Journal of Geosciences ，Osaka City University Vo l. 17 ，Art. 3 ，p. 73 - 86. March ， 1974

Khu 'merspira ，a .new .Genus ' of Bellerophontidae ，and some Middle Permian Gastropods from . Cambodia*

(Contributions (Contributions to the Geology and Palaeontology of Cambodia ， Part 4)

Ken-ichi Ken-ichi ISHU** and Masafumi MURATA***

(With 2 Text-figures and 2 Plates)

Introduction Introduction and Acknowledgments

The Permian limestone in west Cambodia is well known for its yield of a rich fauna of marine invertebra もes. The stratigraphical and paleontological studies of the Permian limestone limestone have been published by DEPRAT (1912 ，1913) ，MANSUY (1913 ，1914) ，GUBLER (1935a ，b) ，DELPEY (1941-1942) ，SKINNER and WILDE (1954) ，SAURIN (1959) ，VIEN (1959) ， CHI-THUAN (1961) ，FONTAINE (1961 ，1965 ，1967) ，SERRA (1966) ， ISHII and NOGAMI (1964) ， ISHII ISHII (1966) ，and by ISHII ，KATO and NAKAMURA (1969). A small ooUection of gastropods including a pecu 1i ar form of the Be Il erophontidae ， was co Il ected from the upper part of the Sisophon Limestone exposed at Phn. **** Bak and Phn. Tup ，west of Sisophon ，Cambodia by the senior author in 1967. The majority of the specimens coUected from the north of Phn. Tup are poorly preserved. However ， nearly nearly a dozen of well preserved bellerophontids were coUected from the east of the Phn. Bak area. As the r: esult of a paleontological study ， the junior author describes six species of gastropods distributed among six genera ，among which one species is described as new to to science ，and one new generic name Khun 'l erspira is proposed. The writers here offer their cordial thanks to Dr. Kotora HATAI ， Professor Emeritus of the Tohoku University ，and Dr. Ichiro HAYASAKA of the Tamagawa University ， for their kind advice on the paleontological study. The writers also express their siDcere thanks to to Professor Koichiro ICHIKAWA ，Osaka City University ， for his kind encouragement and advice advice during this study. Thanks are due to Dr. Makoto KATO ，Hokkaido University for his his co l1 aboration in the field work in 1967.

Stratigraphical Stratigraphical note of the Sisophon Limestone at the foss iI locality

The Lower and Middle Permian Sisophon Limestone is developed typically in the west west of Sisophon ，forming limestone monadnocks ，and was subdivided lithologically by ISHII ISHII (1966) and ISHII ，KATO and NAKAMURA (1969) ， into the four members of A ，s ，C and D in ascending order. Member A consists of pale green to reddish brown andesitic tuff and tuff-breccia over

* Contribution from the Department of Geosciences ，No. 271.

料 Department of Geosciences ，Osaka City University ，Osaka ， Japan. *** *** Institute of Geology and Paleontology ，Tohoku University ，Sendai ， Japan. 料** Abbreviation for Phnom ， meaning mountain in Cambodian language. 74 74 Ken ・ichi ISH1 Il and Masafumi MURAT J¥

N

• i. ・ll.aa: ; I'~ ν 1，.市 ) : Phum pongro

口 Atluvium 1 1t tIt' 田町ember ;・ 一目 Sisophon l limestone

A..member

to to Battambang

t>: t>:

hn. P hn. Lang K . Tom 内 n kvg vu 申 B'

。 C' tI 23 km

Fig. Fig. 1. Geologic map of the Sisophon region ， west Cambodia. (after ISHII ， KATO & NAKAMURA ，1969) ，X ; foss i1 1ocality of gastropods five five meters in thickness. Small1enses of Hmestone are often intercalated in the upper part 0 of the member. This member corresponds to “niveaux 1 " of FONTAINE (1967) in the Phn. Svai area. Several species ' of fusulinids including Pseudodoliolina pseudolepida (DEPRAT) and Pseudodo/. dunbari (GUBLER) ， nine species of corals and eight species of brachiopods were listed by ISH lI， KATO and NAKAMURA (1969) from the Member A. Their distribution distribution charts of the mentioned fossils show that most of the species of the fusulinids and brachiopods range up to a higher horizon. Member B is an alternation of dark gray to milky white ，massive limestone and bedded limestone ，measuring about 40 meters in thickness. The limestone of this member is com- posed of skeletal-micrite and -sparite of fragmental crinoids ， calcareous algae ，corals ， brachiopods and fusulinids. The upper part of the Member B consists of brecciated lime- 0 0 stone stone and stromatolites. FONTAINE'S “niveaux 2 _ 4 " corresponds to Member B in this paper. paper. The Member B yielded a rich fauna of marine invertebrates. ISHII ，KATO and NAKAM .URA (1969) listed several tens of species of fusulinids ， corals and brachiopods from Permian Permian gastropods from Cambodia 75

the the Member B ，and showed that Sumatrina annae longissima DEPRAT ， Colania asiatica (ISHII) ， C. douvillei (OZAWA) ，Neoschwagerina aff. margaritae DEPRAT and Verbeekina sp. occurred occurred from the uppermost part of this member. Member C consists of reddish or brownish calcareous mudstone ， varying in thickness. J. n the Sisophon area ， the thickness of this member attains 60 meters ，but it thins toward the the south to only eight meters at Phn. Takream of the Battambang area. This member may correspond to “ niveaux 50_70" of SAURIN (1959) and “ niveaux 50 and 60" of FONTAINE (1969). (1969). The lower part of this member is considered to be partia l1 y synchronous with the upper part of Member B. Many species of corals and brachiopods were listed from the upper part of this member by ISHII ，KATO and NAKAMURA (1969) ，but the fusulinids were rather rather less numerous than that of Member D. Lepidolina .multiseptata multiseptata (DEPRAT) ， Sumatrina annae longissima DEPRAT and Pseud o. ルsulina cfr. crassa padangensis (LANGE) were reported fvom the top part of this member. Member ID attains 100 meters in maximum thickness. The lower and middle parts of this member consist of gray to dark gray bedded limestone forming a unit ranging from several several tens of centimeters to several meters in thickness. The upper part of the rnember becomes muddy and is intercalated with many layers of siliceous nodules. The Member D yielded a rich fauna of fusulinids and cora 1s. The fusulinid fauna of Member D includes Lepidolina Lepidolina multiseptata multiseptata (DEPRAT) (including Lep. elongata (DEPRAT)*) ， Lep. multiseptata multiseptata gigantea (GUBLER) ， Verbeekina verbeeki (GEINITZ) ， Chusenella cambodgensis (GUBLER) and Chu. globularis (GUBLER). The gastropods described in this article are from the the lower and middle parts of this memlber. Biostratigraphical Biostratigraphical .studies of the Sisophon Limestone have been reported by many

authors. authors. An d recently ，ISHII ，KATO and NAKAMURA (1969) subdivided the Sisophon Lime 四 stone stone biostratigraphically on the basis of the fusulinids ， corals and brachiopods as shown in in Fig. 2.

Paleontological Paleontological Note

The gastropods described in the present article are from two localities ， located about one kilometer east of Ph l1. Bak and nearly 800 meters north of Phn. Tup ， in the west of Sisophon ，Cambodia. The former belongs to the lower part of Member D ， while the latter belongs belongs to the middte part of Member D of the Sisophon Limestone. The gastropods ， treated in this article ， are as follows: ー from the Phn. Bak area Khumerspira ishii MURATA ， n. gen. n. sp. Bellerophon Bellerophon (Bellerophon?) aff. regularis (W AAGEN) from the Phn. Tup area Straparollus Straparollus (Euon ψ halus?) sp. Worthenia Worthenia cf. schirjaevensis (STUCKENBERG) Palaeostylus Palaeostylus (Leptozyga?) sp. λ feekospira sp. Most of the specimens from Phn. Tup area could not be identified to species level because of the unfavorable preservation. However ， the specirnens show much resemblance with

microspheric * microspheric form of Lep. mu!tiseptata mu!tiseptata. ，

σ"3

M !i.dd I.e， Permian tJ pp e r Permian Age

5 十 4 ，0 :l: 8-- 60 . ma X'. 100 Thickness (m ) 主〉 ∞ (') 。 Member 、;' • • 1・ . 的 • ， IV 1' -1' ..__ lE '己 11 ' • 』ー • 一 • • トー .1- . • トー t> トー r-トーーー H 1-一一一一ー 一 11 11 11 、〉/ 』ー 。 ol~lûl~lol~lol~' l o l~lol~lo l b 一 .R 一 一 11 ヤー F 、、 ， 同ー • トー • 一 • 』ー • '--， . 同園ー . 一 • ~Iõl~ løl~lòlo lò 自u 盟刊~戸ト一戸ーー ー ードー- 11 ー"一 11 、、' t> 一 一 11 一 一 トー・ / 、 一ー 同ー『 トー • トー • トー -一 • N ol~lèl~lol~ID) I~ Iò I~lõ~ 1 H H r- トー I--i t-- 一 ー-ー』 11 11 I!... i1th 0 I0 9Y 、、 J • • ~ トーl ・1. 円 11 11 • 同・ ー胴 ;lolõl~lõl!rI U I VoV!' j! iH HH 'H I一一ー -ー 一一 一 一 HII ー 一 ~ 一 • トー • 'ーー・ • 、 • • Itf 自 六 l トートート一一ーー 一 一 11 1Il:ll 11 111 11 ロ ↑ ? .圃圃. ' :;:回ー. - 穴 S annae longissima 一- Y osiatica Zone g 'WYF 向eudo 似 'iolino pseudolepid σ 、v Sumafrina a(1 nae longissima ， Zone Psf. Psf. Zo Yabeino mυ Itiseptata Zone lFusulinid ambiqua ambiqua pursatensis ne Yabeina multiseptata Zone 同 .- . 出回二口ω Tyloplecto Tyloplecto nankingensis Zone Morginifera him aJ ayensis Zone Permophricodo thyris grandis Zone Brachiopod Zone

ーーーー ~ 仏 POlythecalis POlythecalis regularis Zone Euryphyllum 。IIoiteaui Zone Parawentzelella s; sophonensis Zone c 0 ra I Zone 玄一 Jト • g 仰向己 Cyathoxonia Cyathoxonia sp. no 収 Zone SF ー まミ，、』 -J--《 -n-- 豆 HHWAHunvhHVAU ¥ ¥/¥/¥/ AUAU / ¥/¥/ C vAU ¥/¥/ 河〉吋〉 '

AHV ー 。、 ∞ 吋 01 ふ む3 N -・・・・・

Fig.2. Fig.2. Generalized columnar section of the Per .F nian of west Cambodia. (after li SHII ，KATO & NAKA IMURA ， 1969) 1.: 1.: bedded limestone ， 2: muddy limestone or micritic limestone ， 3: crinoidal limestone ， 4: limestone breccias ， 5: 5: tuff bre ∞ ias 01 tuff ， 6: reddish shale ， 7: reddish brownish calcareous mudstone with calcareous nodules ， 8: 8: drusy coating skeletallimestone (grainstone). 1: 1: Khumerspira ishii ，Bellerophon (Bellerophon?) aff. regularis ， 11: Straparollus (Euomphalus?) sp. ，WOl 幼 enia cf. schirjaevensis ，Palaeostylus (Leptozyga?) sp. ， λfeekospira sp. Notes: Notes: For Y. asiatica read Colania asiatica and for Yabeina multiseptata read Lepidolina multiseptata in this paper. paper. 、 Permian Permian gastropods jroln Cambodia 77 the the gastropod fauna described from Calcaires a Productus in Indochine by MANSUY (1913 ， 1914) 1914) and DELPEY (1941-1942). On the other hand ， nearly a dozen of be l1 erophontids from ，PhD '. Bak area are relatively well preserved ，and are additional to the Middle Permian gastropods gastropods of Cambodia.

Systematic Systematic Description of the Gastropoda by Masafumi MURATA

Order A 問 baeogastropoda THIELE ， 1825 Suborder Bellerophontina ULRICH and SCOFIELD ， 1897 Superfam i1 y Bellerophontacea M'CoY ， 1851 Fam i1 y Bellerophontidae M'CoY ， 1851 • Subfamily Subfamily Bellerophontinae M'CoY ， 1851 Genus Bellerophon MONTFORT ， 1808 Subgenus Bellerophon (Bellerophon) MONTFORT ， 1808 Bellerophon Bellerophon (Belterophon?) aff. regularis (W AAGEN) ，1880 Plate Plate 2，figs. 9-11.

Mogulia regularis WAAGEN ，1880 ， p. 157 ， p1. 13 ，figs. 4a-e. Mogulia regularis W AAGBN ，K Nl GHT ，1941 ， p. 200 ， p1. 8，figs. 5a ， b.

Description Description :-Moderately well-rounded ， bellerophontid gastropods having an obscure selenizone selenizone and rounded transverse costae of shell surface; shell medium in size ， well rounded except except for slightly expanded and rather strongly reflected laterallips; sinus and slit unknown; selenizone selenizone rather narrow ， very obscured ， flush with surface in early stages and slightly depressed depressed at maturity; ornamented with numerous ，distinct ，transverse costae with narrow interspaces. interspaces. The measurements of Bellerophon (Bellerophon 7) aff. regularis (W AAGEN) are given in Table 1. Remarks:-Two fragmental specill1ens ，one of which is immature ， are at hand. The present form seems to be closely related to Mogulia regularis W AAGEN ，specially in the immature stages ， but it becomes much larger in size in the mature stage. W AAGEN (1880) distinguished the the genus Mogulia from Bellerophon by the former lacking a proper slit-band. However ， KNIGHT (1941) showed the presence of an obscure selenizone as the result of a re-exa mI na- tion tion of the type specimen of Mogulia WAAGEN. An d he (1944) placed Mogulia WAAGEN in in the synonymy of Bellerophon. With regard to the ontogenical changes of the selenizone of the present form ， the writer agrees with KNIGHT'S (1944) opinion ， but has some questions about the subgeneric position of the specimens. Repository:- OMNH*. E301 ， E302. Occurrence:- Rather rare in the upper part of the Sisophon Limestone (1 ower part of Member D ， Lepidolina mulfisepfata Zone) on the narrow path in the east of Pbn. Bak ， Middle Permian.

* Osaka Museum of Natural History.

， '8 '8 Ken~chi ]SHU and Masatum ¥I MURA 'f A

Table Table 1. MeaS 1J Femen 臼 (io mm) of the specIes.

Bellerophon Bellerophon (Bellerophon 7) aff. reguiaris (W AAGEN)

•

Khumefi'sjJ ~ra お'h~t MWATA ， E!. ge m.. n. sp.

， - ，--- ， -喧- -・・ 4 4

Gen iU 'S、 Khum 飢伊かa MU R. ATA ，s. geo.

Type-speties Type-speties :- Khumer spira ishii MUIRATA ，mew species ，Sisopholil Limestone ，Cambod~ar; M~ddle Permaam. Diagn@sお :-- ßel10~Oph011l!t~d gas ，t~opods with tigh t1 y refle~~d lateral Ii ps so wide as to cever the the latera] slope ，()f ^ whorl; inductu 't a comm0m.~y thtck and pad-like ， noω :tI 'exte 切nddi i加宣ng far beyond al lP 釘ert 岡1lt r陀e; nart U' w s鈴ðlelIl ~zo白且，e 6 1il 1 a sha 倒J喧jrpl 砂y projected med 必ian 阻JCα .r e侭s幻;，t ο例"rm lines lines <0 lilly. ;阻

Remarks:- The 'sl i}el~ @utli :m: e of ぬe proposed genus ， seems ~O incll ftde some species of Bel- lerophon lerophon (Belleropnon) MON ITFORT whlch bave angular shottlders. However ， the shou :1 der of the present new gen .tls ，is the utper margin ef a tigh ，tI y reflexed latetallip ，and an a: ηow groove remains a]ong tne shoulder iffl IJm l]): atNTe stages. 'By the characteristic development om the tbIJghdy reflexed latefa] [i ps ，Kh of the 叩Sabf司i 向am 叫i叫I卜y Bellel 釘r@pnont “i1naa Je 島M"CoY. The new generic 1'l ame is derived from the Khume :u Empire.

Khumer~ψira ishii MWA 1' A， D. S:p.

P】ate ]， figs. l_:~; P~ate 2， ngs. 1-8.

DesC1 ψ fiQn :-Shell 】arge ，anomphalous ，be 1l1 erophomtid gastlopods havimg 3> m extremely wide all1 d tigbdy refi@xed 】ateral lips; w 'horl profile angu l. atea subcardiform ，characterized by a shar"ly projected median crest amd sharply beat shouiders; ，upper surface gen t1 y curved in in early growtb :s~ages and 姐atten m: g witlh matwrity ，bending rdown abruptly to the lateral slopes; slopes; later 叫 sEope S' seems to 掴 be nearly flat ，D 1tl t [s wnk Fl owm Jt reeisely ， because of who~e coverage coverage by tighdy :f1 eftexed latetal lÏJ~s; anterier li J'S' weH curved ，bending back near selen ，izone to forn1 a distinct V-shaped SiOMS; slit narrow ，about 12 per cent (f) f body~'wh⑪ rl in in dep 出; ，selenizone extrem ，ely narrow and flat on p刈 ectea median cres .t; lateral lips ，wide ， Permian Permian gastropods 斤om Cambodia 79 as as twice as anterior lips ， tightly reflexed to cover lateral slopes entirely; ornamented with prominent growth lines. The measurements of Khumerspira ishii MURATA D. sp. are given in Table 1. Remarks:-This species is based on ten specimens ，and includes two immature and two fragmental fragmental specimens. Khumerspira ishii MURATA ， n. sp. seems to have some resemblances in in outline with some species of Bellerophon (Bellerophon) MONTFORT ， such as Bellerophon (B.) (B.) lineatus YOCHELSON ，1960 ，from the Lower Permian of North Central Texas. But ， the the present new species is read i1 y distinguishable from any known species of the genus Bellerophon Bellerophon by its extreme development of the tightly ，reflexed lateral1ips. This species is named after Dr. Ken-ichi ISH 'II of the Osaka City University ，who offered the specimens to the the writer for paleontological study. Repository:-Holotype-OMNH. E303. Paratypes-OMNH. E304 ，E305 ，E306 ，E307 ，E308 ，E309. Occurrence Occurrence :-Rather abundant in the upper part of the Sisophon Limestone (lower part of of Member D ， Lepidolina multiseptata Zone) on the narrow path of the east of Phn. Bak ， Middle Permian.

Subordeli Subordeli Macluritinae Cox and KNIGHT ，1960 Superfamily Superfamily Euomphalacea de KONINCK ， 1881 Fam i1 y Euompha 1i dae de KONINCK ， 1881 Genus Straparollus MONTFORT ，1810 Subgenus Straparollus (Euomphalus) J. SOWERBY ， 1814 Straparollus Straparollus (Euomphalus?) sp. Plate Plate 2 ，fig. 17.

Remarks :-Only a single specimen was examined. Most of the whorl is eroded. Several species species of Euomphalidae have been described from the Permian formations in Cambodia and Yur トNan by MANSUY (1912 ，1913 ， 1914) and DELPEY (1941-1942). Although the generic generic and subgeneric characters are not distinct the present material shows reselnblance with with Straparollus (Euomphalus) subcircularis MANSUY ，1912 ，and Straparollus (Euomphalus) khumerianus MANSUY ，1914 ， in outline. Repository:-OMNH.E310 Occurrence Occurrence :-Rather rare in the upper part of the Sisophon Limestone (middle part of Member D ， Lepidolina multiseptata Zone) at the north of Phn. Tup ，Middle Permian.

Suborder Pleurotomariina Cox and KNIGHT ， 1960 Superfamily Superfamily Pleurotomariacea SWAINSON ， 1840 Family Lophospiridae WENZ ， 1938 Subfamily Subfamily Ruedemanniinae KNIGHT ， 1956 Genus Worthenia de KONINCK ， 1883 Worthenia cf. schirjaevensis (STUCKENBERG) ， 1905 Plate Plate 2 ，figs. 12-14. 80 Ken-ichi IsmI and Masafumi MURATA

1 λ I/ ourlonia schirjaevensis STUCKENBERQ ，1905 ， p. 97 ， p l. 12 ，fig. 11 • Worthenia Worthenia schirjaevensis (SruCKENBERG) ，DELPEY ，1942 ， p. 355 ，fig. 27. Worthenia Worthenia cf. schirjaevensis (STUCKENBERG) ，BAITEN ，1972 ， p. 33 ，fig. 37. DesCl ψ tion :-Trochiform ruedemanniid gastropods with obscure sutural nodes rather domed convexo-concave upper whorl surface; shell sma l1， 14.3 mm in height ， 14.1 mm in width ， spiral spiral angle 85 degrees; nine spiral threads on oonvexo-concave upper whorl surface; se ・ lenizone lenizone with rather prominent growth lines and several fine spiral tbreads; concave outer whorl face with six to eight spiral tbreads; lower margin angulated with a spiral cord; base rather flattened ，with 12 or n10re spiral cords of various strengths; hemiomphalus or cryptomphalus. Remarks: -Only a single specimen is at hand. The present form resembles Worthenia schirjaevensis schirjaevensis of STUCKENBERG (1905) and DELPEY (1942) ，but is not identica l. Because ， the the character of the selen 包one of the former seems to differ from the latter. Worthenia cf. cf. schirjaevensis described by BATTEN (1972) from Malaysia shows a less angulate outline ， but has similarity with the present form in the early growth stages. Repository:-OMNH. E311 Occurrence :-Rare from the upper part of the Sisophon Limestone (middle part of Member D ， Lepidolina multiseptata Zone) ，north of Phn. Tup ，Middle Permian.

Order Caenogastropoda Cox ，1959 Superfamily Loxonematacea KOKEN ，1889 Family Pseudozygopleurridae KNIGHT ，1930

Genus Palaeostylus 恥fANSUY ，1914 Subgenus Palaeostylus (Leptozyga) KNIGHT ，1930 、 Palaeostylus (Leptozyga?) sp. Plate Plate 2 ，fig. 16.

Remarks:-A single fragmental specimen is at hand. The specimen is very small ， 7.9 mm is is height ， 3.5 mm in width ， spiral angle 25 degrees ， slender cyrtoconoid gastropods with straight straight transversa costae. Number of whorls six ， aperture lacking. There is some doubt in in referring the specimen to this subgenus. Repository:- OMNH. E312. Occurrence :-Rare in the upper part of the Sisophon Limestone (middle part of Member D ， Lepidolina multiseptata Zone) ，north of Phn. Tup ，M .iddle Permian.

Superfamily Subulitacea LINDSTROM ，1884

Family 恥1eekospiridae KNIGHT ，1956 Genus Meekospira ULRICH ， in ULRICH and SCOFIELD ，1897

M eekospira sp. 1. Plate Plate 2 ，fig. 15.

Remarks :-A sing1e incomplete specimen ， lacking the spire in the younger stages and ha]f Permian gastropods from Cambodia 81

of the body whorl ， is at hand. The present material ， 16.7 IDDl in height ， 7.6 n1ffi in width ， with spiral angle of nearly 15 degrees is a slender turriculate form with smooth surface. Although some genera of the Subulitacea have been recorded frorn the Pernlian formations in in South Eastern Asia ，the present form may be refered to this genus. R 々pository:-OMNH.E313 Occurrence :-Rather rare in the upper part of the Sisophon Limestone (middle part of Member D ， Lepidolina multiseptata Zone) ，north 'of Phn. Tup ，Middle Permian.

References References

BATTEN ， R.L. (1972): Permian gastropods and chitons from Perak ， Malaysia. Part 1. Chitons ， Belleropbontids ，Euomphalids and Pleurotomarians. Bull. Amer. Mus. Nat. Hist. ，vo 1. 147 ， t. ar t. 1， p. 1-4 4， 52 figs. CIu-THUλN ，Tran ・Thi (1961): Les brachiopodes permiens du Phnm-Tup (Sisophon-Cambodg~). Ann. Ann. Fac. Sci. Sa なon ， p. 167-308 ，pls. 1 -8. DELPEY ，G. (1941 -4 2): Gasteropodes permiens du Cambodge. Jour. de Conchy l.， vo l. 84 ， p.

255-278 ， 19 text-figs.; p. 345-369 ， 23 text 圃 figs.; vo l. 85 ， p. 50- 83 ， 28 text-figs. DEPRAT ， J. (1912): 亘tude des Fusulinides de Chine et 1' lndochine et classification des calcaires a fusu Ii nes. lndochine Serv. Geol. Mem. ，vo l. 1， no. 3， p. 1-76 ，pls. 1-9. (1913): (1913): Ibid. (11 Memoire). Les Fusulinides des calcaires carboniferiens et permiens du Tonkin ，du Laos et du Nord-Annam. Ibil ι，vo l. 2， no. 1， p. 1-74 ，pls. 1-10.

FONTAINE ，H. (1961): Les Madreporaires paleozoiques du Viet 咽 Nam ，du Laos et du Cambodge. Archives Archives Geol. du Viet-Nam ， no. 5， p. 1-276 ，pls. 1-35. (1965): (1965): Quelques Spongiaires du permien superieur du Cambodge. lbid. ， no. 7， p. 39 -4 3. (1967): (1967): Quelques Madreporaires permiens du Viet-Nam et Cambodge. lbid. ， no. 10 ， p. p. 51-58 ，pls. 1-3. GUBLER ， J. (1935a): 亘'I ude geologique an Cambodge occidenta l. Bull. Serv. Geol. Indochine ， vo l. 22 ， no. 2， p. 1- 176. (1935b): (1935b): Les Fusulinides du permien de l'lndochine. 九 1eln. Soc. Geol. France ，N. S. ， no. no. 26 ， p. 1-171 ，pls. 1-8. ISHII ，K. (1966): Permian of the western part of Cambodia-On the straitgraphy of Sisophon and Battambang limestones in Cambodia. “Fossils" ， no. 12 ， p. 29-39. (in Japanese) ，and NOGA 閲， Y. (1964): CO l1 tributions to the geology and palaeontology of Cambodia. Part Part 1: Permian Fusulinids. Jour. Geosci. ，Osaka City Univ. ，vo l. 8 ，art. 2， p. 9-68 ，pIs. 1- 8. ，KATO ，M. and NAKAMURA ，K. (1969): Permian limestones of West Cambodia-Litho- facies facies and Biofacies. (Contribution to the geology and palaeontology of Cambodia ， Part 3). Palaeont. Palaeont. Soc. Japan ， Spec. Paper ， no. 14 ， p. 41 -55 ，pls. 9-1 1. KNIGHT ， J.B. (1930): The gastropods of the St. Louis ，Missouri ， Pennsylvanian outlier . Jour. Palaeont. ，vo 1. 4，Supp l. 1，The Pseudogopleurinae ， p. 1-88 ，pls. 1-5. (1941): (1941): Paleozoic gastropod genotypes. Geol. Soc. America ， Spec. Paper ， no. 32 ， p. 1-510 ，pls. 1-96. (1944): (1944): Paleozoic Gastropoda. ln SHIMER ，H.W. and SHROCK ，R.R. ， Index fossils of 、 North Am erica. New York ， p. 437 -4 79 ，pls. 17 4- 196. (1956): (1956): New families of Gastropoda. Jour. Washington Acad. Sci. ， vo I. 46 ， no. 2， p. 41 ， 42. 42. KONINCK ， L.G. de (1881): Faune du calcaire carbonifere de la Belgique ，3e partie ， Gasteropodes. Mus. Roy. Hist. Nat. Belgique ，Ann. ，ser. paleont. ，vo l. 6， p. 1-170 ，pls. 1- 24. (1893): (1893): Jbid. 4e partie ， Gasteropodes (suite et fin): Ibid. ，vo l. 8， p. 1-240 ，pls. 22-54. 82 Ken-ichi IS Hll and Masafumi MURATA

MANSUY ，H. (1912): 亘tude geologique du YUDnan Oriental ，2 e pt.， Paleontologie. lndochine Serv. Serv. Geol. Mem. ，vo l. 1 ，fasc. 2， p. 1-190 ，pls. 1-10. (1913): (1913): Fau :m es des calc 羽res a Productus de l' Indocbine. lbid. ，vo 1. 2， fasc. 4， p. 104 ， pls. pls. 1- 13. (1914): (1914): Ibid. ，2e ser. Ibid. ，vo l. 3 ，fasc. 2， p. 1-190 ，pls. 1- 1'0. M 'CoY ， F. (1851): On some new Si1 urian Mollusca. Ann. Mag. Nat. Hist. ，ser. 2，vo l. 7， p. 45- 63. 63. SAURIN ， E. (1959): Notes stratigraphiques sur le permien super 允ur de Battambang et Sisophon (Cambodge). Ann. Fac. Sci. Saigon ， p. 121-127. SERRA ， C. (1966): 豆tude anatomique et paleogeographique de quelques espωes Homoxylees du sud Viet ・Nam et du Cambodge. Archives Geol. du Viet-Nam ， no. 8， p. 59-131 ，pls. 1-15. SKINNER ，J.W. and WILDE ，G.L. (1954): The Fusulinid subfamily Boultoniinae. Jour. Paleont. ， vo l. 28 ， no. 4， p. 434 -4 44 ，pls. 42 -4 5. STUCK BNBERG" A. (1 905): Die Fauma der obercarbonischen suite des Wolgadurchbruches bei Samara. Mem. Com. Geol. ，N.S. ，vo l. 23 ， p. 1-144 ，pls. 1-13.

VIEN ， L.T. (1959): 豆tude de Fusulinides du permien du Haut- Laos ，du sud Viet 回 Nam. Ann. Fac. Fac. Sci. Saigon ， p. 99-120. WAAGEN ，W.H. (1880): Productus Limestone fossiIs. Palaeont. lndica ，ser. 13 ， Salt Range fossils ， pt. pt. 2，vo I. 1， p. 73 -183 ，pls. 7- 16. WENZ ，W. (1938): Gastropoda. Allgemeiner Teil und Prosobranchia. ln SC H1 NDEWOLF ，O.H. ， ed. ，Handbuch der Palaozoologie ，Band 6， 1639 p.; Teill ， p. 1- 240. YOCH E:凶 ON ， E.L. (1960): Permian Gastropoda of the southwestern United States. Part 3，Bel- lerophontacea lerophontacea and Pate Il acea. Bull. Amer. Mus. Nat. Hist. ，vo l. 119 ，art ，4 ， p. 205 -294 ，5 text-figs. ，pls. 46-57. :t :t

Plate Plate 11

• 84

E~lanation of Plate 1i

KJ 伽(/:Jfl ersp lI! ra ishti MTUltATA ，日. gen. D. SQ.

日gS. 1-6. Adapertuifal l ，right sid@" ~pertmtal ; left sride ]p osterior and obl li'll ue ad 叩 ert lJl ltral views (i):f (i):f s: olotype ，rO 'M !N H. E303 ，x L

Figs. Figs. 7 ，8. Or oss sectio 11l s of a paratype ，OMNH. E309 ， to sfu. ow the Feflexed la ぬral t:ightJl~ < llips ，X ] .$.

l1~oth S叩JPe 伐c加mZ路6且郎s were co !I日Ueωtt t 飽吃吋i“ecl lb防y Ke 釘閃:Iil吋 1 JDo ぱf もぬ lle Si おSO J})ID，@m_ Limes 鈎ωbωome 河申 (包 Le ゆEβ "， idalina n:l ulfifsep 的 'ta Z~}llle) ， east (f) f Ffu:J!. B'ak ，meati Siso J.D ho lDl， CaJrol~odia.

、 ISHII ISHII & MURATA: Khumer宅pira~from:Cambodia. Plate 1

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K. KUMAGAI & S. OHTOMO Photo. ， Plate 2

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Fi lgs. l-tt K hJ umers JjJ rira ishii M iURALFA ，si gen. fl. SJ!' . 1-4 ，. Adap 、ertufal ，~p(?r~u，Ji al， ø@$teri~r :_a nd left 釘:d e views @f a paraty ，e，'O ，MNH.E

304 ，x 1. 5-$. Ada l' ertural ，a J!' ertuffa[ ，1' 08 ぬf，iof and right side views ， of atl@ther paratype ，0 MNH. E30S ，x 1/2.

Both slf' eo 1m etls Were collec 匂，d frQm the lower )D a1i t of M 創 nlber D ，of th8 Sisophom Lillllestone Lillllestone (LepidoZ~na muttrse ， tata ZO f.nl， e) ，eas を of Ph 唖. :s ak ，tJl eaJ r Sisopmotl ，Cam - bod~a ， ~y Kem-i : c} i}!I ffiSHH.

Fi lgs. 9-[ ，1" BelZero j!) hon (BellerQpnon?) ，aff. regutaris (WAAGEN)

Adape 1i haral l，aper 側 ra 'l aJ md 国;ght s~4e views of a sf'~ci ，me1il" ，O 'MN 回. E3d1 ，xt.

s ，ori ，z i@ ，n (j)F1 d L@ealit ァ; same as :fi gs. [~$.

F I' gs. i2 -- 14. Wortheruia cf. schIJrjtfl(!ve 1fl 的 (5rucKENBERGJ

Ap I' ca] " basaa (bm: cr ~aJiteraJl' view S' of as ;p ecime ，m，'O IMNH !. E311 ，x 2.

Fig. Fig. 15. 九t/ eekospira S1P. 、 Latelial Latelial view of af 1i ag 'Jj)ji} efllt aJ l speci 1men ，Q ，MNH. El rn 3，x 2.

Fig. Fig. 16. Pa!aeostylus (Leptø~ygra ?) sp.

Lateral Lateral view (C) fa specimem ，0 恥1NH. E312 ，. x 3.

Fi lg. 17. S t' Yaparollu S' {Euompli a; lu s: ?) SP " Ap にal view of a specimen ，OMNH. E3 1( ()， x 1.

The s}D ecime l1l s iUtls もrated I1il. sgs . W2-~7 were cQ]lected by K~n-ich[ ]SH Ir( fli@ Ol the

Flil iddle paF t， of Mem lmer 0 of tlle SiS@ :F> !RO m. Liraest@ne (Lep 'idolina multiseptata

ZOlfle) l， Jnl. olith of 1? hn. 'f UIP， hear Sおo，ph@ iIJ1， Camòod~a. IS Hll & MURATA: Khumerspira from Cambodia. Plate 2

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11 11 17

14 14

K. KUMAGAI & S. OHTOMO Photo.