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December 1986

A Catalog of the (Ephemeroptera) of Hong Kong

Michael D. Hubbard Florida A&M University, Tallahassee, Florida

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Hubbard, Michael D., "A Catalog of the Mayflies (Ephemeroptera) of Hong Kong" (1986). Insecta Mundi. 511. https://digitalcommons.unl.edu/insectamundi/511

This Article is brought to you for free and open access by the Center for Systematic Entomology, Gainesville, Florida at DigitalCommons@University of Nebraska - Lincoln. It has been accepted for inclusion in Insecta Mundi by an authorized administrator of DigitalCommons@University of Nebraska - Lincoln. Vol. 1, no. 4, December 1986 INSECTA MUNDI

Michael D. Hubbard Entomology Florida A&M University Tallahassee, Florida 32307 USA

This paper is a catalog of the Ephemerop- Order Ephemeroptera tera known from the British Crown Colony of Hong Kong. It includes a11 known references Family Baetidae Leach, 1815 for each species, along with an indication of the type localities and depositions of Subfamily Baetinae Leach, 1815 type specimens. For each reference the stage (male, female, or nymph) is indicated Leach, 1815 if relevant. Keys which include the species Baetis Leach, 1815:137. also are indicated. The recorded distribu- Type species: fuscata Linnaeus. tion for each species is listed. For each Subsequent designation by Intern. Comm. genus are given the reference to the origi- 2001. Nomenc l., 1966:209. nal description, information on the type Brachyph lebia Wes twood, 1840:25. species, and any synonyms. Type species: Ephemera fuscata Linnaeus Abbreviations of places of type deposition (misidentified as Ephemera bioculata) are as follows: (BMNH), British Museum (Na- [c.f. Intern. Comm. 2001. Nomencl., tural History), London; (Cornel l), Cornel 1 1966:209]. Monotypic. University, Ithaca, New York; (DEI), Deut- Acentrel la Bengtsson, 1912:llO. sches Entomologisches Institut, Eberswalde; Type species: Acentrella laponica Ben- (Hamburg), Zoologisches Staatsinstitut und gtsson. Monotypic. Zoologisches Museum, Hamburg; (Hsu), perso- Heterocloeon McDunnough, 1925:175. nal collection of Y.-C. Hsu [the late Prof. Type species: Centroptilrn curiosum McDun- Hsu indicated (personal communication) that nough. Origina 1 designation. because of events in China over the past few decades these specimens should be considered Baetis sp. T1 destroyed or lost]; (Utah), University of Baetis T1 Dudgeon, 1982:14. Utah, Salt Lake City. Baetis Ti Dudgeon, 1982:225. A1t hough the taxonomic know 1edge of the Baetis Ti Dudgeon, 1983:441. Hong Kong mayflies is still somewhat sket- Baetis TI Dudgeon, 1984:73. chy, there are substantial ecological stu- Baetis T1 Dudgeon, 1984:210. dies ongoing at this time. Therefore, in Distribution: Hong Kong. order to facilitate future studies and com- parisons of data, I have included in this Baetis sp. T2 catalog the "morphospecies" of Dudgeon Bae tis T2 Dudgeon, 1982:14. (1982a, b, 1983, 1984a, b). The reader Baetis T2 Dudgeon, 1982:225. should be aware that many of the taxonomic Baetis T2 Dudgeon, 1983:441. assignments of these morphospecie s are pre- Baetis T2 Dudgeon, 1984:73. liminary and actual placement will have to Baetis T2 Dudgeon, 1984:210. await future taxonomic studies. Distribution: Hong Kong. This paper is one in a continuing series Baetis sp. Tg of catalogs of the Ephemeroptera (cf. Hub- Baetis T3 Dudgeon, 1982: 14. bard & Peters 1978, Hubbard & Pescador 1978, Baetis Tq Dudgeon, 1982:225. Hubbard & Savage 1981, Hubbard 1982a,b). Baetis Tg Dudgeon, 1983:441. I thank Prof. David Dudgeon of the Univer- Baet is T3 Dudgeon, 1984:73. sity of Hong Kong for his helpful comments Baetis T3 Dudgeon, 1984:210. in the preparation of this manuscript. Distribution: Hong Kong. This paper has been partially supported by a grant from the Cooperative States Research Baetis sp. T4 Service, U. S. Department of Agriculture, to Baeti's T4 Dudgeon, 1982:225. the Agriculture Research Programs at Florida Baetis T4 Dudgeon, 1983:441. A&M University. Baetis T4 Dudgeon, 1984:73. 248 INSECTA MUNDI Vol. 1, no. 4, December 1986

Baet is Tq Dudgeon, 1984:210. Austrocloeon Barnard, 1940:616. Distribution: Hong Kong. Type species: africanur Esben- Pe tersen. Origina 1 designation. Baetis sp. T5 Baetis T5 Dudgeon, 1984:73. Cloeon harveyi Kimmins Proc loeon harvey i Kimmins, 1947:94, fig. 2, Bae tis T5 Dudgeon, 1984:ZlO. 6, 10. (male, female). Type: female, Cal- Distribution: Hong Kong. cutta, Bengal (BMNH). Procloeon harveyi Gillies, 1949:176. Baetis sp. Lg - Procloeon harveyi Kimmins, 1971:314. Baetis LC, Dudgeon, 1984:73. - Procloeon harveyi Hubbard & Peters, Baet is Lg Dudgeon, 1984:210. - 1978:ll. Distribution: Hong Kong. Procloeon barveyi - Hubbard & Srivastava, 1984:3. Baetis sp. L6 Cloeon harveyi Mu1 ler-Liebenau & Hubbard, Baetis Lg Dudgeon, 1984:73. - 1986:538. Baetis L6 Dudgeon, 1984:210. Distribution: Hong Kong, Malaysia, India. Distribution: Hong Kong. Genus Pseudocloeon Klapdlek, 1905 Baetis sp. L7 Pseudocloeon Klapdlek, 1905:105. Baetis L7 Dudgeon, 1984:73. Type species: Pseudocloeon kraepelini Kla- Baet is L7 Dudgeon, 1984:210. pslek. Monotypic. Distribution: Hong Kong. Baetiella Weno, 1931:220. Type species: Acentre 1la japonica Imani- Baetis sp. L8 shi. Original designation. Baetis Lg Dudgeon, 1984:73. Baetis Lg Dudgeon, 1984:210. Pseudocloeon sp. Ti Distribution: Hong Kong. Pseudoc loeon Ti Dudgeon, 1982: 14. Pseudoc loeon Ti Dudgeon, 1982:225. Baetis sp. cf. Tj Pseudoc loeon Ti Dudgeon, 1983:441. Baetis cf. T3 Dudgeon, 1984:73. Pseudoc loeon Ti Dudgeon, 1984:73. Baetis cf. T3 Dudgeon, 1984:210. Pseudoc loeon Ti Dudgeon, 1984:210. Distribution: Hong Kong. Distribution: Hong Kong. Sub fami 1y C loeoninae Newman, 1853 Pseudocloeon sp. Tg Pseudocloeon T2 Dudgeon, 1982:14. Genus Centroptilum Eaton, 1869 Pseudoc loeon T2 Dudgeon, 1982:225. Centroptilum Eaton, 1869:132. Pseudoc 1oeon T2 Dudgeon, 1983:441. Type species: Ephemera luteola Mu1 ler. Pseudoc loeon Tp Dudgeon, 1984: 73. Original designation. Pseudoc loeon T2 Dudgeon, 1984:ZlO. Distribution: Hong Kong. Centroptilum sp. L1 Centropti lum L1 Dudgeon, 1984:73. Pseudocloeon sp. L3 Centroptilum L1 Dudgeon, 1984:210. Pseudoc loeon Lg Dudgeon, 1984: 73. Distribution: Hong Kong. Pseudoc loeon Lj Dudgeon, 1984:210. Distribution: Hong Kong. Genus Cloeon Leach, 1815 Cloeon Leach, 1815:137. Pseudoc loeon sp. Type species: Ephemera diptera Linnaeus as Baetella [sic] Ti Dudgeon, 1983:441. Cloeon pal 1idus Leach. Monotypy. Distribution: Hong Kong. Cloe Burmeister, 1839:797. [unjustified eme- ndation of Cloeon Leach]. Type species: Ephemera dipteraLinnaeus, Family Caenidae Newman, 1853 objective synonymy. Cloeopsis Eaton, 1866: 146. Genus Caenis Stephens, 1835 Type species: Ephemera diptera Linnaeus. Caenis Stephens, 1835:60. Monotypy. Type species: Caenis macrura Stephens. Austrocloeon Barnard, 1932:217. [nomen nud- Subsequent designation by Westwood, um] . 1840:47. Neocloeon Traver, 1932:365. Oxycypha Burmeister, 1839:796. Type species: Neocloeon alamance Traver. Type species: Oxycypha luctuosa Burmeis- Original designation. ter. Subsequent designation by Jacob, Vol. 1, no. 4, December 1986 INSECTA MUNDI 249

1974:96. Epheoera (Epherera) serica Eaton Orde 1la Campion, 1923:518. Ephemera serica Eaton, 1871:75, pl. 4 fig. Type species: Caenis macrura Stephens. 12-12a. (male, female). Type: N. China Original designation. (BMNH) . Ephemera serica - MZachlan, 1875: 169. Caenis sp. Ti Ephemera serica - Eaton, 1883:73. (male, Caenis Ti Dudgeon, 1982: 14. female). Caenis Ti Dudgeon, 1982:225. Ephemera serica - Ulmer, 1926:89, fig. 82- Caenis Ti Dudgeon, 1984:73. 83. (male, female, key). Caenis Ti Dudgeon, 1984:210. Ephemera serica - Lestage, 1927:99. (key). Distribution: Hong Kong. Ephemera serica - Navis, 1930:15. Ephemera serica - Navis, 1930: 138. Caenis sp. T2 Ephemera serica - Wu, 1935:248. Caenis T2 Dudgeon, 1983:441. Ephemera serica - Ulmer, 1936:206. Caenis T2 Dudgeon, 1984:73. Ephemera serica - Hsu, 1937:438. (male, Caenis T2 Dudgeon, 1984:210. female, key). Distribution: Hong Kong. Ephemera serica - Kimmins, 1960:309, fig. 53. (male). [designation of lectotype]. Caenis sp. L2 Ephemera serica - Kimmins, 1971:320. Caenis L2 Dudgeon, 1984:73. Distribution: Hong Kong, China, Vietnam. Caenis L2 Dudgeon, 1984:210. Distribution: Hong Kong. Ephemera (subgenus incertus)

Caenis sp. Lg Ephemera sp. TI Caenis L3 Dudgeon, 1984:73. Ephemera Ti Dudgeon, 1982:225. Caenis L3 Dudgeon, 1984:210. Distribution: Hong Kong. Distribution: Hong Kong. Ephemera sp. T2 Family Latreil le, 1810 Ephemera T2 Dudgeon, 1983:441. Distribution: Hong Kong. Genus Ephemera Linnaeus, 1758 Ephemera Linnaeus, 1758:546. Ephemera sp. Type species: Ephemera vulgata Linnaeus. Ephemera sp. Dudgeon, 1984:73. Subsequent designation by Westwood, Ephemera sp. Dudgeon, 1984:210. 1836:439. Distribution: Hong Kong. Nirvius Navis, 1922:56. Type species: lirvius punctatus Navis. Original designation. Fami ly Ephemerel lidae Klapi 1ek, 1909

Subgenus Ephemera Linnaeus, 1758 Subfamily Ephemerellinae Klapslek, 1909

Ephemera (Ephemera) pulcherrima Eaton Genus Ephemerel la Walsh, 1863 Ephemera pulcherrima Eaton, 1892: 185. (fe- Ephemerel la Walsh, 1863:377. male). Type: female, Tenasserim Val ley, Type species: Ephemerel 1a excrucians Burma (BMNH). Walsh. Subsequent designation by Eaton, Ephemera pulcherrima - Ulmer, 1926:86, fig. 1868:87. 80-81. (male, key). Ephemera pulcherrima - Lestage, 1927:99. Ephemerel 1a s p. T 1 (key). Ephemerel la Ti Dudgeon, 1982:14. Ephemera pulcherrima - Wu, 1935: 248. Ephemere 1la Ti Dudgeon, 1982:225. Ephemera pulcherrima - Hsu, 1936:233, fig. Ephemere 1la Ti Dudgeon, 1983:441. 1. (female). Ephemere 1la Ti Dudgeon, 1984:73. Ephemera pulcherrima - Ulmer, 1936:204. Ephemere 1la Ti Dudgeon, 1984:210. Ephemera pulcherrima - Hafiz, 1937: 368. Distribution: Hong Kong. Ephemera pulcherrima - Hsu, 1937:433, fig. 25-26. (male, female, key). Ephemerella sp. T2 Ephemera pulcherrima - Kimmins, 1960:308. Ephemere 1la T2 Dudgeon, 1982: 14. Ephemera pulcherrima - Kimmins, 197 1:319. Ephemere 1la T2 Dudgeon, 1982:225. Ephemera pulcherrima - Hubbard & Peters, Ephemerel la T2 Dudgeon, 1984:73. 1978:17. Ephemere 1la T2 Dudgeon, 1984:210. Distribution: Hong Kong, China, Burma, In- Distribution: Hong Kong. dia. 250 INSECTA MUNDI Vol. 1, no. 4, December 1986 Ephemerella sp. L3 (Iron) sp. Ephemere 1 la L3 Dudgeon, 1984:73. Iron sp. Dudgeon, 1984:73. Ephemere 1 la L3 Dudgeon, 1984:210. Iron sp. Dudgeon, 1984:210. Distribution: Hong Kong. Distribution: Hong Kong.

Family Needham Epeorus (subgenus incertus) in Needham & Betten, 1901 Epeorus sp. Ti Sub fami 1 y Eeptageniidae Needham Epeorus Ti Dudgeon, 1982:14 in Needham & Betten, 1901 Epeorus Ti Dudgeon, 1982:225. Epeorus Ti Dudgeon, 1983:441. Genus Compsoneuria Eaton, 1881 Epeorus T1 Dudgeon, 1984:73. Compsoneuria Eaton, 1881:23. Epeorus T1 Dudgeon, 1984:210. Type species: Compsoneuria spectabilis Distribution: Hong Kong. Eaton. Origina 1 designat ion. Cosmetogenia Eaton, 1883:18, pl. 23. Epeorus sp. T2 Type species: Compsoneuria spectabilis Epeorus T2 Dudgeon, 1984:73. Eaton. Monotypy. Epeorus T2 Dudgeon, 1984:210. Distribution: Hong Kong. Corpsoneuria sp. TI Ecdyonurus Ti Dudgeon, 1982:14; Ecdyonurus Ti Dudgeon, 1982:225. Genus Heptagenia Wal sh, 1863 Ecdyonurus T1 Dudgeon, 1983:441. Eeptagenia Walsh, 1863:197. Ecdyonurus TI Dudgeon, 1984:73. Type species: Palingenia f lavescens Wal sh. Compsoneuria Ti Dudgeon, 1984:210. Subsequent designation by Eaton, 1868, Distribution: Hong Kong. Entomol. Mon. Mag. 5:90. Kageronia Matsumura, 1931:1479. Compsoneuria sp. T2 Type species: Kageronia suzukiel la Mat su- Ecdyonurus T2 Dudgeon, 1982:225. mura. Monotypic. Ecdyonurus T2 Dudgeon, 1984: 73. Eeptagenia ngi Hsu, 1936 Compsoneuria T2 Dudgeon, 1984:210. Heptagenia ngi Hsu, 1936:235, fig. 5-7. Distribution: Hong Kong. (male). Type: male, Shaukiwan, Hong Kong (Hsu). Genus Ecdyonurus Eaton, 1868 Distribution: Hong Kong. Ecdyonurus Eaton, 1868: 142. Type species: Ephemera venosa Fabricius. Genus Paegniodes Eaton, 1881 Original designation. Paegniodes Eaton, 1881:23. Ecdyurus Eaton, 1868:90. Type species: Eeptagenia cupulata Eaton. Type species: Ephemera venosa Fabricius. Or igina 1 designation. Objective synonymy. [unjustifed emendat ion of Ecdyonurus]. Paegniodes cupulatus (Eaton, 187 1) Paracinygmula Bajkova, 1975:54. Heptagenia cupulata Eaton, 1871:138, pl. 6 Type species: Paracinygeula zhilzovae Baj- fig. 14-14a. (male, female). Type: Nor- kova. Origina 1 designation. thern China (BMNH). Paegniodes cupulatus - Eaton, 1881:23. Ecdyonurus herklotsi Hsu, 1936 Heptagenia cupulata - Pa lmCn, 1884:74. Ecdyonurus herklotsi Hsu:233, fig. 2-4. Paegniodes cupulatus - Eaton, 1885:261, pl. (male). Type: male, Pokfulum [Pokfulam], 23 fig. 4, pl. 24 fig. 4. (male, female). Hong Kong (HSU). Paegniodes cupulatus - Ulmer, 1926:107, Distribution: Hong Kong. figs. 98-102. (male, female). Paegniodes cupulatus - Hsu, 1932:39. Genus Epeorus Eaton, 1881 Paegniodes fukienensis - Hsu, 1932:39, fig. Epeorus Eaton, 1881:26. 1-3. (male). Type: male, Ling Sioh, Fuken Type species: Epeorus torrentiur Eaton. Prov. China (Corne 1 1). Original designat ion. Paegniodes cupulatus - Wu, 1935:252. Paegniodes fukienensis - Wu, 1935: 252. Subgenus Iron Eaton, 1883 Paegniodes cupulata - Hsu, 1936:236, fig. 8- Iron Eaton, 1883: pl. 24. 13. (male, female). Type species: Epeorus longimanus Ea ton. Paegniodes cupulatus - Ulmer, 1936:211. Monotypy. (male). Vol. 1, no. 4, December 1986 INSECTA MUNDI 25 1 Paegniodes cupulatus - Kimmins, 1960:283. Type: River Shing Man [Shing Hun River], [designation of lectotype]. near Kowloon, Hong Kong (BHNH). Paegniodes cupulatus - Ugno, 1969:228, fig. Cryptopene 1 la f acialis - peters & Edmunds, 26-36. (male). 1970:202, fig. 46-47, 93-94, 122, 145, Eeptagenia cupulata - Kimrnins, 1971:312. 336-345. (male, female, nymph). Paegniodes cupulatus - Tshernova, 1976:343, Crppt~Penella facialis - Kimmins, 1971:313. fig. 41. (nymph). Distribution: Hong Kong. Paegniodes fukiensis [sic] - Tshernova, 1976:343. Genus Eabrophlebiodes Ulmer, 1920 Distribution: Hong Kong, China, Tibet. ~abrophlebiodesU lmer, 1920:39. Type species: Habrophlebia americana Genus Thalerosphyrus Eaton, 1881 Banks. Original designat ion. Thalerosphyrus Eaton, 1881:22. Type species: Baetis determinatus Walker. Habrophlebiodes gilliesi Peters, 1963 Original designation. Habrophlebiodes gilliesi Peters, 1963:41, Thalerosphyrus sp. TI fig. 1-11. (male, female). Type: male, Fan Thalerosphprus TI Dudgeon, 1982: 14. Ling near Kowloon, Hong Kong (Utah). Thalerosphyrus TI Dudgeon, 1983:441. Habrophlebiodes gi1 liesi - Peters & Edmunds, Thalerosphyrus Ti Dudgeon, 1984:73. 1970:187, fig. 19-21, 77, 112, 135, 323. Thalerosphprus Ti Dudgeon, 1984:210. (male, female). Distribution: Hong Kong. Habrophlebiodes gilliesi - Tsui & Peters, 1970:89, fig. 1-13. (nymph). Fami 1y ~eptophlebiidae Banks, 1900 Eabrophlebiodes gilliesi - Dudgeon, 1982:225. Subfamily Atal~~hlebiinaePeters, 1980 Distribution: Hong Kong.

Genus Choroterpes Eaton, 1881 Genus Isca Gillies, 1951 Choroterpes Eaton, 1881:194. Isca Gillies, 1951:127. Type species: Choroterpes lusitanica Ea- Type species: Isca purpurea Gi 1lies. Ori- ton. Origina 1 designation. gina 1 designat ion.

Subgenus Euthraulus Barnard, 1932 Subgenus Isca Gi1 lies, 1951 Euthraulus Barnard, 1932:249. Type species: Euthrau lus e legans Barnard. Isca (Isca) purpurea Gi11 ies, 1951 Monotypy. Isca purpurea Gi1 lies, 1951:128, fig. 15-22. Thraululus Ulmer, 1939:499. (male, female). Types: male, female, Hong Type species: Thraulus marginatus Ulmer. Kong, 15-20 mi inland from Kowloon (BMNH). Original designation. Isca (Isca) purpurea - Peters & Edmunds, 1970:215, fig. 71, 106, 330, 350-357. Choroterpes (Euthrau lus) sp. (male, nymph). Choroterpes (Euthraulus) sp. Dudgeon, Isca purpurea - Kimrnins, 1971:319. 1984 :73. Isca (Isca) purpurea - Hubbard & Peters, Choroterpes (Euthraulus) sp. Dudgeon, 1978:24. 1984:210. Distribution: Hong Kong, India. Distribution: Hong Kong. Isca (subgenus incertus) Choroterpes (subgenus incertus) Isca sp. T1 Choroterpes sp. Ti Isca T1 Dudgeon, 1982:225. Choroterpes T1 Dudgeon, 1982:14. Isca T1 Dudgeon, 1983:441. Choroterpes T1 Dudgeon, 1983:441. Distribution: Hong Kong. Distribution: Hong Kong. Isca sp. Genus Cryptopenel la Gi11 ies, 1951 Isca sp. Dudgeon, 1984:73. Cryptopenella Gillies, 1951:125. Isca sp. Dudgeon, 1984:210. Type species: Cryptopenella facialis Gil- Distribution: Hong Kong. 1ies. Origina 1 designation. Genus Thraulus Eaton, 1881 Cryptopenella facialis Gillies, 1951 Thraulus Eaton, 1881:195. Cryptopenella facialis Gillies, 1951:127, Type species: Thraulus be1lus Eaton. Ori- fig. 11-12, 14. (male, female, nymph). gina 1 designat ion. INSECTA MUNDI Vol. 1, no. 4, December 1986

------Masharikella Peters, Gi11 ies & ~dmunds, Archiv fur Hydrobiologie Supplement 69:55- 1964:118. 129. Type species: Hagenulus fasciatus Kimmins. Dudgeon, D. 1984b. Longi tudina 1 and tempo- Origina 1 designation. ral changes in functional organization of macroinvertebrate communities in the Lam Thraulus sp. T1 Tsuen River, Hong Kong. Hydrobiologia Thraulus Ti Dudgeon, 1982:225. 111:207-217. Thraulus T1 Dudgeon, 1983:441. Eaton, A. E. 1866. Notes on some species Distribution: Hong Kong. of the orthopterous genus Cloeon, Leach (as limited by M. Pictet). Annals and LITERATURE CITED Magazine of Natural History (3)18:145-148. Eaton, A. E. 1868a. Remarks upon the homo- Bajkova, 0. Ya. 1975. [New genus of logies of the ovipositor. Transactions of from Primoriya.] (in Russian). Izvestia the Entomological Society 1868:141- 144. Sibirskogo Otde leniya Akademii Nauk SSSR Eaton, A. E. 1868b. An outline of a re- l(5) :54-57. arrangement of the genera of Ephemeridae. Banks, N. 1900. New genera and species of Entomologist's Monthly Magazine 5:82-91. Nearctic neuropteroid . Transac- Eaton, A. E. 1869. On Centroptilm, a new tions of the American Entomological Socie- genus of the Ephemeridae. Entomologist's ty 26:239-259. Monthly Magazine 6:131-132. Barnard, K. H. 1932. South African May- Eaton, A. E. 1871. A monograph on the flies (Ephemeroptera). Transactions of Ephemeridae. Transactions of the Entomo- the' Royal Society of South Africa 20:201- logical Society 1871:l-164, 6 pl. 259. Eaton, A. E. 1881a. An announcement of new Barnard, K. H. 1940. Additional records, genera of the Ephemeridae. Entomologist's and descriptions of new species, of South Monthly Magazine 17:191-197. African alder-flies (Megaloptera), May- Eaton, A. E. 1881b. An announcement of new flies (Ephemeroptera), caddis- f 1ies (Tri- genera of the Ephemeridae. Entomologist's choptera), stone-f 1ies (Per laria), and Monthly Magazine 18:21-27. dragon-f lies (Odonata). Annals of the Eaton, A. E. 1883-1885. A revisional mono- South African Museum 32:609-661. graph of recent Ephemeridae or mayflies. Bengtsson, S. 1912. Neue Ephemeriden aus Transactions of the Linnean Society of Schweden. Entomo logisk Tidskrift 33: 107- London, Second Series, Zoology 3:l-352, 65 117. PI Burks, B. D. 1953. The mayflies, or Ephe- Eaton, A. E. 1892. New species of Epheme- meroptera, of Illinois. Bulletin of the ridae from the Tenasserim Val ley. Transa- I1 1inois Natural History Survey 26:l-216. ctions of the Entomological Society of Burmeister, H. 1839. Ephemerina. Handbuch London 1892:185- 190. der Entomologie 2(2):788-804, 1014. Gillies, M. T. 1949. Notes on some Epheme- Campion, H. 1923. On the use of the gene- roptera BaZtidae from India and South-East ric name Brachycercus in Plectoptera and Asia. Transactions of the Royal Entomolo- Orthoptera. Annals and Magazine of Natu- gical Society London 100:161-177. ral History (911 1:515-518. Gillies, M. T. 1951. Further notes on Dudgeon, D. 1982a. Aspects of the microdi- Ephemeroptera from India and South East stribution of macrobenthos in a Asia. Proceedings of the Royal Entomolo- forest stream in Hong Kong. Archiv fur gical Society of London (B)20:121-130. Hydrobiologie Supplement 64:22 1-239. Hafiz, H. A. 1937. The Indian Ephemeropte- Dudgeon, D. 1982b. An investigation of ra (mayf 1ies) of the sub-order Ephemeroi- physical and biological processing of two dea. Records of the Indian Museum 39:351- species of leaf litter in Tai Po Kau Fo- 370, pl. 4-5. rest Stream, New Territories, Hong Kong. Hsu, Y.-C. 1932. Two new species of may- Archiv fur Hydrobiologie 96:l-32. flies from China (Order ~~hemerida).Pe- Dudgeon, D. 1983. An investigation of the king Natural History Bulletin 6:39-41, 1 drift of aquatic insects in Tai Po Kau PI Forest Stream, New Territories, Hong Kong. Hsu, Y.-C. 1936. Mayflies of Hong Kong Archiv fur Hydrobio logie 96:434-447. with descriptions of two new species (Ep- Dudgeon, D. 1984a. Seasona 1 and long- term hemeroptera). Hong Kong ~aturalist7:233- changes in the hydrobiology of the Lam 238. Tsuen River, New Territories, Hong Kong, Hsu, Y.-C. 1937-8. The mayf 1ies of China. with special reference to benthic macroin- Peking Natural History ~ulletin(1937) vertebrate distribution and abundance. 11:129-148, 287-296, 433-440; (1938) Vol. 1, no. 4, December 1986 INSECTA MUNDI 253 12:53-56, 125-126, 221-224. Latrei 1 le, P. A. 1810. Considerations Hubbard, M. D. 1982a. CatBlogo abreviado genet-ales sur 1'ordre nature 1 des animaux de Ephemeroptera da America do Sul. Pa- composant les classes des Crustaces, des p6is Avulsos de Zoologia 34:257-282. Arachnides et des Insectes. Chez I. Hubbard, M. D. 1982b. Catalog of the Ephe- Schoel 1, Paris. 444pp. meroptera: f ami ly-group taxa. Aquatic Leach, W. E. 1815. Entomology. Brewster's Insects 4:49-53. Edinburgh Encyclopaedia 9:57-172. Hubbard, M. D., and M. L. Pescador. 1978. Lestage, J. A. 1927. Une Ephemera nouvel le A catalog of the Ephemeroptera of the du Tonkin et tableau des esp&ces de la Phi 1ippines. Pacific Insects 19:91-99. faune orientale. Annales de la Soci4te Hubbard, M. D., and W. L. Peters. 1978. A Ei~tomologique de France 96:93- 100. catalogue of the Ephemeroptera of the Linnaeus, C. 1758. Systema Naturae. Ed. X. Indian Subregion. Orienta 1 Insects Sup- Holmae. Vol. 1 plement 9:l-43. M'Lachlan, R. 1875. A sketch of our pre- Hubbard, M. D., and W. L. Peters. 1984. sent knowledge of the Neuropterous fauna Ephemeroptera of Sri Lanka: an introduc- of Japan (excluding Odonata and Trichopte- tion to their ecology and biogeography. ra). Transactions of the Entomological Pages 257-274 in C. H. Fernando (ed.), Society 1875:167-190. Ecology and Biogeography in Sri Lanka. Dr. McDunnough, J. 1925. New Canadian Epheme- W. Junk Publishers, The Hague. ridae with notes. 111. Canadian Entomolo- Hubbard, M. D., and H. M. Savage. 1981. gist 57:168-176. The fossi 1 Leptophlebiidae (Ephemeropte- Matsumura, S. 1931. [6,000 I1 lustrated ra): a systematic and phy logenetic review. Insects of the Japanese Empire.] (in Japa- Journal of Paleontology 55:810-813. nese). Tokyo. Hubbard, M. D., and V. D. Srivastava. 1984. Miiller-Liebenau, I., and M. D. Hubbard. Ephemeroptera type-specimens in the Zoolo- 1986. Baetidae from Sri Lanka with some gical Survey of India, Calcutta. Oriental general remarks on the Baetidae of the Insects 18:l-4. Oriental Region (Insects: Ephemeroptera). International Commission on Zoological Nome- Florida Entomologist 68:537-561. nc lature. 1966. Opinion 787. Bul letin of Zoological Nomenclature 23:209-210. Navbs, L. 1922. Efemer6pteros nuevos o Jacob, U. 1974. Zur Kenntnis zweier Oxycy- poco conocidos. Boletin de la Sociedad pba-Arten Hermann Bumeisters (Ephemerop- Entomol6gica de Espana 5:54-63. tera, Caenidae). Reichenbachia 15:93-97. Navis, L. 1930a. Insectos del Museo de Kimmins, D. E. 1947. New species of Indian Paris. 5a S6rie. Brot6ria (Serie Zoologi- Ephemeroptera. Proceedings of the Roya 1 ca) 24:5-24. Entomological Society of London (B)16:92- Navds, L. 1930b. Insectos del Museo de 100. Paris. 6a S6rie. Brot4ria (Serie Zdologi- Kimmins, D. E. 1960. The Ephemeroptera ca) 24: 120-144. types of species described by A. E. Eaton, Needham, J. G., and C. Betten. 1901. Aqua- R. McLachlan and F. Walker, with particu- tic insects in the Adirondacks. Bulletin lar reference to those in the British of the New York State Museum 47:384-612, Museum (Natural History). Bulletin of the 36 pl. British Museum (Natural ~istory)Entomolo- Newman, E. 1853. Proposed division of gy 9:269-318. Neuroptera into two classes. The Zoolo- Kimmins, D. E. 1971. A list of the type- gist 1 l(appendix):181-204. specimens of Ephemeroptera in the British Palmen, J. A. 1884. ijber paarige Ausfihru- Museum (Natural ist tor^). Bu 1letin of the ngsgange der Gesch lectsorgane bei Insec- British Museum (~atural is tor^) Entomolo- ten. Eine morphologische Untersuchung. gy 25:307-324. Dissertation, Helsingfors. 108pp, 5 pl. Klapilek, F. 1905. Plecopteren und Epheme- Peters, W. L. 1963. A new species of Hab- riden aus Java, gesammel t von Prof. K. rophlebiodes from China (Ephemeroptera: Kraepelin 1904. Mittei lungen aus dem Leptophlebiidae). Proceedings of the Ro- Naturhistorischen Museum in Hamburg yal Entomologica 1 Society of London 22:103-107. (B)32:41-43. Klapilek, F. 1909. Ephemerida, Eintag- Peters, W. L. 1980. Phylogeny of the Lep- sf 1iegen. Siisswasserfauna Deutschlands tophlebiidae (Ephemeroptera): an introduc- 8:l-32. tion. Pages 33-31 in J. F. Flannagan and Klyuchareva, 0. A. 1963. [On downstream K. E. Marshall, eds. Advances in Ephemero- and diurnal vertical migrations of benthic ptera Biology. Plenum Press, New York. invertebrates in the Amur.] (in Russian). Peters, W. L., and G. F. Edmunds, Jr. 1970. Zoologicheskii Zhurnal 42:1601-1612. Revision of the generic classification of INSECTA MUNDI Vol. 1, no. 4, December 1986

- ---- the Eastern Hemisphere ~epto~hlebiidae Ueno, M. 1969. Mayflies (Ephemeroptera) (~phemeroptera). Pacific Insects 12:157- from various regions of Southeast Asia. 240. Oriental Insects 3:221-238. Peters, W. L., M. T. Gillies, and G. F. Ulmer, G. 1920. Neue Ephemeropteren. Ar- Edmunds, Jr. 1964. Two new genera of chiv fiir Naturgeschichte (A)85(11):1-80. mayflies from the Ethiopian and Oriental Ulmer, G. 1926. Trichopteren und Ephemero- Regions (Ephemeroptera: Leptophl ebiidae). pteren. In: Beitrage zur Fauna sinaica. Proceedings of the Royal Entomological Archiv fiir Naturgeschichte (A)91(5):19- Society of London (B)33:117-124. 110. Stephens, J. F. 1835. Illustrations of Ulmer, G. 1936. Neue chinesische Ephemero- British Entomology, Mandibulata 6:53-70, pteren, nebst iibersicht iiber die bischer pl. 29. aus China bekannten Arten. Peking Natural Traver, J. R. 1932. Neocloeon, a new may- History Bulletin 10:201-215. fly genus (E~hemerida). Journal of the Ulmer, G. 1939-1940. Eintagf liegen (Ephe- New York Entomological Society 40:365-372, meropteren) von den Sunda-Inseln. Archiv 1 pl. fUr Hydrobiologie Supplement 16:443-692. Tshernova, 0. A. 1976. [A key to the lar- Walsh, B. D. 1863. Observations on certain vae of genera of Heptageniidae (Ephemerop- N. A. Neuroptera. Proceedings of the tera) from the Holarctic and Oriental Entomological Society of Philadelphia Regions.] (in Russian). Entomologicheskoe 2:167-272. Obozrenie 55:332-346. Westwood, J. 0. 1836. Ephemera (Linnaeus). Tsui, P. T. P., and W. L. Peters. 1970. Partington's British Cyclopaedia of Natu- The ' nymph of Habroph 1ebiodes gi 11ies i ral History 2:438-439. Peters (Ephemeroptera: Leptophlebiidae). Westwood, J. 0. 1840. An introduction to Proceedings of the Roya 1 Entomological the modern classification of insects. Society of London 45:89-90. Longman, Orme, Brown, Green, and Longmans, Ugno, M. 1931. Contributions to the know- London. 2 vol. ledge of Japanese Ephemeroptera. Annota- Wu, C. F. 1935. Order VII. Ephemeroptera. tiones Zoologicae Japoneses 13: 189-226, Cata logus Insectorum Sinensium 1:247-253. pl. 12-13.