debate has so far made little use of recent palaeontological advances that provide an independent perspective on deep sponge evolution. This review summarises the available information, particularly where the fossil record reveals extinct character combinations that directly impinge on our understanding of high-level relationships and evolutionary origins. An evolutionary outline is proposed that includes the major early fossil groups, combining the fossil record with molecular phylogenetics. The key points are as follows. (1) Crown-group sponge classes are difficult to recognise in the fossil record, with the exception of demosponges, the origins of which are now becoming clear. (2) Hexactine spicules were present in the stem lineages of Hexactinellida, Demospongiae, Silicea and probably also Calcarea and Porifera; this spicule type is not diagnostic of hexactinellids in the fossil record. (3) Reticulosans form the stem lineage of Silicea, and probably also Porifera. (4) At least some early-branching groups possessed biminerallic spicules of silica (with axial filament) combined with an outer layer of calcite secreted within an organic sheath. -
Morphology and Developmental Traits of the Trilobite Changaspis Elongata from the Cambrian Series 2 of Guizhou, South China
Morphology and developmental traits of the trilobite Changaspis elongata from the Cambrian Series 2 of Guizhou, South China GUANG-YING DU, JIN PENG, DE-ZHI WANG, QIU-JUN WANG, YI-FAN WANG, and HUI ZHANG Du, G.-Y., Peng, J., Wang, D.-Z., Wang, Q.-J., Wang, Y.-F., and Zhang, H. 2019. Morphology and developmental traits of the trilobite Changaspis elongata from the Cambrian Series 2 of Guizhou, South China. Acta Palaeontologica Polonica 64 (4): 797–813. The morphology and ontogeny of the trilobite Changaspis elongata based on 216 specimens collected from the Lazizhai section of the Balang Formation (Stage 4, Series 2 of the Cambrian) in Guizhou Province, South China are described. The relatively continuous ontogenetic series reveals morphological changes, and shows that the species has seventeen thoracic segments in the holaspid period, instead of the sixteen as previously suggested. The development of the pygid- ial segments shows that their number gradually decreases during ontogeny. A new dataset of well-preserved specimens offers a unique opportunity to investigate developmental traits after segment addition is completed. The ontogenetic size progressions for the lengths of cephalon and trunk show overall compliance with Dyar’s rule. As a result of different average growth rates for the lengths of cephalon, trunk and pygidium, the length of the thorax relative to the body shows a gradually increasing trend; however, the cephalon and pygidium follow the opposite trend. Morphometric analysis across fourteen post-embryonic stages reveals growth gradients with increasing values for each thoracic segment from anterior to posterior. The reconstruction of the development traits shows visualization of the changes in relative growth and segmentation for the different body parts. -
Trilobite Evolutionary Rates Constrain the Duration of the Cambrian Explosion
Trilobite evolutionary rates constrain the duration of the Cambrian explosion John R. Patersona,1, Gregory D. Edgecombeb, and Michael S. Y. Leec,d aPalaeoscience Research Centre, School of Environmental & Rural Science, University of New England, Armidale, NSW 2351, Australia; bDepartment of Earth Sciences, The Natural History Museum, London SW7 5BD, United Kingdom; cCollege of Science and Engineering, Flinders University, SA 5001, Australia; and dEarth Sciences Section, South Australian Museum, Adelaide, SA 5000, Australia Edited by Andrew H. Knoll, Harvard University, Cambridge, MA, and approved January 9, 2019 (received for review November 12, 2018) Trilobites are often considered exemplary for understanding the phenotypic and genomic evolution (7, 8). Rapid morphological Cambrian explosion of animal life, due to their unsurpassed di- and molecular evolution during the earliest Cambrian almost versity and abundance. These biomineralized arthropods appear certainly underpinned the pronounced pulses of origination and abruptly in the fossil record with an established diversity, phyloge- diversification throughout the Terreneuvian (3, 9, 10). However, netic disparity, and provincialism at the beginning of Cambrian the question remains as to when evolutionary rates slowed to Series 2 (∼521 Ma), suggesting a protracted but cryptic earlier his- Phanerozoic norms, thus marking the end of the Cambrian ex- tory that possibly extends into the Precambrian. However, recent plosion. For instance, the calibrations used in ref. 7 were mostly analyses indicate elevated rates of phenotypic and genomic evolu- 488 Ma or younger; that analysis therefore only had weak power tion for arthropods during the early Cambrian, thereby shortening to constrain fast early rates further back than that time point. -
Comprehensive Review of Cambrian Himalayan
http://www.diva-portal.org Postprint This is the accepted version of a paper published in Papers in Palaeontology. This paper has been peer- reviewed but does not include the final publisher proof-corrections or journal pagination. Citation for the original published paper (version of record): Popov, L E., Holmer, L E., Hughes, N C., Ghobadi Pour, M., Myrow, P M. (2015) Himalayan Cambrian brachiopods. Papers in Palaeontology, 1(4): 345-399 http://dx.doi.org/10.1002/spp2.1017 Access to the published version may require subscription. N.B. When citing this work, cite the original published paper. Permanent link to this version: http://urn.kb.se/resolve?urn=urn:nbn:se:uu:diva-255813 HIMALAYAN CAMBRIAN BRACHIOPODS BY LEONID E. POPOV1, LARS E. HOLMER2, NIGEL C. HUGHES3 MANSOUREH GHOBADI POUR4 AND PAUL M. MYROW5 1Department of Geology, National Museum of Wales, Cathays Park, Cardiff CF10 3NP, United Kingdom, <[email protected]>; 2Institute of Earth Sciences, Palaeobiology, Uppsala University, SE-752 36 Uppsala, Sweden, <[email protected]>; 3Department of Earth Sciences, University of California, Riverside, CA 92521, USA <[email protected]>; 4Department of Geology, Faculty of Sciences, Golestan University, Gorgan, Iran and Department of Geology, National Museum of Wales, Cathays Park, Cardiff CF10 3NP, United Kingdom <[email protected]>; 5 Department of Geology, Colorado College, Colorado Springs, CO 80903, USA <[email protected]> Abstract: A synoptic analysis of previously published material and new finds reveals that Himalayan Cambrian brachiopods can be referred to 18 genera, of which 17 are considered herein. These contain 20 taxa assigned to species, of which five are new: Eohadrotreta haydeni, Aphalotreta khemangarensis, Hadrotreta timchristiorum, Prototreta? sumnaensis and Amictocracens? brocki. -
Smithsonian Miscellaneous Collections
SMITHSONIAN MISCELLANEOUS COLLECTIONS VOLUME 67, NUMBER 6 CAMBRIAN GEOLOGY AND PALEONTOLOGY IV No. 6.—MIDDLE CAMBRIAN SPONGIAE (With Plates 60 to 90) BY CHARLES D. WALCOTT (Publication 2580) CITY OF WASHINGTON PUBLISHED BY THE SMITHSONIAN INSTITUTION 1920 Z$t Bovb Qgattimote (press BALTIMORE, MD., U. S. A. CAMBRIAN GEOLOGY AND PALEONTOLOGY IV No. 6.—MIDDLE CAMBRIAN SPONGIAE By CHARLES D. WALCOTT (With Plates 60 to 90) CONTENTS PAGE Introduction 263 Habitat = 265 Genera and species 265 Comparison with recent sponges 267 Comparison with Metis shale sponge fauna 267 Description of species 269 Sub-Class Silicispongiae 269 Order Monactinellida Zittel (Monaxonidae Sollas) 269 Sub-Order Halichondrina Vosmaer 269 Halichondrites Dawson 269 Halichondrites elissa, new species 270 Tuponia, new genus 271 Tuponia lineata, new species 272 Tuponia bellilineata, new species 274 Tuponia flexilis, new species 275 Tuponia flexilis var. intermedia, new variety 276 Takakkawia, new genus 277 Takakkawia lineata, new species 277 Wapkia, new genus 279 Wapkia grandis, new species 279 Hazelia, new genus 281 Hazelia palmata, new species 282 Hazelia conf erta, new species 283 Hazelia delicatula, new species 284 Hazelia ? grandis, new species 285. Hazelia mammillata, new species 286' Hazelia nodulifera, new species 287 Hazelia obscura, new species 287 Corralia, new genus 288 Corralia undulata, new species 288 Sentinelia, new genus 289 Sentinelia draco, new species 290 Smithsonian Miscellaneous Collections, Vol. 67, No. 6 261 262 SMITHSONIAN MISCELLANEOUS COLLECTIONS VOL. 6j Family Suberitidae 291 Choia, new genus 291 Choia carteri, new species 292 Choia ridleyi, new species 294 Choia utahensis, new species 295 Choia hindei (Dawson) 295 Hamptonia, new genus 296 Hamptonia bowerbanki, new species 297 Pirania, new genus 298 Pirania muricata, new species 298 Order Hexactinellida O. -
The Spence Shale Lagerstätte: an Important Window Into Cambrian Biodiversity
Downloaded from http://jgs.lyellcollection.org/ by guest on September 24, 2021 Accepted Manuscript Journal of the Geological Society The Spence Shale Lagerstätte: an Important Window into Cambrian Biodiversity Julien Kimmig, Luke C. Strotz, Sara R. Kimmig, Sven O. Egenhoff & Bruce S. Lieberman DOI: https://doi.org/10.1144/jgs2018-195 Received 31 October 2018 Revised 21 February 2019 Accepted 28 February 2019 © 2019 The Author(s). This is an Open Access article distributed under the terms of the Creative Commons Attribution 4.0 License (http://creativecommons.org/licenses/by/4.0/). Published by The Geological Society of London. Publishing disclaimer: www.geolsoc.org.uk/pub_ethics Supplementary material at https://doi.org/10.6084/m9.figshare.c.4423145 To cite this article, please follow the guidance at http://www.geolsoc.org.uk/onlinefirst#cit_journal Downloaded from http://jgs.lyellcollection.org/ by guest on September 24, 2021 The Spence Shale Lagerstätte: an Important Window into Cambrian Biodiversity 1* 1,2 1,3 4 1,2 Julien Kimmig , Luke C. Strotz , Sara R. Kimmig , Sven O. Egenhoff & Bruce S. Lieberman 1Biodiversity Institute, University of Kansas, Lawrence, KS 66045, USA 2 Department of Ecology & Evolutionary Biology, University of Kansas, Lawrence, KS, USA 3Pacific Northwest National Laboratory, Richland, WA 99354, USA 4Department of Geosciences, Colorado State University, Fort Collins, CO 80523, USA *Correspondence: [email protected] Abstract: The Spence Shale Member of the Langston Formation is a Cambrian (Miaolingian: Wuliuan) Lagerstätte in northeastern Utah and southeastern Idaho. It is older than the more well- known Wheeler and Marjum Lagerstätten from western Utah, and the Burgess Shale from Canada. -
Palaeoecology of the Early Cambrian Sinsk Biota from the Siberian Platform
Palaeogeography, Palaeoclimatology, Palaeoecology 220 (2005) 69–88 www.elsevier.com/locate/palaeo Palaeoecology of the Early Cambrian Sinsk biota from the Siberian Platform Andrey Yu. Ivantsova, Andrey Yu. Zhuravlevb,T, Anton V. Legutaa, Valentin A. Krassilova, Lyudmila M. Melnikovaa, Galina T. Ushatinskayaa aPalaeontological Institute, Russian Academy of Sciences, ul. Profsoyuznaya 123, Moscow 117997, Russia bA´rea y Museo de Paleontologı´a, faculdad de Ciences, Universidad de Zaragoza, C/ Pedro Cerbuna, 12, E-50009, Zaragoza, Spain Received 1 February 2002; accepted 15 January 2004 Abstract The Sinsk biota (Early Cambrian, Botoman Stage, Siberian Platform) inhabited an open-marine basin within the photic zone, but in oxygen-depleted bottom waters. Its rapid burial in a fine-grained sediment under anoxic conditions led to the formation of one of the earliest Cambrian Lagerst7tte. All the organisms of the biota were adapted to a life under dysaerobic conditions. It seems possible that the adaptations of many Cambrian organisms, which composed the trophic nucleus of the Sinsk Algal Lens palaeocommunity to low oxygen tensions allowed them to diversify in the earliest Palaeozoic, especially during the Cambrian. Nowadays these groups comprise only a negligible part of communities and usually survive in settings with low levels of competition. Nonetheless, the organization of the Algal Lens palaeocommunity was not simple, it consisted of diverse trophic guilds. The tiering among sessile filter-feeders was well developed with the upper tier at the 50 cm level. In terms of individuals, the community was dominated by sessile filter-feeders, vagrant detritophages, and diverse carnivores/scavengers. The same groups, but in slightly different order, comprised the bulk of the biovolume: vagrant epifaunal and nektobenthic carnivores/ scavengers, sessile filter-feeders, and vagrant detritophages. -
Early and Middle Cambrian Trilobites from Antarctica
Early and Middle Cambrian Trilobites From Antarctica GEOLOGICAL SURVEY PROFESSIONAL PAPER 456-D Early and Middle Cambrian Trilobites From Antarctica By ALLISON R. PALMER and COLIN G. GATEHOUSE CONTRIBUTIONS TO THE GEOLOGY OF ANTARCTICA GEOLOGICAL SURVEY PROFESSIONAL PAPER 456-D Bio stratigraphy and regional significance of nine trilobite faunules from Antarctic outcrops and moraines; 28 species representing 21 genera are described UNITED STATES GOVERNMENT PRINTING OFFICE, WASHINGTON : 1972 UNITED STATES DEPARTMENT OF THE INTERIOR ROGERS C. B. MORTON, Secretary GEOLOGICAL SURVEY V. E. McKelvey, Director Library of Congress catalog-card No. 73-190734 For sale by the Superintendent of Documents, U.S. Government Printing Office Washington, D.C. 20402 - Price 70 cents (paper cover) Stock Number 2401-2071 CONTENTS Page Page Abstract_ _ ________________________ Dl Physical stratigraphy______________________________ D6 I&troduction. _______________________ 1 Regional correlation within Antarctica ________________ 7 Biostratigraphy _____________________ 3 Systematic paleontology._____-_______-____-_-_-----_ 9 Early Cambrian faunules.________ 4 Summary of classification of Antarctic Early and Australaspis magnus faunule_ 4 Chorbusulina wilkesi faunule _ _ 5 Middle Cambrian trilobites. ___________________ 9 Chorbusulina subdita faunule _ _ 5 Agnostida__ _ _________-____-_--____-----__---_ 9 Early Middle Cambrian f aunules __ 5 Redlichiida. __-_--------------------------_---- 12 Xystridura mutilinia faunule- _ 5 Corynexochida._________--________-_-_---_----_ -
An Appraisal of the Great Basin Middle Cambrian Trilobites Described Before 1900
An Appraisal of the Great Basin Middle Cambrian Trilobites Described Before 1900 By ALLISON R. PALMER A SHORTER CONTRIBUTION TO GENERAL GEOLOGY GEOLOGICAL SURVEY PROFESSIONAL PAPER 264-D Of the 2ty species described prior to I(?OO, 2/ are redescribed and 2C} refigured, and a new name is proposedfor I species UNITED STATES GOVERNMENT PRINTING OFFICE, WASHINGTON : 1954 UNITED STATES DEPARTMENT OF THE INTERIOR Douglas McKay, Secretary GEOLOGICAL SURVEY W. E. Wrather, Director For sale by the Superintendent of Documents, U. S. Government Printing Office Washington 25, D. C. - Price $1 (paper cover) CONTENTS Page Abstract..__________________________________ 55 Introduction ________________________________ 55 Original and present taxonomic names of species. 57 Stratigraphic distribution of species ____________ 57 Collection localities._________________________ 58 Systematic descriptions.______________________ 59 Literature cited____________________________ 82 Index __-_-__-__---_--______________________ 85 ILLUSTRATIONS [Plates 13-17 follow page 86] PLATE 13. Agnostidae and Dolichometopidae 14. Dorypygidae 15. Oryctocephalidae, Dorypygidae, Zacanthoididae, and Ptychoparioidea 16. Ptychoparioidea 17. Ptychoparioidea FIGUBE 3. Index map showing collecting localities____________________________ . Page 56 in A SHORTER CONTRIBUTION TO GENERAL GEOLOGY AN APPRAISAL OF THE GREAT BASIN MIDDLE CAMBRIAN TRILOBITES DESCRIBED BEFORE 1900 By ALLISON R. PALMER ABSTRACT the species and changes in their generic assignments All 29 species of Middle Cambrian trilobites -
A Solution to Darwin's Dilemma: Differential Taphonomy of Ediacaran and Palaeozoic Non-Mineralised Discoidal Fossils
Provided by the author(s) and NUI Galway in accordance with publisher policies. Please cite the published version when available. Title A Solution to Darwin's Dilemma: Differential Taphonomy of Ediacaran and Palaeozoic Non-Mineralised Discoidal Fossils Author(s) MacGabhann, Breandán Anraoi Publication Date 2012-08-29 Item record http://hdl.handle.net/10379/3406 Downloaded 2021-09-26T20:57:04Z Some rights reserved. For more information, please see the item record link above. A Solution to Darwin’s Dilemma: Differential taphonomy of Palaeozoic and Ediacaran non- mineralised discoidal fossils Volume 1 of 2 Breandán Anraoi MacGabhann Supervisor: Dr. John Murray Earth and Ocean Sciences, School of Natural Sciences, NUI Galway August 2012 Differential taphonomy of Palaeozoic and Ediacaran non-mineralised fossils Table of Contents List of Figures ........................................................................................................... ix List of Tables ........................................................................................................... xxi Taxonomic Statement ........................................................................................... xxiii Acknowledgements ................................................................................................ xxv Abstract ................................................................................................................. xxix 1. Darwin’s Dilemma ............................................................................................... -
Chemostratigraphic Correlations Across the First Major Trilobite
www.nature.com/scientificreports OPEN Chemostratigraphic correlations across the frst major trilobite extinction and faunal turnovers between Laurentia and South China Jih-Pai Lin 1*, Frederick A. Sundberg2, Ganqing Jiang3, Isabel P. Montañez4 & Thomas Wotte5 During Cambrian Stage 4 (~514 Ma) the oceans were widely populated with endemic trilobites and three major faunas can be distinguished: olenellids, redlichiids, and paradoxidids. The lower–middle Cambrian boundary in Laurentia was based on the frst major trilobite extinction event that is known as the Olenellid Biomere boundary. However, international correlation across this boundary (the Cambrian Series 2–Series 3 boundary) has been a challenge since the formal proposal of a four-series subdivision of the Cambrian System in 2005. Recently, the base of the international Cambrian Series 3 and of Stage 5 has been named as the base of the Miaolingian Series and Wuliuan Stage. This study provides detailed chemostratigraphy coupled with biostratigraphy and sequence stratigraphy across this critical boundary interval based on eight sections in North America and South China. Our results show robust isotopic evidence associated with major faunal turnovers across the Cambrian Series 2–Series 3 boundary in both Laurentia and South China. While the olenellid extinction event in Laurentia and the gradual extinction of redlichiids in South China are linked by an abrupt negative carbonate carbon excursion, the frst appearance datum of Oryctocephalus indicus is currently the best horizon to achieve correlation between the two regions. Te international correlation of the traditional lower–middle Cambrian boundary has been exceedingly difcult primarily due to apparent diachroniety of the datum species used to defne the boundary refecting the endemic faunas. -
10. Brief Notes on the Eodiscids 11, Phylogeny of the Dawsonidea
No. 1.] 43 10. Brief Notes on the Eodiscids 11, Phylogeny of the Dawsonidea. By Teiichi KoBAYASHI. (Comm. by H. YABE,M.I.A., Jan. 12, 1943.) Microdiscus has long been in common use for eodiscids, but as pointed out by Barrande in 1881, Microdiscus quadricostatus Emmons which is the type species of the genus, was a larval form of Trinucleus which was 'found in association with Ordovician graptolites and pele- cypods. Because Microdiscus Emmons, 1885, and the Microdiscidae Koken or Coquin, 196, lost their standing, Raymond adopted Eodiscus Hartt in place of Microdiscus and established Eodiscidae to include, beside the genus, two new ones, Goniodiscus and Weymouthia. While the three genera were all blind, Delgado described eodiscids with eyes in 1904 and Vogdes founded Delgadoia on Microdiscus caudatus Delgado in 1917. One year prior to this, Walcott had described proparian Pagetia as a member of the Eodiscidae, which became a subject of debate among trilobite students. Poulsen mentioned in 1927 that the sole reason for the resemblance of Pagetia to hypoparian eodiscids is adaptative convergence. This opinion is intenable because it is now known in Pagetia, Delgadoia or Al emtejoia that the two forms of each genus which are almost identical except for the presence or the absence of eyes and facial sutures, are found at the same locality. Supporting Beecher's view on the migration of the facial suture, Raymond emphasized in 1917, that Pagetia shows the stage in which the suture just came up to the dorsal side from the ventral in the agnostidian stage, notwithstanding the fact that Dindstrom and others have proven that the agnostid has no suture on the ventral side of its cephalon.