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Open Thesis Final V2.Pdf
The Pennsylvania State University The Graduate School Department of the Geosciences TAXONOMIC AND ECOLOGIC IMPLICATIONS OF MAMMOTH MOLAR MORPHOLOGY AS MEASURED VIA COMPUTED TOMOGRAPHY A Thesis in Geosciences by Gregory J Smith 2015 Gregory J Smith Submitted in Partial Fulfillment of the Requirements for the Degree of Master of Science August 2015 ii The thesis of Gregory J Smith was reviewed and approved* by the following: Russell W. Graham EMS Museum Director and Professor of the Geosciences Thesis Advisor Mark Patzkowsky Professor of the Geosciences Eric Post Director of the Polar Center and Professor of Biology Timothy Ryan Associate Professor of Anthropology and Information Sciences and Technology Michael Arthur Professor of the Geosciences Interim Associate Head for Graduate Programs and Research *Signatures are on file in the Graduate School iii ABSTRACT Two Late Pleistocene species of Mammuthus, M. columbi and M. primigenius, prove difficult to identify on the basis of their third molar (M3) morphology alone due to the effects of dental wear. A newly-erupted, relatively unworn M3 exhibits drastically different characters than that tooth would after a lifetime of wear. On a highly-worn molar, the lophs that comprise the occlusal surface are more broadly spaced and the enamel ridges thicken in comparison to these respective characters on an unworn molar. Since Mammuthus taxonomy depends on the lamellar frequency (# of lophs/decimeter of occlusal surface) and enamel thickness of the third molar, given the effects of wear it becomes apparent that these taxonomic characters are variable throughout the tooth’s life. Therefore, employing static taxonomic identifications that are based on dynamic attributes is a fundamentally flawed practice. -
1.1 První Chobotnatci 5 1.2 Plesielephantiformes 5 1.3 Elephantiformes 6 1.3.1 Mammutida 6 1.3.2 Elephantida 7 1.3.3 Elephantoidea 7 2
MASARYKOVA UNIVERZITA PŘÍRODOVĚDECKÁ FAKULTA ÚSTAV GEOLOGICKÝCH VĚD Jakub Březina Rešerše k bakalářské práci Využití mikrostruktur klů neogenních chobotnatců na příkladu rodu Zygolophodon Vedoucí práce: doc. Mgr. Martin Ivanov, Dr. Brno 2012 OBSAH 1. Současný pohled na evoluci chobotnatců 3 1.1 První chobotnatci 5 1.2 Plesielephantiformes 5 1.3 Elephantiformes 6 1.3.1 Mammutida 6 1.3.2 Elephantida 7 1.3.3 Elephantoidea 7 2. Kly chobotnatců a jejich mikrostruktura 9 2.1 Přírůstky v klech chobotnatců 11 2.1.1 Využití přírůstků v klech chobotnatců 11 2.2 Schregerův vzor 12 2.2.1 Stavba Schregerova vzoru 12 2.2.2 Využití Schregerova vzoru 12 2.3 Dentinové kanálky 15 3 Sedimenty s nálezy savců v okolí Mikulova 16 3.1 Baden 17 3.2 Pannon a Pont 18 1. Současný pohled na evoluci chobotnatců Současná systematika chobotnatců není kompletně odvozena od jejich fylogeneze, rekonstruované pomocí kladistických metod. Diskutované skupiny tak mnohdy nepředstavují monofyletické skupiny. Přestože jsou taxonomické kategorie matoucí (např. Laurin 2005), jsem do jisté míry nucen je používat. Některým skupinám úrovně stále přiřazeny nebyly a zde této skutečnosti není přisuzován žádný význam. V této rešerši jsem se zaměřil hlavně na poznatky, které následovaly po vydání knihy; The Proboscidea: Evolution and Paleoecology of Elephants and Their Relatives, od Shoshaniho a Tassyho (1996). Chobotnatci jsou součástí skupiny Tethytheria společně s anthracobunidy, sirénami a desmostylidy (Shoshani 1998; Shoshani & Tassy 1996; 2005; Gheerbrant & Tassy 2009). Základní klasifikace sestává ze dvou skupin. Ze skupiny Plesielephantiformes, do které patří čeledě Numidotheriidae, Barytheriidae a Deinotheridae a ze skupiny Elephantiformes, do které patří čeledě Palaeomastodontidae, Phiomiidae, Mammutida, Gomphotheriidae, tetralofodontní gomfotéria, Stegodontidae a Elephantidae (Shoshani & Marchant 2001; Shoshani & Tassy 2005; Gheerbrant & Tassy 2009). -
A Report on Late Quaternary Vertebrate Fossil Assemblages from the Eastern San Francisco Bay Region, California
PaleoBios 30(2):50–71, October 19, 2011 © 2011 University of California Museum of Paleontology A report on late Quaternary vertebrate fossil assemblages from the eastern San Francisco Bay region, California SUSUMU TOMIYA*,1,2,3, JENNY L. MCGUIRE1,2,3, RUSSELL W. DEDON1, SETH D. LERNER1, RIKA SETSUDA1,3, ASHLEY N. LIPPS1, JEANNIE F. BAILEY1, KELLY R. HALE1, ALAN B. SHABEL1,2,3, AND ANTHONY D. BARNOSKY1,2,3 1Department of Integrative Biology, 2Museum of Paleontology, 3Museum of Vertebrate Zoology, University of California, Berkeley, CA 94720, USA; mailing address: University of California Berkeley, 3060 Valley Life Sciences Building # 3140, Berkeley, CA 94720-3140, USA; email: [email protected] Here we report on vertebrate fossil assemblages from two late Quaternary localities in the eastern San Francisco Bay region, Pacheco 1 and Pacheco 2. At least six species of extinct mammalian megaherbivores are known from Pacheco 1. The probable occurrence of Megalonyx jeffersonii suggests a late Pleistocene age for the assemblage. Pacheco 2 has yielded a minimum of 20 species of mammals, and provides the first unambiguous Quaternary fossil record ofUrocyon , Procyon, Antrozous, Eptesicus, Lasiurus, Sorex ornatus, Tamias, and Microtus longicaudus from the San Francisco Bay region. While a radiocarbon date of 405 ± 45 RCYBP has been obtained for a single bone sample from Pacheco 2, the possibility that much of the assemblage is considerably older than this date is suggested by (1) the substantial loss of collagen in all other samples for which radiocarbon dating was unsuccessfully attempted and (2) the occurrence of Microtus longicaudus approximately 160 km to the west of, and 600 m lower in elevation than, its present range limit. -
The Easternmost Occurrence of Mammut Pacificus (Proboscidea: Mammutidae), Based on a Partial Skull from Eastern Montana, USA
The easternmost occurrence of Mammut pacificus (Proboscidea: Mammutidae), based on a partial skull from eastern Montana, USA Andrew T. McDonald1, Amy L. Atwater2, Alton C. Dooley Jr1 and Charlotte J.H. Hohman2,3 1 Western Science Center, Hemet, CA, United States of America 2 Museum of the Rockies, Montana State University, Bozeman, MT, United States of America 3 Department of Earth Sciences, Montana State University, Bozeman, MT, United States of America ABSTRACT Mammut pacificus is a recently described species of mastodon from the Pleistocene of California and Idaho. We report the easternmost occurrence of this taxon based upon the palate with right and left M3 of an adult male from the Irvingtonian of eastern Montana. The undamaged right M3 exhibits the extreme narrowness that characterizes M. pacificus rather than M. americanum. The Montana specimen dates to an interglacial interval between pre-Illinoian and Illinoian glaciation, perhaps indicating that M. pacificus was extirpated in the region due to habitat shifts associated with glacial encroachment. Subjects Biogeography, Evolutionary Studies, Paleontology, Zoology Keywords Mammut pacificus, Mammutidae, Montana, Irvingtonian, Pleistocene INTRODUCTION Submitted 7 May 2020 The recent recognition of the Pacific mastodon (Mammut pacificus (Dooley Jr et al., 2019)) Accepted 3 September 2020 as a new species distinct from and contemporaneous with the American mastodon Published 16 November 2020 (M. americanum) revealed an unrealized complexity in North American mammutid Corresponding author evolution during the Pleistocene. Dooley Jr et al. (2019) distinguished M. pacificus from Andrew T. McDonald, [email protected] M. americanum by a suite of dental and skeletal features: (1) upper third molars (M3) Academic editor and lower third molars (m3) much narrower relative to length in M. -
The American Mastodon Mammut Americanum in Mexico
The American Mastodon Mammut americanum in Mexico O.J. Polaco1, J. Arroyo-Cabrales1, E. Corona-M.1, J.G. López-Oliva2 1Laboratorio de Paleozoología, Instituto Nacional de Antropología e Historia, D.F. México - [email protected], [email protected], [email protected] 2Facultad de Ciencias de la Tierra, Universidad Autónoma de Nuevo León, Linares, México - [email protected] SUMMARY: Aim of this study is to clarify the occurrence of the American mastodon Mammut americanum at the southern portion of its distribution. Twenty-three specimens from 15 localities in Mexico have been recorded, most of them from the Central Plateau, up to an altitude of 1500 meters above sea level, and to the east of the country. In addition, the mammoth Mammuthus and the American mastodon are contemporaneous in three localities, and mastodon possible association with man in two localities. Their southernmost record is from Honduras. Our data indicate that the American mastodon is a rare taxon at the southern portion of its distribution, which could be related to the presence and possible feeding overlap with the gomphothere Cuvieronius. 1. INTRODUCTION Central America (Honduras; Lucas & Alvarado 1991). However, the presence of From 1996 to1998, an electronic database American Mastodon is not clearly document- on Late Quaternary mammals of Mexico was ed or is relatively scarce in the fossil record developed with a similar structure to the from México, as it is only known from few FAUNMAP project in the United States of localities in the Mexican Plateau (Polaco et al. America. The purpose was to gather, verify 1998). -
Filling a Gap in the Proboscidean Fossil Record: a New Genus from The
Filling a gap in the proboscidean fossil record: a new genus from the Lutetian of Senegal Rodolphe Tabuce, Raphaël Sarr, Sylvain Adnet, Renaud Lebrun, Fabrice Lihoreau, Jeremy Martin, Bernard Sambou, Mustapha Thiam, Lionel Hautier To cite this version: Rodolphe Tabuce, Raphaël Sarr, Sylvain Adnet, Renaud Lebrun, Fabrice Lihoreau, et al.. Filling a gap in the proboscidean fossil record: a new genus from the Lutetian of Senegal. Journal of Paleontology, Paleontological Society, 2019, pp.1-9. 10.1017/jpa.2019.98. hal-02408861 HAL Id: hal-02408861 https://hal.archives-ouvertes.fr/hal-02408861 Submitted on 8 Dec 2020 HAL is a multi-disciplinary open access L’archive ouverte pluridisciplinaire HAL, est archive for the deposit and dissemination of sci- destinée au dépôt et à la diffusion de documents entific research documents, whether they are pub- scientifiques de niveau recherche, publiés ou non, lished or not. The documents may come from émanant des établissements d’enseignement et de teaching and research institutions in France or recherche français ou étrangers, des laboratoires abroad, or from public or private research centers. publics ou privés. 1 Filling a gap in the proboscidean fossil record: a new genus from 2 the Lutetian of Senegal 3 4 Rodolphe Tabuce1, Raphaël Sarr2, Sylvain Adnet1, Renaud Lebrun1, Fabrice Lihoreau1, Jeremy 5 E. Martin2, Bernard Sambou3, Mustapha Thiam3, and Lionel Hautier1 6 7 1Institut des Sciences de l’Evolution, UMR5554, CNRS, IRD, EPHE, Université de 8 Montpellier, Montpellier, France <[email protected]> 9 <[email protected]> <[email protected]> 10 <[email protected]> <[email protected] > 11 2Univ. -
La Brea and Beyond: the Paleontology of Asphalt-Preserved Biotas
La Brea and Beyond: The Paleontology of Asphalt-Preserved Biotas Edited by John M. Harris Natural History Museum of Los Angeles County Science Series 42 September 15, 2015 Cover Illustration: Pit 91 in 1915 An asphaltic bone mass in Pit 91 was discovered and exposed by the Los Angeles County Museum of History, Science and Art in the summer of 1915. The Los Angeles County Museum of Natural History resumed excavation at this site in 1969. Retrieval of the “microfossils” from the asphaltic matrix has yielded a wealth of insect, mollusk, and plant remains, more than doubling the number of species recovered by earlier excavations. Today, the current excavation site is 900 square feet in extent, yielding fossils that range in age from about 15,000 to about 42,000 radiocarbon years. Natural History Museum of Los Angeles County Archives, RLB 347. LA BREA AND BEYOND: THE PALEONTOLOGY OF ASPHALT-PRESERVED BIOTAS Edited By John M. Harris NO. 42 SCIENCE SERIES NATURAL HISTORY MUSEUM OF LOS ANGELES COUNTY SCIENTIFIC PUBLICATIONS COMMITTEE Luis M. Chiappe, Vice President for Research and Collections John M. Harris, Committee Chairman Joel W. Martin Gregory Pauly Christine Thacker Xiaoming Wang K. Victoria Brown, Managing Editor Go Online to www.nhm.org/scholarlypublications for open access to volumes of Science Series and Contributions in Science. Natural History Museum of Los Angeles County Los Angeles, California 90007 ISSN 1-891276-27-1 Published on September 15, 2015 Printed at Allen Press, Inc., Lawrence, Kansas PREFACE Rancho La Brea was a Mexican land grant Basin during the Late Pleistocene—sagebrush located to the west of El Pueblo de Nuestra scrub dotted with groves of oak and juniper with Sen˜ora la Reina de los A´ ngeles del Rı´ode riparian woodland along the major stream courses Porciu´ncula, now better known as downtown and with chaparral vegetation on the surrounding Los Angeles. -
The Distribution of Proboscidea (Elephants) Professor Dr
The Distribution of Proboscidea (Elephants) Professor Dr. Erich Thenius [In: Kosmos #5, May, pp. 235-242, 1964, Stuttgart] When I speak here about animals with a trunk, I do not mean the tapirs or pigs, but I refer only to the elephants and their ancestors, like the Mastodons and Dinotheria which we call the Proboscidea (after the Greek: proboscis = trunk). Their main characteristic is their remarkable trunk which has been fashioned to become a “gripping” organ. That organ was not present in the geologically oldest ancestors whose skeletons stem from the deposits of the Eocene (old Tertiary) in Africa. Even though we have no “soft tissues” of those animals, their skeletal features suffice to tell the scientist just what their bodily characteristics would have been. Thus also, we are not really going to discuss much about their distribution in historic times, but rather, we will concentrate on the development of these characteristic mammals, from their inception to their distribution in the past. A history of the Proboscidea is necessarily a history of their distribution in time and space. Information of these animals is available from numerous fossil findings in nearly all continents. But, before we even consider the fossil history, let us take a quick look of the current distribution of elephants which is shown in Figure 1. Nowadays, there are only two species of elephants: the Indian and African elephants. They not only differ geographically but also morphologically. That is to say, they are different in their bodily form and in their anatomy in several characteristics as every attentive zoo visitor who sees them side-by-side easily observes: The small-eared Indian elephant (Elephas maximus) has a markedly bowed upper skull; the African cousin (Loxodonta africana) has longer legs and markedly larger ears. -
Educator Guide Presented by the Field Museum
at the San Diego Natural History Museum July 4-November 11, 2013 Educator Guide Presented by The Field Museum INSIDE: Exhibition Introduction • Planning Your Visit Gallery Overviews and Guiding Questions • Focused Field Trip Activities Correlations to California State Content Standards • Additional Resources Mammoths and Mastodons: Titans of the Ice Age at the San Diego Natural History Museum is supported by: City of San Diego Commission for Arts and Culture County of San Diego Board of Supervisors, Community Enhancement Program Walter J. and Betty C. Zable Foundation Qualcomm Foundation The Kenneth T. and Eileen L. Norris Foundation VWR Charitable Foundation Education Sponsor: The Field Museum gratefully acknowledges the collaboration and assistance of the Shemanovskii Regional Museum and Exhibition Complex and the International Mammoth Committee. Walking Map The Field Museum • Mammoths and Mastodons Educator Guide • fieldmuseum.org/mammoths Page 2 www.sdnhm.org/mammoths-mastodons Exhibition Introduction Mammoths and Mastodons: Titans of the Ice Age July 4–November 11, 2013 Millions of years ago, colossal mammals roamed Europe, Asia, and North America. From the gigantic mammoth to the massive mastodon, these creatures have captured the world’s fascination. Meet “Lyuba,” the best-preserved baby mammoth in the world, and discover all that we’ve learned from her. Journey back to the Ice Age through monumental video installations, roam among saber-toothed cats and giant bears, and wonder over some of the oldest human artifacts in existence. Hands-on exciting interactive displays reveal the difference between a mammoth and a mastodon, This sketch shows a Columbian mammoth, an offer what may have caused their extinction, and show African elephant, and an American mastodon how today’s scientists excavate, analyze, and learn more (from back to front). -
(Mammut Americanum) from Fulton County, Indiana
University of Nebraska - Lincoln DigitalCommons@University of Nebraska - Lincoln USGS Staff -- Published Research US Geological Survey 1-2008 The Overmyer Mastodon (Mammut americanum) from Fulton County, Indiana Neal Woodman USGS Patuxent Wildlife Research Center, [email protected] Jon W. Branstrator Earlham College, [email protected] Follow this and additional works at: https://digitalcommons.unl.edu/usgsstaffpub Woodman, Neal and Branstrator, Jon W., "The Overmyer Mastodon (Mammut americanum) from Fulton County, Indiana" (2008). USGS Staff -- Published Research. 581. https://digitalcommons.unl.edu/usgsstaffpub/581 This Article is brought to you for free and open access by the US Geological Survey at DigitalCommons@University of Nebraska - Lincoln. It has been accepted for inclusion in USGS Staff -- Published Research by an authorized administrator of DigitalCommons@University of Nebraska - Lincoln. The American Midland Naturalist January 2008 : Vol. 159, Issue 1 (Jan 2008), pg(s) 125-146 Am. Midl. Nat. 159:125–146 The Overmyer Mastodon (Mammut americanum) from Fulton County, Indiana NEAL WOODMAN1 USGS Patuxent Wildlife Research Center, National Museum of Natural History MRC-111, Smithsonian Institution, Washington, DC 20013 AND JON W. BRANSTRATOR2 Geosciences Department, Earlham College, Richmond, Indiana 47374 ABSTRACT.—In June 1978 the partial skeleton of an American mastodon, Mammut americanum, was salvaged from a drainage ditch in Fulton County, north-central Indiana. The remains were recovered mostly from ca. 170–260 cm below the current land surface in marl overlain by peat and peaty marl. The stratigraphy of the site indicates that the remains were deposited in a small, open-water pond that subsequently filled. The skeleton, which is 41–48% complete, is that of a mature female, ca. -
Paleobiogeography of Trilophodont Gomphotheres (Mammalia: Proboscidea)
Revista Mexicana deTrilophodont Ciencias Geológicas, gomphotheres. v. 28, Anúm. reconstruction 2, 2011, p. applying235-244 DIVA (Dispersion-Vicariance Analysis) 235 Paleobiogeography of trilophodont gomphotheres (Mammalia: Proboscidea). A reconstruction applying DIVA (Dispersion-Vicariance Analysis) María Teresa Alberdi1,*, José Luis Prado2, Edgardo Ortiz-Jaureguizar3, Paula Posadas3, and Mariano Donato1 1 Departamento de Paleobiología, Museo Nacional de Ciencias Naturales, CSIC, José Gutiérrez Abascal 2, 28006, Madrid, España. 2 INCUAPA, Departamento de Arqueología, Universidad Nacional del Centro, Del Valle 5737, B7400JWI Olavarría, Argentina. 3 LASBE, Facultad de Ciencias Naturales y Museo, Universidad Nacional de La Plata, Paseo del Bosque S/Nº, B1900FWA La Plata, Argentina. * [email protected] ABSTRACT The objective of our paper was to analyze the distributional patterns of trilophodont gomphotheres, applying an event-based biogeographic method. We have attempted to interpret the biogeographical history of trilophodont gomphotheres in the context of the geological evolution of the continents they inhabited during the Cenozoic. To reconstruct this biogeographic history we used DIVA 1.1. This application resulted in an exact solution requiring three vicariant events, and 15 dispersal events, most of them (i.e., 14) occurring at terminal taxa. The single dispersal event at an internal node affected the common ancestor to Sinomastodon plus the clade Cuvieronius – Stegomastodon. A vicariant event took place which resulted in two isolated groups: (1) Amebelodontinae (Africa – Europe – Asia) and (2) Gomphotheriinae (North America). The Amebelodontinae clade was split by a second vicariant event into Archaeobelodon (Africa and Europe), and the ancestors of the remaining genera of the clade (Asia). In contrast, the Gomphotheriinae clade evolved mainly in North America. -
Remains of Zygolophodon Turicensis (Proboscidea, Mammutidae) from the Coal Mines Near Bitola, Republic of Macedonia
Remains of Zygolophodon turicensis (Proboscidea, Mammutidae) from the coal mines... 157 Historia naturalis bulgarica, 20: 157-162, 2012 Remains of Zygolophodon turicensis (Proboscidea, Mammutidae) from the coal mines near Bitola, Republic of Macedonia Risto GAREVSKI, Biljana GAREVSKA, Georgi N. MARKOV GAREVSKI R., GAREVSKA B., MARKOV G.N. 2012. Remains of Zygolophodon turicensis (Proboscidea, Mammutidae) from the coal mines near Bitola, Republic of Macedonia. – Historia naturalis bulgarica, 20: 157-162. Abstract. We report Zygolophodon turicensis (Schinz, 1824) from the Miocene deposits in a coal mine near Bitola, Republic of Macedonia, and refer a misidentified molar from Nerezi near Skopje published in the 1930s to the same species. Zygolophodon turicensis is a new taxon to the fossil fauna of Macedonia, and the finds discussed in the paper are among the few fossils of pre-Turolian age from the country. Key words: Proboscidea, Mammutidae, Zygolophodon, Miocene, Macedonia Introduction The specimens described below are an accidental find from the coal mines of the thermal power station near Bitola in the southwest of the Republic of Macedonia. Found in sandy deposits, the molars most probably belong to the same individual. Material and methods Material: Left and right m2-m3, coal mine near Bitola, stored at the Bitola Natural History Museum (BiNHM), coll. no. 13536. Methods: Dental nomenclature follows TASSY (1996). Measurements (in mm) taken by B. Garevska. Systematic palaeontology Order Proboscidea Illiger, 1811 Family Mammutidae Hay, 1922 Genus Zygolophodon Vacek, 1877 Zygolophodon turicensis (Schinz, 1824) All four molars (Fig.1, Fig. 2) are in a rather good condition. The left m2 is preserved very well.