Part I. Montoya and Related Colonial Corals. Part II. Organisms Attached
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MEMOIR 7 PART I Montoya and Related Colonial Corals PART II Organisms Attached to Montoya Corals By ROUSSEAU H. FLOWER TEXT SECTION STATE BU REAU OF MINES AND MINERAL RESOU RCES NEW MEXICO INSTITUTE OF MINING AND TECHNOLOGY CAMPUS STATION SOCORRO, NEW MEXICO NEW MEXICO INSTITUTE OF MINING & TECHNOLOGY E. J. Workman, President STATE BUREAU OF MINES AND MINERAL RESOURCES Alvin J. Thompson, Director THE REGENTS MEMBERS Ex OFFICIO The Honorable Edwin L. Mechem Governor of New Mexico Tom Wiley . ... Superintendent of Public Instruction APPOINTED MEMBERS William G. Abbott .... Hobbs Holm 0. Bursum, Jr. .Socorro Thomas M. Cramer .. .. Carlsbad Frank C. DiLuzio . ............ Albuquerque Eva M. Larrazolo (Mrs. Paul F.). .. Albuquerque Published November 21, 1961 For Sale by the New Mexico Bureau of Mines & Mineral Resources Campus Station, Socorro, N. Mex.-$4.00. Price includes text and plate sections. Contents PART I MONTOYA AND RELATED COLONIAL CORALS Page ABSTRACT 3 INTRODUCTION 3 ACKNOWLEDGMENTS. 6 THE MONTOYA GROUP . 7 General divisions and correlations . 7 Harding-Winnipeg remnants IO The Second Value formation I I Aleman formation 13 Cutter formation 13 FAUNAL ANALYSIS 14 I. The Montoya group 14 II. Western and northern Ordovician faunas . 15 Burnam limestone . 15 Maravillas chert 15 Arizona 15 Colorado 15 Bighorn region 15 British Columbia .. 15 Winnipeg region 16 Red River formation . 16 Stony Mountain formation . 16 Stonewall formation 16 Hudson Bay .. 16 Nelson limestone . 16 Shamattawa limestone 16 Akpatok Island 16 Baffin Island 17 Cape Calhoun region 17 Gonioceras Bay formation 18 Cape Calhoun formation 18 Scattered Arctice occurrences . 18 III. Eastern North America 18 Lake St. John, Quebec 18 Anticosti Island .......... 18 Eastern Quebec . 19 ..r Southern Ontario 19 Cincinnati arch 19 Maquoketa shale .. 19 Fernvale ... 19 Michigan Richmond . 19 Conclusions 20 CORAL MORPHOLOGY AND RELATIONSHIPS 23 General classification 23 General Morphology 23 Colony form 23 Page Corallite . 24 Gross corallite morphology . 24 Septa 24 Tabulae 25 Thecae 25 Fine structure 26 Fibrous wall .. 26 Trabecular wall . 26 Rugosan wall . 28 Tabulae ... 28 Poikiloplasm 28 Evolution and relationships . 31 Envelopes 39 Specific criteria 39 DESCRIPTION OF GENERA AND SPECIES. 42 Cateniform corals 42 Described North American species . 43 MANIPORA Sinclair 45 M. magna n. sp .. 45 M. amicarum Sinclair . 46 M. trapezoidalis n. sp ..... 47 CATENIPORA Lamarck 47 C. workmanae n. sp .. 49 C. cf. workmanae 51 C. sp. (1) . 51 C. sp. (2) 52 Bacular corals . 52 PRAGNELLIA Leith 52 P. delicatula n. sp.. 52 PROTROCHISCOLITHUS Troedsson .. 53 P. magnus (Whiteaves) 55 P. hembrilloensis n. sp .. ' . 56 P. alemanensis n. sp .. 56 COCCOSERIS Eichwald 56 C. asomata n. sp. 56 Cerioid corals with fibrous to trabecular walls. 57 SAFFORDOPHYLLUM Bassler ....... 57 S. newcombae . 60 TRABECULITES n. gen .. 61 T. keithae n. sp. .. 61 T. maculatus n. sp. 62 NYCTOPORA Nicholson . 63 N. mutabilis n. sp. .... 63 N. nondescripta n. sp. 64 N. (?) sp ..... ....... 65 CALAPOECIA Billings 65 ~.r' C. cf. anticostiensis Billings 67 C. cf. ungava Cox . 67 C. coxi Bassler . 68 Cerioid corals with axial planes 69 FOERSTEPHYLLUM Bassler. 69 F. porosum n. sp .. 70 F. minutum n. sp ..... 71 PALEOFAVOSITES Twenhofel .... 71 P. sparsus n. sp .. 73 P. prayi n. sp .. .......... 74 Page P. cf. prayi .. 74 P. kuellmeri n. sp .. 74 P. mccullochae n. sp. 75 FAVISTELLA Dana . 76 FAVISTINA n. gen. 77 F. stellata (Hall) . 79 form from Madison, Ind. 79 form from the Cutter of Tank Canyon .. 80 specimens from the Saluda of Madison, Ind. 80 specimens from the Liberty, near Bardstown, Kentucky 80 F. magister (Bassler) 81 F. crenulata n. sp .. 81 F. discreta (Foerste) .. 82 F. paleophylloides n. sp. 82 F. calicina (Nicholson) .. 82 CYATHOPHYLLOIDES Dybowski .. 83 C. burksae n. sp. 83 C. (?) sp .. 84 CRENULITES n. gen .. 84 C. duncanae n. sp .. 84 C. magnus n. sp. 84 C. rigidus (Billings) .. 85 C. blainvilli (Billings) 86 C. akpatokensis n. sp ... 86 Phaceloid corals . 87 PALEOPHYLLUM Billings. 88 P. gracile n. sp .... 89 P. margaretae n. sp .... 90 P. canteniforme n. sp. 91 P. thomi (Hall) ......... 91 SUMMARY OF THE MONTOYA CORALS .. REFERENCES PART II ORGANISMS ASSOCIATED WITH MONTOYA CORALS ABSTRACT IOI INTRODUCTION ......... IOI I. ORGANIC REMAINS ATTACHED TO COLONIAL CORALS. 105 A Capsules, closed tests and solid bodies . 105 THOLELLA n. gen. 105 T. idiotica n. sp ..... 105 IVESELLA n. gen. .......... 108 ..,t' I. adnata n. sp. 108 MOUNDIA n. gen .. ......... 108 M. fibrosa n. sp. 108 SLOCOMIA n. gen. I I I S. quadrata n. sp. l l l HARJESIA n. gen. l l l H. anomala n. sp. l l l KRUSCHEVIA n. gen. l l l K. verruca n. sp. .. I II MOOREOPSIS n. gen. II2 M. rotundus n. sp. II2 Page ELIASITES n. gen. ... 112 E. pedunculatus n. sp .. 112 CHENEYELLA n. gen. 113 C. clausa n. sp .. 113 NICCUMITES n. gen ...... 113 N. occulatus n. sp. 113 CYSTOSPHAERA n. gen. 113 C. rotunda n. sp .... 114 PEDICILLARIA n. gen. 114 P. bifurcata n. sp. I 14 B. Tube Builders 115 AMPULITES n. gen. 115 A. vasiformis n. sp. .. 115 ANCESTRULITES n. gen. 115 A. tubiformis n. sp. 115 WELLERITES n. gen. 115 W. gracilis n. sp .. 116 C. Coiled Shells or Tests . 116 W ARTHINITES n. gen. ... 117 W. adhaerens n. sp .. 117 GOLDRINGELLA n. gen. 117 G. plana n. sp. I 17 FENTONITES n. gen. I 17 F. irregularis n. sp ... 117 D. Bivalved shells .. 118 Orbiculoidea(?) sp. 118 E. Unnamed bodies ........ 118 II. OTHER TRACES OF FOREIGN ORGANISMS 119 Lichenocrinus (?) minutus .. I 19 Cavities in colonies of Coccoseris . I 19 Annellid burrows 119 REFERENCES ......... I 19 Illustrations TEXT FIGURES 1. Diagrammatic vertical section of corallite, showing relationship of skeleton with the polyp . 24 2. Variations in form of septa . .......... 25 3. Types of wall structure in simpler Ordovician corals, with inferred relationships .......... 27 4. Inferred relationships of some Ordovician coral genera, showing crucial structures ........ 33 Cross-sections showing progressive modification of fibres in the favistinids, modified in accord- 5· -·' ance with the form of the septa . ...... 6. Tholella idiotica, holotype . .. 7. Tholella idiotica paratype 8. Tholella idiotica paratype 9. Tholella idiotica paratype 10. Moundia fibrosa 110 PLATES 1-52; Montoya and related colonial corals. Attached organisms are shown mainly on pis. ~12 .................................................................. Following p. 124 PART I MONTOYA AND RELATED COLONIAL CORALS ' r Abstract Twenty-seven species are described from the Montoya faunas, particularly in western and northern North America. group of New Mexico, 17 from the Second Value formation, Revised concepts of morphology and relationship are re 6 from the Aleman, and 5 from the Cutter, 1 species being quired, notably the derivation of septa as secondary structures common to the Aleman and the Cutter. Only four species from a primitive wall, and the primitive nature of a simple have been described previously; the others are new. Four fibrous wall from which are derived trabecular walls on the species are not determined specifically. One Aleman species one hand and walls with an axial plate on the other, the axial is tentatively identified in the Cutter; others are distinct for plate developing both into the "primary wall" of the higher the three formations. Two new genera, Crenulites and Favositidae, and into the epitheca of the rugosan types. The Trabeculites, are described. Species from other regions de present study fails completely to support division of these scribed and discussed, and in some cases revised, include five corals into Tabulata and Rugosa, but it is felt that alternate species of Favistina, Protrochiscolithus magnus, and Saffordo proposals would be premature. A new structure, poikiloplasm, phyllum crenulatum, three of Crenulites, one of Trabeculites, is described, which may line corallites, be concentrated as and two assigned to Foerstephyllum. spherical knobs, with or without a thin connection, or form Regional analysis strengthens the Red River affinities of groups of discrete spheres. the Second Value formation and the Richmond equivalence A summary is appended, briefly summarizing characters of the Aleman and Cutter formations, but fuller evaluation by which the Montoya species can be differentiated readily. must await more precise study of corals of other Ordovician Introduction This is a study of the colonial corals of the Montoya group. not been suspected previously, near Morenci, Arizona. Some When the Montoya was considered as a formation, the early additional material from our collections from the typical Mon indications (Richardson, 1908) of the presence of a succession toya of New Mexico was included, but there was little reason of distinct faunas were largely ignored. With recognition in to hope that the few specimens were representative taxo the Montoya group of several formations, and with reason to nomically, and they were not particularly well preserved. believe that a considerable interval of erosion separated the The study served to emphasize the need for more informa Second Value deposits from overlying Richmond beds, the tion on the Montoya corals, and to this end a special effort need to take the stratigraphic range of various fossil groups was made in further collecting to obtain more and better into account becomes apparent at once. The material on material. which this study is based is, except for two specimens, of pre When such material was at hand, it remained to find some cisely known stratigraphic origin. one to study it. Dr. Hill was then engaged on other studies Several circumstances have combined to bring attention and could not give these specimens early attention. Further, to the corals. They are locally abundant in several horizons of colonies were large and heavy, and sending them to Queens the Montoya, the large size of the colonies making them con land for study posed some vexing problems. I next turned to spicuous constituents of the faunas. Some years ago it was Miss Helen Duncan, whose knowledge of the corals of west generally believed that Ordovician corals involved relatively ern North America is probably unequaled.