I+cl

Distribution of two species of arrow squid (Nototodarusl around

by P. J. Smith

Fisheries Research Division Occasional Publication No. 49 i;-t'' r "-'i í :i I '.' Distribution of tw arrow squ¡d (Nototodarusl around New Zealand

by P. J. Smith

Fisheries Research Division Occasional Publication No. 49 I 985 Pubtished by the New Zealand Ministry of Agriculture and tr'isheries Iilellington 198s

ISSN 0r10-U65

MAF Libraries cataloguing-in-publication data smith, P.J. Distribution of two species of arrow squid (Nototodarus) around New Zealand / by P.J- Smith. lv. (Occasional publication / N.Z.Fisheries Research Division, ISSN 0110-1765 ; no. 49)

1. Cephalopoda 2. Squid rsBN 0-477-03110-2

Edited by J. L. Taylor and G. G. Baird

Set in l0 on 11 Times Contents

Page Introduction 5

Methods 5

Results 7

Distribution 7

Management implications 9 Acknowledgments ll References 1t Appendix I il 350 s

i* Bay of Plenty

Cape Egmont South Bight 400

dotôt*

Westport

Hokitika Ganyon

\ I ¡ t t I, \-----"

500

Auckland ls S

Fig. l: Places and hydrographic features mentioned in the text. lntroduction The distributions of the two species of arrow squid around New Zealand are described here. Samples have been collected from a wide area The New Zealand arrow squid frshery has expanded within the New Zealand 200-mile exclusive rapidly from insignificant by-catches in the 1960s economic zone to provide information on the tem- to become the most important single fisheries poral and spatial distribution of the two arrow resource in terms of export dollars. Most of the squids. Electrophoresis, because it offers positive catch has been and still is taken by foreign (licensed, identifrcation of all specimens regardless of size or chartered, and joint venture) vessels. A jig fishery sex, was used for species identification. developed by Japanese, Korean, and Taiwanese vessels around the mainland of New Zealand in the 1970s produced 51 315 t in 1983. A trawl fishery developed by Japanese and Russian vessels around the Auckland Islands produced 40 899 t in 1983.

Few biological studies had been made on New Zealand arrow squid before the development of the flshery and the resource was assumed to be of one species (Kawakami 1976, Roberts 1978). A bio- Methods chemical-genetic survey in 1979 found that there 'were two species in the ñshery (Smith, Roberts, and Hurst 1981). The two species were distinguished by their protein types visualised with the technique of Samples of arrow squid were collected aboard gel-electrophoresis. Subsequently a morphological research vessels and commercial fishing vessels difference was found between large males, but small between North Cape and the Auckland Islands males and all females and juveniles could not be (Fig. 1, Fig. 2). Some specimens were frozen whole; separated into species (Smith et al. l98l). for others a piece of body muscle tissue (about 5.9) was dissected out and frozen and the length and, where possible, sex data were recorded. The whole At about the same time Japanese Taiwanese and frozen specimens were measured and sexed in the workers independently described morphologi- the laboratory. A few samples of whole specimens were cal difference between mature males (Tung 1978, ports. Kawakami collected from local coastal vessels at various and Okutani l98l), although Tung (These in Appendix I by port assumed samples are identifred that this variation was intraspecific. name and lack of location data.) Appendix I lists Kawakami and Okutani recognised the two species, all samples of arrow squid collected around New and in addition the difference hectocotylus to in Znaland between January 1979 and September 1984 types between mature males showed they a differ- and tested by electrophoresis. ence in sucker counts on the frrst right arm, which could be used to distinguish females of the two spe- For electrophoresis a small piece cies. They further subdivided the east coast species of body muscle (about 0.1 g) was ground into two types based on the number of suckers on up in an equal volume of distilled water the modified portion of the hectocotylus and on the and the supernatant placed on to a 0.5-cm2 paper club length as a percentage of mantle length. How- filter wick. Twenty-six wicks were placed into starch gels: 24 ever, these latter frndings have not been verifred by unknown specimens and 2 controls, New Zealand workers (R. H. Mattlin pers. comm.). one of each species. The starch gels were made up at a concentration of l2Vo (weight/volume) in a phosphate-citrate buffer at pH 7.0. They were Two species are likely to have different biological run at 100 volts in a refügerator for 4 hours, after properties and exploitation rates and ideally would which the gel was sliced into two and stained for be managed as separate units. This is usually so for the enzyme a-glycerophosphate dehydrogenase the major ñnfrsh species, which are readily distin- (Smith et al. l98l). This enzyme was selected from guished from each other, but the close phenotypic the initial analysis for three reasons: appearance ofthe two species ofarrow squid does It resolved clearly in not permit separate management. However, initial L all specimens regardless of srze or sex; investigations have indicated that the species have different geographical distributions (Kawakami and 2. No heterozygous individuals (both protein types Okurani 1981, Smith et al. t98l), which, if con- together) were found; firmed, would allow the species to be managed 3. It was stable in muscle tissue frozen at -20 "c separately by area. for 2 years or more. 350 s

400

\

Sample size O approx.50 O approx. loo O approx' 5oo f\.t I more than looo

Fig..2; Location of sampling sites, distribution of the two species of arrow squid, and ocean currents around NewZealand. The northern species is shown as white and the southern species as black in the pie charts. Results

Figure 3 is a photograph of a gel stained for the enzyme ø-glycerophosphate dehydrogenase. One species of arrow squid resolves with a fast migrat- ing band and the other species with a slow migrat- ing band. Most individuals were fixed for one or other of these protein bands. Three heterozygous (carrying two different genes) fast types and two heterozygous slow types were found, but these were heterozygous for rare alleles and did not have the common fast and slow bands together. The two species did not share rare alleles.

Distribution Orioin*-1234567 - I The electrophoretically fast species is found off the (-) west coast of the North and South Islands, off the north-east coast of the , in the Bay of Fig. 3: c-Glycerophosphate dehydrogenase phenotypes in a starch gel for the two species ofarrow squid. Samples l- Plenty, offthe south-east coast of the North Island, 4 are from west coast North Island and samples 5-8 from and off the north-east coast of the as the Chatham Rise. far south as Banks Peninsula. It was not found on the eastern Chatham Rise (Appendix l). in a gradual temperature cline off the west coast The electrophoretically slow species is found off with no sharp discontinuity as observed over the the south-east coast of the North Island, off the west Chatham Rise. For the coastal teleosts there is a coast of the South Island, in Tasman Bay and as change in species composition from a northern type far north as Cape Egmont on the west coast, offthe community (gurnard, kahawai, snapper, trevally) to east coast of the South Island, on the Chatham Rise, a southern type community (blue warehou, star- and offthe Snares and Auckland Islands (Appendix gazer, tarakihi) around the Cook Canyon-Hokitika l). Canyon area (J. M. Fenaughty pers. comm.). Thus the two speeies are essentially northern ancl If the two species of arrow squid are differentially southern in distribution, not eastérn and western adapted to water temperature, and they appear to as assumed from the initial analysis (Smith el a/. be so by their general northern and southern dis- l98l). The electrophoretically fast species is the only tribution, a gradual temperature gradient as off the one found north of the Bay of Plenty on the east west coast of the South Island would allow more coast and north of Cape Egmont on the west coast. extensive species mixing. Thus the distribution of The slow species is the only one found south of the two species may vary both between seasons and Banks Peninsula on the east coast. There is overlap between years. The D'Urville Current is thought to around the coasts of central New Zealand (Fig. 2). vary in strength and in strong periods could move On the east coast the division between species more southern squid into the Tasman Bay-South approximates to the Subtropical Convergence zone Taranaki Bight fishery Gig. 2). along the Chatham Rise, with the northern species An indication of seasonal changes in distribution in subtropical water to the north of the zone and can be made from samples collected during a series the southern species in subantarctic water to the of cruises carried out on the research vessel James south. This convergence zono along the Chatham Cookbetween January 1982 and March 1983. The Rise is a hydrological barrier for many species of same stations were covered on each cruise between fish and invertebrates (Robertson, Roberts, and Tasman Bay and Stewart Island. The distribution Wilson 1978). On the west coast the zone is less of the two species appears stable over this time clearly defined, but it is to the south of the South period, particularly off the east coast of the South Island. The southern species is found well to the Island (Fig. a). Most arrow squid caught in Tasman north of this zone, although it forms less than 100/o Bay were the northern species, but the southern was of the squid catch off the north-west coast of the present in small numbers in 16 out of 54 samples South Island (Fig. 2). The Tasman Current diverges and was the only species in 3 samples (Appendix off the west coast of the South Island into southern 1). In two cruises in July-early August and late and northern components, to produce respectively August 1982 (Appendix I and Fig. 4) the propor- the Southland and Westland Currents. This results tion of southern squid rose fo 40o/o and 460/o 1700 E

Jul-Aug 1982 Aug-Sep 1982

Fig.4: Sampling sites and distribution ofthe two species ofarrow squid from January 1982 to March 1983. The northern species is shown as white and the southern species as black in the pie charts. All samples were of more than 20 individuals. respectively, but these samples were small (42 and Management implications 24 individuals) and the squid were mostly juven- iles. Therefore they may not reflect actual abun- dance of fishable adults, but possibly larval drift from a west coast South Island spawning ground. Arrow squid are managed as one species at present. This need no longer continue, for, although the two Samples of squid collected off the west coast of species are not readily distinguished at sea or in the the South Island indicate seasonal changes. One processing shed, they are for the most part spatially hundred and fifty-eight squid collected in January isolated. The four main jig-ñshing, areas are in area 1982 aboard a commercial vessel off Westport H between Tasman Bay and the Taranaki coast during the squid season were all southern species. (essentially the northern species), in area C offBanks Two samples collected aboard a research vessel off Peninsula and in the Canterbury Bight (essentially the central west coast in August 1984 outside the the southern species), in area G off Westport (all commercial season were 30ó southern (in a sample southern species in the 1982 summer frshery), and of 69 individuals) and 5590 southern (in a sample in area F off Stewart Island (all southern species) of 64 individuals). Samples collected by Taiwanese (Fig. 5). The trawl fishery in area E on the Auck- pre- workers between January and April 1977 were land Shelf is based on the southern species. dominantly the northern species (based on male hectocotylus type) off the north-west (991 out of 1003 individuals) and north (1421 out of 1426 indi- Ifthese five frshery areas are treated as single spe- viduals) coasts of the South Island (Tung 1978). cies areas, for the 1981-82 season approximately seven times as much southern squid (74 884 t) as Low numbers of northern squid were caught northern squid (10 931 t) was caught. For the 1982- south and east of (Fig. 2). One sample 83 season approximately 2.5 times as much south- of only seven individuals from the western ern squid (63741t) as northern squid (25 500 t) was Chatham Rise was the northern species, whereas caught. Provisional figures for the 1983-84 season other larger samples along the Chatham Rise have show that approximately 18 times as much south- been of the southern species. Samples collected off ern as northern squid was caught (Table l). These the south-east coast of the North Island show con- differences in catches probably reflect differences in siderable mixing of the two species (Appendix 1), actual abundance. Experimental fishing for squid in which probably reflects the mixed water movement the Bay of Plenty and off the north-east coast of in this area (Fig. 2). Adults collected offCastle Point the North Island has not found large quantities. The in July 1980 were 580/o northern, juveniles in arrow squid in Australia that is the same as the Hawke Bay in February 1981 were 210lo northern, northern species likewise does not support a large and juveniles offWairarapa in February l98l were fishery. Dividing the squid quota by area would also 84oó northern. A sample of juvenile squid from divide the quota by species and would prevent all Cook Strait in April 1979 was 90% northern. There effort being directed on to one species, in particular is no significant fishery off the south-east coast of the less fished northern species. Although there is the North Island where species mixing occurs. a need to monitor species distribution in the com- Howe¡er, further samples of commercial sized mercial fishery, particularly offthe west coast of the squid are needed from this area to establish the South Island, the present results indicate that it proportional distribution of the.two species over would be feasible to manage the two species of the summer-autumn fishing period. arrow squid independently by fishery areas.

TABLE l: Total catch of arrow squi{ (t) and percent of annual catch (in brackets) for each fishery area for the years l98f-84

Fishery l98l-82 l 982-83 I 983-84* area ,Species I Sep-31 Aug I Sep-30 Sep I Oct-30 Sep B northern 0 0 I (0.001) H northern t0 930.2 (t2.7) 25 499.5 (28.6) 5 205.0 (5.6) C southern 2t 193.3 (24.7) 7 869.4 (8.8) l4 608.3 (l 5.8) D southern 208.2 (0.2) 6 359.1 (7.1) 2 s30.9 (2.8) E southern 38 623.7 (45.0) 26 518.1 (29.7\ r2 070.1 (13.1) F southern 3 392.9 (4.0) 9 308.9 (10.4) 56 851.7 (6r.6) G southern l r 4ó5.5 (l 3.9) I 3 685. l (l s.4) r 012.8 (l.l) 85 813.8 89 240.t 92279.8

* Includes provisional figures for trawl-caught squid. 1750 E 1 goo

A

\/---_^ \r------\----__

EE

I I \ \t- Sample size O approx. SO O approx. loo O approx. soo

more than looo O

Fig. 5: Location ofsampling sites and distribution ofthe two species ofarrow squid around New Znaland.in relation to the ûshery zones. The northern species is shown as white and the soutbern species as black in the pie charts.

l0 Acknowledgments Appendix 1

I am gtateful to P. G. Benson for technical assist- Location, date of capture, size range, and electro- ance and to R. H. Mattlin and J. M. Fenaughty for phoretic results for squid collected around New helpful discussions during the course of this work. Zealand between 1979 and 1984. The following staff provided samples of squid: L. Allen (Fisheries Management Division), M.F. Electro- Beardsell, P. G. Benson, S. D. Canning, E. C. Förch, phoresis R. J. Hurst, J. B. Jones, R. H. Mattlin, D. M. Tra- cey, the late W. L. F. van den Broek, and G. J' Voss. Mantle length No. No. Date (cm) fastslow

West coast North Island 39" S 0 38' S 0 39' S 0 39" S 0 39" S 7 38' S t74" 00'E 4.2.84 2l-36 l0 0 References 36" S 173" 46', E 6.2.84 lr-25 I I 0 35' S 173" 03', E 7.2.84 24 1 0 35" S 172" 52', E 7.2.84 24-27 9 0 K¡,wnrcrvrt,T. 1976: The frshery biological study 35" S 172" 25', E 9.2.84 2'1,28 2 0 on a squid Nototodarus sloani sloani (Gray) in 35' S r73" 40', E 10.2.84 7-15 24 0 the New Zealand water. Bulletin of Tokai 35" S 173" 32',8 10.2.84 14-23 6 0 Regional Fisheries Research Laboratory, No. 85: 36' S 173" 59', E 11.2.84 16-22 5 0 31-104. 36" S 173" 59', E 11.2.84 19-22 5 0 38' S 174" 07', 27.5.84 32 1 0 K.cwnKANrt, T., and OKUTANI, T. 1981: A note on E S t73" 49', 27.5.84 9-38 39 0 identity of Ommastrephid squids of the genus 38" E 37" S t74" 34',F. 28.5.84 7 | 0 Nototodarus exploited in the New Znaland waters. S 173" 29.5.84 27-35 3 0 Bulletin of Tokai Regional Fisheries Research 36" 48',8 35' S r72" 54',F, 30.5.84 ll-17 7 0 Laborøtory, No. 105:. 17-29. 36" S 174' 08', E 2.6.84 l8 2 0 Ronnnts, P. E. 1978: New Zealand squid resources. 36" S 174' 01' E 2.6.84 15,33 2 0 (Comps.), Pro- In Habib, G., and Roberts, P. E. 38' S 174' 4',|', E 3.6.84 7-ll l7 0 ceedings of the Pelagic Fisheries Conference, July 38" S 174" 06',8 3.6.84 21,23 2 0 19J7, pp. 90-3. Fisheries Research Division 37" S 173" 59' E 3.6.84 29,34 2 0 Oòcasional Publication, N.Z. Ministy of Agri- 38' S 173'18',E 5.6.84 15-31 9 0 culture and Fisheries, No. 15.. RosrnrsoN, D.4., RoneRts, P. E., and WtlsoN, North Cape J. B. 1978: Mesopelagic faunal transition across 34" 05', S 172" 17', E 23.4.81 17-32 380 the Subtropical Convergence east of New 34" 27',S 172' 15',E 31.5.84 rl-20 s00 Zealand. N.Z. Journal of Marine and Freshwater Research 12 (4): 295-312. North-east coast North Island SMrrH, P. J., RoneRTS, P. E., and HuRsr, R. J. Whangarei 8.12.80 no data 12 0 1981: Evidence for two species of arrow squid in Whangarei 8.12.80 no data 25 0 the New Zealarrd fishery. N.Z. Journal of Marine 35" 07', S 174" 41', E 2t.4.8t 19-26 9 0 and Freshwater Research 15 (3): 247-53. 35'40', S 174" 5l',E 20.4.8t t8-26 36 0 TuNG, I.-H. 1978: On the biology and the frshing 36" tl' s 175' l3',E 3.t2.8r l4-r9 26 0 of the squid, Nototodarus sloani sloani (Gray), in 36' ll' s 175" 13', E 3.12.81 10-16 16 0 the New Zealand \ilaters. Report of the Institute 36" 31', S 176' l3',E 23.2.84 15-30 t2 0 of Fishery Biology of Ministry of Economic Affait's 36" 04',S 175' 58'E 27.2.84 14-28 13 0 and National Taiwan University 3 (3): 44-64. 35" 37'S 175'07',E 26.2.84 16-22 13 0 [English translation held in Fisheries Research 36" 07',S 175' 52',E 23.2.84 26-30 13 0 Division library.l 35" 37',S 175" 07',E 26.2.84 19-27 16 0

l1 Bay of Plenty 40" 14' s 173" 30,E 3.2.81 28-34 23 0 Tauranga 6.1.81 t7-23 93 0 40" 17' s 173" 44' E 7.2.81 l3-16 47 0 36" 47', S 175. 5l'E 24.2.8r l1-16 27 0 40" l7' s 173" 44', E 7.2.8t t9-23 8 0 36" 4g'S 175" 50'E 22.2.8t 6-15 6 0 40" 17' s 173" 44' E 7.2.81 24-29 t3 0 37" 37',S 176. 15'E 22.2.81 10 I 0 40" t7's 174.00'E 20.2.8t 2t-30 t2 0 Tauranga 12.9.84 t0-22 35 0 40" 17' s 173. 23',E 23.2.8t 20-33 12 0 37" 31', S 176" 37' E t7.9.84 t4-26 10 0 40'55',S 173" 25',E 8.1.82 2t,23 2 0 40" 29',S 173" 41',E 8.1.82 7-25 t7 0 South-east coast North Island 40" 20's 173" 51',E 8.1.82 tr-25 39 0 40" 54',S 173" 25',E 3 4 Castle Point I 1.7.80 l8-33 18 13 19.t.82 2-15 40" 29',S 173. 3g',E 19.1.82 2-24 8 0 Hawke Bay 40" 20' s 173" 52', E 20.1.82 t3-27 3 I 40" 2l's 173. 52',E 20.1.82 t3-29 45 3 39" 19', S 177" 44', E 5-7 6 8 27.2.81 40' 55', S 173" 2g'E 16.2.82 l l9 0 39" 22', S 44',8 l-36 ll7" 27.2.8t 5-6 3 2 40" 30', s 173" 39'E t6.2.82 25-37 50 I 39" 26', S 177" 43' E 27.2.81 l-14 0 3 40" 22', S 173" 23' E 16.2.82 8-2t 102 4 39" 28', S t77" 44', E 27.2.8t 2-7 2 31 40" 31', s 173" 40, E 16.2.82 3-8 8 0 39" 35'. S 177" 46' E 3-5 2 5 27.2.8t 40' 55', S 173" 24', E 8.4.82 t7-33 30 0 40" 29',S 173" 4l,E 8.4.82 7 0 Wairarapa tt-21 40" 20',s 173. 53',E 8.4.82 12-22 l8 0 40'55', S 176" 14'E 17.2.81 2-8 I 1 I 40" 2g' s 173" 42'E 21.4.82 2-3 3l 0 40" 52', S 176. 16'E t7.2.81 5,6 2 0 40" 55', S 173. 25',E 29.5.82 t4-27 35 0 40" 53', S 176" 16'E 18.2.81 4-7 t7 0 40" 29',S 173. 4l'E 29.5.82 26-34 5 I 40" 54',S 176. 16'E 18.2.81 2-7 l8 0 40" lg',s 173. 52',E 9.5.82 9-29 15 0 40" 54', S 176. l6',E 18.2.81 4-18 35 l0 40" 20's 173. 50'E 9.6.82 2,3 2 0 40' 53', S 176.20'E r 8.2.81 4-9 4 5 40" 27', S 173. 52'E 9.6.82 2-6 1l 0 40" 52', S 176" 20',E 18.2.81 2-7 4 2 40" 20's 173. 5l,E 24.7.82 l-5 0 T6 40" 18', s 173. 53',E 24.7.82 t-4 7 I Cook Stait 40" 55', S 173" 25,E 5.8.82 t7 2 0 ,7 40" 28'. Island Bay 29.4.'79 1-9 62 S 173" 42',F. 5.8.82 t0-24 l0 0 40" 2l' s 173" 52', E 5.8.82 l0- 19 6 0 West coast South Island 40" 56',S 173" 25'E 26.8.82 10-19 l3 0 40" 29', S 173" 40',E 26.8.82 7-14 0 5 41" 57', S 170. 56',E 15. r.82 l6-18 0 158 40" 22',S 173. 54' E 26.8.82 8-27 0 6 41" 30' s 171" r 0 ll'E 30.8.84 10-28 l 40" 54', S 173" 27', E 12.11.82 4-25 66 I 41" 34', S 171'02',E 30.8.84 rt-26 t4 I 40" 29',S 173" 41',E 12.1t.82 t2-19 3 0 41" 37', S 170' 54',E 30.8.84 13-33 t9 I 40" 20's 173" 51',E t2.tt.82 t7-23 3 0 41" 35', S 170" 48',E 31.8.84 23 0 l0-28 40" 55', S 173" 24' E 8.1.83 l5-34 38 0 42" 27', S 170. 50'E 3.9.84 2 I t7-28 40" 28',S 173" 39,E 8.1.83 t6-37 190 6 42" 30', S 170" 41',E 3.9.84 3 3 t6-40 40" 54',S 173" 25',E 4.3.83 tt-37 25 0 42" 38', S 170' 17',E 4.9.84 7 J l5-33 40" 29', S 173. 40'E 4.3.82 28-37 3 3 42" 51', S 170" Og',E 7.9.84 18-28 2 4 40" 55', S 173. 25',E 15.3.83 no data 8 0 42" 57', S 170. 02',E 0 2 4.9.84 34 40" 37',S 173" 03'E 28.1.84 27-30 l0 0 43" l4',S 169'46',E 5.9.84 3 5 18-33 40" 37',S 173. 00',8 28.1.84 25-32 I l 0 43" 09', S 169" 57',E 5.9.84 13-36 l0 6 40" 4l' s 172. 5l,E 28.1.84 25-33 8 0 43'03', S 170'07',E 5.9.84 30-33 0 6 4l'05' s 173" 04', E 29.1.84 22-30 3 0 43" 13', S 169" 44',E 6.9.84 12,19 I I 43" 13', S 169" 56',E 6.9.84 16-33 l 4 East coast South Island Tasman Bay Kaikoura Tasman Bay -.2.79 t8-27 12 0 4l' 58', S 174" 07', E 9.1.82 t5-25 12 6 Tasman Bay -.2.80 t9-25 24 0 41" 58', S 174. ll',E 9.1.82 25,26 0 2 Tasman Bay -.5.80 8-t2 29 3 42" 00' s 174" 25', E 9.r.82 30-31 0 5 40" 28', S 172" 06',E 30.1.81 t2-16 r5 0 41" 59', S 174" 3l',E 9.1.82 34 0 I 40" 14', S 173" ls',E 5.2.81 t2-t6 26 0 42" 00' s 174" 07' E 18.1.82 z-tt l8 133 40" L4', S 173" l5',E 5.2.81 8-27 t2 I 41" 57', S 174" 34', E 16.2.82 31 0 1 40" 14' s 173. 30',E 3.2.81 t2-17 l8 2 42" 0r' s 174" 2l' E 17.2.82 24 I 0 40" l4',s 173' 30',E 3.2.81 l3-16 24 0 42" 04'. S 174. 03',8 11.4.82 2t 0 I

t2 42" 09', S 174" 04',E 1t.4.82 32 01 43" 45', S 173" 47', E 14.t1.82 t4-27 0 50 42" 03' S 174" 05',E 20.4.82 27 01 43" 45'S 174" 01',E r4.rt.82 20-32 0 4 42" 07',S 174'02',E 30.5.82 t3-37 218 43" 48', S 174" 12' E 14.1t.82 32,35 0 2 42" 12', S 174' 03',8 30.5.82 t0-37 0 1l 43" 50', S 173'09',E 23.1r.82 8 0 l 42" 03',S 174'04',E 8.6.82 4,6 02 43' 50', S 173' 08',E 10.1.83 8-29 0 7 42" 06',S 174'08',E 8.6.82 3-6 020 43" 45', S 173" 46',E 10.1.83 27 0 l 42" 09', S 174' l8',E 8.6.82 3-4 03 43" 45', S 174'00',E 10.1.83 34 0 1 42" 05', S 174'03',E 24.7.82 2-4 019 43" 43',S 174" Ol',E 10.1.83 25-27 0 3 42'05',S 174'08', E 24.7.82 2-4 0 11 43' 50', S 173'08',E 17.1.83 5-9 0 23 42" 05', S 174' 08', E 24.7.82 2-8 08 43" 46', S 173'48',E 18.1.83 7-20 0 3 42" 09', S 174' 17',E 24.7.82 3 0l 43" 47', S 173'49',E 18.1.83 6 0 I 42'09', S 174" 08',E 24.7.82 2 01 43" 50', S 173' 07',E 6.3.83 t7-37 0 70 42" 05', S 174' 03',E 4.8.82 t9-36 04 43" 45',S 173" 48',E 6.3.83 29-37 0 6 42" 00' s 174" 27', E 4.8.82 20-33 03 43" 44', S I74" 02', E 6.3.83 t7-32 0 t2 42'00' s 174" 3l',E 4.8.82 34 01 43' 50',S 174' l4',E 6.3.83 25,3t 0 2 42" 05', S 174' 03',E 31.8.82 33 01 43" 46', S 173" 47' E 13.3.83 2-5 0 12 42" 05',S 174'08',E 31.8.82 34 02 43" 45', S 173'48',E 13.3.83 2-4 0 8 42" 09', S 174'04',E 32 31.8.82 0l Bight 42" 04', S 174" 04',E 13.11.82 29-39 03 Canterbury 42" 07',S 174'07', E 24.1r.82 5-9 0 15 45" 01', S 171" 34',h 22.3.80 9-28 0 78 42" 04', S 174'03',E 9.1.83 27 03 45" 03', S 171" 34', E 5.12.80 no data 0 11 42" 05',S 174" 07',E 9.1.83 25-27 t2 44" zl',S 171" 32',8 6.12.80 no data 0 ll 42" 00' s 174" 26', E 9.1.83 29-32 03 44" 08', S 171' 59',E 6.12.80 no data 0 2l 42" 04', S 174'08'E 18.1.83 2-7 13 44" 36', S 172" 36', E 7.12.80 no data 0 2 42" 05', S 174" 08',E 18.1.83 5-22 04 44" 27', S 172" 52', E 8.12.80 no data 0 l7 42" 04', S 174" 03' E 6.3.83 24-27 238 44" 09', S 172" 47', E 8.12.80 nb data 0 20 42" 03', S 174' 08',E 6.3.83 27-37 29 44" 0l' s 173" 24',8 10.12.80 no data 0 l0 42" 0l' s l'14" 25', E 6.3.83 27-35 03 44" 46', S 172" 16', E -.2.81 no data 0 70 42" 0'7' S 174' 08', E 14.3.83 4-lr tt4 44" 50'S 172' 15'E 1.1.82 24-30 0 5 42" 0l' s 174" 25', E 14.3.83 5 01 44" 47', S 172" l5',E 1.1.82 30 01 44" 47',S 172' ls',E t.t.82 30 0l 45" ll', S l7l" 25',8 5.2.82 t8-29 0 38 Banks Peninsula 44" 37',S 171' 30',E 19.2.82 18-33 0 67 Lyttelton -.8.80 t8-27 037 44" 44',S 171' 55',E t9.2.82 L2-33 0 40 43" t7',S 173" 20', E 9.12.80 no data l 13 44" 48'S 172" ll',E 19.2.82 20-23 0 7 43" 45', S 174" 01',E 10.1.82 t5-29 07 44" 45', S 172' l6',E t9.2.82 24-30 0 9 43" 45',S 173' 39',E 10.1.82 t4-33 04 44" 40',S 171' 33',E 24.2.82 6-11 0 9l 43' 50', S 173" 47', E t8.2.82 8-35 198 44" 44', S 171" 55'E 24.2.82 13-45 0 35 43" 45', S 174" 03',E 18.2.82 8-31 09 44" 38'S 171'29',E 13.4.82 25-31 0 17 43" 48',S 173' 59',E 18.2.82 16-27 016 44" 44',.S 171' 54', E 13.4.82 15-38 0 72 43" 48', S 173" 08',E 18.2.82 l8-34 0 11 44" 47',S 172" 15',E 13.4.82 t2-34 0 59 43" 50', S 173" 08',E 25.2.82 5-9 061 44" 06', S 172" 20', E t3.4.82 27 0 2 43" 48',S 173" 10',E t2.4.82 t9-37 09 44" 38',S 171' 35',E 1.6.82 29-41 0 43 43" 46',S 174'01',E 12.4.82 28 02 44" 43',S 171" 54',E t.6.82 8-12 0 50 43' 50', S 173" 08'E 31.5.82 l8-40 087 44" 47'S 172' 15',E 1.6.82 1l-40 0 40 43" 45', S 173" 48', E 31.5.82 30-32 02 44" 45', S 172" 21', E 1.6.82 13-19 0 3 43" 45',S 174'00',E 31.5.82 15-31 046 44" 43', S 171' 55'E 6.6.82 14-25 0 20 43" 46', S 174'02',E 31.5.82 t7 -33 08 44" 42',S 171' 56',E 6.6.82 3-506 43" 49',S 173" 08',E 25.7.82 5 01 44" 47', S 172" 20', E 7.6.82 3-5 03 43" 45', S 173" 47', E 25.7.82 2-4 28 44" 39', S 171' 34', E 26.7.82 7 01 43? 44', S 173'48',E 25.7.82 2,4 02 44" 42',S 171' 55',E 26.7.82 2-3 0 l8 43" 43', S 174" 02',E 3.8.82 l8-37 014 44" 42', S 171" 54',E 2.8.82 3401 43" 49',S 173'09',E 1.9.82 4-6 034 44" 39',S 171' 30',E 3.9.82 6,902 43" 49',S 173" 49',E 1.9.82 35 0l 44" 45', S 172" 20', E 3.9.82 22-33 0 3 43" 45', S 174'00',E r.9.82 19-30 047 44" 45', S 172" zr', E 3.9.82 2t-35 0 50 43" 49', S 173'09',E 9.9.82 6 01 44" 38',S 171' 30',E 8.9.82 2-6 0 29 43" 48',S 173' 50',E 9.9.82 15-29 034 44" 40',S 171' 35',E 8.9.82 2-4 0 5 43" 47' S 173'09',E 9.9.82 t9-26 09 44" 33', S 171' 35',E 1s.11.82 8-14 0 30

l3 44" 4I',S 171. 56'E 15.1 1.82 t3-29 05 Stewart Island 44" 44', S 172" 24'E 15.1 1.82 23-3t 02 46" 57', S 169. 37' E 2g.l.7g 17-26 0 23 44" 46',S 172" lg',E r 5.l l.82 t6-27 03 no data -.2.80 t6-27 0 61 44" 46', S 172" 25'E l5.r 1.82 t7-34 06 46" 56', S 16g" 51, E 9.2.90 28 44" 43', S 171" 8-13 0 55',E 23.11.82 4 02 47" 13', S 169. lg, E l3.l.g2 3001 44" 39', S 171" 3l'E 1.1.83 l2-t4 022 46" 56',S 16g" 25'E 13.1.92 24,35 0 2 44" 44', S 171. 52'E 1.1.83 35 0l 47" 15', S 167. 46', E 26.1.92 29-30 0 50 44" 46' S 172" 14, E 1.1.83 24-29 09 4',1" 20' s 16g. 17' E 1.2.92 23-3t 0 30 44" 47',S 172. 1g'E 1.1.83 27 01 47" 09', S 169. 25' E 21.2.92 t't-32 0 23 44" 50', S I72" l7'E 1.1.83 26-30 07 47" 00' s 169. 00, E 21.2.92 10-35 0 46 44" 40',S 171" 57'E 17.1.83 8-t2 042 46" 57', S 169. 25', E 21.2.92 tt-34 0 l0 44" 50',S 172" 1g,E 7.3.83 26,33 02 47" 0t' s 168. 55' E t5.4.92 28-30 0 4 44" 46', S 172" lg'E 7.3.83 22-38 013 46" 56', S 16g. 24' E 15.4.92 t9-33 0 6 44" 43' S 171. 56,E 7.3.83 rl-37 034 47" ll' s 169" 22' E 3.6.92 33-40 0 8 44" 40', S 171. 33'E 7.3.83 t7-37 032 47" 0l' s 169. 53' E 3.6.92 2601 44" 39', S 171. 35'E r 1.3.83 2-3 09 46" 56', S 16g. 23' E 3.6.92 30-40 0 50 44" 42',S l7l" 5g,E t2.3.83 6 01 47" 0l' s 16g" 53' E 4.6.92 2-4 0 20 44" 42'S 171" 56'E r2.3.83 9 01 47" 15', S 169. 19, E 30.7.92 27-35 0 19 44" 46', S 172" 2l,E 12.3.83 4 01 47" l0' s 169" 27, E 31.7.92 31 0l Dunedin 47" 15', S 169. lg' E 31.7.92 18-35 0 48 46" 57', S 16g" lg, E 5.9.92 10-23 0 30 46" 07',S 170. 54'E 12.t.82 26 0 I 47" 00' s 1ó9. 43' E 6.9.92 r-404 46" 08' S 170. 57'E 30,31 0 2 t2.l.g2 47" s 16g" 55' 7.9.92 170. 0l' E 2-15 0 4 46'08'S 44',E 23.2.82 20,30 0 2 47" 13', 45" S ló9. 20, E l7.ll.g2 29,32 0 2 59',S 170" 33'E 14.4.82 7-33 0 3 47" 00' s 16g. 55' E l7.ll.g2 2601 46" 07'S 170" 42'E 14.4.82 t6-36 89 0 46" 56', S 169. 22' E l7.ll.g2 5-605 46" 07',S 170" 54'E 14.4.82 18-30 0 76 47" l0's 1ó9.25'E l3.l.g3 2 46" 09',S 170. 54'E 15.4.82 22-37 t2,25 0 0 69 47" 0r's 16g. 55'E 13.1.93 2401 46" 09'S 170. 56'E 15.4.82 29,33 0 2 46" 56',S 169. 22'E 13.1.93 t20l 46" 05', S 170. 04'E t7.4.82 25 0 I 47" 0l' s 16g. 55'E 15.1.93 t2-14 0 5 45" 59',S 170" 31',E 2.6.82 6-17 0 20 47" l4',S 169. 19,E 9.3.93 24-37 0 5 46" 06', S 170" 42',E 2.6.82 27-39 0 6 47" 0l' s 169. 54' E 9.3.93 27,29 0 2 46" 07',S 170" 55',E 2.6.82 29-33 0 3 46" 56',S 169" 22,E 9.3.93 2t0l 46" 06',S 170. 55',E 2.6.82 I l-40 0 t2 46" 57', S 16g" 24' E 10.3.93 206 46" 09',S 170. 56'E 2.6.82 32,41 0 2 47" l6',S 169. 19,E 11.3.93 46' 08', S r70" 42', E 29 0 I 501 5.6.82 47" 16', S 169. 20' E 11.3.93 401 46" 08', S 170" 42', E 5.6.82 1-8 0 t7 46" 06',S 170" 43',E 1.8.82 r9-3t 0 4 46" 08', S 170" 55'E 27 1.8.82 t5-34 0 Chatham Rise 46" 07'.S 170. 57',E 1.8.82 33-36 0 3 46" 07',S 170. 54'E 1.8.82 t0-24 0 54 43" 38', S t77" 23' E 16.4.79 16-22 0 t7 46" 00' s 170" 32',E 4.9.82 17-28 0 28 43" 29' S 175" 00' E 11.12.80 no data 0 13 46" 08', S l70n 54',E 4.9.82 2t-34 0 l8 43" 46' S 174" 57'E 11.12.80 no data 7 0 46" 00' s 170. 2g',E 7.9.82 7 0 I 44" 08', S 175" 57'E 10.2.92 26-29 0 2t 46" 05',S 170. 46'E 7.9.82 2-3 0 6 no data -.8.83 2l-24 0 49 45" 59',S 170" 34',E 16.1 1.82 9,27 0 2 Chatham Islands 27.12.83 24-36 0 42 43" 40', S 175. 46" 05',S 170. 43',E I 6. I I .82 29,32 0 2 53',W 5.4.94 25-35 0 53 46" 0l' s 170. 30'E 12.1.83 2t-25 0 4 44" 22', S 176" 43',W 29.3.94 25-35 0 3l 46" 05',S 170" 45',E 12.1.83 27 0 I 46" l0's 170. 55,E 12.1.83 3l 0 1 46" 03', S 170. 46',E 15.1.83 4,23 0 2 Snares shelf 46" 03',S 170. 47'E 16.1.83 10-15 0 4 48" 15', S 168" l0' E rt.3.g2 15-33 0 36 46" 05'S 170. 56',E 16.1.83 t 0,l I 0 2 46" 04' S 170" 56',E 16.1.83 5 0 2 46" 06',S 170. 43'E 7.3.83 8-2t 0 Aucklands Shelf 46" 07',S 170. 54',E 7.3.83 9 0 49" 45', S 166. 05' E 2g.l.7g t7-30 045 46" 09'S 170" 56'E 7.3.83 3l 0 no data -.2.81 22-31 0 31 45" 58',S 170. 34',E 8.3.83 l9-33 0 49" 4r'. S 166. 49' E 24.3.92 15-33 028

V. R WARD. GOVERNMENT PRINTER, Vr'ELLINGTON, NEW ZEALAND_I985 36 t 79F-85PT