ACTA FAUNISTICA ENTOMOLOGIC A MUSEI NATIONALlS PRAGAE VaL 11, No. 113 Edit. 17. 1. 1966
(Acta faun . ent. Mus. nat. Pragae, 11: 331-372.)
Czechoslovak Species of the Genus Platypalpus .Macquart ( Diptera, Empididae)
LIBUSE Z U S K 0 VA Central Research Institute foT Plant Production, Department of Entomology, Praha-·Ruzyne
Abstract The present paper provides a review oJ the Czechos lovak species of the genus Platypalpus. A key to Czech.oslovak species and a !ist of species with their synonymy, systematic notes and occurrence in Czechoslovakia are included. Fo-rty species arH dealt with, three of them are here described as new tn science.
1. INTR.QD UCT ION The aim of this study is to present a critical review of the Czecho slovak species of the genus Platypalpus, which has up to now been lacking in the literature. The genus Plat,ypalpus represents a group of flies, rich in indivi duals and numerous in species, very difficult to study. In consequence there are very often nomenclatural and systematical obscurities in the older literature. The systematics of the genus was worked out satisfac torily as late as 1926 by Collin, though some important papers or books on the subject were published before that date (Frey, 1907; Melander, 1910, dealing with North American species; Lundbeck, 1910). Collin un dertook the immense work of revising the genus on the basis of study ing the types of almost all the described palaearctic species and ex plaining the synonymy of this group. The afore-mentioned paper is the very successful result of his studies. A further comprehensive publication is the revision by Engel ( 1939). Unfortunately, the appearance of this book did not mean much advancement in our knowledge as it was not written too carefully from the nomenclatural standpoint. Seguy ( 1942) published a rathe·r extensive paper on this genus. containing some de tailed re-descriptions and a key to species, and Frey ( 1943) a key to the palaearctic species of the genus. Recently Collin ( 1961) · reviewed the British species and this. work is also of extraordinary importance for the systematics of the genus P'latypalpus. The faunistic records referring to the Czechoslovak species of the genus Platypalpus are included in the papers by Dalla Torre ( 1878], Kowarz ( 1894], Czizek ( 1906], Landrock ( 1907, 1908, 1910], Brancsik
1 331 ( 1910), Vimmer ( 1913), Fekete ( 1914) and Thaihammer ( 1918 ), which were, however, all published before the appearance of Collin's revision. Therefore all records given in them are most unreliable. The revision of the available materials, having been a basis for those studies (from Czize.k, Landrock, and Vimmer's collections), showed that the greater number of the specimens were misidentified. For these reasons, only the captures revised by myself were considered, no other records were in cluded. However, this does not refer to single records in the papers published after Collin's revision, which are considered here. In the present paper the following abbreviations were used: NMP - deposited in the National Museum in Praha MMB deposited in the Mmavian Museum in Brno' ZZ - collected by L. Zuskovä and J. Zuska and deposited in their col- lection Lit - recmds in the Iiterature Mat - new or revised materials Boh - Bohemia Mor Moravia and Czechoslovak Silesia Slov - Slovakia
2. KEY TO CZECHOSLOVAK SPECIES
The genus Platypalpus is characterized as follows: rather small flies, mostly black (only a few species yellow], predominantly dusted. Eyes separate in both sexes. One or two pairs of vertical bristles develo·· ped. Humeral calli present. Acrostichal bristle·s usually biserial. Middle femora conspicuously stoutened, bearing double ventral row of black sharp spines; middle tibiae with single ventral row of similar bristles, apically possessing process developed to various extent, mostly as long strong black spur. Wings with three closed fields: anterior basal, poste rior basal fused with disc'oidal, and anal. 1 ( 12) Two pairs of vertical bristles: inner convergent, outer divergent, but botlt distinct. 2 ( 5) Acrostichal bristles in four or more rows. 3 ( 4) Black species. Frons strongly widerred behind ocelli, with brownish grey dust. An1ennae short, brownish black; first two joints extraordinarily short, third joint broadly oval. .Palpi very small, brown. Mesonotum black with dark grey dust, also pleurae dusted except for shining sternüpleura and spot on hypo pleura. Acrostichal bristles in four rows, dorso·centrals in two rows at each side of mesonotum. Legs yellow. but coxae and trochanters of second to third pair O>f legs, s-ometimes bases of anterior femora and almost entire middle and posterior femo·ra brown to brownish black; only last two tarsal joints brownish black tO' black; a,pical spur of middle tibiae lang, strong and sharp. Wings with fine yellowish brown tinge. Abdomen black, shining. - Length: body, 2.0 to 2.3 mm; wing, 2.5 to 2.7 mm . . Platypalpus rapidus Meig. 4 [3] Yellow species with black head and brownish black ab-domen. Frons rather wide, with yellowish grey dust. BD'th basal jO'ints of antennae yellow [third joint broken in bo-th specimens examined ). •Palpi small, yellow. Thorax yellow, with fine grey dust, only sternopleura and broad prescutellar pmtion of me· sonotum shining. Mesonotum with median longitudinal black stripe becoming wider befo·re scutellum, with short unifo-rm pale yellow pilosity, without distinct acro·stichal and do-rsocentral bristles. Humeral, no-to•pleural, postalar and scu tellar bristles conspicuous, lang, brownish black. Legs yellow, only trochanter>
332 with bla'ek spots; tips of all femora with rather Iong brorwnish blaek bristles: last three tarsal joints .nf anterior legs eonspieuously widened; apex of middle tibiae only with very slwrt rounded yellow proeess; tops o>f anterior and posterior tibiae with .pale yellow eombs. Wings hyaline. Abdomen brownish blaek, sterna .and terga dusted greyish. Length: body, 2.6 to 3.1 mm; wing, 3.1 to 3.5 mm . . Platypalpus bohemicus n. sp. ~ 5 (2) Aerostiehal bristles in two rows. 6 (9) Third antennai joint not mo•re than twiee as lang as deep at its base. 7 (8) Larger speeies, 2.7 to 3.3 mm. Pleurae mo•re denseiy dus ted, especially hypo pleurae. Legs yellowish, with posterior eo-xae anq troehanters blaek and base of anterior e,o,xae slightly >dar'kened in male, but mure m less darkened in female. r4+s and m rather divergent Sides .of first abdominal segment and anterior margins of seve-ral foHowing segments dusted. Hypopygium !arge and rather staut; left part of epandrium very !arge with dense lang yellowish fringes on left margin meeting and mingling with tuft of similar hairs on right part. - Wing length, 2.8 to 3.0 mm . . Platypalpus agilis Meig. 8 (7] Smaller species, 1.75 to 2.1 m111. >Pleurae less dusted, sternopleura and espe eially hy,popleura extensively bare and shining. Legs yeUowish in both sexes. r4+s and 111 almost parallel. Abdomen without dusted 111argins of segments, shining. Hypopygium rather s111aller, left part of epandrium with only few hairs on its· .Jeft margin, very different from tangled 111ass of yellowish hairs in P. agilis. - Wing length, 2.3 mm . Platypalpus agilellus Co!!. 9 (6) Third antennal joint at least three times as lang as deep at its base. Meso· notum blaek, shining,. only its edges, hu111eral ealli and seutellu111 dusted. Apieal spur of middle tibiae absent. Abdomen blaek, shining. Blaek, moderately !arge speeies. 10 (11) Antennae entirely blaek. - Vertex and narrow stripes along eye margin shin ing, frons otherwise !dusted. Arista slightly Ionger than half of third antennal joint. P!eurae dusted but entire sternopleura and hypopleura shining. Thoraeie bristle>s blae•k. Legs dark yellow, middle femora with four to five lang brown bristles posternventrally. In male posterior tibiae eurved in 111iddle of their length and slightly thiekened in distal third. Wings tinged brownish, veins brown. Hypopygium eonspieuously !arge ahd eharaeteristieally formed (fig. 36J. - Length: body, 2.0 to 2.5 mm; wing, 2.5 mm . Platypalpus nigritarsis Fall. 11 (10] Basal two antennal j.oints yeUowish brown, third joint blaek. - Vertex and front dusted greyi•sh. Artsta very short, one third times as lang as third an tennal joint . .Pleurae dusted, only anterim part of sternopleura shining. Tho raeie bristles brown to brownish blaek . Legs yellow, only last three tarsal joints brownish; middle femo·ra without posteroventral row of bristles. Posterior tibiae of male slender, of normal shape. Wings hyaline, veins yellow basally but getting darker towards apex. Hypopygium small. - Length: body, 1.7 tö 3.0 mm; wing, 2.2 to 2.5 mm . . Platypalpus longicornis Meig. 12 (1) Only one pair of convergent vertieal bristles develo•ped m vertex with unifo·rm hairs without distinet pair of vertieals. 13 (34] Mesonntum shining o•r only very slightly dusted, but ground eolour easily visible. 14 [19) Aerostiehal bristles not or only very narrowly separated from remaining pilo sity nf mesonotum; pilosity of mesonotum short, pale yellow. 15 [18) Speeies entirely yellow, or at least with yellow mesonotum with dark median stripe. 16 (17j Mesonotum entirely yellow, without dark median stripe; seutellum yellow. Third ancl fourth joint of anterior tarsi broadly dilated. - Yellow, rather !arge, but slencler species. Head entirely yellow exeept for black eyes. In lateral aspeet faee eonspicuously protruded; elypeus dusted greyish. Basal antennal joints and base o•f third joint yellow; third jo'int oval, in apieal part brownish blaek similar to arista; this almost three times as lang as third joint. HumeraJ bristle short, preseutellar dorsoc.entral bristles developed. Legs yellow, only tip nf last tarsal joint brown; first tarsal joint of anterioT legs extraordinarily lang, Ionger than half length -of anterinr tibia and almost as lang as four fol lowing joints together; base of this joint bearing lang pale ventral bristle. Double row o>f blaek spines an m:iddle fenwra beginning at a third of its length, at
1* 333 point coinciding with end of apical spur of tibia when this is bent; apical spur short and stüut. Wing.s very lang, veins brown. Abdomen yellow, shining, sometimes irregularly darkened. - Length: büdy, 2.7 to 3.3 mm; wing, 3.5 to 4 mm . Platypalpus luteus Meig. 17 ( 16) M.esonotum yellow with broad median black stripe. Base oJ scutellum usually darkened. All tarsal joints slender. - Moderately !arge pale species with !O'ng narrow wings. Head entirely black. Face extramdinarily narww; clypeus black, shining. Antennae brown o'r sometimes brownish black with paler basal joints: tlürd: joint not oval, about twice· as lang as deep; arista 1.5 times as long as third joint. Humeral bristle lacking, prescutellar dorsocentral bristles incon spicuous. Legs yellow, last tarsal joint dark; first tarsal i'Oint of anterior legs not so developed as in P. luteus. Middle femora with spines beginning as usually, but spines in posterim row Ionger and stronger than in anteriür one; apical projection of middle tibiae very short. Wings slender, veins yellowish brown, point of fusing of c and r1 conspicuously darkened to black. Terga nf ab domen shining black or brownish black, sterna yellowish. - Length: body, 2.2 to 2.25 mm; wing, 3.4 mm . . P'latypalpus pectoralis Fall. 18 ( 15) Black, shining, müderately I arge species. Antennae black, rather lang, more pubescent than usual. VerteXJ and occiput slightly dusted, but two small shining black spots located laterally to ocelli. Palpi conspicuously !arge, 3/4 length of proboscis, yellow, whitish grey pube-scent. Mesonotum shining, only its lateral and posterior edges dusted. Scutellum and pleurae finely dusted except for almost entire sternopleura. Legs yellow, but last four tarsal joints yellowish brown. Middle femora with posternventral row of five to six long conspicuous bristles in apical half. Tip üf middle tibiae only with short pyramidal protuberance. Abdominal terga black, shining, sterna yellowish brown. - Length: body, 2.5 to 3.3 mm; wing, 3.0 to 4.0 mm . . Platypalpus pallipes Fall. 19 ( 14) Acrostichal bristles separated from remaining pilosity of mesonotum by uistinct bare stripes. 20 ( 21) Acrostichal and dorsocentral hairs distinct and numerous; form er biserial, latter uniserial becoming Ionger posteriody. Front strongly widened before anterior ocellus. Occiput with short uniform :Pilosity. Antennae black, short, but arista 3 to 3.5 times as long as third joint. Mesonotum black, shining, only edges dusted. Scutellum and pleurae, except for sternopleurae, also dusted. Legs brightly yellow, only last ta,rsal joints black. Ventral black spines on middle femora in basal half lang and ·distant from one another, in apical half shnrt and crnwded. Middle tibiae thickened and densely whitish grey duste:l apically, their tips blunt, without spur. Wings lang and narrow, veins yellowish brown. Hypopygium conspicuously !arge, globular, finely greyish pilose. - Length: body, 2.5 to 3 mm; wing, 3.5 mm . . Platypalpus ciliaris Fall. 21 ( 20) Mesonotum almost bare. Acrostichal and dors-ocentral hairs inconspicuous, short and sparse. 22 ( 27) Third antennal joint three to- four tim es as long as dBep at its base ( except for female of P. stigmatellus) .. 23 ( 26) Arista white. 24 ( 25) Pleurae shining excei[)t fm dusted narrow stripe connecting anterior coxae and halterae, anterior pa,rt of hypüpleura, whole ptoeropleura and metapleura. ·- Front very narrow as far as to· the ocelli, dusted. !Occiput with triangular dusted spo,t. Almost entire thorax black, shining; edges of mesonotum, whole scutellu:m and meso·phragma dusted. Legs yellowish brown, all tarsal jo·i nts brownish black. Anterior femo:ra bearing posterDventral row of three to fivP. lang black bristles, these Ionger than femora deep. Middle femora with even Ionger pos teroventral black bristles. Tip o.f middle tibiae bearing blunt apical process. Wings slightly tinged brownish, veins dark brown. Hypopygium usu · ally shape, rath'8r small. - Length: body, 2.0 to 2.5 mm; wing, 2.5 to 2.8 mm Platypalpus albiseta Panz. 25 (24) Pleurae dusted greyish but sternopleura shining. - Front dusted, narrow, but distinctly widened in front of anterior ocellus. Occiput finely and uniformly dusted greyish. Mesonotum shining, black, edges of that dusted. Scutellum, mesophragma and pleurae dusted except for shining sternopleura. Legs yellow, tarsal joints brown. Anterior femora with pale brown posteroventral bristles,
334 these shorter than femora deep. Middle femo-ra with posteroventral row of nine ta ten lang blac•k bristles. Middle tibiae without apical spur. Wings practi · cally hyaline, veins dark yellowish. Hypopygium J.onger than in P. albiseta. - Length: büdy, 2.2 to 2.45 mm; wing, 2.75 mm . . Platypalpus niveiseta Zett. 26 (23] Arista dark. - Moderately !arge black species with conspicuously lang and slender wings. Front and vertex shining, occi.put slightly dusted greyish. Vertex uniformly pilose. Antennae black, very lang, especially in male, where third joint almost four times ·as lang as deep at its base, while in female only two to tw'o and one half times as lang. Palpi yellow, Ionger in female. Mesonotum black, shining. Acrostichal and dor.so·central bristles pale yellow, very short, inconspicuous. Scutellum and pleurae dusted greyish except for entire meso pleura and ster.nopleura. Legs yellow, distal parts of middle and posterinr femora and posterior tibiae usually darker. Tarsal joints D'f middle and posterior legs rather distinctly annulated. Middle femora without posterD ventral row o•f bristles, but spines in posterior row conspicuously lang, bristle shaped. Tip of 111iddle tibiae with so111ewhat indistinct blunt prncess. Veins bro•wn, section of c fusing with end of r1 o.ften black. - Length: body, 2.5 to 3.0; wing, 3.5 to 4.0 P'latypalpus stigmatelins Zett. 27 (22] Third antennal joint only twice as lang as deep at its base. 28 (33) Legs yellow. 29 (30) Middle fenwra strongly thic'kened. Front shining, black. Occiput only slightly dusted. Basal antennal jo'ints and basal half of third joint yellow, apical half brown to• black; third joint hardly twice as lang as deep at its base. Brown to black arista about 1.5 times as lang as this jo'int. Mesonotu111 shining, edges co·vered with broad band of brownish-grey dust. •Pleurae dusted, sterno pleura shining. Hairs and b>ristles on 111esonotum slwrter than usual, biserial acrostichals almost indistinct, dorsocentrals in one row. Legs yellow, only tarsal joints darkened at their tips. Middle femora strongly darkened, with double row of ventral spines, but these for111ing single row basally; in apical part two to four Ionge.r spine-like po·steroventral and several anteroventral bristles; anteriorly four to six sharp yellow spines. Tip of 111iddle tibiae bearing conspicu·· ously strong and lang apical spur with sharp darkened a,pex; ventral row of spines on these tibiae accupping only basal two thirds of their Iength. Wings ting ed yellowish, veins yellow basally, yelowish brown apically. Prefurca extending onLy one-third of way back tü hu111e:ral vein. - Length: body, 2.0 to 2.75 mm; wing,2.5 to 2.7 m111 Platypalpus calcaratus Goll. 30 (29) Middle fe111ora slightly thickened. Front dull. '
31 (32] Antennae practically black or basal joints with reddtsh tinge . .Palpi extre111ely . !arge, al111ost as lang as proboscis, o·range, with blunt apex directed posteriorly. ' covered with very fine and short yellowish pile. Vertical bristles conspicuously wide apart. Dusting of thorax less dense, scutellum dusted. Acrostichal and dorsocentral bristles lang, ea'sily visible. Middle knees yelow o·r slightly I tinged bro•wnish. Middle fe111ora less stoutened, their thickness Jargest at or behind half of their length. Middle tibiae langer, only very slightly thickened, their apical spur reaching base of fe111ur; middle 111etatarsi almost one third 'times as lang as middle tibiae (without spur). - Length: body, 1.5 tü 1.9 111m; wing, 1.8 111111 . 'Platypalpus ingenuus Coll. 32 [31) Antennae yellow with brownish black arista. Palpi very !arge but shorter tha11 in previous ~pecies, reddish, with sharp distinct apex directed anteriorly, with whitish pale and several langer whitish hairs. Vertical bristles closer together. Dusting üf thmax dense, disc of scutellu111 shining black. Acrüstichal and dorso central bristles very fine· and short, quite inconspicuous. Middle knees with distinct black spot. Middle femo-ra 111ore stoutened, their thickness largest defore half of their length. Middle tibiae curved, 111ore thickened, their apical spur not 'reaching base bf femur. Middle 111etatarsi one half as lang as middle tibiae (excluding spur). - Length: bO'dy, 1.7 to 2.3 m111; wing, 2.2 to 2.5 111111 Platypalpus ruficornis Macq. 33 (28) Legs predominantly black or brownish black; bases of all femora, al111ost entire tibiae and bases of tarsal joints dark yellow. - Antennae black, very shmt, arista twice as lang as third jO'int. Front rather narrow, faintly dusted J;Jrownish grey. MesO'notu111 1110derately dusted brownish grey, sometimes with broad median shining longitudinal stripe. tPropleura and band connecti'ng this with halterae dusted silvery, metapleura and metanotum 'dusted grey. Bristle' an d haj:rs o-f mesonotum sho-rt, brlownish black to black. Apical spur of midd](~ tibiae sharp, rather •strong. Wings tinged brownish in costal part, veins brown. Abdomen shining, black. - Length: body, 1.5 to 2.0 ; wing, 1.9 mm . Platypalpus nigel' Meig. 34 ( 13) Mes onotum covered with m ore or le·ss dense greyish dust, ground coVour not visible ( except f.or narrow shining median stripe in P. minutus, where anterior tibiae possess a,pical process ). 35 (50) Antennae entirely black. 36 (37) Mesonotum with narrow median longitudinal shining stripe. - Black, rathe1· tiny species. Antennae short. Humeral calli shining laterally. Pleurae dusted except for shinin~ black sternopleura. Legs alm;ost whoHy black, bearing hairs more crowded than usual. Anterior tibiae sligthly stoutened, spindle-like, with small process hearing comb on their tips. Mi ddle tibiae with lang sharp black apical spur. All tarsal joints distinctly annulated, last joints of anterio·r tarsi müderately dilated. Veins brown t o dark brown, only wing bases paler. - Length: llody, 2.0 to 2.5 mm; wing, 2.8 mm . Platypa.lpus minutus Meig. 37 (36) Mesonotum entirely dust-ed, without shining median stripe. ' 38 ( 39) Stronger thoraclc bri·stles black. - -Rather !arge black species. Occiput ancl face duste•cl grey. Third antennal jo•int 3.0 to 3.5 times as lang as deep at its base. Arista shorter than thircl jo-int in male, longei' in female. Mesonotum anrl pleurae except for stern;o,pleura dusted pale grey. Acr ostichal bristles sho-r t, arranged in four irregular rows. Dorsocentral bristles s lightly Ionger ancl darker, in one row at each siele. Legs yellow, only middle and posterior knees conspicuously black. Apical part of posterior tibiae darkened. Tarsal joint3 annulated black. Hairs co-vering legs rather l
336 than usual, espe'Cially anteroventral and pnsteroventral rows of bristles on anterior and middle femora. Posterior femora with conspicuous [ especially in male) rather strong anteroventral bristles. Apical spur of middle tibiae lang, black, strong and sharp, on irs tip bearing very tiny bristle. Veins yellowisil brown. r4+5 and m parallel. Abdomen black, shining, -only first two· joints narrowly dusted t!aterally. - Length:i body, 2.5 to 3.1 mm; wing, 2.8 to 3.0 mm Platypalpus infectus Co!!. 45 [ 42) Third antennal joint only ab out twice as lang as deep at its base and shmter than arista. 46 [ 49) Abdomen black, shining, usually only with narrow dusted stripes at sides of first two segments. Mo·derately large species. 4 7 [ 48) Ocellar bristles as l-ang as vertical; these normaL! y distaut from each other. G!ypeus dusted. Third antennal joint 1.5 times as lang as dee,p at its base; arista twice as lang a·s this joint. Mesonotum comparatively faintly dusted brownish grey. Two rows of acrostichal bristles strongly distaut from each o·ther, Legs yeJl.ow, in males with all or at least middle femora bearing brown sp::J•ts and rings; tarsal jüints sharply annulated. Wings tinged brownish, veins yellow. - Length: 'body, 3.0 to 3.5 mm; wing, 3.8 to 4.0 mm . Platypalpus interstinctus CoH. 48 [ 47) Ocellar bristles half as lang as vertical; these conspicuously di'stant from each other. Clypeus shining. Third antennal joint twice as lang as deep at its base; arista to three times as lang as this joint. Mesonotum covered with rather dense yellowish goiden dust. Two· rows of acrostichal bristles nnrmally distaut from each other. Legs yellow, only knees of middle· and posterior femora in both sexes with blackish spots; tarsal joints annulated. Wings conspicuously yellowish, veins paJe yellow. - Length: body, 2.0 to 3.0 mm; wing, 2.8 to 3.2 mm . Platypalpus luteipes, n. sp. 49 [ 46) Each abdominal segmenth with a pair of parge triangular dusted lateral spots. Arista almost twice as lang as third antennal joint. •Palpi mnderately !arge, yellowish o·range, dusted silverish. Thorax entirely dusted grey except for shining black sternopleura. Acrostichal bristles biserial, dorsocentral in one irregular row at each side. One Ionger and stronger posthumeral bristle de velo·ped, close to humeral callus. Thoraeie bristles yellowish brown, finer and shorter hairs yellow. Legs usually paler in female than in male. Middle and posterio·r coxae brownish black, anterior and middle femora usually with brownish black spots, anteriD'f and middle tibiae darkened in their apical 'parts. Apical spur o.f middle tibiae strong and sharp, blackish. Tarsal joints slender, annulated black. Wings slightly tinged brownish in apical part. Veins brownish blac·k except fo·r yellowish brown bases. r4 +s and m di verging, curved, in apical section 111 approaching to 4 +s. - Length: body, 3,4 to 3.9 mm; wing, 3.1 to 3.3 mm . Platypalpus binderi n. sp. 50 [ 35] At least basal two· joints o·f antennae yellow or yellowish brown. 51 [56) Entire sternopleura dusted greyish. 52 [53) Front very narrow. Tarsal joints only very faint!y annulated, rather unicoloured. Vein m strongly curved, in apical section approaching to r4+5. - Vertical bristles close together. Face wtde, dusted whitish grey, in lateral aspect prominent. Peristom rather wide, distinctly dusted below eyes. Antennae short, their basal joints yellow, third joint brown, about 2.5 times as lang as deep at its base, arista about 1.5 ti111es as lang as third jüint. Palpi !arge, pale yellow, dusted whitish. Tho•rax entirely dusted pale grey, all thoracic bristles and hairs pale ye;l!ow. Legs yellow, apical spur of middle tibiae rather lang, sharp, darkened towards tip. Veins yellowish brown, paler in basal parts. Abdomen brownish black, but üften yellowish brown laterally and ventrally. - Length: body, 3.0 to 4.0; wing, 3.6 mm . Platypalpus candicans Fall. 53 (52) Frons wide. Tarsal jo•ints more or less annulated. Vein 111 not conspicuously curved towards r4 + 5. 54 (55] Large species with lang wings. Dorsocentral bristles not conspicuously lang. Mesonotum with two longitudinal darker stripes, distinct especially in anterior part. - Antennae short, basal jo•ints and base Gof third joint yellow, apical part of third joint brown. Arista wlwlly dark. 1Pa!pi lang, Ionger than wide. Entire thorax dusted pale grey. Legs pale· yellüw. Apical spur of middle tibiae sharp,
337 its tip darkened. Wing almost hyaline, veins yellow basally, torwards apex gra dually becoming brown. r4 + s and m converging. Abdomen black, shining, but first to sixth segment with distinct lateral triangular dusted spots. - Length: body, 3.5 to 4.5 mm; wing; 4.0 to 4.5 mm . . ·Platypalpus cursitans Fabr. 55 (54) Moderately !arge grey species. Dorsocentral bristles c.ouspicuously Jong. Meso· notum without dark longitudinal stripes. - Antenna entirely yellow in male, with third joint e'xcept fo·r its basal ,part brown in female. Arista brown. Palp! wide, oval. Mesonotum duste.d yellowish grey, pleurae dusted pale grey. Legs bright yellow: In male tarsal joints of anterior legs with dark spot at base, seeming to : have double annulations. Apical spur of middle tib!'l.e sharp, darkened towards titp. Wings yellowish, all veins yellow, r4+ s and m almost parallel. Abdominal segments each with pair of lateral triangular spots dusted greyish. - Length: body, 2.2 to 3.0 mm; wing, 2.7 to 3.3 mm . Platypalpus verrali Co11. 56 (51) Sternapleura with distinct shining spo·t. 57 ·(64] Entire antennae yellow, a pex of third joint sometimes brownish. 58 [63) Small species belo·w 3 mm in body length. Anterinr fenwra not much thinner than middle ones. Veins r4 + s and m almost parallel. 59 (60) Palpi !arge, pale yellow, dusted silvery, with conspicuously sharpened tip. Tarsal joints not annulated, only apical half of last joint brown. Tiny species with thnrax dusted ,pale grey. Rather narrow front and vertex dusted pale grey, vertical bristles easily visible, yellow, close to-gether. Antennae yellow, dusted silvery, arista dark yellow towards apex. Third antennal joint about twice as lang as deep at its base. Thnrax entirely dusted silvery except for brownish black shining sternopleura. Acros tichal bristles in two to three irregular rows. All hairs an mesonotum yellow. Legs yellow. Apical spur o.f middle tibiae almosr as lang as tibia deep, rounded, with short hair an its tip. Abdomen brownish black, pilo•se pale yellow, first two segments and ventral side paler. - Length: body, 1.7 to 2.3 mm; wing, 2.5 mm . . 'Platypalpus albicornis Zett. 60 [59] •Palpi broadly oval, their tips rounded. Tarsal joints annulated. Rather small species. Antennae yellow, in female third joint brown apically. Face narrower than frons. 61 ( 62) Face dusted yellowish grey. Vertical bristles close together. Mesonotum dusted 'pale grey. Abdomen black, shining, only with narrow dusted band at its base. - Arista brown. Acrostichal bristles in irregular bi- to triserial rows. Legs pale yellow; tarsal joints, especially in anterior legs, with broad brown annu· lations ~ [less distinct than in P. flavicornis ); apical spur .af middle tibiae short and blunt ( appearing sharp in profile!) in male, shmt but sharpened in female, almost entirely brownish black. - Length: bGdy, 1.8 to 2.5 mm; wing, 2.1 to 2.3 mm Platypalpus pallidicornis Coll. 62 [ 61] Face dusted silvery. Ve rtical bristles rather distant from each other. Me sonotum dusted yellowish grey. Each abdominal segment with a pair of !arge triangular dusted lateral spot s. Aris ta brownish black. Acrostichal bristles biserial, but n nt t :JO distinct. Legs pale yellnw; tarsal jotnts of all legs with sharp brown black annulations ; apical spur of middle tibiae in both sexes distinctly deve loped, sharp, towards tip blackish.- Length: body, 2.2 to 2.5 mm; wing, 2.4 mm Platypalpus flavicornis Meig. 63 (58 ) Large species. Middle fe mora ab out twice as staut as anterinr ones. Vein m conspicuously approaching towards r4+ s in apical section. - Fronsand vertex dusted silvery. Face dusted yellowish grey, in lateral aspect prominent. Ocellar and vertical bristles yellow. Antennae yellow, comparatively short, third join• almost twice as !Gng as deep a t its base. Arista brownish black, about 2.5 times as lang as third joint. Palpi !arge, about half as Ie-ng as proboscis, dusted whitish. Thorax dusted s ilvery, but sternopleura with shining black spot . All bristles and hairs of thorax yellow. Legs yellow, trochanters ventrally with black spD'ts. Apical spur of middle tibiae !arge a nd strong, its tip black. Tarsal joints with very indistinct pale brGwn annulations. Wings slightly yellowisll , veins yellow basally, yellowish brown a pically. Abdo-men black, shining, only base of each segment with nar r ow transverse dusted band. - Length: body, 4.0 to 5.3 mm; wing, 4.0 to 5.5 mm . Platypalpus major Zett. 64 (57) Only basal antennal joints yellow, sometimes base of third joint yellowish,
338 n5 ( 66) Each abdominal segment with a pair of !arge triangular grey dusted spO'ts . Large s,pecies dusted pale greyish. Third antennal joint black, ab out 2.0 to . 2.5 tim es as long as deep at its base, sho-rter than arista. Palpi rather !arge, oval, yel!ow. Tho·rax dusted yellowish grey, only anterior part of sternopleura with small black shining spot. Hairs and bristles of mesonotum pale brown. Legs yellow, trochanters with brownish black spots. Anterior tibiae stoutened, especially in male. Apical spur of middle tibiae sharp, black. Tarsal joints annulated brownish. - Length: body, 4.0 to 5.6 mm; wing, 4.0 to 4.7 mm . Platypalpus fasciatus Meig. 66 (65) Abdomen shining, its segments only with narrow dusted bands at their bases. 67 ( 70) Posthumerat bristle is developed, lang and conspicuous. Anterior and po·sterior tibiae with dark antero-do-rsal bristles. 68 ( 69) Male: Anterior tibiae slender; tarsal joints in both sexes distinctly annulated brownish black, last jo•int almost entirely black. Hypopygium is Ionger than in P. extrtcatus, upper anal papilla is ended by short apical hoo'k. Third antennal joint brownish black, 2.5 times as long as deep at its base; arista similarly coloured. Thorax dusted greyish, mesonotum with yellowish tinge. Acrostichal bristles Ionger than in P. extricatus. Apical spur of middle tibiae lang and sharp. Abdomen black, shining, only base of first and second abdominal segments with! narrow band dus ted greyish. - Length: body, 2.3 to 3.0 mm; wing, 3.0 3.7 mm . ·Platypalpus pallidiventris Meig. 69 [68) Male: Anterior tibiae strongly stoutened; only last tarsal joint black. In female tarsal joints annulated. Upper anal papilla is ended by strong and rather long hook. Third antennal joint brownish black, but with base mure or less coloured yellowish, about 2.5 tim'es as long as deep at its base; arista dark. Thorax dusted pale grey, mesonotum yellowish grey. Acrostichal bristles rather short. Legs yellow. A·pical spur of middle tibiae lang, sharp, darkened. In male abdo men black, shining, its first segment with narrow dusted band at its base; in female base of each segment with narrow transverse band of dust. Length: body, 2.7 to 3.0 mm; wing, 2.7 to 3.0 mm . Platypalpus extricatus Coll. 70 (67) Posthumerat bristle is not developed. Anterior and posterior tibiae antero· dorsal with only pale hairs. 71 (76) Apical spur of middle tibiae lang, strong, sharp, black towards apex. 72 [73 ) All tarsal joints distinctly annulated. - Very small species. Third antennal join·j almos t ·two and one half times as long as deep at its base, black or brownish black. Acro-stichal bristles biserial, rows very close together, almosT in same line, but distinctly divergent. ,Dors-ocentral bristles in one row at each side, but laterally to them several further bristles. Legs yellow, in male anterior coxae and bases of femora sometimes brownish black. Abdomen black, shining. first segment narrowly dusted on its base, following segments only indistinctly dusted. Hypopygium rather elongated, with long hairs. - Length: body, 1.5 to 2.5 mm; wing, 1.9 mm Platypalpus pictarsis Beck. 73 [72) Tarsal joints not annulated. Acrostichals distinctly biserial. 74 [75) Large species. Frons behind ocellar protuberance only slightly widened. Distance between vertical bristles only slightly !arger than width of ocellar protube :r·anee. Third antennal joint alnwst two and one half times as long as deep at its base, black but yellow basally; arista sligthly Ionger than this joint. Thorax densely ·dusted greyish, anterior part of sternopleura shining. Legs mostly yellow. Veins yellow, but brownish in apical -section. Abdomen black. shin'ing, base of its first two segments distinctly dusted greyish, foHowing three joints with narrow band of dust basally. - Length: body, 3.0 to 4.5 mm; wing, 4.2 mm Platypalpus laticinctus Walk. 75 [74) Very small species. Frons behind ocellar protuberance ·strongly widening. Distance between vertical bristles equalling double with of o-cellar protube rance. Third antennal jo-in1 entirely brownish bla·ck, light pubescent, about one and half times as lang as deep at its base; arista one and half times as lang as this joint. Thorax faintly dus ted greyish, sterno,pleura with !arger shining area. Legs yellow, last tarsal joints and apical halves of preceding ones brownish black. Veins yellow. Abdomen brown to brownish black, 'ünly bases o-f first two segments narro-wly dusted greyish. Length: body, 1.5 to 2.0 mm; wing, 2.0 mm Platypalpus calceatus Meig.
339 76 [ 71] Apical spur of middle tibiae blunt. 77 [78 ] Extremely small species. Clypeus dusted silvery. Antennae rather sho-rt, third joint bro-wnish black, hardly 1.5 times as lung as deep at its base; arista black, twice as long as third joint. Mesono•tum moderately dusted dark greyish, pleurae with lighter dusting. Acrostichal bristles biserial; these and dorso centrals short, inconspicuo.us, yellowish brown. Legs brightly yellow; tarsal joints annulated brownish black, especially in anterior pair. Apical spur of middle tibiae short. Wings hyaline; veins yellow basally, yellnwish brown apically. Abdomen black, basal half of each segment dusted greyish Iaterally. - Length: body, 1.2 to 1.8 mm; wing, 1.7 to 2.0 mm . . Platypalpus articulatus Macq. 78 [77} Moderate'ly !arge species. Clypeus black, shirring. Antennae rather Iong, third ju~nt brown to brownish black, 3.0 tu 3.5 times a'S long as deep at its base; arista brownish black, as long as that joint. Mesonotum moderately dusted brownish grey so that the ground colour is visible, pleurae dusted lighter grey. Acrostical bristles in two to three irregular rows; these and dorsocentral bristles very short, yellow. Legs usually yellow, but anterior coxae and fe mora, sometimes also middle femora and a.pical parts of posteri-or femora often brownish; tarsal joints annulated black, oes pecially distinctly on anterior pair of Iegs. Apical spur of middle tibiae long, with short fi"ne protuberance on Hs tup. Wings yelluwish; veins yellow basally and along costal vein, brown apically. Abdomen black, shining, only first two terga basally with narrow transverse bands of dust. - Length: body, 2.2 to 3.0 mm; wing, 3.0 mm . . ·Platypalpus annulipes Meig.
3. REVIEW OF CZECHOSLOVAK SPECIES
Platypalpus Macquart, 1827
Platypalpus Macquart, 1827, Mem. Soc. Sei. Lille, 1827: 92 Phoroxypha Rondani, 1856, Dipt .Ital. Prodr., 1: 146 Cleptodromia Corti, 1907, Wein. ent. Ztg., 26: 101 Tachydromia, authors, nec Meigen Type species: Musca cursitans Fabricius, 1775, for Platypalpus; Ta cliydromia Zongicornis Meigen, 1822, for Phoroxypha; Cl·eptodromia Zon gimana Corti, 1907, for Cleptodromia.
Platypalpus agilellus ( Collin, 1926] Tachydromia agilella Collin, 1926, Ent. mon. Mag., 42: 153 Zoogeographie distribution: Czechoslovakia, England ( Collin, 1961]. Occurrence in Czechoslovakia: Lit: Boh: As ( Collin, 1926).
Platypalpus agilis ( Meigen, 1822) Tachydromia agilis Moeigen, 1822, Syst. Beschr., 3: 80 · Tachydromia curticornis Zetterstedt, 1849, Dipt. Scand., 8: 3004 Platypalpus dubius Walker, 1851, Ins. Brit., 1: 132 Fig. 17, 49. In the materials examined by myself most females have their legs darker than males, as is the general rule. The males and females taken at Rovensko, however, have pale legs while the males and females from Brno have their legs darkened. Prey: Platypalpus longicornis ( Collin, 1961); Metriocnemus sp. -
340 Chironomidae, ?Empis chiroptera Meigen and Bicellaria sp. - Empi didae ( Laurence, 1955). Zoogeographie distribution: Denmark, Finland ( Lundbeck, 1910) ; Sweden ( Lundbeck, 1910; Engel, 1939) ; Belgium ( Goetghebuer, 1931) ; Austria, Hungary, Germany (Engel, 1939); Great Britain ( Collin, 1961); Czeehoslovakia. Occurrence in Czechoslovakia: Mat: Boh: Peat-moss Soos nr. Cheb, 19. v. 1961, 2 <1 2 !i! (ZZ]. - Praha-Krc, 24. vi. 1955, 1 !i! (Macek, MMB]. - Praha-Ruzyne, 18. v. 1958, 1., 4. and 16. vi. 1962, 2
Platypalpus albicornis ( Zetterstedt, 1842) Tachydromia albicornis Zetterstedt, 1842, Dipt. Scand., 1 : 279 Engel ( 1939) synonymizes this species with Plat.ypalpus flavicor nis, but Lundbeek ( 1910) clearly distinguishes the two, and Collin ( 1961] presents perfect characters that these s.pecies differ in. The latter author also gives a detailed explanation of the nomenclature of this speeies. Zoogeographie distribution: Denmark (Lundbeck, 1910); Great Bri tain, Franee ( Collin 1961) ; Czeehoslovakia. Occurrence in Czecho-slovakia: Mat: Mor: Cernovice, 16. v. 1907, 1
Platypalpus albiseta (Panzer, 1806) Tachydromia albiseta Panzer, 1806, Fauna Germ., 103: 17 Tachydromia castanipes Meigen, 1822, Syst. Beschr., 3: 79 Tachydromia vivida Meigen, 1838, Syst. Beschr., 7: 97 Tachydromia fuscimana Zetterstedt, 1842, Dipt. Scand., 1: 292 Fig. 1. Zoogeographie distribution: Denmark, Finland, Italy, North Africa (Lundbeek, 1910); Belgium ( Goetghebuer, 1931); Austria, Hungary, Ger many (Engel, 1939); Sweden [Engel, 1939;. Ardö, 1957); Great Britain ( Collin, 1961) ; Czeehoslovakia. Occurrence in Czechosl>a-vakia: Mat: Boh: . Zbüch nr. Plze>ii, 21. viii. 1962, 1 !i! [ZZ) . - Chyii nr. Praha, 7. ix. 1957, sweeping pond bank, 1 <1 (ZZ]. - ,Praha-'Ruzyne, 26. vii. 1962, 1 !i! [J. Kavalko-va, coll. Zuska.]. · Mm: Vranov nad Dyjf, vii. 1910, 1 !i! ( MMB). - Vrbno pod ·Pradedem, 10. viii. 1960, 1 !i! [ZZ]. - Smrk [apparently Moravia, not specified], 17. viii. 1909, 1 !i! [K. Landrock, MMB].- Brno, 8. vi. 1904, 1 !i! ("Tachydrom. glabrata M.", MMB].
Platypalpus annulatus (Fallen, 1815) Tachydromia annulata Fallen, 1815, Dipt. Suec., Emp.: 7 Tachydromia fascipes Meigen,. 1822, Syst. Beschr., 3: 78 Tachydromia fulvipes Meigen, 1822, Syst. Beschr., 3: 78 As to the synonymy of Platypalpus fascipes and Platypalpus fulvipes, I must agre with Collin's ( 1961) eonclusion. lt is apparent that Engel
341 Wing: Fig. 1, Platypalpus albiseta. - Fig. 2, P. annulipes. - Fig. 3, P. binderi. - .So.Jid lines equal 0.5 mm each,
342 ( 1939) was very perplexed with the synonymy of these names. Platy palpus fuluipes Meigen sensu Engel ( 1939, p. 49] is obviously an un known species, which is described in the present paper on page 344. Zoogeographie distribution: Sweden ( Ardö, 1957], Great Britain ( Collin, 1961), Czechoslovakia. Occurrence in Czechoslovakia: Mat: Hoh: Bohemia, without further data, 7 ,] 2 'i' [A. Binder, NMP). - Plzeii, 25. vi. 1957, 1 ,,] (ZZ).- Dobfichovice, 1 ~ [NMP).- Praha Podbaba, v., 1 'i' ["Tachydromia flavipes Fb."; A. Vimmer, NMP). - Celäkovice, 18. v. 1957, sweeping meadow at Labe River, 1 ,] (ZZ). - Rovensko-, vi., 1 ,] [A. Vimmer, NMP). S!ov: Krälovsky ChYmec, 10. vi. 1960, 1 r! [ ZZ) .
Cerci and sclerite beneath those: Fig. 4, Platypalpus annulipes. - Fig. 5, P. binderi. - Fig. 6, P. articulatus. - Fig. 7, P. extricatus. - Fig. 8, P. ciliaris. - Solid lines equal 0.1 mm each.
343 Platypalpus annulipes ( Meigen, 1822) Tachydromia annulipes Meigen, 1822, Syst. Beschr., 3: 77 Tachydromia infuscata Meigen, 1822, Syst. Beschr., 3: 84 Tachydromia coxata Zetterstedt, 1842, Dipt. Scand., 1: 281 Figs. 2, 4, 9, 16, 50. Zoogeographie distribution: Sweden, Denmark (Lundbeek, 1910); Austria, Hungary (Engel, 1939) ; Great Britain ( Collin, 1961) ; Czeehoslo vakia. · Occurrence in Czechoslovakia: Mat: Boh: Praha-Ruzyne, 6. vi. 1958, 1 !f; 30. v. to l5. vi. 1963, 32 r!, 12 !f ; 23. vi. 1964, 1 !f ( ZZ). Mor: Hrusovany nad JeviSovkou, 22. v. 1964, 1 !f (ZZ]. - Brno, 2 r! ("Tachy· dromia annulipes", MMB ).
Platypalpus articulatus Macquart, 1827 Platypalpus articulatus Macquart, 1827, Mem. Soc. Sei. Lilie, 1827: 98 Tachydromia maculimana Zetterstedt, 1842, Dipt. Scand., 1: 284 Figs. 6, 10. Zoogeographie distribution: Sweden, Finland, Denmark (Lundbeck, 1910) ; Austria (Engel, 1939; Collin, 1961) ; Great Britain ( Collin, 1961) ; Czechoslovakia. Occurrence in Czechoslovakia : Mat: Boh: Praha-Ruzyne, 16. vi. to 1. viii. 1962, 3 r! 7 !f; 14. vi. 1963, 1 r! (ZZ] . Mor: Brusperk nr. Frydek-Mls tek, 8. ix. 1958, 1 !f (J. Beränkovä, coll. Zuska]. - Troubky-Zär!Cl nr. Kromei'lz, 3. vii. 1958, 1 r! 3 !f (P. Lauterer, coll. Zuska].
Platypalpus binderi, n. sp. Figs. 3, 5, 11 ,. H. ö ~. Rather large, black speeies. Head: Frons about as wide as seeond antennal joint deep, dusted greyish. Face narrower, with lighter dust just as in clypeus. Ocellar bristles very short, verticals clearly visible, moderately long; all these bristles black. Other bristles on upper part of occiput with greyish dust short, brown; those on lower part langer, silverish white. Antennae enti rely black, third joint about twice as long as its base deep. Arista 1.5 to 2 times as long as third antennal joint. Palpi yellowish orange, modera tely long, with silvery dust and possessing one long and one short brow nish yellow bristle on their tips. Thorax: Mesonotum, scutellum, metanotum and pleurae dusted greyish, only sternopleura polished, black. Chetotaxy: two rows of acrostichals, two irregular rows of dorsocentrals, their last two pairs Ionger and stronger, laterally between dorsocentrals and notopleura scatte·red small bristles. Notapleura with one long and two shorter brist les ( one of these situated close to humeral callus) and several other shorter hairs. Humeral callus with one long bowed and several shorter bristles. Postalar callus with one long bowed and one shorter bristles. Seutellum bearing one pair of long apical and one pair of shorter sub" apical bristles. All Ionger and stronger bristles yellowish brown, short and fine ones yellow.
344 Legs varying in colouration, especially in that of middle femora. Legs of female usually lighter. Anterior pair: coxae brownish black at the base, their anterior plain with very dense silvery dust and long white pilosity. Femora thickened, posterodorsally with longitudinal brownish black spot of variable width. Hairs of these femora very
9
\ \ \ \ \ I 11
\ '"---
Le ft and right part of epandrium: Fig. 9, Platypalpus annulipes. - Fig. 10, P. articu latus. - Fig. 11, P. binderi: - Solid lines equal 0.1 mm each.
345 lang, pale, especially in ventral, anterior and posterior rows. Tibiae bright yellow, their base a little narrower than darker apex. Anterior row with several conspicuous, s.tronger brownish black bristle.s in both sexes, in males also bearing ventral and posteroventral rows of long whitish yellow hairs ( almost twice as lang as tibia deep], posteroventral row continuing on first tarsal joint. In female these rows of bristles shorter, hardly as lang as tibia deep, without continuation on firs.t tarsal joint. - Middle pair: Whole coxae black, dusted greyish and with lang pilosity anteriorly. Fernara strongly thickened, at le•ast twice as deep as those of anterior pair. In male femora usually wholly brownish black except for apex, but in some specimens colouration lighter, similar to females. In female femora with posteroventral brownish black stripe and in apical half with rounded brownish black spot. Ventral side bear· 1ng usual double row of black short sharp spines, and posteroventral row of lang whitish yellow hairs; those of anteroventral row shorter. In apical third one stronger black bristle. Knees usually black. Tibiae bright yellow, bearing only one row of ventral black short spines; apical spur lang, sharp, almost entirely black, with two extremely short hairs on its top. - Pos.terior pair: Coxae only very slightly dusted greyish below apex and only their apical half with sparse whitish pilosity. Fernara slender, yellow, in dark specimens with brownish black anterior spot behind middle. Whitish yellow hairs more conspicuous in posteroventral and anteroventral row. Tibiae yellow, darkened in apical part. In ante rior row several brownish black stronger bristles.. In male anteroventral and posternventral rows of hairs sp arser and Ionger in basal half while in apical half those more crowded and shorter. This difference less con spicuous .in female. Tarsal joints of all legs annulated brownish blac'k. In dark specimens tarsal joints almost black, especially those of ante rior pair. Ventral side of all metatarsi bearing a row of hairs. All tarsal joints slender. Wings very slightly tinged brownish in apical part. Costa yellowish brown towards end of rt, this almost entirely of same colour. Further veins yellowish brown only in basal part of wing, towards end becorn ing brownish black. Prefurca short, one half as lang as distance bet ween its base and humeral vein. Ve.ins r5 and fit diverging rather strongly in their basal parts, practically parallel in middle ones and strongly converging in their apical parts, but apical section of mt si nuate and therefore top portion of m1 parallel with that of r5. ·First basal cell only slightly shorter than second. Costal bristles brown, be fore end of r1 1onger and sparser, behind it shorter and more crowded. Squamae white with lang white fringes. Halterae yellow with brownish base. Abdomen: Each segment with large triangle-shaped area dusted greyish laterally. Sternites entirely covered with dust of same colour. Abdomen with sparse pale yellow pilosity. In male hairs Ionger at pos.te rior edge of last segment. Hypopygium moderately large, shining. In female seventh segment with yellowish grey dust in apical half, shining at base, eighth segment shining at sides. Fernale cerci slende·r, with exceedingly short pubescence, almost as lang as eighth segment. Length: body, 3.4 to 3.9 mm; wing, 3.1 to 3.5 mm. Type s.e
Platypalpus bohemicus, n. sp. Figs. 18, 48 . .!j!. Moderately large yellow species. Head black. Frons rather wide, relatively short, with yellowish grey dust; face and clypeus similarly dusted, but face only half as wide as frons. Ocellar bristles easily visible, brownish black, about as lang as vertical bristles. The>Se forming two pairs, inner convergent, outer di vergent. Hairs on upper part of occiput yellowish brown, on lower part pale, silvery. Both basal joints of antennae yellow (third joint broken in both specimens). Palpi short, yellow, covered with silvery dust, bear ing lang brown bristle at their top. Proboscis in first examined female shorter, in second Ionger than head. Thorax: Mesonotum yellow with broad longitudinal median black stripe reaching edge of scutellum, where it is broadest. At this point mesonotum shining. Except for this area mesonotum just as scutellum and pleurae with exception of sternopleura fine dusted greyish. Chetotaxy: Whole mesonotum covered with short pale yellow pilosity, acrostichal and dorsocentral bristles not differentiated, only one prescutellar dorso central pair of conspicuous lang brownish black bristles developed. Hu meral bristle slightly langer than those, moderately bowed, postalar bristle of same length; both these bristles and lower of two notopleurals of same colour as humeral, upper natopleural bristle shorter, slightly paler. Scutellum with one pair öf lang brownish black apical bristles and one pair of subapical bristles of same cölour but only half as lang as these. Legs yellow, only trochanters with black longitudinal spots. Anterior pair: Anterior surface of coxae possessing silvery. dust and short greyish white pilosity. Fernara moderately thickened, with short pale yellow pilosity and with ring of six to seven rather lang brownish black bristles at their tip. Tibiae in all parts of their length rather thick (their depth equalling that of femora at their apex), bearing seveoral rows of short pale yellow hairs and one langer yellowish brown posteroventral bristle. Apex of tibia with pale yellow posteroventral small comb. Tarsal joints
2 347 17
Middle leg: Fig. 12, Platypalpus nigritarsis. - Fig. 13, P. ruficornis. - Fig. 14, P. bin · deri. - Fig. 15, P. interstinctus. - Fig. 16, P. annulipes. - Fig. 17, P. agilis. ~ Solid line3 equal 0.2 mm each. yellow. First joint long, almost of same length as following joints to gether, bearing one long (half as long as first joint) yellowish brown ventral bristle at its base, otherwise with dense short yellow pilosity as other joints. Last three joints conspicuously dilated. Tip of second to fourth joints with several langer yellowish brown bristles. Claws long, black, curved; pulvilli silvery grey. - Middle pair: Anterior surface of coxae with several long yellowish brown bristles. Femora strongly thickened, about three times as deep as those of anterior pair, with short yellow pilosity. Ventral rows of black spines on femora beginning as far as behind end of first fourth of length of those; spines composing beginning of these rows shorter, proximally sparse, apically denser than spines in hind row. Fernara possessing also posteroventral row of nine to ten long yellowish brdwn bristles, length of which equalling less than
348 Wing: Fig. 18, Platypalpus bohemicus . - Fig. 19, P. ciliaris. - Fig. 20, T. infectus. - Solid lines equal 0.5 mm each.
2* 349 half width of femur. Five to six Ionger brownish black bristles enclosing tips of femora. Tibiae slightly bowed, short, hardly three-fourth times as long as femora, possessing very tiny rounded protuberance on their apex. Tarsal joints yellow, not thickened, with short yellow pilosity, this more conspicuous ventrally. Hairs on base of first joint ventrally Ionger than those on apical half. Claws curved, shorter than those of anterior pair. Pulvilli of the same size and colour. - Posterior pair: Coxae with short yellow pilosity anteriorly. Fernara rather thick, with their middle and distal part of equal depth, onl y their base thinner, pilo sity short and sparse, yellowish grey. Tip of femora surrounded by about five Ionger conspicuous yellowish brown bristles. Tibiae about as thick as those of anterior pair, with short yellowish grey pilosity, but hairs in apical half denser. Before tip, tibia bearing one rather lang, con spicuous yellowish brown dorsal bristle. Ventrally tibia bearing apical comb being larger than that of ante·rior pair of legs. Tarsal joints slen der, with yellowish grey pilosity. First joint almost as lang as all follow ing joints together, its ventral side more densely pilose. Claws and pul villi similar to those of middle pair of legs. Wings long, hyaline. Veins brown, in halotype subhyaline, yellowish. Costal vein rather strong, especi 350 Platypalpus calcaratus ( Collin, 1926) Tachydromia calcarata Collin, 1926, Ent. man. Mag., 42: 155 Zoogeographie distribution: Italy (Engel, 1939], Great Britain ( Collin 1961), Czeehoslovakia. Oc·currence in Cz-echoslovakia: Mat: Boh: Kundratice nr. Litomei'ice, 18. vi. 1958, 1 id' (ZZ] . Platypalpus calceatus ( Meigen, 1822 ) Tachydromia calceata Meigen, 1822, Syst. Beschr., 3: 87 Zoogeographie distribution: Denmark (Lundbeek, 1910], Great Bri tain ( Collin, 1961), CzeehosJovakia. Occurrence in Czecho·slovakia: Mat: Boh: Zbikh nr. Plzeii, 21. viii. 1962, 1 ~ (ZZ]. - Brdy Mts., Kytfn, vii. 1959, 1 ~ (Macek, MMB].- Nasavrky nr. Tabor, 24. viii. 1959. 1 ~ (ZZ]. - Nove Mesto nad Metuji, vii. 1955, 1 ~ (Macel,, MMB). - Velky Vi'estov. 8. vii. 1957, 1 ~ (Z. Boucek, coll. Zuska]. Mor: Piiovice nr. Litovel, 9. vii. 1958, 1 d' (P. Lauterer, coll. Zuska). Platypalpus candicans (Fallen, 1815) Tachydromia candicans Fallen, 1815, Dipt. Suec., Emp.: 10 Tachydromia ventralis Meigen, 1822, Syst. Beschr., 3: 85 Platypalpus varius Walker, 1851, Ins. Brit., 1: 126 ? Tachydromia oedicnema Strobl, 1898, Mitt. Naturw. Ver. Steierm., 34: 211 The colour of the abdomen varies greatly. In the material investi gated by me both the two types of specimens, with black abdomen and those with yellowish abdomen, were present; and among them inter medial individuals with paler sterna and darker terga. It is not im· probable, however, that Platypalpus oedicnemus, a species characterized by abdomen partially yellowish, is a distinct spec'ies. Plat;ypalpus candicans is a speeies in which males occur extremely rarely. Iil Vimmer collection deposited at NMP is one male with abdo men conspicuously yellow and with the blaek hypopygium. Unfortuna tely, it is not pos~>ible to compare its hypopygium with that of any other specimen and no figure is present in the literature. Zoogeographie distribution: Denmark, Finland, Italy ( Lundbeck, 1910); Sweden (Lundbeck, 1910; Ardö, 1957]; Austria, Albania (Engel, 1939); Holland, Belgium ( Goetghebuer, 1931]; Great Britain ( Collin, 1961), CzeehosJovakia. Occurrence in Czechoslovakia: Mat: Boh: >Prameny nr. Cista, Slavkovsky les, 9. vii. 1962, 4 · ~ (F. Kodys, coll. Zuska]. - Velesin nr. Cesky Krumlov, vii., 1 d' ("Tachydro· mia varius Wlk. det. Vimmer"; A. Vimmer, NNLP]. - Tabor, vi., 2 ~ ["Tachydromia cursitans Fb."; A. Vimmer, NMP].- Blatna, 10. vi. 1958, 1 ~ [ZZ).- Chuchle nr. Praha, 19. vi.., 1 ~ ["Tachydromia varia Wlk., det. Vimmer"; A. Vimmer, NMP]. - Praha-Ru zyne, 14. vi. 1963, 1 ~ [ZZ]. - Modi'any nr. Praha, swept from vegetation along forest path, 8. vi. 1957, 1 ~ [ZZ]. - TiSice nr. Melnik, 29. v. 1964, 4 ~ [F. Kodys, coll. Zuska). - Krkonose Mts., Vitkovice, from hay, 10. viii. 1957, 1 ~ [ZZ]. - Krkonose Hts., Horni Albei'ice, 9. viii. 1958, 1 ~ [P. Peciua, coll Zuska]. - Nnvy Bydzuv, vi., 2 ~ ["Tachy· dromia cursitans Fb., det. Vimmer"; A. Vimmer, NMP] . - Brdy Mts., Kytin, viii. 1957, 1 ~ (Macek, MMB].- Hnrusice nr. Tabo•r, sweeping vegetation on bank of pond, 27. v. 1964, 5 ~ [ZZ). 351 Mor: Cernovice nr. Brno, 20. v., 1 ~; 16. v. 1907, 1 ~; 4. vi. 1907, 2 !? (MMB ). - Dolni Vestonice, 22. v. 1959, 2 !? [R. Rozkosny, MMB). - Sti'elice nr. Brno, 25. v. 1957, 1 !? [MMB). - Olomouc-Cernovir, 1. vii. 1958, 7 !? [P. Lauterer, coU. Zuska).- :Pi'erov, 8. vii. 1957, 1 !? [P . .Pecina, coll. Zuska ). - Strachotin, 10. v. 1908, 1 !? ; 6. vi. 19018, 1 !? ; 8. vi. 1908, 2 !? [K. Landrock, MMB). - Vrbno nr. Bruntal, 30. vi. 1960, 6 !? (ZZ). - Ti'estina nr. Mohelnice, 9. vii. 1958, 1 !? (iP. Lautemr, coll. Zuska). Platypalpus ciliaris (Fallen, 1816) Tachydromia ciliaris Fallen, 1816, Dipt. Suec., Emp.: 33 Platypalpus compungens Walker, 1851, Ins. Brit., 1: 128 Figs. 8, 19, 26. Zoogeographie distribution: Denmark, Finland ( Lundbeek, 1910) ; Sweden ( Lundbeek, 1910; Ringdahl, 1941) ; Belgium ( Goetghebuer, 1931) ; Austria (Lundbeek, 1910; Engel, 1939); Czeehoslovakia (Engel, 1939); Great Britain ( Collin, 1961). Occurrence in Czechoslovakia: Mat: Boh: Pizen, 25. vi. 1957, 2 !? ( ZZ). - Brdy Mts., Kytin, viii. 1957, 1 <1 3 !? ; vii. 1959, 2 rJ (Macek, MMB). - Praha-Ruzyne, Hvezda, 13. vi. 1958, 1 !? [ ZZ). - Krkon ose Mts., Rezek, sweeping vegetation in fmest glade, 4. viii. 1957, 1 !? (ZZ). - IKrkonose Mts., Vii'kovice, 27. vii. 1961, 1 rJ (ZZ) . - Krkonose Mts., Horn! Albei'ice,, 26. vii. 1958, 1 rJ 1 !? [1P. Pecina, coll. Zuska). - Nove Mesto nad Metu ji, vii. 1957, 1 <1 ( Macek, MMB). Mm.: Beskydy Mts., Kycera, 10. vii. 1882, 1 <1 (A. Het'S'chko, NMP). - Pradect, 1 !? [K. Czizek, MMB).- Dolni Vestonice, 30. vi. 1959, 1 !? (R. Rozkosny, MMB) . Platypalpus cursitans ( Fabrieius, 1775) Musca cursitans Fabricius, 1775, Svst. entom.: 782 Zoogeographie distribution: Denmark, Sweden ( Lundbeek, 1910, as ,Jachydromia fasctata"]; Finland (Tuomikoski, 1935; Frey, 1943); Bel gium (Goetghebuer, 1931); Great Britain (Collin 1961); Czeehoslovakia. Occur re·nce in Czechos:ovakia: Mat: Bo'h: Praha-Stvanice, 2 !? ( "Tachydromia cur· sitans, det. Vimmer ·'; Pr ach, NMP]. - Celakovice, sweeping meadow vegetation on bank of Labe River, 18. v. 1957, 1 !? ( ZZ). - Novy Bydzov, vi., 1 !? ( "Tachydromia cur s'tans Fb., det. Vimmer"; A. Vimmer, NM:P). - Cheb, 16. v. 1882, 1 !? (Grad!, NMP). ~ Horusice nr. Tabor, sweeping vegetation on pond bank, 27. v. 1964, 3 !? ( ZZ J. Mor: Cernovice nr. Brno, 16. v. 1907, 1 !? [MMB ). Slov: Trencin, 2 damagecL specimens ( NMP). Platypalpus extricatus ( Collin, 1926) Figs. 7, 27. Ta :· hydromia extricata Collin, 1926, Ent. man. Mag., 42: 188 ? Empis flavipes Fabricius, 1794, Ent. Syst., 4: 406 As to the synonymy of Platypalpus flauipes, refer to the note under Platypalpus pallidiventris. Zoogeogra phie distribution: Great Britain (Engel, 1939; Collin, 1961), Czeehoslovakia. Occurrence in Cz echoslovakia: Mat: Boh: peat-moss Soos nr. Cheb, 16. viii. 1962, 1 <1 [F. Kodys, co.JJ. Zuska). - Pavlov nr. Praha, 7. vii. 1963, 2 !? (ZZ).- J'raha-Ruzyne, 352 vii. 1961, 1 ~; L vi. to 1. viii. 1962, 1 rJ 12 ~ (ZZ] . - Vsetaty, 25. v. 1957,4 ~ (ZZ). Tisice nr. Melnik, 29. v. 1964, 2 rJ 5 ~ (F. Kodys, cool!. Zuska ). - Krkonose Mts., Zaly, vii., 1 ~ ("Tachydrom~a bicol or Fb., det. Vimmer"'; NMP) . Mm: Brno, 20. v. 1907, 1 rJ (MMB). - Cerno,vice nr. Brno, 4. to 5. vi. 1907, 2 ~ (MMB).- 'Piiovice nr. Lito·vel, 9. vii. 1958, 1 ~ (P. Lauterer, coll. Zuska). - Olomouc· Cernovir, 7. vi. 1958, 1 rJ; L vii. 1958, 1 rJ 3 ~ (P. Lauterer, coll. Zuska). Slov: Trenc!n, 1 ~ (NMP). 23 Cerci and sclerit'e beneath those: Fig. 21, Platypalpus fasciatus. - Fig. 22, P. mfectus. - Fig. 23, P. f l auicornis. - Fig. 24, P. luteipes. - Fig. 25, P. minutus. - Solid lines equal 0.1 mm each. 353 Platypalpus fasciatus ( Meigen, 1822) Tachydromia fasciata Meigen, 1822, Syst. Beschr., 3: 86 Figs. 21, 28. My specimens agree fully with Collin's ( 1961) description except for mesonotal hairs and bristles, which are paler, thorax not "with many of the hairs and bristles dark brownish", as Collin writes. Zoogeographie distribution: Belgium ( Goetghebuer, 1931) ; Albania, Germany (Engel, 1939, as "Coryneta cursitans"), Great Britain ( Collin, 1961), Czechoslovakia. Occurrence in Czechoslovakia: Mat: Boh: Teplicka nr. Karlovy Vary, 8. vi. 1964, 2 <1 ( F. Kodys, coH. Zuska ). - Svadov nr. Üst! nad Labern, 1. vi. 1943, 1 ~ ( "Platypalpus fasciatus Mg."); 23. v. 1944, 1 ~ ( "Platypalpus cursitans F."]; 14. v. 1945, 1 <1 1 1 ~ ; 26. v. 1945, 2 <1 (all Pawlik, NMIP] . - Hodkovice nr . .Praha, 5. vi., 1 ~ ("T. maior Zt."; A. Vimmer, NMP].- Oparno nr. Litomei'ice, 26. v. 1964, 3 <1 3 ~ (I. Novak, coll. Zuska]. - Tisice nr. Melnik, 15. v. 1964, 1 <1 1 ~; 29. v. 1964, 1 ~ (F. Kodys, coll. Zuska). -· Bezno nr. Mlada Boleslav, 24. v. 1964, 1 <1 (I. Nüvak, coll. Zuska) . -Kr konose Mts., Horn! Albei'ice, vi. 1957, 1 ~ [fP . .Pecina, coll. Zuska 1. Mor: Loucna nad Desnoru, 25. v. 1956, 1 <1 1 ~ (J. Stehlik, MMB]. -- Bilüvice nad Svitavou, 23. v. 1908, 1 ~ (MMB ]. - Grün nr. Javornik, 9. vi. 1961, 1 ~ [.R. Rozkosny. MMB). S!ov: Trencin, 1 damaged specimen (NMP]. Platypalpus flavicornis ( Meigen, 1822 J Tachydromia flavicornis Meigen, 1822, Syst. Beschr., 3: 83 Figs. 23, 29. Zoogeographie distribution: Denmark, France, Northern Africa (Lund beck, 1910); Austria, Hungary, Germany, Spain, Greece, Tunisia (Engel, 1939) ; Great Britain ( Collin, 1961) ; Czechoslovakia. Occurrence in Czechoslovakia: Mat: Boh: Teplic'ka nr. Karlovy Vary, 8. vi. 1964. 1 <1 [F. Kodys, coil. Zuska]. - Line nr. Pizen, 31. viii. 1962, 1 ~ (ZZ]. - Praha-Ruzyne, 12. vii. 1962, 1 <1 1 ~; 2. vi. 1964, 1 ~ (ZZ) . - Libus nr. tPraha, poultry-farm, 17. vii. 1963, 1 ~ (P. Lastovka, in coll. Central Research Institute of Food Industry, Praha]. Platypalpus infectus Collin, 1926 Tachydromia infecta Collin, 1926, Ent. mon. Mag., 42: 157 Figs. 20, 22, 30. Redescription: <1 ~. Moderately large black species. Head: Frons and occiput dusted greyish. Face and clypeus with lighter greyish silvery dust. Frons about as wide as base of third anten· nal joint, as far as behind ocellar protuberance exceedingly widened. Face narrower than frons. Yellow ocellar bristles clearly visible, almost as lang as vertical bristles. These slightly darker, developed in one pair, only very faintly distant from each other. Upper part of occiput with short yellowish brown hairs, lower with Ionger whitish silvery ones. Antennae rather lang, entirely black. Third joint 2.5 to 3 times as lang as wide at its base, arista slightly longeor than this joint. Palpi modera tely large, egg-shaped, brownish black, with silvery dust and two lang pale yellow bristles at their top. Thorax: Mesonotum, scutellum and metanotum dusted greyish. Also pleurae wholly dusted except for entirely black shining sternopleura. 354 I I I ' , ' 28 ~I I Left and right part of epandrium: Fig. 26, Platypalpus lciliaris. - Fig. 27, P. extricatus. - Fig. 28, P. fas ciatus. - Solid lines a ual 0.1 mm each. Chetotaxy: Bristles and bairs of mesonotum and scutellum light, yellow ish brown. Acrostichal bristle6 relatively sho1:t, in two rows. Dorsocen trals forming two irregular rows at eadh side, last but one and especially last pair strongly prolonged. Laterally to thEl!se rows several scattered short hairs located, extending almost towat ds two lang natopleural 355 l bristles. Between these and humeral callus another long bristle inserted. Humeral calli bearing one very lang curved lmmeral bristle and group of very short bristles close to it. Postalar calli each with one long bristle. Scutellum possessing one pair of lang apical and pair of short subapical bristles. Legs with rather lang pale yellow pilosity, yellow. Tarsal joints with indefinite brownish black annulations, sometimes darkened, last joint entirely black. Also knees of middle legs black. Anterior pair: Coxae anteriorly with dense silvery dust and long white pilosity. Fernara thicke ned, posteroventrally and anteroventrally with rows of lang bristles. Tibiae slender in both sexes, anteriorly with several stronger and darker blistles, posteriorly with bristles of same length, but weaker and paler. Tarsal joints slender. - Middle pair: Coxae anteriorly with lang yellow pilosity. Femora strongly thickened, almost twice as much as those of anterior pair, ventrally bearing double row of black sharp spines, pos teroventrally with row of lang yellow bristles, ante,roventrally with row of slightly shorter bristles. Anteroventrally in apical third of femora three to four darker and stronger conspicuous bristles. Tibiae ventrally bearing only one row of short black spines. Apical spur almost wholly black, strong, lang and sharp, with one hair at its top. - Posterior pair: Coxae bearing only several langer hairs. . Fernara slender, anteroventrally with row of conspicuous stronger bristles, posteroventrally with row of finer bristles of same length. Tibiae very densely pilose, especially ventrally, anteriorly bearing several more conspicuous langer brownish black bristles. Wings hyaline, veins yellowish brown. Costal hairs yellowish brown, before end of r1 langer and sparser, thence towards end of r 5 denser and shorter. Prefurca shorter than distance between its base and hume ral vein. Veins r 5 and m1 parallel, but in apical part r 5 diverging. Anterior basal cell distinctly shorter than posterior one. Halterae yellowish white with darker yellowish brown stem. Abdomen black , shining, with scattered white pilosity. First two seg ments with n arrow greyish dusted areas at their sides basally. In females seventh segment wholly dusted yellowish grey, eighth segment aLso dusted but with one pair of shining spots at sides ante,riorly. Cerci short. Hypopygium moderately large, left part of epandrium bearing lang pale yellow fringes at its left edge, especially in basal part. Collin ( 1926, 1961 ) describes this fly only very briefly, therefore a detailed desciption is given here. Zoogeographie distribution: Great Britain (Engel, 1939; Collin, 1961], Czechoslovakia. Occurrence in Czechoslovakia: Mat: Boh: Radotin nr. Praha, swept from alfalfa, 28 . viii. 195&, '1 !f [J. m abola, co.Jl . Zu ska ). - Praha·Ruzyne, 14. vi. 1963, 2 !f [ZZ). - Horomei'ice nr. ,Praha, vi. 1964, 1 d 2 !f (J. Berankova, coll. Zuska ). - Novy Bydzov, v., 1 r1 ["Platypalpus cothurnat. Maq."; A. Vimmer, NMP ). Mor: Slatinice nr. O!omouc, 9. ix. 1951, 1 d 1 !f; 16. ix . 1951, 1 !f (J. Pa!iisek, MMB) . - Hrusovany nad Jevisovkou n r. Znojmo, 22. v. 1964, 1 r1 ( ZZ ). - Strachotfn nr. Bi'eclav, 1 !f [K. Landrock, MMB) . 356 Platypalpus ingenus (Collin, 1926) Tachydromia ingenua Collin, 1926, Ent. mon. Mag., 42: 155 The previously unknown female is very similar to the male. Abdo men shining blaek with pale pubeseenee, ending blunt eonieally, last three segments overlapping teleseopieally. Zoogeographie distribution: Great Britain (Collin, 1961 - two males only], Czeehoslovakia. Occurrence in 1Czechoslovakia: Mat: Boh: .Praha-Ruzyne, 26. vii. 1962', 1 ~ (J. Ka valkovä, coiL Zuska). Mnr: Cernovice nr. Brno, 30. vi. 1905, 1 c1 ["Tachydr. pallipes FlL", MMB). Platypalpus interstinctus ( Collin, 1926) Tachydromia interstincta Collin, 1926, Ent. mon. Mag., 42: 158 Figs. 15, 51. Zoogeographie distribution: Great Britain ( Collin, 1961), Czeehoslo vakia. Occurrence in Czechoslovakia: Mat: Boh: Praha-Stromovka, 1 ~ ( NMP ). - Bo hemia, without localityl 1 ir:J (A. Binder, NMP]. - Praha-Ruzyne, 21. v. 1957, 3 ~ (ZZ]. - Obi'istvi nr. Meln!k, 28. vii., 1 r:1 ( "Tachydromia flavipes Fb., det. Vimmer"; A. Vim mer, NMP]. Platypalpus laticinctus Walker, 1851 Platypalpus laticinctus Walker, 1851, Ins. Britt., 1: 127 Zoogeographie distribution: Great Britain ( Collin, 1961], Czeeho slovakia. Occurrence in Czechoslo·vakia: Mat: Mor: Strachotin nr. Bi'eclav, 6. v, 1908, 1 'i' ( K Landroc'k, MMB]. Platypalpu longicornis ( Meigen, 1822) Tachydromia Iangteornis Meigen, 1822, Syst. Beschr., 3: 73 Tachydromia pubicornts Zetterstedt, 1838, Ins. Lapp.: 553 A female Platypalpus longicornis was the prey of a small Nabid bug ( Laurenee, 1955). Prey: Gen. sp. Orthoeladiinae - Chironomidae, Ani sopus punctatus F. -- Anisopodidae, Seiara sp. - Sciaridae, Hilara sp. - Empididae ( Laurenee, 1955). Aeeording to Collin ( 1961) Platypalpus agilis preys on Platy;palpus longicornis. Zoogeographie distribution: Denmark, Finland, Spain, Italy (Lund beek, 1910) ; Belgium ( Goetghebuer, 1931) ; Austria, Yugoslavia, Sweden ( Enge•l, 1939] ; Great Britain ( Collin, 1961) ; Czeehoslovakia. Occurrence in Czechoslovakia: Mat: Boh: Zbüch nr. Plzeii, 21. viii. 1962, 1 'i' (ZZ).- Krkonose Mts., Koteine jamy, 13. viii. 1957, 1 ~ (ZZ]. Platypalpus luteus ( Meigen, 1804) Tachydromia lutea Meigen, 1804, Klassif., 1: 238 · Tachydromia glabra Meigen, 1822, Syst. Beschr., 3: 89 Tachydromia pallida Meigen, 1822, Syst. Beschr., 3: 90 ?Platypalpus formalis Walker, 1851, Ins. Brit., 1: 130 Zoogeographie distribution: Denmark, Finland, Italy ( Lundbeek, 1910); Sweden (Lundbeok, 1910; Ringdahl, 1941); Belgium (Goetghebuer, 357 1931]; Germany, Aus.tria (Engel, 1939); Great Britain (C ollin, 1961); Czechoslovakia. Occurrence in Czechoslovakia: Mat: Boh: Brdy Mts., Kytfn, vii. 1957, 1 ~; vii. 1959, 1 ~ (Macek, MMB]. Platypalpus luteipes, n. sp. Figs. 24, 31, 32. c:f'R.. Moderately large black dusted species with yellow legs. Head: Frons at antennal base as wide as third antennal joint deep, posteriorly before ocellar protuberance faintly, behind it rather strongly, widening, densely dusted greyish silvery. Face about half as wide, dusted similarly as frons. In anterior aspect clypeus shining, laterally slightly dusted greyish. Ocellar bristles about half as long as verticals; these represented by one pair of rather distant bristles. These and all other bristles and hairs of upper part of occiput brown, hairs on lower part langer, silverish white. Antennae entirely black, third joint about 29 31 Left and right part of epandrium: Fig. 29, Platypalpus flavicornis. - Fig. 30, P. in· fectus. - Fig. 31, P. luteipes. - Solid lines equal 0.1 mm each. 358 twice as long as deep at its base. Arista very long, up to three times as long as third antennal joint. Palpi small, orange yellow, dusted greyish, with two long silvery white bristles at their tips. Thorax: Mesonotum and scutellum black with golden yellow dust, pleurae dusted greyish, only sternopleura shining, black. Cbetotaxy: Acrestichals biserial, dorsocentrals in one row at each side, their last but one and especially last pairs considerably long; outside dorsocentrals several hairs. Notapleura possessing one very long bristle and three further, but much shorter, bristles situated close to its base. Humeral and postalar calli bearing two bri_stles each. Scutellum with one pair of long apical bristles and with another pair of shorter subapical brist les. All bristles and hairs of thorax palely yellowish. Legs yellow, with brownish black spots on trochanters, tarsal joints narrowly annulated brownish black, last joint dark except for basal third. Anterior pair: Coxae with silvery dust and greyish white pilosity anteriorly. Femora stoutened, with short pale yellow pilosity, bristles in posteroventral and anteroventral rows elongated, especially in male. Tibiae with short pale yellow pilosity, bristles in anterior row stronger, yellowish brown. Tarsal joint slender. - Middle pair: Coxae dusted greyish anteriorly and densely pilose yellow apically. Fernara stoutened, at least twice as deep as those of anterior pair, with short pale yellow pilosity, ventrally with double row of short sharp black spines; poste roventrally and anteroventrally with langer pale yellow b.ristles; tips of femora each with two small brownish black spots anteriorly and posteriorly. Tibiae with very short pale yellow pilosity, ventrally with row of sharp black spines, apically bearing long, strong and sharp, almost entirely black spur, this possessing two minute bristles on its tip. Tarsal joints slender, first joint ventrally w1th yellowish brown hairs being denser and Ionger than those on other tarsal joints. ----' Posterior pair: Coxae only moderately dusted greyish anteriorly, apically with several pale yellow hairs. Fernara slender, with short pale yellow pilo sity and each with two brownish black spots on their knees. Tibia8 slender, almost of same length as femora, covered with short pale yellow pilosity becoming denser towards apex and ventrally; apex of tibia possessing comb ventrally. Tarsal joints slender, first joint long, Ionger than half length of tibia. Wings tinged yellowish, veins palely yellowish brown. Prefurca as long as distance between its base and humeral vein; this very weak. r 5 and m1 parallel, only in apical section slightly converging. Cos.tal hairs palely yellowish brown. Squamae small, yellow, at margin with pale yellowish brown fringes. Halterae yellowish grey. Abdomen shining, black, with scattered pale yellowish brown pilo sity; only its first two segments distinctly dusted dark greyish on their bases; last sternum with long pale yellow hairs at its margin in male. Epandrium moderately large, left part of characteristic shape ( fig. 31], with long pale hairs. In female entire seventh segment dusted yellowish grey, eighth segment dusted greyish; cerci rather long. Length: body, 2.0 to 3.0 mm; wing, 2.8 to 3.2 mm. 359 Wing: Fig. 32, Platypalpus luteipes. - Fig. 33, P. major. - Fig. 34, P. minutus. - Solid lines equal 0.5 mm each. Type series: Holotype, male; Czechoslovakia: Praha-Ruzyne, 23. vi. 1964 (ZZ). - Paratypes: Praha-Ruzyne, 16. vi. 1962, 1 damaged speci men; 4. vi. 1963, 1 ; ~; 7. vi. 1963, 1 rJ; 13. vi. 1963, 5 rJ 3 - ~; 14. vi. 1963, 1 ~ ; 15. vi. 1963, 1 rJ 1 ~ ; 23. vi. 1964, 1 rJ 1 ~ ; 28. vi. 1964, 1 rJ ( ZZ; captured tagether with Platypalpus annulipes]. 360 Engel (1939] knew this species but called it "Coryneta flavipes Fabricius" ( see the note under Platypalpus pallidiV'entris]. Derivation o Platypalpus minutus ( Meigen, 1804) Tachydromia minuta Meigen, 1804, Klassif., 1: 238 Tachydromia exigua Meigen, 1822, Syst. Beschr., 3: 81 Tachydromia pygmaea Macquart, 1823, Mem. Soc. Sei. Lille: 154 Figs. 25, 34, 35. According to the literature, this species varies greatly in the colour of the legs, some specimens having them entirely black [including 361 tarsi), whilst in other individuals ( especially in females) the legs are rather pale. In the materials at my disposal both sexes have practically the same dark leg colour exc'ept for several females with paler legs, PreY': Alloxysta sp. - Cynipidae ( Lundbeck, 1910 J; Metriocnemus sp. - Chironomidae ( Laurence, 1955). Zoogeographie distribution: Denmark, Finland, Spain, Italy ( Lund beck, 1910); Austria, Yougoslavia, Germany (Engel, 1939); Belgium ( Goetghebuer, 1931); Sweden [ Ardö, 1957); Great Britain ( Collin, 1961); Czechoslovakia. Occurrence in Czechoslovakia: Mat: Boh: peat-mos s Soos nL Cheb, 19. v. 1961, 1 '? (ZZ] . - Cistä nr. Sokoiov, 15. viii. 1962, 1 Platypalpus niger ( Meigen, 1804) Tachydromia nigra Meigen, 1804, Klassif., 1: 238 Collin ( 1961) says that the continental individuals he has studied differ from the British in the possession of slightly dusted mesonotum. In Gzechoslovak specimens I found not only rather dusted mesonotum but also front slightly dusted brownish grey, in cantrast to the British flies I have investigated, which agree with Collin's description. Unfor tunately, I had no occasion to compare the hypopygia of both the spe · cies. I am of the opinion, however, that the British and Central Euro· pean specimens are conspecific. Zoogeographie distribution: Great Britain ( Collin, 1961), Czechoslo slovakia. Occurrence in Czechoslovakia: Mat. Boh: •Rove'llsko, vi., 1 6 1 '? ("Tachydromia minutus Mg.", A. Vimmer, NMP] . - Beta pod Bezdezem, vii., 1 6 ("Platypalpus femo ralis"; A. Vimmer , NMP). Mor: Vranov nad Dyji, vii. 1910, 1 6 3 '? (MMB ]. - Bflovice na·d Svitavou, 23. v. 1908, 1 '? (MMB]. - Cerno·vice nr. Brno, 15. vi., 1 '? (MMB].- Olomouc, 3. vii. 1958. 1 '? (P. Lauterer, coll. Zuska). Platypalpus nigritarsis ( Fall€m, 1816) Tachydromia nigrttarsis Fallen, 1816, Dipt. Suec., Emp.: 34 Platypalpus comptus Walker, 1837, Ent. Mag., 4: 228 Figs. 12, 36, 38, 42, 52. This species was found to be the prey of an Asilid fly (Zuskova, unpublished). 362 37 ' ' \ \ \ \ Left and right part of epandrium: Fig. 35, Platypalpus minutus. - Fig. 36, P. nigri· tarsis. - Fig. 37, P. niveiseta. - Solid lines equal 0.1 mm each. 3 363 Wing. Fig. 38, Platypalpus nigritarsis. - Fig. 39, P. pallipes. - Fig. 40, P. pectoralis. - Sohd lines equal 0.5 mm each. 364 Zoogeographie distribution: Denmark, Finland, Italy, Spain (Lund beek, 1910); Sweden (Lundbeek, 1910; Ardö, 1957); Norway (Ardö, 1957); Austria [Engel, 1939); Belgium (Goetghebuer, 1931); Great Britain {Col lin, 1961) ; Czeehoslovakia. Occurrence in Czechoslovakia: Mat: Boh: Cistä nr. Sokolov, 15. viii.- 1962,--2 ~ (J: D!abola, coll. Zuska. - Peat-moss Soos nr. Cheb, 19. v. 1961,2 <1 2 ~ (ZZ), 16. viii. 1962, 2 <1 4 ~ (J. Dlabola, coll. Zuska). - Frantiskovy Lazne, 2 <1 2 ~ [,,Tachydromia compta Wlk."; F. Kowarz, NMP).- Zbilch nr. -Plzeii, 21. viii. 1962, 2 'i! [ZZ).- Dobi'iv nr. Ro kycany, vii., 1 .<1 [" Tachydromia ciliaris Fll., det. Vimmer"; A . Vimmer, NMP ). - Brdy Mts., Kytin, viii. 1957, 1 Ir! 1 ~ [Macek, MMB ). - Markvarec nr. Louny, 2. vi. 1957, sweeping undergrowth in mixed wood, 1 '<1 : (ZZ) . - :Domousice nr. Louny, 12. ix. 1958, 1 'rf 1 <;! ( ZZ ). - Kundratice nr. Litomei'ice, 21. vi. 1958, 1 irf [ ZZ ). - Belä pod Bezde zem, 15. vii., 1 <1 (A. Vimmer, NMP). - Jilemnice, 12. vii. 1962, swept from Holcus sp., 1 ~ (J. Dlabola, coll. Zuska).- Ro·kytnice nad Jizerou, vii. 1954, 1 'i! (Macek, MMB). Krkono·se Mts., Rezek, 13. vi. 1957, sweeping in glade, 1 <1 (ZZ). - Krkono·se Mts., Vit ko·vice, 16. viii. 1957, from hay, 1 ~ [ZZ). Mor: Skorosice nr. Sumperk, 24. viii. 1907, 1 <1 (MMB ). - Skrbo·vice nr. Krnov. 12. viii. 1960, 1 rf ( ZZ ). - Brno, 17. vi., 1 - ~ ( MMB]. - Cernovice nr. Brno, 22. viii. 1919, 1 ~ (MMB) . Platypalpus niveiseta [ Zetterstedt, 1842) Tachydromia niveiseta Zetterstedt, 1842, Dipt. Scand., 1: 311 Figs. 37, 45. Zoogeographie distribution: Sweden [Frey, 1943), Great Britain [ Collin, 1961), Czechöslovakia. Occurrence in Czechoslovakia: Mat: Boh: Brdy Mts., Kytfn, viii. 1957, 1 <1 (Macek, MMB). Slov: .Krälovsky Chrmec nr. Trebisov, sweeping vegetation in wood of Robinia · pseudacacia, 10. vi. 1960, 5 <1 1 ~ ( ZZ ). Platypalpus notatus ( Meigen, 1822) Tachydromia notata Meigen, 1822, Sys. Beschr., 3_: 78 Zoogeographie distribution: Belgium (Goetghebuer, 1931), Great Bri tain ( Collin, 1961), Czeehoslovakia. Occurrence in Czecho-slovakia: Mat: Mor: Hlubo-cany nr. Vyskov, 6. vii. 1908, 1 ~ [K. Landrock, MMB).- Hno-jnik nr. Frydek-Mistek, 22. viii. . 1880, 2 <11 ~ (A. Hetsch ko, NMP) . Platypalpus optivus ( Collin, 1926) Tachydromia optiva Collin, 1926, Ent. man. Mag., 42: 157 Zoogeographie distribution: Germany [Engel, 1939), G.reat Britain ( Collin, 1961), Gzeehoslovakia. Occurrence in Czechoslo·vakia: Mat: Bo-h: Lo-denice nr. Beroun, viii. 1956, 1 <1 (Macek, MMB). Mor: Hlubocany nr. Vyskov, 6. vii. 1908, 1 ~ (K. Landrock, MMB). 365 Platypalpus pallidicornis ( Collin, 1926) Tachydromia pallidicornis Collin, 1926, Ent. man. Mag., 42: 186. Figs. 41, 46. Zoogeographie distribution: Great Britain ( Collin, 1961), Czecho slovakia. Occurrence in Czechoslovakia: Mat: Boh: Protivfn, 19. vii. 1902, 1 'i' (,,Platypalpus flauicornis Mg.", NMP]. - Pi'ednf Kopanina nr. Praha, 16. vi. 1957, sweeping vegetation araund dung heaps, 1 6 (ZZ]. Praha-Ruzyne, 27. v. 1959, 1 'i' ; 16. vi. to• 1. viii. 1962, 11 6 8 'i'; 14. vi. 1963, 2 'i' · ( ZZ] . - Horomei'ice nr. Praha, vi. 1964, 8 'i' (J. Beranl Platypalpus pallidiventris (Mengen, 1822) Tachydromia pallidiuentris Meigen, 1822, Syst. Beschr., 3: 82 Tachydromia dichroa Meigen, 1822, Syst. Beschr., 3: 83 Platypalpus robustus Walker, 1837, Ent. Mag., 4: 228 Tachydromia Iongiseta Zetterstedt, 1842, Dipt. Scand., 1: 278 ? Ernpis flauipes Fabricius, 1794, Ent. Syst., 4: 406 Figs. 44, 47. Collin ( 1961) designated one of the two Fabricius cotypes of Empis flavipes as the lectotype. As this specimen is badly damaged it is not possible to say if it belongs to Platypalpus pallidiuentris or Platypalpus extricatus, according to this author. Nevertheless, it is necessary to re strict the obscurities of the synonymy of Tachydromia flavipes to these two species only. Prey: Urophora solstitialis L. - Trypetidae ( Lundbeck, 1910); Con tarinia tritici Kby. - Cecidomyidae ( Barnes, 1932). Zoogeographie distribution: Denmark, Sweden ( Lundbeck, 1910; Ardö, 1957); Norway (Ardö, 1957) ; Finland, Italy, North Africa [Lund beck, 1910); Gre.at Britain ( Collin, 1961); Belgium ( Goetghebuer, 1931); Austria [Engel, 1939) ; Teneriffe ( Frey, 1931) ; Czechoslovakia. Occurrence in Czechoslnvakia: Mat: Boh: Peat-moss nr. Cheb, 16. viii. 1962, 1 'i' (F. Kodys, cnll. Zus:ka) . - Brdy Mts., Kytfn, vii. 1959, 1 6 1 'i' (Macek, NMPl. - Vräz nr. Beroun, viii. 1955, 1 'i' (Macek, MMB). - Zvol nr. Zbraslav, 17. vii. 1953, 3 'i' (ZZ]. Praha-Ruzyne, 27. v. 1959, 1 'i'; 4. vi. to 1. viii. 1962, 1 6 4 'i' (ZZ]. - Horomei'ice nr. Praha, vi. 1964, 3 'i' (J. Beränko·va, coH. Zuska]. - IKundratice nr. Litomerice, 21. vi. 1958, 1 'i' (ZZ). - Tisice nr. Melnfk, 29. v. 1964, 1 6 (F. Kodys, coll. Zuska]. - Novy Bydzov, vi., 2 'i' :("Tachydromia cursitans Fb., det. Vimmer"; A. Vimmer, NMP]. - Ji · lemnice, 15. vii. 1907, 1 6; 24. vii. 1907, 1 'i' ( NMP); 12. viii. 1962, swept from Holcu ~ sp., 1 'i' (J. Dlabola, coU. Zuska]. - Rokytnice nad Jizerou, vii. 1957, 1 'i' (Macek, MMB). - Krkonose Mts(. , Horn! Albei'ice, 23. viii. 1958, 1 6 ( P. Pecina, coll. Zuska]. Mor: PradM, 1 'i' (K. Czizek, MMB ). - Lublice nr. Opava, 7. vii. 1907, 1 6 (MMB].- Piiovice nr. Litovel, 9. vii. 1958, 3 "6 3 'i' (B. Lauterer, co'll. Zuska].- Troubk.y ~ Zai'icf nr. Kromei'iz, 3. vii. 1958, 1 'i' !(P. Lauterer, coll. Zuska]. - Olomnuc-Cernovir, 30. vi. 1958, 3 ·6 10 'i'; 1. vii. 1958, 4 · 366 Platypalpris pallipes (Fallen, 1815) Tachydromia pallipes Fallen, 1815, Dipt. Suec., Emp.: 8 Tachydromia flavipalpis Meigen, 1822, Syst. Beschr., 3: 74 Fig. 39. Prey: Chorebus discolor Marshai - Braconidae (Lundbeck, 1910). Zoogeographie distribution: Denmark, Sweden ( Lundbeck, 1910; Engel, 1939) ; Finland, France ( Lundbeck, 1910) ; Belgium ( Goetghebuer, 1931) ; Austria (Engel, 1939) ; Great Britain ( Collin, 1961) ; Czechoslo vakia. Occurrence in Czechoslovakia: Mat: B0oh: Brdy Mts., Kytln, viii. 1957, 1 !? [Macek, MMB). - Praha-Ruzyne, 114. vi. 1963, 1 !? '[ZZ). - Tisice nr. Melnlk, 29. v. 1964, 1 'i! (F. Kodys, coll. Zuska). - Jilemnice, vii., 1. t:1 ["Tachydromia fuscicornis Zt., det. Vim mer"; A. Vimmer, NMP). Mor: c'ernovice nr. Brno, 20 . v., 1 ,f [MMB) . - Kerhartice nr. Opava, 20. vii. 1908, 1 !? [ MMB) . - Lublice nr. Opava, 11. vii. 1907, 1 ,f [ MMB) . Platypalpus pectoralis (Fallen, 1815) Tachydromia pectoralis Fallen, 1815, Dipt. Suec., Emp.: 9 Tachydromia varipes. Meigen, 1822, Syst. Beschr., 3: 88 (teste Collin, 1961) Platypalpus pulchellus Walker, 1851, Ins. Brit., 1: 130 Tachydromia straminipes Zetterstedt, 1842, Dipt. Scand., 1: 296 Figs.. 40, 43. Zoogeographie distribution: Denmark, Finland, France ( Lundbeck, 1910); Sweden ( Lundbeck, 1910; Ringdahl, 1941); Belgium ( Goetghebuer, 1931); Austria, Germany (Engel, 1939); Great Britain ( Collin, 1961 ); Cze ehoslovakia. Occurrence in Czechoslovakia: Mat: Boh: Skryje nr. Rakovnlk, 4. x. 1959, 1 !? (K. Benes, CO'll. Zuska).- Brdy Mts., Kytln, viii. 1957, 2 6; vii. 1959, 1!? [Macek, MMB) . - Zatisl nr. •Pizen, 26. v. 1957, 1 ,f 1 !? [Macek, MMB ). - Nove Mesto nad Metujl, vii. 1957,1 !? [Macek, MMB).- Lyseciny nr. Marsov, viii. 1963, 1 t:1 [Macek, coll. Zuska}. Mor: Beskydy Mts.: Kycera, 10. vii. 1882, 1 !? [A. Hetschko, NMP) .- Milotice nr. Krnov, 9. viii. 1960, 1 !? [ZZ} . - Brusperk nr. Frydek-Mlstek, 1. vii. 1958, 1 t:1 [J. Be rankova, coll. Zuska }. Platypalpus pictitarsis ( Becker, 1902) Tachydromia pictitarsis Becker, 1902, Mitt. Zool. Mus. Berl., 2: 44 Platypal pus ruficornis Macquart, 1850, Dipt. exot. Suppl., 4: 401 [nec Tachydromia rufi cornis Roser, 1840} Zoogeographie distribution: Great Britain ( Collin, 1961), Czechoslo vakia. Occurrence in Czechos lovakia: Mat: Boh: ·Praha-Ruzyne, 12. vi. 1962, 1 !? ; 16. vi. 1962, 1 !? ; 12. vii. 1962, 1 ,f [ZZ}. - Praha-Holesovice, slaughterhouse, 27. viii. 1964, 1 .!? [J. Zuska, in coll. Central Re·search Institute of Food Industry, ·Praha). - Horomei'ice nr. Praha vi. 1964, 2 !? [J. Berankova, coll. Zuska). Mor: Hrusovany nad JeviSovkou, 22. v. 1964, 1 ,f (ZZ}. 367 Cerci and sclerite beneath those: Fig. 41, Platypalpus pallidicornis. - Fig. 42, P. ni· gritarsis. - Fig. 43, P. pectoralis. - Fig. 44, P. pallidiventris. - Fig. 45, P. niveiseta. - Solid lines equal 0.1 mm each. 368 49 52 Left and right part of epandrium: Fig. 46, Platypalpus pallidicornis. - Fig. 47, P. palli diventris. - Fig. 48, Anterior leg, P. bohemicus. - Antennae: Fig. 49, P. agilis. - Fig. 50, P. annulipes. - Fig. 51, P. interstinctus. - Fig. 52, P. nigritarsis. - Solid Iines equal 0.1 mm each. 369 Platypalpus rapidus ( Meigen, 1822) Tachydromia rapida Meigen, 1822, Syst. Beschr., 3: 81 Platypalpus mundus Walker, 1837, Entom. Mag., 4: 228 Zoogeographie distribution: Great Britain ( Collin, 1961), Czeehoslo vakia. Occurrence in Czechoslovakia: Mat: Mor: Bflovice nad Svitavou, 25. vi. 1909, 1 ~ [MMB) . Platypalpus ruficornis ( Roser, 1840) Tachydromia ruficornis Roser, 1840, Württemb. Corrbl., 1:, 54 Tachydromia thoracica Lundbeck, 1910, Dipt. Dan., 3: 317 Fig. 13. Zoogeographie distribution: Germany, Denmark (Lundbeek, 1910); Great Britain ( Collin, 1961) ; Czeehoslovakia. Occurrence in Czecho'Slovakia: Mat: Boh: ·Praha-Ruzyne, 1. viii. 1962, 2 · ~ [ZZ]. Platypalpus stigmatellus ( Zetterstedt, 1842) Tachydromia stigmatella Zetterstedt, 1842, Dipt. Scand., 1: 306 Tachydromia lateralis Becker, 1887, Berl. ent. Ztschr., 31: 137 Tachydromia beckeri Mik, 1894, Wien. ent. Zeitg., 13: 166 Zoogeographie distribution: Austria, Czeehoslovakia (Engel, 1939); Seotland ( Collin, 1961) . Occurrence in Czechoslovakia: Lit: Boh: Zelezna Ruda (Engel, 1939). Mat: Boh: Krkonose Mts., Ko1elne jamy, 10. vii. 1922, 1 ~ [,,Tachydromia pallipes Fll., det. Vim mer"; A. Vimmer, NMP]; 13. viii. 1957, 1 ~ [ZZ). Mor: PradM, 1 ·rJ (K. Czizek, MMB) . Platypalpus verraU ( Collin, 1926) Tachydromia verrali Collin, 1926, Ent. mon. Mag., 42: 185 Zooge!Qgraphie distribution: Great Britain (Engel, 1939, Collin 1961), Czeehoslovakia. Occurrence ·in Czechoslovakia: Mat: Boh: As, 30. v. 1878, 1 r!; 8. vi. 1878, 2 ~ [F. Kowarz, NMP]. - Vrane nad Vltavou, 9. v. 1959, 1 ~ [Macek, MMB]. - Horusice nr. Tabm, 27. v. 1964, swept from vegetation on pond bank, 1 ~ (ZZ). Slov: Sulava nr. Nitra, 6. ix. 1953, 1 'r! [Macek, MMB ]. 4. SEX-RATIO OF PLATYPALPUS SPECIES IN CZECHOSLOVAKIA It is a well-known faet that some speeies of the genus Platypalpus show the unbalaneed sex-ratio, whieh is apperently influeneed by their geographie parthenogenesis. Below, I give a reviev of the numbers of males and females of the partieular speeies, whieh were eaptured in Czeehoslovakia and examined by me, so as to eontribute to our know ledge of this phenomenon. 370 I I I Number of examined individuals I No. Species Sex-ratio I Males I ·Females 1. P. agilellus - - - 2. P. agilis 7 26 I 0.3 3. P. albicornis 1 2 0.5 4. P. albiseta 1 6 0.2 5. P. annulatus 11 4 2.8 6. P. annulipes 34 15 2.3 7. P. articulatus 5 11 0.5 8. P. binderi 6 7 0.8 I 9. P. bohemicus - 2 only ~ I 10. P. calcaratus 1 - only <1 11. P. calceatus 1 5 0.2 12. T. cand".cans 1 50 0.02 13. P. ciliaris 7 10 0.7 14. P. cursitans - 9 only ~ 15. P. extricatus 7 32 2.2 I I 16. P. fasciatus 11 13 0.8 17. P. flauicornis 2 4 0.5 18. P. infectus 4 8 0.5 19. P. infenuus 1 1 1.0 20. P. interstinctus 2 5 0.4 21. P. laticinctus - 1 only ~ 22. P. longicornis - 2 only ~ 23. P. luteus - 2 only ~ 24 . T. luteipes 10 7 1.4 25. P. major - 36 ? 26. P. minutus 20 64 0.3 27. T. niger 3 7 0.4 28. P. nigritarsis 14 20 0.7 29 . P. niueiseta 6 1 6.0 30. P. notatus 2 2 1.0 31. T. optiuus 1 1 1.0 32. P. pallidicornis 15 25 0.6 33 . . P. pallidiuentris 13 45 0.3 34. P. pallipes 3 4 :0.8 3~. P. pectoralis 5 6 .0.8 36. T. pictitars!s 2 5 0.4 37. P. rapidus - 1 only ~ 38. P. ruficornis - 2 only ~ ~ "lgmatellu' 1 2 0.5 uerrali 2 4 0.5 Acknowledgements I am very obliged to Dr. J. Moucha (.Praha-Kunratice) and to Dr. J. Steh!Ik (Brnoj for making it possible for me to study the specimens deposited in the Entomology Departmens of the Nati-onal Museum in ·Praha and Moravian Museum in Brno. Dr. L. Lyneborg (Copenhagen) and Dr. K. G. V. Smith (London) kindly lent me some spe cimens from the collections und er their charge. Mr. A. Wootton ( Stone, near Aylesbury, England) was kind enough to check the English of the paper. Dr. C. W. Sabrosky (Washington) sent me a very detailed explanation of the compHcated generic synonymy of Platypalpus. 4 371 References Ar d ö, P., 1957: Studies in the marine shore dune ecosystem with special referen;; c' to the dipterous fauna. Opusc. entom., Suppl. 14: 1-255. B a r n es, H. F., 1932: Studies of fluctuations in insect populations. I. The infestation of broad balk wheat by the wheat blossom midges (Cecidomyidae]. J. anim. Ecol., 1: 12-31. . B r an c s i k, K., 1910: Trencsemvarmegyeben talalt Dipterak felsowlasa. Jahresh. Na turw. 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