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From the Ordovician (Darriwillian) of Morocco
Palaeogeographic implications of a new iocrinid crinoid (Disparida) from the Ordovician (Darriwillian) of Morocco Samuel Zamora1, Imran A. Rahman2 and William I. Ausich3 1 Instituto Geologico´ y Minero de Espana,˜ Zaragoza, Spain 2 School of Earth Sciences, University of Bristol, Bristol, United Kingdom 3 School of Earth Sciences, Ohio State University, Columbus, OH, United States ABSTRACT Complete, articulated crinoids from the Ordovician peri-Gondwanan margin are rare. Here, we describe a new species, Iocrinus africanus sp. nov., from the Darriwilian-age Taddrist Formation of Morocco. The anatomy of this species was studied using a combination of traditional palaeontological methods and non-destructive X-ray micro-tomography (micro-CT). This revealed critical features of the column, distal arms, and aboral cup, which were hidden in the surrounding rock and would have been inaccessible without the application of micro-CT. Iocrinus africanus sp. nov. is characterized by the presence of seven to thirteen tertibrachials, three in-line bifurcations per ray, and an anal sac that is predominantly unplated or very lightly plated. Iocrinus is a common genus in North America (Laurentia) and has also been reported from the United Kingdom (Avalonia) and Oman (middle east Gondwana). Together with Merocrinus, it represents one of the few geographically widespread crinoids during the Ordovician and serves to demonstrate that faunal exchanges between Laurentia and Gondwana occurred at this time. This study highlights the advantages of using both conventional -
Biostratigraphic Precision of the Cruziana Rugosa Group: a Study from the Ordovician Succession of Southern and Central Bolivia
Geol. Mag. 144 (2), 2007, pp. 289–303. c 2007 Cambridge University Press 289 doi:10.1017/S0016756807003093 First published online 9 February 2007 Printed in the United Kingdom Biostratigraphic precision of the Cruziana rugosa group: a study from the Ordovician succession of southern and central Bolivia SVEN O. EGENHOFF∗, BERND WEBER†, OLIVER LEHNERT‡ &JORG¨ MALETZ§ ∗Colorado State University, Department of Geosciences, 322 Natural Resources Building, Fort Collins, CO 80523-1482, USA †Freie Universitat¨ Berlin, Institut fur¨ Geologische Wissenschaften, Fachrichtung Geologie, Malteserstrasse 74-100, D-12249 Berlin, Germany ‡University of Erlangen, Institute of Geology and Mineralogie, Schlossgarten 5, D-91054 Erlangen, Germany §Department of Geology, State University of New York at Buffalo, 772 Natural Sciences and Mathematics Complex, Buffalo, New York 14260-3050, USA (Received 10 October 2005; revised version received 1 May 2006; accepted 22 May 2006) Abstract – Cruziana ichnospecies have been repeatedly reported to have biostratigraphic significance. This study presents a re-evaluation of the arthropod ichnotaxa of the Cruziana rugosa Group from bio- and/or lithostratigraphically well-defined Lower to Upper Ordovician siliciclastic sections of southern and central Bolivia. With the exception of Cruziana rouaulti, the ichnofaunas contain all the members of the Cruziana rugosa Group throughout the Ordovician (Arenig to Caradoc) successions in Bolivia. The Bolivian material therefore indicates that these arthropod ichnofossil assemblages are suitable for recognizing Ordovician strata in Bolivia. These findings cast doubt on their use as reliable indicators for a global intra-Ordovician (Arenig to Caradoc) biozonation of Peri-Gondwanan sedimentary successions. Keywords: Cruziana, biostratigraphy, Bolivia, Ordovician. 1. Introduction to the present study. -
Conodonts in Ordovician Biostratigraphy
View metadata, citation and similar papers at core.ac.uk brought to you by CORE provided by Archivio istituzionale della ricerca - Università di Modena e Reggio Emilia 1 Conodonts in Ordovician biostratigraphy STIG M. BERGSTRÖM AND ANNALISA FERRETTI Conodonts in Ordovician biostratigraphy The long time interval after Pander’s (1856) original conodont study can in terms of Ordovician conodont biostratigraphic research be subdivided into three periods, namely the Pioneer Period (1856-1955), the Transition Period (1955-1971), and the Modern Period (1971-Recent). During the pre-1920s, the few published conodont investigations were restricted to Europe and North America and were not concerned about the potential use of conodonts as guide fossils. Although primarily of taxonomic nature, the pioneer studies by Branson & Mehl, Stauffer, and Furnish during the 1930s represent the beginning of the use of conodonts in Ordovician biostratigraphy. However, no formal zones were introduced until Lindström (1955) proposed four conodont zones in the Lower Ordovician of Sweden, which marks the end of the Pioneer Period. Because Lindström’s zone classification was not followed by similar work outside Baltoscandia, the time interval up to the late 1960s can be regarded as a Transition Period. A milestone symposium volume, entitled ‘Conodont Biostratigraphy’ and published in 1971, 2 summarized much new information on Ordovician conodont biostratigraphy and is taken as the beginning of the Modern Period of Ordovician conodont biostratigraphy. In this volume, the Baltoscandic Ordovician was subdivided into named conodont zones whereas the North American Ordovician succession was classified into a series of lettered or numbered Faunas. Although most of the latter did not receive zone names until 1984, this classification has been used widely in North America. -
International Geological Correlation Programme. Project 591 "The Early to Middle Paleozoic Revolution"
International Geological Correlation Programme. Project 591 "The Early to Middle Paleozoic Revolution". Proceedings of the 3rd IGCP 591 Annual Meeting : Lund, Sweden, 9–19 June 2013 Lindskog, Anders; Mehlqvist, Kristina 2013 Link to publication Citation for published version (APA): Lindskog, A., & Mehlqvist, K. (Eds.) (2013). International Geological Correlation Programme. Project 591 "The Early to Middle Paleozoic Revolution". Proceedings of the 3rd IGCP 591 Annual Meeting : Lund, Sweden, 9–19 June 2013. (International Geological Correlation Programme, UNESCO; Vol. 3). Department of Geology, Lund University. Total number of authors: 2 General rights Unless other specific re-use rights are stated the following general rights apply: Copyright and moral rights for the publications made accessible in the public portal are retained by the authors and/or other copyright owners and it is a condition of accessing publications that users recognise and abide by the legal requirements associated with these rights. • Users may download and print one copy of any publication from the public portal for the purpose of private study or research. • You may not further distribute the material or use it for any profit-making activity or commercial gain • You may freely distribute the URL identifying the publication in the public portal Read more about Creative commons licenses: https://creativecommons.org/licenses/ Take down policy If you believe that this document breaches copyright please contact us providing details, and we will remove access to the work immediately and investigate your claim. LUND UNIVERSITY PO Box 117 221 00 Lund +46 46-222 00 00 ANDERS LINDSKOG | KRISTINA MEHLQVIST Printed by Media-Tryck, Lund 2013 Proceedings of the 3rd IGCP 591 Annual Meeting Proceedings of the 3 Lund, Sweden, 9–19 June 2013 EDITED BY ANDERS LINDSKOG | KRISTINA MEHLQVIST DEPARTMENT OF GEOLOGY | LUND UNIVERSITY The abstracts within this volume were presented at the 3rd IGCP 591 Annual rd Meeting, which was held in Lund, Sweden, in June 2013. -
Preliminary Geologic Map of the Baird Mountains and Part of the Selawik Quadrangles, Alaska By
preliminary Geologic Map of the Baird Mountains and part of the Selawik Quadrangles, Alaska by S.M. Karl, J.A. Dumoulin, Inyo Ellersieck, A.G. Harris, and J.M. Schmidt Open-File Report 89-551 This map is preliminary and has not been reviewed for conformity with the North American stratigraphic Code. Any use of trade, product, or firm names is for descriptive purposes only and does not imply endorsement by the U.S. Government. Table Contents ~ntroduction.......................................... 1 Stratigraphic Framework .................................1 Structural Framework ....................................4 Acknowledgments .......................................6 Unit Descriptions ......................................6 Kc ............................................. 6 KJm .............................................6 JPe .............................................7 ~zb................ '. ... ., ,....................... 8 Mzg ............................................10 MzPzi ..........................................10 MZPZ~.......................................... 11 PMC ............................................12 PD1 ............................................12 Mk0 ............................................12 M1 .............................................13 MDue ...........................................14 Mlt ............................................15 Mk .............................................15 MD1 ............................................16 MDe ............................................16 MD~........................................... -
Crossing the Several Scales of Strain-Accomplishing Mechanisms in the Hinterland of the Central Andean Fold±Thrust Belt, Bolivia
Journal of Structural Geology 24 02002) 1587±1602 www.elsevier.com/locate/jstrugeo Crossing the several scales of strain-accomplishing mechanisms in the hinterland of the central Andean fold±thrust belt, Bolivia Nadine McQuarriea,b,*, George H.Davis a aDepartment of Geosciences, University of Arizona, Tucson, AZ 85721, USA bDivision of Geological andPlanetary Science, California Institute of Technology, Pasadena, CA 91125, USA Received 14 November 2000; revised 28 October 2001; accepted 29 October 2001 Abstract Depictions of structures at outcrop, regional and tectonic scales enforce horizontal shortening and vertical thickening as the predominant style of deformation at all scales within the hinterland of the central Andean fold±thrust belt.Outcrop-scale structures document a progression of strain that created: 01) ¯exural-slip folds, 02) fold ¯attening via axial-planar cleavage, 03) vertical stretching via boudinage and late-stage faulting and, ®nally, 04) kink folding.These examples of intraformational deformation are generally concentrated just beyond the tip lines of thrust faults, where fault-propagation folds and related structures are well developed.Fault-propagation folding accommo- dated the accrual of strain indicated by outcrop-scale structures while the structures themselves indicate how deformation developed within each individual fold.Fault-propagation fold geometries at a regional scale emerge from the construction of regional balanced cross-sections. The sections were drawn with careful attention to: 01) known map relationships, -
CONODONTS of the MOJCZA LIMESTONE -.: Palaeontologia Polonica
CONODONTS OF THE MOJCZA LIMESTONE JERZY DZIK Dzik, J. 1994. Conodonts of the M6jcza Limestone. -In: J. Dzik, E. Olemp ska, and A. Pisera 1994. Ordovician carbonate platform ecosystem of the Holy Cross Moun tains. Palaeontologia Polonica 53, 43-128. The Ordovician organodetrital limestones and marls studied in outcrops at M6jcza and Miedzygorz, Holy Cross Mts, Poland, contains a record of the evolution of local conodont faunas from the latest Arenig (Early Kundan, Lenodus variabilis Zone) to the Ashgill (Amorphognathus ordovicicus Zone), with a single larger hiatus corre sponding to the subzones from Eop/acognathus pseudop/anu s to E. reclinatu s. The conodont fauna is Baltic in general appearance but cold water genera , like Sagitto dontina, Scabbardella, and Hamarodus, as well as those of Welsh or Chinese af finities, like Comp/exodus, Phragmodus, and Rhodesognathu s are dominant in par ticular parts of the section while others common in the Baltic region, like Periodon , Eop/acognathus, and Sca/pellodus are extremely rare. Most of the lineages continue to occur throughout most of the section enabling quantitative studies on their phyletic evolut ion. Apparatuses of sixty seven species of thirty six genera are described and illustrated. Phyletic evolution of Ba/toniodus, Amorphognathu s, Comp/exodus, and Pygodus is biometrically documented. Element s of apparatu ses are homolog ized and the standard notation system is applied to all of them. Acodontidae fam. n., Drepa nodus kie/censis sp. n., and D. santacrucensis sp. n. are proposed . Ke y w o r d s: conodonts, Ordovici an, evolut ion, taxonomy. Jerzy Dzik, Instytut Paleobiologii PAN, A/eja Zwirk i i Wigury 93, 02-089 Warszawa , Poland. -
Carboniferous Formations and Faunas of Central Montana
Carboniferous Formations and Faunas of Central Montana GEOLOGICAL SURVEY PROFESSIONAL PAPER 348 Carboniferous Formations and Faunas of Central Montana By W. H. EASTON GEOLOGICAL SURVEY PROFESSIONAL PAPER 348 A study of the stratigraphic and ecologic associa tions and significance offossils from the Big Snowy group of Mississippian and Pennsylvanian rocks UNITED STATES GOVERNMENT PRINTING OFFICE, WASHINGTON : 1962 UNITED STATES DEPARTMENT OF THE INTERIOR STEWART L. UDALL, Secretary GEOLOGICAL SURVEY Thomas B. Nolan, Director The U.S. Geological Survey Library has cataloged this publication as follows : Eastern, William Heyden, 1916- Carboniferous formations and faunas of central Montana. Washington, U.S. Govt. Print. Off., 1961. iv, 126 p. illus., diagrs., tables. 29 cm. (U.S. Geological Survey. Professional paper 348) Part of illustrative matter folded in pocket. Bibliography: p. 101-108. 1. Paleontology Montana. 2. Paleontology Carboniferous. 3. Geology, Stratigraphic Carboniferous. I. Title. (Series) For sale by the Superintendent of Documents, U.S. Government Printing Office Washington 25, B.C. CONTENTS Page Page Abstract-__________________________________________ 1 Faunal analysis Continued Introduction _______________________________________ 1 Faunal relations ______________________________ 22 Purposes of the study_ __________________________ 1 Long-ranging elements...__________________ 22 Organization of present work___ __________________ 3 Elements of Mississippian affinity.._________ 22 Acknowledgments--.-------.- ___________________ -
Catalog of Type Specimens of Invertebrate Fossils: Cono- Donta
% {I V 0> % rF h y Catalog of Type Specimens Compiled Frederick J. Collier of Invertebrate Fossils: Conodonta SMITHSONIAN CONTRIBUTIONS TO PALEOBIOLOGY NUMBER 9 SERIAL PUBLICATIONS OF THE SMITHSONIAN INSTITUTION The emphasis upon publications as a means of diffusing knowledge was expressed by the first Secretary of the Smithsonian Institution. In his formal plan for the Insti tution, Joseph Henry articulated a program that included the following statement: "It is proposed to publish a series of reports, giving an account of the new discoveries in science, and of the changes made from year to year in all branches of knowledge." This keynote of basic research has been adhered to over the years in the issuance of thousands of titles in serial publications under the Smithsonian imprint, com mencing with Smithsonian Contributions to Knowledge in 1848 and continuing with the following active series: Smithsonian Annals of Flight Smithsonian Contributions to Anthropology Smithsonian Contributions to Astrophysics Smithsonian Contributions to Botany Smithsonian Contributions to the Earth Sciences Smithsonian Contributions to Paleobiology Smithsonian Contributions to Zoology Smithsonian Studies in History and Technology In these series, the Institution publishes original articles and monographs dealing with the research and collections of its several museums and offices and of profes sional colleagues at other institutions of learning. These papers report newly acquired facts, synoptic interpretations of data, or original theory in specialized fields. These publications are distributed by mailing lists to libraries, laboratories, and other in terested institutions and specialists throughout the world. Individual copies may be obtained from the Smithsonian Institution Press as long as stocks are available. -
Lee-Riding-2018.Pdf
Earth-Science Reviews 181 (2018) 98–121 Contents lists available at ScienceDirect Earth-Science Reviews journal homepage: www.elsevier.com/locate/earscirev Marine oxygenation, lithistid sponges, and the early history of Paleozoic T skeletal reefs ⁎ Jeong-Hyun Leea, , Robert Ridingb a Department of Geology and Earth Environmental Sciences, Chungnam National University, Daejeon 34134, Republic of Korea b Department of Earth and Planetary Sciences, University of Tennessee, Knoxville, TN 37996, USA ARTICLE INFO ABSTRACT Keywords: Microbial carbonates were major components of early Paleozoic reefs until coral-stromatoporoid-bryozoan reefs Cambrian appeared in the mid-Ordovician. Microbial reefs were augmented by archaeocyath sponges for ~15 Myr in the Reef gap early Cambrian, by lithistid sponges for the remaining ~25 Myr of the Cambrian, and then by lithistid, calathiid Dysoxia and pulchrilaminid sponges for the first ~25 Myr of the Ordovician. The factors responsible for mid–late Hypoxia Cambrian microbial-lithistid sponge reef dominance remain unclear. Although oxygen increase appears to have Lithistid sponge-microbial reef significantly contributed to the early Cambrian ‘Explosion’ of marine animal life, it was followed by a prolonged period dominated by ‘greenhouse’ conditions, as sea-level rose and CO2 increased. The mid–late Cambrian was unusually warm, and these elevated temperatures can be expected to have lowered oxygen solubility, and to have promoted widespread thermal stratification resulting in marine dysoxia and hypoxia. Greenhouse condi- tions would also have stimulated carbonate platform development, locally further limiting shallow-water cir- culation. Low marine oxygenation has been linked to episodic extinctions of phytoplankton, trilobites and other metazoans during the mid–late Cambrian. -
Version 4.Cdr
THE GEOMAGNETIC POLARITY TIME SCALE FOR EARY PALEOZOIC: DATA AND SYNTHESE 1 2 1 Vladimir Pavlov and Yves Gallet 2 Institute of Physics of the Earth, Institut de Physique Russian Academy of Sciences INTRODUCTION EARLY CAMBRIAN: Kirschvink and LONG STANDING CONROVERSY: Constructing the Geomagnetic Polarity Time Scale (GPTS) through the geological history is of crucial importance to address Rozanov, 1984 TWO PRIMARY POLES FOR ONE SIBERIAN PLATFORM? several major issues in Earth sciences, as for instance the long-term evolution of the geodynamo. The GPTS is also an Khramov et al., 1982 important chronological tool allowing one to decipher the age and duration of various geological processes. Moreover, GPTS 509 Ma is widely used in prospecting for commercial minerals and in petroleum geology Kirschvink’s direction OR Khramov’s direction? Series Regiostage . N N The chronology of the geomagnetic polarity reversals since the Upper Jurassic is rather well known thanks to the magnetic Kirschvink and Pisarevsky anomalies recorded in the sea floor. For more ancient epochs our knowledge of the GPTS is still very fragmentary and much Rozanov, 1984 et al., 1997 more uncertain. However significant information has been obtained for several time intervals, in particular for the Triassic and the beginning of the Phanerozoic. Hirnantian Khondelen We will focus our presentation on recent developments made in the determination of the GPTS during the Early Paleozoic P-T Botoma - Toyon Botoma - Botoma - Toyon Botoma - 516 Ma (Siberia) (Cambrian and Ordovician). Siberian APWP Ashgill Epoch 2 Epoch 2 D3-C1 экватор E30° Е90° Е150° Botmoynak Dolborian (Tianshan) Late Ediacarian - Hyperactivity of the Earth magnetic field? Early Cambrian Kirschvink’s pole Early Cambrian O2-O3 Cm3-O1 Khramov’s pole S30° Moyero N Cm2 N N Low (very low?) reversal frequency (Siberia) Pavlov et al., Pavlov et al., 2018 S60° unpubl. -
ORDOVICIAN FISH from the ARABIAN PENINSULA by IVAN J
[Palaeontology, Vol. 52, Part 2, 2009, pp. 337–342] ORDOVICIAN FISH FROM THE ARABIAN PENINSULA by IVAN J. SANSOM*, C. GILES MILLER , ALAN HEWARDà,–, NEIL S. DAVIES*,**, GRAHAM A. BOOTHà, RICHARD A. FORTEY and FLORENTIN PARIS§ *Earth Sciences, University of Birmingham, Birmingham B15 2TT, UK; e-mail: [email protected] Department of Palaeontology, The Natural History Museum, London SW7 5BD, UK; e-mails: [email protected] and [email protected] àPetroleum Development Oman, Muscat, Oman; e-mail: [email protected] §Ge´osciences, Universite´ de Rennes, 35042 Rennes, France; e-mail: fl[email protected] –Present address: Petrogas E&P, Muscat, Oman; e-mail: [email protected] **Present address: Earth Sciences, Dalhousie University, Halifax, Nova Scotia B3H 3J5, Canada; e-mail: [email protected] Typescript received 25 February 2008; accepted in revised form 19 May 2008 Abstract: Over the past three decades Ordovician pteras- morphs from the Arabian margin of Gondwana. These are pidomorphs (armoured jawless fish) have been recorded among the oldest arandaspids known, and greatly extend from the fringes of the Gondwana palaeocontinent, in par- the palaeogeographical distribution of the clade around the ticular Australia and South America. These occurrences are periGondwanan margin. Their occurrence within a very dominated by arandaspid agnathans, the oldest known narrow, nearshore ecological niche suggests that similar group of vertebrates with extensive biomineralisation of Middle Ordovician palaeoenvironmental settings should be the dermoskeleton. Here we describe specimens of arandas- targeted for further sampling. pid agnathans, referable to the genus Sacabambaspis Gagnier, Blieck and Rodrigo, from the Ordovician of Key words: Ordovician, pteraspidomorphs, Gondwana pal- Oman, which represent the earliest record of pteraspido- aeocontinent, Sacabambaspis, Oman.