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Bassonia Hakelensis (BASSON) Nov 44 44 he A Rei Series A/ Zitteliana An International Journal of Palaeontology and Geobiology Series A /Reihe A Mitteilungen der Bayerischen Staatssammlung für Pa lä on to lo gie und Geologie 44 An International Journal of Palaeontology and Geobiology München 2004 Zitteliana Umschlag 44 1 18.01.2005, 10:04 Uhr Zitteliana An International Journal of Palaeontology and Geobiology Series A/Reihe A Mitteilungen der Bayerischen Staatssammlung für Pa lä on to lo gie und Geologie 44 CONTENTS/INHALT REINHOLD R. LEINFELDER & MICHAEL KRINGS Editorial 3 DIETRICH HERM Herbert HAGN † 5 KAMIL ZÁGORŠEK & ROBERT DARGA Eocene Bryozoa from the Eisenrichterstein beds, Hallthurm, Bavaria 17 THORSTEN KOWALKE Evolution of the Pachychilidae TROSCHEL, 1857 (Chaenogastropoda, Cerithioidea) – from the Tethys to modern tropical rivers 41 HERBERT W. SCHICK The stratigraphical signifi cance of Cymaceras guembeli for the boundary between Platynota Zone and Hypselocyclum Zone, and the correlation of the Swabian and Franconian Alb 51 GÜNTER SCHWEIGERT, RODNEY M. FELDMANN & MATTHIAS WULF Macroacaena franconica n. sp. (Crustaceae: Brachyura: Raninidae) from the Turonian of S Germany 61 JÜRGEN KRIWET & STEFANIE KLUG Late Jurassic selachians (Chondrichthyes, Elasmobranchii) from southern Germany: Re-evaluation on taxonomy and diversity 67 FELIX SCHLAGINTWEIT Calcareous green algae from the Santonian Hochmoos Formation of Gosau (Northern Calcareous Alps, Austria, Lower Gosau Group) 97 MICHAEL KRINGS & HELMUT MAYR Bassonia hakelensis (BASSON) nov. comb., a rare non-calcareous marine alga from the Cenomanian (Upper Cretaceous) of Lebanon 105 ALFRED SELMEIER First record of Matisia (Bombacaceae) and Crudia (Caesalpiniaceae) wood from the Tertiary of Rio Paranaibo, Brazil 113 ULRICH STRUCK, TOBIAS HEYN, ALEXANDER ALTENBACH, JÜRGEN ALHEIT & KAY-CHRISTIAN EMEIS Distribution and nitrogen isotope ratios of fi sh scales in surface sediments from the upwelling area off Namibia 125 Hinweise für Autoren/Instructions for authors 133 Zitteliana A 44 136 Seiten München, 30.12.2004 ISSN 1612-412X Innentitel44 5 18.01.2005, 9:35 Uhr EDITORIAL NOTE HINWEIS DES HERAUSGEBERS As of in 2003, the journal Zitteliana is published in two Vom Jahr 2003 an erscheint die Zeitschrift Zitteliana in zwei series. Reihen. Series A: Mitteilungen der Bayerischen Staats samm lung für Die Reihe A: Mitteilungen der Bayerischen Staats samm lung Paläontologie und Geologie (ISSN 1612-412X) replaces the für Paläontologie und Geologie (ISSN 1612-412X) ersetzt die former „Mit tei lun gen der Bayerischen Staats samm lung für bisherigen „Mit tei lun gen der Bayerischen Staats samm lung für Pa lä on to lo gie und historische Geologie“ (ISSN 0077-2070). Pa lä on to lo gie und historische Geologie“ (ISSN 0077-2070). The num bering of issues is continued (last published: Heft 43, Die Bandzäh lung (zuletzt erschienen: Heft 43, 2003) wird fort - 2003). ge setzt. Series B: Abhandlungen der Bayerischen Staats samm lung für Die Reihe B: Abhandlungen der Bayerischen Staatssammlung Paläontologie und Geologie (ISSN 1612-4138) continues the für aPaläontologie und Geologie (ISSN 1612-4138) führt die pre vious „Zitteliana – Abhandlungen der Baye ri schen Staats - bis he ri ge „Zitteliana – Abhandlungen der Baye ri schen Staats - samm lung für Paläontologie und historische Geologie“ (ISSN samm lung für Paläontologie und historische Geologie“ (ISSN 0373-9627). 0373-9627) fort. Instructions for authors are included at the end of this Hinweise für Autoren beider Reihen sind am Ende dieses volume. Bandes enthalten. Editers-in-Chief/Herausgeber: Reinhold Leinfelder, Michael Krings Production and Layout/Bildbearbeitung und Layout: Lydia Geißler, Manuela Schellenberger Bayerische Staatssammlung für Paläontologie und Geologie Richard-Wagner-Str. 10, D-80333 München, Deutschland http://www.palaeo.de/zitteliana email: [email protected] Für den Inhalt der Arbeiten sind die Autoren allein ver ant wort lich. Authors are solely responsible for the contents of their articles. Copyright © 2004 by Bayerische Staassammlung für Pa lä on to lo gie und Geologie, München Die in der Zitteliana veröffentlichten Arbeiten sind urheberrechtlich geschützt. Nachdruck, Vervielfältigungen auf photomechanischem, elektronischem oder anderem Wege sowie die Anfertigung von Übersetzungen oder die Nut zung in Vorträgen, für Funk und Fernsehen oder im Internet bleiben – auch auszugsweise – vorbehalten und bedürfen der schriftlichen Ge neh mi gung durch die Bayerische Staatssammlung für Paläontologie und Geologie, München. ISSN 1612-412X Cover illustration: Phorcynis catulina THIOLLIÈRE, 1854 (BSP 1990 XVIII 51) from the lower Tithonian of Zandt / Denkendorf (Bavaria), ventral view, 25 cm. Photograph: G. JANßEN (LMU München, Department für Geo- und Umweltwissenschaften, Sektion Paläontologie) Umschlagbild: Phorcynis catulina THIOLLIÈRE, 1854 (BSP 1990 XVIII 51) aus dem unteren Tithon von Zandt / Denkendorf (Bayern), Ventralansicht, 25 cm. Foto: G. JANßEN (LMU München, Department für Geo- und Umweltwissenschaften, Sektion Paläontologie) Innentitel44 1 18.01.2005, 9:35 Uhr Zitteliana A44 105-111 2 Textfi gs, 2 Pls München, 30.12.2004 ISSN 1612-412X Bassonia hakelensis (BASSON) nov. comb., a rare non-calcareous marine alga from the Cenomanian (Upper Cretaceous) of Lebanon By Michael Krings* & Helmut Mayr Bayerische Staatssammlung für Paläontologie und Geologie und GeoBio-CenterLMU, Richard-Wagner-Straße 10, 80333 München, Germany Manuscript received 5 June 2004, revision accepted 9 August 2004. Abstract Schlüsselwörter: Algites SEWARD, Bassonia nov. gen., fossile unkalzifi zierte Alge, Haqel Fischschiefer, Oberkrei- A nearly complete specimen of the Cenomanian (Late de, Pithyopsis tasmanica (SONDER) FALKENBERG (Ceramiales, Cretaceous) non-calcareous alga Bassonia hakelensis (BASSON) Rhodomelaceae) nov. comb. is reported from the Haqel fi sh beds in Lebanon. Bassonia hakelensis was originally described under the name Algites hakelensis BASSON, 1972. Based on the distinctive 1. Introduction morphology of the thallus, we fi nd it compelling to exclude this species from Algites SEWARD, 1894, and assign it to a new Based on the biological and ecological signifi cance of algae genus, for which the name Bassonia nov. gen. is introduced. in many aquatic environments today (ROUND 1981; GRAHAM Bassonia hakelensis displays morphological characters also & WILCOX 2000), a detailed knowledge of their fossil record, found in certain extant rhodophytes, but assignment to any evolution, and the roles they played in biological and ecological particular extant taxon is impossible because important features processes in the past is important in understanding the evolu- required today in order to establish the systematic position of tionary history of ancient and modern aquatic ecosystems. For- an alga cannot be determined. tunately, the fossil record of algae is copious, and many ancient forms are increasingly well-understood relative to their geo- Key words: Algites SEWARD, Bassonia nov. gen., fossil graphical distribution, stratigraphical range, paleobiology and non-calcareous alga, Haqel fish beds, Late Cretaceous, -ecology. Certain groups of microalgae, charophyte oogonia Pithyopsis tasmanica (SONDER) FALKENBERG (Ceramiales, (gyrogonites), and the remains of calcareous macroalgae from Rhodomelaceae) marine and brackish environments are especially abundant and diverse in the fossil record because their thick walls and/or calcium-based skeletons are readily preserved (e.g., FLÜGEL Kurzfassung 1977; TAPPAN 1980; RIDING & VORONOVA 1985; RIDING 1991; TAYLOR & TAYLOR 1993; BROOKE & RIDING 1998; AGUIRRE et Aus den Haqel Fischschiefern (Cenoman, Oberkreide) im al. 2000). On the other hand, non-calcareous macroalgae are Libanon wird ein nahezu vollständiges Exemplar der unkalzi- comparatively rare as fossils, which is primarily the result of fi zierten AlgeBassonia hakelensis (BASSON) nov. comb. vorge- the fact that fossilization potential of the delicate algal bodies stellt. Bassonia hakelensis ist ursprünglich unter dem Namen is low; they are preserved only where the depositional environ- Algites hakelensis BASSON, 1972 beschrieben worden. Aufgrund ment provided ideal conditions for fossilization (e.g., reduced der charakteristischen Thallusmorphologie halten wir es für rates of decomposition, relatively stagnant water, and/or quick angebracht, diese Art von Algites SEWARD, 1894 abzugrenzen embedding) and may sometimes yield exquisite fossils. The fos- und in eine neue Gattung zu stellen, für die der Name Bassonia sil record of non-calcareous macroalgae throughout geological nov. gen. eingeführt wird. Bassonia hakelensis besitzt morpho- time includes spectacular, anatomically preserved Late Neo- logische Merkmale, die auch bei einigen modernen Rotalgen zu proterozoic algae from the Doushantuo Formation in South fi nden sind; eine Zuordnung des Fossils zu einem bestimmten China (XIAO et al. 2004 and literature cited therein), various modernen Taxon ist allerdings nicht möglich, da wesentliche, compressed algal thalli from the Lower Cambrian Kaili Biota für die Bestimmung von Algen generell erforderliche Merkmale (Guizhou Province) in China (YANG et al. 2001a, 2001b) and nicht erhalten sind. Middle Cambrian Burgess Shale (British Columbia) in Can- *Author for correspondence and reprint requests; E-mail: [email protected] Krings_Mayr 105 18.01.2005, 9:50 Uhr 106 ada (WALCOTT 1919), the famous Ordovician Lake Winnipeg a warm and shallow sea. The limestone was deposited
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