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Home , Epialtinae, Ethusidae, Eubrachyura, Majoidea, Monte di Malo, Orgiano

This is anopenaccess article undertheCCBY-NC-SAThis Universidad NacionalAutónoma de México. Peer Reviewing under the responsibility of Manuscript accepted: Corrected manuscript received: Manuscript received: BSGM2021v73n3a120221 http://dx.doi.org/10.18268/A120221. de la Sociedad Geológica Mexicana, 73 (3), (Monti Berici, , NEItaly): Boletín San Feliciano from the late of fossil(, Brachyura) report of De Angeli, A.,Garassino, A.,2021,New How tocitethisarticle: [email protected] * Corresponding author:(A.Garassino) 92350, USA. es, LomaLindaUniversity, LomaLinda,CA 2 giore, Vicenza, . 1 Antonio (, MonteBérico,Vicenza,NEItalia) Nuevo reporte de cangrejos fósiles (Decapoda, Brachyura) del Eoceno tardío en San Hill Feliciano Feliciano Hill(Orgiano,MontiBerici,Vicenza,NEItaly) New report of fossilcrabs (Decapoda, Brachyura) fromthelate Eocene ofSan license(https://creativecommons.org/licenses/by-nc-sa/4.0/) Museo CivicoG.Museo Zannato, Montecchio Mag eateto Earth andBiological Scienc of Department De Angeli 1 , Alessandro Boletín de la Sociedad Geológica Mexicana Geológica Sociedad la de Boletín - - Garassino Eocene, Italy. , ,late Brachyura, ,Dairidae, Keywords:Crustacea, Decapoda, lacking thefronto-orbital margin. the holotype, original description of charactersto the morphological incertae sedis 1991 (Xanthoidea MacLeay, 1838, to well-preserved assigned specimens Berici, Orgiano. Moreover, two San Feliciano Hill, located Monti in Rodriguez, 1986) are reported from bericus MacLeay, 1838)and bretoni Herein, two newcrabs, (2002, 2014). Angeli andGarassino partially Italy) was recorded byDe (Orgiano, MontiBerici, Vicenza, NE San Feliciano hill the late Eocene of richThe decapodassemblage from ABSTRACT Actaeites lobatus n. gen., n. sp. (Epialtidae n.gen., n. sp. (Dairidae Ng and 2,* ) allowed to add some Müller and Collins, Orgianocarcinus Bericirinia /73(3)A120221/2021 Garassino (2002, 2014).Enesta zona hay parcialmenteregistrado por Angeli y De Monte Bérico, Vicenza, NE Italia) fue San Felicianotardío del cerro (Orgiano, El rico conjunto dedecápodos del Eoceno RESUMEN tardío, Italia. Xanthoidea, taxonomia, Eoceno Brachyura, Epialtidae, Dairidae, Palabras Decapoda, clave: Crustacea, frontoorbitario.margen ción original del holotipo donde se carece del algunos caracteres morfológicos a la descrip 1838, y Collins, 1991 (Xanthoidea MacLeay, asignados para y dos ejemplares estan bien conservados del Monte Bérico.Orgiano Además estos SanFeliciano,dos enelcerro localizadoen (Dairidae Ng yRodríguez, 1986)encontra y gen., n.sp. (Epialtidae MacLeay, 1838) cangrejos,dos nuevos Orgianocarcinus bericus incertae sedis Actaeites lobatus ) lo que permitió añadir ) lo que permitió Bericirinia bretoni n. gen., n.sp. Müller 1 n. - -

Fossil crabs (Decapoda, Brachyura) from the late Eocene of San Feliciano, Italy ABSTRACT INTRODUCTION AND GEOLOGICAL Fossil crabs (Decapoda, Brachyura) from the late Eocene of San Feliciano, Italy SETTING / MATERIAL 1. Introductionandgeologicalsetting Figure 1 2 2 and coralsfrom the late Eocene (Priabonian), within the limestones including coralline algae (Figure studied specimens are 1). The preserved MontiBericibetween Orgiano andLonigo side of San Feliciano hill is located on the southwestern including coralline algae, corals, rare mollusc thick, overlap the lower layers. calcarenites These 1822)]. White-yellowishcalcarenites, 6-7 meters [ bivalves, several andrare echinoderms, decapod including manyformation algae, nummulites, greyish marly-limestonewell-stratified by a sented present in the quarry. lowest The layers are repre layers all for age Priabonian the confirmed (2003) fossil analysisby nannofossilsprovided by Beccaro Priabona” (late Eocene, micro Priabonian). The unitisthe so-called “Formazionegraphic di Berici, Vicenza, NEItaly) (Figure strati 2).The located in San Feliciano hill (Orgiano, Monti / / Mapof ofSan Feliciano MontiBerici hill(Orgiano, withlocation(*) MontiBerici, Vicenza,NE Italy). Boletín de la Sociedad Geológica Mexicana Geológica Sociedad la de Boletín Boletín de la Sociedad Geológica Mexicana Geológica Sociedad la de Boletín Palaeocarpilius macrocheilus http://dx.doi.org/10.18268/BSGM2021v73n3a120221 (Desmarest, /73(3)A120221/2021 - - - /73(3)A1202212021 2. Material quarry. the were collected from thiscalcarenite level of thisnote, studiedspecimens,2016). The subjectof Lovato, 2009;DeAngeli Garassino, 2002,2014,inpress;De Angeliand several (Fabiani, authors 1911; De Angeli and lected within the calcarenites were described by Manyechinoderms. decapod crustaceans col including nummulites, bryozoans, bivalves, and upper part is represented by marly limestones and DeAngeli,2019).Finally,Quaggiotto the casts, anddecapodcrustaceans(DeAngeli,2016; tofrontal margin width;wf:frontal width. carapace length; wcxp:carapace width; wof: orbi in millimetres. Anatomical abbreviations –lcxp: (Vicenza, sizes NEItaly) (MCV). areThe expressed Dal Lago”of “Domenico Museo Civico materialThe includes 10specimens, housedinthe et al. , 2010a;DeAngeli, - - 3. Systematicpaleontology Figure 2 Schweitzer Discussion: monotypy. Type species: specimen wascollected.Gender: feminine. Etymology: surface withascabrous-shaped smallgranules. with sometubercles andovate swellings; dorsal regions; cardiac, gastric, andbranchial regions lobes and protogastricrunning between epigastric grooves; frontal region withanaxial depression sal regions slightlyraisedandmarked byweak posterolateral margins, strongly convergent; dor divergent, smooth anterolateral margins; short laterally directed; intercalated spine notpresent; byU-shaped notch; preorbital blunt; angle orbits separated bifid rostrum, front;short, wide dinally; orly, considerablywider thanlong, convex longitu Diagnosis: Close-up ofthelimestonesincluding coralligenous algaeand coralsfromthelateEocene (Priabonian)of SanFeliciano hill. Section deSaintLaurent, 1980 Subsection Guinot,1977 Superfamily MajoideaSamouelle, 1819 Subfamily EpialtinaeMacLeay, 1838 Infraorder Brachyura Latreille, 1802 Family EpialtidaeMacLeay, 1838 Order DecapodaLatreille, 1802 et al. Triangular carapace, larger posteri from MontiBerici where the studied Based upon Davie

(2020), Bericirinia bretoni Bericirinia Boletín de la Sociedad Geológica Mexicana Mexicana Mexicana Geológica Geológica Sociedad Sociedad la la de de Boletín Boletín Bericirinia n.gen. http://dx.doi.org/10.18268/BSGM2021v73n3a120221 n. gen., n. sp., by et al. n.gen.shows (2015) and - - - éadBeo 14-00,i eonto fGérard Breton (1944-2020), in recognition of Etymology: Diagnosis: rostrum.elongate single havingin thecarapace longer thanwiderand Panticarcinus regionsand dorsal withswellings and tubercles. teriorly; oblique, divergent anterolateral margins; genera share the triangular carapace, larger pos shows affinities with UKisthe only genus which Eocene (Ypresian) of ( and Saward, 2006)andoneextant-fossilgenus Müller and Collins, 1991; 2016; Beschin, BusuliniandTessier fossil five genera ( includes which Epialtidae the within and the providedclassificationnew of a altidae MacLeay, 1838.Schweitzer spine allow tothe assign new genus to the Epi intercalated the of lacking the and rostrum bifid and orbits laterally directed. Moreover, the short, pace, larger posteriorly; narrow frontal margin; manyspider crabs, such asthe triangular cara characters shared with the main morphological Eoinachoides Dana,1851), Epialtus Bericirinia bretoni , however, differs from the new genus new the from however,differs , asfor thegenus. /73(3)A120221/2021 / 73(3)A120221/2021 teseisi ae nhnu fof named inhonour the species is H.MilneEdwards,1834; Van Straelen, 1933; Bericirinia Figure 3 Panticarcinus n. gen.,sp. n.gen. Indeed, both Panticarcinus in Beschin from the early et al. Nanomaja Collins Bolcapisa (2020) et al. 3 3 - - - ,

Fossil crabs (Decapoda, Brachyura) from the late Eocene of San Feliciano, Italy SYSTEMATIC PALEONTOLOGY Fossil crabs (Decapoda, Brachyura) from the late Eocene of San Feliciano, Italy SYSTEMATIC PALEONTOLOGY 4. frontal view.Scale bar equals 5mm. Figure 3 4 4 Geological age: Berici, Vicenza, NEItaly). Type locality: monotypy. Holotype: andinvertebratepalaentology fossils. the major contributionshis tothe knowledge of short, bifid rostrum, separated by U-shaped notch; antero- andposterolateral margins;wide front; of long (lcxp/wcxp =0.95),widestbetween angles gitudinally, larger posteriorly, slightlywiderthan Description: 32.8 mm,wcxp: 34.2mm,wof: 18mm). view(MCV.2021/003-I.G.21.23in dorsal –lcxp: Material and measurements: / /

Boletín de la Sociedad Geológica Mexicana Geológica Sociedad la de Boletín Boletín de la Sociedad Geológica Mexicana Geológica Sociedad la de Boletín Bericirinia bretoni MCV.2021/003-I.G.21.23, by Triangular carapace, convex lon San Feliciano hill (Orgiano, Monti late Eocene(Priabonian). n.gen.,sp.,holotype, MCV.21/003-I.G.21.23; 1.dorsalview;2. digitalreconstruction;3.lateral view; http://dx.doi.org/10.18268/BSGM2021v73n3a120221 one carapace /73(3)A120221/2021 - /73(3)A1202212021 anteriorlyby cervical groove withonestrong epi ally, with two swellings; branchial region marked well marked bybranchiocardiac grooves later row, depressed urogastric region; cardiac region regionsraised metagastric with two swellings; nar protogastric regions withthree swellings; relatively lobes withtwo ovateepigastric swellings; raised andprotogastric regions;along epigastric ning depressed axially; frontal axial depression run and marked by shallow grooves; frontal region partially preserved; regions dorsal slightly raised convergent; relatively wide posterior margin, margins;short posterolateral margins, strongly divergent, elongate, anterolateral andsmooth preorbital blunt; angle orbits laterally directed; - - - - Discussion: and wherehave the studied specimens been collected Etymology: sp., by monotypy. Type species: surrounded by smallertubercles. large, obovate or rounded crater-shaped lobules, posterior margin; well-distinct regions dorsal with posterolateral margin with6-7 spines; straight convex fissures; anterolateral marginshort, concave with7spines; narrow two with margin bital axially depressed; round orbitshavingsupraor than long; narrow front, and slightly downturned Diagnosis: brous-shaped smallgranules. metabranchial swelling; surfacewithasca dorsal mesobrachial swelling, andoneweakly transverse branchial swelling, one weakly raised transverse corals. for system camouflage perfect a be could look, coral rounded crater-shaped lobules, resembling the it isconfidentiallyassigned. the Dairidae to which with the representatives of are affinities closer the assignment, systematic its the studied specimen lacks ventral parts, usefulfor in arranged lobules uniformly regionsdorsal withmanysmall mushroom-shaped obovate or rounded crater-shaped tubercles (vs. however,having in regionslargedorsal withsome Angeli several species (De Angeli andGarassino, Veneto1833, well known from the Paleogene with of are shared with the extant andfossil and antero- andposterlateral margins with spines orbital margin; bilobate front, slightly downturned; the orbits, with raised, tuberculate supra shape of Ng andRodriguez, 1986. Dairidae the of those fit specimens studied the of The ornamentation withlarge obovate ornamentation The or Orgianocarcinus Family DairidaeNgandRodriguez, 1986 carcinus Superfamily Dairoidea Serène, 1965 et al. =. Gender:masculine. , 2019). Genus Obovate, convex carapace, wider The main morphological characters mainmorphological The from SanFeliciano hillOrgiano, Orgianocarcinus bericus n.gen. which lived within the Orgianocarcinus Orgianocarcinus Boletín de la Sociedad Geológica Mexicana Mexicana Mexicana Geológica Geológica Sociedad Sociedad la la de de Boletín Boletín Indeed, the rounded Daira http://dx.doi.org/10.18268/BSGM2021v73n3a120221 nov. Daira ). Althoughthe n. gen. differs, gen. n. n. gen., n. n. 2006;De De Haan, - - - Berici, Vicenza, NEItaly). Type locality: Geological age: 010-I.G.21.30. 007-I.G.21.27; MCV.21/009-I.G.21.29; MCV.21/ 005-I.G.21.25;MCV.21/006-I.G.21.26; MCV.21/ Paratypes: Holotype: discovered. Berici wherehave thestudiedspecimens been Etymology: Diagnosis: f6-7tubercles; triangular hepatic regions well of depressed intestinal region with a transverse row with two lobesanteriorly andonelobe posteriorly; one large ovate lobe; subpentagonal cardiac region ovate lobe; well-distinct region metagastric with regionelongate mesogastric withonelongitudinal, protogastric regions withtubercles; subtriangular, lobes with two ovate swellings; slightly raised with medial longitudinal depression;epigastric regions well marked by grooves; frontal region straight posterior margin, as wide as front; dorsal concave posterolateral margin with6-7spines; anterolateral marginshort, slightly with7spines; tubercle; subtriangular extraorbital tooth; convex with one tubercle; medial orbital tooth with one angle preorbital fissures; narrow two with margin axially depressed; roundorbits; raisedsupraorbital protruded beyond orbits, and slightly downturned maximum carapace width; front weakly of half (lcxp/wcxp = 0.71); wide orbitofrontal margin sections, above all longitudinally, widerthanlong Description: I.G.21.30 –lcxp:5;wcxp: 7). MCV.21/009-I.G.21.29 –wcxp: 8;MCV.21/010- wcxp: 6.9; MCV.21/008-I.G.21.28 –wcxp: 9.4; wof: 5.8; MCV.21/007-I.G.21.27 –lcxp:4.9; MCV.21/006-I.G.21.26 –lcxp:7.7;wcxp: 10.8; MCV.21/005-I.G.21.25 –lcxp:8.4;wcxp: 11.8; I.G.21.24 – lcxp:8.2; wcxp: 11.4;wof: 6;wf:4.2; mens withwell-preserved carapace (MCV.21/004- Material and measurements: Orgianocarcinus bericus MCV.21/008-I.G.21.28. asfor thegenus. MCV.21/004-I.G.21.24; MCV.21/ /73(3)A120221/2021 / 73(3)A120221/2021 The trivial name alludes to Monti The Obovate carapace, convex inboth SanFeliciano hill (Orgiano, Monti late Eocene(Priabonian). Figure 4 n.gen.,sp. seven speci 5 5 -

Fossil crabs (Decapoda, Brachyura) from the late Eocene of San Feliciano, Italy SYSTEMATIC PALEONTOLOGY Fossil crabs (Decapoda, Brachyura) from the late Eocene of San Feliciano, Italy SYSTEMATIC PALEONTOLOGY I.G.21.29, dorsalview; 6.paratype,MCV.21/007-I.G.21.27, dorsalview.Scalebar equals 5mm. dorsal view;3.paratype, MCV.21/010-I.G.21.30, dorsalview;4.paratype,MCV.21/006-I.G.21.26, dorsalview;5. paratype,MCV.21/009- Figure 4 6 6 / /

Boletín de la Sociedad Geológica Mexicana Geológica Sociedad la de Boletín Boletín de la Sociedad Geológica Mexicana Geológica Sociedad la de Boletín Orgianocarcinus bericus n.gen.,sp.,1.holotype, MCV.21/008-I.G.21.28,dorsal view;2.paratype,MCV.21/004-I.G.21.24, http://dx.doi.org/10.18268/BSGM2021v73n3a120221 /73(3)A120221/2021 /73(3)A1202212021 bar equals 5mm. Figure 5 50. — Beschin Actaeites lobatus 4c, Pl.4,figs. 9,10. Actaeites lobatus lins, 1991. Fossil species: 1991, by monotypy. Type species: parts notpreserved. lobes. branchial lobes, and three smaller metabranchial chial regions with 4 epibranchial lobes, 4 meso marked bycervical groove withthree lobes; bran — Schweitzer Angeli De — 4. fig. 4, fig.32, Beschin,2008: and Angeli De — 4. Pl. 10, Superfamily XanthoideaMacLeay, 1838(

Cephalic andthoracic appendages and ventralCephalic Actaeites lobatus Genus Actaeites lobatus Actaeites —Karasawa andSchweitzer, 2006: Müller and Collins, 1991: 70, fig. 70, 1991: Collins, and Müller et al. et al. Actaeites lobatus Müllerand Collins,1991, 1.MCV.21/001-I.G.21.21, dorsal view;2.MCV.21/002-I.G.21.22, dorsal view.Scale Actaeites lobatus MüllerandCollins, 1991 MüllerandCollins, 1991 , 2010:129.—Beschin , 2007: 56, Pl. 9, figs. 2a-b, 3, 2a-b, figs. 9, Pl. 56, 2007: , Figure 5 sedis et al. Boletín de la Sociedad Geológica Mexicana Mexicana Mexicana Geológica Geológica Sociedad Sociedad la la de de Boletín Boletín ) , 2010b: 167, fig. 13. fig. 167, 2010b: , Müller and Collins, http://dx.doi.org/10.18268/BSGM2021v73n3a120221 Müller and Col incertae et al. - - - , orbital margins with two fissures; convex antero convex fissures; two with margins orbital separated from thefront by groove;a deep supra small marginal orbits moderatelyspines; wide, than long; bilobate front, depressed axially, with Emended diagnosis: I.G.21.22 –lcxp:15.1,wcxp: 29;wo-f: 14.8;wof: 8). I.G.21.21 – lcxp: 10; wcxp: 12; MCV.21/002- with well-preserved carapace (MCV.21/001- Material and measurements: Berici, NEItaly). Locality: Geological age: b. —DeAngeli Pl. 17, fig. 8. — Beschin fig.7, Beschin Pl. — 93, 4. 2015: Schweitzer, 2006).Fortunately the studied speci thisspecies(Karasawa and tematic of assignment ventral parts did not allow a right sys lacking of orbitofrontalThe margin poorly preserved and the Budapest (Müller and Collins, 1991). bonian of type (MAFI EF-22.1 [M.91-153]) from the Pria the holo characters of upon the morphological Discussion: marked by smoothgrooves. covered by arranged, smalltubercles uniformly the carapace extraorbital regions spine); dorsal of lateral margins withthree (excludingspines the SanFeliciano hill(Orgiano, Monti /73(3)A120221/2021 / 73(3)A120221/2021

Actaeites lobatus et al. late Eocene(Priabonian). 2019:37. Convex carapace, wider et al. , 2018: 196, fig. 128a, wasdescribed based et al. two specimens , 2016:137, 7 7 ------

Fossil crabs (Decapoda, Brachyura) from the late Eocene of San Feliciano, Italy SYSTEMATIC PALEONTOLOGY Fossil crabs (Decapoda, Brachyura) from the late Eocene of San Feliciano, Italy SYSTEMATIC PALEONTOLOGY / CONCLUSIONS / ACKNOWLEDGMENTS / REFERENCES 4. Conclusions 8 8 oee ysaltbrls h oslrgoso covered bysmall tubercles. regions dorsal of The somites. chelipeds The have andpropodus carpus chelipeds, walking legs, and three male pleonal (MCV.21/001-I.G.21.21) preserves incomplete original lacking intheholotype, allowing toamendthe the carapace, preservedmens the anterior part of De Angeliand Lovato, 2009)(Fabiani, 1911; De Garassino,( 2014),andisopods 2016), achelata ( De Angeli,2016; dalpiazi and Alberti,Angeli, Garassino 2010a, ino, 2002], brachyurans ( 2002, 2002, 2002, Longoporcellana lobata petrolisthes blowi thes striatissimus mulleri cantha De Angeli and Garassino, 2002, Palaeomunida defecta 1991, 2002, anomurans [ San Feliciano hill include from the late Eocene of decapodcrustaceans previousThe reports of 2008; De Angeli 2007, 2015,2016,2018;De Angeli andBeschin, di (Vicenza) (Beschin Bernuffiand Soghe of early the from Val LionaandSan Feliciano Hill(Vicenza); and Campolongodi the late Eocene (Priabonian) of Vestenanova and Zovo di Bolca (Verona); from Gecchelina di (Vicenza), sian) of has been recorded from the early Eocene (Ypre marked arranged, byformly smoothgrooves. the carapace are covered bysmall tubercles uni / / Boletín de la Sociedad Geológica Mexicana Geológica Sociedad la de Boletín Boletín de la Sociedad Geológica Mexicana Geológica Sociedad la de Boletín xettePibna fHungary, this species Except the Priabonian of (Müller and Collins, 1991), DeAngeliandGarassino, 2002, A. feldmanni Pisidia dorsosinuosa Spathagalathea minuta Spathagalathea Petrolisthes bittneri diagnosis. Moreover, onestudied specimen Fabiani, 1911, Acanthogalathea parva Galathea berica (Müller and Collins, 1991), DeAngeli and Garassino, 2002, Palinurellus bericus et al. De Angeli and Garassino, 2002, Lőrenthey, 1901, Caporiondolus bericus Caporiondolus DeAngeli and Garassino, , 2010b). DeAngeli and Garassino, De Angeli and Garassino, Eogarthambrus guinotae Eogarthambrus De Angeli & Garassino, DeAngeli and Garass Spathanomus felicianensis Müller and Collins, Sphaeroma gasparellai http://dx.doi.org/10.18268/BSGM2021v73n3a120221 De Angeli and Sadayoshia penta P. multicristata Beripetrolisthes DeAngeli, Phlyctenodes Eopetrolis et al. Lobi De /73(3)A120221/2021 ------/73(3)A1202212021 ,

References Acknowledgements Beschin,2004; Beschin MontiBerici(DeAngeliand side of southeastern duringthelate Eocene onthe acoral reef ence of algae, corals, rare molluscs, pointing out the pres coralline serves the decapod crustaceans isrich of Feliciano fossiliferous hill. The level which pre San thedecapodassemblage of the knowledge of Actaeites lobatus gen., n. sp., Ceccon andDeAngeli(2013). with isopod (bopyrid) infestation was recorded by 2016). Moreover, acarapace of Lovato, 2009;DeAngeli Angeli and Garassino, 2002,2014;DeAngeli and Beschin,Tessier,De Angeli,A., A., C., Busulini, nel stratigrafiche Revisioni 2003, L., Beccaro, review andcriticism. Villalonga, 17B, Tarragona, Catalonia, for careful Coyoacán, Mexico, and ÀlexLlorenç Ossó, de nal Autónoma de México, Ciudad Universitaria, J. Vega, Instituto de Geología, Universidad Nacio SanFeliciano hill;and Francisco of information dell’Università di Padova, for the stratigraphical study;Livia Beccaro, Dipartimento diGeoscienze (Vicenza), tomakeavailable forthe specimens Dal Lago”diValdagno“Domenico Civico We Pallozzi, to wish thank Bernadetta Museo by Hungary Müller and Collins(1991). Eocene of ciano hillwere previouslyreported from the late SanFeli recorded inthe decapodassemblage of Three new brachyurans G., 2007,IDecapodi dell’Eocene inferiore Thesis.Geofisica. Unpublished Dipartimento di Geologia, Paleontologia e XVI°, Università studidiPadova, degli di Ricerca inScienze della Terra, Ciclo Paleocene del occidentale: Dottorato Orgianocarcinus bericus Müller and Collins, 1991)increase et al. et al. , 2018).Some species , 2010a;DeAngeli, Bericirinia bretoni n. gen., n. sp. and Lobipetrolistes blowi n. - - - - Ceccon, L., De Angeli, A., 2013, Segnalazione Beschin, C., Busulini, A.,Tessier, G., Zorzin,R., Beschin,Tessier,A., C., Busulini, G., 2015,Nuova Beschin, C., Busulini, A.,Fornaciari, E., Papazzoni, De Angeli, A., Beschin, C., 2004, De Angeli, A., 2016,Nuovi crostacei Cymonomidae Davie, P.J.F., Guinot,D., Ng, P.K.L., 2015, Dana, J.D., 1851, Conspectus Crustaceorum Collins, J.S.H., Saward, J., 2006,Three new genera associati acorallidell’Eocene superiore di C.A., Tessier, G., 2018,Lafaunadicrostacei Zannato”, 9-76. 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Fossil crabs (Decapoda, Brachyura) from the late Eocene of San Feliciano, Italy REFERENCES