Three New Species of Dactylicapnos (Fumariaceae) and a Synopsis of the D

Total Page:16

File Type:pdf, Size:1020Kb

Three New Species of Dactylicapnos (Fumariaceae) and a Synopsis of the D Nordic Journal of Botany 28: 656Á660, 2010 doi: 10.1111/j.1756-1051.2010.00872.x, # 2010 The Author. Nordic Journal of Botany # 2010 Nordic Society Oikos Subject Editor: Martin Cheek. Accepted 5 November 2010 Three new species of Dactylicapnos (Fumariaceae) and a synopsis of the D. macrocapnos complex Magnus Lide´n Magnus Lide´n ([email protected]), Uppsala Univ. Botanic Gardens, Villava¨gen 8, SE-75236 Uppsala, Sweden. The Dactylicapnos macrocapnos complex is revised, and D. platycarpa Lide´n, D. odontocarpa Lide´n and D. macrocapnos subsp. echinosperma Lide´n are recognised as new taxa. The complex consists of a chain of 4 vicariant PECIAL ISSUE taxa from northwestern India (Garhwal) to western Bhutan (Thimphu). Dactylicapnos cordata Lide´n (eastern Nepal, S Darjeeling) is described and contrasted with its close relative, the geographically disjunct D. burmanica (western Yunnan, Burma). Dactylicapnos Wallich is a very natural and easily circum- flowers are held apex upwards, although they are actually scribed genus, recognised by i.a. the scandent habit with pendent. the terminal leaflet (except of the basalmost leaves) AXONOMY transformed into a branched tendril,9 corymbose pendent T inflorescences, yellow bisymmetric flowers and unique The Dactylicapnos macrocapnos complex stigma, stamen and seed structures. It has at times been (Fig. 1) included in Dicentra Bernh. (Khanh 1972), solely based on the bisymmetric flowers which is a plesiomorphy at family LANT The Dactylicapnos scandens group is characterised by having level. Morphological and molecular data show that P two alternate lateral primary leaflets (opposite in other Dactylicapnos is not closely related to Dicentra, but rather perennial species of the genus) and comparatively thin and the sister group to the genus Corydalis DC (Lide´n et al. soft leaflets (Lide´n 2008). The variable species D. scandens 1997). (D. Don) Hutchinson is vegetatively similar to the Dactylicapnos is distributed in the Sino-Himalayan area D. macrocapnos complex. The latter is nevertheless easily from Garhwal to southern China with some annual distinguished from D. scandens when flowering or fruiting species extending north as far as Sichuan. There are 15 or ASCULAR 16 species in two sections: sect. Dactylicapnos (perennials) by the slightly oblique inflorescence with basally slightly reflexed (vs not reflexed) pedicels, sepals that are abruptly V and sect. Minicalcara (annuals) (Khanh 1972, Lide´n 2007). broadened at the base (vs lanceolate), the strongly up- The perennial species (sect. Dactylicapnos) fall into four curved nectary with long thin apex and usually with a groups: a) the D. grandifoliolata group (four spp.) swelling or wart 1/3 to 2/3 from the base (vs short sigmoid characterised by two opposite primary leaflets with few evenly thick, only shortly acute, not up-curved), lanceolate SIATIC firm strongly veined ultimate leaflets, fleshy fruits opening flat membranous green capsules (vs [when fresh] ovoid A with valves, persistent placentas with tooth-like funiculae, thick fleshy, red yellow or purple), and low broad flat very large easily caducous elaiosomes made up of funiculi (vs thick annular). free tubular cells, and a branched narrow rhizome; Dactylicapnos scandens s.l. is found from central Nepal to b) D. schneideri with two pairs of opposite primary leaflets, southern China (Guizhou) whereas the Dactylicapnos narrow fruits with persistent spongy placentas and seeds macrocapnos complex has an overlapping, but more western with small elaiosomes; c) the D. scandens group (four or distribution from northwestern India (Garhwal) to western five spp, incl. the D. macrocapnos complex, the main Bhutan. subject of this paper); d) the D. burmanica group with two There is a clearcut geograpical variation within the species (one described here). D. macrocapnos complex, with four vicariant taxa along The analysis presented here is based on material at Himalaya. This pattern is not recognised in the excellent ASSAM, BM, CAL, E, K, KATH, PE, TI and UPS. Note monograph by Khanh (1972), as it was revealed by recent that, in the description of floral parts, it is assumed that the collections from eastern Nepal, India and Bhutan. 656 A SIATIC Figure 1. Fruits, seeds, stigmas and flowers of the Dactylicapnos macrocapnos complex (note that flowers and fruits are actually pendent, V but here illustrated with the pedicel attachment at page bottom). (A) D. macrocapnos supsp. macrocapnos (fruit and seed), (B) D. ASCULAR macrocapnos subsp. echinosperma, (C) D. odontocarpa (fruit, seed and stigma), (D) D. platycarpa. Key to the Dactylicapnos macrocapnos complex usually with a median wart below. Capsule narrowly lanceolate, flattened, 25Á356Á7 mm; style 5Á6 mm. 1. Margin of fruit grossly dentate; seeds faintly colliculate, Seeds colliculate to aculeate, 1.6Á1.8 mm; elaiosome small. 1.2mm ................... D. odontocarpa Flowering time: AprÁNov, fruiting time: JulÁNov. P Á Margin of fruit even; seeds sharply colliculate to Ecology: climber in forest understories at an altitudinal LANT aculeate, 1.6Á1.8mm.................... 2 range of 900Á2700 m a.s.l. 2. Capsule twice as long as broad . D. platycarpa Á Capsule 3.5Á5.0 times as long as broad . 3 3. Seeds spinulose/aculeate . D. macrocapnos subsp. Dactylicapnos macrocapnos subsp. macrocapnos T echinosperma Margin of fruit usually without or with indistinct rim; seeds AXONOMY Á Seeds colliculate, but not spinulose . D. macro- 1.6Á1.7 mm, sharply colliculate. capnos subsp. macrocapnos Distributed in Garhwal, Kumaon and extreme west of Nepal. Dactylicapnos macrocapnos (Prain) Hutchinson (1921, p. 105). Dactylicapnos macrocapnos subsp. echinosperma Lide´n subsp. nov. S Basionym: Dicentra macrocapnos Prain (1896, p. 12). Type: A D. macrocapno subsp. macrocapno seminibus aculeatis PECIAL ISSUE India, Garhwal, above Ghat, 2100Á2400 m a.s.l., 20 Sep differt. 1885, J. F. Duthie s.n. (lectotype: DD, isotype: LE; fide Khanh 1972). Type: Nepal, Bakhri Kharka north of Pokhara, 1700 m a.s.l., twining amoung bushes on open slope, ‘‘flowers Perennial with stout subterranean rootstock. Stems 2Á8m, yellow’’, 23 Apr 1954, Stainton, Sykes and Williams 5009 thin and weak, more or less terete (appearing slightly sulcate (holotype: BM!, isotypes: E!, GH!, UPS!). in the dry state). Leaves slightly reflexed on zigzag stem; petiole 0.5Á3.0 cm; blade glaucous below, green above, with Margin of fruit usually with distinct rim; seeds 1.6Á1.8 1 pair of alternate primary leaflets; these bi- (rarely mm, spinulose. It inhabits central Nepal and southern Tibet tri-)ternate; ultimate leaflets ovate, 10Á257Á20 mm. (Nyalam). Raceme 5Á14-flowered, pendent, usually slightly obliquely so; peduncle 2Á4 cm; bracts 3Á101Á2 mm, entire, sub- acute. Pedicels 10Á20 mm, in fruit to 30 mm, reflexed at Additional specimen examined (selection of paratypes) base, sometimes partly fused to subtending bracts. Sepals Tibet: Nyalam, near Zhangmu, 2300 m a.s.l., 13 Aug 1972, 2Á32.0Á2.5 mm, abruptly tapering to narrow apex from unknown 1060 (PE); Zhangmu, 2700 m a.s.l., 24 Aug broad base, slightly dentate. Corolla yellow, often turning 1972, unknown 1472 (PE); Nepal: Lumle west Pokhara, reddish with age, oblong-cordate to obtusely triangular in 1500 m a.s.l., 2 Nov 1954, Stainton et al. 8279 (BM); profile, 18Á208Á12 mm; outer petals strongly keeled or central Nepal, cultivated in Uppsala Univ. Bot. Gard. Aug usually winged at base; sinus at petiole insertion 3 mm deep; 2007, Lide´n s.n. (UPS). dorsal crest of inner petals overtopping apex 1(Á2) mm; This taxon is available from several garden nurseries, nectary strongly curved upwards, tapering to slender apex, often under the name ‘‘Dicentra scandens’’ (Lide´n 2008). 657 Dactylicapnos odontocarpa Lide´n sp. nov. insertion 3Á4 mm deep. Inner petals: claw 10Á13 mm; blade ca 5 mm; dorsal crest overtopping apex 1 mm. A D. macrocapno capsulis irregulariter grosse dentatis semi- Nectary tapering to curved apex, usually with prominent nibus multo minoribus leviter colliculatis differt. gibbosity or peg 1/3Á1/2 as long as distal part of nectary (in D. macrocapnos up to 1/5). Stigma trapezoidal (in D. Type: east Nepal, Janakpur zone (Ramechhap Distr.); macrocapnos subsquarish), with small but distinct papillae in Sagarmatha zone (Solukhumbu Distr.), Bhandar (2300 m apical corners and larger papillae in basal corners. Ovary a.s.l.); Kinja (1650 m a.s.l.); Sheptom (2050 m a.s.l.); with ca 100 ovules, apparently more thick-walled than in D. Nemar (2400 m a.s.l.); Sete (2550 m a.s.l.), in shrubberies, macrocapnos. Capsule broadly elliptic-lanceolate, 22Á23 ‘‘flowers yellow’’, 18 Aug 1985, H. Ohba, M. Wakabaya- 10Á11 mm (in D. macrocapnos 25Á356Á7 mm); style 7Á9 shi, M. Suzuki, N. Kurosaki, K. R. Rajbandhari and S. K. mm. Seeds strongly colliculate, ca 1.6 mm; elaiosome small. Wu 8571764 (holotype: E!, isotypes: KATH, TI). Climbing perennial herbs, with stout subterranean root- Additional specimens examined (paratypes) stock. Stems 2Á8 m (?), thin. Petioles 0.5Á3.0 cm. Leaves Bhutan: Thimphu district, above end of road along with 1 pair of alternate primary leaflets; primary leaflets Thimphu Chu just below Changri monastery, 2500 m biternate; ultimate leaflets 5Á9 per primary leaflet, ovate, a.s.l., 9 Aug 1987, J. R. I. Wood 5675 (E); Thimphu subobtuse, 10Á207Á15 mm, glaucous below. Raceme 1Á district, J. R. I. Wood 6091 (E); Chendebi, among shrubs 3 cm, 5Á8-flowered, pendent. Peduncle 2Á4 cm. Bracts on cliff face, 2700 m a.s.l., 5 Aug 1949, Ludlow, Sherriff linear to narrowly oblanceolate, ca 51 mm, entire, and Hicks 19580 (E, BM, UPS). India: Darjeeling, Senchal subacute. Pedicels 10*20 mm, reflexed at base. Sepals 2Á WLS, way to Tung, 14 Oct 2008, A. K. Ghosh 41501 32.0Á2.5 mm, abruptly tapering to narrow apex from (CAL). PECIAL ISSUE broad base, slightly dentate. Corolla yellow, oblong-cordate S to obtusely triangular, 18Á208Á12 mm. Outer petals keeled or winged at base; sinus at petiole insertion narrow, The Dactylicapnos burmanica group (Fig.
Recommended publications
  • Newsletter 123 May 2012
    TheTheThe Irish Garden Plant Society Newsletter Number 11123123 May 2012 The Annual General Meeting 2012 The Annual General Meeting will be held on Sat 12th May 10.00 a.m for 10.30 a.m., at Hillsborough Courthouse, The Square, Hillsborough, BT26 6AG. As always, it will be followed by a series of garden visits on Saturday & Sunday and a meal on Saturday evening. The meal will be held at 8:00pm in La Mon Hotel & Country Club, 41 Gransha Road, Comber, BT23 5RF. See the January 2012 newsletter for details of the gardens to be visited. If you haven’t already booked contact Patrick Quigley, 24 Areema Drive, Dunmurry, Belfast, BT17 0QG. tel: +44 (0) 7801 299263 [email protected] for further information. A.G. M. Agenda 1. Apologies 2. Minutes of AGM 2011 3. Matters arising 4. Chairman’s report 5. Treasurer’s report 6. Election of Committee Members 7. Any other business Front cover : Solanum crispum ‘Glasnevin’. Photograph : Pearse Rowe In this issue 2 Editorial 3 Southern Climbers for Northern Walls by John Joe Costin 11 Worth a Read by Paddy Tobin 14 The Palm House, a review by Mary Bradshaw 16 Cheers to Chiltern Seeds and to the Sole Survivor of Seed Project 1997/98 by Michael Kelleher 17 Seed Distribution Report 2011 and 2012 by Stephen Butler 19 Regional Reports 28 Spring at Kilmacurragh by Seamus O’Brien 37 Propagation of Arbutus by Kevin Line 41 Tulipa ‘Molly Bloom’ - an new Tulip for 2012 1 Editorial Spring time and new plants are synonymous, and as Christopher Lloyd said in Garden Flowers from Seed, seed sowing is “one of life’s big thrills”.
    [Show full text]
  • Plant List 2016
    Established 1990 PLANT LIST 2016 European mail order website www.crug-farm.co.uk CRÛG FARM PLANTS • 2016 Welcome to our 2016 list hope we can tempt you with plenty of our old favourites as well as some exciting new plants that we have searched out on our travels. There has been little chance of us standing still with what has been going on here in 2015. The year started well with the birth of our sixth grandchild. January into February had Sue and I in Colombia for our first winter/early spring expedition. It was exhilarating, we were able to travel much further afield than we had previously, as the mountainous areas become safer to travel. We are looking forward to working ever closer with the Colombian institutes, such as the Medellin Botanic Gardens whom we met up with. Consequently we were absent from the RHS February Show at Vincent Square. We are finding it increasingly expensive participating in the London shows, while re-branding the RHS February Show as a potato event hardly encourages our type of customer base to visit. A long standing speaking engagement and a last minute change of date, meant that we missed going to Fota near Cork last spring, no such problem this coming year. We were pleasantly surprised at the level of interest at the Trgrehan Garden Rare Plant Fair, in Cornwall. Hopefully this will become an annual event for us, as well as the Cornwall Garden Society show in April. Poor Sue went through the wars having to have a rush hysterectomy in June, after some timely results revealed future risks.
    [Show full text]
  • Diversity and Distribution of Vascular Epiphytic Flora in Sub-Temperate Forests of Darjeeling Himalaya, India
    Annual Research & Review in Biology 35(5): 63-81, 2020; Article no.ARRB.57913 ISSN: 2347-565X, NLM ID: 101632869 Diversity and Distribution of Vascular Epiphytic Flora in Sub-temperate Forests of Darjeeling Himalaya, India Preshina Rai1 and Saurav Moktan1* 1Department of Botany, University of Calcutta, 35, B.C. Road, Kolkata, 700 019, West Bengal, India. Authors’ contributions This work was carried out in collaboration between both authors. Author PR conducted field study, collected data and prepared initial draft including literature searches. Author SM provided taxonomic expertise with identification and data analysis. Both authors read and approved the final manuscript. Article Information DOI: 10.9734/ARRB/2020/v35i530226 Editor(s): (1) Dr. Rishee K. Kalaria, Navsari Agricultural University, India. Reviewers: (1) Sameh Cherif, University of Carthage, Tunisia. (2) Ricardo Moreno-González, University of Göttingen, Germany. (3) Nelson Túlio Lage Pena, Universidade Federal de Viçosa, Brazil. Complete Peer review History: http://www.sdiarticle4.com/review-history/57913 Received 06 April 2020 Accepted 11 June 2020 Original Research Article Published 22 June 2020 ABSTRACT Aims: This communication deals with the diversity and distribution including host species distribution of vascular epiphytes also reflecting its phenological observations. Study Design: Random field survey was carried out in the study site to identify and record the taxa. Host species was identified and vascular epiphytes were noted. Study Site and Duration: The study was conducted in the sub-temperate forests of Darjeeling Himalaya which is a part of the eastern Himalaya hotspot. The zone extends between 1200 to 1850 m amsl representing the amalgamation of both sub-tropical and temperate vegetation.
    [Show full text]
  • Dicentra Formosa
    Plant Propagation Protocol for [Dicentra formosa] ESRM 412 – Native Plant Production Spring 2015 Protocol URL: https://courses.washington.edu/esrm412/protocols/DIFO.pdf Adding information from: http://depts.washington.edu/propplnt/Plants/bleeding_heart.htm North America Distribution Washington State Distribution From the USDA Plants Database6 TAXONOMY Plant Family Scientific Name Fumariaceae Common Name Fumitory Species Scientific Name Scientific Name Dicentra formosa (Haw.) Walp. Varieties Dicentra formosa (Haw.) Walp. var. brevifolia L.F. Hend. Dicentra formosa (Haw.) Walp. var. brevipes L.F. Hend. Sub-species Dicentra formosa (Haw.) Walp. ssp. formosa Dicentra formosa (Haw.) Walp. ssp. oregona (Eastw.) Munz Dicentra formosa (Haw.) Walp. ssp. nevadensis (Eastw.) Munz Cultivar Common Synonym(s) Common Name(s) Bleeding heart, Pacific bleeding heart, Oregon bleeding heart, Sierra bleeding heart Species Code (as per DIFO6 USDA Plants database) GENERAL INFORMATION Geographical range Southern British Colombia to Central California, mid-elevation Cascades and below7; See maps above for North American and Washington State distribution6. Ecological distribution Moist woody to dry open areas, shade preferred7. Climate and elevation Low-middle elevations, mild climate7. range Local habitat and Pseduotsuga menziesii, Tusga heterophylla; typically found in abundance coniferous forests11. Plant strategy type / Late successional plant7. successional stage Plant characteristics Perennial forb growing from a rhizome, pink-purple heart-shaped flowers on leafless stems12. Leaves are divided and fern-like4. PROPAGATION DETAILS Ecotype Propagation Goal Plants Propagation Method Seed5 Product Type Seeds and containers Stock Type Seeds or container plants Propagation Method Vegetative5 Product Type Bareroot and cuttings Stock Type Bareroot Propagule Collection After parent plant has finished blooming and is preparing for Instructions winter, collect seeds for storage; plant seeds in late fall.
    [Show full text]
  • (Largeflower Triteleia): a Technical Conservation Assessment
    Triteleia grandiflora Lindley (largeflower triteleia): A Technical Conservation Assessment © 2003 Ben Legler Prepared for the USDA Forest Service, Rocky Mountain Region, Species Conservation Project January 29, 2007 Juanita A. R. Ladyman, Ph.D. JnJ Associates LLC 6760 S. Kit Carson Cir E. Centennial, CO 80122 Peer Review Administered by Society for Conservation Biology Ladyman, J.A.R. (2007, January 29). Triteleia grandiflora Lindley (largeflower triteleia): a technical conservation assessment. [Online]. USDA Forest Service, Rocky Mountain Region. Available: http://www.fs.fed.us/r2/ projects/scp/assessments/triteleiagrandiflora.pdf [date of access]. ACKNOWLEDGMENTS The time spent and the help given by all the people and institutions mentioned in the References section are gratefully acknowledged. I would also like to thank the Colorado Natural Heritage Program for their generosity in making their files and records available. I also appreciate access to the files and assistance given to me by Andrew Kratz, USDA Forest Service Region 2. The data provided by the Wyoming Natural Diversity Database and by James Cosgrove and Lesley Kennes with the Natural History Collections Section, Royal BC Museum were invaluable in the preparation of the assessment. Documents and information provided by Michael Piep with the Intermountain Herbarium, Leslie Stewart and Cara Gildar of the San Juan National Forest, Jim Ozenberger of the Bridger-Teton National Forest and Peggy Lyon with the Colorado Natural Heritage Program are also gratefully acknowledged. The information provided by Dr. Ronald Hartman and B. Ernie Nelson with the Rocky Mountain Herbarium, Teresa Prendusi with the Region 4 USDA Forest Service, Klara Varga with the Grand Teton National Park, Jennifer Whipple with Yellowstone National Park, Dave Dyer with the University of Montana Herbarium, Caleb Morse of the R.L.
    [Show full text]
  • A Dissertation Submitted for Partial Fulfillment Of
    DIVERSITY OF NATURALIZED PLANT SPECIES ACROSS LAND USE TYPES IN MAKWANPUR DISTRICT, CENTRAL NEPAL A Dissertation Submitted for Partial Fulfillment of the Requirmentment for the Master‟s Degree in Botany, Institute of Science and Technology, Tribhuvan University, Kathmandu, Nepal Submitted by Bhawani Nyaupane Exam Roll No.:107/071 Batch: 2071/73 T.U Reg. No.: 5-2-49-10-2010 Ecology and Resource Management Unit Central Department of Botany Institute of Science and Technology Tribhuvan University Kirtipur, Kathamndu, Nepal May, 2019 RECOMMENDATION This is to certify that the dissertation work entitled “DIVERSITY OF NATURALIZED PLANT ACROSS LAND USE TYPES IN MAKWANPUR DISTRICT, CENTRAL NEPAL” has been submitted by Ms. Bhawani Nyaupane under my supervision. The entire work is accomplished on the basis of Candidate‘s original research work. As per my knowledge, the work has not been submitted to any other academic degree. It is hereby recommended for acceptance of this dissertation as a partial fulfillment of the requirement of Master‘s Degree in Botany at Institute of Science and Technology, Tribhuvan University. ………………………… Supervisor Dr. Bharat Babu Shrestha Associate Professor Central Department of Botany TU, Kathmandu, Nepal. Date: 17th May, 2019 ii LETTER OF APPROVAL The M.Sc. dissertation entitled “DIVERSITY OF NATURALIZED PLANT SPECIES ACROSS LAND USE TYPES IN MAKWANPUR DISTRICT, CENTRAL NEPAL” submitted at the Central Department of Botany, Tribhuvan University by Ms. Bhawani Nyaupane has been accepted as a partial fulfillment of the requirement of Master‘s Degree in Botany (Ecology and Resource Management Unit). EXAMINATION COMMITTEE ………………………. ……………………. External Examiner Internal Examiner Dr. Rashila Deshar Dr. Anjana Devkota Assistant Professor Associate Professor Central Department of Environmental Science Central Department of Botany TU, Kathmandu, Nepal.
    [Show full text]
  • A Checklist of Vascular Plants Endemic to California
    Humboldt State University Digital Commons @ Humboldt State University Botanical Studies Open Educational Resources and Data 3-2020 A Checklist of Vascular Plants Endemic to California James P. Smith Jr Humboldt State University, [email protected] Follow this and additional works at: https://digitalcommons.humboldt.edu/botany_jps Part of the Botany Commons Recommended Citation Smith, James P. Jr, "A Checklist of Vascular Plants Endemic to California" (2020). Botanical Studies. 42. https://digitalcommons.humboldt.edu/botany_jps/42 This Flora of California is brought to you for free and open access by the Open Educational Resources and Data at Digital Commons @ Humboldt State University. It has been accepted for inclusion in Botanical Studies by an authorized administrator of Digital Commons @ Humboldt State University. For more information, please contact [email protected]. A LIST OF THE VASCULAR PLANTS ENDEMIC TO CALIFORNIA Compiled By James P. Smith, Jr. Professor Emeritus of Botany Department of Biological Sciences Humboldt State University Arcata, California 13 February 2020 CONTENTS Willis Jepson (1923-1925) recognized that the assemblage of plants that characterized our flora excludes the desert province of southwest California Introduction. 1 and extends beyond its political boundaries to include An Overview. 2 southwestern Oregon, a small portion of western Endemic Genera . 2 Nevada, and the northern portion of Baja California, Almost Endemic Genera . 3 Mexico. This expanded region became known as the California Floristic Province (CFP). Keep in mind that List of Endemic Plants . 4 not all plants endemic to California lie within the CFP Plants Endemic to a Single County or Island 24 and others that are endemic to the CFP are not County and Channel Island Abbreviations .
    [Show full text]
  • PERENNIAL PLANTS Plant Name Common Name Height Colour Bl Time Special Conditions Country S
    PERENNIAL PLANTS Plant Name Common Name Height Colour Bl Time Special Conditions Country S. Europe, NW Acanthus mollis Bear's Breeches to 5' (1.5m) white fls. with purple shaded bracts l summer z7 sun/pt.shade,well drained, moist good soil Africa Acanthus spinosus Bear's Breeches to 5' (150cm) white flowers with purple bracts lsp-msum z5 sun/pt.shade, good soil, tolerates dry heat Italy to W Turkey Aconitum Monkshood large dark blue flowers l summer z5 sun/part shade, cool moist fertile soil Aconitum Monkshood dark blue flowers l summer z5 sun/part shade, cool moist fertile soil Monkshood (all parts are Aconitum carmichaelii to 6' (190cm) violet or blue flowers l sp to fall z3 sun/pt.shade, cool, moist, fertile soil Russia poisonous) Monkshood (all parts are Aconitum carmichaelii 'Barker's Variety' to 6' (190cm) deep violet flowers early fall z3 sun/pt.shade, cool, moist, fertile soil poisonous) Aconitum 'Ivorine' (syn.A.septentrionale Monkshpood (all parts are to 36" (90cm) ivory flowers l spring z5 sun/pt.shade, cool, moist, fertile soil garden origin 'Ivorine') poisonous) Aconitum lycoctonum ssp.vulparia Monkshood (all parts are to 5' (1.5m) pale yellow flowered form sum/e fall z4 sun/pt.shade, cool, moist, fertile soil Europe (A.orientale of gardens) poisonous) Acorus gramineus 'Variegatus' Variegated Japanese Rush to 10" (25 cm) creamy white and green striped leaves summer z5 full sun, wet or very moist soil E Asia z4 shade/pt.sh.moist mod-fertile soil.Survives under Actaea erythrocarpa (syn. A.spicata var. rubra) 24" (60cm) racemes of white flowers,red berries late spring Euro.
    [Show full text]
  • COMPREHENSIVE CROP REPORTS Purchased the Plant from a Nursery in Increased Dry Weight, Or Both Plants Remain Vegetative and May flower Shanghai
    Bleeding heart is native to the cool, moist woodlands of northeast- ern Asia, growing in dappled shade. It does not bloom well in dense shade and will not tolerate poorly drained soils, especially in the winter. It is adapted to U.S. Department of Agri- Comprehensive culture hardiness zones 2 to 9. Plants can tolerate more sun in the more northern latitudes if kept reliably moist. Partial shade is required in Crop Reports the south or where summer temper- atures are high. High, diffuse light under cool, moist, well-drained con- ditions in a neutral to slightly alkaline soil high in humus are optimal con- Bleeding Heart: A Review for Growers ditions. Plants need to be protected from wind. Plants can form clumps 1 m across and almost as tall. Anec- Laurie Hodges1 dotal reports indicate stems approach- ing 2 m height. If plants are cut back in midsummer, no new leaves will form ADDITIONAL INDEX WORDS. Dicentra spectabilis, Lamprocapnos spectabilis, perennial, cut flower, floriculture, breeding (Hanchek, 1989), whereas cutting flowering stems in the spring will re- SUMMARY. A favorite garden flower for centuries, bleeding heart or old-fashioned sult in new floral racemes developing bleeding heart (Lamprocapnos spectabilis) is also used as a potted plant and in floral from axillary buds on the stem or from arrangements. Most general gardening guides include information on growing crown buds. Once growing, long pho- conditions but provide few specifics regarding plant growth and development that toperiods of 18 h promote growth and are important to those interested in commercial cultivation and use in the floral industry.
    [Show full text]
  • Family: Papaveraceae) Leaf * Sarma Dhrubajyoti, Thakuria Arup Jyoti, Sarma Hiranmoy, Sarma Nipan, Kakoty Dipu, Mazumder Zahid, Dr
    International Journal of Research in Pharmacy and Pharmaceutical Sciences ISSN: 2455-698X; Impact Factor: RJIF 5.22 www.pharmacysciencejournal.com Volume 1; Issue 3; July 2016; Page No. 21-23 Pharmacognostic studies and physicochemical properties of the Dactylicapnos scandens (Family: Papaveraceae) leaf * Sarma Dhrubajyoti, Thakuria Arup Jyoti, Sarma Hiranmoy, Sarma Nipan, Kakoty Dipu, Mazumder Zahid, Dr. Kamal Barman State Drug Testing Laboratory (AYUSH), Jalukbari, Guwahati, Assam, India Abstract A large number of the plants are claimed to possess the medicinal properties in the traditional system and are also used extensively by the tribal people worldwide. The present study deals with the macroscopic and microscopic studies of the leaf of Dactylicapnos scandens (Family: Papaveraceae). The anatomy of the leaf was studied by taking transverse section which showed anomocytic stomata, unicellular trichomes, etc. Physicochemical parameters of the powder were also carried out. The present investigation on Dactylicapnos scandens leaf might be useful to supplement information in regard to its identification parameters. Such studies are important in the way of acceptability of herbal drugs in present scenario of lacking regulatory laws to control quality of herbal drugs. Keywords: Dactylicapnos scandens, Family: Papaveraceae, anomocytic stomata, unicellular trichomes 1. Introduction 2. Materials and methods Plants have formed the sophisticated traditional medicine 2.1 Plant collection and extraction systems since thousands of years [1]. Pharmacognosy basically Fresh leaves of D actylicapnos scandens (Family: deals with the standardization, authentication and study of Papaveraceae) were collected from Nagaland, North Eastern natural drugs. In villages ethnic people depend on this region, India. The plant was identified by Botanical Survey of medicinal plant for their primary healthcare.
    [Show full text]
  • Threatened Ecosystems of Myanmar
    Threatened ecosystems of Myanmar An IUCN Red List of Ecosystems Assessment Nicholas J. Murray, David A. Keith, Robert Tizard, Adam Duncan, Win Thuya Htut, Nyan Hlaing, Aung Htat Oo, Kyaw Zay Ya and Hedley Grantham 2020 | Version 1.0 Threatened Ecosystems of Myanmar. An IUCN Red List of Ecosystems Assessment. Version 1.0. Murray, N.J., Keith, D.A., Tizard, R., Duncan, A., Htut, W.T., Hlaing, N., Oo, A.H., Ya, K.Z., Grantham, H. License This document is an open access publication licensed under a Creative Commons Attribution-Non- commercial-No Derivatives 4.0 International (CC BY-NC-ND 4.0). Authors: Nicholas J. Murray University of New South Wales and James Cook University, Australia David A. Keith University of New South Wales, Australia Robert Tizard Wildlife Conservation Society, Myanmar Adam Duncan Wildlife Conservation Society, Canada Nyan Hlaing Wildlife Conservation Society, Myanmar Win Thuya Htut Wildlife Conservation Society, Myanmar Aung Htat Oo Wildlife Conservation Society, Myanmar Kyaw Zay Ya Wildlife Conservation Society, Myanmar Hedley Grantham Wildlife Conservation Society, Australia Citation: Murray, N.J., Keith, D.A., Tizard, R., Duncan, A., Htut, W.T., Hlaing, N., Oo, A.H., Ya, K.Z., Grantham, H. (2020) Threatened Ecosystems of Myanmar. An IUCN Red List of Ecosystems Assessment. Version 1.0. Wildlife Conservation Society. ISBN: 978-0-9903852-5-7 DOI 10.19121/2019.Report.37457 ISBN 978-0-9903852-5-7 Cover photos: © Nicholas J. Murray, Hedley Grantham, Robert Tizard Numerous experts from around the world participated in the development of the IUCN Red List of Ecosystems of Myanmar. The complete list of contributors is located in Appendix 1.
    [Show full text]
  • Habitat Management Guide for Pollinators
    PENNSYLVANIA NATURAL HERITAGE PROGRAM HABITAT MANAGEMENT FOR POLLINATORS Atlantis Fritillary (Speyeria atlantis) on forget-me-nots (Myosotis sp.) Tri-colored bumble bee (Bombus ternarius) on common milkweed (Asclepias syriaca) 1 Contents Introduction ............................................................................................................................................ 3 Management Priorities ........................................................................................................................... 4 Inventory Habitats and Plants................................................................................................................. 4 Promote Habitat Variety to Support All Life Stages of Pollinators ......................................................... 5 Maintain Open Habitats .......................................................................................................................... 7 Control Invasive Plants ............................................................................................................................ 8 Protect Pollinator Diversity and Rare Species ........................................................................................ 9 Find Native and Local Plants ................................................................................................................. 14 Books and Other Publications ............................................................................................................... 20 Websites ..............................................................................................................................................
    [Show full text]