STATE AND FEDERALLY LISTED ENDANGERED AND THREATENED OF

July 2021

State of California Natural Resources Agency Department of and Biogeographic Data Branch California Natural Diversity Database (CNDDB)

Table of Contents

Overview ...... i Background ...... i Listing Status Summary Table ...... ii List Content ...... iii Abbreviations ...... iv Additional Resources...... iv

Overview

This document contains a list of taxa found within California or off the coast of the State that have been classified as endangered or threatened by the California Fish and Commission (FGC; state listed) or by the U.S. Secretary of the Interior or the U.S. Secretary of Commerce (federally listed). This list also includes taxa that are official candidates for state or federal listing, or have been officially proposed for federal listing, as well as taxa that were once listed but have since been delisted.

Background

State listing is pursuant to the California Endangered Act of 1984 (CESA; California Code of Regulations, Title 14, Chapter 6, §§783.0-787.9; Fish and Game Code Chapter 1.5, §§ 2050-2115.5). The designations “Endangered” and “Rare” were first established in 1970 by the original California Act, and taxa with a state list date of June 27, 1971 were protected under this regulation. In 1984, CESA was amended, at which time the “Rare” designation was changed to “Threatened,” and on January 1, 1985, all animal species previously designated as “Rare” were reclassified as “Threatened.” The official California listing of Endangered and Threatened animals is contained in the California Code of Regulations, Title 14, §670.5.

Federal listing is pursuant to the Federal Endangered Species Act of 1973, as amended (16 USC §§1531-1544; 50 CFR §§17.1-17.108). The federal agencies responsible for listing are the Fish and Wildlife Service (USFWS) and the National Marine Fisheries Service (NMFS). Prior federal regulations include the Endangered Species Conservation Act of 1969, and the Endangered Species Preservation Act of 1966, under which all species with a federal list date of March 11, 1967 were listed. The official federal listing of Endangered and Threatened animals is published in the Federal Register, 50 CFR §17.11

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Listing Status Summary Table

Totals include subspecies, Distinct Segments, and Ecologically Significant Units when listed separately.

Abbreviation Designation Totals SE State Listed - Endangered 52 ST State Listed - Threatened 41 SC State Candidate for Listing 10 SCD State Candidate for Delisting 0 SDR State Delisted (Recovered) 2 SDE State Delisted (Extinct) 2 FE Federally Listed - Endangered 87 FT Federally Listed - Threatened 42 FPE Federally Proposed - Endangered 2 FPT Federally Proposed - Threatened 1 FC Federal Candidate for Listing 3 FPDE Federally Proposed for Delisting (currently Endangered) 0 FPDT Federally Proposed for Delisting (currently Threatened) 1 FDR Federally Delisted (Recovered) 13 FDE Federally Delisted (Extinct) 2 # Animal Taxa State Listed Only 43 # Animal Taxa Federally Listed Only 79 # Animal Taxa State AND Federally Listed 51 Total # State Listed Taxa (SE, ST, SCD) 93 Total # Federally Listed Taxa (FE, FPDE, FT, FPDT) 130 Total # Listed Animal Taxa 173

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List Content

Taxa Group Page Gastropods 1 1 2 5 11 Reptiles 15 16 19

Common and scientific names are shown as they are in current usage, typically based standards used by the NatureServe Natural Heritage Network, unless otherwise noted. If current nomenclature differs from that in state and federal listing documents, the nomenclature at the time of listing is provided in the notes. Synonyms, name changes, and other clarifying points are also noted. Where state and federal listings apply to different ranges, subspecies, or , each taxa will be listed separately, and statuses that apply to only a portion of the taxon, or that also encompass other taxa, will be shown in parentheses. Where state and federal listings differ in name, but represent the same biological unit, the common name will be listed using the California state listing; the federal name will be listed in the notes.

The “List Date” for final federal listing is the date the listing became effective. This is typically not the date of publication of the rule in the Federal Register; it is usually about 30 days after publication, but may be longer.

If an animal was previously listed and no longer has any listing status, the entry text is grey. If an animal was previously proposed or a candidate for listing, but the listing was not warranted or was revoked, the record has been removed from the table.

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For taxa having more than one status entry, the current status is in bold and underlined. All dates are in the “YYYYMMDD” format.

Abbreviations

• CCR: California Code of Regulations • CDFW: California Department of Fish and Wildlife (previously Department of Fish and Game (DFG)) • CESA: California Endangered Species Act • DPS: Distinct population segment • ESA: Endangered Species Act (Federal) • ESU: Evolutionarily significant unit • FGC: California Fish and Game Commission • NMFS: National Marine Fisheries Service • NOAA: National Oceanic and Atmospheric Administration • USFWS: United States Fish and Wildlife Service

Additional Resources

• The California Fish and Game Commission publishes notices relating to changes to Title 14 of the California Code of Regulations • Title 14 of the California Code of Regulations can be accessed through The Office of Administrative Law • The U.S. Fish and Wildlife Service is responsible for protecting endangered and , and conserving candidate and at-risk species so that ESA listing is not necessary • The National Marine Fisheries Service (a.k.a. NOAA Fisheries) Office of Protected Resources is responsible for protecting marine mammals and endangered and threatened

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State and Federally Listed Endangered and Threatened Animals of California Last updated July 6, 2021

State State List Federal Federal Taxa Group Taxa Subgroup Taxon Common Name Status Date Status List Date Notes Invertebrates Haliotis cracherodii Black abalone FE 20110413 Listed by NMFS in 2009 and by USFWS in (snails, slugs, and 20090213 2011. abalone) Invertebrates Gastropoda Haliotis sorenseni White abalone FE 20051116 Listed by NMFS in 2001 and by USFWS in (snails, slugs, and 20010628 2005. abalone) Invertebrates Gastropoda Helminthoglypta Morro shoulderband FE 19950117 The 2006 five year review should be consulted (snails, slugs, and walkeriana (=banded dune) snail to better understand the status of this species. abalone) See Federal Register 85(143): 44821-44835, 20200724 for downlisting proposed rule.

Invertebrates Gastropoda infumata Trinity bristle snail ST 19801002 Listed by the State of California as Monadenia (snails, slugs, and setosa setosa . abalone) Invertebrates Crustacea - Anostraca Branchinecta Conservancy fairy FE 19940919 (fairy shrimp) conservatio shrimp

Invertebrates Crustacea - Anostraca Branchinecta Longhorn fairy shrimp FE 19940919 (fairy shrimp) longiantenna

Invertebrates Crustacea - Anostraca Branchinecta lynchi Vernal pool fairy FT 19940919 (fairy shrimp) shrimp

Invertebrates Crustacea - Anostraca Branchinecta San Diego fairy FE 19970203 (fairy shrimp) sandiegonensis shrimp

Invertebrates Crustacea - Anostraca Streptocephalus Riverside fairy shrimp FE 19930803 (fairy shrimp) woottoni

Invertebrates Crustacea - Notostraca Vernal pool tadpole FE 19940919 (tadpole shrimp) shrimp

Page 1 of 25 State and Federally Listed Endangered and Threatened Animals of California Last updated July 6, 2021

State State List Federal Federal Taxa Group Taxa Subgroup Taxon Common Name Status Date Status List Date Notes Invertebrates Crustacea - Shasta crayfish SE 19880226 FE 19880930 (crayfish and shrimp) ST 19801002

Invertebrates Crustacea - Decapoda pacifica California freshwater SE 19801002 FE 19881031 (crayfish and shrimp) shrimp

Invertebrates Insecta - Orthoptera Trimerotropis Zayante band-winged FE 19970224 (grasshoppers, infantilis grasshopper katydids, and crickets)

Invertebrates Insecta - Coleoptera Cicindela ohlone Ohlone FE 20011003 () Invertebrates Insecta - Coleoptera Desmocerus Valley elderberry FT 19800915 (beetles) californicus longhorn beetle dimorphus Invertebrates Insecta - Coleoptera Dinacoma caseyi Casey’s June beetle FE 20111024 (beetles) Invertebrates Insecta - Coleoptera Elaphrus viridis Delta green ground FT 19800915 (beetles) beetle Invertebrates Insecta - Coleoptera Polyphylla barbata Mount Hermon June FE 19970224 (beetles) beetle Invertebrates Insecta - Diptera Rhaphiomidas Delhi Sands flower- FE 19930923 (flies) terminatus loving fly abdominalis Invertebrates Insecta - Apodemia mormo Lange’s metalmark FE 19760608 ( and moths) langei

Invertebrates Insecta - Lepidoptera mossii FE 19760608 Synonymous with fotis bayensis and (butterflies and moths) bayensis butterfly Callophrys fotis bayensis .

Page 2 of 25 State and Federally Listed Endangered and Threatened Animals of California Last updated July 6, 2021

State State List Federal Federal Taxa Group Taxa Subgroup Taxon Common Name Status Date Status List Date Notes Invertebrates Insecta - Lepidoptera Danaus plexippus Monarch - California FC 20201217 (butterflies and moths) pop. 1 overwintering population Invertebrates Insecta - Lepidoptera battoides El Segundo blue FE 19760608 (butterflies and moths) allyni butterfly

Invertebrates Insecta - Lepidoptera Euphilotes enoptes Smith’s blue butterfly FE 19760608 Synonymous with Philotes enoptes smithi and (butterflies and moths) smithi Shijimiaeoides enoptes smithi .

Invertebrates Insecta - Lepidoptera Euphydryas editha Bay checkerspot FT 19871019 (butterflies and moths) bayensis butterfly

Invertebrates Insecta - Lepidoptera Euphydryas editha FE 19970116 Synonymous with Euphydryas editha wrighti (butterflies and moths) quino butterfly

Invertebrates Insecta - Lepidoptera Euproserpinus Kern primrose sphinx FT 19800509 (butterflies and moths) euterpe moth

Invertebrates Insecta - Lepidoptera Glaucopsyche FE 19800801 (butterflies and moths) lygdamus butterfly palosverdesensis Invertebrates Insecta - Lepidoptera Lycaena hermes Hermes copper FPT 20200108 (butterflies and moths) butterfly

Invertebrates Insecta - Lepidoptera icarioides Mission blue butterfly FE 19760608 Synonymous with (butterflies and moths) missionensis missionensis .

Invertebrates Insecta - Lepidoptera Plebejus idas lotis Lotis blue butterfly FE 19760608 Synonymous with Plebejus anna lotis and (butterflies and moths) Lycaeides argyrognomon lotis .

Page 3 of 25 State and Federally Listed Endangered and Threatened Animals of California Last updated July 6, 2021

State State List Federal Federal Taxa Group Taxa Subgroup Taxon Common Name Status Date Status List Date Notes Invertebrates Insecta - Lepidoptera Pseudocopaeodes Carson wandering FE 20020807 (butterflies and moths) eunus obscurus skipper

Invertebrates Insecta - Lepidoptera Pyrgus ruralis Laguna Mountains FE 19970116 (butterflies and moths) lagunae skipper

Invertebrates Insecta - Lepidoptera Speyeria callippe Callippe silverspot FE 19971205 (butterflies and moths) callippe butterfly

Invertebrates Insecta - Lepidoptera Speyeria zerene Behren’s silverspot FE 19971205 (butterflies and moths) behrensii butterfly

Invertebrates Insecta - Lepidoptera Speyeria zerene Oregon silverspot FT 19801015 (butterflies and moths) hippolyta butterfly

Invertebrates Insecta - Lepidoptera Speyeria zerene Myrtle’s silverspot FE 19920622 The USFWS and others have not yet (butterflies and moths) myrtleae butterfly determined if the taxonomic expansion by Emmel and Emmel (1998) into S. z. myrtleae and S. z. puntareyes is warranted. Speyereia zerene along the coast of Marin and Sonoma counties are Federally Endangered under the subspecies concept in the 1992 listing.

Invertebrates Insecta - Bombus crotchii Crotch bumble SC 20190618 (ants, , and wasps)

Invertebrates Insecta - Hymenoptera Bombus franklini Franklin's bumble SC 20190618 FPE 20190813 (ants, bees, and bee wasps)

Page 4 of 25 State and Federally Listed Endangered and Threatened Animals of California Last updated July 6, 2021

State State List Federal Federal Taxa Group Taxa Subgroup Taxon Common Name Status Date Status List Date Notes Invertebrates Insecta - Hymenoptera Bombus occidentalis western bumble bee SC 20190618 (ants, bees, and wasps)

Invertebrates Insecta - Hymenoptera Bombus suckleyi Suckley's cuckoo SC 20190618 (ants, bees, and bumble bee wasps)

Fishes Acipenseridae Acipenser medirostris Green sturgeon FT 20060606 Includes all spawning populations south of the (sturgeon) [southern DPS] Eel River. Fishes Cyprinidae crassicauda SDE 19801002 Extinct ( and carp) SE 19740110 Fishes Cyprinidae Gila elegans Bonytail SE 19740110 FE 19800523 Federal common name: . (minnows and carp) ST 19710627 Fishes Cyprinidae Lavinia exilicauda chi Clear Lake hitch ST 20140806 (minnows and carp) Fishes Cyprinidae Ptychocheilus lucius pikeminnow SE 19710627 FE 19670311 (minnows and carp) Fishes Cyprinidae Siphateles bicolor Mohave tui chub SE 19710627 FE 19701013 Listed by the State of California as Gila bicolor (minnows and carp) mohavensis mohavensis . Fishes Cyprinidae Siphateles bicolor Owens tui chub SE 19740110 FE 19850904 Listed by the State of California as Gila bicolor (minnows and carp) snyderi snyderi . Fishes microps SE 19801002 FDR 20160107 Recovered (suckers) ST 19740110 FE 19850711 Fishes Catostomidae Catostomus FT 20000512 Populations in the Los Angeles, San Gabriel, (suckers) santaanae and Santa Ana River basins. Fishes Catostomidae Chasmistes Shortnose sucker SE 19740110 FE 19880817 (suckers) brevirostris ST 19710627 Fishes Catostomidae Deltistes luxatus Lost River sucker SE 19740110 FE 19880817 (suckers) ST 19710627 Fishes Catostomidae Xyrauchen texanus SE 19740110 FE 19911122 (suckers) ST 19710627

Page 5 of 25 State and Federally Listed Endangered and Threatened Animals of California Last updated July 6, 2021

State State List Federal Federal Taxa Group Taxa Subgroup Taxon Common Name Status Date Status List Date Notes Fishes Osmeridae Delta SE 20100120 FT 19930305 20161202 USFWS Annual Notification of (smelt) transpacificus ST 19931209 Findings indicates uplisting to Federally Endangered (original uplisting petition received 20060308) is "warranted-but-precluded," with a Listing Priority Number of 2.

Fishes Osmeridae Spirinchus Longfin smelt ST 20090405 FC 20120402 Federal candidacy is only for Bay- (smelt) thaleichthys Delta distinct population segment. Fishes Osmeridae Thaleichthys Pacific eulachon FT 20110413 Eulachon was listed as Threatened by NMFS in (smelt) pacificus [southern DPS] 20100517 2010 and by USFWS in 2011. Fishes Salmonidae clarkii Lahontan cutthroat FT 19750716 Early Federal Register notices spelled "clarkii " (trout and salmon) henshawi trout FE 19701013 with only one "i" Fishes Salmonidae Oncorhynchus clarkii Paiute cutthroat trout FT 19750716 (trout and salmon) seleniris FE 19670311

Page 6 of 25 State and Federally Listed Endangered and Threatened Animals of California Last updated July 6, 2021

State State List Federal Federal Taxa Group Taxa Subgroup Taxon Common Name Status Date Status List Date Notes Fishes Salmonidae Oncorhynchus Coho salmon [south SE 20050330 FE 20050829 The Federal listing is for the Central California (trout and salmon) kisutch of Punta Gorda FT 19961130 Coast Coho ESU and includes populations from (Humboldt County), Punta Gorda south to, and including, the San California] Lorenzo River as well as populations in tributaries to , excluding the Sacramento- system. Coho south of San Francisco Bay were state listed in 1995. In February 2004 the Fish and Game Commission determined that coho from San Francisco to Punta Gorda should also be listed as Endangered. This change was finalized by the Office of Administrative Law on 20050330. NMFS completed a comprehensive status review in 2005 reaffirming the status, and uplisting the Central Coast ESU from threatened to endangered. NMFS reaffirmed the FE status again 20140723.

Fishes Salmonidae Oncorhynchus Coho salmon [from ST 20050330 FT 19970605 The Federal listing is for the Southern Oregon- (trout and salmon) kisutch Punta Gorda Coast ESU, and includes (Humboldt County), populations in coastal between Cape California to the Blanco, Oregon and Punta Gorda, California. northern border of The Fish and Game Commission determined California] that coho from Punta Gorda to the Oregon border should be listed as Threatened on 20040225. This determination was finalized by the Office of Administrative Law on 20050330. NMFS completed a comprehensive status review 20050829 reaffirming the status.

Page 7 of 25 State and Federally Listed Endangered and Threatened Animals of California Last updated July 6, 2021

State State List Federal Federal Taxa Group Taxa Subgroup Taxon Common Name Status Date Status List Date Notes Fishes Salmonidae Oncorhynchus Steelhead [Southern FE 19971017 Coastal basins from the Santa Maria River (trout and salmon) mykiss irideus California DPS] (inclusive), south to the U.S.-Mexico Border. NMFS completed a comprehensive status review 20060206 reaffirming the status. Fishes Salmonidae Oncorhynchus Steelhead [South FT 19971017 Coastal basins from the Pajaro River (inclusive) (trout and salmon) mykiss irideus Central California south to, but not including, the Santa Maria Coast DPS] River. NMFS completed a comprehensive status review 20060206 reaffirming the status.

Fishes Salmonidae Oncorhynchus Steelhead [California FT 19980518 The Sacramento and San Joaquin Rivers and (trout and salmon) mykiss irideus Central Valley DPS] their tributaries. NMFS completed a comprehensive status review 20060206 reaffirming the status. Fishes Salmonidae Oncorhynchus Steelhead [Central FT 19971017 Coastal streams from the Russian River (trout and salmon) mykiss irideus California Coast (inclusive) to Aptos Creek (inclusive), and the DPS] drainages of San Francisco, San Pablo, and Suisun Bays eastward to at the confluence of the Sacramento and San Joaquin Rivers; and tributary streams to including Suisun Creek, Green Valley Creek, and an unnamed tributary to (commonly referred to as Red Top Creek), exclusive of the Sacramento-San Joaquin River Basin of the California Central Valley. NMFS completed a comprehensive status review 20060206 reaffirming the status.

Page 8 of 25 State and Federally Listed Endangered and Threatened Animals of California Last updated July 6, 2021

State State List Federal Federal Taxa Group Taxa Subgroup Taxon Common Name Status Date Status List Date Notes Fishes Salmonidae Oncorhynchus Steelhead [Northern FT 20000807 Naturally spawned populations residing below (trout and salmon) mykiss irideus California DPS] impassable barriers in coastal basins from Redwood Creek in Humboldt County to, and including, the Gualala River in Mendocino County. NMFS completed a comprehensive status review 20060206 reaffirming the status.

Fishes Salmonidae Oncorhynchus Steelhead [summer- SC 20190618 CNDDB-tracked taxa includes northern (trout and salmon) mykiss irideus run] California coastal streams south to Middle Fork Eel River, within range of Klamath Mountains Province DPS and Northern California DPS. Proposed CESA listing status applies only to Northern California DPS summer-run steelhead.

Fishes Salmonidae Oncorhynchus Little Kern golden FT 19780515 Originally listed as Salmo aguabonita whitei . (trout and salmon) mykiss whitei trout The Salmo was reclassified as Oncorhynchus changing the name to Oncorhynchus aguabonita whitei . However, recent studies indicate this is a subspecies of , therefore Oncorhynchus mykiss whitei . Fishes Salmonidae Oncorhynchus Chinook salmon SE 19890922 FE 19940323 The federal designation is for the Sacramento (trout and salmon) tshawytscha [winter run] FT 19901130 River winter-run ESU, and described as winter- run populations in the and its tributaries in California. NMFS completed a comprehensive status review 20050829 reaffirming the status.

Fishes Salmonidae Oncorhynchus Chinook salmon SC 20190212 FC 20180227 Spring-run Chinook salmon in the Trinity River (trout and salmon) tshawytscha [Upper Klamath- and the upstream of the mouth of Trinity River Spring the Trinity River. ESU]

Page 9 of 25 State and Federally Listed Endangered and Threatened Animals of California Last updated July 6, 2021

State State List Federal Federal Taxa Group Taxa Subgroup Taxon Common Name Status Date Status List Date Notes Fishes Salmonidae Oncorhynchus Chinook salmon FT 19991115 Rivers and streams south of the Klamath River (trout and salmon) tshawytscha [California Coastal to the Russian River. NMFS completed a ESU] comprehensive status review 20050829 reaffirming the status. Fishes Salmonidae Oncorhynchus Chinook salmon ST 19990205 FT 19991115 The State listing is for “Spring-run chinook (trout and salmon) tshawytscha [spring-run of the salmon (Oncorhynchus tshawytscha ) of the Sacramento River Sacramento River drainage.” The Federal listing drainage] is for the Central Valley spring-run ESU, and includes populations of spring-run Chinook salmon in the Sacramento River and its tributaries including the Feather River. NMFS completed a comprehensive status review 20050829 reaffirming the status. Fishes Salmonidae Salvelinus Bull trout SE 19801002 FT 19991201 Considered to be extirpated in California. (trout and salmon) confluentus Fishes Cyprinodontidae SE 19801002 FE 19860430 (killifishes) macularius Fishes Cyprinodontidae Cyprinodon Tecopa pupfish SDE 19870609 FDE 19820216 Extinct (killifishes) nevadensis calidae SE 19710627 FE 19701013 Fishes Cyprinodontidae Cyprinodon radiosus SE 19710627 FE 19670311 (killifishes) Fishes Cyprinodontidae Cyprinodon salinus Cottonball Marsh ST 19740110 (killifishes) milleri pupfish Fishes Gasterosteidae Gasterosteus Unarmored SE 19710627 FE 19701013 (sticklebacks) aculeatus williamsoni threespine stickleback Fishes Cottidae Cottus asperrimus Rough sculpin ST 19740110 (sculpins) Fishes Gobiidae Eucyclogobius Tidewater goby FE 19940307 See Federal Register 79(49):14340-14362, (gobies) newberryi 20140313, for down-listing proposed rule.

Page 10 of 25 State and Federally Listed Endangered and Threatened Animals of California Last updated July 6, 2021

State State List Federal Federal Taxa Group Taxa Subgroup Taxon Common Name Status Date Status List Date Notes Amphibians Ambystomatidae Ambystoma California tiger ST 20100819 (FE), The State listing applies to the species as a (mole ) californiense (FT) whole throughout its range; federal statuses apply to Distinct Population Segments (see below). Amphibians Ambystomatidae Ambystoma California tiger (ST) FT 20040903 The 2004 federal Threatened status originally (mole salamanders) californiense salamander [Central applied to the species throughout its range; California DPS] subsequent legal action resulted in reclassification of other DPSs to Endangered; the central California DPS remained listed as Threatened. Amphibians Ambystomatidae Ambystoma California tiger (ST) FE 20000915 In 2004 the California tiger salamander was (mole salamanders) californiense salamander [Santa federally listed as Threatened statewide. The Barbara County DPS] Santa Barbara County and Sonoma County Distinct Vertebrate Population Segments (DPS), formerly listed as Endangered, were reclassified to Threatened. On 20050819 U.S. District court vacated the down-listing of the Sonoma and Santa Barbara populations from Endangered to Threatened. Therefore, the Sonoma & Santa Barbara populations were once again listed as Endangered. Amphibians Ambystomatidae Ambystoma California tiger (ST) FE 20030319 In 2004 the California tiger salamander was (mole salamanders) californiense salamander [Sonoma federally listed as Threatened statewide. The County DPS] Santa Barbara County and Sonoma County Distinct Vertebrate Population Segments (DPS), formerly listed as Endangered, were reclassified to Threatened. On 20050819 U.S. District court vacated the down-listing of the Sonoma and Santa Barbara populations from Endangered to Threatened. Therefore, the Sonoma & Santa Barbara populations were once again listed as Endangered.

Page 11 of 25 State and Federally Listed Endangered and Threatened Animals of California Last updated July 6, 2021

State State List Federal Federal Taxa Group Taxa Subgroup Taxon Common Name Status Date Status List Date Notes Amphibians Ambystomatidae Ambystoma Santa Cruz long-toed SE 19710627 FE 19670311 (mole salamanders) macrodactylum salamander croceum Amphibians Batrachoseps major Desert slender SE 19710627 FE 19730604 Listed by the State of California as (lungless salamanders) aridus salamander Batrachoseps aridus and originally listed by the USFWS as B. aridus . USFWS 5-year review refers to B. major aridus . Amphibians Plethodontidae Batrachoseps Kern Canyon slender ST 19710627 (lungless salamanders) simatus salamander

Amphibians Plethodontidae Batrachoseps Tehachapi slender ST 19710627 (lungless salamanders) stebbinsi salamander

Amphibians Plethodontidae shastae Shasta salamander ST 19710627 (lungless salamanders)

Amphibians Plethodontidae Hydromantes brunus Limestone ST 19710627 (lungless salamanders) salamander

Amphibians Plethodontidae Plethodon asupak Scott Bar salamander ST 19710627 As recognized by the FGC, the Scott Bar (lungless salamanders) salamander is currently protected under the CESA as a sub-population of the Siskiyou Mountains salamander (Plethodon stormi ) (Calif. Regulatory Notice Register, No. 21-Z, p. 916, 20070525).

Page 12 of 25 State and Federally Listed Endangered and Threatened Animals of California Last updated July 6, 2021

State State List Federal Federal Taxa Group Taxa Subgroup Taxon Common Name Status Date Status List Date Notes Amphibians Plethodontidae Plethodon stormi Siskiyou Mountains ST 19710627 The common name is spelled incorrectly in Title (lungless salamanders) salamander 14 of the CCR as “Siskiyou mountain salamander.” Was a State Candidate for Delisting on 20050930. No action was taken by the FGC after the CDFW presented a Department report on 20061103; SMS was tabled at the 20070503 FGC meeting, and there was nothing to report regarding the Department’s environmental documents at the 20071011 meeting. Therefore, with respect to Fish & Game Code 2075, it is assumed that this is no longer a candidate for delisting.

Amphibians Bufonidae Anaxyrus californicus FE 19950117 At the time of listing, arroyo toad was known as (true toads) Bufo microscaphus californicus , a subspecies of southwestern toad. In 2001, it was determined to be its own species, Bufo californicus . Since then, many species in the genus Bufo were changed to the genus Anaxyrus , and now arroyo toad is known as Anaxyrus californicus .

Amphibians Bufonidae Anaxyrus canorus FT 20140630 (true toads) Amphibians Bufonidae Anaxyrus exsul Black toad ST 19710627 Listed by the State of California as Bufo exsul . (true toads) Amphibians Ranidae Rana boylii Foothill yellow-legged SE 20200320 CESA listing status varies by clade; listing of the (true frogs) frog (Southwest / Northwest/North Coast clade is not warranted. South Coast clade)

Page 13 of 25 State and Federally Listed Endangered and Threatened Animals of California Last updated July 6, 2021

State State List Federal Federal Taxa Group Taxa Subgroup Taxon Common Name Status Date Status List Date Notes Amphibians Ranidae Rana boylii Foothill yellow-legged SE 20200320 CESA listing status varies by clade; listing of the (true frogs) frog (West / Central Northwest/North Coast clade is not warranted. Coast clade)

Amphibians Ranidae Rana boylii Foothill yellow-legged SE 20200320 CESA listing status varies by clade; listing of the (true frogs) frog (East / Southern Northwest/North Coast clade is not warranted. Sierra clade)

Amphibians Ranidae Rana boylii Foothill yellow-legged ST 20200320 CESA listing status varies by clade; listing of the (true frogs) frog (northeast / Northwest/North Coast clade is not warranted. Northern Sierra clade)

Amphibians Ranidae Rana boylii Foothill yellow-legged ST 20200320 CESA listing status varies by clade; listing of the (true frogs) frog (Feather River Northwest/North Coast clade is not warranted. clade)

Amphibians Ranidae Rana cascadae Cascades frog SC 20171017 Date of FGC vote to advance to candidacy was (true frogs) 20171011. Amphibians Ranidae Rana draytonii California red-legged FT 19960624 Synonymous with Rana aurora draytonii . (true frogs) frog Amphibians Ranidae Rana muscosa Southern mountain SE 20130401 (FE) Though the scientific name Rana muscosa is (true frogs) yellow-legged frog not disputed, the State uses this common name, whereas the USFWS listing refers to two distinct population segments. This species is also known by the common name Sierra Madre yellow- legged frog (Vredenburg et al. 2007).

Amphibians Ranidae Rana muscosa Mountain yellow- (SE) FE 20020801 San Gabriel, San Jacinto, and San Bernardino (true frogs) legged frog [Southern Mountains only. California DPS]

Page 14 of 25 State and Federally Listed Endangered and Threatened Animals of California Last updated July 6, 2021

State State List Federal Federal Taxa Group Taxa Subgroup Taxon Common Name Status Date Status List Date Notes Amphibians Ranidae Rana muscosa Mountain yellow- (SE) FE 20140630 North of the Tehachapi Mountains from the (true frogs) legged frog [Northern Monarch Divide to portions of the Kern River California DPS] drainage.

Amphibians Ranidae Rana pretiosa Oregon spotted frog FT 20140929 (true frogs) Amphibians Ranidae Rana sierrae Sierra yellow- ST 20130401 FE 20140630 (true frogs) legged frog Reptiles Cheloniidae Caretta caretta Loggerhead sea FE 20111024 The 1978 listing was for the worldwide range of (sea turtles) turtle [North Pacific FT 19780728 the species. The 20111024 final rule is for the DPS] North Pacific DPS (north of the equator & south of 60 degrees north latitude). Reptiles Cheloniidae Chelonia mydas Green turtle FT 20160506 Alternate common name: green . (sea turtles) FT 19780728 Current FT status refers to East Pacific DPS. FE 19701013 Reptiles Cheloniidae Lepidochelys Olive (=Pacific) ridley FT 19780728 (sea turtles) olivacea sea turtle Reptiles Dermochelyidae Dermochelys Leatherback sea SCE 20200825 FE 19700602 (leatherback turtles) coriacea turtle Reptiles Testudinidae Gopherus agassizii SCE 20201019 FT 19900402 (land tortoises) ST 19890803 Reptiles Gekkonidae Coleonyx switaki Barefoot gecko ST 19801002 Alternate common names: Switak's banded (geckos) gecko, barefoot banded gecko. Reptiles Crotaphytidae Gambelia sila Blunt-nosed SE 19710627 FE 19670311 Synonymous with Gambelia silus . Originally (collared and leopard lizard listed under the ESA as Crotaphytus wislizenii lizards) silus . Reptiles Phrynosomatidae Uma inornata Coachella Valley SE 19801002 FT 19801027 (spiny lizards) fringe-toed lizard Reptiles Xantusiidae Xantusia riversiana Island night lizard FDR 20140501 Recovered (night lizards) FT 19770811 Reptiles Boidae Charina umbratica Southern rubber boa ST 19710627 Synonymous with Charina bottae umbratica . (boas)

Page 15 of 25 State and Federally Listed Endangered and Threatened Animals of California Last updated July 6, 2021

State State List Federal Federal Taxa Group Taxa Subgroup Taxon Common Name Status Date Status List Date Notes Reptiles Colubridae Masticophis lateralis whipsnake ST 19710627 FT 19971205 Synonymous with Coluber lateralis euryxanthus . (egg-laying snakes) euryxanthus

Reptiles Natricidae Thamnophis gigas Giant garter snake ST 19710627 FT 19931119 Listed by State of California as Thamnophis (live-bearing snakes) couchi gigas . Reptiles Natricidae Thamnophis sirtalis San Francisco garter SE 19710627 FE 19670311 (live-bearing snakes) tetrataenia snake Birds Anatidae Branta hutchinsii Cackling (=Aleutian FDR 20010320 Recovered. At time of federal listing, known as (ducks, geese, and leucopareia Canada) goose FT 19910111 Branta canadensis leucopareia . swans) FE 19670311 Birds Diomedeidae Phoebastria albatrus Short-tailed FE 20000830 Synonymous with Diomedea albatrus . Listed as () FE 19700602 Endangered in one of the original species lists, but “due to an inadvertent oversight” when the 1973 ESA repealed the 1969 Act, short-tailed albatross was effectively delisted. Proposed listing to fix this error in 1980, with final rule in 2000. Birds Pelecaniidae Pelecanus California brown SDR 20090603 FDR 20091217 Recovered. Federal nomenclature: Brown (pelicans) occidentalis pelican SE 19710627 FE 19701013 pelican (Pelecanus occidentalis ). californicus Birds Cathartidae Gymnogyps SE 19710627 FE 19670311 (New World vultures) californianus Birds Accipitridae Buteo swainsoni Swainson’s hawk ST 19830417 (hawks, kites, harriers, and eagles) Birds Accipitridae Haliaeetus SE (rev) 19801002 FDR 20070808 The Post-delisting Monitoring Plan will monitor (hawks, kites, harriers, leucocephalus SE 19710627 FT 19950811 the status of the bald eagle over a 20 year and eagles) FE (rev) 19780316 period with sampling events held once every 5 FE 19670311 years. Birds Falconidae Falco peregrinus American peregrine SDR 20091104 FDR 19990825 Recovered (falcons) anatum falcon SE 19710627 FE 19700602

Page 16 of 25 State and Federally Listed Endangered and Threatened Animals of California Last updated July 6, 2021

State State List Federal Federal Taxa Group Taxa Subgroup Taxon Common Name Status Date Status List Date Notes Birds Falconidae Falco peregrinus Arctic peregrine FDR 19941005 Recovered (falcons) tundrius falcon FT 19840419 FE 19700602 Birds Rallidae Laterallus California black ST 19710627 (rails, coots, and jamaicensis gallinules) coturniculus Birds Rallidae Rallus obsoletus Light-footed SE 19710627 FE 19701013 Formerly light-footed clapper rail, Rallus (rails, coots, and levipes Ridgway's rail longirostris levipes gallinules) Birds Rallidae Rallus obsoletus California Ridgway's SE 19710627 FE 19701013 Formerly California clapper rail, Rallus (rails, coots, and obsoletus rail longirostris obsoletus gallinules) Birds Rallidae Rallus obsoletus Yuma Ridgway's rail ST 19780222 FE 19670311 Formerly Yuma clapper rail, Rallus longirostris (rails, coots, and yumanensis SE 19710627 yumanensis gallinules) Birds Gruidae Grus canadensis Greater sandhill ST 19830417 (cranes) tabida Birds Charadriidae Charadrius nivosus Western snowy FT 19930405 Federal status applies only to the Pacific coastal (plovers and relatives) nivosus plover population.

Birds Laridae Sternula antillarum California least SE 19710627 FE 19700602 Listed by the State of California and federal (gulls and ) browni government as Sterna antillarum browni . Birds Alcidae Brachyramphus Marbled murrelet SE 19920312 FT 19920928 (auklets, puffins, and marmoratus relatives) Birds Alcidae Synthliboramphus Scripps’s murrelet ST 20041222 At the time of listing, this species was known as (auklets, puffins, and scrippsi (=Xantus’s murrelet) the Xantus’s Murrelet (Synthliboramphus relatives) hypoleucus , with California breeding populations ascribed to Synthliboramphus hypoleucus subsp. scrippsi ).

Page 17 of 25 State and Federally Listed Endangered and Threatened Animals of California Last updated July 6, 2021

State State List Federal Federal Taxa Group Taxa Subgroup Taxon Common Name Status Date Status List Date Notes Birds Alcidae Synthliboramphus ST 20041222 At the time of listing, this species was known as (auklets, puffins, and hypoleucus (=Xantus’s murrelet) the Xantus’s Murrelet (Synthliboramphus relatives) hypoleucus , with breeding populations from Baja California ascribed to Synthliboramphus hypoleucus subsp. hypoleucus ).

Birds Cuculidae Coccyzus Western yellow-billed SE 19880326 FT 20141103 Federal listing is for the Western DPS of (cuckoos and relatives) americanus cuckoo ST 19710627 Coccyzus americanus . occidentalis Birds Strigidae Micrathene whitneyi Elf SE 19801002 () Birds Strigidae Strix nebulosa Great gray owl SE 19801002 (owls) Birds Strigidae Strix occidentalis Northern spotted owl ST 20190318 FT 19900723 (owls) caurina Birds Picidae Colaptes chrysoides Gilded (=Gilded SE 19880317 Listed by the State of California as Colaptes (woodpeckers) northern) flicker auratus chrysoides . Birds Picidae Melanerpes Gila woodpecker SE 19880317 (woodpeckers) uropygialis Birds Tyrannidae Empidonax traillii Willow flycatcher SE 19910102 State listing includes all subspecies. (tyrant flycatchers) Birds Tyrannidae Empidonax traillii Southwestern willow (SE) FE 19950329 (tyrant flycatchers) extimus flycatcher Birds Laniidae Lanius ludovicianus San Clemente FE 19770912 (shrikes) mearnsi loggerhead shrike

Birds Vireonidae Vireo bellii arizonae Bell’s vireo SE 19880317 (vireos) Birds Vireonidae Vireo bellii pusillus Least Bell’s vireo SE 19801002 FE 19860602 (vireos) Birds Hirundinidae Riparia riparia Bank ST 19890611 ()

Page 18 of 25 State and Federally Listed Endangered and Threatened Animals of California Last updated July 6, 2021

State State List Federal Federal Taxa Group Taxa Subgroup Taxon Common Name Status Date Status List Date Notes Birds Polioptilidae Polioptila californica Coastal California FT 19930330 (gnatcatchers) californica gnatcatcher

Birds Emberizidae Artemisiospiza belli San Clemente sage FT 19770912 Federal nomenclature at time of listing: (sparrows, buntings, clementeae sparrow Amphispiza belli clementeae . warblers, and relatives)

Birds Emberizidae Melospiza melodia Santa Barbara song FDE 19831012 Extinct. This status refers specifically to the (sparrows, buntings, graminea sparrow FE 19730604 Santa Barbara song sparrow, which was later warblers, and relatives) reclassified as a subspecies (Channel Islands song sparrow) with the same scientific name, but which also combined two additional groups formerly classified as their own subspecies.

Birds Emberizidae Melozone crissalis Inyo California SE 19801002 FPD 20131104 Listed by the State of California and federal (sparrows, buntings, eremophilus towhee FT 19870902 government as Pipilo crissalis eremophilus . warblers, and relatives)

Birds Emberizidae Passerculus Belding’s savannah SE 19740110 Listed by the State of California as Passerculus (sparrows, buntings, sandwichensis sparrow sandwichensis beldingii warblers, and relatives) beldingi

Birds Icteridae Agelaius tricolor ST 20190318 (blackbirds) Mammals Soricidae Sorex ornatus relictus Buena Vista Lake FE 20020405 (shrews) ornate shrew Mammals Phyllostomidae Leptonycteris Lesser long-nosed FDR 20180518 Recovered. (leaf-nosed bats) yerbabuenae bat FE 19881031 Mammals Leporidae Sylvilagus bachmani Riparian brush rabbit SE 19940529 FE 20000324 (rabbits and hares) riparius Mammals Aplodontidae Aplodontia rufa nigra Point Arena mountain FE 19911212 (mountain beavers) beaver

Page 19 of 25 State and Federally Listed Endangered and Threatened Animals of California Last updated July 6, 2021

State State List Federal Federal Taxa Group Taxa Subgroup Taxon Common Name Status Date Status List Date Notes Mammals Sciuridae Ammospermophilus Nelson’s (=San ST 19801002 (squirrels and relatives) nelsoni Joaquin) antelope squirrel Mammals Sciuridae Xerospermophilus Mohave ground ST 19710627 Listed by the State of California as (squirrels and relatives) mohavensis squirrel Spermophilus mohavensis .

Mammals Heteromyidae Dipodomys Morro Bay kangaroo SE 19710627 FE 19701013 (kangaroo rats, pocket heermanni rat mice, and kangaroo morroensis mice) Mammals Heteromyidae Dipodomys ingens Giant kangaroo rat SE 19801002 FE 19870105 (kangaroo rats, pocket mice, and kangaroo mice) Mammals Heteromyidae Dipodomys merriami San Bernardino SC 20190813 FE 19980924 Federal nomenclature: San Bernardino (kangaroo rats, pocket parvus kangaroo rat Merriam’s kangaroo rat. mice, and kangaroo mice) Mammals Heteromyidae Dipodomys Fresno kangaroo rat SE 19801002 FE 19850301 (kangaroo rats, pocket nitratoides exilis ST 19710627 mice, and kangaroo mice) Mammals Heteromyidae Dipodomys Tipton kangaroo rat SE 19890611 FE 19880808 (kangaroo rats, pocket nitratoides nitratoides mice, and kangaroo mice) Mammals Heteromyidae Dipodomys stephensi Stephens’ kangaroo ST 19710627 FE 19881031 See Federal Register 85(161):50991-51006, (kangaroo rats, pocket rat 20200819, for down-listing proposed rule. mice, and kangaroo mice)

Page 20 of 25 State and Federally Listed Endangered and Threatened Animals of California Last updated July 6, 2021

State State List Federal Federal Taxa Group Taxa Subgroup Taxon Common Name Status Date Status List Date Notes Mammals Heteromyidae Perognathus Pacific pocket mouse FE 19940926 (kangaroo rats, pocket longimembris mice, and kangaroo pacificus mice) Mammals Cricetidae Microtus californicus SE 19801002 FE 19841217 (New World mice, rats, scirpensis and voles) Mammals Cricetidae Neotoma fuscipes Riparian woodrat FE 20000324 (New World mice, rats, riparia and voles) Mammals Cricetidae Reithrodontomys Salt-marsh harvest SE 19710627 FE 19701013 (New World mice, rats, raviventris mouse and voles) Mammals lupus Gray SE 20170101 FDR 20201103 (, , and FE 19780410 ) Mammals Canidae littoralis Island ST 19710627 (FT) State listing includes all 6 subspecies on all 6 (foxes, wolves, and islands. Federal listing is for only 4 subspecies coyotes) on 4 islands. Mammals Canidae Urocyon littoralis Santa Catalina Island (ST) FT 20160912 (foxes, wolves, and catalinae Fox FE 20040405 coyotes) Mammals Canidae Urocyon littoralis San Miguel Island (ST) FDR 20160912 (foxes, wolves, and littoralis Fox FE 20040405 coyotes) Mammals Canidae Urocyon littoralis Santa Cruz Island (ST) FDR 20160216 (foxes, wolves, and santacruzae Fox FE 20040405 coyotes) Mammals Canidae Urocyon littoralis Santa Rosa Island (ST) FDR 20160216 (foxes, wolves, and santarosae Fox FE 20040405 coyotes)

Page 21 of 25 State and Federally Listed Endangered and Threatened Animals of California Last updated July 6, 2021

State State List Federal Federal Taxa Group Taxa Subgroup Taxon Common Name Status Date Status List Date Notes Mammals Canidae macrotis San Joaquin ST 19710627 FE 19670311 (foxes, wolves, and mutica coyotes) Mammals Canidae Vulpes vulpes Sierra Nevada red ST 19801002 FPE 20200108 Proposed federal status applies only to the (foxes, wolves, and necator fox Sierra Nevada DPS. coyotes) Mammals Enhydra lutris nereis Southern sea FT 19770114 ( and relatives)

Mammals Mustelidae Gulo gulo North American ST 19710627 Listed by the State of California as Gulo (weasels and relatives) gulo .The distinct population segment of the North American wolverine (Gulo gulo luscus ) occurring in the contiguous U.S. was proposed for listing on 20130204 and was withdrawn on 20201013. Mammals Mustelidae Martes caurina Humboldt (=Coastal) SE 20190318 FT 20201008 Federal status refers to the coastal DPS of (weasels and relatives) humboldtensis Pacific marten (Martes caurina ), which includes coastal populations in Oregon and the Humbodt marten. Mammals Mustelidae Pekania [=Martes] Pacific ST 20190318 FE 20200615 California listing is under Martes pennanti and (weasels and relatives) pennanti [Southern Sierra common name Pacific fisher, whereas the Nevada ESU/DPS] USFWS refers to Martes pennanti and common name fisher. Previous USFWS candidacy referred to the West Coast DPS in California, Oregon, and Washington. On 20190318, the Southern Sierra ESU (defined as California south of the Merced River) was officially listed as Threatened under CESA; the Southern Sierra Nevada DPS was listed as federaly endangered 20200615.

Page 22 of 25 State and Federally Listed Endangered and Threatened Animals of California Last updated July 6, 2021

State State List Federal Federal Taxa Group Taxa Subgroup Taxon Common Name Status Date Status List Date Notes Mammals concolor Mountain SC 20200421 Not currently tracked by CNDDB. (cats and relatives) (Southern California/Central Coast ESU) Mammals Otariidae Guadalupe ST 19710627 FT 19860115 Not currently tracked by CNDDB. (sea and fur townsendi FE 19670311 seals) Mammals Otariidae Eumetopias jubatus Steller FDR 20131204 Recovered. Delisted by NMFS. (sea lions and fur [Eastern DPS] FT 19970604 On 19901204, the was listed as seals) federally Threatened throughout its entire range. In 1997, NMFS reclassified Steller sea lions into two distinct population segments: the Western DPS (west of 144 degrees longitude) was listed as Endangered; the Eastern DPS (east of 144 degrees longitude) was listed as Threatened, and subsequently delisted in 2013.

Mammals Bovidae Ovis canadensis Peninsular bighorn ST 19710627 FE 19980318 Listed by the State of California as Ovis (sheep and relatives) nelsoni sheep [Peninsular CA canadensis cremnobates . The subspecies O.c. DPS] cremnobates has been synonymized with O.c. nelsoni . The desert bighorn sheep in the Peninsular Ranges, the Peninsular bighorn sheep, is now considered to be a Distinct Population Segment of O.c. nelsoni . Mammals Bovidae Ovis canadensis Sierra Nevada (= SE 19990827 FE 20000103 Listed by the State of California as California (sheep and relatives) sierrae California) bighorn ST 19710627 bighorn sheep (Ovis canadensis californiana ). sheep Mammals Cetacea Balaenoptera Sei FE 19700602 Not currently tracked by CNDDB. ( and dolphins) borealis

Page 23 of 25 State and Federally Listed Endangered and Threatened Animals of California Last updated July 6, 2021

State State List Federal Federal Taxa Group Taxa Subgroup Taxon Common Name Status Date Status List Date Notes Mammals Cetacea Balaenoptera FE 19700602 Not currently tracked by CNDDB. (whales and dolphins) musculus

Mammals Cetacea Balaenoptera FE 19700602 Not currently tracked by CNDDB. (whales and dolphins) physalus

Mammals Cetacea Eubalaena japonica North Pacific right FE 20080407 Originally listed as part of the northern right (whales and dolphins) whale FE 19700602 whale (Eubalaena glacialis ) species concept under the federal ESA. and nomenclature were updated and clarified in the Federal Register Vol. 68, No. 69, 20030410.In 2006, NMFS completed a status review of right whales in the N. Pacific and N. Atlantic Oceans and in 2008, reclassified the previously Endangered northern (Eubalaena spp.) as two separate Endangered species: North Pacific right whale (E. japonica ) and North Atlantic right whale (E. glacialis ). Not currently tracked by CNDDB.

Mammals Cetacea Eschrichtius robustus [Eastern FDR 19940616 Recovered. NMFS delisted the California (whales and dolphins) North Pacific DPS] FE 19700602 population (Eastern North Pacific DPS), while keeping the Western North Pacific DPS as Endangered. Mammals Cetacea Megaptera FE 20161011 Also known as Hump-backed whale. 2016 ruling (whales and dolphins) novaeangliae [Central America FE 19700602 by NMFS established 14 Distinct Population DPS] Segments, and revised listing status by DPS. Both the Mexico DPS and Central America DPS feed and travel off the coast of California. Not currently tracked by CNDDB.

Page 24 of 25 State and Federally Listed Endangered and Threatened Animals of California Last updated July 6, 2021

State State List Federal Federal Taxa Group Taxa Subgroup Taxon Common Name Status Date Status List Date Notes Mammals Cetacea Megaptera Humpback whale FT 20161011 Also known as Hump-backed whale. 2016 ruling (whales and dolphins) novaeangliae [Mexico DPS] FE 19700602 by NMFS established 14 Distinct Population Segments, and revised listing status by DPS. Both the Mexico DPS and Central America DPS feed and travel off the coast of California. Not currently tracked by CNDDB.

Mammals Cetacea Orcinus orca FE 20070404 The Southern Resident DPS of killer whale was (whales and dolphins) [Southern Resident 20060216 listed as Endangered by NMFS on 20060216 DPS] and by USFWS on 20070404. Not currently tracked by CNDDB. Mammals Cetacea Physeter Sperm whale FE 19700602 Federal nomenclature at time of listing: Physeter (whales and dolphins) macrocephalus catodon . Not currently tracked by CNDDB.

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