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Evidence of Lingual-Luring by an Aquatic Snake
Journal of Herpetology, Vol. 34 No. 1 pp 67-74, 2000 Copyright 2000 Society for the Study of Amphibians and Reptiles Evidence of Lingual-luring by an Aquatic Snake HARTWELL H. WELSH, JR. AND AMY J. LIND Pacific Southwest Research Station, USDA Forest Service, 1700 Bayview Dr., Arcata, California 95521, USA. E-mail: hwelsh/[email protected] ABSTRACT.-We describe and quantify the components of an unusual snake behavior used to attract fish prey: lingual-luring. Our earlier research on the foraging behavior of the Pacific Coast aquatic garter snake (Thamnophis atratus) indicated that adults are active foragers, feeding primarily on aquatic Pacific giant salamanders (Dicamptodon tenebrosus) in streambed substrates. Juvenile snakes, however, use primarily ambush tactics to capture larval anurans and juvenile salmonids along stream margins, behaviors that include the lingual-luring described here. We found that lingual-luring differed from typical chemosensory tongue-flicking by the position of the snake, contact of the tongue with the water surface, and the length of time the tongue was extended. Luring snakes are in ambush position and extend and hold their tongues out rigid, with the tongue-tips quivering on the water surface, apparently mimicking insects in order to draw young fish within striking range. This behavior is a novel adaptation of the tongue-vomeronasal system by a visually-oriented predator. The luring of prey by snakes has been asso- luring function (Mushinsky, 1987; Ford and ciated primarily with the use of the tail, a be- Burghardt, 1993). However, Lillywhite and Hen- havior termed caudal luring (e.g., Neill, 1960; derson (1993) noted the occurrence of a pro- Greene and Campbell, 1972; Heatwole and Dav- longed extension of the tongue observed in vine ison, 1976; Jackson and Martin, 1980; Schuett et snakes (e.g., Kennedy, 1965; Henderson and al., 1984; Chizar et al., 1990). -
Methane Cold Seeps As Biological Oases in the High‐
LIMNOLOGY and Limnol. Oceanogr. 00, 2017, 00–00 VC 2017 The Authors Limnology and Oceanography published by Wiley Periodicals, Inc. OCEANOGRAPHY on behalf of Association for the Sciences of Limnology and Oceanography doi: 10.1002/lno.10732 Methane cold seeps as biological oases in the high-Arctic deep sea Emmelie K. L. A˚ strom€ ,1* Michael L. Carroll,1,2 William G. Ambrose, Jr.,1,2,3,4 Arunima Sen,1 Anna Silyakova,1 JoLynn Carroll1,2 1CAGE - Centre for Arctic Gas Hydrate, Environment and Climate, Department of Geosciences, UiT The Arctic University of Norway, Tromsø, Norway 2Akvaplan-niva, FRAM – High North Research Centre for Climate and the Environment, Tromsø, Norway 3Division of Polar Programs, National Science Foundation, Arlington, Virginia 4Department of Biology, Bates College, Lewiston, Maine Abstract Cold seeps can support unique faunal communities via chemosynthetic interactions fueled by seabed emissions of hydrocarbons. Additionally, cold seeps can enhance habitat complexity at the deep seafloor through the accretion of methane derived authigenic carbonates (MDAC). We examined infaunal and mega- faunal community structure at high-Arctic cold seeps through analyses of benthic samples and seafloor pho- tographs from pockmarks exhibiting highly elevated methane concentrations in sediments and the water column at Vestnesa Ridge (VR), Svalbard (798 N). Infaunal biomass and abundance were five times higher, species richness was 2.5 times higher and diversity was 1.5 times higher at methane-rich Vestnesa compared to a nearby control region. Seabed photos reveal different faunal associations inside, at the edge, and outside Vestnesa pockmarks. Brittle stars were the most common megafauna occurring on the soft bottom plains out- side pockmarks. -
8.4 the Significance of Ocean Deoxygenation for Continental Margin Mesopelagic Communities J
8.4 The significance of ocean deoxygenation for continental margin mesopelagic communities J. Anthony Koslow 8.4 The significance of ocean deoxygenation for continental margin mesopelagic communities J. Anthony Koslow Institute for Marine and Antarctic Studies, University of Tasmania, Hobart, Tasmania, Australia and Scripps Institution of Oceanography, University of California, SD, La Jolla, CA 92093 USA. Email: [email protected] Summary • Global climate models predict global warming will lead to declines in midwater oxygen concentrations, with greatest impact in regions of oxygen minimum zones (OMZ) along continental margins. Time series from these regions indicate that there have been significant changes in oxygen concentration, with evidence of both decadal variability and a secular declining trend in recent decades. The areal extent and volume of hypoxic and suboxic waters have increased substantially in recent decades with significant shoaling of hypoxic boundary layers along continental margins. • The mesopelagic communities in OMZ regions are unique, with the fauna noted for their adaptations to hypoxic and suboxic environments. However, mesopelagic faunas differ considerably, such that deoxygenation and warming could lead to the increased dominance of subtropical and tropical faunas most highly adapted to OMZ conditions. • Denitrifying bacteria within the suboxic zones of the ocean’s OMZs account for about a third of the ocean’s loss of fixed nitrogen. Denitrification in the eastern tropical Pacific has varied by about a factor of 4 over the past 50 years, about half due to variation in the volume of suboxic waters in the Pacific. Continued long- term deoxygenation could lead to decreased nutrient content and hence decreased ocean productivity and decreased ocean uptake of carbon dioxide (CO2). -
Coastal and Marine Ecological Classification Standard (2012)
FGDC-STD-018-2012 Coastal and Marine Ecological Classification Standard Marine and Coastal Spatial Data Subcommittee Federal Geographic Data Committee June, 2012 Federal Geographic Data Committee FGDC-STD-018-2012 Coastal and Marine Ecological Classification Standard, June 2012 ______________________________________________________________________________________ CONTENTS PAGE 1. Introduction ..................................................................................................................... 1 1.1 Objectives ................................................................................................................ 1 1.2 Need ......................................................................................................................... 2 1.3 Scope ........................................................................................................................ 2 1.4 Application ............................................................................................................... 3 1.5 Relationship to Previous FGDC Standards .............................................................. 4 1.6 Development Procedures ......................................................................................... 5 1.7 Guiding Principles ................................................................................................... 7 1.7.1 Build a Scientifically Sound Ecological Classification .................................... 7 1.7.2 Meet the Needs of a Wide Range of Users ...................................................... -
AUV Adaptive Sampling Methods: a Review
applied sciences Review AUV Adaptive Sampling Methods: A Review Jimin Hwang 1 , Neil Bose 2 and Shuangshuang Fan 3,* 1 Australian Maritime College, University of Tasmania, Launceston 7250, TAS, Australia; [email protected] 2 Department of Ocean and Naval Architectural Engineering, Memorial University of Newfoundland, St. John’s, NL A1C 5S7, Canada; [email protected] 3 School of Marine Sciences, Sun Yat-sen University, Zhuhai 519082, Guangdong, China * Correspondence: [email protected] Received: 16 July 2019; Accepted: 29 July 2019; Published: 2 August 2019 Abstract: Autonomous underwater vehicles (AUVs) are unmanned marine robots that have been used for a broad range of oceanographic missions. They are programmed to perform at various levels of autonomy, including autonomous behaviours and intelligent behaviours. Adaptive sampling is one class of intelligent behaviour that allows the vehicle to autonomously make decisions during a mission in response to environment changes and vehicle state changes. Having a closed-loop control architecture, an AUV can perceive the environment, interpret the data and take follow-up measures. Thus, the mission plan can be modified, sampling criteria can be adjusted, and target features can be traced. This paper presents an overview of existing adaptive sampling techniques. Included are adaptive mission uses and underlying methods for perception, interpretation and reaction to underwater phenomena in AUV operations. The potential for future research in adaptive missions is discussed. Keywords: autonomous underwater vehicle(s); maritime robotics; adaptive sampling; underwater feature tracking; in-situ sensors; sensor fusion 1. Introduction Autonomous underwater vehicles (AUVs) are unmanned marine robots. Owing to their mobility and increased ability to accommodate sensors, they have been used for a broad range of oceanographic missions, such as surveying underwater plumes and other phenomena, collecting bathymetric data and tracking oceanographic dynamic features. -
Manual on Hydrometry
Government of India & Government of The Netherlands DHV CONSULTANTS & DELFT HYDRAULICS with HALCROW, TAHAL, CES, ORG & JPS VOLUME 4 HYDROMETRY DESIGN MANUAL Design Manual – Hydrometry (SW) Volume 4 Table of Contents 1 INTRODUCTION 1 1.1 GENERAL 1 1.2 DEFINITION OF VARIABLES AND UNITS 2 2 PHYSICS OF RIVER FLOW 5 2.1 GENERAL 5 2.2 CLASSIFICATION OF FLOWS 5 2.3 VELOCITY PROFILES 8 2.4 HYDRAULIC RESISTANCE 10 2.5 UNSTEADY FLOW 14 2.6 BACKWATER CURVES 17 3 HYDROMETRIC NETWORK DESIGN 21 3.1 INTRODUCTION 21 3.2 NETWORK DESIGN CONSIDERATIONS 22 3.2.1 CLASSIFICATION 22 3.2.2 MINIMUM NETWORKS 22 3.2.3 NETWORKS FOR LARGE RIVER BASINS 23 3.2.4 NETWORKS FOR SMALL RIVER BASINS 23 3.2.5 NETWORKS FOR DELTAS AND COASTAL FLOODPLAINS 23 3.2.6 REPRESENTATIVE BASINS 24 3.2.7 SUSTAINABILITY 24 3.2.8 DUPLICATION AVOIDANCE 24 3.2.9 PERIODIC RE-EVALUATION 24 3.3 NETWORK DENSITY 24 3.3.1 WMO RECOMMENDATIONS 24 3.3.2 PRIORITISATION SYSTEM 25 3.3.3 STATISTICAL AND MATHEMATICAL OPTIMISATION 26 3.4 THE NETWORK DESIGN PROCESS 26 4 SITE SELECTION OF WATER LEVEL AND STREAMFLOW STATIONS 28 4.1 DEFINITION OF OBJECTIVES 28 4.2 DEFINITION OF CONTROLS 28 4.3 SITE SURVEYS 29 4.4 SELECTION OF WATER LEVEL GAUGING SITES 31 4.5 SELECTION OF STREAMFLOW MEASUREMENT SITE 31 5 MEASURING FREQUENCY 34 5.1 GENERAL 34 5.2 STAGE MEASUREMENT FREQUENCY 35 5.3 CURRENT METER MEASUREMENT FREQUENCY 36 6 MEASURING TECHNIQUES 38 6.1 STAGE MEASUREMENT 38 6.1.1 GENERAL 38 6.1.2 VERTICAL STAFF GAUGES 39 6.1.3 INCLINED STAFF OR RAMP GAUGES 42 6.1.4 CREST STAGE GAUGES 43 6.1.5 ELECTRIC TAPE GAUGES 44 -
MARINE ENVIRONMENTS Teaching Module for Grades 6-12
MARINE ENVIRONMENTS Teaching Module for Grades 6-12 Dear Educator, We are pleased to present you with the first in a series of teaching and learning modules developed by the DEEPEND (Deep-Pelagic Nekton Dynamics) consortium and their consultants. DEEPEND is a research network focusing primarily on the pelagic zone of the Gulf of Mexico, therefore the majority of the lessons will be based around this topic. Whenever possible, the lessons will focus specifically on events of the Gulf of Mexico or work from the DEEPEND scientists. All modules in this series aim to engage students in grades 6 through 12 in STEM disciplines, while promoting student learning of the marine environment. We hope these lessons enable teachers to address student misconceptions and apprehensions regarding the unique organisms and properties of marine ecosystems. We intend for these modules to be a guide for teaching. Teachers are welcome to use the lessons in any order they wish, use just portions of lessons, and may modify the lessons as they wish. Furthermore, educators may share these lessons with other school districts and teachers; however, please do not receive monetary gain for lessons in any of the modules. Moreover, please provide credit to photographers and authors whenever possible. This first module focuses on the marine environment in general including biological, chemical, and physical properties of the water column. We have provided a variety of activities and extensions within this module such that lessons can easily be adapted for various grade and proficiency levels. Given that education reform strives to incorporate authentic science experiences, many of these lessons encourage exploration and experimentation to encourage students to think and act like a scientist. -
Aging Techniques & Population Dynamics of Blue Suckers (Cycleptus Elongatus) in the Lower Wabash River
Eastern Illinois University The Keep Masters Theses Student Theses & Publications Summer 2020 Aging Techniques & Population Dynamics of Blue Suckers (Cycleptus elongatus) in the Lower Wabash River Dakota S. Radford Eastern Illinois University Follow this and additional works at: https://thekeep.eiu.edu/theses Part of the Aquaculture and Fisheries Commons Recommended Citation Radford, Dakota S., "Aging Techniques & Population Dynamics of Blue Suckers (Cycleptus elongatus) in the Lower Wabash River" (2020). Masters Theses. 4806. https://thekeep.eiu.edu/theses/4806 This Dissertation/Thesis is brought to you for free and open access by the Student Theses & Publications at The Keep. It has been accepted for inclusion in Masters Theses by an authorized administrator of The Keep. For more information, please contact [email protected]. AGING TECHNIQUES & POPULATION DYNAMICS OF BLUE SUCKERS (CYCLEPTUS ELONGATUS) IN THE LOWER WABASH RIVER By Dakota S. Radford B.S. Environmental Biology Eastern Illinois University A thesis prepared for the requirements for the degree of Master of Science Department of Biological Sciences Eastern Illinois University May 2020 TABLE OF CONTENTS Thesis abstract .................................................................................................................... iii Acknowledgements ............................................................................................................ iv List of Tables .......................................................................................................................v -
Lake Ecology
Fundamentals of Limnology Oxygen, Temperature and Lake Stratification Prereqs: Students should have reviewed the importance of Oxygen and Carbon Dioxide in Aquatic Systems Students should have reviewed the video tape on the calibration and use of a YSI oxygen meter. Students should have a basic knowledge of pH and how to use a pH meter. Safety: This module includes field work in boats on Raystown Lake. On average, there is a death due to drowning on Raystown Lake every two years due to careless boating activities. You will very strongly decrease the risk of accident when you obey the following rules: 1. All participants in this field exercise will wear Coast Guard certified PFDs. (No exceptions for teachers or staff). 2. There is no "horseplay" allowed on boats. This includes throwing objects, splashing others, rocking boats, erratic operation of boats or unnecessary navigational detours. 3. Obey all boating regulations, especially, no wake zone markers 4. No swimming from boats 5. Keep all hands and sampling equipment inside of boats while the boats are moving. 6. Whenever possible, hold sampling equipment inside of the boats rather than over the water. We have no desire to donate sampling gear to the bottom of the lake. 7. The program director has final say as to what is and is not appropriate safety behavior. Failure to comply with the safety guidelines and the program director's requests will result in expulsion from the program and loss of Field Station privileges. I. Introduction to Aquatic Environments Water covers 75% of the Earth's surface. We divide that water into three types based on the salinity, the concentration of dissolved salts in the water. -
Chapter 101 Minnesota Statutes 1941
MINNESOTA STATUTES 1941 101.01 DIVISION OF GAME AND FISH; FISH 846 CHAPTER 101 DIVISION OF GAME AND FISH; FISH Sec. Sec. 101.01 Manner of taking flsh 101.21 Sale of flsh caught In certain counties; o).her 101.02 Manner of taking minnows for bait flsh not bought or sold at any time 101.03 Open season for black bass and yellow bass 101.22 Prohibited methods and equipments 101.04 Open season for trout, except lake trout; 101.23 Polluting streams hours for taking 101.24 Fish screens; removal 101.05 Fishing in trout streams 101.25 Dark houses or fish houses, when used; 101.06 Open season for lake trout licenses 101.07 Open season for pike, pickerel, and muskel- 101.26 Open season for whiteflsh, tullibees, and her lunge ring 101.08 Closed season for sturgeon, hackleback, 101.27 Open season for frogs spoonbill, or paddleflsh. 101.28 Turtles and tortoises 101.09 Open season for crappies 101.29 Fishways; construction; fishing near flshways 101.10 Fishing in boundary waters forbidden 101.11 Open season for fishing in boundary waters 101.30 Fish may be taken and sold from certain lakes 101.12 Open season for sunflsh, rock bass, and other 10.1.31 Regulations by director / varieties 101.32 Restriction 101.13 Open season for sunflsh in Goodhue county 101.34 Sections 101.30 to 101.32 supplementary 101.14 Open season for carp, dogfish, redhorse, 101.35 Disposition of dead flsh sheepshead, catfish, suckers, eelpout, garfish, 101.36 Open season for fishing in Lake of the Woods bullheads, whiteflsh, and buffaloflsh 101.37 Open season for suckers and other rough flsh 101.153 Propagation of game flsh in Lake of the Woods 101.16 When and where artificial lights may be used 101.38 Fishing from towed boats prohibited in spearing certain fish 101.39 Taking of fish in natural spawning beds 101.18 Placing carp in waters prohibited prohibited 101.19 Fishing in Minneapolis 101.40 Fish screens; permits 101.20 Limit of catch 101.01 MANNER OF TAKING FISH. -
Real-Time Monitoring for Toxicity Caused by Harmful Algal Blooms and Other Water Quality Perturbations EPA/600/R-01/103 November 2001
United States Office of Research and EPA/600/R-01/103 Environmental Protection Development November 2001 Agency Washington, DC 20460 Real-Time Monitoring for Toxicity Caused By Harmful Algal Blooms and Other Water Quality Perturbations EPA/600/R-01/103 November 2001 Real-Time Monitoring for Toxicity Caused By Harmful Algal Blooms and Other Water Quality Perturbations National Center for Environmental Assessment-Washington Office Office of Research and Development U.S. Environmental Protection Agency Washington, DC DISCLAIMER This document has been reviewed in accordance with U.S. Environmental Protection Agency policy and approved for publication. Mention of trade names or commercial products does not constitute endorsement or recommendation for use. ABSTRACT This project, sponsored by EPA’s Environmental Monitoring for Public Access and Community Tracking (EMPACT) program, evaluated the ability of an automated biological monitoring system that measures fish ventilatory responses (ventilatory rate, ventilatory depth, and cough rate) to detect developing toxic conditions in water. In laboratory tests, acutely toxic levels of both brevetoxin (PbTx-2) and toxic Pfiesteria piscicida cultures caused fish responses primarily through large increases in cough rate. In the field, the automated biomonitoring system operated continuously for 3 months on the Chicamacomico River, a tributary to the Chesapeake Bay that has had a history of intermittent toxic algal blooms. Data gathered through this effort complemented chemical monitoring data collected by the Maryland Department of Natural Resources (DNR) as part of their pfiesteria monitoring program. After evaluation by DNR personnel, the public could access the data at a DNR Internet website, (www.dnr.state.md.us/bay/pfiesteria/00results.html), or receive more detailed information at aquaticpath.umd.edu/empact. -
Hydrological Measurements
Water Quality Monitoring - A Practical Guide to the Design and Implementation of Freshwater Quality Studies and Monitoring Programmes Edited by Jamie Bartram and Richard Ballance Published on behalf of United Nations Environment Programme and the World Health Organization © 1996 UNEP/WHO ISBN 0 419 22320 7 (Hbk) 0 419 21730 4 (Pbk) Chapter 12 - HYDROLOGICAL MEASUREMENTS This chapter was prepared by E. Kuusisto. Hydrological measurements are essential for the interpretation of water quality data and for water resource management. Variations in hydrological conditions have important effects on water quality. In rivers, such factors as the discharge (volume of water passing through a cross-section of the river in a unit of time), the velocity of flow, turbulence and depth will influence water quality. For example, the water in a stream that is in flood and experiencing extreme turbulence is likely to be of poorer quality than when the stream is flowing under quiescent conditions. This is clearly illustrated by the example of the hysteresis effect in river suspended sediments during storm events (see Figure 13.2). Discharge estimates are also essential when calculating pollutant fluxes, such as where rivers cross international boundaries or enter the sea. In lakes, the residence time (see section 2.1.1), depth and stratification are the main factors influencing water quality. A deep lake with a long residence time and a stratified water column is more likely to have anoxic conditions at the bottom than will a small lake with a short residence time and an unstratified water column. It is important that personnel engaged in hydrological or water quality measurements are familiar, in general terms, with the principles and techniques employed by each other.