DOCSLIB.ORG

Análise Cronoestratigráfica Baseada Em Conodontes Da Formação Itaituba (Pedreira Calminas), Atokano Da Bacia Do Amazonas - Brasil

Total Page：16

File Type：pdf, Size：1020Kb

Download full-text PDF Read full-text
Análise Cronoestratigráfica Baseada Em Conodontes Da Formação Itaituba (Pedreira Calminas), Atokano Da Bacia Do Amazonas - Brasil UNIVERSIDADE FEDERAL DO RIO GRANDE DO SUL INSTITUTO DE GEOCIÊNCIAS PROGRAMA DE PÓS-GRADUAÇÃO EM GEOCIÊNCIAS ANÁLISE CRONOESTRATIGRÁFICA BASEADA EM CONODONTES DA FORMAÇÃO ITAITUBA (PEDREIRA CALMINAS), ATOKANO DA BACIA DO AMAZONAS - BRASIL CASSIANE NEGREIROS CARDOSO ORIENTADORA: DRª. VALESCA BRASIL LEMOS COMISSÃO EXAMINADORA: Dr. Claiton Marlon dos Santos Scherer Drª. Deusana Maria Costa Machado Dr. Nilo Siguehiko Matsuda Dissertação de Mestrado apresentada como requisito para obtenção do Título de Mestre em Geociências. Porto Alegre 2010 Livros Grátis http://www.livrosgratis.com.br Milhares de livros grátis para download. DEDICATÓRIA Aos meus amados pais, Luiz e Vera. Dedico a vocês mais essa conquista! AGRADECIMENTOS Aos meus pais, Luiz e Vera, por estarem sempre ao meu lado, por todo amor, incentivo e dedicação. Ao meu namorado Junior, pelo amor, cumplicidade e por ser extremamente paciente, principalmente em meus momentos de fraqueza (na finalera, uns dois, três, quatro por dia... rsrsrsrs). À Cris, mana. Por perceber que da graduação para cá, as coisas até que melhoraram. Antes, ela não dormia direito, agora, graças a mim, ela nunca se atrasa para o trabalho, mas também não dorme! A minha orientadora Valesca Brasil Lemos, pelo acolhimento, conhecimento compartilhado e oportunidade. As minhas amigas, companheiras de ralação no lab., Sara e Cris. “Sara, café!” E foram muitos cafés, mesmo! Obrigada pelas tardes agradáveis no LAPROM, regadas a calor e a poeira gostosa e, também, por manter sempre esse bom humor (realmente não sei como você consegue!). Cris, meu oráculo digital! O que seria de mim sem a sua prestatividade e parceria?! Ah, obrigada por me deixar esmagar, apertar, afagar e esmagar de novo, compulsivamente, os seus gatinhos lindos! Valeu meninas!!! À Aninha, pela amizade, dedicação, parceria, críticas, aquele perfeccionismo que dá trabalho (hehehe) e por despender, de grátis, parte de seu tempo comigo, até mesmo em um feriadão! Enfim, a todos aqueles não citados, mas que, talvez mesmo inconscientemente, com certeza contribuíram de alguma forma, seja por um abraço, um sorriso ou um simples bom dia, tornando essa jornada mais leve e mais prazerosa. “Sedimento nos olhos dos outros é refresco!” (autor desconhecido) RESUMO Conodontes são microfósseis potencialmente úteis para bioestratigrafia e em reconstruções paleoecológicas de rochas sedimentares paleozóicas e do Triássico. Desta forma, este estudo apresenta a classificação taxonômica e o estabelecimento das condições biocronoestratigráficas e paleoecológicas nas quais estes organismos viveram, bem como uma revisão do estágio atual do conhecimento a respeito dos conodontes e de seu aparelho alimentar. A seção analisada pertence à Formação Itaituba, Grupo Tapajós, Pensilvaniano da Bacia do Amazonas. A área de estudo localiza-se a Nordeste da cidade de Itaituba, às margens do Rio Tapajós, em um afloramento da Pedreira Calminas. A distribuição dos conodontes reconhecida define duas biozonas: uma Zona de Amplitude Diplognathodus orphanus e uma Zona de Concorrência Idiognathodus incurvus-Idiognathoides sinuatus. As espécies Diplognathodus coloradoensis, Diplognathodus orphanus e Idiognathodus incurvus formam uma associação tipicamente atokana. Baseando-se na associação descrita, a seção analisada foi definida como atokana. A espécie de melhor resolução bioestratigráfica é Diplognathodus orphanus, sendo sua distribuição restrita ao Atokano. O ambiente deposicional dominante na Bacia do Amazonas durante o Pensilvaniano é caracterizado por uma rampa carbonática rasa. Os depósitos estudados refletem uma sequência predominantemente regressiva, com a presença de gêneros de águas mais rasas, tais como Adetognathus, Diplognathodus e Ellisonia. A ocorrência de conodontes associados a braquiópodes, briozoários, crinóides e fragmentos de peixes sugere que estes estratos foram depositados em paleoambiente marinho raso, com energia de sedimentação baixa a moderada, em águas calmas, límpidas e quentes. No regime de flutuações de alta freqüência do nível do mar, os principais ambientes deposicionais eram laguna (submaré) e planície de maré (intermaré superior-supramaré inferior com tapetes microbiais). Palavras-chave: conodontes, bioestratigrafia, Pensilvaniano, Bacia do Amazonas. ABSTRACT Conodonts are useful microfossils for biostratigraphy and paleoecological reconstructions in Paleozoic and Triassic sedimentary rocks. This research aims to their taxonomic classification and to establish the biochronostratigraphic and paleoecological conditions in which these organisms lived. Furthermore, this study reviews the current knowledge about conodonts and their feeding apparatus. The section analyzed belongs to Itaituba Formation, Tapajos Group, Pennsylvanian of the Amazon Basin. The study area is located northeast of Itaituba city, on the banks of the Tapajós River, in an outcrop of Calminas Quarry. The distribution of conodonts recognized defined two biozones: Diplognathodus orphanus Amplitude Zone and Idiognathodus incurvus-Idiognathoides sinuatus Competition Zone. The species Diplognathodus coloradoensis, Diplognathodus orphanus and Idiognathodus incurvus are typically an Atokan association. Based on the association described, the section analyzed was defined as Atokan. The specie with better biostratigraphic resolution is Diplognathodus orphanus, because its distribution is restricted to the Atokan. The depositional environment prevailing in the Amazon Basin during the Pennsylvanian is characterized by a shallow carbonate ramp. The deposits studied reflect a predominantly regressive sequence, with the presence of shallow water genera, such as Adetognathus, Diplognathodus and Ellisonia. The occurrence of conodonts associated with brachiopods, bryozoans, crinoids and fish fragments suggests that these strata were deposited in a shallow marine environment, with low to moderate energy, in calm, clear and warm waters. In high frequency sea level fluctuations regime, the main depositional environments were lagoon (subtidal) and tidal flat (upper intertidal-lower supratidal with microbial mats). Keywords: conodonts, biostratigraphy, Pennsylvanian, Amazonas Basin. LISTA DE FIGURAS Figura 1 - Mapa de localização da Bacia do Amazonas (modificado de MILANI; THOMAZ FILHO, 2000). - 16 - Figura 2 - Mapa de localização do afloramento estudado (balão em azul, GOOGLE EARTH, 2005). - 17 - Figura 3 - Seção colunar da Pedreira Calminas. - 18 - Figura 4 - Mapa de localização das bacias intracontinentais paleozóicas da América do Sul. (MILANI; THOMAZ FILHO, 2000). - 22 - Figura 5 - Estratigrafia das bacias intracontinentais paleozóicas da América do Sul. (MILANI; ZALÁN, 1999). - 24 - Figura 6 - Representação esquemática dos principais tipos de bacias fanerozóicas interiores do Brasil (SILVA et al., 2003). - 25 - Figura 7 - Perfil Norte-Sul da Bacia do Amazonas (MILANI; THOMAZ FILHO, 2000).- 31 - Figura 8 - Representação artística do organismo conodonte (modificado de PURNELL et al., 1995). - 40 - Figura 9 - Relações dos conodontes entre si e dos conodontes, invertebrados e outros vertebrados (SWEET; DONOGHUE, 2001). - 42 - Figura 10 - Impressão fóssil de Clydagnathus windsorensis. Organismo completo, bem preservado, com 3,75 cm de comprimento (SWEET; DONOGHUE, 2001). - 43 - Figura 11 - Impressão fóssil de Promissum pulchrum (GABBOTT et al., 1995). Região cefálica posicionada para a esquerda e dorsal direcionada para o topo. - 43 - Figura 12 - Impressão fóssil de Clydagnathus windsorensis. Região cefálica, detalhando os lobos orgânicos escuros (olhos) e as estruturas associadas, incluindo uma assembléia natural de elementos conodontes (SWEET; DONOGHUE, 2001). - 44 - Figura 13 - Impressão fóssil de Clydagnathus windsorensis. Detalhe da cauda e de raios sugestivos de uma nadadeira caudal (SWEET; DONOGHUE, 2001). - 44 - Figura 14 - Impressão fóssil de Clydagnathus windsorensis. Detalhe da região do tronco apresentando duas linhas axiais paralelas, interpretadas como resquícios de uma notocorda e marcas em chevrons sobrepostos, interpretados como resquícios de músculos segmentados em forma de “V” (SWEET; DONOGHUE, 2001). - 45 - Figura 15 - Reconstrução da anatomia de Clydagnathus windsorensis (PURNELL, 1995a). - 46 - Figura 16 - Filogenia dos vertebrados primitivos, localizando os conodontes como organismos mais evoluídos do que mixinóides (peixes-bruxa) e lampréias, posicionando-os como os vertebrados mais primitivos (ARMSTRONG, 2005). - 47 - Figura 17 - Assembléia natural de elementos conodontes do gênero Idiognathodus (Ordem Ozarkodinida) preservado em rochas pensilvanianas de Illinois. As letras em preto indicam alguns dos componentes das posições P, S e M do aparelho alimentar (SWEET; DONOGHUE, 2001). - 49 - Figura 18 - Seções transversais de: (1) elemento protoconodonte. (2) elemento paraconodonte. (3) elemento euconodonte. As diferentes tonalidades refletem as diferentes camadas de tecido nos elementos. As camadas de tecido dos protoconodontes não têm homólogos nos paraconodontes e/ou nos euconodontes. No euconodonte, a tonalidade rosa representa a coroa, enquanto que a marrom representa a base, homóloga nos paraconodontes (SWEET; DONOGHUE, 2001). - 50 - Figura 19 - Morfologia de elementos coniformes (CLARK et al., 1981). - 51 - Figura 20 - Orientação e terminologia de elementos coniformes (CLARK et al., 1981). - 52 - Figura 21 - Morfologia de um elemento ramiforme Alate visto sob as perspectivas (1a) anterior e (1b) lateral (CLARK et al., 1981). - 54 - Figura 22 - Morfologia de elementos ramiformes
Load more

Recommended publications
  • CONODONTS of the MOJCZA LIMESTONE -.: Palaeontologia Polonica
    CONODONTS OF THE MOJCZA LIMESTONE JERZY DZIK Dzik, J. 1994. Conodonts of the M6jcza Limestone. -In: J. Dzik, E. Olemp ska, and A. Pisera 1994. Ordovician carbonate platform ecosystem of the Holy Cross Moun­ tains. Palaeontologia Polonica 53, 43-128. The Ordovician organodetrital limestones and marls studied in outcrops at M6jcza and Miedzygorz, Holy Cross Mts, Poland, contains a record of the evolution of local conodont faunas from the latest Arenig (Early Kundan, Lenodus variabilis Zone) to the Ashgill (Amorphognathus ordovicicus Zone), with a single larger hiatus corre­ sponding to the subzones from Eop/acognathus pseudop/anu s to E. reclinatu s. The conodont fauna is Baltic in general appearance but cold water genera , like Sagitto­ dontina, Scabbardella, and Hamarodus, as well as those of Welsh or Chinese af­ finities, like Comp/exodus, Phragmodus, and Rhodesognathu s are dominant in par­ ticular parts of the section while others common in the Baltic region, like Periodon , Eop/acognathus, and Sca/pellodus are extremely rare. Most of the lineages continue to occur throughout most of the section enabling quantitative studies on their phyletic evolut ion. Apparatuses of sixty seven species of thirty six genera are described and illustrated. Phyletic evolution of Ba/toniodus, Amorphognathu s, Comp/exodus, and Pygodus is biometrically documented. Element s of apparatu ses are homolog ized and the standard notation system is applied to all of them. Acodontidae fam. n., Drepa­ nodus kie/censis sp. n., and D. santacrucensis sp. n. are proposed . Ke y w o r d s: conodonts, Ordovici an, evolut ion, taxonomy. Jerzy Dzik, Instytut Paleobiologii PAN, A/eja Zwirk i i Wigury 93, 02-089 Warszawa , Poland.
    [Show full text]
  • Two New Early Balognathid Conodont Genera from the Ordovician Of
    Two new early balognathid conodont genera from the Ordovician of Oman and comments on the early evolution of prioniodontid conodonts Miller, Giles; Heward, Alan; Mossoni, Angelo; Sansom, Ivan DOI: 10.1080/14772019.2017.1314985 License: Other (please specify with Rights Statement) Document Version Peer reviewed version Citation for published version (Harvard): Miller, G, Heward, A, Mossoni, A & Sansom, I 2017, 'Two new early balognathid conodont genera from the Ordovician of Oman and comments on the early evolution of prioniodontid conodonts', Journal of Systematic Palaeontology, pp. 1-23. https://doi.org/10.1080/14772019.2017.1314985 Link to publication on Research at Birmingham portal Publisher Rights Statement: This is an Accepted Manuscript of an article published by Taylor & Francis in Journal of Systematic Palaeontology on 05/05/2017, available online: http://www.tandfonline.com/10.1080/14772019.2017.1314985 General rights Unless a licence is specified above, all rights (including copyright and moral rights) in this document are retained by the authors and/or the copyright holders. The express permission of the copyright holder must be obtained for any use of this material other than for purposes permitted by law. •Users may freely distribute the URL that is used to identify this publication. •Users may download and/or print one copy of the publication from the University of Birmingham research portal for the purpose of private study or non-commercial research. •User may use extracts from the document in line with the concept of ‘fair dealing’ under the Copyright, Designs and Patents Act 1988 (?) •Users may not further distribute the material nor use it for the purposes of commercial gain.
    [Show full text]
  • BIASES in INTERPRETATION of the FOSSIL RECORD of CONODONTS by MARK A
    [Special Papers in Palaeontology, 73, 2005, pp. 7–25] BETWEEN DEATH AND DATA: BIASES IN INTERPRETATION OF THE FOSSIL RECORD OF CONODONTS by MARK A. PURNELL* and PHILIP C. J. DONOGHUE *Department of Geology, University of Leicester, University Road, Leicester LE1 7RH, UK; e-mail: [email protected] Department of Earth Sciences, University of Bristol, Wills Memorial Building, Queens Road, Bristol BS8 1RJ, UK; e-mail: [email protected] Abstract: The fossil record of conodonts may be among and standing generic diversity. Analysis of epoch ⁄ stage-level the best of any group of organisms, but it is biased nonethe- data for the Ordovician–Devonian interval suggests that less. Pre- and syndepositional biases, including predation there is generally no correspondence between research effort and scavenging of carcasses, current activity, reworking and and generic diversity, and more research is required to bioturbation, cause loss, redistribution and breakage of ele- determine whether this indicates that sampling of the cono- ments. These biases may be exacerbated by the way in which dont record has reached a level of maturity where few genera rocks are collected and treated in the laboratory to extract remain to be discovered. One area of long-standing interest elements. As is the case for all fossils, intervals for which in potential biases and the conodont record concerns the there is no rock record cause inevitable gaps in the strati- pattern of recovery of different components of the skeleton graphic distribution of conodonts, and unpreserved environ- through time. We have found no evidence that the increas- ments lead to further impoverishment of the recorded ing abundance of P elements relative to S and M elements spatial and temporal distributions of taxa.
    [Show full text]
  • Late Devonian Conodont Fauna of the Gümüflali Formation
    TurkishJournalofEarthSciences (TurkishJ.EarthSci.),Vol.9, 2000,pp.69-89. Copyright©TÜB‹TAK LateDevonianConodontFaunaoftheGümüflali Formation,theEasternTaurides,Turkey fiENOLÇAPKINO⁄LU&‹SMETGED‹K KaradenizTeknikÜniversitesi,JeolojiMühendisli¤iBölümü,TR-61080Trabzon,TURKEY (e-mail:[email protected]) Abstract: TheLateDevonianGümüflaliformationoftheeasternTauridesisaterrigenous-carbonaterocksequence about600mthick,consistingmainlyofquartzsandstone,quartzsiltstone,shale,andcarbonaterocks. Palaeontologicandsedimentologicdatamainlyindicateashallowsubtidaldepositionalenvironment.Thissequence generallyrepresentstheshallow-waterpolygnathid-icriodidbiofacies,andcontainsconodontfaunasthatrange fromtheUpperfalsiovalis ZoneintotheUpperpraesulcata Zone.However,theydonotcorrelatewelltotheLate Devonianstandardconodontzonationbecauseofthelackofzonallydiagnosticspeciesandtheirregularvertical distributionsofthepresenttaxa.Herein,54taxabelongingtoninegeneraaredescribedandillustratedfromthe studiedsection.Icriodusadanaensis,Icriodusfekeensis,andPolygnathusantecompressus arethenewlydescribed species. KeyWords: LateDevonian,conodont,Gümüflaliformation,easternTaurides,Turkey. GümüflaliFormasyonu’nun(Do¤uToroslar,Türkiye)GeçDevoniyen KonodontFaunas› Özet: Do¤uToroslarboyuncayayg›nyüzeylemeleriolanGeçDevoniyenyafll›Gümüflaliformasyonu,yaklafl›k600 metrekal›nl›¤aulaflanbirk›r›nt›l›-karbonatkayadizisidir.Litolojisinibafll›cakuvarskumtafl›,kuvarsmiltafl›,fleylve karbonatkayalar›n›noluflturdu¤ububiriminpaleontolojikvesedimantolojiközellikleri,çökelmeninbafll›cas›¤,gel- gitalt›ortamdageliflti¤ineiflareteder.Konodontfaunas›genelliklek›y›-yak›n›polygnathid-icriodidbiyofasiyesini
    [Show full text]
  • Information to Users
    INFORMATION TO USERS This manuscript has been reproduced from the microfilm master. UMI films the text directly from the original or copy submitted. Thus, some thesis arxi dissertation copies are in typewriter face, while others may be from any type of computer printer. The quality of this reproduction is dependent upon the quality of the copy submitted. Broken or indistinct print, colored or poor quality illustrations and photographs, print bleedthrough, substandard margins, and improper alignment can adversely affect reproduction. In the unlikely event that the author did not send UMI a complete manuscript and there are missing pages, these will be noted. Also, if unauthorized copyright material had to be removed, a note will indicate the deletion. Oversize materials (e.g., maps, drawings, charts) are reproduced by sectioning the original, beginning at the upper left-hand comer and continuing from left to right in equal sections with small overlaps. Photographs included in the original manuscript have been reproduced xerographically in this copy. Higher quality 6" x 9" black and white photographic prints are available for any photographs or illustrations appearing In this copy for an additional charge. Contact UMI directly to order. Bell & Howell Information and Learning 300 North Zeeb Road, Ann Arbor, Ml 48106-1346 USA 800-521-0600 UMI NOTE TO USERS This reproduction is the best copy available. UMI Stratigraphy, Conodont Taxonomy and Biostratigraphy of Upper Cambrian to Lower Silurian Platform to Basin Facies, Northern British Columbia by Leanne Pyle B. Sc., University of Saskatchewan, 1994 A Dissertation Submitted in Partial Fulfillment of the Requirements for the Degree of DOCTOR OF PHILOSOPHY in the School of Earth and Ocean Sciences We accept this dissertation as conforming to the required standard , Supervisor (School of Earth and Ocean Sciences) Dr.
    [Show full text]
  • Synchrotron-Aided Reconstruction of the Conodont Feeding Apparatus and Implications for the Mouth of the first Vertebrates
    Synchrotron-aided reconstruction of the conodont feeding apparatus and implications for the mouth of the first vertebrates Nicolas Goudemanda,1, Michael J. Orchardb, Séverine Urdya, Hugo Buchera, and Paul Tafforeauc aPalaeontological Institute and Museum, University of Zurich, CH-8006 Zürich, Switzerland; bGeological Survey of Canada, Vancouver, BC, Canada V6B 5J3; and cEuropean Synchrotron Radiation Facility, 38043 Grenoble Cedex, France Edited* by A. M. Celâl Sxengör, Istanbul Technical University, Istanbul, Turkey, and approved April 14, 2011 (received for review February 1, 2011) The origin of jaws remains largely an enigma that is best addressed siderations. Despite the absence of any preserved traces of oral by studying fossil and living jawless vertebrates. Conodonts were cartilages in the rare specimens of conodonts with partly pre- eel-shaped jawless animals, whose vertebrate affinity is still dis- served soft tissue (10), we show that partial reconstruction of the puted. The geometrical analysis of exceptional three-dimensionally conodont mouth is possible through biomechanical analysis. preserved clusters of oro-pharyngeal elements of the Early Triassic Novispathodus, imaged using propagation phase-contrast X-ray Results synchrotron microtomography, suggests the presence of a pul- We recently discovered several fused clusters (rare occurrences ley-shaped lingual cartilage similar to that of extant cyclostomes of exceptional preservation where several elements of the same within the feeding apparatus of euconodonts (“true” conodonts). animal were diagenetically cemented together) of the Early This would lend strong support to their interpretation as verte- Triassic conodont Novispathodus (11). One of these specimens brates and demonstrates that the presence of such cartilage is a (Fig. 2A), found in lowermost Spathian rocks of the Tsoteng plesiomorphic condition of crown vertebrates.
    [Show full text]
  • Back Matter (PDF)
    Index Page numbers in italic denote Figures. Page numbers in bold denote Tables. Acadian Orogeny 224 Ancyrodelloides delta biozone 15 Acanthopyge Limestone 126, 128 Ancyrodelloides transitans biozone 15, 17,19 Acastella 52, 68, 69, 70 Ancyrodelloides trigonicus biozone 15, 17,19 Acastoides 52, 54 Ancyrospora 31, 32,37 Acinosporites lindlarensis 27, 30, 32, 35, 147 Anetoceras 82 Acrimeroceras 302, 313 ?Aneurospora 33 acritarchs Aneurospora minuta 148 Appalachian Basin 143, 145, 146, 147, 148–149 Angochitina 32, 36, 141, 142, 146, 147 extinction 395 annulata Events 1, 2, 291–344 Falkand Islands 29, 30, 31, 32, 33, 34, 36, 37 comparison of conodonts 327–331 late Devonian–Mississippian 443 effects on fauna 292–293 Prague Basin 137 global recognition 294–299, 343 see also Umbellasphaeridium saharicum limestone beds 3, 246, 291–292, 301, 308, 309, Acrospirifer 46, 51, 52, 73, 82 311, 321 Acrospirifer eckfeldensis 58, 59, 81, 82 conodonts 329, 331 Acrospirifer primaevus 58, 63, 72, 74–77, 81, 82 Tafilalt fauna 59, 63, 72, 74, 76, 103 ammonoid succession 302–305, 310–311 Actinodesma 52 comparison of facies 319, 321, 323, 325, 327 Actinosporites 135 conodont zonation 299–302, 310–311, 320 Acuticryphops 253, 254, 255, 256, 257, 264 Anoplia theorassensis 86 Acutimitoceras 369, 392 anoxia 2, 3–4, 171, 191–192, 191 Acutimitoceras (Stockumites) 357, 359, 366, 367, 368, Hangenberg Crisis 391, 392, 394, 401–402, 369, 372, 413 414–417, 456 agnathans 65, 71, 72, 273–286 and carbon cycle 410–413 Ahbach Formation 172 Kellwasser Events 237–239, 243, 245, 252
    [Show full text]
  • Exceptionally Preserved Conodont Apparatuses with Giant Elements from the Middle Ordovician Winneshiek Konservat-Lagerstätte, Iowa, USA
    Journal of Paleontology, 91(3), 2017, p. 493–511 Copyright © 2017, The Paleontological Society 0022-3360/16/0088-0906 doi: 10.1017/jpa.2016.155 Exceptionally preserved conodont apparatuses with giant elements from the Middle Ordovician Winneshiek Konservat-Lagerstätte, Iowa, USA Huaibao P. Liu,1 Stig M. Bergström,2 Brian J. Witzke,3 Derek E. G. Briggs,4 Robert M. McKay,1 and Annalisa Ferretti5 1Iowa Geological Survey, IIHR-Hydroscience & Engineering, University of Iowa, 340 Trowbridge Hall, Iowa City, IA 52242, USA 〈[email protected]〉; 〈[email protected]〉 2School of Earth Sciences, Division of Earth History, The Ohio State University, 125 S. Oval Mall, Columbus, Ohio 43210, USA 〈[email protected]〉 3Department of Earth and Environmental Sciences, University of Iowa, 115 Trowbridge Hall, Iowa City, IA 52242, USA 〈[email protected]〉 4Department of Geology and Geophysics, and Yale Peabody Museum of Natural History, Yale University, New Haven, CT 06520, USA 〈[email protected]〉 5Dipartimento di Scienze Chimiche e Geologiche, Università degli Studi di Modena e Reggio Emilia, via Campi 103, I-41125 Modena, Italy 〈[email protected]〉 Abstract.—Considerable numbers of exceptionally preserved conodont apparatuses with hyaline elements are present in the middle-upper Darriwilian (Middle Ordovician, Whiterockian) Winneshiek Konservat-Lagerstätte in northeastern Iowa. These fossils, which are associated with a restricted biota including other conodonts, occur in fine-grained clastic sediments deposited in a meteorite impact crater. Among these conodont apparatuses, the com- mon ones are identified as Archeognathus primus Cullison, 1938 and Iowagnathus grandis new genus new species. The 6-element apparatus of A.
    [Show full text]
  • Durham E-Theses
    Durham E-Theses Biotic recovery of conodonts following the end-Ordovician mass extinction Radclie, Gail How to cite: Radclie, Gail (1998) Biotic recovery of conodonts following the end-Ordovician mass extinction, Durham theses, Durham University. Available at Durham E-Theses Online: http://etheses.dur.ac.uk/4686/ Use policy The full-text may be used and/or reproduced, and given to third parties in any format or medium, without prior permission or charge, for personal research or study, educational, or not-for-prot purposes provided that: • a full bibliographic reference is made to the original source • a link is made to the metadata record in Durham E-Theses • the full-text is not changed in any way The full-text must not be sold in any format or medium without the formal permission of the copyright holders. Please consult the full Durham E-Theses policy for further details. Academic Support Oce, Durham University, University Oce, Old Elvet, Durham DH1 3HP e-mail: [email protected] Tel: +44 0191 334 6107 http://etheses.dur.ac.uk Biotic Recovery of Conodonts following the End-Ordovician Mass Extinction Gail Radcliffe Department of Geological Sciences University of Durham The copyright of this thesis rests with the author. No quotation from it should be published without the written consent of the author an information derived from it should be acknowledged. Volume 2 1? ..... .A thesis submitted in partial fulfilment of the degree of Doctor of Philosophy , University of Durham 1998 / Volume 2 Table of Contents Title Page i Table of Contents ii Appendix A: Systematic Palaeontology A.1 Introduction Al A.
    [Show full text]
  • Fossils, Histology, and Phylogeny: Why Conodonts Are Not Vertebrates
    234 by Alain Blieck1, Susan Turner2,3, Carole J. Burrow3, Hans-Peter Schultze4, Carl B. Rexroad5, Pierre Bultynck6 and Godfrey S. Nowlan7 Fossils, histology, and phylogeny: Why conodonts are not vertebrates 1Université Lille 1: Sciences de la Terre, FRE 3298 du CNRS «Géosystèmes», F-59655 Villeneuve d’Ascq cedex, France. E-mail: [email protected] 2Monash University, Geosciences, Box 28E, Victoria 3800, Australia 3Queensland Museum, Geosciences, 122 Gerler Road, Hendra, Queensland 4011, Australia 4Kansas University, Biodiversity Institute & Natural History Museum, 1345 Jayhawk Blvd., Lawrence, KS 66045-7593, USA 5Indiana Geological Survey, 611 North Walnut Grove, Bloomington, IN 47405-2208, USA 6Institut Royal des Sciences Naturelles de Belgique, Département de Paléontologie, Rue Vautier, 29, B-1000 Bruxelles, Belgium 7Geological Survey of Canada, 3303 – 33rd Street NW, Calgary, AB T2L 2A7, Canada The term vertebrate is generally viewed by help resolve the phylogenetic relationships of conodonts systematists in two contexts, either as Craniata and chordates, the analysis should be extended to (myxinoids or hagfishes + vertebrates s.s., i.e. basically, include non-chordate taxa. animals possessing a stiff backbone) or as Vertebrata (lampreys + other vertebrae-bearing animals, which Historical background we propose to call here Euvertebrata). Craniates are characterized by a skull; vertebrates by vertebrae A recent paper by Sweet and Cooper (2008), within the classic paper series of Episodes, drew our attention and prompted this (arcualia); euvertebrates are vertebrates with hard response. Their paper concerned the discovery and the concept of phosphatised tissues in the skeleton. The earliest conodonts by Christian Heinrich Pander (1856). He was the first known possible craniate is Myllokunmingia (syn.
    [Show full text]
  • Durham E-Theses
    Durham E-Theses The palaeobiology of the panderodontacea and selected other euconodonts Sansom, Ivan James How to cite: Sansom, Ivan James (1992) The palaeobiology of the panderodontacea and selected other euconodonts, Durham theses, Durham University. Available at Durham E-Theses Online: http://etheses.dur.ac.uk/5743/ Use policy The full-text may be used and/or reproduced, and given to third parties in any format or medium, without prior permission or charge, for personal research or study, educational, or not-for-prot purposes provided that: • a full bibliographic reference is made to the original source • a link is made to the metadata record in Durham E-Theses • the full-text is not changed in any way The full-text must not be sold in any format or medium without the formal permission of the copyright holders. Please consult the full Durham E-Theses policy for further details. Academic Support Oce, Durham University, University Oce, Old Elvet, Durham DH1 3HP e-mail: [email protected] Tel: +44 0191 334 6107 http://etheses.dur.ac.uk The copyright of this thesis rests with the author. No quotation from it should be pubHshed without his prior written consent and information derived from it should be acknowledged. THE PALAEOBIOLOGY OF THE PANDERODONTACEA AND SELECTED OTHER EUCONODONTS Ivan James Sansom, B.Sc. (Graduate Society) A thesis presented for the degree of Doctor of Philosophy in the University of Durham Department of Geological Sciences, July 1992 University of Durham. 2 DEC 1992 Contents CONTENTS CONTENTS p. i ACKNOWLEDGMENTS p. viii DECLARATION AND COPYRIGHT p.
    [Show full text]
  • Papers in Press
    Papers in Press “Papers in Press” includes peer-reviewed, accepted manuscripts of research articles, reviews, and short notes to be published in Paleontological Research. They have not yet been copy edited and/or formatted in the publication style of Paleontological Research. As soon as they are printed, they will be removed from this website. Please note they can be cited using the year of online publication and the DOI, as follows: Humblet, M. and Iryu, Y. 2014: Pleistocene coral assemblages on Irabu-jima, South Ryukyu Islands, Japan. Paleontological Research, doi: 10.2517/2014PR020. doi:10.2517/2017PR023 Missourian (Kasimovian, Late Pennsylvanian) conodonts from limestone boulders, Mizuboradani Valley, Gifu Prefecture, central Japan AcceptedTAKUMI MAEKAWA1, TOSHIFUMI KOMATSU2, GENGO TANAKA3, MARK WILLIAMS4, CHRISTOPHER P. STOCKER4, MASATOSHI OKURA5 and AKIHIRO UMAYAHARA6 1Center for Water Cycle, Marine Environment, and Disaster Management, Marine Science Laboratory, Kumamoto University, 2-39-1 Kurokami, Chuo-ku, Kumamoto 850-8555, Japan (e-mail: [email protected]) 2Faculty of Advanced Science and Technology, Kumamoto University, 2-39-1 Kurokami, Chuo-ku, Kumamoto 860-8555, Japan ([email protected]) 3Institute of Liberal Arts and Science, Kanazawamanuscript University, Kakuma, Kanazawa, Ishikawa 920-1192, Japan (e-mail: [email protected]) 4School of Geography, Geology and Environment, University of Leicester, University Road, Leicester, LEI 7RH, UK ([email protected]) 5Naka 86, Minamiyama-cho, Konan, Aichi 483-8155, Japan 6Graduate School of Science and Technology, Kumamoto University, 2-39-1 Kurokami, Chuo-ku, Kumamoto 860-8555, Japan Abstract. Two Late Pennsylvanian conodont species, Gondolella sublanceolata Gunnell and Idiognathodus sulciferus Gunnell, were extracted from limestone boulders in the Mizuboradani Valley, Fukuji district, central Japan.
    [Show full text]