Amiiforms from the Iberian Peninsula: Historic Review and Research Prospects

Total Page:16

File Type:pdf, Size:1020Kb

Amiiforms from the Iberian Peninsula: Historic Review and Research Prospects Mesozoic Fishes 5 – Global Diversity and Evolution, G. Arratia, H.-P. Schultze & M. V. H. Wilson (eds.): pp. 73-86, 6 figs., 1 tab. © 2013 by Verlag Dr. Friedrich Pfeil, München, Germany – ISBN 978-3-89937-159-8 Amiiforms from the Iberian Peninsula: historic review and research prospects Hugo MARTÍN-ABAD and Francisco José POYATO-ARIZA Abstract The fossil record of amiiform fishes from the Iberian Peninsula is known from 12 different sites. It extends along the three periods of the Mesozoic era, being much more abundant during the Early Cretaceous. The majority of these sites has yielded isolated remains, mostly teeth; complete and articulated specimens are known from three Konservat-Lagerstätten only: Montral-Alcover, El Montsec, and Las Hoyas. Generic and specific level assessments are possible in these three localities only. The historical taxonomical problems of the cited amiiform taxa are commented. Special mention deserves the case of the genus Urocles (= Megalurus), traditionally cited from El Montsec and Las Hoyas. This genus is invalid since 1998, but its taxonomic history goes back to the 1830’s. The amiiform fishes from Las Hoyas have traditionally been assigned to the same taxa as those coming from El Montsec, but they have not been studied in detail yet, so their taxonomical assessment is in need of confirma- tion. These specimens, characterized by a great quality of preservation and an abundant record, may provide significant information concerning the ontogenetic development of the fishes of this order. Introduction The amiiform fishes: Amiiformes HAY (1929) is an order of neopterygian fishes with an only extant species of the type genus, Amia calva; their peak of diversity occurred during the Mesozoic. Since there are very few living non-teleosts neotperygians, this order is considered to be as a very relevant group from a systematic and paleobiogeographic point of view, especially when dealing with its phylogenetic relationships with the Teleostei. In the last thirty years diverse hypotheses about the relationship be- tween amiiforms and teleosts have been offered, in an attempt to solve the debate on whether they are sister-groups or not. GRANDE & BEMIS (1998) provided a cladistic analysis resolving a clade containing both teleosts and amiiforms as sister-groups; this clade has traditionally been called Halecostomi. They redescribed the order Amiiformes, which they diagnosed by the following three evolutionary novelties: phylogenetic reduction in the number of ossified ural neural arches to two or less (character secondarily lost within Caturidae); phylogenetic loss of the opisthotic bone (character apparently convergent with lepisosteids and more derived teleosts); and phylogenetic loss of the pterotic bone. GRANDE & BEMIS (1998) also divided the amiiforms into two superfamilies, Amioidea and Caturoidea, each of which is in turn divided into two families, respectively: Amiidae and Sinamiidae, and Caturidae and Liodesmidae. Although it is considered as a principally Mesozoic order, Amiiformes present a relatively long chrono- stratigraphical record, which extends at least from the Jurassic, or even the Triassic, if the identifications made by BELTAN (1972, 1984) and LIU et al. (2002) are correct, to the Recent Amia calva, that inhabits freshwaters systems of the eastern North America. The amiiform fossil record also presents a wide-ranged geographical distribution that could be considered almost cosmopolite; however, the vast majority of the known fossil material has been discovered in sites from the North Hemisphere, probably as a result of bias in paleontological field work. This is partly evidenced by their distribution reaching very high latitudes (ESTES & HUTCHINSON 1980, LEHMAN 1951). 73 MESOZOIC The Spanish record: A number of nominal PICKS genera have been described within the Amii- PERIOD EPOCH AGE (Ma) 12. Cacém-Alcântara 65.5 formes, mainly in the family Amiidae. The MAASTRICHTIAN 70.6 present paper is included in a revision project CAMPANIAN 11. Las Hoyas on the amiiforms from Las Hoyas fossil site LATE SANTONIAN 83.5 85.8 CONIACIAN (Cuenca, Spain), currently in progress. In this 89.3 TURONIAN 93.5 12. Buenache de la Sierra sense, the main objective of the present paper CENOMANIAN 99.6 is to offer the most relevant references on ALBIAN 9. Alcaine amiiform fishes from the Iberian Peninsula, in 112 order to provide a biogeographical and chro- APTIAN EARLY 125 noestratigraphical framework for the amiiforms BARREMIAN 8. Galve CRETACEOUS 130 HAUTERIVIAN of Las Hoyas. Some of these references appear 136 VALANGINIAN 140 in the few compilations of the Spanish material BERRIASIAN 145.5 7. Vega de Pas I TITHONIAN published in the second half of the past century, 151 LATE KIMMERIDGIAN which are faunal lists mainly focused on the 156 OXFORDIAN 161 CALLOVIAN 6. El Montsec Cretaceous period (BATALLER 1960, POYATO- 165 BATHONIAN MIDDLE 168 BAJOCIAN ARIZA & WENZ 1990, POYATO-ARIZA et al. 172 AALENIAN 176 5. Cuenca de Cameros 1999). Those concerning only amiids that were TOARCIAN 183 published before 1998 were already included in EARLY PLIENSBACHIAN JURASSIC 190 SINEMURIAN GRANDE & BEMIS (1998); the posterior refer- 197 4. Tereñes & Lastres HETTANGIAN 201.6 ences to this family in the Iberian Peninsula RHAETIAN 204 are compiled in the present paper for the first 3. Guimarota time, together with all available references to NORIAN LATE the other families of this order. 228 2. Sot de Chera & Loriguilla Chronostratigraphicly, the Iberian record of CARNIAN 235 LADINIAN amiiform fishes extends from the Oxfordian- MIDDLE 241 TRIASSIC ANISIAN 245 1. Montral-Alcover Kimmeridgian (Late Jurassic) to the middle- OLENEKIAN EARLY 250 INDUAN 251.0 to-late Cenomanian (Upper Cretaceous), being Fig. 1. thus exclusively Mesozoic (Fig. 1). The first Chronostratigraphic distribution of the Iberian sites that appearance of the order could be traced back have yielded remains of amiiform fishes. The assessment of to the late Ladinian (Middle Triassic) accord- the material from Montral-Alcover to the order Amiiformes ing to BELTAN (1972, 1984) and VÍA-BOADA is doubtful. et al. (1977), but other authors (CARTANYÀ Table 1. Distribution of the taxonomic assessments of the amiiform fishes cited from Mesozoic localities of the Iberian Peninsula. Montral-Alcover Sot de Chera & Loriguilla Guimarota Tereñes & Lastres Cuenca de Cameros El Montsec Vega de Pas I Galve Alcaine Buenache de la Sierra Las Hoyas Cacém-Alcântara Amiiformes indet. %% %% Caturidae indet. %% Caturus sp. ? %%% Caturus tarraconensis % Amiidae indet. %% %% % Amiopsinae indet. % Amiopsis woodwardi %% Vidalamia catalunica %% 74 4 7 5 6 1 8 10 9 11 3 2 12 Cretaceous Jurassic Triassic Fig. 2. Mesozoic Iberian sites that have yielded remains of amiiform fishes. Numeration of localities corresponds to that from Figure 1. 1995, 1999) considered their assessment doubtful and in need of a revision. Geographically, the sites that have yielded remains of amiiform fishes are mainly concentrated in the northern and oriental regions of Spain and in the occidental coast of Portugal. To date, no insular outcrops have been reported from Spain or Portugal. Figure 2 shows the Iberian localities from Triassic, Jurassic, and Cretaceous beds. In general terms, the Iberian record could be divided into two sets. On one hand there are several sites whose remains appear to be more or less isolated and disarticulated, thereby creating problems with taxonomy. This set includes remains that have been reported as Actinopterygii indet., Neopterygii indet., Halecostomi indet., Halecomorphi indet., or Holostei indet., but they will not be considered in the present paper, since their assessment to the order Amiiformes would require a detailed revision that lies outside of the scope of the present work. On the other hand, there are a few Konservat-Lagerstätten which present an extensive and excellently-preserved fish record that usually allows a much more precise taxonomic assignment. Among these last ones, there are two localities that are especially relevant: El Montsec and Las Hoyas. In some cases, the material was originally described under names that have been changed in the course of history. In other cases, the remains do not really belong to the amiiform taxa they were initially assigned to or the corresponding taxa are not included in the Amiiformes anymore. Each particular case will be presented and discussed in detail below, following a systematic order (Table 1), with special reference to the genus Urocles (= Megalurus), of which the taxonomic history is quite complex. Vernacular suffixes and institutional abbreviations: The vernacular suffixes are used in this paper following GRANDE & BEMIS (1999). Institutional abbreviations used here are as follow: MCCM, Museo de las Ciencias de Castilla-La Mancha, Cuenca, Spain (LH, Las Hoyas Collection; MSE, El Montsec Collection); MMGB, Museo Municipal de Geología de Barcelona, Barcelona, Spain; MNHN, Muséum National d’Histoire Naturelle (Institut de Paléontologie), Paris, France. 75 The amiiform record from the Iberian Peninsula Taxa no longer assigned to the Amiiformes: Two different genera have been traditionally included within the Amiiformes, but subsequently were transferred to other orders. These are Eoeugnathus (i. e. BELTAN 1984, CARTANYÀ 1995) and Ophiopsis (i. e. CARTANYÀ 1999). Amiiformes indet.: In many localities the remains of amiiform fishes have been cited, of which the quality and/or integrity of preservation only allows an identification
Recommended publications
  • A New Microvertebrate Assemblage from the Mussentuchit
    A new microvertebrate assemblage from the Mussentuchit Member, Cedar Mountain Formation: insights into the paleobiodiversity and paleobiogeography of early Late Cretaceous ecosystems in western North America Haviv M. Avrahami1,2,3, Terry A. Gates1, Andrew B. Heckert3, Peter J. Makovicky4 and Lindsay E. Zanno1,2 1 Department of Biological Sciences, North Carolina State University, Raleigh, NC, USA 2 North Carolina Museum of Natural Sciences, Raleigh, NC, USA 3 Department of Geological and Environmental Sciences, Appalachian State University, Boone, NC, USA 4 Field Museum of Natural History, Chicago, IL, USA ABSTRACT The vertebrate fauna of the Late Cretaceous Mussentuchit Member of the Cedar Mountain Formation has been studied for nearly three decades, yet the fossil-rich unit continues to produce new information about life in western North America approximately 97 million years ago. Here we report on the composition of the Cliffs of Insanity (COI) microvertebrate locality, a newly sampled site containing perhaps one of the densest concentrations of microvertebrate fossils yet discovered in the Mussentuchit Member. The COI locality preserves osteichthyan, lissamphibian, testudinatan, mesoeucrocodylian, dinosaurian, metatherian, and trace fossil remains and is among the most taxonomically rich microvertebrate localities in the Mussentuchit Submitted 30 May 2018 fi fi Accepted 8 October 2018 Member. To better re ne taxonomic identi cations of isolated theropod dinosaur Published 16 November 2018 teeth, we used quantitative analyses of taxonomically comprehensive databases of Corresponding authors theropod tooth measurements, adding new data on theropod tooth morphodiversity in Haviv M. Avrahami, this poorly understood interval. We further provide the first descriptions of [email protected] tyrannosauroid premaxillary teeth and document the earliest North American record of Lindsay E.
    [Show full text]
  • From the Crato Formation (Lower Cretaceous)
    ORYCTOS.Vol. 3 : 3 - 8. Décembre2000 FIRSTRECORD OT CALAMOPLEU RUS (ACTINOPTERYGII:HALECOMORPHI: AMIIDAE) FROMTHE CRATO FORMATION (LOWER CRETACEOUS) OF NORTH-EAST BRAZTL David M. MARTILL' and Paulo M. BRITO'z 'School of Earth, Environmentaland PhysicalSciences, University of Portsmouth,Portsmouth, POl 3QL UK. 2Departmentode Biologia Animal e Vegetal,Universidade do Estadode Rio de Janeiro, rua SâoFrancisco Xavier 524. Rio de Janeiro.Brazll. Abstract : A partial skeleton representsthe first occurrenceof the amiid (Actinopterygii: Halecomorphi: Amiidae) Calamopleurus from the Nova Olinda Member of the Crato Formation (Aptian) of north east Brazil. The new spe- cimen is further evidencethat the Crato Formation ichthyofauna is similar to that of the slightly younger Romualdo Member of the Santana Formation of the same sedimentary basin. The extended temporal range, ?Aptian to ?Cenomanian,for this genus rules out its usefulnessas a biostratigraphic indicator for the Araripe Basin. Key words: Amiidae, Calamopleurus,Early Cretaceous,Brazil Première mention de Calamopleurus (Actinopterygii: Halecomorphi: Amiidae) dans la Formation Crato (Crétacé inférieur), nord est du Brésil Résumé : la première mention dans le Membre Nova Olinda de la Formation Crato (Aptien ; nord-est du Brésil) de I'amiidé (Actinopterygii: Halecomorphi: Amiidae) Calamopleurus est basée sur la découverted'un squelettepar- tiel. Le nouveau spécimen est un élément supplémentaireindiquant que I'ichtyofaune de la Formation Crato est similaire à celle du Membre Romualdo de la Formation Santana, située dans le même bassin sédimentaire. L'extension temporelle de ce genre (?Aptien à ?Cénomanien)ne permet pas de le considérer comme un indicateur biostratigraphiquepour le bassin de l'Araripe. Mots clés : Amiidae, Calamopleurus, Crétacé inférieu4 Brésil INTRODUCTION Araripina and at Mina Pedra Branca, near Nova Olinda where cf.
    [Show full text]
  • Clay Diagenesis and Low-Grade Metamorphism of Tithonian and Berriasian Sediments in the Cameros Basin (Spain)
    View metadata, citation and similar papers at core.ac.uk brought to you by CORE provided by EPrints Complutense Clay Minerals (2001) 36, 325–333 Clay diagenesis and low-grade metamorphism of Tithonian and Berriasian sediments in the Cameros Basin (Spain) J. F. BARRENECHEA1,*, M. RODAS1 ,M.FREY2 , J. ALONSO-AZCA´ RATE3 4 AND J. R. MAS 1 Departamento de Cristalografı´a y Mineralogı´a, Universidad Complutense de Madrid, 28040 Madrid, Spain, 2 Mineralogisch-Petrographisches Institut, Basel University, CH 4056 Basel, Switzerland (deceased), 3 Facultad de Ciencias del Medio Ambiente, Universidad de Castilla-La Mancha, Fa´brica de Armas, 45071 Toledo, Spain, and 4 Departamento de Estratigrafı´a, Universidad Complutense de Madrid, 28040 Madrid, Spain (Received 29 February 2000; revised 13 October 2000) ABSTRACT: The clay mineral assemblages of the Tithonian and Berriasian sediments (Tera and Oncala Groups) in the eastern part of the Cameros basin are investigated at seven localities. The lowest-grade assemblage, located on the southern border of the basin, contains calcite + quartz + hematite + kaolinite + mixed-layer illite-smectite (R = 1, 65À85% illite layers) + discrete illite (IC = 0.5À0.65D82y). Systematic increases in the illite and chlorite crystallinities suggest increasing metamorphic grade from the northwest part of the basin to the southeast. This trend does not follow the pattern previously described for the overlying late Berriasian–early Aptian sediments (Urbio´n and Enciso Groups), which exhibit a higher metamorphic grade. This may result from local variations in sedimentary facies, as well as the circulation of hot migratory fluids. Tertiary compression occurring long after the main metamorphic event is considered to be responsible for the enhanced illite and chlorite crystallinities measured in the SE extreme of the basin.
    [Show full text]
  • An Overview of the Lower Cretaceous Dinosaur Tracksites from the Mirambel Formation in the Iberian Range (Ne Spain)
    Khosla, A. and Lucas, S.G., eds., 2016, Cretaceous Period: Biotic Diversity and Biogeography. New Mexico Museum of Natural History and Science Bulletin 71. 65 AN OVERVIEW OF THE LOWER CRETACEOUS DINOSAUR TRACKSITES FROM THE MIRAMBEL FORMATION IN THE IBERIAN RANGE (NE SPAIN) D. CASTANERA1, I. DÍAZ-MARTÍNEZ2, M. MORENO-AZANZA3, J.I. CANUDO4, AND J.M. GASCA4 1 Bayerische Staatssammlung für Paläontologie und Geologie and GeoBioCenter, Ludwig-Maximilians-Universität, Richard-Wagner-Str. 10, 80333 Munich, Germany. [email protected]; 2 CONICET - Instituto de Investigación en Paleobiología y Geología, Universidad Nacional de Río Negro, General Roca 1242, 8332 General Roca, Río Negro, [email protected]; 3 Departamento de Ciências da Terra, Geobiotec. Departamento de Ciências da Terra. Faculdade de Ciências e Tecnologia, FCT, Universidade Nova de Lisboa, 2829-526. Caparica, Portugal. Museu da Lourinhã. [email protected]; 4 Grupo Aragosaurus-IUCA, Paleontología, Departamento de Ciencias de la Tierra, Facultad de Ciencias, Universidad de Zaragoza, Calle Pedro Cerbuna, 12, 50009, Zaragoza, Spain. [email protected]; [email protected] Abstract—Up to now, the ichnological vertebrate record from the Barremian Mirambel Formation (NE Spain) has remained completely unknown despite the fact that osteological findings have been reported in recent years. Here we provide an overview of 11 new dinosaur tracksites found during a fieldwork campaign in the year 2011. The majority of these tracksites (seven) preserve small- to medium-sized tridactyl tracks here assigned to indeterminate theropods. Only one footprint presents enough characters to classify it as Megalosauripus isp. Ornithopod tracks identified asCaririchnium isp. and Iguanodontipodidae indet.
    [Show full text]
  • The Strawberry Bank Lagerstätte Reveals Insights Into Early Jurassic Lifematt Williams, Michael J
    XXX10.1144/jgs2014-144M. Williams et al.Early Jurassic Strawberry Bank Lagerstätte 2015 Downloaded from http://jgs.lyellcollection.org/ by guest on September 27, 2021 2014-144review-articleReview focus10.1144/jgs2014-144The Strawberry Bank Lagerstätte reveals insights into Early Jurassic lifeMatt Williams, Michael J. Benton &, Andrew Ross Review focus Journal of the Geological Society Published Online First doi:10.1144/jgs2014-144 The Strawberry Bank Lagerstätte reveals insights into Early Jurassic life Matt Williams1, Michael J. Benton2* & Andrew Ross3 1 Bath Royal Literary and Scientific Institution, 16–18 Queen Square, Bath BA1 2HN, UK 2 School of Earth Sciences, University of Bristol, Bristol BS8 2BU, UK 3 National Museum of Scotland, Chambers Street, Edinburgh EH1 1JF, UK * Correspondence: [email protected] Abstract: The Strawberry Bank Lagerstätte provides a rich insight into Early Jurassic marine vertebrate life, revealing exquisite anatomical detail of marine reptiles and large pachycormid fishes thanks to exceptional preservation, and especially the uncrushed, 3D nature of the fossils. The site documents a fauna of Early Jurassic nektonic marine animals (five species of fishes, one species of marine crocodilian, two species of ichthyosaurs, cephalopods and crustaceans), but also over 20 spe- cies of insects. Unlike other fossil sites of similar age, the 3D preservation at Strawberry Bank provides unique evidence on palatal and braincase structures in the fishes and reptiles. The age of the site is important, documenting a marine ecosystem during recovery from the end-Triassic mass extinction, but also exactly coincident with the height of the Toarcian Oceanic Anoxic Event, a further time of turmoil in evolution.
    [Show full text]
  • Abstract Book.Pdf
    Welcome! Welcome to the VI Symposium on Dinosaur Eggs and Babies, the return of this periodic gathering to the Iberian Peninsula, when it hatched eighteen years ago. From the slopes of the Pyrenees, we have followed the first steps of dinosaurs through France, Argentina, the United States and China. Today, we come back and see the coast where the first theropod embryos were discovered twenty years ago. Since the end of the last century, Paleoology, much like other branches of palaeontology, has evolved thanks to the advance of new methodologies and analytical tools, becoming a progressively more interdisciplinary area of knowledge. Dinosaur babies and embryos, rare findings back when these meetings started, seem to be everywhere now that we learn to look for them under the light of the microscope. New astonishing specimens allow us to understand how Mesozoic dinosaurs mate and reproduce. Oology, our parent discipline in the modern world, has made great advances in understanding the form and function of the egg, and its applications on poultry industry are countless. More than thirty contributions evidence that our field remains small but alive and healthy. We hope that you find in this Symposium an opportunity to share knowledge and open new lines of collaboration. And do not forget to enjoy your stay in Portugal. The host committee CONTENTS How to get to the FCT 6 Acknowledgements 10 PROGRAM 11 ABSTRACTS 14 THE FIRST ORNITHOMIMID EMBRYO IN A SHELL WITH A SINGLE STRUCTURAL LAYER: A CHALLENGE TO ORTHODOXY 15 Araújo R., Lamb J., Atkinson P., Martins R. M. S., Polcyn M.J., Fernandez V.
    [Show full text]
  • Pterosaur Distribution in Time and Space: an Atlas 61
    Zitteliana An International Journal of Palaeontology and Geobiology Series B/Reihe B Abhandlungen der Bayerischen Staatssammlung für Pa lä on to lo gie und Geologie B28 DAVID W. E. HONE & ERIC BUFFETAUT (Eds) Flugsaurier: pterosaur papers in honour of Peter Wellnhofer CONTENTS/INHALT Dedication 3 PETER WELLNHOFER A short history of pterosaur research 7 KEVIN PADIAN Were pterosaur ancestors bipedal or quadrupedal?: Morphometric, functional, and phylogenetic considerations 21 DAVID W. E. HONE & MICHAEL J. BENTON Contrasting supertree and total-evidence methods: the origin of the pterosaurs 35 PAUL M. BARRETT, RICHARD J. BUTLER, NICHOLAS P. EDWARDS & ANDREW R. MILNER Pterosaur distribution in time and space: an atlas 61 LORNA STEEL The palaeohistology of pterosaur bone: an overview 109 S. CHRISTOPHER BENNETT Morphological evolution of the wing of pterosaurs: myology and function 127 MARK P. WITTON A new approach to determining pterosaur body mass and its implications for pterosaur fl ight 143 MICHAEL B. HABIB Comparative evidence for quadrupedal launch in pterosaurs 159 ROSS A. ELGIN, CARLOS A. GRAU, COLIN PALMER, DAVID W. E. HONE, DOUGLAS GREENWELL & MICHAEL J. BENTON Aerodynamic characters of the cranial crest in Pteranodon 167 DAVID M. MARTILL & MARK P. WITTON Catastrophic failure in a pterosaur skull from the Cretaceous Santana Formation of Brazil 175 MARTIN LOCKLEY, JERALD D. HARRIS & LAURA MITCHELL A global overview of pterosaur ichnology: tracksite distribution in space and time 185 DAVID M. UNWIN & D. CHARLES DEEMING Pterosaur eggshell structure and its implications for pterosaur reproductive biology 199 DAVID M. MARTILL, MARK P. WITTON & ANDREW GALE Possible azhdarchoid pterosaur remains from the Coniacian (Late Cretaceous) of England 209 TAISSA RODRIGUES & ALEXANDER W.
    [Show full text]
  • Phylogeny and Biogeography of Iguanodontian Dinosaurs, with Implications from Ontogeny and an Examination of the Function of the Fused Carpal-Digit I Complex
    Phylogeny and Biogeography of Iguanodontian Dinosaurs, with Implications from Ontogeny and an Examination of the Function of the Fused Carpal-Digit I Complex By Karen E. Poole B.A. in Geology, May 2004, University of Pennsylvania M.A. in Earth and Planetary Sciences, August 2008, Washington University in St. Louis A Dissertation submitted to The Faculty of The Columbian College of Arts and Sciences of The George Washington University in partial fulfillment of the requirements for the degree of Doctor of Philosophy August 31, 2015 Dissertation Directed by Catherine Forster Professor of Biology The Columbian College of Arts and Sciences of The George Washington University certifies that Karen Poole has passed the Final Examination for the degree of Doctor of Philosophy as of August 10th, 2015. This is the final and approved form of the dissertation. Phylogeny and Biogeography of Iguanodontian Dinosaurs, with Implications from Ontogeny and an Examination of the Function of the Fused Carpal-Digit I Complex Karen E. Poole Dissertation Research Committee: Catherine A. Forster, Professor of Biology, Dissertation Director James M. Clark, Ronald Weintraub Professor of Biology, Committee Member R. Alexander Pyron, Robert F. Griggs Assistant Professor of Biology, Committee Member ii © Copyright 2015 by Karen Poole All rights reserved iii Dedication To Joseph Theis, for his unending support, and for always reminding me what matters most in life. To my parents, who have always encouraged me to pursue my dreams, even those they didn’t understand. iv Acknowledgements First, a heartfelt thank you is due to my advisor, Cathy Forster, for giving me free reign in this dissertation, but always providing valuable commentary on any piece of writing I sent her, no matter how messy.
    [Show full text]
  • Fast-Running Theropods Tracks from the Early Cretaceous of La Rioja, Spain
    Fast-Running Theropods Tracks From the Early Cretaceous of La Rioja, Spain Pablo Navarro-Lorbés ( [email protected] ) Universidad de La Rioja (UR), C/ Luis de Ulloa Javier Ruiz Universidad Complutense de Madrid Ignacio Díaz-Martínez Universidad Nacional de Río Negro-IIPG Erik Isasmendi Universidad del País Vasco/Euskal Herriko Unibertsitatea (UPV/EHU) Patxi Sáez-Benito Centro de Interpretación Paleontológica de La Rioja, C/ Mayor, 10, 26525, Igea, La Rioja Luis Viera Centro de Interpretación Paleontológica de La Rioja, C/ Mayor, 10, 26525, Igea, La Rioja Xabier Pereda-Suberbiola Universidad del País Vasco/Euskal Herriko Unibertsitatea (UPV/EHU) Angélica Torices Universidad de La Rioja (UR), C/ Luis de Ulloa Research Article Keywords: Fast-running theropods tracks, Early Cretaceous, La Rioja, Spain, Theropod behaviour, biodynamics, aleontology, biomechanical models Posted Date: August 5th, 2021 DOI: https://doi.org/10.21203/rs.3.rs-764084/v1 License: This work is licensed under a Creative Commons Attribution 4.0 International License. Read Full License Page 1/23 Abstract Theropod behaviour and biodynamics are intriguing questions that paleontology has been trying to resolve for a long time. The lack of extant groups with similar bipedalism has made it hard to answer some of the questions on the matter, yet theoretical biomechanical models have shed some light on the question of how fast theropods could run and what kind of movement they showed. The study of dinosaur tracks can help answer some of these questions due to the very nature of tracks as a product of the interaction of these animals with the environment. Two trackways belonging to fast-running theropods from the Lower Cretaceous Enciso Group of Igea (La Rioja) are presented here and compared with other fast-running theropod trackways published to date.
    [Show full text]
  • Mcabee Fossil Site Assessment
    1 McAbee Fossil Site Assessment Final Report July 30, 2007 Revised August 5, 2007 Further revised October 24, 2008 Contract CCLAL08009 by Mark V. H. Wilson, Ph.D. Edmonton, Alberta, Canada Phone 780 435 6501; email [email protected] 2 Table of Contents Executive Summary ..............................................................................................................................................................3 McAbee Fossil Site Assessment ..........................................................................................................................................4 Introduction .......................................................................................................................................................................4 Geological Context ...........................................................................................................................................................8 Claim Use and Impact ....................................................................................................................................................10 Quality, Abundance, and Importance of the Fossils from McAbee ............................................................................11 Sale and Private Use of Fossils from McAbee..............................................................................................................12 Educational Use of Fossils from McAbee.....................................................................................................................13
    [Show full text]
  • Curriculum Vitae
    CURRICULUM VITAE JOSÉ M. GASCA, PERSONAL SUMMARY Surname Gasca Name José Manuel Date of birth April 8, 1981 Place of birth Zaragoza, Spain CONICET‐ Museo Olsacher. Affiliation Zapala (8340 Neuquén, Argentina) Mobile number +54 2942 660 880 (Argentina) Mobile number (Spain) +34 699 697707 and WhatsApp E‐mail [email protected] Personal blog cretaceousfossils.blogspot.com.es RG profile https://www.researchgate.net/profile/Jose_Gasca ACADEMIC Ph.D. in Geology. University of Zaragoza, Spain. March, 2015. PhD Title: Contributions to the knowledge of the lower Barremian (Lower Cretaceous) dinosaurs from Teruel, Spain: fossil associations, systematics, palaeobiodiversity and palaeobiogeographical affinities. Director: José Ignacio Canudo (University of Zaragoza). Score: Sobresaliente Cum Laude M.Sc. in Geology. University of Zaragoza, Spain, September, 2008. B.A. in Geology. University of Zaragoza, Spain, February, 2007. RESEARCH FELLOWSHIPS PostDoctoral Fellowship of CONICET , Argentina. Since April 2016 until now. Director: Dr. Leonardo Salgado. Workplace: Natural Science Museum of Zapala (Neuquén province, Argentina). Doctoral Fellowship of the regional Government of Aragón, Spain. September 2008‐ August 2012. Director: José I. Canudo. Workplace: University of Zaragoza (Spain). OTHER FELLOWSHIPS, CURATION AND PROFESSIONAL EXPERIENCE Technical assistance and coordination in the installation of the new exhibition of the Natural Science Museum of the University of Zaragoza, Spain. Position: freelancer. Client: University of Zaragoza. September, 2014 – December, 2014. Fossil preparation (dinosaur bones). Position: freelancer. Client: Deparment of Cultural Heritage, Government of Aragón, Spain. September, 2014 – December, 2014. Management tasks, social networks, guided tours and curation in the Natural Science Museum of the University of Zaragoza, Spain. Position: fellowship. October, 2012 – September, 2013 (1 year, 30 hours a week).
    [Show full text]
  • High European Sauropod Dinosaur Diversity During Jurassic–Cretaceous Transition in Riodeva (Teruel, Spain)
    CORE Metadata, citation and similar papers at core.ac.uk Provided by RERO DOC Digital Library [Palaeontology, Vol. 52, Part 5, 2009, pp. 1009–1027] HIGH EUROPEAN SAUROPOD DINOSAUR DIVERSITY DURING JURASSIC–CRETACEOUS TRANSITION IN RIODEVA (TERUEL, SPAIN) by RAFAEL ROYO-TORRES*, ALBERTO COBOS*, LUIS LUQUE*, AINARA ABERASTURI*, , EDUARDO ESPI´LEZ*, IGNACIO FIERRO*, ANA GONZA´ LEZ*, LUIS MAMPEL* and LUIS ALCALA´ * *Fundacio´n Conjunto Paleontolo´gico de Teruel-Dino´polis. Avda. Sagunto s ⁄ n. E-44002 Teruel, Spain; e-mail: [email protected] Escuela Taller de Restauracio´n Paleontolo´gica II del Gobierno de Arago´n. Avda. Sagunto s ⁄ n. E-44002 Teruel, Spain Typescript received 13 December 2007; accepted in revised form 3 November 2008 Abstract: Up to now, more than 40 dinosaur sites have (CPT-1074) referring to the Diplodocidae clade. New been found in the latest Jurassic – earliest Cretaceous remains from the RD-28, RD-41 and RD-43 sites, of the sedimentary outcrops (Villar del Arzobispo Formation) of same age, among which there are caudal vertebrae, are Riodeva (Iberian Range, Spain). Those already excavated, assigned to Macronaria. New sauropod footprints from the as well as other findings, provide a large and diverse Villar del Arzobispo Formation complete the extraordinary number of sauropod remains, suggesting a great diversity sauropod record coming to light in the area. The inclusion for this group in the Iberian Peninsula during this time. of other sauropods from different contemporaneous expo- Vertebrae and ischial remains from Riodevan site RD-13 sures in Teruel within the Turiasauria clade adds new evi- are assigned to Turiasaurus riodevensis (a species described dence of a great diversity of sauropods in Iberia during in RD-10, Barrihonda site), which is part of the the Jurassic–Cretaceous transition.
    [Show full text]