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THE LEAF FAUNA (COLEOPTERA, CHRYSOMELIDAE) OF THE RODNA MOUNTAINS (TRANSYLVANIA-MARAMURE Ş, )

Sanda MAICAN *

* Romanian Academy Institute of Biology of Bucharest, Department of Ecology, Taxonomy and Nature Conservation, Independen ţei Street 296, Bucharest, Romania, RO-060031, [email protected]

KEYWORDS : Romanian Carpathians, Rodna Mountains, Coleoptera, Chrysomelidae, diversity.

ABSTRACT The paper presents data on the species diversity of the leaf beetles fauna from the Rodna Mountains on the basis of the previously published information and on the material collected from this area between 1995 and 2004. Taxonomical structure of the Chrysomelidae family consists of 43 genera and 138 species, representing about 64% from the total number of leaf beetles species known in the Maramure ş region and 24% from the Romanian chrysomelid fauna. Sclerophaedon carpathicus Weise, Chrysolina weisei (Frivaldszky), Psylliodes frivaldszkyi Weise and Neocrepidodera transsilvanica (Fuss) are Carpathian endemic species.

RÉSUMÉ : La faune des chrysomelides (Coleoptera: Chrysomelidae) dans les Montagnes de Rodna (Transylvanie-Maramure ş, Roumanie). Ce travail présente des données sur la diversité de la faune de coléoptères chrysomelides dans le Montagnes de Rodna, sur la base de sources bibliographiques et de l’étude du matériel collecté au cours de la période 1995 et 2004. La structure taxonomique de la famille Chrysomelidae comprend 43 genres et 138 espèces, représentant presque 64% du total des espèces de crisomelides connues pour la région de Maramure ş et 24% du total des espèces de crisomelides citées dans la faune de Roumanie. Les espèces Sclerophaedon carpathicus Weise, Chrysolina weisei (Frivaldszky), Psylliodes frivaldszkyi Weise and Neocrepidodera transsilvanica (Fuss) sont endémiques dans les Carpathes.

REZUMAT : Fauna de crisomelide (Coleoptera: Chrysomelidae) a Mun ţilor Rodna (Transilvania-Maramure ş, România). Lucrarea prezint ă date, referitoare la diversitatea specific ă a faunei de crisomelide, din Mun ţii Rodna, pe baza informa ţiilor bibliografice şi a materialului colectat, în perioada 1995- 2004. Structura taxonomic ă a familiei Chrysomelidae, include 43 genuri şi 138 specii, ceea ce reprezint ă circa 64% din num ărul speciilor semnalate în regiunea Maramure ş şi 24% din totalul speciilor, citate în fauna României. Speciile Sclerophaedon carpathicus Weise, Chrysolina weisei (Frivaldszky), Psylliodes frivaldszkyi Weise şi Neocrepidodera transsilvanica (Fuss) sunt endemice în Mun ţii Carpa ţi.

124 S. Maican – The Leaf fauna of the Rodna Mountains (123 ~ 140)

INTRODUCTION The first information concerning the coleopterans from the Rodna Mountains, situated in the northern part of the Romanian Eastern Carpathians, were published in the second half of the XIX th century by Frivaldszky (1871) and Bielz (1887). A number of beetle species recorded from this area are found in the catalogues of Petri (1912, 1925). The Csiki’s paper (1951) presents a list with 1343 taxa (collected by the author, together with previous records for this area) from 56 families, including the Chrysomelidae family. Most of the chrysomelid species mentioned here were found on the southern mountainous slopes, on the tributaries of the Some şul Mare River: B ăilor Valley, M ăriilor Valley, valley of Ro şu Rivulet and Saca Valley. Szél et al. (1995) presents few leaf beetles species collected from Bor şa. Latest, some papers concerning the coleopterans from Maramure ş, also including the Rodna Mountains, were published by: Serafim (1997) - for Cerambycidae and Coccinellidae families; Proche ş (1998) - for Curculionidea; Stan (2002) - for Staphylinidae; Maican and Serafim (2001, 2004) and Maican (2007) - for Chrysomelidae; Nitzu et al. (2008), Merkl (2008). Investigations on the biodiversity of Maramure ş region and the National Park “Pietrosul Rodnei” were performed since 1995, within the “Knowledge of the invertebrates of Maramure ş” project, developed by the “Grigore Antipa” National Museum of Nature History of Bucharest. The objectives of this project were published by G ăldean and Pârvu (1997). Beetwen 1995 and 2004 several collecting trips were performed in Maramure ş area: the Maramure ş Depression, the catchment areas of the Iza, Mara and S ăpân ţa rivers, the Igni ş Plateau, the Ţible ş Mountains, the Maramure ş Mountains. Two expeditions were carried out in 1995 in the Pietrosul Rodnei Nature Park and on the valley of Iza River.

MATERIALS AND METHODS The paper presents data about the distribution of the chrysomelid species in the Rodna Mountains, based on the study of the material collected during the expeditions made by “Grigore Antipa” National Museum’s team, in the period 1995-2004, and also on records from faunal papers published since 1887 up to 2008. The subfamilies within Chrysomelidae family are listed in the phylogenetic order, according to Seeno and Wilcox (1982). For each recorded species, the collecting sites and localities, the general distribution and the literature cited are given. The geographic distribution, the nomenclature and systematics are presented after Warchałowski (2003). The zoogeographical classification of the taxa is presented in accordance with information from the Gruev’s papers (2006 a, b). Abbreviations: S - Siberian complex (EAP - Euro Asiatic Palaearctic: SibE - Sibero-European; SsibE - Southsibero-European; TrPal - Transpalaearctic; Hpal - Holopalaearctic; H - Holarctic; EAAP - Euro-Asiatic-African Palaearctic). E - European complex (CE - Central European; CEMount - Central European Mountain; Emount - European Mountain; SbM - Submediterranean; EsbM - Eastsubmediterranean); M - Mediterranean complex (HM - Holomediterranean; EM - Eastmediterranean). End - Endemic (CarpEnd - Endemic species to the Carpathians; AlpCarpEnd - Endemic species to the Alps and Carpathians).

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RESULTS AND DISCUSSION On the basis of the information obtained until now, 138 chrysomelid species belonging to 11 subfamilies and 43 genera are reported from the Rodna Mountains, representing about 24.2% from the number of leaf beetles species mentioned in the Romanian fauna and 64.4% from the total species recorded in Maramure ş region (Tab. 1). Among them, aproximatively 26% are mountainous species (e.g. Chrysolina globipennis , C. hemisphaerica , C. carpathica , C. marcasitica , C. umbratilis , Oreina coerulea , O. intricata , O. Virgulata , O. viridis , Sclerophaedon carniolicus , S. carpathicus , Psylliodes glaber, Orestia aubei ). According to the information from the coleopterological literature, the following European endemics of the Alticinae have been recorded in the fauna of the Rodna Mountains: Orestia aubei , Minota carpathica , Longitarsus rubellus , Longitarsus pallidicornis , Neocrepidodera melanostoma and Phyllotreta christinae - all of them being distributed in most part of the European mountains. Neocrepidodera transsilvanica , Psylliodes frivaldszkyi , Chrysolina weisei and Sclerophaedon carpathicus are endemic species in the Carpathians. Also, Neocrepidodera cyanescens is endemic in the Alpine-Carpathian chain. Among the species mentioned in the older literature, Chrysolina weisei is a rare mountainous species, possible endemic to the Romanian Carpathians. The Minota obesa presence (Petri, 1912; from the Rodna Mountains) in the Romanian fauna is questionable. According to Gruev et al. (1993), the records of this species in Romania were possible based on the misidentified specimens of Minota halmae or M. carpathica .

Table 1: The leaf beetles species recorded from the Rodna Mountains. Distribution/ Taxa Collecting sites Literature cited Zoogeography Kirby, 1837 semicuprea Băilor Valley Csiki, 1951 British Islands, Panzer, 1796 Southern Scandinavia, Central Donacia thalassina Valley of Ro şu Rivulet Csiki, 1951 Europe, Siberia, Germar, 1811 North western China, Japan S/EAP/TrPal Plateumaris sericea Valley of Ro şu Rivulet; Csiki, 1951; Europe, Caucasus, (Linnaeus, 1761) Săcel, Izvorul Izei (1 sp., Maican and Asia Minor, 6.VII.1995, P. C.) Serafim, 2004 Transcaspia, Siberia, Japan S\EAP\TrPal Plateumaris consimilis Rodna Mts. (without Petri, 1925; from Western Europe (Schrank, 1781) other specifications); Maican and to Japan Săcel, Izvorul Albastru al Serafim, 2001; S\EAP\TrPal Izei (3 sp., 6.VII.1995, P. Merkl, 2008 C.); S ăcel, Rodna Mts., Izvorul Izei, 850 m altitude

126 S. Maican – The fauna of the Rodna Mountains (123 ~ 140)

Distribution/ Taxa Collecting sites Literature cited Zoogeography Orsodacninae Thomson, 1866 Orsodacne cerasi Rodna Mts., Vinului Csiki, 1951; Europe, Caucasus, (Linnaeus, 1758) Valley, Valley of Saca Maican and Asia Minor, Western Rivulet, Valley of Ro şu Serafim, 2004 Siberia Rivulet; National Park S\EAP\SibE “Pietrosul Rodnei”, Laboratory House 1370 m altitude (1 sp., 11.VII.1995, P. C.); Izvorul Izei (1 sp., 6.VII.1995, P. C.) Zeugophorinae Chûjô, 1952 Zeugophora flavicollis Rodna Mts.; Valley of Csiki, 1951; Central and Western (Marsham, 1802) Iza River, 6 km upstream Maican and Europe Săcel (1 sp., VI.1998) Serafim, 2001 E/CE Criocerinae Latreille, 1807 Lilioceris merdigera Rodna Mts.; B ăilor Petri, 1925; from Iberian (Linnaeus, 1758) Valley Csiki, 1951 Peninsula, France to Japan S/EAP/TrPal Oulema erichsonii Săcel, Rodna Mts., Merkl, 2008 Central and Northern (Suffrian, 1841) Izvorul Izei, 850 m Europe altitude Oulema gallaeciana Săcel, Rodna Mts., Merkl, 2008 Europe, Danube (Heyden, 1870) Izvorul Izei, 850 m basin, western altitude Siberia, European S/EAP/SibE Oulema melanopus Bor şa, Rodna Mts., Merkl, 2008 from Ireland, (Linnaeus, 1758) Izvorul Bistri ţei, 1,700 southern and m; Sta ţiunea Bor şa, 1,544 Morocco to western m; S ăcel, Rodna Mts., Siberia and Mongolia Izvorul Izei, 850 m S/EAP/SibE altitude Oulema septentrionis Bor şa (Rodna Mts.), Szél et al., 1995 Central and Northern (Weise, 1880) 1,200 m altitude Europe, Volga basin Clythrinae Kirby, 1837 Labidostomis longimana Măriilor Valley (Some şul Csiki, 1951 Europe, Asia Minor, (Linnaeus, 1761) Mare Basin) Caucasus, Siberia, Mongolia S/EAP/SibE Smaragdina flavicollis Vinului Valley (Some şul Csiki, 1951 from France and (Charpentier, 1825) Mare Basin) Northern to and Northern Turkey; , E\CE Transylv. Rev. Syst. Ecol. Res. 9, (2010), "The Rodna Mountains National Park" 127

Distribution/ Taxa Collecting sites Literature cited Zoogeography Cryptocephalinae Gyllenhal, 1813 Cryptocephalus Rodna Mts., Ineu Peak, Petri, 1912; , , aureolus Ro şu Mountain, Vinului Csiki, 1951 Romania, Balkan Suffrian, 1847 Valley, M ăriilor Valley, Peninsula, Băilor Valley E\SbM\ESbM Cryptocephalus flavipes Corongi ş Peak (Rodna Csiki, 1951 Europe, Asia Minor, Fabricius, 1781 Mts.) Caucasus, Central Asia, Siberia S/EAP/SibE Cryptocephalus frenatus Valley of Iza River, 6 km Maican and Western France, Laicharting , 1781 upstream S ăcel (1 sp., Serafim, 2001 , , VI.1998) Danube and Nistru basins E\CE Cryptocephalus frontalis Rodna Mts.; Corongi ş Petri, 1912; Central Europe Marsham, 1802 Peak (Rodna Mts.) Csiki, 1951; E/CE Ro şca, 1973 Cryptocephalus Rodna Mts.; Saca Valley, Petri, 1912; Europe, hypochoeridis Vinului Valley, Csiki, 1951; Asia Minor, Southern (Linnaeus, 1758) MăriilorValley; Pietrosul Maican and Siberia Rodnei (1 sp., Serafim, 2001 S\EAP\SSibE 12.VII.1995, S. A.); Săcel, Izvorul Izei (1 sp., 6.VII.1996, P. C.); Bor şa, Rodna Mts. (2 sp., 21.VIII.1998, V. A.) Cryptocephalus moraei Rodna Mts., Vinului Petri, 1912; Europe (excepting (Linnaeus, 1758) Valey, valley of Ro şu Csiki, 1951; Northern Rivulet; S ăcel, Rodna Ro şca, 1973; Scandinavia) Mts., Izvorul Izei, 850 m Merkl, 2008 E/CE altitude Cryptocephalus Rodna (Some şul Mare Csiki, 1951 Europe, Asia Minor, ocellatus Drapiez, 1819 Basin) Iran, Kazakhstan, Western Siberia S/EAP/SibE Cryptocephalus Izvorul Albastru al Izei (1 Maican and Europe, Kazakhstan, octopunctatus (Scopoli, spec, 17.VI.1998) Serafim, 2001 Western Siberia 1763) S\EAP\SibE Cryptocephalus Vinului Valley (Some şul Csiki, 1951 Mid, East and South quadriguttatus Mare Basin) Eastern Europe, Richter, 1820 Caucasus, Kazakhstan, Western Siberia S\EAP\SibE Cryptocephalus Rodna Mts.; Corongi ş Petri, 1912; Central, East and quadripustulatus Peak (Rodna Mts.) Csiki, 1951 South Eastern Europe Gyllenhal, 1813 E/CE

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Distribution/ Taxa Collecting sites Literature cited Zoogeography Cryptocephalus querceti Vinului Valley (Some şul Csiki, 1951 North and Central Suffrian, 1848 Mare Basin) Europe, southwards to the E\CE Cryptocephalus sericeus Rodna Mts.; valley of Petri, 1912; Europe, Asia Minor, (Linnaeus, 1758) Saca Rivulet (Some şul Csiki, 1951; Kazachstan, Siberia, Mare Basin); Bor şa Szél et al., 1995 North Western China (Rodna Mts.), 1,200 m S/EAP/SibE altitude Cryptocephalus Săcel, Rodna Mts., Merkl, 2008 Northern Italy, Alps, transiens Franz, 1949 Izvorul Izei, 850 m basin of Danube altitude Pachybrachis Rodna (Some şul Mare Csiki, 1951 Europe, Asia Minor, hieroglyphicus Basin) Kazakhstan, Siberia (Laicharting, 1781) S/EAP/SibE Pachybrachis sinuatus Rodna Mts. Petri, 1912, Southern France, (Mulsant and Rey, 1859) 1925; Csiki, Central Europe, 1951 Balkans, Asia Minor E/CE Eumolpinae Thomson, 1859 Bromius obscurus Săcel, Rodna Mts., Iza Merkl, 2008 Europe, Urals, (Linnaeus, 1758) Valley, 850 m altitude Siberia, Kazakhstan, Central Asia, Mongolia, North China, Japan, North America S\H Chrysomelinae Latreille, 1802 Chrysolina carpathica Rodna Mts. Petri, 1912; Carpathians, Sudetes, (Fuss, 1856) Panin, 1944; Stara Planina Ro şca, 1974 EMount Chrysolina cerealis Rodna Mts. Panin, 1944 from Great Britain (Linnaeus, 1767) and Northern Spain to basins of Amur and Ussuri TrPal Chrysolina coerulans Bor şa (Rodna Mts.), Szél et al., 1995 from Central France, (Scriba, 1791) 1,600 m altitude Italy, Balkan to Asia Minor; E/CE Chrysolina cuprina Vinului Valley, valley of Csiki, 1951 in mountainous (Duftschmid, 1825) Saca Streamlet (Some şul regions from Central Mare bBsin) Europe; E/CEMount Chrysolina fastuosa Rodna Mts.; Vinului Petri, 1912; Europe, Caucasus, (Scopoli, 1763) Valley, valley of Saca Panin, 1944; Asia Minor, Central Rivulet, valley of Ro şu Csiki, 1951; and Western Siberia Rivulet, Rodna; S ăcel, Merkl, 2008 S/EAP/SibE Rodna Mts., Izvorul Izei, 850 m altitude Transylv. Rev. Syst. Ecol. Res. 9, (2010), "The Rodna Mountains National Park" 129

Distribution/ Taxa Collecting sites Literature cited Zoogeography Chrysolina fimbrialis Rodna Mts. Panin, 1944; Northern Italy, (Küster, 1845) Ro şca, 1974 Balkans, Danube River basin Chrysolina geminata Vinului Valley (Some şul Csiki, 1951 Europe, Caucasus, (Paykull, 1799) Mare Basin) Asia Minor E\CE Chrysolina globipennis Rodna Mts.; Vinului Petri, 1912; Western, Eastern and (Suffrian, 1851) Valley; S ăcel, Rodna Csiki, 1951; Southern Carpathians Mts., Izvorul Izei, 850 m Ro şca, 1974; CarpEnd altitude Merkl, 2008 Chrysolina Rodna Mts. Petri, 1912; Alps, Sudetes, hemisphaerica Panin, 1944; Carpathians, Balkans, (Germar, 1817) Csiki, 1951 EMount Chrysolina herbacea Săcel, Rodna Mts., Merkl, 2008 Europe, Caucasus, (Duftschmid, 1825) Izvorul Izei, 850 m Asia Minor, Iran, altitude Afghanistan, Altai S/EAP/SSibE Chrysolina lichenis Rodna Mts.; Ineu Peak Petri, 1912; Alps, Sudetes, (Richter, 1820) (Rodna Mts.); Valley of Csiki, 1951 Carpathians Ro şu Rivulet (Some şul EMount Mare Basin) Chrysolina marcasitica Rodna Mts.; Vinului Petri, 1912; Slovenia, Alps, (Germar, 1824) Valley, B ăilor Valley, Panin, 1944; Sudetes, Carpathians Ro şu Mountain, Corongi ş Csiki, 1951 EMount Peak (Rodna Mts.) Chrysolina marcasitica Săcel, Rodna Mts., Merkl, 2008 Sudetes, Carpathians turgida (Weise, 1882) Izvorul Izei, 850 m EMount altitude Chrysolina olivieri Rodna Mts. Csiki, 1951 Alps, Carpathians, (Bedel, 1892) Dinaric Alps, Stara Planina EMount Chrysolina polita Rodna Mts., Rodna Csiki, 1951 Europe, Western (Linnaeus, 1758) (Some şul Mare basin) Siberia, Mongolia S/EAP/TrPal Chrysolina Rodna Mts. Petri, 1912 Alps, Tatra, purpurascens Carpathians, Sudetes (Germar, 1822) EMount Chrysolina rufa Săcel, Rodna Mts., Merkl, 2008 Alps, Sudetes, crassicollis Izvorul Izei, 850 m Carpathians, Northern (Suffrian, 1851) altitude Balkans; EMount Chrysolina staphylaea Ro şu Mountain (Rodna Csiki, 1951 from Ireland and (Linnaeus, 1758) Mts.) Iceland to Kamchatka S/EAP/TrPal

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Distribution/ Taxa Collecting sites Literature cited Zoogeography Chrysolina umbratilis Rodna Mts.; valley of Petri, 1912, Carpathians, Sudetes, (Weise, 1887) Saca Rivulet; Bor şa, Panin, 1944; Alps, Dinaric Alps Rodna Mts., 2,000 m; Csiki, 1951; EMount Săcel, Rodna Mts., Ro şca, 1974; Izvorul Izei, 850 m Szél et al., altitude 1995; Merkl, 2008 Chrysolina varians Rodna Mts.; Vinului Petri, 1912; Europe, Asia Minor, (Schaller, 1783) Valley, valley of Saca Csiki, 1951; Siberia Streamlet, valley of Ro şu Szél et al., S\EAP\SibE Rivulet, Corongi ş Peak, 1995; Maican Ineu Peak, B ăilor Valley, and Serafim, Pietrosul Rodnei (1 sp., 2001; Merkl, VII.1995, M. I.); Bor şa 2008 (Rodna Mts.), 1,200 m altitude; S ăcel, Rodna Mts., Izvorul Izei, 850 m Chrysolina weisei Rodna Mts. (without Panin, 1944; Southern Carpathians (Frivaldszky, 1883) other specifications) Ro şca, 1974 CarpEnd Chrysomela collaris Rodna Mts. Csiki, 1951 from eastern France Linnaeus, 1758 to Siberia, Mongolia, North Eastern China S/EAP/TrPal Chrysomela cuprea Rodna; S ăcel, Izvorul Csiki, 1951; from eastern France Fabricius, 1775 Albastru al Izei (1 sp., Maican and to North-eastern Asia VI.1998) Serafim, 2001 S/EAP/TrPal Chrysomela populi Rodna Mts. Csiki, 1951 Palaearctic Linnaeus, 1758 S/EAP/HPal Chrysomela Rodna, Vinului Valley Csiki, 1951 From Eastern France vigintipunctata to Japan (Scopoli, 1763) S/EAP/TrPal Gastrophysa polygoni Bor şa (Rodna Mts.), Szél et al., 1995 Europe, Caucasus, (Linnaeus, 1758) 1,600 m altitude Asia Minor, Kazakhstan, Middle Asia, Siberia, Mongolia, China, Korea, North Africa. S\EAP\Hpal Gonioctena Vinului Valley Csiki, 1951 from the British Isles decemnotata to Japan (Marsham, 1802) S/EAP/TrPal Gonioctena pallida Săcel, Rodna Mts., Merkl, 2008 Western Palaearctic (Linnaeus, 1758) Izvorul Izei, 850 m altitude

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Distribution/ Taxa Collecting sites Literature cited Zoogeography Gastrophysa viridula Valley of Ro şu Rivulet, Csiki, 1951; Europe, Caucasus, (De Geer, 1775) Vinului Valley, B ăilor Szél et al., Asia Minor, Valley; Bor şa (Rodna 1995; Maican Kazakhstan, Mts.), 1,200-1,600 m; and Serafim, Mongolia, Kamchatka Pietrosul Rodnei, 800- 2001, 2004; S\EAP\TrPal 1,300 m altitude (18 spec, Merkl, 2008 11.VII.1995, M. I.); Bor şa (4 sp., 4.VIII.1995, P. S.); Izvorul Izei (9 sp., 9.IX.1995, P. C.); Lab. house “Pietrosul Rodnei”, 1,370-1,400 m altitude (32 sp., 12.IX.1996, P. C.; 26 sp., 11.-13.VII.1995, P. C.; 4 sp., 14.IX.1995, P. C.); Săcel, Izvorul Izei Gonioctena linnaeana Rodna Csiki, 1951 Europe, Asia Minor, (Schrank, 1781) Kazakhstan, Siberia, Mongolia S/EAP/TrPal Gonioctena Pietrosul Rodnei, Vinului Csiki, 1951 Northern, Central and quinquepunctata Valley, B ăilor Valley, Southern Europe (Fabricius, 1787) valley of Ro şu Rivulet Hydrothassa glabra Vinului Valley, B ăilor Csiki, 1951; Europe, West Siberia, (Herbst, 1783) Valley, valley of Ro şu Merkl, 2008 Morocco Rivulet; S ăcel, Rodna S\EAP\SibE Mts., Valea Izei, 850 m altitude Leptinotarsa Bor şa (Rodna Mts.), Szél et al., introduced in Europe decemlineata 1,200 m; Izvorul Albastru 1995; Maican from North America (Say, 1824) al Izei (1 sp., 6.VII.1995, and Serafim, P. C.); S ăcel, Rodna 2004; Merkl, Mts., Iza Valley, Izvorul 2008 Izei, 850 m altitude Oreina alpestris Rodna Mts.; Vinului Csiki, 1951 all mountains of (Schumell, 1824) Valley, valley of Saca Europe except Rivulet, B ăilor Valley, Scandinavia and Corongi ş Peak, Ineu Iberian Peninsula Peak EMount Oreina alpestris Săcel, Rodna Mts., Merkl, 2008 Romania, Poland, banatica (Weise, 1884) Izvorul Izei, Iza Valley, Ukraine 850 m altitude EMount Oreina bifrons decora Rodna Mts. Csiki, 1951; Carpathians, Sudetes (Richter, 1820) Petri, 1912 EMount

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Distribution/ Taxa Collecting sites Literature cited Zoogeography Linaeidea aenea Rodna Mts., Vinului Csiki, 1951; Europe, Palaearctic (Linnaeus, 1758) Valley, B ăilor Valley, Maican and Asia (including valley of Ro şu Rivulet, Serafim, 2001, Japan) valley of Saca Rivulet; 2004; Merkl, S\EAP\TrPal valley of Iza River, 6 km 2008 upstream S ăcel (4 sp., VI.1998, M. I.); Izvorul Albastru al Izei (1 sp., 17.VI.1998); Izvorul Izei (1 sp., 6.VII.1995, P. C.); Bor şa, Rodna Mts., Izvorul Bistri ţei, 1,665- 1,710 m altitude Oreina cacaliae Rodna Mts., Ineu Peak, Petri 1912; Carpathians, Sudetes senecionis Ro şu Mountain (Rodna Csiki, 1951; EMount (Schummel, 1844) Mts.), valley of Saca Merkl, 2008 Rivulet; S ăcel, Rodna Mts., Izvorul Izei, 850 m altitude Oreina coerulea Rodna Mts.; Vinului Petri 1912; mountains of (Olivier, 1790) Valley, Ineu Peak, valley Csiki, 1951; Southern Germany of Saca Rivulet; Pietrosul Maican and Sudetes, Carpathians, Rodnei, 1,600-2,050 m Serafim, 2001 Dinaric Alps, altitude (1 spec, 12.VII.1995, S. A.) EMount Oreina intricata Rodna Mts.; Ineu Peak, Petri, 1912; Carpathians, Sudetes, (Germar, 1824) Ro şu Mountain, Vinului Csiki, 1951; Alps, Balkans Valley; Bor şa (Rodna Szél et al., EMount Mts.), 1,600 m altitude; 1995; Maican Pietrosul Rodnei (1 sp., and Serafim, 13.VII.1995, P. C.); 2004; Merkl, National Park “Pietrosul 2008 Rodnei”, Laboratory House, 1,370 m altitude (1 sp., 12.IX.1996, P. C.); Săcel, Rodna Mts., Izvorul Izei, 850 m altitude Oreina plagiata Rodna Mts.; Corongi ş Petri, 1912; Apennines, Alps, (Suffrian, 1861) Peak (Rodna Mts.) Csiki, 1951 Balkans, Carpathians EMount Oreina speciosissima Rodna Mts.; Ineu Peak, Petri, 1912; Carpathians, Sudetes, (Scopoli, 1763) Ro şu Mountain, Vinului Csiki, 1951 Alps, Pyrenees, Valley, valley of Saca Balkans Rivulet Emount

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Distribution/ Taxa Collecting sites Literature cited Zoogeography Oreina virgulata Ineu Peak, valley of Saca Csiki, 1951 Carpathians, Sudetes, (Germar, 1824) Rivulet, valley of Ro şu Alps, Apennines, Rivulet, Vinului Valley Balkans; EMount Oreina virgulata ab. Săcel, Rodna Mts., Merkl, 2008 EMount praefica (Weise, 1884) Izvorul Izei, 850 m altitude Oreina viridis Pietrosul Rodnei; Bor şa Csiki, 1951; Alps, Vosges, (Duftschmid, 1825) (Rodna Mts.), 2,000 m Szél et al., 1995 Carpathians altitude EMount Phaedon cochleariae Rodna Mts.; valley of Petri, 1912; Europe, Palaearctic (Fabricius, 1792) Ro şu Rivulet, Vinului Csiki, 1951; Asia Valley; Izvorul Izei, Maican and S/EAP/TrPal Săcel (1 sp., 6.VII.1995, Serafim, 2001; P. C.); S ăcel, Rodna Merkl, 2008 Mts., Izvorul Izei, 850 m altitude Phaedon laevigatus Rodna Mts. Petri, 1912; Central, South and (Duftschmid, 1825) Csiki, 1951 East Europe; E\CE Phaedon segnis Pietrosul Rodnei (Rodna Petri, 1912; Carpathians, Alps, Weise, 1884 Mts.); Ineu Peak (Rodna Csiki, 1951; Dinaric Alps Mts.); Izvorul Izei (1 sp., Maican and EMount 6.VII.1995, P. C.) Serafim, 2001 Phratora atrovirens Rodna Mts. Petri, 1912; Alps, Fennoscandia, (Cornelius, 1857) Csiki, 1951 Germany, Poland, Russia, North Eastern Asia Phratora tibialis Bor şa (Rodna Mts.); Csiki, 1951; Europe, Asia Minor (Suffrian, 1851) Pietrosul Rodnei, 1,500- Maican and E\CE 2,050 m altitude (2 sp., Serafim, 2001 12.VII.1995, M. I.); Izvorul Albastru al Izei (2 sp., 9.IX.1995, P. C.) Phratora vitellinae Rodna, Vinului Valley, Csiki, 1951; Europe, (Linnaeus, 1758) Bor şa, Ineu Peak (Rodna Merkl, 2008 Palaearctic Asia Mts.); S ăcel, Rodna Mts., (excl. Japan) Izvorul Izei, 850 m S\EAP\TrPal altitude Plagiodera versicolora Rodna; valley of Iza Csiki, 1951; Palaearctic, India, (Laicharting, 1781) River, 1 km upstream Maican and Taiwan Săcel (1 sp., VI.1995, M. Serafim, 2001 S/EAP/HPal I.); Vadul Izei (2 sp., 9.VII.1995, M. I.); valley of Iza River, 6 km upstream S ăcel (2 sp., VI.1998)

134 S. Maican – The Leaf Beetle fauna of the Rodna Mountains (123 ~ 140)

Distribution/ Taxa Collecting sites Literature cited Zoogeography Sclerophaedon Rodna Mts.; Vinului Petri, 1912; Carpathians, Dinaric carniolicus (Germar, Valley, valley of Ro şu Csiki, 1951; Alps, Alps, Sudetes 1824) Rivulet, Corongi ş Peak; Merkl, 2008 EMount Săcel, Rodna Mts., Izvorul Izei, 850 m altitude Sclerophaedon Rodna Mts.; Vinului Petri, 1912; Eastern Carpathians carpathicus Valley (Some şul Mare Csiki, 1951 CarpEnd (Weise, 1875) basin) Timarcha metallica Ineu Peak (Rodna Mts.); Csiki, 1951; mountains of Central (Laicharting, 1781) “Pietrosul Mare” Nitzu et al., Europe and Balkans Scientific Reserve, 1755 2008 E/CEMount m altitude Timarcha rugulosa Rodna Mts.; S ăcel, Csiki, 1951; Southern Poland, Herrich-Schäffer, 1838 Rodna Mts., Valea Izei, Merkl, 2008 Moldavia, Southern 850 m altitude Carpathians, , Ukrainian Carpathians, Romania Latreille, 1802 Agelastica alni Valley of Ro şu Rivulet, Csiki, 1951; Europe, from Ireland (Linnaeus, 1758) Vinului Valley, valley of Maican and to Southern Finland Saca Rivulet, Corongi ş Serafim, 2001 and from the Peak (Rodna Mts.); Pyrenees to Caucasus, Săcel, Izvorul Albastru al Asia Minor Izei (1 sp., VI.1998) E\CE Galeruca tanaceti Valley of Saca Rivulet; Csiki, 1951; from Ireland and (Linnaeus, 1758) Bor şa, Rodna Mts., Pasul Merkl, 2008 Portugal to Korea; Prislop, 1,014 m a.s.l.; introduced in North Săcel, Rodna Mts., America Izvorul Izei, 850 m S/EAP/TrPal altitude; Bor şa, Rodna Mts., Pasul Prislop Galerucella lineola Vinului Valley (Some şul Csiki, 1951 from Ireland to Japan (Fabricius, 1781) Mare Basin) S/EAP/TrPal Lochmaea caprea Vinului Valley, valley of Csiki, 1951; from Spain and (Linnaeus, 1758) Saca Rivulet; S ăcel, Merkl, 2008 Ireland to Southern Rodna Mts., Izvorul Izei, Norway and Japan 850 m altitude S/EAP/TrPal viridipennis Borşa (Rodna Mts.), Szél et al., 1995 Alps, Carpathians, Germar, 1824 1,600 m altitude Balkans, Ural, Centr. Asia S\EAP\SibE Alticinae Newman, 1834 Altica helianthemi Pietrosul Rodnei, 1,400- Maican and Europe, Asia (Allard, 1859) 1,900 m altitude (1 sp., Serafim, 2004 S/EAP/SibE 14.IX.1995, P. C.)

Transylv. Rev. Syst. Ecol. Res. 9, (2010), "The Rodna Mountains National Park" 135

Distribution/ Taxa Collecting sites Literature cited Zoogeography Altica lythri Aubé, 1843 Rodna Mts. Gruev et al., Europe, Turkey 1993 E/CE Altica oleracea Rodna Mts. Csiki, 1951 Europe, Palaearctic (Linnaeus, 1758) Asia; S/EAP/TrPal Altica tamaricis Vinului Valley, valley of Csiki, 1951 Europe, Palaearctic Schrank, 1785 Saca Rivulet Asia S/EAP/TrPal Aphtona atrocaerulea Pietrosul Rodnei Gruev et al., Northern and Central (Stephens, 1831) 1993 Europe, South to Northern Italy and Romania Aphtona ovata Valley of Saca Rivulet Csiki, 1951 Central and Southern Foudras, 1860 Europe, Caucasus, Asia Minor E/CE Aphtona violacea Săcel, Izvorul Albastru al Maican and Europe, Siberia, (Koch, 1803) Izei (2 sp., 6.VII.1995, P. Serafim, 2001 Caucasus C.) S/EAP/SibE Batophila rubi Rodna Mts.; Vinului Petri, 1912; Europe, Caucasus, (Paykull, 1799) Valley; S ăcel, Rodna Csiki, 1951; Western Siberia Mts., Izvorul Izei, 850 m Konnerth , 1963; S\EAP\SibE altitude Merkl, 2008 Chaetocnema concinna Rodna Mts. Csiki, 1951 Europe, Palaearctic (Marsham, 1802) Asia, North Western Africa; S/EAAP/HPal Chaetocnema hortensis Rodna Mts. Petri, 1912; Europe, Sudan, (Geoffroy, 1785) Csiki, 1951 Palaearctic Asia, Madeira, North Western Africa S/EAP/HPal Chaetocnema tibialis Rodna Mts.; Ineu Peak Petri, 1912; Europe, Palaearctic (Illiger, 1807) Csiki, 1951; Asia, Northern Africa Konnerth- S/EAP/SibE Ionescu, 1963 Crepidodera aurata Pietrosul Rodnei; Rodna Csiki, 1951 Europe, Palaearctic (Marsham, 1802) Mts. Asia (except Japan), Morocco S/EAAP/HPal Dibolia occultans Baia Bor şa Gruev et al., Europe, Caucasus, (Koch, 1803) 1993 Asia Minor, North Western Africa; E/CE Longitarsus aeneicollis Măriilor Valley Csiki, 1951 Europe, Caucasus, (Faldermann, 1837) (Some şul Mare Basin) Asia Minor, Iran, Iraq, Syria, Central Asia, North Western Africa; E/CE

136 S. Maican – The Leaf Beetle fauna of the Rodna Mountains (123 ~ 140)

Distribution/ Taxa Collecting sites Literature cited Zoogeography Longitarsus brunneus Pietrosul Rodnei Csiki, 1951 Europe, Palaearctic (Duftschmid, 1825) Asia (excluding Japan); S\EAP\TrPal Longitarsus exsoletus Rodna (Some şul Mare Csiki, 1951 Europe, Caucasus, Linnaeus, 1758 basin) Asia Minor, Iran, Syria, Cyprus E/CE Longitarsus longiseta Săcel, Rodna Mts., Merkl, 2008 Europe, Siberia, Weise, 1889 Izvorul Izei, 850 m Japan and Northern altitude China Longitarsus lycopi Pietrosul Rodnei, 1,400- Maican and Europe, Caucasus, (Foudras, 1860) 1,900 m altitude (1 sp. Serafim, 2004 Asia Minor, Iran, 14.IX.1995, P. C.) Central Asia, Northern Africa; E/CE Longitarsus Pietrosul Rodnei Csiki, 1951 Alps, Pyrénées, pallidicornis Kutschera, Carpathians, Dinaric 1863 Alps; EMount Longitarsus parvulus Bor şa (Rodna Mts.), Szél et al., 1995 Europe, Caucasus, (Paykull, 1799) 1,600 m altitude Asia Minor, Israel, Siberia; S/EAP/SibE Longitarsus pinguis Rodna Mts. Petri, 1912, Northern Italy, Weise, 1888 1925 Danube River basin, Balkan Peninsula, Ukraine; E/CE Longitarsus rubellus Pietrosul Rodnei Csiki, 1951 Alps, Sudetes, (Foudras, 1860) Carpathians EMount Longitarsus suturellus Pietrosul Rodnei, Rodna Petri, 1912; Europe, Palaearctic (Duftschmid, 1825) Mts., Vinului Valley, Csiki, 1951; Asia Rodna Veche; S ăcel, Konnerth-I., S/EAP/TrPal Rodna Mts., Izvorul Izei, 1963; Gruev et 850 m altitude al., 1993; Merkl, 2008 Minota carpathica Rodna Mts., Pietrosul Petri, 1912; Carpathians Sudetes, Heikertinger, 1911 Rodnei, Ineu Peak, Csiki, 1951; Alps Corongi ş Peak, Ro şu Szél et al., 1995 EMount Mountain, Valley of Saca Rivulet; Bor şa (Rodna Mts.), 1,600 m altitude Minota obesa Pietrosul Rodnei, Rodna Konnerth- mountains and (Waltl, 1839) Mts. Ionescu, 1963 submountains areas of (with older Western and Central data) Europe excepting Pyrenees; EMount Transylv. Rev. Syst. Ecol. Res. 9, (2010), "The Rodna Mountains National Park" 137

Distribution/ Taxa Collecting sites Literature cited Zoogeography Mniophila muscorum Rodna Mts.; Bor şa Petri, 1912; Europe, Caucasus (Koch, 1803) (Rodna Mts.), 1,600 m; Szél et al., E/CE Săcel, Rodna Mts., 1995; Merkl, Izvorul Izei, 850 m 2008; Nitzu et altitude; “Pietrosul al., 2008 Mare” Scientific Reserve, 2,047 m altitude Neocrepidodera Pietrosul Rodnei Frivaldszky, Alps, Carpathians cyanescens 1871 AlpCarpEnd (Duftschmid, 1825) Neocrepidodera Rodna Mts.; S ăcel, Csiki, 1951; boreo-alpine species; femorata (Gyllenhal, Rodna Mts., Izvorul Izei, Konnerth- mountainous areas of 1813) 850 m altitude Ionescu, 1963; Europe and Siberia Merkl, 2008 Neocrepidodera Pietrosul Rodnei (1 sp., Maican and Alps, Apeninnes, melanostoma 14.IX.1995, P. C.) Serafim, 2001 Carpathians, Dinaric, (Redtenbacher, 1849) Vosges; EMount Neocrepidodera Pietrosul Rodnei; Ineu Petri, 1912, Endemic species to transsilvanica Peak (Rodna Mts.); Csiki, 1951; the Eastern and (Fuss, 1864) valley of Saca Rivulet; Konnerth- Southern Carpathians Bor şa (Rodna Mts.), Ionescu, 1963; 1,600 m altitude Szél et al., 1995 Orestia aubei Pietrosul Rodnei, Rodna Petri, 1912; Carpathians, Allard, 1859 Mts.; valley of Saca Csiki, 1951; Transylvania, Dinaric Rivulet Konnerth- Alps Ionescu, EMount 1963 Phyllotreta atra Rodna Mts.; Ineu Peak, Petri, 1912; Europe, Palaearctic (Fabricius, 1775) Valea Vinului, Valley of Csiki, 1951; Asia, Morocco, Saca rivulet; Bor şa Konnerth- Yemen (Rodna Mts., 1,600 m Ionescu, 1963; S/EAAP/HPal altitude) Gruev et al., 1993; Szél et al., 1995 Phyllotreta christinae Bor şa (Rodna Mts.), Szél et al., 1995 mountains of Central Heikertinger, 1941 1,600 m altitude Europe and Balkan Peninsula; E/Mount Phyllotreta cruciferae Rodna Mts.; Vinului Petri, 1912; Europe, Caucasus, (Goeze, 1777) Valley Csiki, 1951; Asia Minor, Central Konnerth- Asia, Siberia, Ionescu, 1963; Mongolia, Northern Gruev et al, Africa; S/EAP/SibE 1993

138 S. Maican – The Leaf Beetle fauna of the Rodna Mountains (123 ~ 140)

Distribution/ Taxa Collecting sites Literature cited Zoogeography Phyllotreta diademata Rodna Mts.; Vinului Petri, 1912; Europe, Central Asia, Foudras, 1860 Valley Csiki, 1951; India Konnerth-I., S/EAP/SibE 1963; Gruev et al., 1993 Phyllotreta flexuosa Rodna Mts.; Vinului Petri, 1912; Boreal part of (Illiger, 1794) Valley Csiki, 1951; Palaearctic Region Konnerth-I., 1963; Gruev et al., 1993 Phyllotreta nemorum Vinului Valley; Pietrosul Csiki, 1951; Europe, Palaearctic (Linnaeus, 1758) Rodnei Gruev et al., Asia (excl. Japan) 1993 S/EAP/SibE Phyllotreta striolata Pietrosul Rodnei (1 sp., Gruev et al., Europe, Asia, (Fabricius, 1803) 14.IX.1995, H. C.) 1993; Indonesia Maican and S/EAP/TrPal Serafim, 2001 Phyllotreta undulata Rodna Mts., Vinului Petri, 1912; Europe, Palaearctic (Kutschera, 1860) Valley, valley of Saca Konnerth-I., Asia Rivulet, Pietrosul Rodnei, 1963; Gruev et S/EAP/TrPal Săcel, Izvorul Izei (1 sp., al., 1993; Pausescu 6.VII.1995, P. C.) Maican and Serafim, 2001 Phyllotreta vittula Rodna Mts., Ineu Petri, 1912; Europe, Palaearctic (Redtenbacher, 1849) Peak; valley of Saca Csiki, 1951; Asia (excluding Rivulet; Pietrosul Konnerth- Japan) Rodnei; Bor şa (Rodna Ionescu, 1963; S/EAP/TrPal Mts.), 1,600 m altitude; Gruev et al., Pietrosul Rodnei (1 sp., 1993; Szél et 14.IX.1995, H. C.) al., 1995; Maican and Serafim, 2001 Psylliodes aereus Rodna Mts. Gruev et al., Emount Foudras, 1860 1993 (probably ssp. austriaca Heikertinger, 1911) Psylliodes affinis Rodna Mts.; Rodna Csiki, 1951; Europe, Kazakhstan, (Paykull, 1799) Veche; Vinului Valley Gruev et al., Caucasus, Siberia, 1993 Altai, Morocco S/EAP/SibE Psylliodes attenuatus Rodna Mts. Petri, 1912; Europe, Palaearctic (Koch, 1803) Csiki, 1951; Asia Konnerth-I., S/EAP/TrPal 1963; Gruev et al., 1993 Transylv. Rev. Syst. Ecol. Res. 9, (2010), "The Rodna Mountains National Park" 139

Distribution/ Taxa Collecting sites Literature cited Zoogeography Psylliodes cucullatus Pietrosul Rodnei Gruev et al., TrPal (Illiger, 1807) 1993 Psylliodes frivaldszkyi Rodna Mts.; Pietrosul Petri, 1912; Carpathians Weise, 1888 Rodnei; Ineu Peak Csiki, 1951; CarpEnd (Rodna Mts.); Rodna Konnerth-I., Veche; valley of Saca 1963; Gruev et Rivulet al., 1993 Psylliodes glaber Pietrosul Rodnei Csiki, 1951 Alps, (Duftschmid, 1825) Carpathians, Dinaric Alps EMount Psylliodes napi Săcel, Rodna Mts., Merkl, 2008 Europe, (Fabricius, 1792) Izvorul Izei, 850 m Palaearctic Asia altitude (except Japan), North Western Africa S/EAAP/HPal Psylliodes subaeneus Rodna Mts. Petri, 1912; Alps, Eastern and Kutschera, 1867 Csiki, 1951; Southern Carpathians, Gruev et al., Slovenia 1993 EMount Cassidinae Gyllenhal, 1813 Cassida murraea Vinului Valley Csiki, 1951 Palaearctic (excluding Linnaeus, 1767 northern Africa) S/EAP/TrPal Cassida viridis Vinului Valley, valley of Csiki, 1951; Palaearctic Linnaeus, 1758 Saca Rivulet; S ăcel, Merkl, 2008 S/EAAP/HPal Rodna Mts., Izvorul Izei, 850 m altitude Cassida vittata Rodna Mts. Petri, 1912; from Morocco and Villers, 1789 Csiki, 1951 Portugal to Japan TrPal

CONCLUSIONS Integrating the data obtained from the study of material preserved in the collections of “Grigore Antipa” National Musem of Nature History from Bucharest, with those from the available coleopterological literature, a synthesis concerning the diversity of the leaf beetles fauna in the Rodna Mountains (Maramure ş, northern Romania) is given. The chrysomelid fauna from this area is relatively well known, being represented by 138 species from 43 genera, belonging to 11 subfamilies. Among them, Neocrepidodera transsilvanica (Fuss), Psylliodes frivaldszkyi Weise, Chrysolina weisei (Frivaldszky) and Sclerophaedon carpathicus Weise are endemic species to the . From a zoogeographical point of view, the Siberian and European complexes are dominant in the Rodna Mountains.

140 S. Maican – The Leaf Beetle fauna of the Rodna Mountains (123 ~ 140)

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