California Terrestrial and Vernal Pool Invertebrates of Conservation Priority1
Species Federal ESA CA ESA G-Rank S-Rank Other Statuses GASTROPODA (Snails and slugs) Ammonitella yatesii None None G1 S1 IUCN - Vulnerable tight coin (=Yates' snail) Ancotrema voyanum None None G1G2 S1S2 BLM-Sensitive hooded lancetooth Assiminea infima None None G1 S1 IUCN - Vulnerable Badwater snail Binneya notabilis None None G1 S1 IUCN - Data Deficient Santa Barbara shelled slug Eremarionta immaculata None None G1 S1 IUCN - Vulnerable white desertsnail Eremarionta millepalmarum None None G1 S1 IUCN - Vulnerable Thousand Palms desertsnail Eremarionta morongoana IUCN - Near None None G1G3 S1 Morongo (=Colorado) desertsnail Threatened Eremarionta rowelli bakerensis None None G3G4T1 S1 IUCN - Data Deficient Baker's desertsnail Eremarionta rowelli mccoiana None None G3G4T1 S1 IUCN - Data Deficient California Mccoy snail Haplotrema catalinense None None G1 S1 Santa Catalina lancetooth Haplotrema duranti None None G1G2 S1S2 ribbed lancetooth Helminthoglypta allynsmithi None None G1 S1 IUCN - Vulnerable Merced Canyon shoulderband Helminthoglypta arrosa monticola None None G2G3T1 S1 mountain shoulderband Helminthoglypta arrosa pomoensis None None G2G3T1 S1 IUCN - Data Deficient Pomo bronze shoulderband Helminthoglypta ayresiana sanctaecrucis None None G1G2T1T2 S1S2 Ayer's snail Helminthoglypta callistoderma None None G1 S1 IUCN - Endangered Kern shoulderband
1 This list pertains to Scientific Collecting Permit regulations (Title 14, Section 650, California Code of Regulations). The species on this list include vernal pool, terrestrial, and aquatic invertebrates with a terrestrial phase that are listed under the Federal and California Endangered Species Acts, or have a NatureServe rank of G1, G1G2, S1, S1S2, SH or SX. For a full list of invertebrates of conservation concern, see the California Natural Diversity Database Special Animals List: https://www.wildlife.ca.gov/Data/CNDDB/Plants-and-Animals.
List generated on June 12, 2017 from a query of the California Natural Diversity Database.
California Department of Fish and Wildlife Page 1 of 18 California Terrestrial and Vernal Pool Invertebrates of Conservation Priority
Species Federal ESA CA ESA G-Rank S-Rank Other Statuses Helminthoglypta coelata None None G1 S1 IUCN - Vulnerable mesa shoulderband Helminthoglypta concolor None None G1G2 S1S2 whitefir shoulderband Helminthoglypta fontiphila None None G1 S1 Soledad shoulderband Helminthoglypta greggi None None G1 S1 Mohave shoulderband Helminthoglypta hertleini None None G1 S1S2 BLM-Sensitive Oregon shoulderband Helminthoglypta milleri None None G1 S1 peak shoulderband Helminthoglypta mohaveana IUCN - Near None None G1 S1 Victorville shoulderband Threatened Helminthoglypta nickliniana awania None None G3T1 S1 IUCN - Data Deficient Peninsula coast range shoulderband Helminthoglypta nickliniana bridgesi None None G3T1 S1S2 IUCN - Data Deficient Bridges' coast range shoulderband Helminthoglypta sequoicola consors None None G2T1 S1 IUCN - Data Deficient redwood shoulderband Helminthoglypta stiversiana williamsi None None G2G3T1 S1 IUCN - Data Deficient Williams' bronze shoulderband Helminthoglypta taylori None None G1 S1 westfork shoulderband Helminthoglypta traskii pacoimensis None None G1G2T1 S1 Pacoima shoulderband Helminthoglypta traskii traskii None None G1G2T1 S1 Trask shoulderband Helminthoglypta uvasana None None G1 S1 Grapevine shoulderband Helminthoglypta vasquezi None None G1 S1 Vasquez shoulderband Helminthoglypta walkeriana IUCN - Critically Morro shoulderband (=banded dune) Endangered None G1 S1S2 Endangered snail Herpeteros angelus None None G1 S1 Soledad desertsnail Hesperarion plumbeus None None G1 S1 leaden slug Megomphix californicus None None G1G2 S1S2 Natural Bridge megomphix Micrarionta facta None None G1G2 S1S2 IUCN - Vulnerable Santa Barbara islandsnail Micrarionta feralis IUCN - Critically None None G1 S1 San Nicolas islandsnail Endangered Micrarionta gabbi None None G1 S1 IUCN - Vulnerable San Clemente islandsnail
California Department of Fish and Wildlife Page 2 of 18 California Terrestrial and Vernal Pool Invertebrates of Conservation Priority
Species Federal ESA CA ESA G-Rank S-Rank Other Statuses Micrarionta opuntia None None G1 S1 IUCN - Vulnerable pricklypear islandsnail Monadenia callipeplus None None G1G2 S1S2 downy sideband Monadenia circumcarinata BLM-Sensitive | IUCN None None G1 S1 keeled sideband - Vulnerable Monadenia cristulata None None G1G2 S1S2 crested sideband Monadenia fidelis leonina None None G4G5T1T2 S1S2 A terrestrial snail Monadenia fidelis pronotis None None G4G5T1 S1 rocky coast Pacific sideband Monadenia infumata ochromphalus None None G2T1 S1 yellow-based sideband Monadenia infumata setosa None Threatened G2T2 S2 IUCN - Vulnerable Trinity bristle snail Monadenia marmarotis None None G1 S1 marble sideband Monadenia mormonum buttoni None None G2T1 S1S2 Button's Sierra sideband Monadenia mormonum hirsuta None None G2T1 S1 BLM-Sensitive hirsute Sierra sideband Monadenia troglodytes troglodytes IUCN - Data Deficient | None None G1G2T1T2 S1S2 Shasta sideband USFS - Sensitive Monadenia troglodytes wintu IUCN - Data Deficient | None None G1G2T1T2 S1S2 Wintu sideband USFS - Sensitive Monadenia tuolumneana None None G1 S1 BLM-Sensitive Tuolumne sideband Monadenia yosemitensis None None G1 S1S2 Yosemite Mariposa sideband Pristiloma shepardae None None G1 S1 Shepard's snail Prophysaon coeruleum None None G3G4 S1S2 Blue-gray taildropper slug Punctum hannai None None G1G2 S1S2 Trinity Spot Radiocentrum avalonense IUCN - Critically None None G1 S1 Catalina mountainsnail Endangered Rothelix warnerfontis None None G1 S1 USFS - Sensitive Warner Springs shoulderband Sterkia clementina IUCN - Near None None G1 S1S2 San Clemente Island blunt-top snail Threatened Trilobopsis roperi None None G1 S1 USFS - Sensitive Shasta chaparral Trilobopsis tehamana BLM-Sensitive | USFS None None G1 S1 Tehama chaparral - Sensitive
California Department of Fish and Wildlife Page 3 of 18 California Terrestrial and Vernal Pool Invertebrates of Conservation Priority
Species Federal ESA CA ESA G-Rank S-Rank Other Statuses Vespericola pressleyi BLM-Sensitive | USFS None None G1 S1 Big Bar hesperian - Sensitive Vespericola scotti None None G1 S1 Benson Gulch hesperian Vespericola shasta None None G1 S1 USFS - Sensitive Shasta hesperian Vespericola sierranus None None G2 S1S2 Siskiyou hesperian Xerarionta intercisa None None G1 S1 IUCN - Vulnerable horseshoe snail Xerarionta redimita None None G1G2 S1 IUCN - Vulnerable wreathed cactussnail Xerarionta tryoni None None G1 S1 IUCN - Vulnerable Bicolor cactussnail ARACHNIDA (Spiders and relatives) Aphrastochthonius grubbsi None None G1G2 S1S2 Grubbs' Cave pseudoscorpion Aphrastochthonius similis None None G1G2 S1S2 Carlow's Cave pseudoscorpion Archeolarca aalbui None None G1G2 S1S2 Aalbu's Cave pseudoscorpion Banksula californica None None GH SH Alabaster Cave harvestman Banksula galilei None None G1 S1 Galile's cave harvestman Banksula grubbsi None None G1 S1 Grubbs' cave harvestman Banksula incredula None None G1 S1 incredible harvestman Banksula martinorum None None G1 S1 Martins' cave harvestman Banksula melones None None G1 S1 IUCN - Vulnerable Melones Cave harvestman Banksula rudolphi None None G1 S1 Rudolph's cave harvestman Banksula tuolumne None None G1 S1 Tuolumne cave harvestman Banksula tutankhamen None None G1 S1 King Tut Cave harvestman Calicina arida None None G1 S1 San Benito harvestman Calicina breva None None G1 S1 Stanislaus harvestman Calicina cloughensis None None G1 S1 Clough Cave harvestman
California Department of Fish and Wildlife Page 4 of 18 California Terrestrial and Vernal Pool Invertebrates of Conservation Priority
Species Federal ESA CA ESA G-Rank S-Rank Other Statuses Calicina conifera None None G1 S1 Crane Flat harvestman Calicina diminua None None G1 S1 Marin blind harvestman Calicina dimorphica None None G1 S1 Watts Valley harvestman Calicina macula None None G1 S1 marbled harvestman Calicina mesaensis None None G1 S1 Table Mountain harvestman Calicina minor None None G1 S1 Edgewood blind harvestman Calicina piedra None None G1 S1 Piedra harvestman Calileptoneta briggsi None None G1 S1 Briggs' leptonetid spider Calileptoneta oasa None None G1 S1 Andreas Canyon leptonetid spider Calileptoneta ubicki None None G1 S1 Ubick's leptonetid spider Calileptoneta wapiti None None G1 S1 Mendocino leptonetid spider Fissilicreagris imperialis None None G1 S1 IUCN - Vulnerable Empire Cave pseudoscorpion Hubbardia idria None None G1 S1 Idria short-tailed whipscorpion Hubbardia secoensis None None G1 S1 Arroyo Seco short-tailed whipscorpion Hubbardia shoshonensis None None G1 S1 BLM-Sensitive Shoshone Cave whip-scorpion Larca laceyi None None G1G2 S1 Lacey's Cave pseudoscorpion Meta dolloff None None G1 S1 IUCN - Vulnerable Dolloff Cave spider Microcina edgewoodensis None None G1 S1 Edgewood Park micro-blind harvestman Microcina homi None None G1 S1 Hom's micro-blind harvestman Microcina jungi None None G1 S1 Jung's micro-blind harvestman Microcina leei None None G1 S1 Lee's micro-blind harvestman Microcina lumi None None G1 S1 Lum's micro-blind harvestman Microcina tiburona None None G1 S1 Tiburon micro-blind harvestman
California Department of Fish and Wildlife Page 5 of 18 California Terrestrial and Vernal Pool Invertebrates of Conservation Priority
Species Federal ESA CA ESA G-Rank S-Rank Other Statuses Neochthonius imperialis None None G1 S1 Empire Cave pseudoscorpion Pseudogarypus orpheus None None G1G2 S1 Music Hall Cave pseudoscorpion Socalchemmis gertschi None None G1 S1 Gertsch's socalchemmis spider Socalchemmis icenoglei None None G1 S1 Icenogle's socalchemmis spider Socalchemmis monterey None None G1 S1 Monterey socalchemmis spider Talanites moodyae None None G1G2 S1S2 Moody's gnaphosid spider Talanites ubicki None None G1 S1 Ubick's gnaphosid spider Telema sp. None None G1G2 S1S2 Santa Cruz telemid spider Texella deserticola None None G1 S1 Whitewater Canyon harvestman Texella kokoweef None None G1 S1 Kokoweef Crystal Cave harvestman Texella shoshone None None G1 S1 Shoshone Cave harvestman CRUSTACEA, Order Anostraca (fairy shrimp) Branchinecta campestris None None G2 S1 pocket pouch fairy shrimp Branchinecta conservatio Endangered None G2 S2 IUCN - Endangered Conservancy fairy shrimp Branchinecta longiantenna Endangered None G1 S1S2 IUCN - Endangered longhorn fairy shrimp Branchinecta lynchi Threatened None G3 S3 IUCN - Vulnerable vernal pool fairy shrimp Branchinecta sandiegonensis Endangered None G2 S2 IUCN - Endangered San Diego fairy shrimp Linderiella santarosae None None G1G2 S1 Santa Rosa Plateau fairy shrimp Streptocephalus woottoni Endangered None G1G2 S1S2 IUCN - Endangered Riverside fairy shrimp CRUSTACEA, Order Notostraca (tadpole shrimp) Lepidurus packardi Endangered None G4 S3S4 IUCN - Endangered vernal pool tadpole shrimp CRUSTACEA, Order Anomopoda (water fleas) Dumontia oregonensis None None G1G3 S1 hairy water flea INSECTA, Order Plecoptera (stoneflies) Capnia lacustra None None G1 S1 Lake Tahoe benthic stonefly
California Department of Fish and Wildlife Page 6 of 18 California Terrestrial and Vernal Pool Invertebrates of Conservation Priority
Species Federal ESA CA ESA G-Rank S-Rank Other Statuses INSECTA, Order Orthoptera (grasshoppers, katydids, and crickets) Aglaothorax longipennis IUCN - Critically None None G1G2 S1S2 Santa Monica shieldback katydid Endangered Ammopelmatus kelsoensis None None G1G2 S1S2 IUCN - Vulnerable Kelso jerusalem cricket Ammopelmatus muwu None None G1 S1 IUCN - Vulnerable Point Conception jerusalem cricket Idiostatus kathleenae None None G1G2 S1S2 Pinnacles shieldback katydid Idiostatus middlekauffi IUCN - Critically None None G1G2 S1 Middlekauff's shieldback katydid Endangered Macrobaenetes algodonensis None None G1G2 S1S2 Algodones sand treader cricket Macrobaenetes valgum None None G1G2 S1S2 IUCN - Vulnerable Coachella giant sand treader cricket Psychomastax deserticola None None G1G2 S1S2 IUCN - Vulnerable desert monkey grasshopper Stenopelmatus cahuilaensis None None G1G2 S1S2 IUCN - Vulnerable Coachella Valley jerusalem cricket Tetrix sierrana None None G1G2 S1S2 IUCN - Vulnerable Sierra pygmy grasshopper Trimerotropis infantilis Endangered None G1 S1 IUCN - Endangered Zayante band-winged grasshopper Trimerotropis occidentiloides None None G1G2 S1S2 IUCN - Endangered Santa Monica grasshopper Trimerotropis occulens None None G1G2 S1S2 IUCN - Endangered Lompoc grasshopper INSECTA, Order HeteropteraHemiptera (true bugs) Oravelia pege None None G1 S1 Dry Creek cliff strider bug Saldula usingeri None None G1 S1 Wilbur Springs shorebug INSECTA, Order Coleoptera (beetles) Aegialia concinna BLM-Sensitive | IUCN None None G1 S1 Ciervo aegilian scarab beetle - Vulnerable Agrilus harenus None None G1G2 S1S2 Harenus jewel beetle Anomala carlsoni None None G1 S1 Carlson's dune beetle Anomala hardyorum None None G1 S1 Hardy's dune beetle Anthicus antiochensis None None G1 S1 Antioch Dunes anthicid beetle Anthicus sacramento None None G1 S1 IUCN - Endangered Sacramento anthicid beetle
California Department of Fish and Wildlife Page 7 of 18 California Terrestrial and Vernal Pool Invertebrates of Conservation Priority
Species Federal ESA CA ESA G-Rank S-Rank Other Statuses Cicindela gabbii None None G2G4 S1 western tidal-flat tiger beetle Cicindela hirticollis abrupta None None G5TH SH Sacramento Valley tiger beetle Cicindela latesignata latesignata None None G2G4T1T2 S1 western beach tiger beetle Cicindela ohlone Endangered None G1 S1 Ohlone tiger beetle Cicindela senilis frosti None None G2G3T1T3 S1 senile tiger beetle Cicindela tranquebarica ssp. None None G5T1 S1 San Joaquin tiger beetle Cicindela tranquebarica viridissima None None G5T1 S1 greenest tiger beetle Coelus globosus None None G1G2 S1S2 IUCN - Vulnerable globose dune beetle Coelus gracilis BLM-Sensitive | IUCN None None G1 S1 San Joaquin dune beetle - Vulnerable Coenonycha clementina None None G1G2 S1S2 San Clemente Island coenonycha beetle Cyclocephala wandae None None G1G2 S1S2 Wandae dune beetle Deltaspis ivae None None G1 S1 marsh-elder long-horned beetle Desmocerus californicus dimorphus Threatened None G3T2 S2 valley elderberry longhorn beetle Dinacoma caseyi Endangered None G1 S1 Casey's June beetle Elaphrus viridis IUCN - Critically Threatened None G1 S1 Delta green ground beetle Endangered Juniperella mirabilis None None G1 S1 juniper metallic wood-boring beetle Lepismadora algodones None None G1 S1 Algodones sand jewel beetle Lichnanthe albipilosa None None G1 S1 white sand bear scarab beetle Lytta hoppingi None None G1G2 S1S2 Hopping's blister beetle Lytta insperata None None G1G2 S1S2 Mojave Desert blister beetle Lytta morrisoni None None G1G2 S1S2 Morrison's blister beetle Microcylloepus formicoideus None None G1 S1 Furnace Creek riffle beetle Nebria darlingtoni None None G1 S1 South Forks ground beetle
California Department of Fish and Wildlife Page 8 of 18 California Terrestrial and Vernal Pool Invertebrates of Conservation Priority
Species Federal ESA CA ESA G-Rank S-Rank Other Statuses Nebria gebleri siskiyouensis None None G4G5T4 S1S2 Siskiyou ground beetle Nebria sahlbergii triad None None G1T1 S1 Trinity Alps ground beetle Onychobaris langei None None G1 S1 Lange's El Segundo Dune weevil Polyphylla anteronivea None None G1 S1 Saline Valley snow-front June beetle Polyphylla barbata Endangered None G1 S1 Mount Hermon (=barbate) June beetle Polyphylla erratica None None G1G2 S1S2 Death Valley June beetle Polyphylla nubila None None G1 S1 Atascadero June beetle Prasinalia imperialis None None G1G2 S1S2 Algodones white wax jewel beetle Pseudocotalpa andrewsi None None G1 S1 Andrew's dune scarab beetle Trachykele hartmani None None G1 S1 serpentine cypress wood-boring beetle Trichinorhipis knulli None None G1 S1 Knull's metallic wood-boring beetle Trigonoscuta brunnotesselata None None G1G2 S1S2 brown tassel trigonoscuta weevil Trigonoscuta dorothea dorothea None None G1T1 S1 Dorothy's El Segundo Dune weevil Trigonoscuta rothi algodones None None G1G2 S1S2 Algodones dune weevil Trigonoscuta rothi imperialis None None G1G2 S1S2 Imperial dune weevil Trigonoscuta rothi punctata None None G1G2 S1S2 Punctate dune weevil Trigonoscuta rothi rothi None None G1G2 S1S2 Roth's dune weevil Trigonoscuta sp. None None G1Q S1 Doyen's trigonoscuta dune weevil Trigonoscuta stantoni None None G1 S1 Santa Cruz Island shore weevil Vandykea tuberculata None None G1 S1 serpentine cypress long-horned beetle INSECTA, Order Mecoptera (scorpionflies) Orobittacus obscurus None None G1 S1 gold rush hanging scorpionfly INSECTA, Order Diptera (flies) Ablautus schlingeri None None G1 S1 Oso Flaco robber fly
California Department of Fish and Wildlife Page 9 of 18 California Terrestrial and Vernal Pool Invertebrates of Conservation Priority
Species Federal ESA CA ESA G-Rank S-Rank Other Statuses Andrena subapasta None None G1G2 S1S2 an andrenid bee Apiocera warneri None None G1G2 S1S2 Glamis sand fly Argochrysis lassenae None None G1 S1 Lassen cuckoo wasp Bombus caliginosus None None G4? S1S2 IUCN - Vulnerable obscure bumble bee Bombus crotchii None None G3G4 S1S2 Crotch bumble bee IUCN - Critically Bombus franklini Endangered | None None G1 S1 Franklin's bumble bee XERCES - Critically Imperiled Bombus morrisoni None None G4G5 S1S2 IUCN - Vulnerable Morrison bumble bee Bombus occidentalis USFS - Sensitive | None None G2G3 S1 western bumble bee XERCES - Imperiled Bombus suckleyi None None GU S1 Suckley's cuckoo bumble bee Brennania belkini None None G1G2 S1S2 IUCN - Vulnerable Belkin's dune tabanid fly Ceratochrysis bradleyi None None G1 S1 Bradley's cuckoo wasp Ceratochrysis gracilis None None G1 S1 Piute Mountains cuckoo wasp Ceratochrysis longimala None None G1 S1 Desert cuckoo wasp Ceratochrysis menkei None None G1 S1 Menke's cuckoo wasp Chrysis tularensis None None G1G2 S1S2 Tulare cuckoo wasp Cleptes humboldti None None G1G2 S1S2 Humboldt cuckoo wasp Dufourea stagei None None G1 S1 Stage's dufourine bee Efferia antiochi None None G1G2 S1S2 Antioch efferian robberfly Efferia macroxipha None None G1G2 S1S2 Glamis robberfly Eucerceris ruficeps redheaded sphecid wasp None None G1G3G1G2 S1S2 Metapogon hurdi Hurd's metapogon robberfly Euparagia unidentata Algodones euparagiaParacoenia calida None None G1G2G1 S1S2S1 Wilbur Springs shore fly
California Department of Fish and Wildlife Page 10 of 18 California Terrestrial and Vernal Pool Invertebrates of Conservation Priority
Species Federal ESA CA ESA G-Rank S-Rank Other Statuses Habropoda pallida white faced beeRhaphiomidas None None G1G2G1T1 S1S2S1 terminatus abdominalis Endangered Delhi Sands flower-loving fly Hedychridium argenteum Riverside cuckoo waspRhaphiomidas None None G1G2G1T1 S1S2S1 terminatus terminatus El Segundo flower-loving fly Hedychridium milleri Borax Lake cuckoo waspRhaphiomidas None None G1 S1 trochilus Valley mydas fly INSECTA, Order Lepidoptera (butterflies & moths) Areniscythris brachypteris None None G1 S1 Oso Flaco flightless moth Eucosma hennei None None G1 S1 Henne's eucosman moth Carolella busckana None None G1G3 SH Busck's gallmoth Pyrgus ruralis lagunae XERCES - Critically Endangered None G5T1 S1 Laguna Mountains skipper Imperiled Carterocephalus palaemon magnus None None G5T5 S1 Sonoma arctic skipper Pseudocopaeodes eunus obscurus XERCES - Critically Endangered None G3G4T1 S1 Carson wandering skipper Imperiled Hesperia miriamae longaevicola None None G2G3T1 S1 White Mountains skipper Polites mardon USFS - Sensitive | None None G2G3 S1 mardon skipper XERCES - Imperiled Euphyes vestris harbisoni None None G5T1T2 S1S2 dun skipper Euchloe hyantis andrewsi None None G3G4T1 S1 Andrew's marble butterfly Lycaena rubidus incana None None G5T1 S1 White Mountains copper Lycaena hermes IUCN - Vulnerable | Candidate None G1 S1 Hermes copper butterfly USFS - Sensitive Callophrys comstocki None None G3G4 S1S2 XERCES - Imperiled desert green hairstreak Callophrys thornei None None G1 S1 BLM-Sensitive Thorne's hairstreak Callophrys mossii bayensis XERCES - Critically Endangered None G4T1 S1 San Bruno elfin butterfly Imperiled Callophrys mossii hidakupa None None G4T1T2 S1S2 USFS - Sensitive San Gabriel Mountains elfin butterfly Callophrys mossii marinensis None None G4T1 S1 Marin elfin butterfly
California Department of Fish and Wildlife Page 11 of 18 California Terrestrial and Vernal Pool Invertebrates of Conservation Priority
Species Federal ESA CA ESA G-Rank S-Rank Other Statuses Euphilotes battoides allyni XERCES - Critically Endangered None G5T1 S1 El Segundo blue butterfly Imperiled Euphilotes enoptes smithi XERCES - Critically Endangered None G5T1T2 S1S2 Smith's blue butterfly Imperiled Euphilotes mojave None None G2G3 S1S2 XERCES - Imperiled Mojave dotted-blue Euphilotes baueri USFS - Sensitive | None None G2G4 S1S2 Bauer's dotted-blue XERCES - Imperiled Philotiella speciosa bohartorum None None G3G4T1 S1 Boharts' blue butterfly Glaucopsyche lygdamus palosverdesensis XERCES - Critically Endangered None G5T1 S1 Palos Verdes blue butterfly Imperiled Plebejus idas lotis XERCES - Critically Endangered None G5TH SH lotis blue butterfly Imperiled Plebejus saepiolus aureolus None None G5T1 S1 USFS - Sensitive San Gabriel Mountains blue butterfly Plebejus saepiolus albomontanus None None G5T2 S1S2 White Mountains saepiolus blue butterfly Plebulina emigdionis None None G1G2 S1S2 USFS - Sensitive San Emigdio blue butterfly Plebejus icarioides missionensis XERCES - Critically Endangered None G5T1 S1 Mission blue butterfly Imperiled Plebejus icarioides parapheres None None G5T1T2 S1S2 Point Reyes blue butterfly Apodemia mormo langei XERCES - Critically Endangered None G5T1 S1 Lange's metalmark butterfly Imperiled Speyeria nokomis carsonensis None None G3T1 S1 Carson Valley silverspot Speyeria zerene sonomensis None None G5T1 S1 Sonoma zerene fritillary Speyeria zerene hippolyta XERCES - Critically Threatened None G5T1 S1 Oregon silverspot butterfly Imperiled Speyeria zerene behrensii XERCES - Critically Endangered None G5T1 S1 Behren's silverspot butterfly Imperiled Speyeria zerene myrtleae XERCES - Critically Endangered None G5T1 S1 Myrtle's silverspot butterfly Imperiled Speyeria callippe callippe XERCES - Critically Endangered None G5T1 S1 callippe silverspot butterfly Imperiled Speyeria adiaste adiaste None None G1G2T1 S1 unsilvered fritillary Chlosyne leanira elegans None None G4G5T1T2 S1S2 Oso Flaco patch butterfly Euphydryas editha bayensis XERCES - Critically Threatened None G5T1 S1 Bay checkerspot butterfly Imperiled Euphydryas editha monoensis None None G5T2T3 S1S2 USFS - Sensitive Mono checkerspot butterfly
California Department of Fish and Wildlife Page 12 of 18 California Terrestrial and Vernal Pool Invertebrates of Conservation Priority
Species Federal ESA CA ESA G-Rank S-Rank Other Statuses Euphydryas editha quino XERCES - Critically Endangered None G5T1T2 S1S2 quino checkerspot butterfly Imperiled Coenonympha tullia yontockett None None G5T1T2 S1 Yontocket satyr Cercyonis pegala carsonensis None None G5T1T2 S1S2 Carson Valley wood nymph Danaus plexippus None None G4T2T3 S2S3 USFS - Sensitive monarch Hesperopsis gracielae None None G2G3 S1S2 XERCES - Vulnerable Macneill's sootywing Euproserpinus euterpe XERCES - Critically Threatened None G1G2 S1 Kern primrose sphinx moth Imperiled INSECTA, Order Trichoptera (caddisflies) Cryptochia denningi None None G1G2 S1S2 Denning's cryptic caddisfly Cryptochia excella None None G1G2 S1S2 Kings Canyon cryptochian caddisfly Cryptochia shasta None None G1G2 S1S2 confusion caddisfly Diplectrona californica None None G1G2 S1S2 California diplectronan caddisfly Ecclisomyia bilera None None G1G2 S1S2 Kings Creek ecclysomyian caddisfly Farula praelonga None None G1G2 S1S2 long-tailed caddisfly Goeracea oregona None None G3 S1S2 Sagehen Creek goeracean caddisfly Lepidostoma ermanae None None G1G2 S1S2 Cold Spring caddisfly Limnephilus atercus None None G3G4 S1 Fort Dick limnephilus caddisfly Neothremma genella None None G1G2 S1S2 golden-horned caddisfly Neothremma siskiyou None None G1G2 S1S2 Siskiyou caddisfly Parapsyche extensa None None GH SH King's Creek parapsyche caddisfly Rhyacophila lineata None None G1G3 S1S2 Castle Crags rhyacophilan caddisfly Rhyacophila mosana None None G1G2Q S1S2 bilobed rhyacophilan caddisfly Rhyacophila spinata None None G1G2 S1S2 spiny rhyacophilan caddisfly INSECTA, Order Hymenoptera (ants, bees, & wasps) Andrena subapasta None None G1G2 S1S2 an andrenid bee
California Department of Fish and Wildlife Page 13 of 18 California Terrestrial and Vernal Pool Invertebrates of Conservation Priority
Species Federal ESA CA ESA G-Rank S-Rank Other Statuses Argochrysis lassenae None None G1 S1 Lassen cuckoo wasp Bombus caliginosus None None G4? S1S2 IUCN - Vulnerable obscure bumble bee Bombus crotchii None None G3G4 S1S2 Crotch bumble bee IUCN - Critically Bombus franklini Endangered | None None G1 S1 Franklin's bumble bee XERCES - Critically Imperiled Bombus morrisoni None None G4G5 S1S2 IUCN - Vulnerable Morrison bumble bee Bombus occidentalis USFS - Sensitive | None None G2G3 S1 western bumble bee XERCES - Imperiled Bombus suckleyi None None GU S1 Suckley's cuckoo bumble bee Ceratochrysis bradleyi None None G1 S1 Bradley's cuckoo wasp Ceratochrysis gracilis None None G1 S1 Piute Mountains cuckoo wasp Ceratochrysis longimala None None G1 S1 Desert cuckoo wasp Ceratochrysis menkei None None G1 S1 Menke's cuckoo wasp Chrysis tularensis None None G1G2 S1S2 Tulare cuckoo wasp Cleptes humboldti None None G1G2 S1S2 Humboldt cuckoo wasp Dufourea stagei None None G1 S1 Stage's dufourine bee Eucerceris ruficeps None None G1G3 S1S2 redheaded sphecid wasp Euparagia unidentata None None G1G2 S1S2 Algodones euparagia Habropoda pallida None None G1G2 S1S2 white faced bee Hedychridium argenteum None None G1G2 S1S2 Riverside cuckoo wasp Hedychridium milleri None None G1 S1 Borax Lake cuckoo wasp Lasioglossum channelense None None G1 S1 Channel Island sweat bee Metapogon hurdi None None G1G2 S1S2 Hurd's metapogon robberfly Microbembex elegans None None G1G2 S1S2 Algodones elegant sand wasp
California Department of Fish and Wildlife Page 14 of 18 California Terrestrial and Vernal Pool Invertebrates of Conservation Priority
Species Federal ESA CA ESA G-Rank S-Rank Other Statuses Myrmosula pacifica None None GH SH Antioch multilid wasp Neolarra alba None None GH SH white cuckoo bee Oravelia pege None None G1 S1 Dry Creek cliff strider bug Paracoenia calida None None G1 S1 Wilbur Springs shore fly Paranomada californica None None G1 S1 California cuckoo bee Parnopes borregoensis None None G1G2 S1S2 Borrego parnopes cuckoo wasp Perdita algodones None None G1G2 S1S2 Algodones perdita Perdita frontalis None None G1G2 S1S2 Imperial Perdita Perdita glamis None None G1G2 S1S2 Glamis perdita Perdita scitula antiochensis None None G1T1 S1 Antioch andrenid bee Philanthus nasalis None None G1 S1 Antioch specid wasp Protodufourea wasbaueri XERCES_DD-Data None None G1 S1 Wasbauer's protodufourea bee Deficient Protodufourea zavortinki None None G1 S1 Zavortink's protodufourea bee Rhaphiomidas terminatus abdominalis Endangered None G1T1 S1 Delhi Sands flower-loving fly Rhaphiomidas terminatus terminatus None None G1T1 S1 El Segundo flower-loving fly Rhaphiomidas trochilus None None G1 S1 Valley mydas fly Rhopalolemma robertsi None None G1 S1 Roberts' rhopalolemma bee Sedomaya glamisensis None None G1G2 S1S2 Glamis night tiphiid Sphaeropthalma ecarinata None None G1G2 S1S2 Glamis night mutillid Sphecodogastra antiochensis XERCES - Critically None None G1 S1 Antioch Dunes halcitid bee Imperiled Stictiella villegasi None None G1G2 S1S2 Algodones sand wasp Trachusa gummifera None None G1 S1 San Francisco Bay Area leaf-cutter bee
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NatureServe Element Ranking
All Heritage Programs, such as the California Natural Diversity Database (CNDDB) use the same ranking methodology, originally developed by The Nature Conservancy and now maintained and recently revised by NatureServe. It includes a Global rank (G-rank), describing the rank for a given taxon over its entire distribution and a State rank (S-rank), describing the rank for the taxon over its state distribution. For subspecies and varieties, there is also a “T” rank describing the global rank for the infraspecific taxon. The next page of this document details the criteria used to assign element ranks, from G1 to G5 for the Global rank and from S1 to S5 for the State rank. Procedurally, state programs such as the CNDDB develop the State ranks. The Global ranks are determined collaboratively among the Heritage Programs for the states/provinces containing the species. NatureServe then checks for consistency and logical errors at the national level. Because the units of conservation may include non-taxonomic biological entities such as populations or ecological communities, NatureServe refers to the targets of biological conservation as elements rather than taxa.
An element rank is assigned using standard criteria and rank definitions. This standardization makes the ranks comparable between organisms and across political boundaries. NatureServe has developed a “rank calculator” to help increase repeatability and transparency of the ranking process. The three main categories that are taken into consideration when assigning an element rank are rarity, threats, and trends. Within these three categories, various factors are considered including:
Range extent, area of occupancy, population size, total number of occurrences, environmental specificity and number of good occurrences (ranked A or B).
Overall threat impact as well as intrinsic vulnerability (if threats are unknown).
Long-term and short-term trends.
Detailed information on the newest element ranking methodology can be found here: https://connect.natureserve.org/publications/StatusAssess_Methodology
With the above considerations in mind, refer below for the numerical definitions for G1-5 and S1-5. An element’s ranking status may be adjusted up or down depending upon the considerations above.
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Element Ranking
GLOBAL RANKING The global rank (G-rank) is a reflection of the overall status of an element throughout its global range. Both Global and State ranks represent a letter and number score that reflects a combination of Rarity, Threat, and Trend factors, with weighting being heavier on Rarity than the other two.
SPECIES OR NATURAL COMMUNITY LEVEL G1 = Critically Imperiled—At very high risk of extinction due to extreme rarity (often 5 or fewer populations), very steep declines, or other factors. G2 = Imperiled—At high risk of extinction due to very restricted range, very few populations (often 20 or fewer), steep declines, or other factors. G3 = Vulnerable—At moderate risk of extinction due to a restricted range, relatively few populations (often 80 or fewer), recent and widespread declines, or other factors. G4 = Apparently Secure—Uncommon but not rare; some cause for long-term concern due to declines or other factors. G5 = Secure—Common; widespread and abundant.
SUBSPECIES LEVEL Taxa which are subspecies or varieties receive a taxon rank (T-rank) attached to their G-rank. Where the G-rank reflects the condition of the entire species, the T-rank reflects the global situation of just the subspecies. For example: the Point Reyes mountain beaver, Aplodontia rufa ssp. phaea is ranked G5T2. The G-rank refers to the whole species range i.e., Aplodontia rufa. The T-rank refers only to the global condition of ssp. phaea.
STATE RANKING The state rank (S-rank) is assigned much the same way as the global rank, but state ranks refer to the imperilment status only within California’s state boundaries.
S1 = Critically Imperiled—Critically imperiled in the state because of extreme rarity (often 5 or fewer populations) or because of factor(s) such as very steep declines making it especially vulnerable to extirpation from the state. S2 = Imperiled—Imperiled in the state because of rarity due to very restricted range, very few populations (often 20 or fewer), steep declines, or other factors making it very vulnerable to extirpation from the state. S3 = Vulnerable—Vulnerable in the state due to a restricted range, relatively few populations (often 80 or fewer), recent and widespread declines, or other factors making it vulnerable to extirpation from the state. S4 = Apparently Secure—Uncommon but not rare in the state; some cause for long-term concern due to declines or other factors. S5 = Secure—Common, widespread, and abundant in the state.
Notes: 1. Other considerations used when 3. Other symbols: ranking a species or natural community include the pattern of GH All sites are historical; the element has not been seen distribution of the element on the for at least 20 years, but suitable habitat still exists (SH landscape, fragmentation of the = All California sites are historical). population/stands, and historical extent as compared to its modern GX All sites are extirpated; this element is extinct in the range. It is important to take a bird's wild (SX = All California sites are extirpated). eye or aerial view when ranking sensitive elements rather than simply GXC Extinct in the wild; exists in cultivation. counting element occurrences. G1Q The element is very rare, but there are taxonomic questions associated with it.
T Rank applies to a subspecies or variety.
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Listing and Special Status Information
CALIFORNIA ENDANGERED SPECIES ACT (CESA) LISTING CODES: The listing status of each species is current as of the date of this list. The most current changes in listing status will be found in the “Endangered and Threatened Animals List,” which the CNDDB updates and issues quarterly.
SE State listed as Endangered ST State listed as Threatened SCE State candidate for listing as Endangered SCT State candidate for listing as Threatened SCD State candidate for delisting
FEDERAL ENDANGERED SPECIES ACT (ESA) LISTING CODES: The listing status is current as of the date of this list. The most current changes in listing status will be found in the “Endangered and Threatened Animals List,” which the CNDDB updates and issues quarterly. Federal listing actions contained in the Federal Register are also available at: http://www.regulations.gov
FE Federally listed as Endangered FT Federally listed as Threatened FPE Federally proposed for listing as Endangered FPT Federally proposed for listing as Threatened FPD Federally proposed for delisting FC Federal candidate species (former Category 1 candidates)
Section 4(c)(2)(A) of the Act requires the U.S. Fish and Wildlife Service to conduct a review of listed species at least once every five years. Five year reviews from the Sacramento Fish and Wildlife Office are available at: https://www.fws.gov/sacramento/es/Five-Year-Reviews/es_five-year.htm
BLM Sensitive: Bureau of Land Management (BLM) Manual §6840 states that “BLM sensitive species are: (1) species listed or proposed for listing under the Endangered Species Act (ESA), and (2) species requiring special management consideration to promote their conservation and reduce the likelihood and need for future listing under the ESA, which are designated as Bureau sensitive by the State Director(s). All Federal candidate species, proposed species, and delisted species in the 5 years following delisting will be conserved as Bureau sensitive species.” The California-BLM Sensitive Animals list is available at: https://www.blm.gov/programs/fish-and-wildlife/threatened-and-endangered/state-te-data/california
IUCN - The International Union for Conservation of Nature (IUCN): The IUCN assesses, on a global scale, the conservation status of species, subspecies, varieties and even selected subpopulations in order to highlight taxa threatened with extinction, and therefore promote their conservation. Detailed information on the IUCN and the Red List is available at: http://www.iucnredlist.org
Xerces Society Red List: The Xerces Society is an international non-profit organization dedicated to protecting biological diversity through invertebrate conservation. The Society advocates for invertebrates and their habitats by working with scientists, land managers, educators, and citizens on conservation and education projects. Their core programs focus on endangered species, native pollinators, and watershed health. More information on the Red List is available at: http://www.xerces.org.
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