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(966I) tSZ-902r(Z)0telsl,{nN G( NuytsiaVol. 10,No. 2 (1995)
Introduction
,,Flora The principal aims of this paperare to describenew taxa aheadof their inclusion in the of Australia' lxeatmeotof Acaci4, and to presentre-descriptions of some previously described taxa related to the new ones; we have also lectotypified nameswhere necessary.
Methods
Arrangement of the text. The taxa included here are all refenable to Acacia secttonplurinerves (Benth.) ,,A. Maiden & Betche and are treated in five groups, the densiflora Group,,, the "A. ,'A. ,A. ancistrophyLlaGroup",the"A. enenra Group',,the fragitis Group" andthe dielsil Group,,. Apart from their globular heads on axillary peduncles these taxa have rn common onlv their microneurousphyllodes (cl Maslin & Pedtey 1988for definition), having usually numerous,closely parallel, longitudinal neryeswith anastomosesabsent or rare. It should be noted that these,,GrouDs', are simply mnemonic devices adoptedto bring togethertaxa that are consideredto be most closelv related to one anotherfor purposesof communication. Theseare informal groupingsand not all thl taxa which could be referredto them are necessarilyincluded herein. Furthermore,the,,Grouos', are sometimesseparated by only small differences,especially in the case of the firsr thre€, and future classiticationsmay well combine theseinto a single infragenerictaxon.
Teenomic rank. orr approachto the application of rank is briefly discussedin cowan & Maslin (l995).
Typncafion. Our approachto typification is discussedin Maslin & Cowan (l994b).
Conservationstatus. We have assessedconservation status of eachtaxon included in this treatment. In the caseof western Australiantaxa we have usedthe conservation codes for western Austalian Flora (seeend of this issue). For taxa occuning elsewherein Australia we haveused the conservation codingsof Briggs andLeigh ( t988.).
Measurements. All measurementsand observationswere made from dried specimensunless stated otherwlse.
The ..Acacia densiflora Group',
Although they have no unique distinguishing characters,the speciesof this,,GrouD" can be recognizedin sectionPlurtnemes by acombination oftheir tomentulosebranchlets (excepi glabrous rn A. papulosa and appressed-puberulousin A. tlissona and somematerial ofA. erenophiial-with the indumentum often extendingto the adaxialsurface ofthe pulvinus, cucullatebasal peduncular bracts and their small headson short peduncles. The taxa refened to this Group include the following: A. densiflora Morrison, A. drssora R.S. cowan & Maslin sp. nov. (including var. dissona andv;r. indoloria R S. cowan & Masrin var. nov.), A. eremoph aw. Fitzg. (including var. eremoph a and var. variablis Maid'en& Blakely), A. hadrophyua R.s. cowan & Maslin sp. nov., A. kargoorriensis R.S. cowan & Maslin sp. nov.,A. mackeyanaEwart& Ieanwhlte, A. papuiosaR.s.cowan & Maslin sp. nov. and A. undosa R.S. Cowan & Maslin sp. nov. " psopun v'8 ruult-I x st-02 tt ,zttrqdolwq.v.b ...... ::]::::::*::::I :] ::::::::::llff H-: 'ctdl I::Tl o-Suotqosopo ,4qd:eplnpun lou spod €I snorq"ltssepunped zl orclnsuap y I"""" ,(Ir?qselcunped'Zl leu sapoll^rld 'l ' ouosstp're|Duossrp V UZ 3uo1urur g-7 selcunped:sdg tueSund oqsol pelc?Iluocxade oql qlr,$Buol wx V-Zsapo11,{q6 11 'V nsuagtooS\o>19""""""""""" 3uo1ruur g I-90.{lprurousalcunpad :slurod esourds Suololur potrsrlordxedu eqt qlr,,n3uo1 ruc 1-g sepolli(qd'tt 'lq8rcrls pere/olJ-ZZ-gI speaq :3uol uul 0r-02 'xede lo/puvsnur,rlnd o1 peredel ,{lpnper8 sepo11,tq4 91 "" ouot lJDw V 9""""""" pete/holJ-Zl-LspEeq 'pe^JnooJ 'xode :3uoluruJ 0g-, ,{luoururoc pu? snunlndqloq luo{ peluredasd11dn.rqu sopulq opoll,{qd 0I lue8undfld.mqs sepolli(q4 1 'lr-n 'i(rrcq flUqollD^ olltqdowatay qg" Euo1ruc 11-9sepol.tqd :etelnpun spo4 6 'll't "' Dllqdotuata o1lqdowatay E€ Suoyuc 9-g sepollfqd:e1e1npun 1ou'snorqelE spo4 6 Jnolocur urroJrunsecuds o,rreu-ro1ur pue selteu 'posseJdop 'aleJel + se^JeuiEuol uJur €-I snur^lnd 8 'JcA 'qZ ollqopu! DuosslpV sacudse,uau-lalur ueq reled,{poruruoc se^Jau pesluJ qtla sepolt,(qd 'esuq eJ$erx:Euol urur Z-9 0 lu pepuedxesnur^lnd 8 lue8und,{1d.mqstou sepollfqd , os rsolup.ro a:r; spda5 g 'y """ """'ser\Jeu (luau€^pe^Jeu-snoJplu',nru) Dllqdouala € uBr{tJopri\i ,{lacrecssacuds e,r:eu-relur 'snoreurnu seAJeN 'g """ ',{rl€q s !qD!.tD^'lr-rollrqdowata y qg 3uo1urc 11-gsepollfqd ielulnpun spod 9 ,(J 'V '"€ """" " " ollrldowatan^ Dlllldouah Euo1ruc 9-7 sepo[,(qdlelBlnpun lou 'snorqElSspod '9 ,(t, '(se^reu secudss,ueu-:elur pesrcr Jo qlpr,r sarup peorq^roleq pesserdep'01 r saAJeN'g ,,,, lae^es) eqt ,(rnq ro snorqelEselcunped :sd! (lq8tets seruneuos) gaA apurcun-qnsol opurcunluo8und-uou qlr,r sopollrqd t otoyfisuap'V I eleFpunlou'.&leq spod:^Ir€q salcunpad :sdnlq8rcr1s + lua8und,(lasreoc o1 fldreqs qr,,n sopoy1,4g4 y 43ue1rraqt V lseel$ roJpelrun spdeg g frmq slelqcuerg g nso|ndod'y 7""" sno;qe16slalqcuerg 6 ")rui elaJet-qnsol eleJelsepo[fqd 'I Joe
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L. Acacia densiflora Morrison, Scott.Bot. Rev. 1: 96 (1912): Trans.& Proc. Bot. Soc.Edinbureh 26: 51 0917) II tl Typas: Kellerbenin, E Avon district, Westem Australia,September 1897,R.8. Iea,te (iso: K, NSW, d PERTH 01017489fragment ex E). al p Dense,rounded or obconic s/irubs (0.2)0.5-I.Z(1.7) m tall, few- to many-sternmedfrom ground Ievel. Bark grey, smooth, sometimesfissured basally. Branchlets tomentulosewith short, dense, crisped white hairs, sometimesreddish resin-hairs intermixed. New shoots with grey, a matted pubescenceand with many rust-coloured resin-hairs. Stlprles persistent,subulate to triangular, 0.5-2.5 mm long, t puberulous. Phyllodes terete, sub-tercteor compressedto flat and linear, 25-50(60) mm long, l-2 mm wide, rigid, inclined to ascending,straight, glabrous except tomentulose on pulvinus and baseof phyllodes, sometimessparingly appressed-puberulous but glabrescent,dark tr green; apexending in a straightor slightly curved,sharply to coanely pungentpoint;t 16 neruesplane p to slightly raised, closely parallel, sometimesinegularly venuculose, stomataraised; gland small, 1 located on adaxial margin near the middle of blade, occasionally a second gland on upper a 1/2 of blade. Peduncles2 per axil,0.5-1.5 mm long, crisped-tomenttiose; basal peduncular bract T caducous,cucullate, 1.5-3mm long,villose; heads globular, 4-5 mm diam.,(l l ) l5- l8(25)-flowered: I bracteoles oblanceolateto obovate,arcuate, puberulous and ciliolate, Flowers 5-merous.Sepals less I than half petals, pods as long as l/2-3/4 ufited. Petals 112united. linear, raised over and slightly r constricted between seeds, to 4.5 cm long and 2.5 mm wide, thin-crustaceous,slightly curved, sometimes slightly tlexuose, tomentulose and with many red resin hairs. ,Seedslongitudinally anangedin pods,efliptic, c. 3 mm long, 2 mm wide, I mm thick, glossyblack pleurogramlJ-shaped; areole smalli arll terminal, short-tubularwith scallopedmargin, 2 mm long, white.
Selectedspecimens examined. WESTERN AUSTRALIA: 30 km W of90 mile Tank, Frank Hann National Park, T.E.H. Aplin andM.E. Trudgen5918 (K, MO, NY, PERTH); about0.5 miles [0.8 km] N of Bendering on road to Narembeen,B.R. Maslin 5l'7 (MEL, PERTH); l8 miles [28.9 km] W of Pederahon road to Kdin, B.R. Mqslin 532 (K, MEL, PERTH); 8 km dueW of M er:Iedin,B.R. Maslin 2352 (CANB, K, NY, PERTH); l0 km due SW of ChiddarcoopingHill, B.R. Mastin 6379 (CANB. K, PERTH); 15 km SSW of QueenVictoria Rocks, c. 60 km SSW of Coolgardie, K. Newbey 56gl (PERTH); 4.6 km E of GrahamRocks turn-off on Hyden-Newdegateroad, ,/.C. antJM.H. Simmons 1325 (PERTH); 97 miles u56 kml E of SouthernCross, E. Wittwer 1298 (PERTH).
Distribution. southwest wesrern Australia from near wyalkatchem and chiddarcooping Nature Reserve(c. 80 km north-northeastof Merredin) south to ongerup and Frank Hann National park (which is located 30-110 km east northeastofLake King). A few collectionshave beenmade e ast of the main distribution in the Coolgardie area (near Coolgardie and eueen Victoria Rocks).
Habitat. h sand and loam, occasionallyclay, mostly in open mallee shrubland.
Flowering and.fruiting periods. Flowers from June-September;pods with matur€ seedshave been collected in December.
Afrnities. Acacia densiflora differs from most of its relativesin its partly united sepalsand petals; other charactersuseful in recognizing the speciesinclude its very sho , tomentulosepeduncles, denselytomentulose branchlets and many-nervedphyllodes with commonly straight,sharply pungent Doints.
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9l ,{uellerslwrrrrey'ullsew U g pu! ue^\oCS U 270 NuytsiaVol. 10,No. 2 (1995) pedunclesare glabrous and tie perianthparts are free. Acaciamackeyana, however, has phyllodes whicharecommonly recurved and abruptly contxacted at boththe apex and pulvinus, as well ashaving 7-12-floweredheads and sub-teretei thick-walled, crustaceous pods. ll Thereis considerablesimilarity also between A. dissona,particulaxly the typical variety,and A. kalgoorliensisR.S. Cowan & Maslin(see below) which has generally longer phyllodes with the apexdrawn out into a longspinose tip, mosdyshorter peduncles, pods with appressed,red resin-hairs 7 andseeds with an aril thatdoes not investthe seedaDex. (r
Infraspecific taxa. The differencesseparating the two vadetiescomprising the speciesare mostly to 1 befound in thepods and seeds. There are, however, qualitative differences in thedegree ofpungency l of the phyllodesand the colourof thenerves. A t " Etymology.The namegiven refers to its discordantnature, in relationto its closerelatives (from t dissonus,Lath for discordantor different). '1I ( 2a. Acaciadissona R.S. Cowan & Maslinvar. dissona, s I Phyllodes2-4 cm long,sub-terete, contracted to short,more or lesssharply pungent tips, nerves andinter-neNe spaces uniform in colour,stomata sometimes tuberculate. Peduncles 2-5 mm long. Padsdistinctly constricted between the seeds, sparingly appressed-puberulous. Seeds 4.5 mm long; aril long-conical.
Sekctedspecimens examined. WESTERN AUSTRALIA: 45 km E of Norseman,T.E.H.Aplin L8l1 (PERTH);Muntadgin, E.T. Bailey 690 (PERTH); 2l.6 kmENE of Merredinby road, R. Coveny8340 andB. Haberley(PERTH); 5 km SEof Coorow,A. Doley4 (PERTH);Rabbit Proof Fence, Wanara, C.A.Gardner 13897 (PERTH); New Zealand Gully, Southern Cross, S. Grayling | (PERTH);14 km N of LakeGrace towards Kulin,8.R. Maslln 4067 (PERTH);c. 4 kn S of Karlgadnon PederahRoad, B.R.Maslin6738 (PERTH): 9 km WSW of LakeCairlocup, K. Newbey4333 (PERTH);4.1 km from Hydenon road to DragonRocks,,/.G. and, M.H. Simmonsl3l9 (PERTH);between Hyden and Kondinin,D.l.E. Whibley3355 (PERTH).
Distribution. Discontinuousin southwestWestern Australia in the Menedin-SoutheroCross area southto nearOngerup with outliersin theCoorow-Wanarra area (c. 250km northwestof Menedin) andin the Norsemanarea (c. 350 km northeastof Ongerup).
Habitat. Growson clay,loam, and sandy soils in eucalyptwoodland or mallee,sometimes dominated by Eucalyptussalmonophloia.
Flov)eringand fruiting periods. Most floweringspecimens have been collected in Septemberand October,but the no hernones (from theCoorow-Wanara area) were collected in July andAugust. Podswith matureseeds have been collected in December.
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Dense, rounded to obconic shrubs usually 0.4-2 m tall and about the same across. Bark grey, fi smooth. Branchletsterete, slightly ribbed, white- or grey- sub-appressed-tomentulose(often with dark resin-hairs intemixed), appressed-puberulousor cdspate-sericea\rs.Stipules persistent,subulate- triangular,0.4-0.8 mm long. PhyLlodesterete , 2-11 cm long, 0.6-1.5 mm diam., rigid, patentto erect, s shaight, glabrous except tomentuloseon pulvinus and phyllode base, green to greyish-greenor 7 yellowish green, sometimest viscid when young; dpex commonly reflexed uncinate,rarely straight 1 or sub-uncinats;pulvinustercte,l-3 mm long; nervesmostly 9 or 10,closely parallel, depressed below ,t. raisedinter-nerve spaces bearing raised stomata; glard small, mostly at or below middle ofphyllode. I Peduncles 2 per axil, 1-2 mm long, t tomentulose,sometimes also with red resin-hairs; Dasal I\ peduncular bracts caducous,cucullate, 1-2 nrn long, glabrous or hairy, red resin-hairspresent or E absent;lreads globular, light-golden,3-4 mm diam., l0-25-flowered,;bracteoler spathulate,ciliolate. r Flowers 5-merous. Sepalsmore or lessunited to 1/2 their lengthbut sometimesonly basally,or some 2 even free, narrowly oblong, ciliolate. Petals free, oblancaolateto obovate. PodJ linear, raised over L and somewhatconstricted between seeds, 2-5 cm long, 1.5-3 mm wide, thinly crustaceous,staight ri or undulate, tomentulose, appressed-puberulousor glabrous, marginal nerve distinct. Seeds I longitudinally aranged in pods,elliptic to oblong-ovate,2.5-3 mm long, L5- 2 mm wide, dark brown; 1 ariL white, terminal, to 2/3 length of seed. It
Distribution. Widely distribut€d throughoutsouthem inland W€stern Australia. L n Notes. When Ewart & Jean White published A. LeptoneurqBenth. var. eremophila, they cited n A. eremophiLaW.V. Fitzg. as being in manuscript;at the varietal rank we would cite Ewart & Jean White as the publishing authorsbut the first availablename at the specific rank is Fitzgerald's name.
As is otien the casewith Max Koch collections,there is someconfusion because the numbershe I attachedto collections were taxon numbers,not collection numbers. Although the protologue lists c Koch 1024 and lO24a as the basis lor A. eremophiLa,this is clearly an eror; Koch's own notes in archivesof the Western Australian Herbarium llsts ^ Crevillea speciesfor 1024 andA. eremophila t for lO24a. Further discussionmay be found in the paperon Fitzgeraldtypes by Maslin & R.S. Cowan l (1994^). F u A|finities. Related to A. densillora Morrison (see above) which has c. l6-nerved phyllodes and 1/2- to 3|4-!Jnitedperianth parts. Acacia eremophiLaoccurs as two varieties,separated on pod shape and hairinessas well as phyllode length, pedunclepubescence and number of llowers per head. S I Variation. This species is reasonablyconstant morphologically, although there is a variant, not encompassedby the abovedescription, which differs from typical A. eremophilain having phyllodes with more numerousnerves and consequentlynarrower inter-nervespaces. This "numerous-nerved variant" hasa sporadicdistribution and hasbeen recorded fiom Edjudina Station(c. 130 km northeast of Kalgoorlie) (M. Blackwell 47 - PERTH), ths Great Victoria Desert betweenNeale Junction and Plumridge Lakes (8.R. Maslin 5101 - K, MO, NSW, PERTH) and from between Norseman and Balladonia (K Newbey1429 - PERTH). It was treatedas A. rlgens Cunn. ex Don by Maslin (1981) "Flora in the of Central Australia" and soecimenswere distributed as such.
3a. Acacia eremophila W. Fitzg. var. eremophila
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Shrubs1.-1.6 mtall. Phyllodes5-11 cm long,1-1.5 mm diwn. Peduncles2 mn long,sparsely to denselytomentulose or appressed-puberulous,resin hafus rarely Vesentiheals 20-25-flowercd.Pods to 5 cm long, 1.5-3mm wide,distinctly constricted between seeds, loosely undulate, tomentulose or appressed-puberulous.Seeds oblong-ovate.
Otherspecimens exanized. WESTERNAUSTRALIA: 94 miles[151 km] E of Norsemanon Eyre Highway,I. Arrnitage562 (PERTH);between Balladonia and Norseman, 20.3 km E of tumoff to NewmanRocks, R. J. Chinnock3002 (PERTH); 20 miles [32 km] S of Menzies,1975, N. Pratt s.n. (PERTH00700541); 34.4 km W of Balladoniaon EyreHighway, M.H. Sirnnons1162 (PERTII); 5 km E of Zanthus,P.G. Wilson7623 (PERTH).
Distribution: This poorly collectedvariety of southernWesten Aushalia is known from only three widelyseparated localities, near Balladonia, near Zanthus (c. 160km northofBalladonia) and Comet Vale (c. 85 km north-northwestof Kalgoorlie).
Habitat. Prcbablygrows in sandor sandyloam in low woodlandor scrub.
Flowering andfruiting periods. Flowersin September;pods with matureseeds have been collected in December.Additional fruiting collectionsare needed to confirm the separationof this vadety ftom the typicalone.
Typification.Choice ofa lectotypeis madebecause several collections were cited in theProtologue, somein flower,others in fruit, eventhough all apparentlyrepresent the sametaxon.
Variation. One specimenof this variety(Pratt s.n. from southof Menzies- PERTH)records a maximumheight of 3-4.5m for the taxon,which is considerablylarger than is tyPical.
Conseryationstatus. A Priority3 taxonin theDepartment of Conservationand Land Management's DeclaredRare and Priodty Flora List.
4. Acaciahadrophylla R.S.Cowan & Maslin,sP. nov.
Frutex (20\30-50(70) cm altus, ramulis tomentulosiset cum micro-piliis nigris, interdum plus minusvesericeis. Stipul.ae persistentes, anguste ffiangulares, 0.5-0.8 mm longae.Phyllodia plana, oblongo-elliptica,obtusa vel subobtusaet cum paryomuuo, pulvino0.8-1.5 mm longo,a lamina abruptesepamto, valde crassa, rigida, (5)7 -25 mmlonga,2,5-5.5 mln lata,patentia ad inclinata,saepe leviter incurvata,glabra praeter pulvinum tomentulosum, uniglandulosa, in quoquesuperficie 5-7 nervata,nervis valde elevatis, distantibus. Pedunculi binati, glabri, 0.5-2 mm longi,pedunculorum bracteisbasalibus cucullatis, rostratis, 2.5-4 mm longis;capitula 3-3.5 mm diametroin sicco,14-25- floribus,fores5-meri, sepdlis discretis vel Paxtimconnatis, petalis plus minusve discretis. Legumina linearia,inter semina plus minusveconstricta, L2-22 mm longa,2 rm lata,puberula, plus minusve cttllatA. Seminqlongitudinalia, oblongo-elliptica, 2.5-3 mm longa, L-Z mm lata, brunneo-nigra, arillo terminali.
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'S'd 9t {uDIImsrWtTrrry'urtstnl U g pue tl0.^\o3 216 NuytsiaVol. 10,No.2 (1995) R.S curyatum attenuata,pulvino 2-3 mm longo, cdspato-sericeo,laminis 3-7 cm longis, plus minusve 1.5 Nc mm diametro,rigidis, erectis,rcctis, glabris, 20-nervatis,nervis indistinctis,stomatibus plus minusve M( elevatis,glande saepe2. Pedunculi 2 in quoqueaxilla, 0.5-L5(5) mm longi, glabri vel resinoso-pilis thr dispersisrubris ornatis; capitula globularia ad late ellipsoidea,3-3.5 mrn diametro, l5-22-floribus, fro bracteolis linearibus ad spathulatis,tomentulosis et ciliolatis. Flores S-meri. Sepala longitudrne 1/2 petali partes aequantia,discreta, villosulosa. Petala disqeta. Ovarium paptlloso-puberulum. Legumina linearia, supra semina elevata ct inter semina constricta, ad 7.5 cm longa, 3 mrn lata, ott chartacea,leviter curvata,appresso-puberula, pilis minutis,resinosis. Seminalongitudinalia, anguste elliptica ad oblongo-elliptica, 4-4.5 rnm longa, 1.8 nm lata, I mm crassitie,hebetato-nigra, arillo Afi terminali. un pr( Typus: 4 rniles [6 kml N of Broad Arrow towardMenzies, Western Australia, 10 August 1971, wt B.R.MasLin 19lO (holo. PERTH 00194905;lso: CANB, K, NSW, NY). it. ph Dense, rounded shrubs l-3 m tall, many-stemmedfiom ground level, the main stems slightly twisted. Bark grey. llranchLets sub-appressed-puberulous,the hairs white, curved antrorsely or in somewhattwistedand with redresin-hairs intermixed. Phyllodes terete,3-7 cmlong,c. 1.5mmdiam., lm dgid, erect, straight, green, glabrous except ibr the crispate-sericeouspulvinus; ap€x tapering into long, hard,dark, straightor rarelycurved tip, sharplypungent; pulvinus 2-3 mm longi t 20 nerves slightly raised,closely parallel;stomata somewhat raised; glands usually 2, one nearmiddle, the other near the phyllode apex. PeduncLes2 per axil, 0.5- I .5 mm, rarely to 5 mm long, glabrous or with Dr scatteredred resin-hairlets;basal peduncular bract cucullate,rostrate,2.5-3 mm long, appressed- puberulous;heads globular to widelyellipsoid, golden, 3-3.5 mm diam.,15-22-flowered; bracteoles EI linear to spathulate,tomentulose and ciliolate. Flowers S-merous.Sepals 112lengthof petals,free, Ki linear to spathulate,villose. Perals fiee. Pod,rlinear, raisedover and constrictedbetween seeds, to 7.5 cm long,3 mm wide, chartaceous,shallowly curved, appressed-puberulous with minute,red- brown resinhairs. Seedslongitudinally aranged in pods,nanowly elliptic to oblong-elliptic,4-4.5 mm long, 1.8 mrn wide, I Inm thick, dull, black;pleurogram U-shaped; areole raised, shiny; aril LI terminal. (\ Cr Selected specimens examined. WESTERN AUSTRALIA: Hampton Hill Station, 23 km E of 0( Kalgoorlie, R. Coveny 8426 and B. HaberLey(NSW, PERTH); near Kanowna, C.A. Cardner 783 (PERTH); 22.4 krrr E of Kalgoorlie along road to Zanthus, January 1991, M. McDonald 1338 (PERTH); 8 rniles 12.8 kml SW of Kalgoorlie towardsCoolgardte, B.R. Maslin 1897 (MEL, MO, [ \1 PERTH); about 16 km NNE of Kalgoorlie on road to Edjudina Station,B.R. Maslin 4847 (PERTH); 28.5 km SSE of Marvel Loch on track towards Mount Day, B.R. Mqslin 5514 (PERTH). pl Distribution. Restrictedto nearKalgoorlie (with collectionswithin a 50 km radiusof Kalgoorlie) and BI the Marvel Loch area (c. 200 km wsst-southwestof Kalgoorlie), with a variant scatteredc. 320-560 al km NW of Kalgoorlie (near Wubin and Noongal and Yuinmery Stns),W.A. h: br Habitat. Rocky loam andclay on slopesof low hills in eucalyptwoodland. The variantgrows in sandy c( loam and loam over calcretein eucalyptopen woodland andAcacia open scrub. st al Flowering and Jiuiting periods. Flowers from August to October;pods with matureseeds have been S( collected in January. pl 2. Variants. Four collectionsare clearly relatedto A. kaLgoorlierxsi$but may representadistinct species. Thesecollections differ by having widely ellipsoid, 3O-4O-floweredheads on peduncles2-4 mm long:
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218 N'.ryl\i4vol. 10,No. 2 (1995) to 1/2 as long as petals,free, linear,ciliolate. Petals ftee, elliptic, glabrous. Podr sub-terete,oblong- linear, 15-35 mm long, 2-2.5 mm wide, straight or slightly curved, thick-crustaceous,glabrous, longitudinally nerved. Seedlongitudinal, narrowly oblong, 3-4 mm long, 1-2 mm wide, dull, dark brown to black, the adl terminal with scallopedmargin.
Selectedspecimens examined. WESTERN AUSTRALIA: 5.5 km E of Tammin, R. Coveny8311 and B. Haberley (PERTH);2.5 miles [4 km] N ofcadoux towardsKalannie, R. Cumming 1868(PERTH); I I miles u7.6 kml N of Wyalkatchemtowards Koorda, B.R. Masltn 155 (PERTH);7 miles [11.2 km] W of Moorine Rock on Great EasternHighway, B.R. Maslin 583a (NSW, PERTH); 5.5 km NW of Wongan Hills towardsPiawaning, B.R. Maslin4203 (PEP.TH);29km NW of Kulin towardsCodgin, B.R.MasLin 43'7 2 (PERTH); nearSnake Soak, c. 11 km dueSW of Wialki, B.R.Maslin 4464 (PERTH); I km E of 90 Mile Tank, 32' 39'S, 120' Al'F,,B.R. Maslin 5'189(MEXU, PERTH); Chiddarcooping Nature Reserve,B.R. Masllr 6380 (CANB, MEL, NSW, PERTH, Z); Davies property, l0 km due N of Coorow,B.R. Maslin 6577 (PERTH);Nugadong Reserve 12614,21 km SEof Wubin, B.C. Muir 206 (1.5)(PERTH);3 niles [4.8kml E of Billericay,K Newbey3237 (CANB, K, MEL);35 km from Ravensthorpetowards Lake King, R. Perry 674 (PERTH);CemeteryRoad, Yorkrakine, M.H. Simmons 1274 (PERTH); Mullewa, 3 July 1952, N.H. Speck (PERTH 00884677); 20 km NE of Ongerup, N. SteyensKRN9520- I (MELU, PERTH); 5.9 km SE of Bruce Rock on main road to Narembeen, M.D. Tindale 3736 (CANB, K, MEL, NSW, PERTH, US).
Distribution, Widely distributedin southwestWest€rn Australia from nearCoorow southeastto near Corrigin and Moorine Rock (which is c. 30 km west-southwestof SouthemCross on Creat Eastern Highway), with a north-western outlier at Mullewa and south-easternoutliers in the Ongerup, Ravensthorpeand Frank Hann National Park areas(c. 200 km southeastofthe Conigin-Moorine Rock area).
Habitat. Gravelly soil, loam, loamy clay and sometimessand in eucalypt woodland and mallee communities and also in MelaLeucauncinata thickets.
Flowering and fruiting periods. Flowers f'rom June to Augusti pods with mature seedshave been collected in Decemberand January.
Typirtca on. In the protologueof A. mackeyanaEwart& J€anWhite cite only onecollection, M. Koclr 1013,even though both flowers and fruits were described. As was noted underA. eremophila above, Koch regularly followed the practiceof assigningtaxon numbersto his collectionsand consequently the samenumber appearson quite diff€rent collections;in this instance,although both collections are from the samelocality and certainly the sametaxon, it is desirableto fix the applicationof the name. Consequently,we have selectedthe flowering specimenon MEL sheet#l l? 106 as lectotype.
Affinities. Acdcia mackeyanuis related to A. densiflora which has partly united petals and sepals, r tomentulose pedunclesand staight, generally longer, terete to tlat phyllodes. Also related to A. dissona which has straight phyllodes, 15-20-floweredheads and linear, thinly crustaceousor coriaceouspods.
Discussion. Although the authorsof the protologuedescribed the petals as 3/4 united, we have not seenthis condition in any of the specimens,including the type collection.
ConservqtionJtal,rs. Widely distdbuted, not under threat.
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9 I ,{uellorsrr{,,rrry'urlwt{ U gpuv uE^loJ S f, Nuytsiavol.10, No. 2(1995) tipped phyllodes, heads3-4 Inm in diameter, spathulatebracteoles, sho er, hairy peduncles,non- resinouspods, elliptic to oblong-ovateseeds and it grows in loam or sand.
Discussion. C.A. Cardner's two collectionsof the species,as nearly as we can decipherhis notesin his fieldbook (in the PERTH library archives),were made along theFitzgerald River liom the crossing of Colletts Road north about 6.5 km. That therehave only beentwo collectionsin the FitzgeraldRiver National Park is surprisingin view ofrelatively thoroughcollecting in the areaover the pastdecade, especiallyby K. Newbey who madetwo collectionsfrom a secondpopulation nearBoxwood Hill.
Conseruationstutus. A Priority 2 taxon on the DepartmentofConservation and Land Management's Declared Rare and Priority Flora List.
EtymoLogy.The specific epithet is basedon the conspicuouspapulose surface of the branchletsand the inter-nerve spaceson the phyllodes (from papulosus,Latin for pustulateor pimply).
8. Acacia undosaR.S. Cowan & Maslin.sD. nov.
Frater densustholifbrmis vel obconicus0.3- 1.5 m altus,ramulis tomentulosis et cum micro-piliis nigris. StipuLaepersistentes, subulatae ad subulato-triangulares,0.6-2 mm longae.Phyllodialineatia ad lineari-oblanceolata,20-45 mm longa, 1-4 mm lata, oblique mucronata, innocua ad grosse pungentia, raro acute pungentia,rigida, patentia, recta ad leviter incurvata, glabra, saepe 2-glandulosa,nervis omnino circa 20, arcte parallelis,stomatibus valde elevatis. Pedunculi binati, l-1.5 mm longi, glabri; pedunculorumbracteae basales ovatae, cucullatae, rostratae, 1.5-3.5 mm longae,villosae et ciliolatae;capitula globularia, 3-3.5 mm diametro, 18-20-floribus. F/or€s pentameri; sepafadiscr€ta vel l/2-connata.Legumina linearia, valde undulata, 10-40 mm longa, 2.5 mm lata, papillata. Semina longitudinalia, elliptico-oblonga ad lat€ oblongo-elliptica, 2.2-2.7 mm longa, 1.2-1.8 mm lata, atro-brunnea.
Typus: 0.5 km S of Belka Siding (between Bruce Rock and Merredin), Western Australia, 14 December1971, B.R. Masl.in 2361 (lolo: PERTH 00191450;lso: CANB, K, NSW).
Dense,domed or obconicshnrbs 0.3- I .5 m tall andspreading to aboutthe sameacrcss. Brqnchlets tomentulose,the very short hairscurved to crispedand with black micro-hairletsintermixed. St,pules persistent,subulate to subulate-triangular,0.6-2 mm long. Phyllodes linear to linear-oblanceolate, 20-45 mm long, l-4 mm wide, rigid, spreading,straight to shallowly incurved, dull green, glabrous except for tomentulosepulvinus; apexobliquely mucronateto sub-uncinatewith a hard, innocuousto coarsely pungent point, rarely with a straight,sharply pungentpoint; pulvinus 1-2 mm long; nerves c. 9 per face, 20 in all, closely parallel with very occasionalanastomoses, plane or slightly impressed, the inter-neryespaces with stronglyraised stomata; glands not prominent,I or 2, one at or nearmiddle ofphyllodesand the other on upperhalfofphyllode. Peduncles2peraxil, 1-1.5mm long,glabrous; basal peduncular bract ovate, cucullate, rostrate, 1.5-3.5 mm long, villose and ciliolate; heads globular, showy, golden, 3-3.5 mm diam., 18-20-flowered; bracteoles oblanceolate to linear- oblanceolate,ciliolate. FLowers5-merous. Sepals linear to oblong,free or to half-united,puberulous at apex. Pods linear, strongly undulate, 10-40 mm long, 2.5 mm wide, papillate,the marginal nerve distinct, much lighter in colour. Seedslongitudinally arrangedin pods, elliptic-oblong to broadly oblong-elliptic, 2.2-2.'7mm long, 1.2-1.8 mm wide, dark brown, the aril terminal.
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'urlsBh] I I ,{u[llersrr,\lDrrr, y U gpnD u!^\oJ.S U (s6 222 NuytsiaVol. 10, No. 2 (1995)
"Group" The is closelyrelated to the'4. enerviaGrotp" whosemembers lack the red-brownresin- "A. hairs so common in the ancistrophylLaGrotp", as well as having longer pods.
Key to taxa of '14. ancistrophylla Grotp"
1. Floweringpeduncles 6-15 mm long;pods nanowly oblong, 5-7 mm wide; seedsoblique in pods;phyllodes 2.5-8 mm wide. South Australia .....3.A. whibletana 1. Floweringpeduncles 1.5-7 mm long; podslineiu, 2-3 mm wide; seedslongitudinal in pods;phyllodes 1-5 mm wide. Western Australia (except A. qncistrophyllu var. Ii.ssophylla) 2. Phyllode tip sharply to coarselypungent, the nervesstrong, discreteand palerthan inter-nerve spaces ...... l. A. amyctica 2. Phyllodetip not pungent,the nervesobscure to a evident, not paler than intgr-nervgspaces 3. Podsstrongly arcuate to openlyonce-coiled; phyllode tip roundedwith a minutemucro ...... 2c.A. ancistrophylLa\at. perarcuata 3. Podsstraight to shallowlycurved; phyllode tip acuteto subacute 4. Phyllodes(7)10-20 tines long€rthan wide; podsglabrous. Mostly SouthAustralia, infrequent in WesternAustralia and northwestVictoria...... 2b. A. ancistrophylla var. Lissophylla 4. Phyllodesmostly 3-8 timeslonger than wide; pods t appressed-puberulous.Western Australia ...... 2a. A. ancistrophyLlayqr. ancistrophyLla
l. Acacia amyctica R.S.Cowan & Maslin,sp. nov.
Frrter obconicus0.7-1.5 m altus, coronaaperte ramosa, l-1.5 m diametro,ramulis leviter angulatissparse vel parceappresso-puberulis. StipuLae cadtcae, Phyllodia angusteoblanceolata ad elliptico-oblanceolata,acuta vel sub-obtusa,mucronata, grosse ad acute pungentia, versus basem angustataad pulvinum0.5- I mm longum,laminis l5-25 mm longis,2.5-4 mm latis,ratione horum 3.5-7,rigidis, ascendentibus ad erectis,rectis ad levitercurvatis, glabris, obscure medio-viridibus; nervi numerosi,arcte paralleli, valde discreti,pallidiores quam spatiis internervis,stomatibus distinctis,plus minusveelevatis, glande una, parva, inconspicua, basali. PeduncuLi bir'ati in quoque axilla,4-7 mm longi,plus minusve appresso-puberuli vel glabri.Capitula globularia, aurea,3-3.5 mm diametro, 18-25-floribus, bracteolis obovatis ad spathulatis, ciliolatis. Flores 5-meri, sepaLis longitudine l/3 petali aequantia,discretis oblongo-spathulatisciliolatis, petalis discretis glabris. Ovarium albo-appresso-puberulurn.Legumin& (rmmatura et valvis apertis)linearia, inter seminanon constricta,ad 6 cm longa, 3 rnm lata,firme chartacea,valde curvata, glabra, marginibus pallidioribus. Semlna longitudinalia, non visa.
Typusr I km N of SalmonGums on Coolgardie-EsperanceHighway, Western Australia, 25 September 1983,B.R. Maslin5448 (iolo: PERTH 00700517;lso:MO, NY, distdbutedas Acacia ancistrophJlla).
Obconic sirubs 0.7-1.5 m tall with openlybranching crown 1-1.5 m diam. 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Acacia ancistrophylla C.R.P. Andrews, J. WesternAustralia Nat. Hist. Soc. 1: 40 (1904) Typas.'near Dundas, Western Australia, October 1903,Herb. CecilAndrews (holo: NSW,.fde Maslin & Cowan 1994; lso: K, NSW, PERTH 00739995,00740403 and 00740411). Dense,rounded or flat-crowned.rhrrbJ 0.6-2.5m tall, spreading3-6 m diam, rarely obconic,small trees 1.3-1.6mtall. Bart grey, smoothor fissuredat baseofmain stems. Branchlcts angledor terete or scarcely ribbed, sometimesglabrous but usually more or less appressed-puberulousand soon glabrescent.New shootsbronzish-green, sparingly to denselyinvested with minute, red-brownresin- hairs. Slrpulescaducous, Phrllodes oblanceolate,nanowly oblanceolate,oblong-oblanceolate, linear or linear-oblanceolate,12-48 mm long, (1)1.5-5 mm wide, l:w = 3-20, coriaceousto rigid-coriaceous, patentto erect, straight to slightly curved, glabrous,normally green;apex staight to uncinate,acute, mucronate, sometimes rounded-obtusewith blunt mucro or apiculum; nerves numerous,closely parallel, obscure to a distinct but then concolorouswith the inter-neryespaces; stomata distinct to obscure;gland single, ator n€ar baseofhlade. Peduncles2 per node, 1.5-5.5mm long, glabrousor appressed-puberulouswith few to many red resin-hairsas well as white or grey non-resinoushairs; basalpeduncular bractrounded, concave, appressed-puberulous; heads globular, mid- to dark-golden, 6-7 mm diam (fresh), 3-4 mm diam. (dry), 11-23-flowered;bracteoles obovate, puberulous with red resin-hairs. Flowers mostly 5-merouswith some4-merous ones intermixed. Sepols l/4-ll2 as long as petals,free or rarcly connatebasally, oblong, obtuseor truncate,apex with red resin-hairs. Petals free, glabrous. Ovdr) papillose, puberulous or appressed-puberulous.Pods linear, somewhat irregularly raised over seeds,slightly constrictedor not betweenseeds, to 4 cm long and 2.5-3 mm wide, chartaceousto thinly coriaceous,straight to strongly curved or openly once-coiled,appressed- puberulouswith many red resin-hairs,sometimes also with appressedwhite hairs,or glabrous,brown, themargins obscure. Seeds longitudinally aranged in pods,oblong or widelyelliptic, 3-3.5 mm long, l-2.2 mm wide, I mm thick, dull tan or dark brown; pleurogramnanowly U-shaped;areole small, c. l/6 aslong as seed; aril terminal, conical, l/2 or moreas long as seed or a small,sub-terminal, closely appressed,thin plate of tissue,white or cream. Distribution. Widespread in southwest Western Australia, with one variety, var. lissophylla, extendins to southernSouth Australia and northwestVictoria. Affinities. Within its group, A. ancistrophyLlais nearestA. whibleyanawhich has longer, glabrous pedunclesand broaderpods with obliquely orientedseeds. Acacia whibleyana andA. ancistrophylla var, Lissophyllaaretheonly membersof the"A. ancistrophyllaGroup" occurringoutside W.A. Acacra "4. ancistrophylla is also rclated to A. lineolqta Benth. (see eneryia Group" below) which has phyllodes with one to three, mostly more than one, gland along the adaxial margin. The differencesamong the threevarieties comprising the speciesare subtle and dilfrcult to express: var. ancistrophyllahasstraight or slightly curved,puberulous pods and its phyllodeshave the tip often curved to uncinate;it is most similar to var.perarcuata v'/hichhas rounded,obtuse phyllodes with a short mucro or apiculum and t coiled pods. The typical variety is also quite similar to A, amyctica which has sharply to coarselypungent phyllodes. Variety lr sophylla drffersfrom the other varieties by its phyllode proportions,glabrous pods and seedshape. Discussion. In the protologue,Andrews appears confused regarding whether the flowers are4-merous or S-merousbut examination of representativespecimens explains the apparentconfusion, for the flowers vary in number of parts. Although they are mostly 5-merous,one specimenhas been noted with a preponderanceof 4-merousflowers (see under yariety anclstrophrUa). I .]trnu .Vf l'GL6i LEZ:Z €rrotrr^ Id qpuuH'sllll,^dH I ul unoJ g.V:.qlueg uou DJqoau '6nlnZEL6 '0gg .uESSorprV,"lnsurued ev) u s raddal O 9T l requratdeg ,plnsurue .qroH ,088I ?qroL(E)'.(ZEZDZ896eV) u s )trolg ,{T qreH de[roL:(Igl9Z, L6 OV) arr., 'uessoJpJv'Blnsurued '6Lgl ,Blnsuruad a{ro^ (Z) :(/000€g16 (fV) u s nddal O.g.t loqruo^oN 'lurod 'eruo.roolrllN .(qy elro^ aurd lpeu (D iadtJopaloto; xe lueur8B4-glgz9600 HJUgd :os? .Erl€Itsnv ,slnsurued :3e,1uo-dlls qlr/r\Uel uo lalqcuu.rq3ur:e ^olJ'ELIZZVL6 eV) u s awJ d' qtnos e{rolJolsBorA'turodeurdrEeu'enro^\oolnt{:(petcelesereg)ad&oJra1.GZ6D6gE:rt"rlErtsnv '{cEIg qlnos cos fog cord T' sue{ trl I Di1Ktldossqtut falput'Ittll(qdonps Dlrocy wKuorsog 'Q66D 'BrlsJlsnv 907, S cecv 'uotu,{5 ry ,(e1qtq16ur urTsehtrd, us^\oJ S U ({culg t{.i) Blldqdosslt re^ elt.tqdortslruu BlrBrV .qZ 'luerql .r3punloN snto$ uotlDMasuoJ 'Iunsn u?ql ra8uol e.rc luql salcunpedsno:qsl8 suqpetrr uorlJalloJqco) eql .prrd,{t st ucql sie^\og snoreu-yro uor odo:d 'pe^Jnc :aq8rq u pua ,{ll€ptoru8rsue^o elllos ,sepol1{qdpe,unc {18uo.qssurJ o^oqr pellc uorlcalloc uoaJo'ld faqJ ollKqdoqsnuD tu^ ur s? erncsqoeJu se ^reu eWlnq,oluJtut .Vlo esoql ol lpl[urs 'sdu .uoltotloA epoll^qd lue8und ssel ro erour seq (Surru8urdwotl gtrg futa{ W) uorlcelloc eu} ',{:unur1 pue Joqureceour petcelloc uoeq e^?q speoseJnl"u qlr,r spod :Jequeld?S-lsn8nvur sJe,rolC spouad 3u14n$ puo SuUa^^olC ',{BIc,(pu?s Jo 'ruBol .{slc ur se8prrpue seprsllrq'sluu uo (o\o1tldouow1os.A btaldlp g ,suqryDs.E ,ttuDlqpaUs 'outauotqrfua A sntdKl"Jrg) seqrununloc eallErupuE puulpoo,,n ur uoururoc,{ra,,rotuowwoJ tollqDH 'PrlE.usnv uJelsel\ lse^rqlnos 'surn9 .r) uorules .reaupua (el1roo31u)Jo lsee tul 002 uorssrhl eal??punJ ol lsue (eceJo e)€.I .uo1nqlrJsl(I Io lsue rrr{ 0S ,) ap8ep,reN puu (nurlle,rluq {o rluou ru>lZZ r) urqni& tuo4 perelrcs .(HJd!ld) .s,8fl ;vtt L aun^as I99 lu ,CV) ,ssotJ ., teeqsd?u [000 092 :l ] aeleepunl uo :(HJUAd gr09 ,trq4{aNN uraqlnosJo gS uDl Z6 'unc ,IcoU Jo S uDl9t:(HLdAd) tlVg ultsDl4U €, Euolcr^ ueen|;o g peor,{quq 9.29:(HIdgd) 'urrelloht .W,3ur,roc,roJ gllV ullsow Ug sprp,rol E[tn>IJo g uq 0t :(HJUAd) VEOI l.trox '14 'elu8epara51 .?.s :(HJUAd lgV, uaar) 7 Jo 1Y\uDl L.Z,I :GEVOOLOOHJUAd) nupn) V o '9761 'erpn:31oo3 .tequeldeg rceu :VI.IVUJSnV NUAJSA,41 pauwoxa suawlrads papaps 'alrq,r'pees ,lsurtuJel,lpcruor ,uul .epr,r ,3uo1 se 3uo1su oJcurrc Z lus eqt llnp rurug.I-y 'Suolqo 'pa,rrnc ulur S'€-€ spaas snolnJeqnd ,{v'ro11eqso1 lqSrBJls ,epr,r ruru g.Z spodr .eeq ,sleteds€ 'perc/V'og,gz-gt .eJncsqo 3uo1sz77yc slodag sp€eq:8uoJurw V-Zsapufipad Blpruols:eJncsqo se^leu 'eleurcun ,elncE :stBuortnur o] pe^Jnc xedE:tq8re4s ,Surpuerse ,(gl)g-g =,r:l ,eprmu[u g.y-9.g .Buo1 'alu1oecue1qo-8uo1qo rutu (0t)92-ZI ot eluloaruBlqo.sapol/fqJ srrErl-ursalu,rorq-per r.Dr^\paJe^of, .Aafi ,{yasuepslooqs,tre11 lnoq3norqt qloours qtog ruprpul Eol Burpuelds,llelur S.Z-9.0.sqnr{S e11,{qdorlsnuuIs^ s,rerpuv 4 g 3 u11,{qdo:lsgrueErrerv .BZ 9ZZ g1Iur1l.r.rsry1nr:,"ryurlsxr1 U Ipuc ucmoJS U 226 NuytsiaVol.10, No. 2 ( 1995) Shrubs 1-3 m tall, spreadingto 5 m diam. Branchlets angled,glabrous or sparselyappressed- puberulousand soonglabrescent. New growth only sparinglyinvested with red resin-hairs.Phyllodes linearor linear-oblanceolate,(10)15-48 mm long,(1)1.5 3.5(4) mm wide,l:w = (7)10-20,patent to erect, straight to slightly curved; apex curyed, acute; nerves and stomata more or less distinct. Peduncles2-5.5 mm long, appressed-puberulouswith mixture of red resin-hairsand white non-resin hairs; heads 11-18-flowered. Sepals ll4-113 as long as petals,free or rarely connatebasally. Pod.t linear, raisedover and slightly constrictedbetween seeds, to 3 mm wide, straightto shallowly curved, glabrous. Seedswidely elliptic, 3 mm long, 2.2 mm wide, dark-brown,the aril sub-terminal,cream- coloured, consistingof a thin plate of tissueclosely appressedto seed. Selectedspecimens examined. WESTERN AUSTRALIA: 28 km SW of Balladonia Hotel, Dundas Nature Reserve,K. Newbey 11745 (PERTH). SOUTH AUSTRALIA: 15 miles [24 km] fiom Poochera towards Minnipa on Eyre Highway, E.M. CanningW A/68 2267 (PERTH); c. 20 km E of Minnipa, N.N. Donner 25O1(PERTH); 40.7 km W of Kyancuttatowards Ceduna on Eyre Highway,M llall H80/62 (BRI, PERTH); l8 miles [28.8 km] SE ofCedunaon FlindersHighway,23 August 1974, H. Hendersons.n. (PERTH 00694711); c. 24 km due E of Kadina, I).R. Maslin 4525 (CANB, MEL, NSW, PERTH); c. 24 km E of Kimba on Eyre Highway,A.E. Orchartl2345 (PERTH); c. l0 km NE of Port Neill on Lincoln Highway, D.J.E. Whibley 1977(K, PERTH). VICTORIA: Goschenarea, l4 September1989,7. ktngdon s.n.(MEL ll83l8); Rosebery,September 1913, D.C. Trainors.n. (MEL 1500550). Distribution. Western Australia, South Australia and Victoria: known by three collections fiom Western Australia, iiom near Balladonia, Kalannie and liom betweenLake Grace and Newdegate; in South Australia scatteredtiom Cedunaon westernEyre Peninsulaeast to Pine Point on the Yorke Peninsula;and in Victoria by two collectionsfiom the north-westernregion, We have not seen specimensfiom the Murray region of South Australia, although Whibley (1980) recorded A. scLerophyLktvar. lissophylLa,the basionym, f'rom this area. A.B. Court (1913:237) recorded A. lineolata Benth. as occurring in Victoria but this apparentlyrefe[ed to the presenttaxon. HabitaL Crows mostly on flats in sandy loam over limestoneusually in mallee communities. Flowering and fruiting periods. Flowers liom August to Ootober;pods with matureseeds have been collected in Septemberand November. None of the WesternAustralian collectionsare fruiting and only very f'ew of those fiom South Australia; additional fruiting material would be valuable. Authorship oJ the name. During the courseof David Symon's revision of Whibley's (1980) Acacia handbook, we supplied him with our noteson severaltaxa, as he acknowledges. At the time, our manuscript of the present paper which incorporatedthe above combination was in press and the relevant pageswere sentto Symon. Unfortunately,publication ofour paper was delayedbutSymon attdbuted the combination to us, acknowledgedthe source,and indicated the combination was "in "in" press". Consequently, is th€ correct connectingword, as seemsclear lrom Art.46.2, Note I of the Code (Greuter et al. 1994\. Typirtcation. The authorresponsible for the basionymfailed to cite any collectionsin the protologue, only remarking that it was from the Yorke Peninsula. There are severalsheets at AD of this vafiety but the one chosenas lbctotypebest fits the protologuemorphologically and geographicallyand also "sclerophylLa is annotatedby Black on a small slip-on label as Lndley var. lissophylla". On another AD sheet(97830007) there are two other branchlets,one in flower, the other in fruit from the type \ 'sJeIllno pu€ ecnlg ot qlnos ulpnqul{nhtr uog eJuqJrq,r Jo o^\l eerql qll^\ ulqtJJuJ ol lsueqlnos IcoU 'urpeJJet{ 'uo4nq1J$1CI ',{leurEu uro4 uI{ 0/-09 lnoqE Jo snlpEJt utqlr"n '{psotu SuIunJJO '(HJuad '-If 'uaequereN gS ury :(HIUgd ^D 'l{lSN) gELt alDpulJ C /,{ ol p€or ul€rx uo {cou ecnrg Jo 6 9 :(Hluad) uns0w ag LLLLtaq aN x'ueuresroN.toasuDI9lI ,'^\erpuvlunol JoNruI L zLtg 'Idg 'cv) ecnrgJo 'ttrH seurH enp uq :(HJuad) tr€z puE(HJugd € l8I u!/s'y'\I d''g'Icou Jo as 0l 'ulpnqur{nhl 'O.IAnI) gN "np selllu r :(8Zt00L gd) u's raupflD VJ :(HJUAd t,"rl t tll 6 HId 'IASN) '3uru,4er1 g uq :(HIUgd f iraq'H B .SZZ Aur*.nC 7 y splB^\ol tuaqcl€)ll€^l!\Jo € ZI 'urq€IJBc 'uuuresloN qrnos tu{ :(HJdad) pii-1ggg tuu,ro3 d' :(Hldad) SoLl lrlruoJ a d Jo 0t 'fe,r,rq341 q ed sp$/\dol)lco1 auIIooI tuorJuDl 6€ yg77 ggly 1y13uruuo37,1 g urelscglteJC Suole t t 'uv1oo31t16 suawtradspaPaPS itnjogal iots prnag 5 1 :vl'IvuJsnv Nua"1s11yPaulwDxa 'pees 'lue su 3uo1sE eJoul Jo Z/l ellq,a\'Fuluuel 'lucruocqlr,r (arntDu-qns)sPaas seuoellq,$ ro/puEslluq-ulsel pa.l^uetu qll/t\ snolnJoqnd-pesseldde 'p"lto"-"cuo 'spees pel3lJlsuoc (ernlutu-qns)tpod rnq^"-oi ,{luedool elunare,(13uor1s ues,tleq lou :ea:1'r1n1ed'rt peJqll"\ snolnleqnd-pesseldde 3uo1, B tll c spdas peJe^\og-sl-l t spveq:sJreq-ulsel '8uol eJnJsqoElelllols pu? sa^Jau tBq,4{euos ruru E L sapunpad luapt,ta fllua,n ssel Jo aroul ol '3pl^\ '8uol ^Je^ :Surpuacsu'opl^\ uEqlJe8uol seuJll L-E tutu S-92 wu tZ-Zl'oJcnu uoqs lunlq qlr,4 'esnlqo 'pePunol 'elEloatuElqo,{[,t\oreu ol elelo?cuelqot2Poii'(qd srteq-utsa:u'\\o]q-Pel qllt\ 'ssoJcu '114 -9 pela^oc flesuep ql/(oJ3 .^eN ru g ol Surpuerds ut 9 1 0 sq'rl?y'scluocqo Jo pepunou 'SNVJ '(IN ') :ost :0i600/00 HIU3d :otott) 89LI unsoW 8-B 'J 'erlErlsnv 'lltH selttu ' :r'd'tl 16l lsn8nv g urelselN saurHspr",\\ol laqro) Jo 1(NN ltul 9 9] 9 € 'otluoc',lpuluJJel'oql€ ' eJJlplluErodns slulues aulpnltSuol z/l 'slleJJslp ollu€ 'srlueqnd-ossarddue,tsnutu snld'snqllurrds-1 euod? pBslle^Jnc JPI€^ sfiqt'ru'l'8al ;rnqtrog-g1-I 'stlsalluuul pu enbsnqlle*tuols ",lrar" I sllnllduc euJgul e^snulu snld sunJsqo sr^Jeu'snqluopuecsP'r-f,uJnJoq euoll€l'sllBI tutu 9-97'st3uo1 tuul tu-zl snEInJtdtIe^ sllEuoJcnu 'slsuvdxe JeL s,srJqo srlupunloJsltpoll(,q(l ru 9 pE le sql€ ru 9 I-9 0 aJ!JW.{Dll{lldoJrslxuD V '^ou 'ulls?trAtr 'JZ Jll'\ T' ue'^oJ S U elunl:e'rad lu^ Bll'{rldoJlslJuE BItBtV '(8861) ';,1g Sut8uv: ^lePtiA s/r'ts uouo^rasuo) q8rel ry s33rrg 3o ErJelrJceql Sulsn :u,{\ou)l^llood 1nq spprq-JJ^\o[lallssJs lsourlP rcq patrr uorloallor{e1qtq16 '(ueJE 'H'w) eqa ,(lerru,reql ur sapo11,(qdlsepl{\ eql saq e8utu Jel,^^oDeql tuo.U99LI suouwls 'e^oq€ uollolJoA uorlralloc usrpJlsnv qlnos JeqlouEsB lle,^ds€ paltc uollrelloJ uosJepueHegl 'snlBls Jreql eAIoseJol pepeeusl "xel eseqllo uolluSllse^uIlequnJ pue e,\ISnlcuoJuIale euo lEctd'(1aq1pue 'fietru't 'elElnlsnd 'snou ,rql u""^t"q auou"ullslp eqJ sdll lelqcu€Jq ,{llEnsn lseJSl r puE(arcl red sa'ueu g at) 'DllKtldotlslJuo einlenreu epoly,{qdsll uI tlullslp f:o^ T v ol paleler sl satcadst€ql q8noqlp 'y :peoEtdsru,{lurcueJ se^\I e:Fq\ o Kqdolalr.r uJo{ palleJsuutls1o11(qdossrl ^leuu L uolssnJslQ 'en8o1o1o:d 'ueeq eql ut spod uollueu lou ptp lc€lg ecurs a,teqlou ,{eureuo SuDInUeql lnq IBIJelEtll :redd{'o ,(q pelcelloc "&Ilscol leur8rroeql;o ued pereprsuocsr lelqcuelq Suue,tog eql ls€31l€ C I gpu! LZZ SI ^ur o.sll,.,trrrDy'ullsrl l U ur^{oC S U 228 NuytsiaVol.10, No. 2(1995) 35-40km southof Norsemanand one near Mt Andrewwhich is c. 116km southeastof Norseman- !n,pi southwestWestem Austxalia. :l* Habitat. lnlow Eucolyptus(E. salmonophloia,E. salubris,E. diptera, E. erernophila)woodland on oth red sand,clayey loam and loam. con (gE Floweringand fruiting periods. Flowersfrom August to September;pods with nearlymature seeds -CAI havebeen collected in December.Mature pods and seeds are needed, especially from theNorseman area,to confirmthe distributiongiven above, since some of the specimensare assigned here largely Dis on the basisof phyllodecharacters. Aut Conservationstatus. A Priority3 taxonin theDepartment of Conservationand Land Management's Hg DeclaredRare and Priority Flora List. Fla Etymology.The varietalepithet refers to thestrongly arcuate pods, from per, Latin plefix denoting De, an extremecondition and arcuatus, Latin for curved, Afr 3. Acacia whibleyanaR.S. Cowan & Maslin,sp. nov. wh Fh Frrtsr densus1-2.5 m altus,2.5-4 m diametroexpansus, ramulis initio anguladbusdemum tltl teretibusglabris, foliorum cicatricibus elevatis ornatisi Stipulnenon v\sae. Phyllodiaelliptica ad It. oblanceolata,plus minusveinaequilateralia, ad apicemarcuata ad rostriformia,aliquando recta, sp( apiculata,pulvino 0.5 mm longo,laminis 9-30 mm longis, 2.5-8 mm latis, rigidis, crassis, vulgo rectis lYIt sedinterdum leviter curvatis, glabris, nervis numerosis, arcte parallelis tenuibus, immersis, glande he inconspicua,laminarum prope basem. Pedunculi 2 in quoqueaxilla,6-15 mm longi,glabri; capitula hp globularia,atrea,2.5-5 mm diametro,18-19-floribus, bracteolis plus minusvespathulatis, ciliatis. Flores 5-mer| Sepala113-ll2 petalis breviora, discreta, oblongo-oblanceolata. Petala elliptica, $t discreta,glabra, pate ia. Leguminaanguste oblonga, ad 4.5 cm longa,5-7nm lata,coriacea, plus IF' minusveundulata et arcuataad circinnata, glabra, apiculata. Semina obliqna,2.5-3 mm longa,2-2.2 p mm lata, 1.2mm crassitie,sub-nitida, atrate brunneo-nigra; areola elevata, arillo magno,terminali, funiculi 2-Dlicatumformata. Fl c0 Typus: Eyre Peninsula[precise locality withheld for conseryationreasons], South Australia, a! 3 December1965, C.R. ALcock831 (holo: PERTH 00721786; iso: AD, CANB, K). hr ks Denseshrubs l-2.5 m tall, spreading2.5-4 m diam. Branchletsangular at first but soonterete, glabrous,with raised,prominent old phyllode-scars.Stipules not seen. Phyllodeselliptic to J\ oblanceolate,more or lessinequilateral, 9-30 mm long, 2.5-8 mm wide, 2-4.5 times longer than wide, l rigid, thick,ascending, mostly straight, occasionally slightly curued, glabrous, dull mid-green; apex qi curvedto rostriformor occasionallystraight, apiculate; pulvinus mm 0.5 long: nervesnumerous, t(l closelyparallel, fine, immersed,anastomoses inftequent; gland small, inconspicuous, near base of blade.Peduncles 2 pernode,6- 15 nm long,glabrous, slender;basal peduncular bracts persistent until about anthesis,t semicircular,ciliolate; heads globular, bright golden,2.5-5 mm diam., 18-19-flowered;bracteoles spathulate with theblade ciliate. Flowers 5-merous. Sepals 113-ll2 petal length, free, oblong-oblanceolate.Petals elliptic, free, glabrous,spreading. Ovary minttely puberulous.Pods narrowly oblong, slightly raised over but not constuictedbetween seeds, to 4.5 cm long, 5-7 mm wide, coriaceous,slightly undulate,shallowly to markedlycurved, sometimes circinnate,smooth, glabrous, apiculate, the margins somewhat thickened. Seeds obliquely arranged 'Aou dsqns uIIS"I I ? uE,toJ S U o1rt41dra'dsqnsDl^Jaua V'olgaua dsqr\s^lerylg ? uapl?t{ 'sercedsqns 'selceds ohJaua V'Aleureu'dnoJCeql uI papnlJur o,a\lSulsudujoc qcBa esJqleJl? eJeqJ 'reeurl puB luu 'selcunpad ere spod eq1 ear; ,41meuro ee:gslrud qluet:edeq1 q1t,tr snotelll-g Jo -? e.resle,,t\ou 3ql pue ruJoJrlg uo euJoq 'llBurs reqp.r eJEspsoq eqJ pesrcr ,{lsnonctdsuoJol elncsqo uorJ ^JuA se^Jeu 1e1pred,(Ieso1csnoJeunu oql:dn Sul Jnc elBcllepe uI lno u,{\PJppue eleulunJe-uoqs ol alnlc xede eql e,ruq,4luouluoc qcrq,r sepoll,{qdteg ol oloJalfq pezlJelcuruqcaru ,dnorg,, slql Jo ExPleqJ ,dnot11 mttaua p!?ttJv, aqtr '^8,$ srql uI suounqllluoc srq SurzruSocu puz arueu stq SuDcntedJadut ernseeldtsar8 alu1 eid sBuLt,iEJpaul[ se IIe,t se sqdurSogoqdlrquqqt1,Lr petu.4snlg puu ,{llculccnslnq ,{leslcerdpaqrrcsap selceds qcue qlll looqpuBq '2661 st en]l ? Jo elduexe Surpu?lslnouB sI ut uolllpe pest,tu uour,45puu felqlqlN eql sE [a,\\ 'lcelqnseql uo uorlEcrlqndgg61 stH urle]lsnvqlnosur salllrlA eql]o eSpelmoullno ol uoonqtnuoc rofrru u epuut sgq oqa ,4elqlqld A I pl^E(I Jnouoq ol ue,rt8 st eureu clJlcads aqJ KSopwtlg '( tutuoJ sled'uesuaJosg puE 'IN) srl?rsnf u3J3 pazrlucol ^Je^ u ut sa8le^puoJ uo SuL,\\oJAsluBId 09 ueq]ssal uorJ u,tou{ ^llueJJnosI '(pereSuBpug) serrads,aAeueqJ (g86 t ) q8re'l T s33rrg lo er:eltroeql Sutsn AZ sntDlsuotttl^.tzsuo) 'fqpurpnlr8uol pe8utue speesaql qll.^Ale,,\\oJJgu ein lcql spod pur sprJq pare,rou-9g-81 'BlvruolspeslE.I 'sacuds e^leu-JeluI 3ql ueql lelBd aJPleql se^JeupesIBJ lcullslp e^uq puDlue8und ,{lesJEocel? sepoll,{qdsll lnq pelBleJoslE sl (a^oqB?es) uEllBJlsnV urelsa/(\ tlloJJ uIIsBhl '9'6 'aduqs T' uu,roc \ct\cKwo olcocy lualeJJlpe Jo sapoll.{qdre,\\ojrsu ,{11uraua8s1r .{q selceds ,reu eql lrjo4 peqslnEugstpst fleug^ slql Pelou ^peallu sJelcuJEqaaql ol uolllppE ul'DuDfalqlq$ V ',olltLldollspuD su uorSer praueS aurs eqt ulqlr.r sncco D11(qdosstltr-^ v'IO selleue^ aeJql 'slecs eqlJO ,{lpulPnlleuol po8uslr" speeseql qll.^\spod ]a,,!\oJ.luupu" selcunpedlelroqs epo11,{qd qc]q,t pesrBJIte[ l€ql slelqcuoJqsnoln:eqnd-pesse;dds ^ll€nsn Sut,requl ,{lsnot,rqolsoul sla.}Ilp (a^oqs aes)s.teJpuv d d )ttlltqdottslJu, yolpeluleJ,{lesolclsotns\DuotalqtlltltllJwv'sallluL[V ',{renuel pue JequJeceq 'spouad ur petcalloc ueeqelcq spaeseJnttur qlr.^\ sPod :tsn8nv uI 8uI nlt'olg Suutntlputt Suuauolg 'sdluE.{\s llEs Jeeusetuqeulos'ureol puE euolserulluo sll'oJ9 |D|!qDH 'PrluJlsnv ^ qlnos ur BlnsuruodaJfg eql uo ,(eg ,(qun1;o qlnos BeJululseoc-.rEeu E 01pelculsau uo4nqulsto u 'Czg\ot s L6 puB28980126'02980T16 ov) 'r1'.r 'W '(HJdAd'CgJ) pu€(CV) 86.raxro,4ig t'USngnOIO HIUgd pue€€69088DgJ) u s uqel 'B'V '(OV '7971 uno) p\e OOE:HI^tolpeH B IP) LI6V pttu gl6t ta|do3 B" [suosreruollE^Iasuot roJ pleqqtlt\ satllp.ol eslcerdl Elnsuruadar^A :VI'MISnV HJnOS paulwDxasualuDads taqt) 'eycrun; 'eBJEl fqseg 1o sdooyluurural o,tr1,(q peuJoJ IuE :pesIEJeloaJE :pedErls-n ureJSoJneld ',{sso1t-qns 'e||,/,t\vrut 'Euol 'cudqle 'spod :{cBlq-u,{orq lrep 1crq1urur Z I Z Z-Z unu g-9 7 ,{leptan ut 6ZZ SI {uc r)sr^ n/.,r'.)yurlsr!.! U I pucuetoJ S U 230 NuytsiaVol. 10, No. 2 (1995) A. inceene Domin subsp.incean.t, A. inceana subsp.conformis R.S. Cowan & Maslin subsp.nov., A. lineolata Benth. subsp. lineolata, A. lineoLatas\bsp. muLtilineata(W. Fitzg.) R.S. Cowan & "Acaciq Mqslin, comb. et stat. nov. (this last taxon was fbrmerly refened by us to the multilineata Group", cf Cowan & Maslin 1990) "A. This inibrmal Croup is closely allied to the ancistrophylla Group" whosemembers differ in having new growth, branchletsand other parts, puberulouswith white and/or red-brown resin-hairs and much shorter, often curved to coiled pods. Key to Taxa of ".4. enervia Group" 1. Flowers4-merous; pods 3-4 mm wide 2. Phyllodes perfectly terete; pods tan; plants from Kalgoorlie areaSE to BinneringieStation (E of 120' 30' long.)...... 2a . A, inceenastbsp. inceana 2. Phyllodessub-terete to tlat; podspale dull brown;plants from Morawa to Kalannie SE to Hines Hill and Boorabbin (W of l2O' 30'long.)...... 2b.A. inceana subsp. conformis 1. Flowers5-merous; pods 2-3 mm wide 3. Phyllodeglands (inconspicuous) 0 or l, whenpresent situatedfiom distalend of pulvinusto 0.5 (l) mm above; seedaril white; nervesof phyllodes weak to obscure 4. Phyflodes terete to sub-terete,often with pustulatestomata .....la. A. eneruia siubsp.enervia 4. Phyllodes tlat, linear to narrowly oblanceolate,stomata not pustu1ate...... lb. A. enerviasubsp, ex.plicata 3. Phyllodeglands (inconspicuous) l-3 alongadaxial margin, the lowermost(3)5-21 mm abovepulvinus; seed aril yellow; phyllodenerves distinct, slightly raised 5. Phyllodeslinear to Iinear-oblanceolat€or oblong-elliptic, t 20 times longer than broad, curved-acute,not pungent; sepalsfree or to l/2-united; plants often of saline situations on sandyor stonyloam ...... 3a.A. lineolotasrbsp. lineolata 5. Phyllodesoblong-oblanceolate to oblong-elliptic, 10-13 times longer than broad, acute to short-aouminate, coarsely to sharply pungent,rigid-coriaceous; sepals alwaysfree; plants of sandplains or on rocky clay...... 3b. A. Iineolatastrbsp. multilineata l. Acacia enervia Maiden & Blakely, J. Roy. Soc.Western Australia l3: 8, pl. 8, figs 12-19 (1928) "September" Typas: Hines Hill, WesternAustralia, 2 October(.rphalm. in protologue) 1923, M. Koch 2806 (holo: NSW 195719; lso: K, MEL, PERTH 00969737, 00969729and 00750484-frasmentex NSW). Dense, rounded shrubs, often becoming obconic, rarely small trees, 0.6-3.5 m tall, spreading 1-3.5 m diam.. Bark grey, fibrous, fissured. Branchlets slightly angular, glabrous or somewhat appressed-puberulousapically and glabrescent.Phyllodes fTatand linear to nanowly oblanceolateor '(I9€g8900 'utddounun;q HJdAd) usroplA ?g'lI6I ^rlnrqeC8 :(HJdSd) Zlt suowruls HW 'aere 'uEuesroN gSA 8ur{rEd sellsqJ l€ed eplstno lsnf :(HIUgd) 9691 taqnall , Jo ur{ 'lloi[ 'IN) 'puor 'IJ?d 0, r {ee]is r€elJ Jo N uq 8 :(HIUSd 9099 fa4,{aN , 8uI) e{8'I-u€ures.IoN '{uuJ 'X'ueluJesroNJo ur{ I"uoDuNuuBH Iu?rC elII I06lo l&S uDtt€ : (HJUAd) SEIg [aq$aN lAl\l S/ ', rllrH "Iabl') 'SNVJ) o^r?cJogs ur{ 9:(HJdad 90i9 ?1isr7yy g'etprcSlooJ ol PEoruo lrou 'AN '-IAW ') 'SNVJ) euolcr^ ueano Jo N tu{ 9 t :(HJUad 196€uusDw & {J'qcur.+llngspJE.{\ol "IglN) ssorJulaqlnos Jo N url S :(Hldgd ZtLl ultsDWUg'llIHseurHlogSenp[u)8lsalrut E r 'qcuIJIIng :(HJdAd) 8902 8lliuulJ U sPrl?,{\olUIIBI€rrviNIo g [(u) S t] sellruZ Z :(HIUAd) VgtZ€ 'SuIp"rEnO 'SNVJ) t s Itutrolg g A puEurStlroJ uee^\leq 1(t€h98900 pue 98n€8900HIdAd') Katlog'Jg'tt6lreqrrleldes5lcodetnrg:VI-IVUISnVNdAJSA/&'paurw2rasuatulradspatrzlas '3uo1 '3uo1qospaaS pe^Jnc,{l^\olpqs ulc g g ol sPod ultlp luttr i-€ spueH e}ulnlsnd '8uol 'aleJel-qns -oJrru ueUoulelrjols :ornosqose,ueu :opl,tl tuul Z-8 0 tuc a-Z ol e1€Jelsaponfqd ulr.raua dsqns,41e1u19A uepIEI tr ul^Jauo SIJBJY'BI 'Elutuols eql.+oosn€ceq elulnlsnd-orctu rueddeuago qclq,\\ sepo11,{qde1ere1 putr spueq Jellutus fpq8rls s1t,(q 'rxot pezrusoceJsr euo 1vrrd,{1eql qc1q,,vt1o setcedsqnso,tl sest:dtuocnta"ra&a ncncy c\ficadsntlul '(DiDpawt '69€9 '89fg dsqns ullsDw A B puuDtoJlldtz dsqns unsow'U'B f :s1ltgue8uol6 'oldurl?xe eql Jo lso^\ uq Z lnoqu luu aullps E uo lol) xllecutEdul^s.rncco ,{aql uaq,lt ,{1:ulncqrud 'pesnluoaaq ,(lls"e uuc pul?lEllutls ,411utcadseere olDloaull dsqnsDlopaull V puq'oJortldxa dsqtrs '(otDpault ohJauaolzDcv yur,ro1le,{PuE serceds slqlul olrq^\)llJu eql Jo Jnoloceql ul su lle,t su 'ernl?^Jeu eql Jo ecueuturordeql pul-(o$4oau y ur snut,tlndeq1 a^oqu tuul IZ-g(E) uIEJeuIIEIxEpu 'u^raua eq13uo1e erour ro euo y uI snur^lndeql e^oqe utu I o1dn ro pue ls]slp eql le euo) spualS epo11,{qdalo go uolllsod puu requnu eql ut Suuagtp lnq qluag Dwloaull V Jf.eu["n6 sautul[[y 'Jequeldes se ,{1e1u1gpue uepl€I ,(q peqr.Icsue:1Xlualulsttu ,{pue.ruddess,,lt ll lnq ..€Z6t x 2,, s" uoDcalloJJo elEp eql selEls,{1:ue1c puuq s,qoo;4ur ed,(to1oqeql uo leqPleqJ uouw{ldtJ '(ueurasJoN Jo lseeqlnos-lsueuq 0, , sr tlclq,t) IIe^& IueJlS JBoIJlltJu ol lsce (Jdloqlsua^cu ro 'eslllc lsBeqtJou-ls€eur{ 08 J) ulua8l?tr1 e{e'l pue e{E I JBeuol lseaqlnos(nulllE^\lE(I Io lsBeqlou uq 0g , sl qorq^\)SulpJeqqrl JBeu ruo4 €rlEJlsnVuretsad\ lso^\qlnos ul SulSuer-epl16 uo!trnEJis1Q 'ellq,l\'luuruJel IIJEeql')3tlq'epl,/Y\ ultu s l '3uo1 ruur 7-g'3uo1qo o1 cr1d111efyrrorruu'spod uI paSul?fie,{1luurpn1r8uol s/22rs snoJqBIS'pe^Jnc 'snoeceuoo 'epr,,r, '3uo1 'sPees ,{1mol1eqsro lq8rz4s ,(lurqt ww S Z-Z utc a 3 ol uee,'$leqpalclJlsuot 'J€eurl 'eeJJ 'qfuel l"ql\euos (eJnleu-qns) spod Suturoceqtnq Jla.raqot A11tudseuqeuos lo ee{ 'alEInqleds ppdEIZ-Zll spdas t,{luosoru0auos sleladro sledas'snoreu-gIllsoul tlr,lrold elqo tlrJ 'ur€lp ',to11e,{ 'JEInqolS + ol etB^oqo soloels€Jq: peJe,,!\ot+-EE-8] ulur 9-6 ueploS deep ol tunrPeu spEeq:snoJqEIS ro snoln:eqnd-pessa]ddB'snolnjeqnd,{lesJEds'TUJoJIIIJ'3uo1uru 6-7';txc rad g-1 'snonlldsuocul'lFuls sapunpad luesqero snur,rynda^oqD urul ( I)g 0 ol puo plslp lE puEIS:lJuIlsIpuI '1a11ered 'snolaunu tnq aletqsnd pq^\euos ,(flEuolseaJo?lEluo]s :eJnasqool lcuqslpul {1aso;r '1uecserqu18'snoln:eqnd se^Jau:pe^Jncpuv lepuels etEururncE-lJorlsot elncExedE:uae:8 )I.Iep o1 elud 'epl^ 'eleJel -pesserddessel ro uoru ro sno.rqe18lq8leJts'tJoJa ol peullcul ulu.I9-8 9'3uol urc a 8-Z EI ,{tll]lle.sll,1lr?rrry'urlscw d gpulr uc^\oJ S U NuytsiaVol.10, No.2 (1995) Distribution. Found from Kununoppin (c. 60 km northwestof Meredin) eastto nearCoolgardie and south to Frank Hann National Park (located30- 110 km east-northeastof Lake King) and nearClear Streak Well (which is c. 70 km east-southeastof Norseman)in southwestWestern Austxalia. Hcrbitat. ln sand or loam, rarely clay, in open eucalypt woodland or open mallee scrub. Flowering and fruiting periods. Flowering in Septemberand October with one collection in December;immature pods have beencollected in November,December and February. Conservqtionf,tqtrs. Not under threat. lb. Acacia en€rvia subsp.explicata R.S. Cowan & Mtslin, subsp.nov. Phyllodia plana,lineariaad angusteoblanceolata, 2.5-8.5 cm longa, l-6 mm lata; capitula4-5 mm diamelj.oi legumina ad 7 cm longa, saepe leviter curvata].semina angusteelliptica vel oblongo- elliptica. "floodway", Typus: 19 km N of Wongan Hills near Kondut in broad saline Westem Australia, 30 September1984, G. Craig 1608 (holo: PERTH 01469924;lso: CANB, PERTH 00697850). Phyllodes flat,\near to naffowly oblanceolate,2.5-8.5 cm long, l-6 mm wide; nervesmore or less distinct. Heads4-5 mm diam. Pods to7 cm long, olten slightly curved. Seedr narowly elliptic or oblong-elliptic. Selectedspecimens examinetl. WESTERN AUSTRALIA: WalyahmoningRock,Baynes Museum56 (PERTH);Bencubbin, W.E. BLackaLLS32 (PERTH); l2- l4 km E ofLake Gracetowards Newdegate, G. Craig 1512 (PERTH); 9.6 km N of Ballidu towardsPithara, G. Craig 1595b(PERTH); 9 km E of Pithara towards Kalannie, G. Craig 1599 (K, PERTH); 18.2 km W of Koorda towards Cadoux, R.J.Cumming23l l (PERTH);I km N of"Koobabbie"House, Coorow, A. Doley9B (PERTH);10 km E of Mollerin towards Beacon, B.R. Maslin 4140 (PERTH); 4 km W of Wyalkatchem,B.R. Maslin 4449 (PERTH, TLF); about2 km W of Wongan Hills on road to Piawaning,LR. Maslin 5368 (MEL, PERTH); 4 km fiom Coolgardie on road to QueenVictoria Rock, B.R. Maslin 54O4(AD, K, NSW, NY, PERTH); t km S ofYellowdine, K. Newbel5985 (PERTH); 1 km S ofGhooli microwavestation, c. 20 km E of Southern Cross, M.11. Simmons 1220 (PERTH); 2l.l km S of Pingrup turnoff on Newdegate-LakeMagenta road, M.H. Simmons1343 (PERTH). Distribution. Extendsfrom betweenJibberding and Whitewells homesteads(c. 50-70 km northeast of Dalwallinu) south to nearLake Graceand eastto Coolgardie,common in the Coorow to Wongan Hills region, southwestWestern Australia. One collection, B.R. Maslin 5EOl from 18 kmeastof the Norseman-Esperanceroad on Quast Road about 27 km NNE of Salmon Gums (PERTH), is far southeastof the main range of the subspeciesand indicatesa considerablywider distribution. Habitat. Mostly in saline situationsaround salt marshes,flats and lakes;occasionally in rocky clay on hills or in sandy loam near saline areas. Flowering and fruiting periods. Flowers from August to October with the main flush apparentlyin September;pods with mature seedshave been collected in January. '(HJdSd) ,[erlroo8lu) .r] 6LLI suoaw1s.l.I.r{ Jo IINN ru) 0€ uoqcunl .g ,erFoo8pl;o lpul^ rr]og peor .{ralleg urel uo N ur{ g e :(HJdAd) llLg auD^as A tuq Z€l 'Suolng ,serzuey{ satrur0Z :(HIdAd) 6109 ufFow l' B' ot pror uo erTroo8le)Jo N uDt g I :(HJX!Id) 'uoll"ts tLZg plDag ST at8uueuutg :VI'IVdISnV NUILLSA/d paulruDxasuawroads papaps 'uees lou spzzs :u?l '3uo1 ur: 9 g-g spod:Suolqo,([,rorJEU s/pdas :pera,roU-0€-02 speoq :8uol ruru g-t .rapunpad,.snotqel? '3uo1 'lue8und ruc A g-7 ,{lasJgocol lua8undlq8r€'lls seurrlsruosJo pe^rnadrl eqt,e:r,Jetsapoll(qd euealuldsqns unuoC BuBaJurBTJeJV .BZ 'sercodsqns 'ouoaiu 'V o,til sesrrdruoctr1 v Jeprll' 'setceds ,elqplru^u o,rl eql uaa,^^leqqlpr^\ ur ra3;rpspod eql lurp s:uedde1r pellurl eql uloJg '(luesqE leuel€ur pu"l8 ro) epelq er{lJo osBqer{l le pue13e13urs r q1r,,rrsepo11,{qd pue sre,roU snoreueluadsBq qcrq,r (e^oqe ees)flelelg T ueprcI^l2,iatauay oy jl-ltws llpcr8oloqdrou sr sanads eq1 .sag\uyffy ',{aup,{S puu en8er4 le slueu8ur; qtr/' /t\a) l[ st uorlcelloc uruu eq] snqJ .Burrupulol uuol uo relel uorlJellor aql pe^receJueprcI I puo ^\e){ ol lrsr^ u Suunp lueul8e{ u peurulqoururoc :,ra) lu uorlcelloceuuseqluopesuqeta^\^la)plg?uopruhllotuqlpusatueus,urutoqqlog.uopocr[trl(1 '?IleJlsnv uJelseid lse,rqlnos'seru arlJoo8l€)Ieql ol lsEe pue (,{u.$q3rguJelsEa lcetD 'r) .uo!nqlqs!O uo urpaJJehl.lolsa,rqlnos uI 0Z IIrH seurHol lweqlnos E,rEJoltr leeu tuoJl sJncco 'luururJel 'epr,r IU?oql'u^\oJq {rup "{ssol8-qns uru g 7-7.3uo1uru g-g 1,8uo1qo,rpoa u,p"a",illil .u,rorq ,{1purpn1r8uo1Glw.totuoT dsqns) spaa5 IInpelgd Jo uel ,sno.rqul8,lq8rens ,snoacsuor 'epr^\ 'tuol ,spaos ,{lulql ruurt-€ uc A g-y ueo,rlaqpolarJlsuoc lou lnq Ja,\opesr€J ,{pq8rls ,.rveur1 spoT 'el€lnql"ds-J€eurl ,eeq ,qlSuaflBled .sno]Fw+ ro etvlnqluds+ o13uo1qo,(l^ oJJuu Z/l slDdaS sla^oll 'pere,rou-0€-01 'ru€rp ,rBlnqolS ,3uol ,1rxu ruru t-g € sp€eq :snorqulS ruul 9-g red Z sapunpad 'snur^lnd ,z ,lsllBJ€d e^oqEuru sl-Ol euo lsa,rol lo l spupJS:pesrcl seurqeuosslBLuols :tcuusrpur '?urJ 'snoJorunu ,{lesolc se^JoullueBund + ol snoncouur,u,roJq ,lq8rEJls,(1erur ro pe,,rrnc,eug xade 'snur,rlnd :ueeJ3-e^rlool -lq8r.rq ldecxeluecselqulS lsJrJ t€ snolueqnd-pesseJdduro snorqulf ,po,,rrnc ,{pq8qs ,pr8u ,l€U .uterp .t ,Buo1 ro lq8re4s lJaJe ol Surpuec$ Jeqtej ueq^{ept,r\ ruru Z ol Jo ruru g. 1-1 rrjc a 8-t l"U ol oletet sapo tqd tuarseJqEJ?,snolnJeqnd-posseldde :o snorqul8 ,e:trJ€lsJafpuotg 'lcuDsrpur 'snocnpEc ,,{€JA .l]r.t ll"rus salndllg eseqlc q8nor ro qtoorus )trDg u t_I sqnqs '(;J ,ndSN ,.t xe lueru8er.l-116r29Ud :ost :opq) acul uretselA :.rrdfJ '^ou'uoufs'(gZ6l)91-1s8rg,g.1d,g:gI€rlellsnvuretsald.ros.,{odH A4,Erlullsnv I,,{le)plgn neprcWaD u!.V '(X ,iASN .H xe lueu8B4-r/61.29 Ud tos! :>I:olotl) ul A4,ErT€rlsnvulatse^A .In€N .ucelods .l[D{ :sndq (EZ6I) €t :Z JrJdjjul J -rI eISeC utsa^ ,ultuo( BuBaJu!srcBJV .Z 'pepuudxe ro 1no peerds$I u\e-I,suttrlldxa 'uoxel ,pepuedxe ruo4 sql Jo sepoll,{qdpeue €U eql ol JeJeJol uesoqJsr erxeu eqJ .l.to1owK1g 'l€aJql 'sn|Dts repun eq ot u,rlou) loN uo!1Duasuo) SI {uBllorsrr\,r/ry'urlsDIAI U gpuv uB^r\oJ.S! NnytsiaVol.10, No. 2(1995) a Distribution. All known collections are from north and east of Kalgoorlie within a 35 km radius, excluding one from Binneringie Station, c. 100 km southeastof Kalgoorlie. Habitat. On rc(l soil with scatteredeucalypts and low semi-denseshrub association,rarely in saline habitats. Flowering and fruiting periods. Flowers in August-Septemb€r;pods with mature seedsnot seen. Affinities. This subspeciesis quite similar to subsp.conformts which differs by its sub-tereteto flat phyllodes, fewer flowers per headand pale dull brown pods, as well as by its habitatpreference and distribution. There is also a strongsuperficial resemblancetoA. kalgoorliensis R.S. Cowan &Maslin "A (seeabove under . densiflora Group") liom the samegeneral region but that specieshas branchlets with sub-appressed,white hairs and intermixedred resin-hairs,phyllodes with about20 slightly raised nerves,between which raised stomataare clearly evident, and 5-merousflowers. Discussion. It must be noted that Maiden describedthe speciesas 5-merousbut all the material seen by us has predominantly4-merous flowers,5-merous ones rarely being ibund among the 4-merous ones. Conservation status. Not known to be threatened. 2b. Acacia inceana subsp.conformis R.S. Cowan & MasLin,subsp. nov. Phyllodia sub-terctiaad plana,apice innocuo,ourvato,4-7 cm longa, l-2 mm lata, initio appresso- puberulaglabrescentia praeter pulvinum puberulumipeduncuLi 5-6 mm longi, capitulis l0-15- floribus,.repalis plus minusvespathulatis; Legumina 4-7 cm longa. Typas. betweenMorawa and Perenjori Ipreciselocality details withheld for conservationreasons], WesternAustralia, 3l August |,982,R.J. Cumming 2199 (hoLo: PERTH 00191043;lso: CANB, K). Phyllodes sub-terete to flat, 4-7 cm long, l-2 mm wide, at first appressed-puberulousbut glabrescentexcept for pulvinus;tip curved,t innocuous,peduncles 5-6 mm long, bearingl0-15- flowered heads;sepals more or less spathulate;pods 4-'7 cn long, pale dull brown. Selectedspecimens examined. WESTERN AUSTRALIA: Ipreciselocalities withheld forconservation reasonsl,N of Kalannie, G. Craig 1600 (NY, PERTH) and 1632A (NSW, PERTH); NE of Kulja, G. Craig 1,622(MEL, PERTH); betweenMukinbudin and Bencubbin,B.R. M.lslinfot R.J. Cumming 2303 (MELU, PERTH);between Hines Hill andNungarin, B.lt. Maslin2339 (PERTH)and 23394 (CANB, K, PERTH);Cowcowing, F. S/oward311 (PERTH). Distribution. Extendsfrom nearMorawa southeastto Hines Hill and eastto Boorabbin (c. 90 km west of Coolgardie on Great EasternHighway), southwestWestern Australia. Habitat. Ahighly salt-tolerantspecies on alkalinedark red-brown,coarse clay-sand overlying yellow- brown coarseclay-sand or sandy loam at margins of salt pans or lakes. Flowering and fruiting periods. Flowers in August-September;pods with mature seed have been collected from SeDtemberto December. '(uorssncsrp roJ a^oqBees) .{1e1e1gry ueptel/\LDl^taua y o} J€FurS saUML[fV 'relurrs 'uoxul ,{11ercr;radnssr 1r q8noqlp srqt lou sr lr :perJrlueprueeq lou seq (adflotcetcmd) '.V uollcelloc roqlo eqJ ed,(1o1ca1su laeqs Jo{ooH qJeH eql uo EI puouunt6' ;o ueurrceds aql Jo aJroqc aql acuaq'taeqs ur€q]u3g qJeH eql uo pesre,\alolll sJequrnuuoDcelloc eql d pu€ 'CXO 'C lp ^lel"Jadespatunour selecrldnp,{q u/roqs su'stueurele o,rt eqlJo qceerol peuBrssesr requnu 'suorl3elloc uollcalloJ laeJJoceqt sleeqseseql Jo lsrrJ oql uO :t puB ZL :V puowwue',{lelupu ',,unuurJe{ooH luareJJrp o,rt Jo slelqcuurq 3^"q qloq :(.unuElur€qtueg unrJuqreH,, Jsqto eql 'prreluu 'qreH 'uoltwL[ldKJ unrJEqJoH,,peduels euo ed,{1Sutreeq ; tE steeqso,rl eJEaJeqJ 'dn:8ur4 o1 'uounqllste qlnos (uolduEq oNJolsuatu{S€ J) "un^ uroJlurlEllsnv uJelssl&lse^\qlnos ur sJnJOO ',rrro11a{ 'pcruoc 'pees lqBuq sr 3uo1 'lEurulel 'pees se trc-Ell lue eql :u.,!\o:qreled su Suol s€ €/l -tlT lnoqB ?looJu:ped€qs-n u"rSoJneld 'fssol8 'pr8rnl 'ept,r ,3uo1ulur ,Suolqo :Iculq-u,roJq Jo u/r\oJq Jo pesseJduoc lxrx 7-71 t-€ Jo JqdlIle-Suolqo'cDdr11e 'spod ut poSuelu ,{l1vurpn1rBuo1spaas snolqslSro spaasuae,rleq snolnraqnd 'pe,rrnc -passe.tddete q,teuos ,{1no1pqsot lq8lels 'snoeJuuuqcJo snoersuoc ,{1urql 'epr,tr aiai 'speas 'mal|ll g-7 pu? Suoyrur 0l ol uear!\laqpalcrJlsuoJ ,{pq8rls putu ra^o pesrur spod snolnraqnd -el?llldud Jo snolrueqnd-pesserddu'snolnJeqnd K|D^O 'pe:r]un-Z':lol Jo eou slo|ad pe:nun-Zll 'sFlad 'uJJoJrsnJ o1 dn ro ee:; sB Suol sE EIZ- I s/odas' snoJeu-g ean ol! e]I-lo\ta ol Jceull "tuurp seloelcerq:pe.re,roU-Sg-81 lutu 9-7'ueplo8'proBuolqo,(pq8rls ro .ru1nqo13spueq :elEIIncnc 'snocnpcc '3uol 'lrxe slcuq lulncunpedI"sEq :snolnteqnd-pessetddu ssel Jo eJourol snoJqElS urur I I -Z 'snur,rlnd 'urErsur 'snonJrdsuocur red Z(l) sapunpad e,roquturu tZ-S(g) lsoure,{lol eql lerxepe uo 'llEtus 't-I 'elu(uols (s)pu€13:lsuorsBJco Jo luesqe sasouolsEue pesrBJseuDetuos !cullsrp Jo s,roJ '(elels ^q pelJ€ur,(t]?atc pus peJnolocta{tEp secrdsa^Jau-Jalur eql'peslcJ ol pesJauul ,(Jpur lseal le),{\ollef puEluepr^e fFeelc'1e1p.red flssolc 'snoJeurnusa^Jau:tue8und ol snoncouut'el€ulunau -Uoqs 'pe^rnc ot elnt" sseIJo eJou ro lq8rurls xadu :ueol3-{Jvp11np ro ueer8-e,rr1oluecseJq"lS pu? 'po^Jnc Sunof ueq,trsnolnreqnd-pesserdde :o snoJqslS ,{ll?proruBrs.rope,uncur t€q,{euos ol lq8r€Jls 'SuIpuersB 'snoer€uoo-pr8u 'apr.r '3uol 'tug ol luelpd ol snoecEuoc urr.ug-Z( I) rur (g r)9 9-g .{1pnsn 'ct1dr11a-3uo1qo ,J€ln8u€rJl'1ue1srs:ed o1e1€loecuElqo-JBeurl ot mewl sapoIKUd 3uo1ruu I uuql ssel salrd?,l5 snoeJues,{a13,{;elys ro ,(Je^lrs-uollrcqt,lrolS ,ne1q sno:qu13selEuu eq1 qlrm snoynreqnd 'xede ,qloorus,,(eJB -posseJddEJo snolqelS tu palSuele.q/Y|awos svlqJuDtB es€qlp patnssrJserulleuos 'rnetp ,esuaq ,Uog u g ol u^\oJc ^qsnq Jo esuepqlr/'\ \el w L'l-g O sqwqs cruocqoJo papunoJ 'nldtJojJal 'DJoloaull ,ueurJads nsuas y lou sr ll lnq srql ol allluu e ,{1dduo1 eyqeun a;B e/!\ :(,roleq uorssncsrpees 'adtlonapsl pue a.l,1olcalqtr,r pelunou ,tu€qluog 'qroH 'qreH '){) puE relooH Zl :l puoawnrc I'lEllE.nsnv uJelsol\] ra^ru ue^l'g .ar[(Jotca1ot24 '(ACI') 'd '!CXO .tu€qlueg ,g ..otralos! ,.t"IlooH xe 1uau3u.ry-g1ggg800HJUIId qreH-) 'qroH '8t81 'lEltErlsnv )) tI :V puowwnte f urelsed\] re^ru uE.,!\S:(pelceles erct1) ad(1o1ca7 ' .e G98I) 929 :92 eeuuur-J qlueg "lEloaull BlruJv 'JBlrrurs JoJ vt\e1'stu.uoluoJ ruo4'sJe,rou snoreu-g 3ur,\rq ur sJaJJrpqcrq,\\ ,DtD)lldra .dsqts 'V .t?o1outt1g o!^Jaua ot uoxel srrll Jo acue.reeddu.tulrturs oql ot uorsnlle ut uesoqasr taqlrde eq1 'lsl'l t'rolJ ^llrolJdpue eJEdpJJ"lJeC .sn\o$ s,lueure8euetr{ pu€.I puBuorlE^JesuoJ Jo luerugudaq eql ur uoxel 1f1uou6y uotlD^,rasuo) gtz SI ,{wllacsrtr{,?rrry'urlsDl U gpuc ur^\oC Sd a NuytsiaVol. 10, No.2 (1995) Infraspecific taxa. Comprisingtwo subspecieswith the typical one differing from s'rbsp.multilineqt.l in having proportionately much longer than wide, often sigrnoidallycurved phyllodes with the apex recurved. 3a. Acacia lineolata Benth. subsp.lineolata Phyllodes linear tolinear-oblancaolate,occasionally i terete,2-5(7.5) cm long, (l)2-4 mm wide, c. 20 times longer than wide, more or lesscurved, ofien sigmoidally; apexcurved-acute and not at all pungentor rigid. Peduncles2-6 mmlong. Sepalsfree or to half-united. Seedsglossy brown-black. Selectedspecimens examined. WESTERN AUSTRALIA: betweenPitharaandMlbng W.E.Blackall 2893 (PERTH); Pingrup, W.E BIackeLL298'7 (PERTH); l l miles [17.6 km] N of Dandaragan, W.E. BLackall 3671 (PERTH); 0.7 km S of Tammin on road to GardnerReserve, R.S. Cowan 4744 and B.R. Maslin (PERTH); l4.l km from junction of Corge road with Ajana-Kalbari road, Kalbarri National Park, Il.S. Crnan A-821 and R.A. Cowan (PERTH); east of Coblinine River crossingon Warren Road (eastof Katanning),G. Craig 1586 (PERTH); 7 km S of Miling on Lyons East Road, G. Craig l59O (PERTH); 41,6 km fiorn Brand Highway towards Coorow, R.J. Cumming 1319 (PERTH); Wilroy, -/. Galbraith34T (PERTH); 12 miles [ 19.2km] E of Katanningon road to Nyabing, B.R. Maslin'784 (MEL, PERTH); 32 km W of Kununoppintowards Wyalkatchem, B.R. Mqslin3400 (CANB, K, NSW, P, PERTH); 14 km E of Kulja towardsBeacon, B R Maslln 3975 (CANB, K, MEL, NSW, PERTH); about 2 kn W of the Wongan Hills on road to Piawaning,B.R. MasLin 5369 (NY, PERTH); 27 km due NNW of Coomalling, Mortlake River North, B.R. MasLin 6200 (BM, BRI, PERTH); 3 miles [4.8 km] N of Brookton, K. Newbey891 (PERTH). Distribution. Occurs fion Kalbarri National Park (c. 100 kn north of Ceraldton) southeastto near Katanning and Pingrup (c. 50 km north of Ongerup), southwestWestern Australia. Habitat. In sandyor stony,oflen saline,loam in associationwith Casuarina spp andHalosarcia spp' or in eucalyptwoodland. Flowering arul fruiting periods. Flowerc in August-September;pods and mature seedshave been collected in late November. Variqnt. In lhe \ofth-central part of the rangefrom Yuna southto Yorkakine (c. 25 km due southeast of Wyalkatchem) and east to Kulja (c. 45 km due north-northwestof Koorda), a form with narrow phyllodes ( l- 1.5 mm wide) occurs,some so narrow as to appearmore or lessterete, e.g. B.R. Maslin 3975 (CANB, K, MEL, NSw, PERTH) and6200 (BM, BRI, PERTH). Conservcttion,rtal&,r. Not under threat. 3b. Acacia lineolata subsp.multilincata (W. Fitzg.) R.S. Cowan & Maslin, comb. et stat. nov. A. multilineotq W. Fitzg., J. Western Australian Nat. Hist. Soc. 1: 13 (1904). Lectotype: Anino, WesternAustralia, September1903, W.V. FitzgerlU s.n.(NSW 216915,left-hand specimen on sheet, lde Maslin & Cowan 1994a,isolecto: NSW 216915 (right hand specimen),PERTH 00765813, 00765791(ex herb.C A. Gardnerno.13068),00765805(Fragment, presumably ex herb.W.E.BlackaLl). ateretsepollfqd I 'V DqJtldltt ecEJqcee uo se^Jeuluulpnll8uol oarqt 'l qll^\ (,,'\ou€u,{Ja^uaq,t elaJel-qnsSuuuedddP) l?U sepolt,{qd (s!p1r!^ottt2 dsqnss4lur1ss, y o1 ,{ttrulturtslPlal]adns sll Jo 'a^oqe asnBceq,{eI slql uI pepnlcul sI lnq peur;apsu dnorg eql Jo laqulauJ3 lou sl DqJtldul olcDcv) ,dnot} u.113o.tl y,, Jo BxBl ol ,{a){ '\tuerz 'S aulrun VpuB ^ou ds ullsEt{ T, uB,ii\oC d?rlqiuolqdo y',{le{Elg d' uepIBlNsI!8D$ V ' 'ds ^ou ullsul^l ? u€^\oJ SU,4 lfl3tlasaoly' ,rou dsutlssl T' uE,,r\oCS d oll&ldoJolno V"r''ou 'dsqns 'Si ur1su1,t1? uB.{oJ S'ld stplJt^otlo dsqnssilpalssa V'snlalsso dsqnsoJooIAI slllwrsso V :3ur,tro1yo;aql epnlcut ,dnorg,, stql Sutst:druocvx4 aq1 ,{lqueptrteql Sutpm8e:uolsn;uoc lsed eq1 ,(1ue1cog 1dure11e ue ur ereqpeqlJcsep-ar elc sxut peqlrcsopflsnol^eJd eql Jo eulos spodpesseJduoc 'r€eurl puE:spdes ee{ qlr/$sra.{\ou snoletu-9 :op?tq eql Jo prlql-ouoIDs€q eql uI puulSu qlIA sopotl,(qd elerel ol alaral-qns :uoluruoc ur sJalsaJellc,treg e,Teq SutdnoJ3 PurloJul slql ;o setcedseqtr .dno41 stltSotl mcDrv,, }qI 'lsI'I €Jolc ,{luolld pu€ erBu pal?[ceo s.luoueS?u?hl pu?'I puB uou?AJesuoc;o lueuludeq eql uo I ^luoud v su0$ uotpMasuoc 'satcads '(se1ndt1s o,rrt1eseql;o uorlducsepJoJ (066I) ullsenl d, uB,,!\oCeas esoutdssuq qclq,{\)ults€I,\I 4' u€^o)'S'\DUclngun V Pu€(elnp^Iau epo11,{qd1ue:a4tp ,{re^ s€qqclq,,\\) ults€ld 'S 'sary?u{lv ? u?,4{o3 d DtDlSotod Vol ecuelquesel ptcgredns E smeqovaullq\w selcedsqns 'reqtuaJac puu Jeque^oN uI pelcelloJ 'spouatl ueaq a^€q speeseJnleul qll/|\ spod :lsn8nv ol eunl tuo4 snholg 3u!Jlu! puo qulol$oll ',{e1c,(1co.ruo pue suwldpuusuO lBtIquH '(Bggg ullsDl! U g) ereql ser.odseql Jo ecuesardpenuquoc eq1 srrJJguosuorlcelloc luocors lnq selllpcol uo4cellos Jeqlooql Jo lsotrlIo lsJ^\ pue qlnos ,(lqElJptsuoc '?,uellnt{ sl ,{lrl€col ad,{l eqJ "rIEJlsnV uJelseid lse/\{qlnos Jo ^llurat^ eql uI sellllEcol ^\eJ 3 uro.U,(luo u^\ou{ sr sercedseql 'ouuJv ruo4 palcellocsg,t\ qcq,,l\ 'ed,{t eql ol uoqrypEuI uoltnqlJ$!Q 'GIUSd) 'ouruv 8899 ullsDw y 6f r?eu :(HJdiId ,1.0S "Ight ')') 'a '(Hruad) pr? (HJdgd Ltgtultson s ztdc sdlltlltd D'(Hrda 9t9? puP(HJdsd) '(HIUAd 'SNVJ) '(HIUAd) '(HJdAd) 9t9t 9I9t 9I9t 1291tqqsy 1,t1'y'fsuoseeruollE^]esuor IoJ 'pautaoxa pleqqlr^\ saqllEcolesrceJdl sala B,\'\allnIAI:vITVuIsnv NdaISaN\ suawlradspaDalas 'u,{lorq ',{sso[S 'ee4 spaas s,{u,,nps1nda5 3uo1unu11-g saicanpad ]ue8und,{ldrqs 01,{lesr€o.puu 'snoecBuoc-pr8u'opr,r 'apl.\\ lq8prs lurod pcrdE :pe^JnJurol tq8lEJls uuql Je8uolserup €I-0I ultu '3uol 'epuruncB-uoqs 'crldllla-Suolqo S-€ ruc g'9-e ol eln'? ol etEloacuElqo-8uo1qosapoqttl4 -U SI ,{uvll6csrj^1'rrrry'ullspt{ g pueus^\oJ S U NuytsiaVol. 10,No. 2 (1995) 2. Phyllodes with numerous(>8), closely parallel nerves 3. Phyllodes commonly light green, mostly 50-95 mm long; pulvinus slender,cylindric, not llared towards base; headsmostly 30-40-ilowered...... la. A. assimilissubsp. assimilis 3. Phyllodes dark green, mostly 80-140 mm long; pulvinus flared towards its base,often strongly compressed dorsi-ventrally (adaxial or abaxial view); heads mostly50-70-flowered...... lb. A. assimilissubsp. atroviridis 2. Phyllodes 4- or 8-nerved,the nelves sometimesobscure 4. Phyllodes(6)1.5-12(16.5) cm long,deeply sulcate between the 8 stongly raisednerves...... 2. A. aulacophylla 4. Phyllodes shorter and/or not deeply sulcate 5. Pulvinus tlared at its base,appressed-puberulous on its upper sudace; bracteolesacute; phyllodes mostly 25-43 mm long (rarely to 70 mm) with glabrousapical points;heads mostly 20-25-flowered ...... 3 . A. consanguinea 5. Pulvinus i terete,glabrous (except occasionallyhairy in A, uncinella\: bracteolesrounded 6. Headsl0-15-flowered on veryslender peduncles (5)7-13 mm long;phyllodes mostly 25-36 mm long,nerves scarcely discernible,apical point minute and glabrous...... 5. A. ophiolithica 6. Headsl6-31-flowered on peduncles4-6 mm long; phyllodes with discerniblenerves (x10 mag,) and distinct apical Points ?. Phyllodes3.5-7(8) cm long,the apexdrawn out in a delicate, curved,long-acuminate (mostly 2-6 mm), normally pe$istentlyhairy tip ...... ,,,,...... 4.A. fragilis 7. Phyllodesl-3.5 cm long,the apexnot as above(i.e, shorter and/orglabrous)...... 6. A. uncinella 1. Acacia assimilisS. Moore,J. Linn. Soc.,Bot.45:172 (1920) Lectotype(here selected):Bruce Rock, WesternAustralia, 1915,F. Stoward l16,BM; isolecto:BM, PERTH00741477-fragmentexBM. PdraLectotype.Cowcowing,WestemAustralia,1916, F- Stoward 405 (BM, PERTH 00741485and 00741469). Low, rounded, dense,spreading shrrbs or small trees(0.5)l-2.5(4) m tall. Ba* on main stems grey, brown or grey-brown, fissured at base,smooth above. Branchlets terete, light brown, red, grey-redor red-brown,glabrous. New shootsdensely yellow pubescent,the hairs appressed.Stlpules caducous,scarious-chartaceous, narrowly oblong. Phyllodes terete,filiform, (40)50-140(175) mm long, (0.7)0.8-1.3mm diam.,patent or ascendingto occasionallyerect, straight to gently curving, light to Cark green, glabrous (except apex); apex delicately curved, caudate-acuminate,t plumose or appressed-puberulousand glabrescent;pulvinus cylindric or fladng towards base, 1-2.5(3.5) mm long; nervesnumerous, closely parallel, slightly raisedand frequently lighter green than inter-nerve spaces;gland single,2-24(30) mm abovepulvinus, widely elliptic, depressed.Peduncles 2 per axll, (3)6-11(14) mm long, hoary appressed-puberulousor glabrous; basal peduncular bract broadly ',ldSN '-Igy{ 'rug) :(HJU!Id t€61 Sriutu rJ I y'e^\ellnt{ ro g [uDl9l ] selru 0 t :(?9269900HJdAd 't) 'Zl.6I 's8uudg 'J u s uowdDLlCj ]sn8nv 97 eerql Jo r\\ ur{ 0I tnoqB :(HJdad) BBZbllog A 'ulSp?lunt{ 'paulwDra Jo N luq 9 6] selru 9 :VI.MJSnV NUAJSA,^N suawlrads papaps 'spted patrun-679-glleql se Suol s€ Z/l uuql erou sp2das peuasxa 11am xedu eql tseel l" qll^ {rep seloelcBrq:pere.,rou{it)0t-0€(Ll ) spl?eH ^llusuq Surl€U tou 'cupullfr '(fJlerx"p" 'ropuals :o .{11urxuqupe,ner,r) asrq epo11,{qdeql Jo releuerp oqt tnoqr snur^lnd ' lueer8lq8trq ro 1q311,{luouluoc ruerpruur (€'I)I-80(r'0)'8uolruur (09T)96-09{4V)sapo tqd 'BI sJlJurlssedsqns erool S slgurlss€rrlJBJv 'peeq ,qlSuel red sre^\ouJo JequJnueql pw snur^lnd eql Io uuol eqt pu? JnoloJ opoll^qd ur,(1rmur:d ragrp qcrqrrrsercadsqns o,Lr1 sosuduroc solcads oqJ 'oxDt){tJadsDrtul 'sepoll,{qd slr Jo so^teu pesrurf13uo:1s tqSre eql ul {1pe>puulsragrp rnq palular 'pa:nun ,{1asolcos1u sr (,r,io1eqees) urlset{ ry u€^\oJ Sdo KqdoJDUDDtiDxV eroru;o Jpq ,{lprurou 'sludes en sllrwlssD y ur sluled aql spereq,r puu sluted eo4 suq sut?p{ y :Slrudqluvrrad eq1;o 'uorlrppE uorun eql ur reJJrp,{aql uI se,r:eu{uuur se acr,rl lsBel1u q}r,n sepoll,(qdre8uol ,{11ureua8 'saqlulfiv s"q srlralrs.r, y lnq Julnurs,{ll€rJrJlodns eJu ,{leIBIg ? ueprclNsnt?Dtl V pue soraodssrqtr 'ad,{lolcolerud 'ggy 'seurl u se p:u8e: e,r.rqcrq,n p./p,{ols.lo lrcuod,(q paleJEdaslrJ3rr eql uo lelqcuurq Jeqloeuo pue laeqsaql Jo elpprtueql ur 91 1 prr,,ror5 oslu ,{lqeurns-eJdst luq,r Jo slelqcuulqJeqlo eeJql eJpeJeqJ (91I ou) ed,(1o1ce1sr leeqs eql Jo oprsl.+el eql lsarueuueuroeds aql patuu8rsepe^uq e,r puc IAIq1e leeqs ed,{g eq1 uo reeddvsuoqcolloc qloq lo l?rJslr?ur:3ur^\oc,roJ utol} gOt puuItod ecnrq rnotl9Il ptDltrols dr'uoxEl eruEseql Suquese:de:suogcell0c o,rrr1 s1sll en301010 et11 uouqttfid(l 'sueutceds pele^rllnc lueseJdeJ,{eu xeql Jo ,41r1eco1o1lcedser tlll.,'\pelloqpl ,{ltceuoJurflqrssod elB puu sarcedseq1.}o eSuuJ eqt Jo ^le^rlcedseJlse,r puu lse,rqlnos 'e8uug {lqureplsuoc er€ (dn8ur11e,nqulo:1 utl Z€ Surpeq eqt luu.ott'u'sllD\cDlB g11 puu dnuuup 'r Jo tse.^qlnos rrJ{ gz l.IJorttz6tl nuuo1 N N) senadsqnsIsJrd,{l eql Io suoDcellocHlugd regunJ o,tJ serJedsqnso^\l eql uee.^\lequoDEzrprrq{q alqlssod SurlseSSns 'scrlsrrelcurEqc eterpeurJalur suq 'sumo (HJUgd uorulos Jo N uDl 9 I uJor! LVVZunsow y g) uoqcelloc euo (usuasJoN Io qlnos) sveJEuerxesJoN-slunD uolrrlps pu€ (a rC sW.I Jo lsEa) alE8ep,reN-3ur) oIe.I eql ur snonSrluoc 's7p1t1rtot1a eJBselcedsqns o^\t eql Jo se8uelcrqdBJSoe? eq1 dsqns,4q Xla,trsnlcxe lsorulu perdncco '"rTEJlsnV sI e8uur luql Jo pue uJeqlnose{L ulelsai6 lse,{\qtnosJo ue:e ucppEcs-ueurasJoNeql pue IIIH poo^\xog ol ls€eqlnos€^\ellntrltr uro{ uorutuocpue peeJdsepltlsr serJadsa\L uounqu$re 'IBUrUJel-qns Ir.rE .llnp .spr/$ eql :,{uq eloelu :pedpqs-n Jo .rulnc.rrcruesueJSomeld :Iculq o] u,roJq-{Jep urur g T-g l '3uo1 'cqdr11a-3uo1qo ,spod unu E-9 7',{llecrde petrutsuoc f1p:e1u|un ro e1u,,roo1 cr1dr11a ,{1epr,t ur ,paloDBcs pa8uerm ,{lleurpnlr8uol,rpaas furqs JoqtEJ'slelJrEqpessa:dde qlr/h.ro snoJqBISlq8rEJls ',{reded'eptm '3uo1ulu luJeql urur y-g gg-08 uee^\laqpelcrJlsuoc IIE 1E lou ro ,{11q8r1spue spees:a,to pJsIeJrJBeurl ,rpod snolnreqnd-eso1|dudro snolnreqnd-pesserddq,QD^O.otBIoeJuplqo ,ee4,(leJEJ 'Wttutr- 'atulorlrc 'pepunoJ E-Zll sloed /l.orJBu'elB8uoleedqs eql epplq eql ,et€lnqlcds-JeeulJol elelnqrds ',(1pseqpetrun fl otisfJa^ ro eer;'spled;o qfuaIZllr\oqu sf2daS pesredsrelursnoreu-tr ^\eJE ,(leJer'snoJeur-g ,([puuou sJa/i.oll 'pnq ur pegesxe,{lsnor,rqo seruqaluos xedu eql ,painoloJ 'al€rlrc 'e^BcuoJ 'pepunoJ -IJEp saruqeuos Fq^\eruos epelqeql ,repualspuE Suol edrlseqt ,olelnqlsds ' ,,{\o11e,4-uoruel saloalcErq:pare,rou{08)0/-0€(fI) urBrprutu (9)9-t ro ueplo8 1Inpor rq8rrq,prolqo ro ptosdlle dleprrrro1 rc1nqol8speeq :,tllerelul snopreqnd-pessa:ddu,{pq8rts ol snorqels ,pepunol sI ^rrEllmsrl rrrr,y urlschl d g pueue/aoJ s u 240 NuytsiaVol.10, No.2 (1995) Bendering, C.A. Gardner 9461 (CANB, G, NY, PERTH); 8.7 km N from SouthernCross towards Bullfinch, N. HaU H76/45 (PERTH); Wongan Hills, 16 August 1925, E.H. lsing r.a. (PERTTI 00689203);8 km due SSW of Bencubbin,B.R. Maslin 2001 (PERTH);6.5 miles [10.4 km] S of Meredin towards Bruce Rock, B.R. Maslin 2299 (AD, BRI, CANB, K, MEL, PERTH); 4 km N of Daniell Siding, Norseman-SalmonGums road, B.R. Maslin2464 (MEL, PERTH); 10 miles [16 km] W of Moorine Rock, Great EasternHighway, B.R. Maslin 3201 (AD, NSW, PERTH); 7 km N of Tammin towards Korelocking, B.R. Maslin 4420 (PERTH); 2.5 km W of Kulja towards Burakin, B.R. Maslin4445 (MEL, PERTH, TLF);6 miles [9.6 km] N of Lake Biddy, r(. Newbe] 1050(PERTH); 8 miles [2.8 kmlNE of Hyden,K NewDe;y1067 (PERTH);4 km SSW of McDermid Rock, K Newbey 5281 (PERTH); Mount Gibbs, K NewDey5486 (PERTH); StennetRock, c. 48 km SSW of Norseman, K. Newbey1679(PERTH);Wallaroo Rock, c. 72 km WNW ofCoolgardie,K Newbe] 8830 (PERTH); 22.5 km W of Paynes Find, R.A. Saffrey 854 (MEL, PERTH, TLF); i miles [1.6 km] SW of Manmanning, B.H. Smith 385 (AD, BRI, CBG, NSW, PERTH). Distribution. Commonfrorn Mullewa south-southeastto nearLakeBiddy (which is c, 45 km northeast ofLake Crace) andeastto nearNorseman and Coolgardie. Two distributionallyanomalous specimens are referred to above in the discussionof distribution. Habitat. Grows in sand,sandy or gravelly loam oflen in associationwith granite, less frequently in laterite, in mallee heathand scrubon wet, low-lying areason hillocks and mountain slopesand open sandplains. Flowering and fruiting periods. Flowers fiom July to October; pods with mature seedshave been collectedin Decemberand January. Conservationstdt&J. Widespreadand not under threat. lb. Acacia assimilis subsp.atroviridis R.S. Cowan & Maslin, subsp.nov. A subsp.a,rsirnillrp hyltodiis (60)80-140(175) mm longis, l- L5 mm diametro,aeque atroviridibus, pulvino versus basem dilato, saepevalde dorsi-ventralitercompresso (adaxialiter vel abaxialiter visus);capitula cum 50-70(80)floribus, bracteolis non exsertisdiffert. Typas: 83.9 miles [ 135kn] tiorn Albany on Borden-Pingruproad, WesternAustralia, 25 May 1973, A.M.Ashby 4724 (holo:PERTH 00686670;lsa:BRI, CANB, K,MEL, NSW,NY, PERTH00688789). Phyllodes (60)tlo-140(175) rnm long, l- l .5 mm diam.,drying uniformly dark-green,the pulvinus flared towards its base,otten strongly compresseddorsi-ventrally (adaxial or abaxial view). Heads 50-70(80)-flowered;bracteoles neither dark-coloured nor obviously exsened. Sepalshalfor less as long as 2/3-3l4-united petals. Selectedspecimens examined. WESTERN AUSTRALIA: 30 km W of 90 Mile Tank, T.E.H. Aplin andM.E. Trudgen5917 (K, PERTH); 88 miles [14] km] from Albany on Borden-Pingruproad [c. 20 km N of Bordenl, A.M. Ashby 4733 (8, BM, MEL, NY, PERTH, W); 28 km due E of Lake King Gate on Rabbit ProofFence,33" 04' 30"S, 120" 19'8, K. Bradby 39 (PERTH); 32 km ESE ofMuckinwobert Rock on West Point Road,33' 28'S,120'37'8, M.A. Burgman1050 ar'dS. McNee (PERTH); Boxwood Hill-Toompup road, 11 km NW from Chillilup Pool turnoff, 34' l7'S, 118' 3l'8, M.D. Crisp 5163 (PERTH); Fitzgerald River, CA. Gardner 9259 (PERTH); Oldfield Location 1002, near Dallinup '(^N ') 'BNVJ I'll :of?:ELZ18l00 HJUgd :oi04) 'erruqlnleehl uDt g :s'd'q V69Eullsory U'g'VL6l,{lnf 0t'€Ilurlsnv ulelseiA spnr.^dolenJJoN T 'IputlrrJe]-qnsolluB tr8lu eJncsqo'ElelsuctuepulpnllSuol tuex! uIncJIc'ulel ultu 'e8uol 'Essalduroa'ucDdllla 'u11uutpn1t3uo1 'stlcutlsrpsnqtluur8ruur g Z uIuI g t elDI aararas slpllll?d 'Bttal 'eeJ€uoc 'elnpuad 'e1e1 'e8uol 'elclqsuoc 'olJEeuII sr^reu '€JqBl8 urur 9-9 tutu 00I-09 uou 'srlBrlrt 'suElnqleds 'slleJcslp 'slulpntlSuol DuMrnBaT sItBuuoJ-t/€-ZlI slp@d srlpllrJlp tJJrc 'sp€tled-qns 'else8uoa unrelelad stlDdas,uew-g satolC, sqprlr3'sllnJeqnd p€ sllulnqlEdsslloelcBJq 'snqIJoU-18-0?'orteruulp 'eIJEInqolS esuep uul 9-9'eelnl tln1rdu3 urqel3 1e,lulnraqnd-ossardde 'IImaqnd-osserddu'serorBuol '€lullrc 'vl€ nJnc'ElE^o attl '"cnpPaill?s€q €elcerq unJolnounped orer 'plllxe 'unul^lnd 'rguolrurrl (0I)8-g(t) enbonbur Zle^ | ry Junpad Ecudllle-ol€lIo^ llElnJrrr uldns 'B]cunslp 'sqe,ra1e 'snqIpIJI^-oIpetu -o1n1rp'srrqc13 'srtr:le pe rutuS [-Z apuoi3 erutssrpp^sI^Jeu 8 le^ 'oJleruElp 'slSuol -Sr(09) 'o8uol snquuepuacs"'slplSu-Irues tutu (9 1)I rutx (S9I )02 I slu[uEl uru t-Z 'elnJeqnd-osseJdde-qns 'cluululnoE 'lII'eJ8 oul^lnd 'equecserqulSuluud runpJalul Ie^ -osseJdde 'snl€punloJ -o8uol elBncJ"'e tle JetoryollK\d sI.IqElS'snqlleJel sllnurPJ edens'snlle u t I ra''lJ '^ou 'Z ds'uIIS€I ? uu,no3 5 g u11.{qdoJBlnEBIJBJY 'ueel8 JoI pJo/r'e 'slpu!^ pus 'llep Suluteur sr.,rllD 'spro,r urlu'I o.rt ruo4 'uoxel srqlroJ aureu eq1 pe1sa88ns sepoll,{qd poup eqlJo rnopc eq; KSopa{Jg 'snl?its ']eeJqlJepun 1ou pue puardsaptl6 uollD^JasuoJ 'ex!l 'sueJEJ9llulls uI lncco qloq Jo suopcellocSut,fttlueplsltu Jo ,{llllqlssod lual u sl eJeqlos '(ecuj 'lul+ ,(eql ',{lleclqdurSoeg pe^leulllnu puo eleJelSuleq uEql Jeqlel led se^Jau€) p3^leu-8 eJu 'fSoloqdrou ot\ctldltg 'y p sapoll.(qdeql t?ql uI IeJJrp.(eqJ seloalccJqpuu slcerq;o utro; Surpnlcur '1en111',goqcKrlyt y o1 rclttuts altnb sl Je,rog Jo slrplep ur ue^e puB ecusJ?eddBuI qluag xo slpyt^ulo sorledsgns 'selsadseql Jepunpessncslp aJ? selcedsqns eql u?e^ueqs atuanlyq sattrutlly 'Je/$ouur osle eJElEql slueld uo Jncco{eur spod Sutdole,req Jaqlllecac pw Jeque^oN ut pelcellos 'sPouad ueeq e^eq speeselnl€u qll^\ spod :JEa,{eql lnoqSnolq! srewold Suun$ purt 3ulno\oll 'qleaq puE pu"lpoo/r\ 'qnJcsuI ellJelfl 'luuol 'puESrrt r11r,lrfluouruoc ssal'elru?r8 qtt,^A uorl"IcossE ul uelJo ,{lle^"J8ro fpuEs s/1'oJ9 JDIlqoH '€llEJlsnv uJelsei[ '?ele lse,rqlnos u"ppBcs-I]Burnl eq1ur ,(u,tq341acuuredsg-etp:e81ooJ ol lsEepuD (dnSunurBJJef Jo lsa^\qlnos-qlnosuI0g ,) IIIH poo,t\xogJBeu ol qlnos3un,(elqunc Jtau u]o4 uoluuto) uounqu$lo '(HIUAd) lggg g91ytg ta131t14,t1 1 'etu8ap1(eN '|Jlreln>l spJe..l\olSutl elel uror; uq € I I :(HIUAd) u4,rDIAtA g pue 991 alDpulJ'O'W 'd'8'196I sprB/r\olSun,telquno Jo gN [uD{ Il,l] sellur 601 :(811.68900HJUgd) ?./'ssuaqov ')..rDd 'sqqro ,(€N 5lcou lloH-,{olre^ aIE'I :(HtrUAd) VVS,{aq$aN X IeuolluN uuBH{ueIC lunol^lIo 'lt\SN '>I S ur{ 9 :(HJUAd) tBEg bqi/'aN )/'Irud puolleN salEIlJ {Eod'€rouEetg {"ed :(HJdSd 'C'SNVC) g Kaq aN'X'eaerCe){?-I Jo S [uDI9 6] sellul91(HJUAd'gNVJ) €9?9ullsow V EZgt 'l,'AV) ',(u^\q8rHecuBredsg-arpE8looJ uo sunC uoules.Jo S UDI0Z :(HJUAd 999Z unsDw Ag 'dnuISunI 'OI,,{) 't 'S,tZ 'deruorl uDl :(HIUAd Z 8LEtraqSDX 9'g.,lt .6I I .f € tluoN Jo S I Jo td 0I 'S,t€ ll]){ Z:(HJUAd) Z0I K.nqroH V 7'tll.wryt:(HJuAd) €ZrSl28roa9'S V'A,}V.0ZI o€€learC gl j(uullorsll rrrrty'utlsBl,{ u gpue ue^\oJ Su a 242 Nuyrsiavol.10, No.2 (1995) Often roundedshrrbs l-4 m tall, bushy but becomingopenly branchedwith age,single-stemmed or branchedat ground level. Bark grey, smooth throughoutor fissuredonly at baseof main stems. Branchlets tercte, glabrous,red-brown ageinggrey. New shootsoften golden appressed-puberulous. Strpulesonly on very young branchlets,lanceolate-oblong, 1.5-2 mm long, 0.6-1 mm wide, scarious, appressed-puberulous.Phyllodes terete, (60)75-120(165)mm tong, 1(1.5) nn diam., semi-rigid, ascendingto erect, straight, glabrous,light to medium green; apex long-acuminate,the delicate tip arcuate,appressed to subappressed-puberulousand sometimespartly glabrescent;pulvinus slender, 2-4 mm long; nerves8, very stronglyraised and deeplysulcate betweent gland distinct, 2- 15 mm above pulvinus, circular or widely elliptic. PedunclesI or 2 per axil, (4)5-8(10) mm long, rarely longer, appressed-puberulous;basal peduncular bract caducous,broadly ovate,cucullate, ciliate, appressed- puberulousor glabrous; headsglobular, golden, 5-6 mm diam., 40-87-flowered,densely congested; bracteolesspathulate to subpeltate,puberulous, ciliate. Flowers 5-merous. SepcLrabout half as long as petals,fiee, spathulate,ciliate. Petals ll2-314-united. Ovarl denselyappressed-puberulous. Pods linear, not strongly raised over nor constricted between seeds,60-100 mm long, 5-6 mm wide, pendulous, coriaceous, straight, glabrous, the marginal n€rves distinct, light-coloured. Seeds Iongitudinallyarranged in pods,widely elliptic,compressed,4.5 mm long,2.5 mm wide, crested around long axis, dull, black; pleurogramcrescent-shaped; the aril sub-terminal. Selectedspecimens examined. WESTERN AUSTRALIA: Watheroo,August 1903,C. Antlrews s.n. (PERTH 00187283); Mary Spring, Kalbarri National Park boundary, D.R. BelLairs 1609 (MEL, PERTH); East Yuna Reserve,NE of Geraldton,A.C.Burns 9 (PERTH); No. 8 Tank, Tching Range, Nookawarra Station, R./ CranJield 5181 (G, PERTH); 1.5 km S of Jingemarra Homestead, R.J. CranJield 5255 (PERTH); 3 I .3 miles [50. I km] S of Yalgoo towardsPaynes Find, R. Cumming 1993(AD, NSW, PERTH);6 miles[9.6 km]E of Morawa,J.R. Kro.r 650813 (PERTH); about 6.5 km Nof MountMagnet towards Cue, B.R. Maslltr 3582 (PERTH);Gnows NestRange,50 km SEofYalgoo towards PaynesFind, B.R. Maslin 4254 (NY, PERTH); 5 km E of Mullewa on road to Yalgoo, B.R. Maslin 5075 (PERTH); 4l km fiom Byro Homesteadon the track to Milly Milly Srarion, B.R. Muslin 5175 (BM, MO, PERTH); Donkey Paddock,Coodardy Station, A.A. Mitchell 759 (PERTH); 9 miles [l4.4 km] N of MadoongaHomestead, N,H. Speck 1072 (BRI, DNA, NSW, PERTH)- Distibution. Found from Byro Station(.. 250 km west-northwestofMeekatharra) southto Mull€wa and Morawa and extendinginland to Cue (c. 65 krn north of Mount Magnet),Western Australia. Two collsctionswere made slightly outsidethis distributionat Mary Spring,c. 120 km northwestof Mullewa and at Watheroo,c. 120km southof Morawa. Habitet. On latefite or granite hills, outcropsand breakawayson sand,rocky sand,clay or loam in open scrub, often dominated by Acacia species. The occunence of this specieson hilltops and breakawaysmay in largepart accountfor its discontinuousdistribution. Flowering andfruiting periods. Flowers mostlyfrorn April to August but one collection (8.R. Maslin 5075), with maturs flowers and fruit, was made in December;pods with mature seedshave been collected in December. Affinities. The deeply furowed phyllodes with strongly raisednerves, distinguish A, aulncophylla from all its.relatives,of which A. asslallls is perhapsclosest, in spite of the difference in number of nervesin the phyllodes; superficially the two taxa can easily be confused. Acacia acuminata srbsp. burkittti (F. Muell. ex Benth.) Tindale & Kodela (ms) which is common in the geographicrange of the new specieshas a similar habit and foliage colour but is readily distinguishedby its spicate inflorescencesand t moniliform pods which ue 4-7 mrn wide. \ '8ury luDl € l l selru 8 :(H 1Ag4) 9gg7 Sutwwtt3 y eqel spJ€/t\ot[.r€lxn) Jo A\ [uDt 8 29] selltu Z 6t 'fA 'A,EV 'S,t€ 'ulltEpog :(HJdgd) gStZ plr {uDtJ .BI I " l€ Jo l!\S tu)t 9Z :(HJUAd) KaUaqoH B 'S,02 'reBuooN pue g LEB[ua^o3 y'4,89 "8t I .l t :(HJdAd) Z99Z 89/V76 Sutuurt3 713'.{u.tq8t11 urelsugl€ero SuolEqued sprP.{\otlcod eulloot{ ruo4 uq / 9t:(HJUAd fgtz 89lVl{^SutuuDJ Wg ',(p,\\q8rH uralseg l€eJD 3uo1e sso:3 ureqlnos sprE^\ol slprE8looJ urorJ tul 6 tS :(HIUAd '),'g) 'A VZOVllt trqg AI'uIqErrBJ :VI-lVdJSnV NdSISAA\. paulw2ra suawnads patraps 'palseJc'l!]alul-qns II.IE:elnurru sloale :mlncJlclluosure]SoJnold :u,\\olq ',{ssol8'lrrql 'epl,!\ '8uol 'slxu -,{er8 elud uo u,^\orqJe)llcp pelllour luul I I rutu 8 l-9 I urtu Z Z-Z 'spod 3uo[ punolE pelseJc'unlrq JBaupetcrJlsuoc 'crtdr1le ,{1apr,tr ut pa8uu.ue{11uulpn1t8uo1 spaag ',(ssol8 'atElnpun 'snoacl?lnqc-ulql'ePL\\ '3uo1 'snoJqelS pue pe,\Jnc,{Fq8rls ulur g-E 7 tuul 09-09 'ruoql uee^\lsq pelclJlsuoc ,{lslclepotu puu sp33s la^o seprs SuIluuJell" uo pesl€J 'eeU 'elqoaculllqo ,{lSuoJls'JEeurl .!pod snolnraqnd-orctruf-rD^O snoJqelB'pellun-JIuq ^lelEJ s/rlad 'elElorlrc'elplnqlpds-JEourl'aa4'qlSuel 'rttdr;1e lEtedeqtJlEq lnoqc.tlarlag snoreru-9 s'raaroiS a1n:rc 'Pa ssel Jo eJou Euruel eql 'e]elllc 'snolnroqnd '€lnfi 'elulnqlEds-oll?loacu€lqo esxe seloolculq :pare.rot'-(62)SZ-02(tI) "ruulp urur g-tr 'uep1o3'rnInqo13 spteq :snolueqnd-pesserddoflelnulru 'pepunor'el"llncnc lcrrq rclncunpedlusuq :snolnreqnd-pesserddu,4lesruds,{llcuolsBcco ro sno;qv18 '3uol 'lIru 'pesseJdep'snur^ -9(?)'elqrsr^ ruur(8)9-€ nd7sapunpa4 cqdlIle lnde^oqt uu ( € I )0l lou ueuo'elnunu puulS:s,rollnj JplnBeJJtssel ro eiolll'deep ^q pelvrvdespue pesrcr3ut:eeddu ue13o e1e1s 'tuutsrp'euEld 'arncsqo ?ftpur uauo'8 se^Jau:oauJtnsIuIXtpE uo snolrueqnd-pesserddupua esuq lu 'pel{uu,4^ 'eluuoJcnur'pe,unc pelBlrp ^les:e,rsue4,(putul ol qloous'3uoltutu Z-S I snul^lnd :snoJq€13 'po^Jncul{laolluqs xad€:uaeJ8-{JEp'snoJqulS'proru8ls,{1,no1pqsro lq8ru.tts sellrneulos ,{luotutuoc 'Surpuocse 'pl8u 'tuelp '3uol -SZ6d 'elpJ€t ol luelud trut (l, t)S I-Z I(I) urur (ODt sapolltuld 'snocnpzc sairdu5 ,{e.r3'snoln:eqnd-pesseJdde'elerel,ualqJuwg'qloouts'^at37-rtrg1e,r.e1punor8 'llul 'popunoj 'potlcuuJq l€ paqruuJq-qcnu ru 9 t-? 0 sqzlqs Sutpuerds ue o fluedo ro esueq '(IN 'tdSN ') '9 'SNVC '.E6LZVE0O 'A'g ;ost Hld3.d:olo1l) 0681 uusow 'lZ6T 'Bllullsnv 'IcoU lsn8nv 8 ulatselN et,{ueu3 spre,nol 3ur11nqu11ng.+o,14.N uDl Ol :sndKJ 'olusllJ'rluelEI-qnsollrJf, 'slauunlq-oasr:3-otnltp ul sltuunlq 'BprJnl'arllssoJc 'Elul 'e8uol 'crt1dt11e srlnceu runc Blulncuur tutu 9l tutu 8 l-9 1 ww Z'Z-Z atul 'uJqBIS'eeJe 'Blulnpun 'ulul 'e\eulpnl\3uol oulwas ?prcnl "qc Jeltnuel le "tur\Jnclell^el tutu €-9 2 'B8uol 'ultlJlsuoc '€lD^ala 'utrceutl turu 0g-0€ Jell\el eurues Jalul eplE^ tutleulollE cutues udns DutwngaT 'squcslp enbslizlad slpdas 'ueuJEtuedsanl! e1j.\aB'ecgdt11ee,tsnutu snld uululel 'srlErlrs 'suesxe 'sngltow-Gdsz-0zol) lo srlnreqnd'srl€lnrllEds-oleloeJuBlqo orlsEqBluur slloalcurq 'BrJEInqolS la oJleuerp tuur g-? ulnlrdec :uqBl8 anbulnleld le^ IlnJeqnd-osseJddua,tsnulru snld'r8uoluur (8)9-E'BIlrxtrenbonb uI Zlprunpad tunur^lnd€Idns rutu (€ I )01-9(t) ElnuItuepu€13 'elE 'EluncJu 'occrs ur srl€,\eleaple,r enburn:e1dpos sruBldsl\Jeu Jsu-8 eproul8ls lelL\el [eA ?lJaJ 'ElluapuoJsu 'epl8u 'oJleuIEIp unp.rolur pas ulp^JnJurJell^el oBIn,\ pu cqueled ullu (/ I)9 I-Z I(t) 'u8uol ruru (ODw-gZ6d 'ole8nJJocletll€sra^su€.It lell^el pB I^eEl 'olElrpl-tlescq pB le olnJJqnd -osserdduruelrxupe ruuut8ud pe out,llnd 'oJqEIS'oluuoJtnul 'olencJE ec]dD 'ulleJet olpontuld sreJ€Iu]x 'srlnJeqnd-osseJddB'snqrloJal 'EeJeuIc'I^eEl 'snlp srlmuuj eJIlJoJ ur I l-t 0 sIInecDInlrJralr./C ' '€ ou ds'urlszl ? ue,a\oJ S U "auF8uesuoJ elJaJY 'IEel ',solorlo ',sptot rcI UollKlld puE ,'!\oJJnJE Jo Suruueu eql qI/\\ '(3oplatlq I?erC pazrurlBlo,rl ruo4 laqlde clJlcedseql.IoJ slsEq eql epr,rordsepo11,{qd pa,lr ounl eqJ- 'FuelseJql loN Swols uorlD\lasuo) 9I,{uyllarsrli,rrrty'ullser{ U gpueue,toJ S d NuytsiaVol. 10,No. 2 (1995) S of Bonnie Rock, J. Goodwin 87(1262) (PERTH); 25 km W of Bullabulling on Great Eastem Highway, A.R. Maslin 2413 (AD, CANB, K, MEL, NSW, NY, PERTH); 10 km by road S of Queen Victoria Rock, B.R. Maslin 5408 (PERTH); about l2 km N of SouthemCross on road to Bullfinch, B.R. Mastin 6711 (PERTH): 2 km S of Karalee, c. 50 km E of SouthernCross, ,f(. Newbey 6O32 (PERTH); 0.5 kn E of Marvel Loch turn-off on Great Eastern Highway, M.H. Simmons 1204 (PERTH); Muntadgin, T.W. Stone andE.T. Bailey 816 (PERTH) andT.W. Stone866 (PERTH); 12 milesu9.2 kml NW of Wialki,4 October1958, G.M. Stdrrs.t. (PERTH00750158); Ghooli, 15 km Eof SouthernCross along Great EasternHighway, A. Strid20035 (AD, BRI, CANB, G, MEL, MO, NSW. PERTH). Distribution. Locally common from Muntadgin (c. 40 km southeastof Merredin) east to near Coolgardie with most collections from along the Great Eastern Highway, southwest Western Australia. There are two outlying populations,the first representedby G.M. Storr s.n. (PERTH 00750158) fiom nearWialki (c, 150 km north-northwestof Muntadgin) and the secondrepresented by R. Cumming 2556 (PERTH) from c. 63 km west of Kumarl (Kumarl is c. 200 km south of Coolgardie). Habitat. Found on sand,sandy loam or sandy clay in heathor densescrub. FLoweringandfruiting periocls.Flowers mainly in September,occasionally August; podswith mature seedshave been collected in December. Affinities. The new speciesis closely relatedto A.tcgills Maiden & Blakely andA' uncinellaBenth. (seebelow) from both of which it difTersin having the pulvinus appressed-puberulouson its adaxial surfaceand flared towardsits base,gland minute,bracteoles acute and slightly exsertedin the bud and seedsnearly rotund. The phyllode apex ofA. consanguineais more like A. uncineLlain form and in being glabrousbut the headsare larger as in A. fragiLis. Flower morphologyalso links the new species more closely with A. fragilis; both have free sepalsand petals in the vast majority of specimens examined. GeographicallyA. /ragilis andA. consanguineaoccur in morc northem areasthan does A. uncineLla- Conservqtionstaras. Not under threat. Etymology. The closerelationship ofthe specieswith other speciesofthis alliancesuggests the name, frorn consctnguineus,Latin fbr kindred, relatedby blood. 4. Acacia fragilis Maiden & Blakely, J. Roy. Soc.Western Australia 13: 5, pl. 4, figs l-l 1 (1928) Based on A. triptycha var. tenuis Maiden, J. & Proc. Roy. Soc. New South Wales 53: 178 (1920). ?ypas: Tammin, West€rnAustralia, September [sPralm. Novemberin protolog'te] 1909,J'H' Maiden s.n. (holo: NSW; lso: K, NSW, PERTH 00963828 and 00859435-fragmentex NSW). Dense, rounded shruls, becoming more openly branchedwith aEe,0.5-2.5 m tall, spreading 2.5-3 macross; stemsto 10 cm diam. atbase. Bark grey,finely fissuredbasally. Branchlets tercte, glabrous,red-brown, becoming grey with separationof epidermis from lower layers. New shoots golden-sericeous.Stlprles only on very young shoots,lanceolatelinear, scarious, obtuse. 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They are similar to A. uncinella in having a half-unitedcorolla and, to some extent, in the phyllode apex which is otten glabrous but also often elongate as in A. fragilis. There is a need to examinemore closely the significanceof the petal union characterin a comprehensivereview of the A. fragiLis-A. uncinelLarelatlorlship. Another variant occurs near Bungalla,c. l0 kmeastof Kellerberrrn(e.g. B.R. Maslln 3394);itis characterizedby havingphyllodes near the tip of the branchletswith the apex glabrous but some of the phyllodes farther down the branchletswith plumose tips. Affinities. A highly variable speciesdistinguishable, somewhat arbitrarily, from A. uncinellaBenth. (seebelow), its closestrelative, which generallyoccurs farther south,although the two overlap in the central wheatbeltfrom Bungalla to near Boorabbin (Boorabbin is c. 80 km east of SouthernCross). All charactersvruy to someextent but, in general,A. fragilis haslonger phyllodes(mostly 3.5-7 cm), the apex of which terminatesin a long to very long, delicate,gently curved, hairy tip (mostly 2-6 mm but up to 7 mm long); the pub€scenceon the tip variesfrom appressedto spreading:it is alwayspresent onjuvenile phyllodes and usually persistson at leastsome ofthe matureones. Acacia uncinella has shorter phyllodes (1-3.5 cm) generallywith a shorter,sub-uncinate-mucronate tip (0.8-1.5mm long) that is only minutely appressed-puberulousat filst but commoniy soon glabrescent. The petals of A. uncinella are mostly half-united but in A. /ra8ilis this characterappears sporadically without any obvious patternexcept tbr the Ravensthorpevariant discussed above. The possibility ofhybridization betweenthetwo speciesshould receive serious attention, for thereare collections in the zoneofoverlap (i.e. the central wheatbeltfrom Bungallato Boorabbin)which cannotbe cont'identlyassigned to either taxon. Clearly much more detailed study is neededto elucidate this complex group and in the meantimeit is consideredprudent to retainthe currentclassification and not pre-emptivelyeffect name cnanges. ConseryationsldtaJ. Widespreadand not consideredrare or endangered. 5. Acacia ophiolithica R.S.Cowan & Maslin,sp. nov. Fr&te,vrotundatus 0.3-2 rn altus, ramulis teretibusad leviter angulatis,glabris rubello-castaneis. Phyllodia teretia, oblique et abruptemucronata, saepe rostriformia vel subuncinata,pulvino gracili, 1.5-2mm longo,croceo, a laminamabrupte separato, laminis (20)25-36(45) mm longis,0.7-l mm diametro,erectis ad asoendentibus,rectis, glabris, pallido- ad vivido- vel atro-viridibus,4 vel 8 nervis immersis perobscuris,glande saepenon manifesta,minuta, 6-14 mm supra pulvinum. Pedunculi 2 in quoqueaxilla, (5)7- l3 mrn longi, filifbrmes, glabri, pedunculorumbractea basali caduca, cucullata, glabra. Capitula globularia,5 rnm (3-3.5 mm in sicco) diametro, l0-15-floribus,bracteolis spathulatis,brevi-stipitatis et laminisellipticis, ovatis vel lanceolatis,plus minusve concavis, ciliatis, Fktres 5-meri, petalarum sepalis minus quarn dinidiis longitudinis, discretis, linead-spathulatis usquead spathulatis,parce ciliatis, petalis discretis. Legumina lrr'eatra,30-37 mmlonga,2.5-3.5 mm lata, deflexa, ohartacea,recta vel leviter curvata, glabra, lucida, marginibus distincte incrassatis. Seziza longitudinalia, oblonga usque ad elliptica, ad apicem unilateraliter constricta,2.4-2.8 mm longa, 1.3-1.6mm lata,subcompressa, lucido-brunnea, arillo sub-terminali. Typas: adjacent to JerdacuttupRiver on North JerdacuttupRoad, 10.5 km E of Ravensthorpe- Hopetounroad, WesternAustralia, 3l August 1980,B.R. Maslin 4789 (holo: PERTH 00149381:i.so; AD, BRI, CANB, K, MEL, NSW, NY). lA. uncinella auct. non Benth.: G. Bentham,Fl. Austral. 2: 341 (1864), as to the Maxwell specimen cited.l '1ue1deq1 go (1oor aurluedresuroq pe^rJepsllos) elersqns 'lelrqeq luenbe4 eql rog ur1u1'sncryqotqdo tuo4 s1ro1 sre;er sercedssql Jo etupu eq1- [SolowKig '1\r-l€Joll {tr.ror.16pun JrnU pJrrlJeC s,luerueS"uuJ,^{pu"'I pue uorlc,\JesuoJJo lueu!.lt?deoeql uo uoxr} Z ,{]rJoudv sntDlsuorto^tasuoJ 'spod peuoc sselJo eJo(upuE seuec€JpapEeq-Z oqs ur spyeqra8Jl?l 'sepol1,{qd 'pe^Jeu-€ esnlqo flerncsqo suq qcrq.r,r(Z0Z :8L61) urFuINDJDulq y ,{e,{\lereue8 e uI selqueser puB qgr,rrcqedur,{s st 1l o1awJun y se "ar" re^r1 plerJplo eql ruoJ.}serceds aql ;o uoncollo? lle,\\xul4l 3 palrc,{llEnlrB(lt€;t98I) ur€qtu3gpuu (.roteq aos) qTuegDlpulJunye1l1 lsoursr lt ecuureedde 'llcrus 'sepo11,{qd IleJe^ouI spueqpera,roU-^\eJ fla^qelar puc selcunpedJepuels ?uo1 pa,r.reu-gro -y ,(leJncsqof.re^ 'Jepuols'lglrqvq sllJo esnu??qdnor8 srqtur i^or.odsa rlcurtsrp ,{JeAu sr slrll salt1u![[V 'JequeJecur palcallotueaq e^eq speesarnlEu r{lr/( spod:Jaqolcoo}reqruordeS (sn8ny) uo.g sl.,i'olJ spouadqulqw[puD 3u1Ja$olc 'auquedrasuorJ pa^uepslros uo ^DuonbeJl'suorlulndodesuep Suturrol ualJo 'sld{lBcne eelpu qlEeueqpw $Ju"q ro^rJ uo ',{lcoJ seluqeuos '!uBol-^ElaJo AElxrO lDtlqDH 'BrleJlsnv 're^rU uratselAls3/hqtnos dnuncEpJeleql lo ,{trurcr^eqt ur (drqsu,rol edroqti^uo^l?ulo lsue ,4llq8Its) oBUEUedJoqlsue^uu eql Io lsse BoJEllprus B ur uouluror X11uco1puu ol peloulsex uolnqu$lo '(o€€6ntoo Pur Eot6tloo ',,\\SN '1ElN) 'Ie^rU 'HIUAd 'Ohl 'D 'ANVJ HJUAd u s ad\ww g pler.lplo Jo ^rutnqul ,^(\:(Z 'r\Ig) $VgUUSvW yg'acuuredsgo1 1 o1q,{u,,nq8rguo ed.roqlsua,rrXlo g ur)tg ZZ :(HId!Id) €90? 'HIU!Id 'VNe'g) 'eJueradsg pup (l\\ lZ6EultsDW y g sp.u7,ro1adroq1sue,,iuX1o g ur)t I I :(HJUAd) ',roqle E80rl JauWDg yg raarg dntrnrnp:e1:(HJUAd) urspw I!B pulT lgLV uDA4o) SA 'acuBjedsg 'y spru,,vroledroqlsue,red Jo g uq I I :(HJUAd) aaN)W S puu OVLZ uowtung 1,a'1 'edroqlsue^lu 'IN '-Ightr ') 'CV) '6L61 Jo gS ru{L 8Z :Qzt6fl00 HIUAd ?r's auuagW g 'adroqlsue^Eu lsn8nY 0Z Jo A tul gl , :VI"IVUJSnV NUA J,sgl.]l. paulluDxasuawnddr^ PanaPS 'l€uluJefqns '^ssol3'PesseJdluoc lue:,{uUeloers ipadpqs-n urerSornald :u,toJq 'epl^\ '3uo1 'xedB '.rtdrlla 'spod -qns urur 9 t-€ l ruru g 6-2 7 lE ,{llErolBlrunpolcrlsuo. ol 8uo1qo uI pe8uuru ,{1purpn1r8uo1.rpaas peue{crqt'snoncrdsuoc sur8rulU eql'u,roJq-peJ'3urultls'snojqtlS 'pa^Jnc 'snoec€u€qc' 'S 9t ^u?llcrsrtrAl,Drry urtsrtr^t U g puDuE^1oJ d 248 NuytsiaVol.10, No.2 (1995) 6. Acacia uncinella Benth., Linnaea 26: 6l3 (1855) ?)prs: interior of WesternAustralia, -/.S. Roe s.n. (hoLo.K; iso: PERTH 008500i2-fragment ex K). Low, rounded s/rrabs0.3-3 m tall, spreading1-1.5 m across,with stemsto 7 cm diam. Barklight grey, fibrous, snrooth. Brunchletstercte to slightly angled,glabrous or denselyvillose to more or less appressed-villoseor appressed-puberulous,glabrescent, tan tobrown-red atfirst, becominggrey. New shoots white- ol yellow-appressed-puberulousor villose. Stipules only seen on youngest shoots, lanceolate-linear,obtuse, scarious, light brown. Phyllodes terete to compressed,l0-35 mm long, 0.?-1.5 mm diam., spreadingto ascendingor erect, straightor shallowly curved, sometimes sigrnoidally, glabrous, glossy, light to dark Sreen; apex mucronate,sometimes abruptly so, sub- uncinate,the tip 0.8-1 .5 mm long,glabrous, minutely appressed-puberulous or puberulous and more or less glabrescent;pulvinus ofien abruptly separated,l-2 mm long, glabrous or puberulous to appressed-puberulous;nerves 8, distant, raisedto plane,olten obscuredby furrows in drying; gland 4-11(20) mrn abovepulvinus, depressed, elliptic, usually in angleof two coalescingadaxial nerves. Peduncle.s2 per axil, (2)4-5 mrn long, puberulous to appressed-puberulousto glabrous; basal peduncular bract cucullate, rounded, glabrous to appressed-puberulous;heads globular, golden, 2.8-4.5mm diarn., l6-30-flowered:bracteoles spathulate to peltate-spathulate,the bladerounded, puberulous,ciliate, concave. F/owers 5-merous. SepaLsfree, about half as long as petals,naffowly spathufate,ciliate. Petals about l/2-united, occasionallyfree. Ovary appressed-puberulous.Pods linear,slightly rrised over and constrictedbetween seeds,40-50 mm long,2.5-3.5mm wide, thin- crustaceous,stfaight, glabrous or sparinglyappressed-puberulous. S€sdr afianged longitudinally in thepods, elliptic or oblong-ellipticwith terminalconstriction, 2.5-3.5 mm long, 1.8-2mm wide,sub- nitid, mottleddark and pale blown, nigrescent;aril sub-terminal,crested. Selectedspecimens examined. WESTERN AUSTRALIA: nearBoorabbin, T.E.H.APlin 1935 (8, G, K, NSW, PERTH);Pingrup, W.E Blackall3033(PERTH);0.2 km E of VarleyGate on RabbitProof Fence,32'45'S, ll9" 43'30'8, K. Bratlby24 (PERTH);King Rocks,P.E Conrick1563 (PERTH); Wittenoom Hills, N.M Donner 2888 (PERTH); c. 95 km WSW of Coolgardie on Great Easten Highway, N.N. Dctnner 4556 (PERTH); c. 3 km N of Bungalla towardsWyalkatchem, B.R. Maslin 3390 (AD, BRI, CANB, G, MO, NSW, PERTH); 27 km E of Newdegatetowards Lake King, B.R.Maslin3863A (NY, PERTH);c. 2.5 km dueNW ofNo TreeHill, 20 km dueNW ofHopetoun, B.R. Maslin 3892 (CANB, K, MEL, NSW, PERTH); I km E of Lake King, B.R. Maslin 4064A (PERTH); l8 miles {21J.8kml N of Ongerup,K Newbey2474 (PERTH);Hatter H1ll,K. Newbey3295 (MEL, MO, PERTH); 6.5 km E of Lake Ace, K. Newbey8039 (PERTH); 33 km SW of Buningonia Spring,c. 70 knr SSW ofzanthuq K. Newhef 8251 (PERTH); 18.5km N ofHyden on roadto Anderson Rocks, /.G. an<\M.H. Simmon.s1309 (PERTH); 54 km W of Kumarl, P.G. Wilson 5698 (MEL, PERTH): 33 km E of Lake King at No. I Rabbit Proof Fence,P.G. Mlson 5750 (K, NSW, PERTH). Distribution. Most common in an areabounded by Ongerup,east to the Wittenoom Hills (c. 50 km north-northeastof Esperance)and north to Anderson Rocks (c. 30 km north of Hyden), southwest Western Australia. Four populationshave been recorded from slightly north of this area(and within the rcnge of A. .frdgllis) fiom nearMerredin, Trayning, Bungalla (c. 10 km eastof Kellerberin) and Boorabbin (c. 80 krn eastof SouthernCross); a collection from BuningoniaSpring, southofZanthus (c. 185 km no heastof Norseman),represents the most easterlyrecord for the species. Hebitat. Sand,loarn anclrocky loam along road verges,mallee scruband proteaceousshrubland, on slopes,hilltops and sandplains,often near salt llats or lakes. '(uorssncsrp ' 'IAhl 't061 IoJ€t66I u€,roJ ? urls"hl ees ^ r tnq qpq) 866qrox n requeldes 'BIIuIlsnV '3ur,roc,,AoJ puvlsn8ny urolsel[ :adK]uKS(ll6D L-SsSlJ'09 Id'L8Z:Z td'€Z res u euogctn cog ,4og coJd'poo/!|g T, elrql[ uEeI'IB^\g I V uI allql[ ul?elx€ 8ztlC l\\ttuDluD6raV '(g xa luetu8"{-t988tr00 HJUSd '7 '106I 'ururupJ ios!)6982 qal1 ,{EII IZ'erlBJlsnvurelsa/d Jo S url I'tcutslpuo^VuI :st?dtrJ ' 'I G06l) V6Z:9t lsfs qrq€t rog zrud g [qa!p BIJETV " """ Dsaqov E elrssesspreH I "" Datlu V Z SnOUIS9J '?lrlnpun stalqcueJq:speas elrllrr€ eqt uee^\leqpelclJlsuoc lou spod z !!sla!p'V'l snoursbJtou stalqcuPrq:spees ettllu€Yo eql 'elclnpun uee,rleq suoqculsuoc,rorrEu 1u ,(|prer Suqee.rq lou spod Z etslncunpeospeeH I 'y,, ,dnot5 11s7a1p ;o sagcadso1 fay Josdrqsuou'rau spod eqr ur purpnlr'uol spees puu :snoJau-,.r"^"ni[:]ilil^iil tllll?;:tlH 'snoncouur'e1ere1 '(3uo1 spEeq:snolqslSflpur.rou pue eterlsrllnu ulur S€-01)goqs sapol1,{qd :3ur^\olloJeql epnlcurexul eseql ,{q peJtrqssJalJsrurlC ou ds UIISEIAI? ue,,lroJ S U Da u V PUE ',tou 'V ''ztrJd'if!tsptp ds ur1sey41ryuu,roJ S'U psaqo y'sercedseerql sest:dutoc,dnorg,, srql ,.dnorg lrslalp DIJDJV,,aq; 'l€elql lepun loN .trinrl uollDlJasuoJ 'suorl€AJ?sqoplerl pelcnpuoc,{llnleJ€c uro{ ,4Fulncqrud'Elup eJou qcmu 'st1r?ot! 'selcadb^ ',{JlllelJ go uorlrsrnbcueql qlrm fluo ,(11n;pootsrepun eq llr,\\ y r4l/(' BuolB srql 'ruJoJ IBJrd,{loql qll,'\ delJe^o 'sJe,rou 'Jell€us sr eJeqlsJelceJ"qJ eseql ur lnq Je,reJr.llr,r alllrl p eq ol luul.n?^aqt Jo spEeqaql loJ ,(cuapuale oslBsr oJoqJ 'tunuetunpurasuap E uroJ seuneruospue Sutpuerdso1 pessarddu uto:y ,{re,r sJrvqeql :eJnl"rueJE spod eql aurrleql,{q snorqulB,{11e:eua81nq Suuea.rog 1e ,furuq erv osle selcunped eqtr qltror8 s,ree,{ guerrnceqt puo,{eq ue,^.esetrlrlalUos 'slsrsred snul^lnd eqt uo puE stelqcu"Jq eql uo eJuecseqndeql Jo lsohl stelqruprq 3uno,{ aseq}uo euroq eJ€ sp€eqeql :ellq^\ Suturoceq ,{plcrnb f:e,r 1nq ueplo8 SurequortEqrur tE sJrEqeqt'q}4\oJB /heu snolnJeqnd-pesserddeo1 esoll,l flesuep slr ^q pezruSoreJsr lI lueserdlu pegqsnfun sr esJnoce qcns tBql elqBlrp^Berl suollcelloc /$eJos tnq uonruSocerpruro; errnbu feu 8ur,y e1e1 ;o ,{1rurcr,reqt ur uoDtlndod y uollDtroA 'sll13D4 'V Jepun^[nJ pessnasrpoJv saauateJJl( slelceJPqclu€lsuoc ^le^Il€lel ',reJ ,{ra^ ,{q ,(le)plg ? uepruhl sll!?Dtl V ulequou erour ,(lle.reue8eql luo4 sreJJrC saltlu$V 'JequsJo(I uI pelcelloc 'spouad uaoqoABq spees eJ$eru q1r,tr spod ::eqo1c6o1 lsn8ny ruorgs JFM}IC 8ulllntl puo SurJa oli SI truellersrt\rrrrry'urlstw )i gpuc uu^\o3 S-U 250 NuytsiaVol. 10,No. 2 (1995) A. ewartiana W. Fitzg., J. Bot.50: 19 0912). Lectotype:Cowcowing, W.A., August 1904,M. Koch 998 (BM,lde Maslinand Cowan, 1994a); isolecto:K,MEL (2 sheets,r?.v.), NSW, PERTH00751618. Paralectotlpe. Cowcowing, W.A., September 1904, M. Koch 998 (MEL - n.v., NSW, PERTH 00751596). Shrubs 0.5-2 m tall. Branchlets terete or slightly angled,glabrous, occasionally tomentulose or asericeous. Phyllodestercte, rclnded- or tluncate-obtuse,minutely t uncinate-mucronate,10-35 mm lor,g, 0.1-1.2 mm diarn., inclined to erect, straight to curved, glabrous, sometimesglaucous, with 14-18, distant, raised nerves. Peduncles2-4(7) mm long, 2 per axil, slender,glabrous; basal bracts persistent, cucullate, often glaucous; heads globular, 2.5-3.5 mm diam., dark-golden, aromatic, loosely (8)10-13(17) flowered. Flowers 5-merous;sepaLs 113-213-unrted, rurely free;petals ll2-3/4- rnited. Legume 'aised over seedsand constrictedto nanow isthmusbetween seeds, readily breaking into l-seededarticles at the constrictions,to 3 cm long, 1,2-1.8mm wide, thin-papery,straight to slightly curved, sparingly appressed-puberulous,articles fusiform, 4-5 mm long. Seeds ananged longitudinally in the pods, nanowly oblong-elliptio, 2.5-3 mm long, slightly glossy, mottled tan, exarillate. Selectedspecimens examined. WESTERN AUSTRALIA: c. 4 km W of Newdegateon road to Lake Grace,E.M. CanningW 41687407B(PERTH); Mou ntMdden, J. Goodwln224 (PERTH);9 km NNE of Bruce Rock on road to Meredin, B.R. Maslin 64'73(CANB, PERTH, Z); Murchison River, c. 15 km E of North West Coastal Highway, l0 November 1984, M. Whiting andJ. Coxon s.I.(PERTH 00684031). Distribution. Occuls in a belt fiom the Mufchison River (eastof Kalbarri) south to the Newdegate area (Newdegateis c. 50 krn eastof Lake Grace), southwestWestern Australia. Hqbitat. Grows on sandy, loamy and lateritic soils in open scrub and shrubland. Variation. There is considerablevariation in some of the characters,especially the vestitureof the branchletsand the degreeof union of perianthparts. In the north coastalpart of the range, many of the collectionsare fiom plantswith more or lessappressed-puberulous branchlets. Only a little farther inland the more common, glabrouscondition is found and it rangessouth to the limit of the species; thetypeofA. e\rartianuisof this variant.The typecollection of A, dielsii isfrom themiddle of the geographicrange and it has tomentulosebranchlets; one other collection at PERTH, labelled Diels and,Pritzel 157 and made in the sam€month and year at the samelocation (possibly an isotype or paratype)has exactly the samemorphology as the type. Affinities. ,\ very distinct speciesby virtue of the deeply constricted,easily fragmentedpods and exarillate seeds,characters that reliably separateit fiom its nearestrelative, A. nlvea R.S. Cowan & Maslin (seeunder A. niuea below for discussion). Conservationstatus. Not under threat. 2. Acacia nivea R.S. Cowan & Maslin. sp. nov. Frutex 40-100 cm altus, 60-170 cm expansus,ramulis teretibus,dense puberulis vel strigulosis, pilis appressisvel sub-ercctis,in resina inclusis, resina sicco saepealba. Stiprrlae plusminusve persistentes,minutae, triangulares,fuscatae, circa 0.5 rnm lorlgae. 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It is not surpdsingthat this specieshas been confused with A. dielsiiE. Pritz. (seeabove), for they are superficially very similar, the most diagnosticfeatures being the type of fruit. In A. d.ielsii the pods ale very thin and brittle, lessthan 2 nxn wide, not undulate,and breakreadily at the very deep constrictionsbetween the exarillate seeds. Sterile specimensofA. dielsii can be distinguishedfiom A. nivea by the latter's resinousbranchlets, with the resin commonly drying white, and the tendency for the phyllodesto havea more obvioustip. In flower the two speciescan be separatedby the normally united sepals and petals ol A. dietsii, as well as by its larger, cucullate basal peduncular bracts. Furthermore,in A. dleLriithe branchletsare commonly glabrous. FromA. obesaR.S. Cowan & Maslin (seebelow), anotherrelative, the new speciescan be distinguishedby its pedunculateheads, free petals and white-resinous branchlets. AIso superficially similar to A. pinguiculosa subsp. teretifolia R.S. Cowan & Maslin (ms name,in prep.) which hasthicker, 6-8-nerved phyllodes and biconvexpods. ConservationslatrJ. Not consideredto be threatened. Etymology. The white-resinousnature of the branchletsprovides the basisfor the name,fiom nlveas, Latin for snowyor snowywhitc. 3, Acacia obesaR.S. Cowan & Maslin,sp. nov. Frutex 0.3-O.6(l) rn itltus, ramulis teretibus,tomentulosis vel interdumappresso-puberulis. Stipulae percistentes,0.6-1.8 mm longae. PhytLodia teretia, rotundato-obtusaet saepe minute mucronulata,vulgo leviter ad valdecurvata sed laro aliquot recta interspersa, pulvino tomentuloso vel appresso-puberulo,laminis (10)t5 25 mm longis,1.2-1.'75 mm diametro,glabris, raro tomentulosis et glabrescantibus,I 2- I 6-nervatis,nervis elevatis, gJande minuta, basali. Capitula sessilia, globularia, diluto-lutea,3.5-4 nrrn diarr.retro,9-14-floribus, bracteolis unilateraliter peltatis. Florer 5-meri. Sepala 213-3/4-connata,zona marginali appresso-puberula,pilis rubris, minutissima. Legumina linearia,biconvexa, 20-40 rnrnlonga, 2-2.5 mrn lata,ooriacea, valde curuata, glabra vel ad basem puberulavel ubiquepuberula. Scrxlna longitudinalia, quadrato-rotundata, 1.5-1.8 mm longa,1.3-1.5 mm lata, I rnm crassitie,lucida, maoulata, arillo terminali,magno. Typus:21 miles133.6 krnl E of Hyden,Western Australia, 14 July 1970,B.R. MasLin 550 (holo: PERTH00158488; isc,: BRI, CANB, K, MEL,NY). Low, spreadingshrab,i 0.3-0.6(l) m tL\ll. Brqnchletsterete, tomentulose or occasionally appressed-puberulous(hails straight)with intermingledred or black micro-hairs. St,p&lespersistent, triangularto subu)ate,0.6-1.8mmlong, tomentulose.Ph!llodes terete, (10)15-25 mm long, 1.2-1.75 mm diam.,nornally thick,ascending, shallowly to stronglycurved but sometimes a few t staightones interspersed,glabrous, r'arely tomentulose and glabrescent;apex rctund-obtuse, often minutely mucronulate;pulvinus l-1.5 mm long, indumentumas on branchlets;nerves l2-16, raised;gland basal, not always evident, minute. Heads sessile,globular, pale golden, 3.5-4 mm diam., 9- l4-flowered; bracteolesunilaterally peltate, the stipe short, the bladeoblate-oyate,ciliolate. Flowers 5-merous. SepctLsll2-213 petal length, 213-3/4-tnited,lobes more or less triangular, acute to sub- obtuse,with a marginal band ofred, appressedmicro-hairs. Petals ftee or weakly coherentin lower half.. Ovary papillate-puberulous,appressed, red micro-hairssometimes also present. Pods linear, biconvex, 20-40 mm long, 2-2.5 mm wide, coriaceous,strongly curved, becoming once- or twice- coiled afterdehiscence, glabrous exaept puberulous basally or generallypuberulous. Seeds longitudinally arrangedin pods, quadrate-rotund,1.5 1.8 mm long, 1.3-1.5mm wide, I rnm thick, the lateral faces flattened,glossy, mottled dark-tanon very pale tan; pleurogramU-shaped; aril nearly as long as the seed. 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(1864). "Flora Ausaaliensis" vol. 2. (Irvell Reeve:l,ondon.) Briggs,J.D. &tligh, J.H.(1988). "RarE orThreatenedAustralianPlanb", AushalianNational Parks andWildlifeSewice, Special Publicationu4l. Court,A,B. (1973). /rr J.H.Willis. "A handbookto the Plants in Victoria"vol.2. ((Melboume Univercity Prcss onbehalfolThe Maud GibsonGardens Trusl Melboume) Cowan, R.S. & Ma.slin,B.R. (1990). Acacio Miscellany 3. Somenew miffoneurous taxa of WestemAustalia rclated io A. multilineata(l3gtrninosae:Mmosoideae: section Pltrlin?rve.t). Nuytsi a.l:209-219. Cowan,R.S. & Maslin,B.R. (1995), Acacja Miscellany 10. New taxa and notes on previously described taxa of,Acoci4, mosdy sectionJuliflorae(LEgnminosae: Mimosoideae), in WestemAustsalia. Nuytsia l0i 15-62, Greuter,W. ?tdl. ( 1994).Intemational Code ofBotanical Nomenclaturc; Regnum Vegetabile I 3 1. IntemationalAssociation for PlantTaxonomy@urop€). Maslin,B,R. (1978a). Studies in thegenus, cacla (Mimosaceae) -7. Thetaxonomyofsome diaphyllodinous species. Nuytsia2i 200-219. Maslin,B.R. (1981). Acacl4. 1n Jqssop, J, (ed.), "Flora of CentalAustralia." (A.H. and A.W. Reed: Sydney.) Maslin,B.R. & Cowan,R.S. (1994). C.R,P. Andrews' sp€cies ofAcdcid (lrguminosae: Mimosoideae): typification ofthe narnes. Nuytsia9:383-385. Maslin,B.R. &Cowan, R.S. (1994a). WilliamVincentFitzgerald's species ofAcdcia(lrguminosae:Mimosoideao); typification ofthenames.Nuytsia 91 38?-398. Maslin,B.R. & Cowan,R,S, (1994b). What Type OfType? Austral, Syst. Bot, Soc. Newsletter 81r 2-7, Maslin,B.R. & Pedley,L. (1988).Patterns ofdistribution ofAcacia in Austalia, Austual.J. Bot.361 385-393, Whibley,D.J.E. (1980), "Acacias ofSoulh Australia" (South Austalian GovemmentPrinter: Adelaide,) Whibley,D.J.E, & Synon, D.B. (1992).Acacias ofSouth Australia. (South Australian Covemment Printer: Adelaide.)