SOUTHERN CALIFORNIA ASSOCIATION OF MARINE INVERTEBRATE TAXONOMISTS NEWSLETTER

July, 1982 Vol. 1, Ho. k

Next Scheduled Meeting: August 16, 1982

Place: Marine Biological Consultants 947 NewhalJ Street Costa Mesa, CA 92627

Guest Speakers: Dave Montagne on Phyllodocids (postponed from July meeting ? due to Dave's injury from a rockf tsh). Fred Pilz on Typosyl1i s

Specimen Exchange Group: Phoxocephalidae

Top,ic Taxonomic Group: Hestonidae, Pilargidae, Typosyl1 is

MINUTES FROM JULY 19, 1982:

Charter

The Charter was discussed and generally was acceptable, Three areas were modified at the meeting.

The position of Curator was changed from an elected position to an appointed office. A provision was added that curators should house the collection at the institute where they are employed and 0*> to move the collection upon change of curators. The definition of charter membership was changed to include all persons who join in the first year. {Motion by Don Cadien and Ann ^•^^ Martin, second by Larry Lovetl, passed by 2/3 majority.)

- Elections for officers will take place in April, the first official #t election will be in April, 1983 when current officers will run as incumbents. {Motion by Don Cadien, second by Don Maurer, passed by 2/3 majori ty.)

After the meeting was adjourned, more changes to the charter were brought up within the Charter Committee. These will be discussed at the August meeting before a vote of final acceptance will be held. Karen Green of Pholoides aspera

Karen looked at all 20 specimens of P_. aspera from the exchange. She found that seta! types were the same in all specimens and that P_. aspera is a valid name for the organism.

Literature Donations

Because the literature distributed from Rick Klink's collection was much appreciated and successfully distributed, future literature donations will be auctioned as fund raising efforts, (Motion by Don Cadien, second by Susan Hamilton and John Shisko, passed by 2/3 majority.)

Literature Committee

Leslie Harris has joined the Literature Committee to fill out the member­ ship of three along with John Dorsey and Don Cadien.

Logo

Decision on logo approval was postponed a month in order to give more members a chance to submit art work. Two examples are given below. Every­ one is welcome to participate. If you cannot attend the meeting just send a copy to Ann Martin.

Southern California Association of

Marine Invertebrate Taxonomists CALIFORNIA

INVERTEBRATE Helpful Hints

A reference that should be helpful to people:

Mikkelsen, Paul S. and Robert W. Virnsteln. 1982. An illustrated glossary of polychaete terms. Harbor Branch Foundation, Inc., Technical Report No. 46, R.R. 1, Box 196, Fort Pierce, Florida.

Membership

Membership is going very well. There are 35 members as of July 28. If you have not sent in your application yet, please do so. It is impor­ tant because only members will receive this Newsletter and other notices of SCAMIT activities.

APPLICATION FOR MEMBERSHIP TO SOUTHERN CALIFORNIA ASSOCIATION OF MARINE INVERTEBRATE TAXONOMISTS

June 1982 - May I983 membership fee is $5.00. Make checks payable to SCAMIT and mai1 to Ann Martin at 10844 Ellis Avenue, Fountain Val ley, CA. 92708.

Type of membership is:

CHARTER ( ) PARTICIPATING ( ) CORRESPONDENT ( ) VOUCHER SHEET

IDENTIFIED AS: Arctonoe vi ttata (Grube, 1855)

SPECIMEN CODE: AHF 3

KEYS USED: Hartroan, 0. 1968 (Atlas) - p. ^9 Fauchald, K. 1977 - p. 59

OTHER TEXTS CONSULTED: Banse, K. : K. Hobson. 197^ - p. 26 Pettibone, M.H. 1953 - p. 57 Skogsberg, T. 19^2 - p. ^89 Ushakov, P.V. 1965 - p. 113

IMPORTANT CHARACTERS: Elytra greater than 21 pairs, leaving middorsum bare; two types of neurosetae; superior setae blunt with bifid or notched tips, interior setae falcate; segments 7-8 with dark brown dorsal band.

RELATED AND Arctonoe f ragi 1 i s and A_. pulchra both have CHARACTER DIFFERENCES: only one type of neurosetae, falcate; A. fragi1 is has elytra with frilled posterior margins; elytra of A_. pulchra and A. vi ttata s imilar; slightly wavy at margins; no brownish band across dorsum on A_. fragi1 is and A. pulchra.

VARIABILITY: Degree of pigmentation can vary from dark brown to nearly matching dorsum color.

COMMON SYNONYMS: Polynoe vi ttata Grube, T855

AIDS TO IDENTIFICATION: Dorsal pigment band can usually be seen without a scope; if band is faint, prepare parapodium to check neurosetae.

STATION DATA: AHF N7420 VOUCHER SHEET

IDENTIFIED AS: Ha_l osy dna lain or Chamber 1 in, 1919 .

SPECIMEN CODE: AHF k

KEYS USED: Hartman, 0. 1968 (Atlas)-p. 67 Fauchald, K. 1977 - p. 59.

OTHER TEXTS CONSULTED: Chamberlin, R.V. 1919. - P- 1

IMPORTANT CHARACTERS: Elytra with heavy lateral fringe; simple neurosetae, body depressed; ]8 pairs eyltra.

RELATED SPECIES AND Halosydna brevi setosa wi th vivid CHARACTER DIFFERENCES: pigmentation on anterior segment; elytra with sparse marginal fringe; noticeable distal swelling on dorsal cirri; H. johnsoni has bifid neurosetae.

STATION DATA: AHF N3761

COMMENTS: There are problems with the definition of "heavy" or "slight" elytral fringe, in distinguishing a pale H_. brevi setosa from a M.' 1at_ior; and in the identification of juvenile Halosydna. VOUCHER SHEET

IDENTIFIED AS: Lagisca pedroens i s Hartman, I960

SPECIMEN CODE: SCCWRP 1

KEYS USED: Hartman, 0. 1968 (Atlas) - p. 107 Fauchald, K. 1977 - p. 57

OTHER TEXTS CONSULTED: Hartman, 0.; J.L. Barnard. I960 - 80

IMPORTANT CHARACTERS: Ventral prostomial antennae; 15 pairs elytra; posterior setigers not covered by elytra; elytra with marginal fringe; notosetae coarser than neurosetae; neurosetae of two types; superior ones bifid with widely spaced teeth and the crotch with rugosity; inferior ones di s ta11y enti re.

RELATED SPECIES AND L. lame 11 i fer and L. yokohami ens i s both CHARACTER DIFFERENCES: have two types of neurosetae, however L. pedroensis has a widely spaced distal tooth with rugosity in the crotch and both L_. 1 ame 11 i fer and L_. yokohami ensi s have a narrow gap between the distal tooth and shaft. The elytra of L_. lamel 1 ifer have drop-like papillae on the posterior half; L. pedroensis does not. L. yokohamiensi s has elytra uniform!ly covered with horny papillae; L. pedroensi 5 has elytra which appear smooth, with uneven distribution of two types of papillae.

VARIABILITY: The range in the number of segments can vary in specimens. Large specimens with greater than 50 segments may key to Polynoe. Juvenile specimens do not show the same number of exposed posterior setigers as larger specimens; this could cause some confusion with specimens of Harmothoe.

STATION DATA: SCCWRP 5-8. Orange County Deep Survey 118° 71 2" W. 33o 27' I" N. 627m trawl 26 January 1982. Soft bottom.

COMMENTS: The species key in Hartman, )968 implies that pedroens is is the only Lagisca wlth two types of neurosetae, however, the description for L. lamel1i fer in Hartman (5968, p. 103) states "neurosetae longest, with transverse rows of spines (fig. 5); others increasingly shorter, distally bifid (fig. 6) or entire". The description for L. yokohamens is in Hartman (1968, p. 109) states "neurosetae with a prominent accessory tooth (fig. 2} and 20 to 30 pairs of pectinated plates subdi s ta 1 ly; i nferior most neurosetae lack the accessory tooth". Izuka (1912) also describes two types of neurosetae for Harmothoe yokohaminensis. VOUCHER SHEET

IDENTIFIED AS: Pholoides aspera (Johnson, 1897)

SPECIMEN CODE: SCCWRP 2

KEYS USED: Hartman, 0. 1968 (Atlas) - p. 1^7 Fauchald, K. 1977 - p. 66

Banse, K.; K.D. Hobson. 1974 - p. 33 OTHER TEXTS CONSULTED: Hartman, 0.; K. Fauchald. 1971. - p. 29 Johnson, H.P. 1897. - p. 184

Elytra on alternate segments, each with IMPORTANT CHARACTERS: fringe, concentric rings often with central dark spot; single median antenna, filiform and fimbriated at tip; dorsal cirri fimbriated at tip, present only on first setiger; notosetae simpie; neurosetae composite dentate falcigers.

RELATED SPECIES AND P_. tuberculata (Hartmann-Schroder, 1965) CHARACTER DIFFERENCES: Apparently identical but not synonymized with P_. aspera. P_. bermudensi s Hartman £ Fauchald, 1971; 30-32 segments instead of 35_38; neurosetae falciger smooth,instead of dentate. Phlos aspera has geniculate superior notosetae, prostomium with single median antenna; the median antenna is not fimbriated and the elytra have no concentric rings and they are pale white with sparse marginal papi1 la.

COMMON SYNONYMS: Peis1di ce aspera Johnson, 1897•

AIDS TO IDENTIFICATION: Fimbriated median antenna, concentric rings on elytra.

SCCWRP 8.3-60 Santa Monica Bay STATION DATA: 118° 29' 5" W. 33° 52' 2" N. 9 May 1979 1.0mm screen - soft-bottom

Fauchald (1977) states "fringed elytra COMMENTS: alternate with dorsal cirri in all setigers"; Johnson (1397) specifically states "no dorsal cirri", only one median antenna and one pa i r peristomial cirri; Hartmann and Fauchald (197') state "dorsal cirri and branchia are absent", "parapodia of first segment are directed forward, at sides of prostomium; each has a pair of long dorsal cirri resembling the median prostomial antenna". VOUCHER SHEET

IDENTIFIED AS: Stenelais tertiaglabra Moore,1910

SPECIMEN CODE: OCSD 2

KEYS USED: Hartman. 0. 1968 (Atlas) p, 165 Fauchald, K. 1977 - p. 70

OTHER TEXTS CONSULTED: Moore, J.P. 1910. - p. 395 Pettibone, M.H. 1971a. 1971b Banse, K. : K.D. Hobson. 1974 - P- 35 Hartman, 0. 1939 - p. 65 (S. hancocki)

IMPORTANT CHARACTERS: Ventral surface smooth; triangular elytra! imcrotubercles encrusted with extracuticular rusty material; neurosetae with long, slender, minutely bidentate falcigers with few stout bifid falcigers in median parapodia; body pale except for anterior portion of elytra.

RELATED SPECIES AND S. berkeleyi ; ventral surface densely CHARACTER DIFFERENCES; papillated, elytra pale to rusty brown; S_. verruculosa; neurosetae with multi- articulated simple setae, elytra with small wart-like tubercules on dorsum, elytra with fingerlike patpodes on margin; S_. fusca; elytra survace with conical microtubercles, mottled with dark pigment.

STATION DATA: 60m, sitty-sand k February I98O 330 34' I&" n8° 01 ' 30" VOUCHER SHEET

IDENTIFIED AS: Stenelanella uni formi 5 Moo re, 1910

SPECIMEN CODE: OCSD 1

KEYS USED: Hartman, 0. 1968 (Atlas) p. 169 Fauchald, K. 1977- p. 69

Moore, J.P. 1910 - p, 391 OTHER TEXTS CONSULTED: Pettibone, M.H. 1969 - p. 431 Hartman, 0. 1939 - p. 69

Approximately the first 10 elytra with IMPORTANT CHARACTERS: mottled rusty-brown pigment, the rest colorless; stout, short-appendaged, compound neurosetae; notopodiai spi nni ng glands emergent with silky thread; elytra lack fringe, except first pair; first pair elytra orbicular, with the rest; ctenidia between setigers 2:3, with additional clenidia ventrally on setiger 3.

RELATED SPECIES AND S_. ehlersi (Horst, 1916) Pettibone, I969, CHARACTER DIFFERENCES: anterior elytra banded, not mottled; middle and posterior elytra with deeply sinuous external margins; ctenidia on dorsal bases of tentacular parapodia of setiger I, none between setigers 2-3- (Dutch East Indies, South Africa.)

AIDS TO IDENTIFICATION; Only sigalionid in California with spinning glands.

STATION DATA: 60m, silty-sand 13 July 1978 33° 34' k€>" 118° 01' 30" tdmZ*

Some Taxonomic Literature on the from the Eastern Pacific Region: Families Amphinomidae, , Euphro- sinidae, Hesionidae, Pilargiidae, and Genus Typosyllis (All literature containing the original descriptions of genera and species recorded from Southern California has been included.)

Banse, K. 1963. Polychaetous from Puget Sound and the San Juan Archipelago, Washington. Proc. Biol. Soc. Wash. 76: 197-208. Includes description of the new subspecies Eteonides coineaui difficilis.

Banse, K. 19 72. On some species of Phyllodocidae, Syllidae, Nephtyidae, Goniadidae, Apistobranchidae, and Spionidae from the northeast Pacific Ocean. Pac. Sci. 26: 191-222. Eteone pacifica Hartman and E. tuberculata Treadwell are re- described; additions to descriptions are given for (Eulalia) quadrioculata Moore, Syllis (Typosyllis) fasciata Malmgren, S. (Ty_pos^y_ll1i5^) pulchra Berkeley & Berkeley, S_. (Typosyllis) adamantea adamantea (Treadwell), and S. (Typosyllis) stewarti Berkeley & Berkeley.

Banse, K. 1973. The ventral parapodial cirrus of the benthic Phyllodocidae (Polychaeta), with special reference to Clavadoce Hartman and Bergstroemia Banse. J. Nat. Hist. 7: 683-389. Discussion of large reniform ventral cirri as generic chara­ cters; new diagnosis forClavadoce; Bergstroemia elevated to rank of genus. Banse, K. and K.D. Hobson. 1968. Benthic polychaetes from Puget Sound, Washington, with remarks on four other species. Proc. U.S. Natn. Mus. 125: 1-53. Accounts of Eulalia (Hypoeulalia) bilineata (Johnston)?, E. (Eulalia) levicornuta Moore, E. (Ptercirrus) parvoseta, n.sp. , (Anaitides) nr. multiseriata Rioja, P. (Anaitides) williamsi (Hartman), Gyptis brevipalpa (Hartmann-Schroder) , Micropodarke dubia (Hessle) , Sigambra tentaculata (Treadwell) , and Syllis (Typosyllis) harti Berkeley & Berkeley.

Bergstrom, E. 1914. Zur Systematik des Polychaeten Farnilie per Phyllodociden. Zoologiska Bidrag fran Uppsala, .3: 37-224, 81 figs. Monograph on the family, including a extensive discussion of the genera, descriptions of the new genera Austrophy11urn and Steggoa. Berkeley, E. and Berkeley, C. 1938. Notes on polychaeta from the coast of western Canada. Ann. Mag. Nat. Hist. ser. 11, Vol. 1: 33-49, 12 figs.

Syllis pulchra and Syllis spenceri (now synonymized w/Typosy His adamantea Treadwell, 1914 ) are described. - Berkeley, E. and Berkeley, C. 1941. On a collection of polychaeta from southern California. Bull. So. Calif. Acad. Sci., 40:16-60, 1 pi. Refers to 4 amphinomids & euphrosinids, 2 phyllodocids, 3 species of Typosyllis, and 1 hesionid, plus the description of Loandalia fauveli. Berkeley, E. and Berkeley, C- 1948. Canadian Pacific Fauna. 9. Annelida. 9b(l) Polychaeta . Uni. Toronto Press, Toronto, Canada, 100 pp. Discussions of 3 species of Euphrosine, 12 phyllodocids, 2 pilargrids, and 6 species of Typosyllis also found in southern California. Blake, J.A. and C.P. Walton. 1977. New species and records of polychaeta from the Gulf of the Farallones, California, pp. 307-321. I_n D.J. Reish and K. Fauchald (Eds), Essays on Poly- chaetous Annelids in Memory of Dr. Olga Hartman. Allan Hancock Foundation, University of Southern California, Los Angeles, Ca. Phyllodoce hartmanae and Typosyllis farallonensis are newly described. Chamberlin, R.V. 1919. The Annelida Polychaeta. Mem. Mus. Cornp. Zool., Harvard, 48:1-514.

Report on polychaetes from Pacific Mexico south to the Galapagos Islands. Includes type description of Chloeia entypa, an account of Eurytho'e complanata (Pallas) , and the description of the genus Synelmis. Chamberlin, R.V. 1919. New polychaetous annelids from Laguna Beach, California. J. Entomol. Zool. Pomona Coll., 11:1-23. Includes description of Hesperophyllum tectum, n.g., n.sp. , Steggoa gracilor n.sp. , Sige_ californiensis n.sp., Anaitides heterocirrus n.sp., Typosyllis be11a n.sp., Pionosyllis pigmentata n.sp. (now Typosyllis), Pionosyllis lucida n.sp. (now Typosyllis) , Hesperalia californiensis n.sp. (?Odontosyllis), Hesperalia nans n.sp. (?Odontosyllis), and Campesyllis minor n.g., n.sp.

Claparede, E. 1868. Les Annelides Chetopodes du Golfe de Naples. Mem. Soc. Phys. Geneve, 19(2): 313-584, 16 pis. Includes the new genus Pterocirrus.

Czerniavsky, V. 1882. Materialia ad zoographiam Ponticam comparatam. Bull. Soc. Imp. Nat. Moscow, 57:146-198. Description of the genus Anaitides.

Ditlevsen, H. 1917. Annelids. I. Danish Ingolf-Expedition, 4(4):1-71, 6 pis. 24 figs.

Report on polychaetes from Greenland, Iceland and the Faroe Islands, Includes discussion of Phyllodoce groenlandica Oersted and Eteone longa (Fabricius). — ~ Dorsey, J.H. 1978. A first report of Heteropodarke heteromorpha Hartman-Schroder, 1962 (Polychaeta: Hesionidae) from California. Bull. So. Calif. Acad. Sci. 77:82-84. The genus Heteropodarke is amended plus additions to the description of H. heteromorpha.

Ebbs, N.K. Jr. 1966. The coral-inhabiting polychaetes of the northern Florida Reef Tract. Part I. Aphroditidae, Polynoidae, Amphinomidae, Eunicidae, and Lysaretidae. Bull. Mar. Sci. 16:485-555. An extensive account of Eurythoe complanata is given.

Ehlers, E. 1864. Die Borstenwurmer, nach systematischen und anatomischen Untersuchungen dargestellt. Leipzig, pp. 1-268. Includes description of Orseis, new genus.

Emerson, R.R. and K. Fauchald. 1971. A revision of the genus Loandalia Monro with description of a new genus and species of pilgariid polychaete. Bull. So. Calif. Acad. Sci., 70{l}:18-22.

Maintains generic status of Ancistargis Jones, describes Parandalia ocularis n.g., n.sp., and transfer Loandalia fauveli to Parandalia. Essenberg, C. 1917. New species of Amphinomidae from the Pacific Coast. Univ. Calif. Publ. Zool. 18:61-74.

Presents general description of the family and new species: Euphrosyne calypta, E. multibranchiata, E. kyllosetosa, and Eurythoe spirocirrata.

Fabricius, 0. 1780. Fauna Groenlandica, systematice sistens, Animalia Groenlandica occidentalis hactenus indagata, quod nomen specificum, triviale, vernaculumque; synonyma auctorum plurium, descriptionem, locum, vieturn, generationem, mores, usum, capturamque singuli , pront detegendi occasio fuit, maximaque parti secundum proprias observationes, Hafniae, XVI and 452 pp.

Descriptions of Nereis flava and Nereis longa (both now Eteone)

Fauchald, K. 1972. Benthic polychaetous annelids from deep water off western Mexico and adjacent areas in the eastern Pacific Ocean. Allan Hancock Monogr. Mar. Biol. 7:575 pp., 69 pis. Geographic monograph with accounts of 1 new amphinomid, 1 euphrosinid, 5 new and 1 old phyllodocids, including AustrophyHum exsilium, and 4 new pilargiids, including Sigambra setosa.

Fauchald, K. 197 7. The Polychaete Worms. Definitions and Keys to the orders, families and genera. Nat. Hist. Mus. Los Angeles Co., Sci. Ser. 28:1-190.

Fauvel, P. 1923. Polychaetes errantes. Faune de France, 5:1-488. Fauvel, P. 1932. Annelida polychaeta of the Indian Museum, Calcutta. Mem. Indian Mus.Calcutta, 12(l):l-262, 9 pis., 40 figs.

Description of Pseudeurythoe new genus Fauvel, P. 1934. Sur quelques Syllidiens du Japon. Annot. Zool. Japon, 14 (3):301-316, 2 figs. Grube, A.-E. 1860. Beschreibung neur Oder wenig bekannter Anneliclen. Beitrag: Zahlreiche Gattungen. Arch. Naturg. Berlin, 26:71-118, 3 pis. Description of Syllis variegata (now Typosyllis)

Grube, A.-E. 186 3 . Beschreibung neur oder"weinig bekannter Anneliden . Beitrag: Zahlreiche Gattungen. Arch. Naturg. Berlin, 29:37-69, 3 pis. Description of Syllis hyalina (now Typosyllis) Gustafson, G. 19 30. Anatomische Studien uber die Polychaeten Fami lien Amphinomidae und Euphrosynidae. Zool. Bidrag Uppsala, 12:305 —471, pis. 1-36, 64 figs. Monograph, primarily on anatomy, also has generic description Of Pareurythoe.

Hartman, 0. 19 36. Nomenclatorial changes involving California poly- chaete worms. J. Wash. Acad. Sci., 26(l):31-32.

Eteone pacifica is proposed for^E. maculata Treadwell, preoccupied.

Hartman, 0. 1936. A review of the phyllodocidae {Annelida Polychaeta) of the coast of California, with descritpions of nine new species. Univ. Calif. Publ. Zool. 41:117-132.

A key to 29 species from California is given. New species include Anaitides williamsi, Clavadoce splendida (also new genus), Eteone dilatae, E. lighti, E. californica, E. balboensis, Eulalia avisuliseta, Sige montereyensis, and Steggoa californiensis.

Hartman, 0. 1939. New species of polychaetous annelids from southern California. Allan Hancock Pac. Exped. 7 (2}:157-172, 2 pis.

Type description of Hesionella mccullochae, new genus and species.

Hartman, 0. 1940. Polychaetous annelids. Part II. Chrysopetalidae to Goniadidae. Allan Hancock Pac. Exped. 7:173-286.

Report on polychaetes from tropical and subtropical eastern Pacific Ocean; includes families Amphinomidae (7 species), Euphrosynidae (3 species)f and Hesionidae {3 species). Hartman, 0. 1945. The marine annelids of North Carolina. Bull. Duke Univ. Mar. Sta., no. 2:1-51, 10 pis. Discription of Sigambra bassi. Hartman, 0. 1947. Polychaetous annelids. Part VIII. Pilargiidae. Allan Hancock Pac. Exped. 10:483-522. Monograph on the family Pilargiidae; keys to genera and species; Pilargis maculata, Ancistrosyllis bassi, and Loandalia americana are newly described.

Hartman, 0. 1948. The Polycahetous Annelids of Alaska. Pac. Sci., 2(1):3-58. Report includes information on 5 phyllodocids and 3 species of Typosyllis found in southern California.

Hartman, 0. 1960. Systematic account of some marine invertebrate from the deep basins off southern California. Allan Hancock Pac. Exped., 22:69-214, 19 pis. Describes Pilargis hamatus (sic), (now Ancistargis) and Euphrosine paucibranchiata, plus new reocrds for Pseudeurytho'e ambigua (Monro). Hartman, 0. 1961. Polychaetous annelids from California. Allan Hancock Pac. Exped., 25:226 p., 34 pis.

Type descriptions of Orseis lagunae, Amphiduros pacificus , Oxydromus arenicolus n. ssp. glabrus (now Gyptis brevipalpa (Hartman-Schroder)) and Oxydromus brunnea (now Gyptis brunnea) , plus accounts of Ophiodromus pugettensis (Johnson) Eumida bifoliata (Moore) new combination, and new records for Eulalia viridis (Linnaeus) and Anaitides multiseriata Rioja.

Hartman, O. 1963. Submarine canyons of southern California, 3: Systematics: Polychaetes. Allan Hancock Pac. Exped., 27:1-93, 4 figs. Includes description of Ancistrosyllis breviceps.

Hartmann-Schroder, G. 1958. Einige Polychaeten aus dem Kusten- grundwasser der Bimini-Inseln (Bahamas). Kieler Meeresforsch, 14:233-240.

Includes generic description of Hesionura.

Hartmann-Schroder, G. 1959. Zur okologie der polychaeten des mangrove- estero-gebietes von El Salvador. Beitrage zur Neotropischen Fauna, 1(2);69-183.

Descriptions of Oxydromus brevipalpa (now Gyptis brevipalpa) and Loandalia gracilis (possibly a synonym of Parandalia fauveli (Berkeley and Berkeley). Hartmann-Schroder, G. 1962. Zweiter Beitrag zur Polychaetenfauna von Peru. Kieler Meeresforschungen, 18 (1) :109-147. Type description of Heteropodarke heteromorpha, new genus and species.

Hartmann-Schroder, G. 1965. Zur kenntnis der eulitoralen poly­ chaetenf auna von Hawaii, Palmyra und Samoa. Naturwiss. Vereins Hamburg, Abh. Verh., N.F.9:81-161, 85 figs.

Has brief account of Typosyllis hyalina (Grube)

Hessle, C. 1925, Einiges uber die Hesioniden und die Stellung der Gattung Ancistrosyllis. Ark. Zool., 17:1-37. Describes Kefersteinia dubia (now Micropodarke dubia).

Imajima, M. 1966. The Syllidae (Polychaetous Annelids) of Japan. (V) Syllinae (2) . Publ. Seto Mar. Biol. Lab. 14:253-294.

Descriptions are given for 15 species of Typosyllis of which X* hyalina, T. alternata, and T. varigata are common to both Japan and North America/

Imaj ima, M. and 0. Hartman. 19 64. The polychaetous annelids of Japan. Part 1. Allan Hancock Fdn. Pub., Occas. Pap. 26:1-237, 35 pis. Transfers kefersteinia dubia Hessle into Micropodarke, dis­ cusses Ophiodromus pugettensis (Johnson), Typosyllis adamantea (Treadwell), Ty^osy_ll_is fasciata (Malmgren) , Typosyllis variegata (Grube), and describes Typosyllis aciculata orientalis n. ssp. Information on Eurythoe complanata (Pallas), Anaitides madeirensis (Langerhans), Anaitides groenlandica (Oersted), Eteone longa (Fabricius), Eulalia bilineata (Johnston) E. viridis (Linnaeus), Eumida sanguinea (Oersted) and~Genetyllis castanea (Marenzeller) .

Izuka, A. 1912. The errantiate polychaeta of Japan. J. Coll. Sci. Tokyo, 30(2):l-262, 24 pis.

Discussions of Phyllodoce groenlandica Oersted, Carobia castanea Marenzeller (now Genetyllis castanea), Eumida sanguinea (Oersted) and Eulalia viridis (Miiller) .

Johnson, H.P. 1897. A preliminary account of the marine annelids of the Pacific coast, with descriptions of new species. Euphrosynidae, Amphinomidae, Palrnyridae, Polynoidae and Sigalionidae. Proc. Calif. Acad. Sci., Zool., 1:153-190, pis. 5-10.

Account on mostly California material, including new species Euphrosyne (sic) aurantiaca, E. arctia and Eurythoe californica {now Pareurythoe). Johnson, H.P. 1901. The Polychaeta of the Puget Sound Region. Proc. Boston Soc. Nat. Hist., 29:381-437, 19 pis.

Has description of Podarke pugettensis, new species. Johnston, G. 1840. Miscellanea Zoologica. The British Nereides. Ann. Mag. Nat. Hist. London, ser. 1, vol. 4:224-232, pis. 6-7 . Includes description of Phyllodoce bilineata (now Eulalia) Jones, M.L. 1961. Two new polychaetes of the families Pilargiidae and Capitellidae from the Gulf of Mexico. Amer. Mus. Nov., 2049:1-18. Discription of Ancistargis, new genus. Kinberg, J.G.H. 1857 . Nya slagten och arter af Annelider. Forh. Oefv. Vet. Akad. Stockholm, 14:11-14. Description of new genus Eurythoe.

Kinberg, J.G.H. 1866. Annulata nova. Forh. Oefv. Vet, Akad. Stockholm, 22:239-258.

Description of Phyllodoce longipes.

Kravitz, M.J. and H.R. Jones. 19 79. a Systernatics and ecology of benthic Phyllodocidae (:Polychaeta) off the Columbia River, U.S.A. Bull. So. Calif. Acad. Sci. 78:1-19.

Seven species of Eteone, three of Anaitides, and a single each of Paranaitides, Eulalia, and Eumida are reported. New species include Eteone fauchaldi, E. spilotus, E. columbiensis, and Anaitides multipapillata.

LaGreca, M. 1946. Studii sui Policheti del Golfo di Napoli. Pubbl. Staz. Zool. Napoli, 20:270-280, figs. 1-11. Has the description of Oxydromus arenicolus, the stem species for Gyptis arenicola glabra, and the original drawings Hartman modified for the Atlas (1968).

Langerhans, P. 1879. Die Wurmfauna von Madeira. Pt. 1. Zeits. wiss. Zool., Leipzig, 32:513-592, pis. 31-33.

Description of the new genus Typosyllis.

Langerhans, P. 1880. Die wurmfauna von Maderia. Pt. 2. Zeitschrift fur Wissenschaftliche Zoologie, 33:267-316, pis. 14-18. Includes description of Phyllodoce madeirensis.

Langerhans, P. 1881. Uber einige canarische Anneliden. Nova Acta Ksl. Leopoldino-Carolina Deutschen Akad. Naturf., 42:93-124, 2 pis. Describes Ancistrosyllis albini (now Synelmis) Linnaeus, C 1766-68. Systema naturae. 12th edition.

Includes description of Nereis viridis (now Eulalia)

Malmgren, A.J. 1865. Nordiska Hafs-Annulater. Forh. Oefv. K. Vetensk. Akad. Stockholm, 21:51-110, pis. 8-15 Includes generic descriptions of Eumidia, Genetyllis, Sige, and Anaitis (now Paranaitis)

Malmgren, A.J. 1867. Annulata Polychaeta Spetsbergiae, Groenlandiae, Islandiae et Scandinaviae hactenus cognita. Oefv. Svenska Vetensk. Akad. Forh., 24:127-235, pis. 2-15.

Description of Syllis fasciata (now Typosyllis) Marenzeller, E. 1879. Sudjapanische Anneliden. I. Akad. Wiss. Wien, Denkschr., 41(2): 109-152, pis. 1-6.

Description of Carobia castanea (now Genetyllis)

Marion, A.F. and N. Bobretsky. 1875. Etude des Anne1ides du golfe de Marseille. Ann. Sci. Nat. Paris, ser. 6, vol. 2:1-106, 12 pis.

The genus Gyptis is newly described.

McCammon, J.A. and D.E. Montagne. 1979. Some species of the genus Phyllodoce {Polychaeta} from southern California. Zool. J. Linn. Soc. 66:353-368. A key to species of Phyllodoce from California is given; Phyllodoce (Anaitides) cuspidata and the subgenus Aponaitides are newly described. Mcintosh, W.C. 1879. On the Annelida obtained during the cruise of H.M.S. Valorous to Davis Strait in 1875. Trans. Linn. Soc. London, 1:499-511.

Includes descriptions of AncistrosyHis groenlandica, new genus and species.

Metschnikow, E. 1865. Beitrage zur kenntniss der Chaetopoden. Zeits. wiss. Zool. Leipzig, 15:328-341, pis. 24-25.

Includes description of Microphthalmus, new genus. Monro, CCA. 1933. On a new species of polychaeta of the genus Pilargis from Friday Harbour, Washington. Ann. Mag. Nat. Hist., ser. 10, vol. 11:673-675, 4 figs.

Description of Pilargis berkeleyae.

Monro, CCA. 1933. The Polychaeta Errantia collected by Dr. Crossland at Colon in the Panama region and the Galapagos Islands during the expedition of the S.Y. St. George. Proc. Zool. Soc. London, pt. 1, 1-96, figs. 1-36. Moore, J.P. 1905. New species of polychaeta from the North Pacific, chiefly from Alaskan waters. Proc. Acad. Nat. Sci. Phil., 57:525-554. Descriptions of Euphrosyne (sic) bicirrata and E. hortensis.

Moore, J.P. 1906. Additional new species of polychaeta from the North Pacific. Proc. Adad. Nat. Sci., Phil., 58:217-260, pis. 10 Report on polychaetes from Alaska, with descriptions of the new species Notophyllum imbricatum, Eulalia quadrioculata and Eulalia longicornuta {now Euroida). Moore, J.P. 1908. Some polychaetous annelids of the northern Pacific coast of North America. Proc. Acad. Nat. Sci. Phil., 60:321-364. Includes description of Syllis alternata (now Typosyllis), information on Syllis armillaris (Muller), 5 phyllodocids, 3 euphrosinids and 1 hesionid.

Moore, J.P. 1909. Polychaetous annelids from Monterey Bay and San Diego, California. Proc. Acad. Nat. Sci. Phil., 61:235-295. pis. 7-9.

Type description of Phyllodoce medipapillata.

Moore, J.P. 1909. The polychaetous annelids dredged by the U.S.S. "Albatross" off the coast of southern California in 1904. I. Syllidae, Sphaerodoridae, Hesionidae and Phyllodocidae. Proc. Acad. Nat. Sci. Phil., 61:321-351, pis. 15-16.

Information on Syllis alternata Moore, Podarke pugettensis Johnson, Phyllodoce mucosa Oersted, P. medipapillata Moore, and P. (Carobia) castanea, Marenzeller. Newly described are Phyllodoce ferruginea, Anaitis polynoides (now Paranaitis), Eumida tubiformis, Eulalia nigrimaculata (now Genetyllis), Eulalia levicornuta and Eulalia (Sige)bi^foliata.

Moore, P.J. 1911. The polychaetous annelids dredged by the U.S.S. "Albatross" off the coast of southern California in 1904. III. Euphrosynidae to Goniadidae. Proc. Acad. Nat. Sci. Phila. 63: 234-318.

Euphrosyne dumosa, E. limbata, and Chloeia pinnata are newly described.

Muller, F. 1858. Einiges uber die Annelidenfauna der Insel Santa Catharina an der brasilianischen kuste. Arch. Naturg. 24; 211-220.

Includes description of Sigambra, new genus. Muller, O.F. 1771. Von Wurmern des siissen und salzigen Wassers. Heinich Mumme und Faber, Copenhagen, 200 pp., 16 pis.

Description of Syllis armillaris {now Typosyllis^) Oersted, A.S. 1843. Gr^nlands Annulata Dorsibranchiata. Afh. K. Danske Vidensk. Selsk. Nat. Mat. K. , 10:15.3-216.

Information on Typosyllis armillaris (Muller) and description of Phyllodoce groenlandica.

Oersted, A.S. 1843. Annulatorum danicorum conspectus, Fasc. 1. Maricolae. Copenhagen, pp. 1-52, 7 pis.

Descriptions of the new genus Notophyllum, and new species Phyllodoce mucosa and Eulalia sanguinea (now Eumida). Okuda, S. 193 8. Polychaetous annelids from the vicinity of the Mitsui Institute of Marine Biology. Japan. J. Zool., 8:75-105, 15 figs. Description of Micropodarke, new genus.

Pallas, P.S. 1766. Miscellanea Zoologica. Hagae Comitum, xii + 224 pp. Description of Aphrodita complanata (now Eurythoe).

Pettibone, M.H. 19 54. Marine polychaeta worms from Point Barrow, Alaska, with additional records from the North Atlantic and North Pacific. Proc. U.S. Natl. Mus., 103:203-356, figs. 26-39.

Information on Eteone longa (Fabricius), Phyllodoce (Anaitides) groenlandica Oersted and Syllis (Typosyllis) fasciata (Malmgren) .

Pettibone, M.H. 1966. Revision of the Pilargiidae (Annelida: Polychaeta), including descriptions of new species, and redescription of the pelagic Podarmus ploa Chamberlin (Polynoidae). Proc. U.S. Natn. Mus. 118:155-208.

Review of the family, including keys to genera and species.

Reish, D.J. 1950. Redescription of the polychaetous annelid Typosyllis aciculata Treadwell. Amer. Mus. Novitates, no. 1466:1-5.

Redescription of T. aciculata and comparisons with other typosyllids with similar characteristics.

Rioja, E. 1941. Estudios anelidologicos. III. Datos para el concimiento de la fauna de poliquetos de las costas del Pacifico de Mexico. Anal. Inst. Biol. 12:669-746.

Review of taxonomic polychaete studies from the Pacific coast of Mexico; account of polychaetes collected around Acapulco. Phyllodoce multiseriata is newly described. Rowe, R. 1980. Polychaeta. pp. 78-177. In D.Straughan & R.W. Klink (compilers) ,A Taxonomic Listing of Common Marine Invertebrate Species from Southern California. Allan Hancock Foundation, Technical Rept. 3:281 pp. Saint-Joseph, A. 1899. Annelides polychetes de la rade de Brest et de Paimpol. Ann. Sci. Nat. Paris, ser. 8, vol. 10:161-194.

Description of Pilargis, new genus. Sars, M. 1862. Beretning om en i Sommeren 1859 foretagen zoologisk Reise ved Kysten af Romsdals Amt. Nyt. Mag. Naturv. Oslo, 11(3):241-263.

Includes description of Ophiodromus, new genus.

Savigny, J.C. 1818. Annelids. In J.B. deLamarck, Histoire Naturelie des Animaux sans Vertebres, v + 612 pp.

Type descriptions of the new genera Chloeia, Euphrosine, Eteone, Eulalia and Phyllodoce.

Southern, R. 1914. Archiannelida and Polychaeta. Proc. Roy. Irish Acad., Dublin, 31 (pt. 47): 1-160, 15 pis.

Proposal of Paranitis to replace Anaitis Malmgren, preoccupied.

Treadwell, A.L. 1914. New Syllidae from San Francisco Bay, collected by the U.S.S. "Albatross". Univ. Calif. Pub. Zool., 13(9): 235-238, 7 figs. Describes Trypanosyllis adamanteus (now Typosyllis). Treadwell, A.L. 1941. Polychaetous annelids from the New England region, Porto Rico and Brazil. Amer. Mus. Novit. no. 1138:1-4, 12 figs. Describes Ancistrosyllis tentaculata (now Sigambra).

Treadwell, A.L. 1945. New species of polychaetous annelids from Massachusetts and California. Amer. Mus. Novit., no. 1280: 3 pp. Description of Typosyllis aciculata.

Uschakov, P. and Wu, B.L. 1959. The polychaetous annelids of the families Phyllodocidae and Aphroditidac from the Yellow Sea. Archives Institute Oceanology Sinica, l(4):l-40.

Type description of Phyllodoce papillosa. Ushakov, P. 1972. Polychaeta 1. Polychaetes of the suborder Phyllodociformia of the polar basin and the northwestern part of the Pacific, (in Russian) Akad. Nauk. SSSR. Zool. Inst.; Fauna of the SSSR,n.s. 102, 271 pp. (translated 1974 by the Israel Program for Scientific Translation, Jerusalem).

Monograph of the family; includes morphological and ecological aspects of the suborder; description of Pterocirrus imajimai n.sp.

Webster, H.E. 1879. The Annelida Chaetopoda of New Jersey, Ann. Rep. N.Y. State Mus. Nat. Hist., 32:101-128.

Description of Eteone alba